identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E04B4F00FFE2FFC3FF4FF9C77F4F576A.text	E04B4F00FFE2FFC3FF4FF9C77F4F576A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleolissus perlucens (Bates 1878)	<div><p>Coleolissus perlucens (Bates, 1878)</p> <p>(Figs. 1–8)</p> <p>Hypolithus perlucens Bates, 1878: 715.</p> <p>Type material. Holotype: female, “Jhelum Vall., Cashmere”, “ Hypolithus perlucens Bates ” [Bates’ handwriting], “ Ex Musaeo H.W. Bates, 1892” (MNHN).</p> <p>Additional material examined. Pakistan. 1 male, 2 females, Islamabad, The National Forest Park, 12.VII.2003, S. Ovchinnikov leg. (ZIN); 1 female, Kaghan Valley, Balakot, ~ 1000 m, 21/ 24.VII.2003, W. Heinz leg. (SMNS). India. Uttarakhand: 1 female, India bor., Uttar Pradesh bor., env. Karnaprayas, 20.VII.1994 (cFCCH).</p> <p>Re-description (1 male and 5 females measured). Body medium-sized for the genus, length 8.6–9.1 mm, width 3.7–3.9 mm. Habitus as in Figs. 1, 2.</p> <p>Body dark brown to black, shiny and iridescent, with labrum externally, mandibles basally and very narrow lateral margins of pronotum reddish brown; abdominal sternites reddish black; palps, antennae and legs yellowish brown; femora slightly paler than tibiae and tarsi; coxae blackish brown or blackish yellow.</p> <p>Head medium-sized (HWmax/PWmax 0.61–0.64, HWmin/PWmax 0.50–0.52), impunctate. Eyes large, moderately convex (HWmax/HWmin 1.20–1.27), almost touching ventrally margins of buccal fissure. Genae glabrous. Tempora moderately long, slightly convex, somewhat steeply sloping to neck. Labrum flat, with almost straight anterior margin. Clypeus very slightly convex, very shallowly emarginated along anterior margin, with one setigerous pore at each outer angle and with very fine and short longitudinal wrinkles adjacent medially to the pore. Fronto-clypeal suture very fine, superficial, almost straight. Frontal foveae oval, more or less deepened, without prolongation on clypeus. Fronto-ocular furrows fine and short, not reaching supraorbital furrows. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by distance less than width of antennomere 2 basally. Mentum separated from submentum by complete transverse suture, with a prominent, rather wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite moderately widened distally, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, narrow, markedly projecting beyond ligular sclerite and separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere as long as apical one. Mandibles elongate, somewhat narrow, evenly curved apically, with very fine oblique wrinkles in apical half dorsally; left mandible acute, at most only slightly blunted at tip. Dorsal microsculpture distinct at anterior margin of clypeus, obliterated on remaining surface, consisting of mixture of isodiametric and slightly transverse meshes. Antennae slender, surpassing pronotal basal edge by approximately one and a half apical antennomeres (in female slightly shorter than in male), pubescent from middle of antennomere 3, with antennomeres 4–8 about 2.5 times as long as wide and basal antennomere about as long as antennomere 3.</p> <p>Pronotum wider than long (PWmax/PL 1.38–1.45), widest just before the middle, slightly more strongly narrowed apically than basally (PWmax/PWmin-ap 1.49–1.53, PWmax/PWmin-bas 1.24–1.32), with one lateral seta inserted slightly before widest point. Sides evenly rounded along entire length; lateral bead complete, very narrow throughout. Apical margin markedly emarginated, almost straight medially, very narrowly bordered along entire length. Apical angles protruding anteriorly, narrowly rounded at apex. Basal margin almost straight or slightly concave in middle portion, slightly rounded laterally, bordered along entire length, slightly longer than apical margin (PWmin-bas/PWmin-ap 1.14–1.21) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions beginning from apical angles as grooves; these depressions markedly widened from lateral setae and fused basally with basal foveae forming deep oblique laterobasal depressions isolated from each other by a convex area. Median line fine, superficial, not reaching apical margin and basal margins. Anterior transverse depression very shallow or indistinct. Disc densely punctate, more coarsely along margins and very finely in middle portion, with coarsest and confluent punctures in laterobasal depressions. Microsculpture not recognized under 56x magnification.</p> <p>Elytra convex, oval (EL/EW 1.48–1.50, EL/PL 2.62–2.71, EW/PWmax 1.22–1.29), widest before the apical third; sides rounded apically and basally, almost straightly diverging before the middle; subapical sinuation very shallow, almost indistinct. Humerus angularly rounded, smooth or with indistinct denticle at apex recognizable from behind. Sutural angle slightly blunted at tip, not extended posteriorly. Basal edge markedly arched, forming an obtuse angle with lateral margin. Striae with granulate microsculpture on bottom, impressed along entire length, reaching anteriorly basal elytral edge and widened apically. Intervals glabrous, strongly narrowed apically, slightly convex on disc and rather strongly convex before apex; two or three lateral intervals finely punctate, more distinctly in apical half. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 7–10 small discal setigerous pores adjoining stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series without distinct gap at middle, consisting of 20–23 setigerous pores. Lateral groove narrow and flat, slightly widened apically, with granulate meshes. Microsculpture on intervals highly obliterated, at most with indistinct transverse lines. Wings fully developed.</p> <p>Pro- and metasternum with very fine and short setae. