taxonID	type	description	language	source
03B087BDFF14FFACB5F3FDD1C0C2F9A5.taxon	type_taxon	Type species: Ceratostigma gracilis Li, 2002: 1595, by original designation.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFACB5F3FDD1C0C2F9A5.taxon	diagnosis	Diagnosis. Forewing with veins on basal two-thirds yellowish or grayish white; Sc usually ending free; Rs and M meeting in a point or fused for a short distance; CuA 1 - 2 and M + CuA 1 nearly in a line. Male genitalia: Hypandrium of asymmetrical, simple plate, medially with a ridge. Female genitalia: Epiproct subtriangular. Paraproct subtriangular. Subgenital plate with a short and distally rounded egg guide; pigmented area slightly sclerotized, roughly T-shaped, stem of T-shaped marking more than two times as long as basal width. Dorsal valve of gonapophyses broad, pointed at tip, inner margin with a strongly sclerotized area at base; external valve with a triangular posterior lobe.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFACB5F3FDD1C0C2F9A5.taxon	distribution	Distribution. China, Indonesia.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFACB5F3FDD1C0C2F9A5.taxon	discussion	Key to species of Ceratostigma	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFAFB5F3F9EDC657FE53.taxon	description	(Figs 1 – 15)	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFAFB5F3F9EDC657FE53.taxon	diagnosis	Diagnosis. Male. Forewing (Fig. 1) basal one-fifth with a slightly curved greyish white stripe; radial fork with branches diverging at an angle about 90 °. Epiproct (Figs 3 – 4) nearly triangular, pointed at apex, with a nearly Ushaped sclerotized area at base; hypandrium (Figs 5 – 6) with a broad ridge at middle. Female. Forewing (Fig. 8) with a grayish white striped marking at one-fifth base; radial fork with branches diverging at an angle about 90 °. Pigmented area of subgenital plate (Figs 14 – 15) roughly T-shaped, distally flat; dorsal valve (Figs 12 – 13) anterior one-third with a slightly sclerotized area.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFAFB5F3F9EDC657FE53.taxon	distribution	Distribution. China (Guangxi).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFAFB5F3F9EDC657FE53.taxon	materials_examined	Material examined. Holotype male, CHINA: Guangxi, Longsheng, Tianpingshan [25 ° 39 ’ N, 109 ° 55 ’ E], 680 m, 12. VII. 1982, Fasheng Li (CAU); Paratype: 1 female, same as holotype; 1 male, 1 female, CHINA: Guangxi, Jiuxiu, Dayao Mountain [23 ° 58 ’ N, 110 ° 07 ’ E], 720 m, 23. VI. 1982, Fasheng Li (CAU).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF14FFAFB5F3F9EDC657FE53.taxon	discussion	Remarks. Ceratostigma gracilis is similar to C. macrostigmatum in male forewing basal one-fifth with a slightly curved greyish white stripe; radial fork with branches diverging at an angle less than 90 °; epiproct nearly triangular, pointed at apex, with a nearly U-shaped sclerotized area at base; hypandrium with a broad ridge in the medium. C. gracilis can be distinguished from C. macrostigmatum by the hypandrium rounded apically, while the hypandrium of C. macrostigmatum is narrowed distally.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF16FFA9B5F3FAEBC164FE0B.taxon	diagnosis	Diagnosis. Male. Forewing radial fork with branches diverging at an angle of about 30 °. Epiproct rounded at apex; hypandrium with a narrowed ridge medially. Female. Forewing radial fork with branches diverging at an angle of less than 90 °; cells r 3 and r 5 almost hyaline; basal area with irregular hyaline areas. Pigmented area of subgenital plate roughly T-shaped, nearly pointed at apex; dorsal valve anterior one-third with a small slightly sclerotized area.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF16FFA9B5F3FAEBC164FE0B.taxon	distribution	Distribution. Indonesia (Bali, Flores, Java).