taxonID	type	description	language	source
2A00879F6B24583BFF49FCEAFD66E91E.taxon	description	(Figs 1 a – l)	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B24583BFF49FCEAFD66E91E.taxon	type_taxon	Type species: Sulcolotis seminigra Miyatake, 1994, by monotypy.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B24583BFF49FCEAFD66E91E.taxon	diagnosis	Diagnosis. This genus can be easily distinguished from all other genera of the tribe Sticholotidini by its sulcate genae, narrow intercoxal process of prosternum and short maxillary palpi.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B24583BFF49FCEAFD66E91E.taxon	description	Description. Body broadly oval, moderately to strongly convex; dorsal surface apparently glabrous, head, lateral margins of pronotum and elytra and basal areas of elytra indistinctly pubescent (Figs 2 a – c). Head relatively broad and large (Fig. 2 c) with frons flat and punctate (Fig. 1 a). Clypeus short and transverse with anterior margin emarginate medially (Fig. 1 b). Eyes relatively large, finely faceted, slightly emarginate in front of antennal socket, inner orbital striae fine and inconspicuous (Fig. 2 c). Antennae moderately long, composed of 11 antennomeres, with a narrowly dilated 3 antennomere club (Figs 1 a, c). Labrum transverse, rounded anteriorly and covered with long dense setae (Fig. 1 d). Mandible subtriangular with two apical teeth, sharp and smooth; mola well developed without distinct molar teeth (Figs 1 e, f). Maxillary palp with four palpomeres, palpomere 1 small, palpomere 2 geniculate and at least two times as long as wide, palpomere 3 obviously short, terminal one slender, sub parallel sided, with obliquely truncate apex (Fig. 1 g). Labial palp with three palpomeres, palpomere 1 tiny, palpomere 2 swollen, terminal palpomere slender, strongly conical, pointed apically; ligula membranous; insertion of labial palps visible ventrally on prementum; mentum trapezoidal, distinctly narrowed basally (Fig. 1 h). Pronotum moderately convex, broadly and gently emarginate anteriorly; anterior portion laminate and transparent; lateral margin narrowly reflexed; posterior corners subrectangular; base somewhat sinuously produced posteriorly, basal margin with distinct and complete carina (Figs 2 a – c). Scutellar shield triangular (Figs 2 a, c). Elytra with prominent humeral angles, convex, anterior margin almost equal in width to hind margin of pronotum, lateral margins rounded, broadly explanate (Figs 2 a – c). Elytral epipleurae moderately wide at basal half, slightly inclined and gradually narrowing to apex. Prosternum T – shaped, with inconspicuous punctures; prosternal process with carinae, narrow, less than half width of procoxal cavity diameter; hypomeron broad without foveae (Fig. 1 i). Mesoventral process broad, as wide as mesocoxal diameter, with golden pubescence and inconspicuous punctures; metaventral postcoxal lines recurved, complete laterally, joined on metaventral process and forming a straight line (Fig. 1 j). Abdomen with six ventrites, ventrite 1 moderately long, around 3 x as long as ventrite 2, with postcoxal lines incomplete laterally, not reaching posterior margin of ventrite 1 but running parallel in short distance to it; ventrites 2 – 4 successively shorter, ventrite 5 as long as the preceding two combined, its posterior margin broadly rounded in female (Fig. 2 g). Female genitalia: coxites elongate (Fig. 1 l), subtriangular, without stylus. Spermatheca C-shape, sclerotised (Fig. 1 k).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B24583BFF49FCEAFD66E91E.taxon	distribution	Distribution. Philippines (Mindanao, Luzon).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	description	(Figs 1 a – l)	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	diagnosis	Diagnosis. This species can be easily separated from S. seminigra by the following characters: elytron yellow, with a large black spot situated at 1 / 3 elytral length to base. In S. seminigra, elytra are black in basal half and yellow in apical half, without dark spots. It is also similar to S. ovalis but can be separated from the latter by the bright yellow head and penis guide nearly as long as parameres. In S. ovalis, head is reddish brown and penis guide shorter than parameres.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	description	Description. TL: 2.73 – 3.07 mm, TW: 2.30 – 2.52 mm, TH: 1.51 – 1.56 mm, TL / TW: 1.19 – 1.22, PL / PW: 0.26 – 0.29, EL / EW: 0.88 – 0.89, HW / PW: 0.56 – 0.63, PW / TW: 0.63 – 0.66, HW / TW: 0.38 – 0.40, Eye W / HW: 0.54 – 0.59. Head yellow. Pronotum, scutellar shield, and elytra bright yellow, with small dense punctures. Underside yellowish brown, except metaventrite and the middle part of the first ventrite blackish brown. Body rounded, weakly convex, dosally shiny and glabrous (Figs 2 a – c). Head 0.39 times of elytral width (HW / TW = 1: 2.6). Eyes finely faceted interocular distance 0.57 times of head width. Frons broad with punctures uniform and dense (Fig. 2 c). Pronotum 0.65 times of elytral width (PW / TW = 1: 1.5), moderately transverse, punctures uniform (Figs 2 a, c). Elytra with uniform and dense punctures (Figs 2 a – c). Ventrite 6 of posterior margin at middle slightly concave and ventrite 7 subtrianglar (Fig. 2 g). Male genitalia (Figs 2 d – f): penis long and slender, apex pointed, penis capsule with inner arm reduced and outer arm well developed; tegmen with penis guide as long as parameres, in lateral view stout, widest at base, gradually narrowing to a pointed apex.