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</p> <p>Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side. Metafemur ventrally with two long setae at posterior margin and with several very short setae at anterior margin. Protibia with longitudinal sulcus on dorsal side, with one preapical spine on outer margin in male and with three such spines in females. Tarsi almost glabrous dorsally, with a few very short and fine, faintly visible setae on male pro- and mesotarsi. In male, protarsomeres 1–4 and mesotarsomeres 2–4 markedly widened, with biseriate adhesive scales ventrally; mesotarsomere 1 weakly widened, with scales only apically, shorter (in female longer) than mesotarsomeres 2 and 3 combined. Metatarsus slender, markedly longer than HWmax, with metatarsomeres 2–4 slightly widened distally; metatarsomere 1 elongate, about as long as metatarsomeres 2–3 combined. Tarsomere 5 with two (occasionally three) pairs of ventro-lateral setae.</p> <p>Abdominal sternites finely pubescent; apex of last visible sternite (VII) in both sexes rounded (in male more widely than in female, almost subtruncate), with two pairs of marginal setae; these setae not distant from margin.</p> <p>Female genitalia (Figs. 3, 4). Laterotergite symmetrical, longer than wide, with two thick setae apically. Gonosubcoxite shorter than laterotergite, markedly widened posteriorly, with one preapical spine on outer margin. Gonocoxite narrow, about 0.7 times as long as gonosubcoxite, moderately curved, with relatively narrow base and with a tiny spine (short thick seta) on both ventral and dorsal edges of outer margin.</p> <p>Aedeagus (Figs. 5–8) robust, with basal bulb and parameres comparatively large (Figs. 7, 8). Median lobe of aedeagus in lateral view (Fig. 5) arcuate on dorsal side, with ventral margin almost straight in middle portion and slightly convex in the apical quarter; oblique apical capitulum slightly hooked ventrally; in dorsal view median lobe (Fig. 6) almost straight, with terminal lamella comparatively narrow, about as long as wide, rounded apically. Apical orifice in dorsal position, wide apically. Internal sac with three groups of small spines apically.</p> <p>Comparison. Coleolissus perlucens, originally described as Hypolithus perlucens, the type species of Coleolissus, is recognizable by the following combination of characters: pronotal basal angles widely rounded, protibia sulcate on dorsal side, elytra clearly punctate on lateral intervals and at apex, and abdominal sternites finely pubescent. In general habitus with widely rounded basal angles of pronotum and oval elytra, this species is somewhat similar to C. nakajimai Ito, 2016 described from one female from the “Animalai” (= Anamalai) Hills in the Indian state of Tamil Nadu, but the latter species, according to the original description (Ito, 2016), is larger (body length 12.5 mm), with vestigial wings, elytra without distinct punctation on lateral intervals (only very sparsely and microscopically punctate on disc), protibia not sulcate and abdominal sternites not pubescent.</p> <p>Distribution. This species has been described from one female collected in Jhelum Valley (Jammu and Kashmir state, India) and has been known for a long time only from this female holotype. According to recent data (Kataev &amp; Wrase, 2017; examined material listed above), this species also occurs in the Indian state of Uttarakhand and in Pakistan and seems to be endemic to the West Himalaya.</p></div> 	http://treatment.plazi.org/id/E04B4F00FFE2FFC3FF4FF9C77F4F576A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
E04B4F00FFE6FFC1FF4FFF3479C65374.text	E04B4F00FFE6FFC1FF4FFF3479C65374.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleolissus kejvali Kataev 2022	<div><p>Coleolissus kejvali sp. n.</p> <p>(Figs. 9, 11–14)</p> <p>Type material. Holotype: male, “S-India, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.083336&amp;materialsCitation.latitude=8.75" title="Search Plazi for locations around (long 77.083336/lat 8.75)">Kerala St.</a>, nr. Kallar, 30 km NE Trivandrum, 400 m Kallar riv., 29.6.99, 77°05′E, 8°45′N leg. Kejval, Trỳzna”, “coll. S. Facchini Pia Senza / Italy ” (cFCCH).</p> <p>Description (male). Habitus as in Fig. 9. Morphological character states are similar to those as described for C. perlucens, but differing as follows.</p> <p>Body slightly smaller: length 8.1 mm, width 3.3 mm.</p> <p>Head (HWmax/PWmax 0.67, HWmin/PWmax 0.48), with more convex eyes (HWmax/HWmin 1.29). Labrum very shallowly concave anteriorly. Microsculpture on clypeus, frons and vertex highly obliterated, very fine, more or less isodiametric meshes recognized only under and behind eyes.</p> <p>Pronotum (PWmax/PL 1.38, PWmax/PWmin-ap 1.54, PWmax/PWmin-bas 1.29, PWmin-bas/PWmin-ap 1.20), with shallower laterobasal depressions; basal foveae more distinct, very narrow, situated at inner margins of laterobasal depressions.</p> <p>Elytra (EL/EW 1.50, EL/PL 2.54, EW/PWmax 1.22), with subapical sinuation very shallow, but slightly more distinct. Intervals, including lateral ones, impunctate. Interval 3 with a series of 5 (left elytron) and 8 (right elytron) small discal setigerous pores adjoining stria 2 along its entire length. Marginal umbilicate series consisting of 21 (left elytron) and 20 (right elytron) setigerous pores.</p> <p>Protibia without longitudinal sulcus on dorsal side.</p> <p>Pro- and metasternum and abdominal sternites almost glabrous, only with a few scattered very fine and short, almost indistinct setae. Apex of last visible sternite (VII) (in male) subtruncate, with one (inner) pair of marginal setae; these setae not distant from margin.