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF16FFA9B5F3FAEBC164FE0B.taxon	discussion	Remarks. This species was originally placed in Metylophorus Pearman, 1932 (Metylophorini Roesler, 1943) because of its similar color of wings and venations with most species of Metylophorus. Metylophorus can be diagnosed by having the forewing veins entirely brown; hypandrium broad, divided into three lobes, median lobe strongly sclerotized, usually with strongly sclerotized projections; subgenital plate with a long egg guide; dorsal valve of gonapophyses with a round apex. The type species of Metylophorus, M. nebulosus (Stephens, 1836), can be separated from C. lisae by male hypandrium asymmetrical, divided into three lobes, median lobe with two left processes well developed and two small right processes close to the base; female egg guide of subgenital plate elongate (Endang et al. 2002; Li 2002; Kido & Yoshizawa 2019; Pearman 1932; Thornton 1984). C. lisae can be distinguished from M. nebulosus by having the forewing with veins on the basal two-thirds yellowish or grayish white; male hypandrium a simple plate, medially with a ridge; female subgenital plate with a short egg guide; dorsal valve of gonapophyses broad, pointed at tip. Thus, it seems reasonable to transfer this species to Ceratostigma, and the new combination proposed above. Prior to this study, all the species of Ceratostigma were only known from China, while the new combination represents the first record of this genus from Indonesia. Ceratostigma lisae can be distinguished from congeners by having the male forewing radial fork with branches diverging at an angle of about 30 °, epiproct with a rounded apex, hypandrium medially with a narrowed ridge, female forewing radial fork with branches diverging at an angle less than 90 ° and cell r 3 and r 5 almost hyaline.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA8B5F3FF01C7FDFD0A.taxon	description	(Figs 16 – 22)	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA8B5F3FF01C7FDFD0A.taxon	diagnosis	Diagnosis. Forewing (Fig. 16) basal one-fifth with a slightly curved greyish white stripe; radial fork with branches diverging at an angle of about 60 °. Epiproct (Figs 18 – 19) nearly triangular, pointed at apex, with a nearly U-shaped sclerotized area at base; hypandrium (Figs 21 – 22) narrowed at apex, with a broad ridge in median area.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA8B5F3FF01C7FDFD0A.taxon	distribution	Distribution. China (Fujian).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA8B5F3FF01C7FDFD0A.taxon	materials_examined	Material examined. Holotype male, CHINA: Fujian, Chong’an, Guidong [27 ° 41 ’ N, 117 ° 58 ’ E], 1. VII. 1979, Chikun Yang (CAU).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA8B5F3FF01C7FDFD0A.taxon	discussion	Remarks. This species is known only from the male. It can be distinguished from congeners by the hypandrium narrowed at apex, medially with a broad ridge.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	description	(Figs 23 – 44)	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	diagnosis	Diagnosis. Male. Forewing basal one-fifth with a slightly curved grayish white stripe; radial fork with branches diverging at an angle of about 90 °. Epiproct pointed at tip, with a nearly U-shaped sclerotized area at base; hypandrium distally with a nearly hat-shaped sclerotized area. Female. Forewing basal one-fifth with a nearly V-shaped grayish white marking. Pigmented area of subgenital plate roughly T-shaped, distally nearly drop-shaped, pointed at tip; dorsal valve of gonapophyses ventro-distally with a slightly sclerotized area.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	description	Description. Male. Coloration (in 95 % ethanol). Head (Figs 25 – 27) pale brown; vertex with two grayish white markings; epicranial suture and inner margin of compound eyes dark brown; postclypeus pale brown with brown stripes; anteclypeus pale brown. Ocelli black, an irregularly shaped brown marking between median ocellus and anteclypeus. Compound eyes black. Antennae dark brown, except scape, pedicel, and first and second flagellomere pale yellow. Maxillary palpus grayish brown, distally brown. Thorax (Figs 25 – 27) dark brown, median area, scutellum of mesothorax and median area of postnotum pale brown. Legs brown. Forewing (Fig. 23) brown, veins with proximal two-thirds