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	materials_examined	Type material. Holotype: ♂, Philippines, Baguio, Benguet, Baker (USNM). Paratypes: 6 ♀, same data as holotype (USNM, ANIC); 1 ♂ 4 ♀, Philippines, Luzon – Benguet LaTrinidadv. 1914, G. Boellcher 6500 ’ (MNHB); 1 ♀, Philippines 1600 m, Mindanao, 30 km W of MARAMAG, 28 – 30. Dec. Bolm 1 gt. 1990 (BNHM).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	distribution	Distribution. Philippines (Mindanao, Luzon).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B27583AFF49FA52FACAEA1A.taxon	etymology	Etymology. The specific name threadis is adopted as meaning thread, referring to its thin and long penis.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	description	(Figs 3 a – g)	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	diagnosis	Diagnosis. This species is most similar to S. threadis sp. nov. in external appearance but can be distinguished from the latter by the penis guide shorter than parameres. In S. threadis sp. nov., the penis guide is nearly as long as parameres in lateral view.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	description	Description. TL: 2.63 – 2.86 mm, TW: 2.26 – 2.37 mm, TH: 1.37 – 1.59 mm, TL / TW: 1.16 – 1.21, PL / PW: 0.28 – 0.30, EL / EW: 0.94 – 0.99, HW / PW: 0.56 – 0.58, PW / TW: 0.63 – 0.64, HW / TW: 0.35 – 0.37, Eye W / HW: 0.43 – 0.46. Head reddish brown. Elytra reddish brown with a long ovate black spot situated at 1 / 3 elytral length to base. Underside yellowish brown, except metaventrite and the middle part of the first abdominal ventrite reddish brown. Body rounded, weakly convex, shiny and glabrous. Eyes finely faceted, interocular distance 0.45 times width of head. Frons broad with irregular transparent spots, punctures uniform and dense (Fig. 3 c). Pronotum 0.64 times of elytral width (PW / TW = 1: 1.6), moderately transverse, punctures uniform and dense (Fig. 3 c). Elytra with uniform and dense punctures (Figs 3 a – c). Ventrite 6 subhemisphenical and ventrite 7 nearly straight (Fig. 3 g). Male genitalia: penis relatively short and slender, apex pointed, penis capsule with nearly same length of inner arm and outer arm. Tegmen with penis guide in lateral view slender, widest at base, narrowing to apex, distinctly shorter than parameres.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	materials_examined	Type material. Holotype: ♂, Philippines, Butuan Mindanao, Baker (USNM). Paratypes: 2 ♀♀, same data as holotype (USNM).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	distribution	Distribution. Philippines (Mindanao).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B26583DFF49F92BFF65EAAA.taxon	etymology	Etymology. The specific epithet is formed from the Latin noun ovalis, referring to elytra with a long ovate black spot.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	description	(Figs 4 a – g)	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	diagnosis	Diagnosis: This species can be distinguished from S. seminigra Miyatake and S. ovalis sp. nov by the nearly entirely black elytron.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	description	Description. TL: 2.31 – 2.79 mm, TW: 1.75 – 2.18 mm, TH: 1.25 – 1.82 mm, TL / TW: 1.28 – 1.32; PL / PW: 0.23 – 0.24; EL / EW: 1.05 – 1.10; HW / PW: 0.71 – 0.73; PW / TW: 0.66 – 0.67; HW / TW: 0.47 – 0.48; Eye W / HW: 0.40 – 0.45. Head and pronotum yellowish brown. Scutellar shield, and elytra blackish brown with yellowish brown margin. Underside yellowish brown; except metaventrite and the middle part of ventrite 1 reddish brown. Body rounded, strongly convex (Figs 4 b, c). Head 0.48 times of elytral width (HW / TW = 1: 2.1), with sparse pubescence. Eyes finely faceted, interocular distance 0.43 times of head width. Frons broad, punctures uniform and dense (Figs 4 c, d). Pronotum 0.67 times of elytral width (PW / TW = 1: 1.5), moderately transverse, punctures uniform (Figs 4 a, c). Elytra with uniform and dense punctures (Figs 4 a – c). Ventrite 6 subhemisphenical and posterior margin of ventrite 7 at middle strongly concave (Fig. 4 g). Male genitalia: Penis relatively short, pointed and apically curved, capsule with nearly same length of inner and outer arm; tegmen with penis guide distinctly shorter than parameres, in lateral view widest, then gradually tapering to apex; penis guide in ventral view gradually narrowing to apex.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	materials_examined	Type material. Holotype: ♂, Philippines, Mt. Makiling, Luzon, Baker (USNM). Paratype: 2 ♂ 2 ♀, same data as holotype (USNM); 1 ♀, Philippines Malinao Tayabas Baker (USNM); 1 ♀, Philippines, S Luzon Quezon N. P (Lucena) 8 – 10. Jan. 1991 Bolm lgt., 250 m (ANIC).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	distribution	Distribution. Philippines (Luzon).	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
2A00879F6B21583CFF49F8CFFE56EB3A.taxon	etymology	Etymology. The specific epithet is formed from the Greek noun Xanthus and Latin noun marginatus, referring to elytra with yellowish margin.	en	Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng, Wang, Xingmin (2022): Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines. Zootaxa 5168 (1): 75-82, DOI: https://doi.org/10.11646/zootaxa.5168.1.6