</p> <p>Aedeagus (Figs. 11–14) slenderer, with relatively smaller basal bulb. Median lobe of aedeagus in lateral view (Fig. 11) arcuate on dorsal side, with ventral margin rounded in middle portion and slightly convex in the apical quarter, without apical capitulum; in dorsal view median lobe (Fig. 12) more clearly curved to the right apically; terminal lamella rather wide, wider than long, more widely rounded apically. Apical orifice divided into two parts by narrow oblique sclerified bridge: a larger proximal part in dorsal position and a smaller distal one shifted to the left. Internal sac without sclerotic armature.</p> <p>Etymology. Named after Zbynĕk Kejval (Domažlice, Czech Republic), a specialist on Anticidae, one of the collectors of the type specimens.</p> <p>Comparison. Very similar in habitus to C. perlucens, but differs from it in having elytral intervals (including lateral ones) impunctate, protibia not sulcate, abdominal sternites not pubescent, apex of last visible abdominal sternite of male subtruncate, with one pair of marginal setae, median lobe of aedeagus with apical orifice divided into two parts by narrow oblique bridge, without apical capitulum and without spines in internal sac, and in other distinctive characters mentioned in the description. The new species differs from C. nakajimai Ito, 2016 described based on one female from Tamil Nadu and known to me only from the original description at least in its smaller size (body length 8.1 mm versus 12.5 mm), more convex eyes, non-crenulate elytral striae and fully developed wings.</p> <p>Distribution. Known only from the type locality (Kallar river, 30 km NE of Trivandrum) in Kerala State, India.</p></div> 	http://treatment.plazi.org/id/E04B4F00FFE6FFC1FF4FFF3479C65374	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
E04B4F00FFE4FFCEFF4FFF3479C65280.text	E04B4F00FFE4FFCEFF4FFF3479C65280.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleolissus deformipenis Kataev 2022	<div><p>Coleolissus deformipenis sp. n.</p> <p>(Figs. 10, 15–18)</p> <p>Type material. Holotype: male, “ India, Kerala, 7 km N Munnar, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.066666&amp;materialsCitation.latitude=10.15" title="Search Plazi for locations around (long 77.066666/lat 10.15)">Eravikulam</a>, 10°09′N 77°04′E 1740 m, 12– 18.IV.1997 ” (ZIN).</p> <p>Description (male). Body elongate, medium-sized for the genus, length 8.4 mm, width 3.3 mm. Habitus as in Fig. 10.</p> <p>Body black, shiny and iridescent, with labrum, mandibles basally and very narrow lateral margins of pronotum reddish brown; elytral epipleura reddish black; palps, antennae and legs yellowish brown; femora slightly paler than tibiae and tarsi; coxae blackish brown.</p> <p>Head medium-sized (HWmax/PWmax 0.69, HWmin/PWmax 0.53), impunctate. Eyes large, strongly convex (HWmax/HWmin 1.30),almost touching ventrally margins of buccal fissure. Genae glabrous.Tempora short, slightly convex, somewhat steeply sloping to neck. Labrum somewhat flat, very shallowly concave anteriorly. Clypeus very slightly convex mediobasally, very shallowly emarginated along anterior margin, with one setigerous pore at each outer angle and with very fine and short longitudinal wrinkles adjacent medially to each pore. Fronto-clypeal suture very fine, superficial, almost straight. Frontal foveae markedly deepened, without prolongation on clypeus. Frontoocular furrows fine, impressed at clypeus, vaguely reaching supraorbital furrows. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by distance less than width of antennomere 2 basally. Mentum separated from submentum by complete transverse suture, with a prominent, rather wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite moderately widened distally, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, narrow, markedly projecting beyond ligular sclerite and separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere about as long as apical one. Mandibles elongate, somewhat narrow, evenly curved apically, with very fine oblique wrinkles in apical half dorsally; left mandible acute, at most only slightly blunted at tip. Dorsal microsculpture visible throughout, very distinct at anterior margin of clypeus, finer on remaining surface, consisting of mixture of isodiametric and slightly transverse meshes. Antennae slender, surpassing pronotal basal edge by approximately two and a half apical antennomeres, pubescent from middle of antennomere 3, with antennomeres 4–8 about 2.5 times as long as wide and basal antennomere about as long as antennomere 3.</p> <p>Pronotum comparatively narrow (PWmax/PL 1.30), widest just before the middle, slightly more strongly narrowed apically than basally (PWmax/PWmin-ap 1.45, PWmax/PWmin-bas 1.36), with one lateral seta inserted in widest point. Sides rounded along entire length, more widely in basal half; lateral bead complete, throughout very narrow. Apical margin markedly, almost roundly emarginated, very narrowly bordered along entire length; apical border slightly obliterated at middle. Apical angles protruding anteriorly, narrowly rounded at apex. Basal margin almost straight medially, slightly rounded laterally, bordered along entire length, almost as long as apical margin (PWmin-bas/PWmin-ap 1.07) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions beginning from apical angles as grooves, distinctly widened from lateral setae and fused basally with basal foveae forming flat and wide laterobasal depressions isolated from each other by convex area; basal foveae distinct, very narrow, situated at inner margins of laterobasal depressions. Median line fine, superficial, not reaching apical and basal margins. Anterior transverse depression very shallow, almost indistinct. Disc densely and finely punctate, slightly more coarsely in laterobasal depressions and with very small, indistinct punctures in central portion; punctures in laterobasal depressions confluent. Microsculpture highly obliterated, consisting of indistinct transverse meshes.