yellow and distal one-third brown; a slightly curved grayish white stripe at proximal one-fifth; pterostigma reddish brown. Hind wing (Fig. 24) brown. Morphology. Head (Figs 25 – 27) inverted trapezoidal; compound eyes ovoid. Forewing (Fig. 23) membranous, Sc ending free; Rs and M meet in a point, or fused for a short distance; radial fork with branches diverging at an angle of about 90 °; pterostigma subtriangular with apex rounded; discoidal cell nearly rectangular; CuA 1 - 2 and M + CuA 1 nearly in a line. Genitalia (Figs 28 – 33): Epiproct (Figs 28 – 29) nearly semicircular, pointed at apex, with a nearly U-shaped sclerotized area at base, medially with setae. Paraproct distally with a spinous projection slightly curved at apex; sensory field with about 46 trichobothria. Hypandrium (Figs 30 – 31) of asymmetrical, simple plate, broad median ridge laterally with several creases; a dumbbell-like sclerotized area present at base. Phallosome (Figs 32 – 33) simple, ring-like. Measurements. B: 3.31 mm, BW: 4.99 mm. IO: 0.53 mm, d: 0.19 mm, IO / d: 2.9. f 1: 1.36 mm. FWL: 4.32 mm, FWW: 1.75 mm, HWL: 3.23 mm, HWW: 1.28 mm. t 1: 0.47 mm, t 2: 0.22 mm. Female. Coloration (in 95 % ethanol) (Figs 34 – 38). Same as in male, but marking between median ocellus and anteclypeus paler. Forewing (Fig. 34) separated with numerous irregular pale brown markings and a V-shaped pale brown marking at proximal one-fifth. Morphology. Adult: Generally similar to male except for smaller compound eyes. Genitalia (Figs 39 – 44): Epiproct (Figs 39 – 40) subtriangular, with a nearly H-shaped sclerotized area, distally rounded, antero-medially with setae. Paraproct (Figs 39 – 40) triangular, sensory field with about 38 trichobothria. Subgenital plate (Figs 43 – 44) with short and distally rounded egg guide, pigmented area slightly sclerotized, roughly T-shaped, apex nearly drop-shaped, pointed at tip, transverse region of pigmented area nearly dumbbelllike with numerous setae. Gonapophyses (Figs 41 – 42): Ventral valve slender, apex narrowed ventrally with short setae; dorsal valve broad, pointed at tip, with strongly sclerotized area at inner margin, ventro-distally with a slightly sclerotized area; external valve with a triangular posterior lobe. Measurements. B: 4.16 mm, BW: 6.32 mm. IO: 0.84 mm, d: 0.17 mm, IO / d: 4.94. f 1: 1.32 mm; f 2: 1.32 mm; f 3: 0.50 mm. FWL: 5.36 mm, FWW: 2.21 mm, HWL: 2.26 mm, HWW: 1.37 mm.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	materials_examined	Material examined. Holotype female, CHINA: Xizang, Zayu County, Upper Zayu, Cibacun [28 ° 63 ’ N, 97 ° 41 ’ E], 2265 m, 17. VI. 2019, Rongrong Shen & Yingnan He (CAU). Paratypes: 2 males, 1 female, same as holotype (CAU).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	distribution	Distribution. China (Xizang).	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	etymology	Etymology. The epithet “ stagona ” refers to the drop-shaped apex of the pigmented area of the female subgenital plate.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
03B087BDFF10FFA5B5F3FD5BC495FE2E.taxon	discussion	Remarks. The new species appears to be related to C. gracilis and C. macrostigmatum based on similarity of the following characters: male forewing basal one-fifth with a slightly curved grayish white stripe; epiproct pointed at apex with a nearly U-shaped sclerotized area at base; hypandrium medially with a broad ridge; female forewing radial fork with branches diverging at an angle of about 90 °; pigmented area of subgenital plate roughly T-shaped. The new species can be distinguished from them by the male forewing radial fork with branches diverging at an angle of more than 90 °, the hypandrium distally with a nearly hat-shaped sclerotized area, the female forewing basal one-fifth with a nearly V-shaped grayish white marking, and the pigmented area of the subgenital distally nearly drop-shaped, pointed at tip.	en	Jie, Lulan, Liang, Feiyang, Liu, Xingyue (2022): Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China. Zootaxa 5168 (2): 196-206, DOI: https://doi.org/10.11646/zootaxa.5168.2.6