</p> <p>Elytra convex, elongate oval (EL/EW 1.62, EL/PL 2.61, EW/PWmax 1.23), widest behind middle; sides rounded along entire length, very broadly medially; subapical sinuation shallow, but distinct. Humerus angularly rounded, with very small, almost indistinct denticle at apex faintly visible from behind. Sutural angle narrowly rounded at tip, not extended posteriorly. Basal edge markedly arched, forming an obtuse angle with lateral margin. Striae with granulate microsculpture on bottom, impressed along entire length, reaching anteriorly basal elytral edge, comparatively wide, widest apically. Intervals glabrous and impunctate, very strongly narrowed apically, moderately convex on disc and rather strongly convex before apex. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 5 small discal setigerous pores adjoining stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series with very short gap at middle, consisting of 23 setigerous pores. Lateral groove flat, narrow basally, gradually widened apically, with granulate meshes. Microsculpture on intervals highly obliterated, not forming distinct meshes. Wings fully developed.</p> <p>Thoracic sternites punctate, with very fine and short setae on prosternum anteriorly. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</p> <p>Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side distally. Protibia without longitudinal sulcus on dorsal side, with one preapical spine (in male) on outer margin. Metafemur ventrally with two long setae at posterior margin and with a few very short setae at anterior margin. Tarsi glabrous dorsally. Pro- and mesotarsomeres 1–4 (in male) markedly widened, with biseriate adhesive scales ventrally; mesotarsomere 1 slightly longer than mesotarsomere 2. Metatarsus slender, markedly longer than HWmax; metatarsomeres 1–3 only slightly, metatarsomere 4 markedly widened distally; metatarsomere 1 elongate, very slightly longer than metatarsomeres 2 and 3 combined. Tarsomere 5 with two pairs of ventro-lateral setae.</p> <p>Abdominal sternites glabrous, with only obligatory setae; apex of last visible sternite (VII) concave (in male), with one pair of marginal setae; these setae not distant from margin.</p> <p>Aedeagus (Figs. 15–18) with basal bulb and parameres comparatively large (Figs. 15, 18). Median lobe of aedeagus strongly asymmetrical, sinuously curved, in lateral view (Fig. 16) arcuate, with ventral margin slightly convex in middle portion; almost transverse apical capitulum slightly hooked dorsally; in dorsal view apical portion (Fig. 17) sharply bent to the right; terminal lamella narrow, much longer than wide. Apical orifice vast and strongly curved along median lobe. Internal sac without sclerotic elements.</p> <p>Etymology. The specific name is a Latin noun at apposition, referring to the unusual (deformed) form of the median lobe of the aedeagus.</p> <p>Comparison. This new species is somewhat similar in habitus to C. perlucens and C. kejvali sp. n., but well differs from them in having body more elongate, fronto-ocular furrows vaguely reaching supraorbital furrows, elytra with wider striae and more convex intervals, apex of last visible abdominal sternite in male concave and median lobe of aedeagus strongly asymmetrical. In addition, C. deformipenis sp. n. differs from C. perlucens in elytral intervals impunctate, abdominal sternites not pubescent, last visible abdominal sternite of male with one pair of setae and protibia not sulcate, and from C. kejvali sp. n. in having more distinct microsculpture on head and visible throughout, and male pro- and mesotarsi more strongly widened.</p> <p>Distribution. Known only from the type locality (7 km N of Munnar) in the Western Ghats, Kerala State, India.</p></div> 	http://treatment.plazi.org/id/E04B4F00FFE4FFCEFF4FFF3479C65280	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
E04B4F00FFEAFFCCFF4FFD597D295252.text	E04B4F00FFEAFFCCFF4FFD597D295252.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleolissus iris Andrewes 1924	<div><p>Coleolissus iris Andrewes, 1924</p> <p>(Figs. 19, 21–30)</p> <p>Coleolissus iris Andrewes, 1924: 135.</p> <p>Coleolissus andrewesi Alluaud, 1932: 19 (unnecessary replaced name for Coleolissus iris Andrewes, 1924).</p> <p>Calathomimus splendens Ito, 1997: 518, syn. n.</p> <p>Coleolissus (Tenuistilus) turturensis Ito, 2016: 360, syn. n.</p> <p>Type material examined. Holotype of Coleolissus iris: male, labelled “Nagpur. C. P. India. 1,000 ft., 30.7.1918. E. A. D’Abreau. ”, “Type”, “ Coleolissus iris Andr., Type, H.E. Andrewes det.”, “ Ex. Compared with perlucens Bates [Andrewes’ handwriting]”, “ Central Mus. Nagpur, C. P.”, “ H.E. Andrewes Coll., B.M. 1945–97” (BMNH).</p> <p>Holotype of Calathomimus splendens: male, labelled “N. Indien, Uttar Pradesh, Shiwalik-Kette, S. Seite, 19 km SW Dehra Dun ”, “ 600 m, 20.8.1985, leg. Hieke ”, “ HOLOTYPE Calathomimus splendens N. ITO ” (MFNB); and paratype: 1 female, labelled “South India, Pondicherry State, Karikal ”, “ PARATYPE Calathomimus splendens N. ITO ” (MFNB).</p> <p>Additional material examined. Nepal. 1 female, Narayani / <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.36639&amp;materialsCitation.latitude=27.551943" title="Search Plazi for locations around (long 84.36639/lat 27.551943)">Chitwan</a>, 13 km W Sauraha, Kasara, Chitwan-NP, 27°33′07″N 84°21′59″E, 180 m, deciduous forest, 20.VI.2005, A. Weigel leg. (cJS); 1 male, Sauraha, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=84.49166&amp;materialsCitation.latitude=27.585835" title="Search Plazi for locations around (long 84.49166/lat 27.585835)">Hotel</a> “Sweet home”, 27°35′09″N 84°29′30″E, 180 m, LFF, 5.VI.2017, A. Weigel leg. #17–22 (NME). Pakistan. Islamabad: 3 males, 1 female, Islamabad “SW from Garni”, 1500 m, 5–15.VII.2003, V. Gurko &amp; S. Ovchinnikov leg. (ZIN); 1 male, Islamabad, National Forest Park, 12.VII.2003, S. Ovchinnikov leg. (ZIN); 1 male, Islamabad, 10.VII.2003, S. Ovchinnikov leg. (ZIN). North-West Frontier Prov.: 1 female, Khyber AG, middle stream Kabul river, 700–900 m, 15–22.VIII.2005, V. Gurko leg. (cWR). Balochistan: 1 female, Suleiman Mts., Zhob Valley, 1700 m, 8.V....[?] (SIZK). India. Punjab: 1 female, Hoshiarpur, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=76.21611&amp;materialsCitation.latitude=31.197416" title="Search Plazi for locations around (long 76.21611/lat 31.197416)">Rormazara</a>, 31°11′50.7″N 76°12′58.0″E, 308 m, 6–9.VII.2004, M. Uhler leg. (cDST). Kerala: 1 female, 7 km N of Munnar, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.066666&amp;materialsCitation.latitude=10.15" title="Search Plazi for locations around (long 77.066666/lat 10.15)">Eravikulam Nature Park</a>, 10°09′N 77°04′E, 1740 m, 12– 18.IV.1997 (ZIN); 1 male, near Kallar, 30 km NE Trivandrum, 400 m, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.083336&amp;materialsCitation.latitude=8.75" title="Search Plazi for locations around (long 77.083336/lat 8.75)">Kallar River</a>, 8°45′N 77°05′E, 29.VI.1999, Kejval, Tryzna leg. (cFCCH). Tamil Nadu: 1 male, Vilupparam Distr., <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=79.8&amp;materialsCitation.latitude=12.0" title="Search Plazi for locations around (long 79.8/lat 12.0)">Auroville</a>, 12°0′N 79°48′E, 1.VIII.–5.X.2012, F. Burger leg. (NME); 4 females, Vilupparam Distr., Auroville, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=79.7995&amp;materialsCitation.latitude=12.011666" title="Search Plazi for locations around (long 79.7995/lat 12.011666)">Discipline Vill.</a>, 12°0.7′N 79°47.97′E, 1.VII– 31.VIII.2013, local collector leg. (NME); 1 female, same but on Zizyphus mauritianus, 19.IX–30.X.2017, local collector leg. (NME). Sri Lanka. 1 female, Ratnapura Distr., Panamure env., 15.XII.1995, S. Bečvář leg. (cKM); 1 female, “ Hambantota, 10.XII.1979, Ex. coll. Viggo Mahler” (ZMUC); 1 female, Polonaruwa, 23.XI.1984, P. Beron &amp; S. Andreev leg. (NMNHS).</p> <p>Diagnosis. This species is similar to C. perlucens in general habitus (Fig. 19) and in many other characters including fore tibiae with longitudinal sulcus, abdominal sternites finely pubescent, last visible abdominal sternite rounded and with two pairs of setae in both sexes, but well differs from it as following: head very finely punctate, with fronto-ocular furrows reaching supraorbital furrows; mentum with a very wide and short, less prominent tooth; pronotum relatively narrower, with wider base, more widely rounded sides and more narrowly rounded basal angles; elytra relatively longer, their basal border more strongly arched, forming almost right or acute angle with lateral border; all elytral intervals finely punctate; pro- and mesotarsomeres in male less widened; meso- and metatarsomeres 4 in most specimens with three pairs of ventro-lateral setae; median lobe of aedeagus (Figs. 23–30) much more slender, with button-like apical capitulum and without sclerotic spines in internal sac. In addition, pronotum of C. iris is with wider yellowish brown margins, and elytra along sides and suture in apical half are also yellowish brown. Female genitalia as in Figs. 21, 22.</p> <p>Body length 7.9–9.2 mm, width 3.2–3.8 mm. Proportions (7 males and 6 females measured): HWmax/PWmax 0.62–0.68; HWmin/PWmax 0.47–0.53; HWmax/HWmin 1.25–1.38; PWmax/PL 1.34–1.43; PWmax/PWmin-ap 1.46–1.64; PWmax/PWmin-bas 1.14–1.20; EL/EW 1.54–1.61; EL/PL 2.68–2.74 in male and 2.73–2.86 in female; EW/PWmax 1.23–1.27 in male and 1.25–1.31 in female.</p> <p>Distribution. This species is widely distributed over India, Pakistan, Nepal and Sri Lanka. Andrewes (1924, 1930) recorded it from the Indian states Uttarakhand, Madhya Pradesh, Maharashtra, Karnataka and Tamil Nadu, and from Sri Lanka. Ito (1997) recorded it (as Calathomimus splendens) from the Indian states of Chandigarh, Delhi, Puducherry and West Bengal, and from Nepal. It is recorded here from the Indian states of Punjab and Kerala for the first time.</p> <p>Remarks. Andrewes (1924) has described C. iris from many specimens collected in different localities in India and Sri Lanka (type locality: Nagpur, India). Ito (1997) has described Calathomimus splendens from the series collected in the Indian states of Uttarakhand, Chandigarh, Delhi, Puducherry, Tamil Nadu and West Bengal, also in Nepal and Sri Lanka (type locality: Dehra Dun, Uttarakhand, India) as similar to Coleolisssus teradai Habu, 1978 from Taiwan without comparison of the former species with C. iris. More recently, the same author (Ito, 2016) has described C. turturensis based on one male from Nepal as similar to C. puncticollis Ito, 2008 from Laos; he also did not compare it with either C. iris or C. splendens. Examination of the original descriptions of C. splendens and C. turturensis and the holotype and one paratype of the former species revealed that both these species are conspecific with C. iris.</p> <p>Coleolissus iris demonstrates geographical variation in body length. Specimens examined from southernmost localities in India (Kerala and Tamil Nadu) and Sri Lanka are, on average, smaller (7.9–8.7 mm, mean 8.3 mm) than the specimens examined from other part of the species range (8.6–9.2 mm, mean 9.0 mm). The status of the populations from the Indian states of Kerala and Tamil Nadu and from Sri Lanka needs further study based on additional material.</p> </div>	http://treatment.plazi.org/id/E04B4F00FFEAFFCCFF4FFD597D295252	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
E04B4F00FFE8FFD7FF4FF94E7EAC5454.text	E04B4F00FFE8FFD7FF4FF94E7EAC5454.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coleolissus monstrosipenis Kataev 2022	<div><p>Coleolissus monstrosipenis sp. n.</p> <p>(Figs. 31–45)</p> <p>Type material. Holotype: male, India occ., Maharashtra st., Wai env., 70 km S of Pune, 3–6.X.2005, F. &amp; L. Kantner leg. (SMNS).</p> <p>Paratypes: 4 males, 7 females, same data as holotype (SMNS, ZIN); 1 male, 2 females, India occ., Maharashtra state, Mulshi env., 40 km W of Pune, 7–11.X.2005, F. Kantner leg. (SMNS); 1 male, 2 females, India occ., Maharashtra st., Bhushi Dam env., 4 km S of Lonavala, 500 m, 12–15.X.2005, F. &amp; L. Kantner leg. (SMNS).</p> <p>Description (5 males and 5 females measured). Body elongate, comparatively large for the genus, length 10.6– 12.4 mm, width 4.0– 4.9 mm. Habitus as in Figs. 31, 33.</p> <p>Body dark brown to black, shiny and iridescent on dorsal side, with labrum, base of mandibles and very narrow lateral margins of pronotum reddish brown; ventral side in most specimens reddish black; palps, antennae and legs yellowish brown; tibiae, tarsi, apical palpomeres and antennomeres 3–11 very slightly infuscate.</p> <p>Head medium-sized (HWmax/PWmax 0.63–0.67, HWmin/PWmax 0.51–0.56), very finely punctate dorsally. Eyes large, moderately convex (HWmax/HWmin 1.18–1.28), separated ventrally from buccal fissure by distance approximately equal to width of antennomere 1 apically. Genae glabrous. Tempora short, oblique, gently sloping to neck. Labrum somewhat flat, very shallowly and widely concave anteriorly. Clypeus very slightly convex, very shallowly emarginated along anterior margin, with one setigerous pore at each outer angle and with very fine and short longitudinal wrinkles adjacent medially to each pore. Fronto-clypeal suture fine, superficial, almost straight. Frontal foveae very small and shallow, almost punctiform, without prolongation on clypeus. Fronto-ocular furrows very fine, vaguely reaching supraorbital furrows. Frons with more or less distinct, very small and shallow medial depression. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated markedly before level of posterior margin of eyes and removed from supraorbital furrows by distance equal to width of antennomere 2 basally. Mentum separated from submentum by complete transverse suture, with a prominent acutangular median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite almost parallel-sided, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, comparatively narrow, slightly projecting beyond ligular sclerite and markedly separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere slightly longer than apical one. Mandibles moderately elongate, evenly curved apically, with oblique wrinkles in apical half dorsally; left mandible acute, at most only slightly blunted at tip. Dorsal microsculpture highly obliterated, not forming distinct meshes. Antennae slender, surpassing pronotal basal edge by approximately two and a half apical antennomeres, pubescent from apical 0.7 antennomere 3, with antennomeres 4–8 about three times as long as wide and basal antennomere slightly longer than antennomere 3.</p> <p>Pronotum only slightly wider than long (PWmax/PL 1.03–1.15), widest in the middle or just before it, slightly more strongly narrowed apically than basally (PWmax/PWmin-ap 1.41–1.55, PWmax/PWmin-bas 1.29–1.39), with one lateral seta inserted slightly before the middle. Sides evenly rounded along entire length; lateral bead complete, very narrow throughout. Apical margin moderately emarginated, almost straight medially, very narrowly bordered along entire length. Apical angles protruding anteriorly, narrowly rounded at apex. Basal margin almost straight in middle portion, slightly rounded laterally, bordered along entire length, slightly longer than apical margin (PWmin-bas/PWmin-ap 1.03–1.15) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions beginning from apical angles as grooves, markedly widened from lateral setae and fused basally with basal foveae forming deep oblique laterobasal depressions isolated from each other by undepressed area. Median line fine, superficial, rather long, almost reaching apical margin and basal margins. Anterior transverse depression very shallow. Punctation distinct and dense, restricted to lateral furrows and laterobasal depressions. Microsculpture highly obliterated, not forming distinct meshes.</p> <p>Elytra convex, elongate, slightly wider than pronotum (EL/EW 1.55–1.66, EL/PL 2.39–2.61, EW/PWmax 1.11–1.19), widest before the apical third; sides slightly rounded, with very weak sinuation before the middle; subapical sinuation distinct, but very shallow. Humerus rounded, without denticle at apex. Sutural angle blunted or narrowly rounded at tip, not extended posteriorly. Basal edge slightly arched, forming a very obtuse angle with lateral margin. Striae crenulate, impressed along entire length, reaching anteriorly basal elytral edge, with granulate microsculpture on bottom of striae in their apical portion. Intervals glabrous, convex, strongly narrowed apically, sparsely micropunctate. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 9–13 small discal setigerous pores adjoining stria 2 along entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series without distinct gap at middle, consisting of 23–31 setigerous pores. Lateral groove narrow and flat, slightly widened apically. Microsculpture highly obliterated, at most with indistinct transverse lines on intervals, with obliterate isodiametric meshes in lateral groove. Wings fully developed.</p> <p>Ventral side of thorax smooth or micropunctate, glabrous, except for sparse very fine short setae on prosternum anteriorly. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</p> <p>Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side. Metafemur ventrally with two long setae at posterior margin and with two or three very short setae at anterior margin. Protibia on dorsal side without longitudinal sulcus, on outer margin preapically with two stouter and one thiner spines in male and with three comparatively stout spines in female. Tarsi with very fine and short setae dorsally. In male, protarsomeres 1–4 and mesotarsomeres 2–4 markedly widened, with biseriate adhesive scales ventrally; mesotarsomere 1 weakly widened, with scales only apically, slightly shorter (in female, slightly longer) than mesotarsomeres 2 and 3 combined. Metatarsus (Figs. 32, 34) slender, in male slightly longer, in female slightly shorter than HWmax, with metatarsomeres 2–4 very short, markedly widened distally; metatarsomere 1 elongate, weakly widened distally, longer than metatarsomeres 2–4 combined. Tarsomere 5 in most specimens with three pairs of ventro-lateral setae.</p> <p>Abdominal sternites glabrous, smooth or micropunctate; apex of last visible sternite (VII) in male subtruncate, with one pair of marginal setae, in female rounded, with two pairs of setae; these setae not distant from margin.</p> <p>Female genitalia (Figs. 35, 36). Laterotergite asymmetrical, longer than wide, with four (occasionally five) thick setae apically. Gonosubcoxite shorter than laterotergite, widened posteriorly, with a preapical thick seta on outer margin. Gonocoxite short, strongly curved, about two times as long as gonosubcoxite, with a tiny spine (short thick seta) on ventral edge of outer side. Vagina membraneous, no specific features observed.</p> <p>Aedeagus (Figs. 37–45) with comparatively large parameres (Figs. 39, 40), moderately large basal bulb and highly modified, hypertrophic apical portion. Median lobe of aedeagus (Figs. 37, 38) narrow basally, slightly arcuate in lateral view; apical portion greatly enlarged laterally, irregular-shaped. Apical orifice divided into two isolated portions: proximal portion very vast, beginning from basal bulb on left side and going along dorsal and right side of median lobe up to its modified apical portion; four of five examined males having probably constantly everted elongate saccule “ a ” of internal sac (Figs. 37, 38) at distal margin of membranous area; this saccule absent in one of the examined males (Fig. 41); distal portion small, oval, situated on dorsal side just at apex of median lobe. Internal sac without sclerotic elements, everting mostly through distal portion of apical orifice; in everted condition (Figs. 41–45), small, with two apical lobes and with two narrow saccules “ b ” (Fig. 45) on the right side prebasally.</p> <p>Etymology. The specific name is a Latin noun at apposition, referring to the monstrous form of the median lobe of the aedeagus.</p> <p>Comparison. Having elongate, almost parallel-sided elytra, Coleolissus monstrosipenis is somewhat similar in appearance to the members of the genus Oxycentrus Chaudoir, 1854, for example, to O. fulgens Ito, 1997 from Vietnam.The new species is treated as belonging to Coleolissus since it possesses the following characters typical for this genus: frontal foveae without prolongation on clypeus, parascutellar striole long, interval 3 with a row of 9–13 setigerous pores, and protibia with two preapical spines on outer margin (see Kataev, 2021). Within Coleolissus, C. monstrosipenis sp. n. is similar to C. iridipennis Ito, 1999 from Laos in having elongate body with widely rounded basal pronotal angles, but well differs from it in having metatarsomeres 2–4 very short and markedly widened distally, apex of last visible abdominal sternite (VII) in male with one pair of marginal setae and median lobe of aedeagus strongly modified. In metatarsus with shortened tarsomeres 2–4, the new species is distinguished from all other congeners, which have all these tarsomeres slenderer, with combined length greater than metatarsomere 1. The new species can be additionally distinguished from C. inessae Kataev, 2021, which is also described from the Western Ghats and also has elongate body and only one pair of setae on last abdominal sternite of male, in greater size (body length 10.6–12.4 mm versus 7.8 mm) and pronotum markedly depressed laterobasally and with more widely rounded sides and basal angles.</p> <p>Distribution. The type series has been collected in Western India, the state of Maharashtra, on the western margin of the Deccan Plateau at the eastern slopes of the Western Ghats at altitudes of about 500 m.</p> <p>Remarks. The very unusual shape of the median lobe of the aedeagus in this new species is far beyond the known variability of genital structures not only in Harpalini but also in other ground beetles, and at first glance gives the impression of an individual anomaly; however, since all males examined have such genitalia, there is no doubt that this unusual structure is a constant feature of the species. Particular attention should be given to the similarity of the modified apical portion of the median lobe of C. monstrosipenis sp. n. with inverted internal sac to the apical portion of the median lobe of many carabids with fully everted internal sac. In everted condition, the internal sac of carabids looks as an apical extension of the sclerified median lobe (Deuve, 2021). The shape and sclerotic armature of the fully everted internal sac are very variable in different species and are considered the most reliable morphological basis both for distinguishing similar species and for grouping species into natural units. The special investigations indicate that the internal sac of carabids is the real copulatory organ that not only forms spermatophore, but also transfers it to the vagina of the female during mating (Jeannel, 1955); it acts as an intromittent apparatus and its features apparently correspond as a counterpart with the vagina of the same species (Ishikawa, 1978, 1987; Sasabe et al., 2010). During genital coupling, males mould the spermatophore within the vagina using the everted internal sac. Strong adhesion of spermatophore to a particular site of the vagina seems to be unique for each species (Takami, 2002). Although these investigations were carried out on the Carabus species, their results are apparently suitable for other carabids. It is possible that the modified apical portion of the aedeagus of C. monstrosipenis sp. n. took over some of the mechanical functions of the internal sac, which is quite small in this species (Figs. 41–45). Small internal sac everting in two places of the median lobe (before the modified apical portion and at its apex) counts in favor of this assumption. The genetic base of such converting is obscure and needs special study. The members of Harpalini demonstrate several cases of definite modifications of the aedeagus as a result of probably hormonal imbalance during the embryonic development (Kataev, 1995; Kataev &amp; Wrase, 2019). It has been suggested that in the case of the genetic determination of such modifications, the aberrant specimens under certain conditions can be considered as material for speciation (Tichomirova, 1979, 1991) and give rise to species with sharply different, including monstrous, genitalia (Kataev &amp; Wrase, 2019). Among ground beetles, strong modifications of the male genitalia are quite rare, but in some other beetle families, for example in Catopidae, Leiodidae and Pselaphidae, they seem to be more common (Jeannel, 1955). The discovery among representatives of the genus Coleolissus of several species with an unusual structure of the aedeagus suggests that this genus has a genetic predisposition to such genital transformations.</p> </div>	http://treatment.plazi.org/id/E04B4F00FFE8FFD7FF4FF94E7EAC5454	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
E04B4F00FFF3FFD4FF4FFAB57CB95134.text	E04B4F00FFF3FFD4FF4FFAB57CB95134.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siopelus (Neosiopelus) liangi Kataev 2022	<div><p>Siopelus (Neosiopelus) liangi sp. n.</p> <p>(Figs. 20, 46–49)</p> <p>Type material. Holotype: male, China, Yunnan, Jinping: border Mengla and <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.141556&amp;materialsCitation.latitude=22.649166" title="Search Plazi for locations around (long 103.141556/lat 22.649166)">Nafa</a>, 323 m, 22°38′57.0″N 103°08′29.6″E, 15.XII.2003, B. Kataev leg. (ZIN).</p> <p>Description (male). Body comparatively wide, medium-sized for the genus, length 6.5 mm, width 2.9 mm. Habitus as in Fig. 20.</p> <p>Body black, shiny, not iridescent, with labrum, mandibles basally and clypeus apically yellowish brown, very narrow margins of pronotum reddish brown; palps, antennae and legs yellowish brown; antennae from antennomere 3 and tarsi partly slightly infuscate; coxae blackish brown.</p> <p>Head medium-sized (HWmax/PWmax 0.66, HWmin/PWmax 0.52). Eyes large, moderately convex (HWmax/ HWmin 1.31), separated ventrally from buccal fissure by distance less than width of antennomere 2 basally. Genae with a few fine and short setae. Tempora flat, gently sloping to neck. Labrum very shallowly concave at middle of apical margin. Clypeus slightly flattened and very shallowly arcuately emarginated along apical margin and labral base partly visible in this emargination; one setigerous pore at each outer angle of clypeus. Fronto-clypeal suture fine and slightly arched. Frontal foveae punctiform. Fronto-ocular furrows very fine and short. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by distance approximately equal to width of antennomere 2 basally. Mentum without median tooth, separated from submentum by complete transverse suture; epilobes comparatively narrow; submentum with one pair of long lateral setae. Ligular sclerite moderately narrowed distally, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, wide, markedly projecting beyond ligular sclerite and not separated from it. Basal labial palpomere not carinate; penultimate labial palpomere narrower and shorter than apical one. Mandibles not elongate, sharply curved in apical quarter; apex of left mandible subtruncate in frontal view. Dorsal surface glabrous, finely and irregularly punctate and with fine longitudinal wrinkles on clypeus laterally. Dorsal microsculpture highly obliterated, consisting of indistinct slightly transverse meshes. Antennae slender, surpassing pronotal basal edge by approximately two apical antennomeres, pubescent from middle of antennomere 3, with antennomeres 4–8 about two times as long as wide, and basal antennomere slightly longer than antennomere 3.</p> <p>Pronotum markedly transverse (PWmax/PL 1.48), widest at the middle, more strongly narrowed apically than basally (PWmax/PWmin-ap 1.48, PWmax/PWmin-bas 1.15), with one lateral seta inserted in the end of anterior third. Sides rounded along entire length, more widely in basal half; lateral bead complete, throughout very narrow. Apical margin moderately emarginated, almost straight medially, very narrowly bordered along entire length. Apical angles protruding anteriorly, narrowly rounded at tip. Basal margin almost straight medially, slightly rounded laterally, bordered along entire length, longer than apical margin (PWmin-bas/PWmin-ap 1.29) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles obtuse, rounded at apex. Disc convex, widely depressed laterobasally. Basal foveae wide and very shallow, vaguely outlined. Median line fine, superficial, not reaching apical and basal margins. Anterior transverse depression very shallow. Disc densely and finely punctate, more coarsely in laterobasal depressions, more finely in remaining surface, with finest punctures in central portion; punctures in laterobasal depressions confluent. Microsculpture not recognized under 56x magnification.</p> <p>Elytra convex, comparatively wide (EL/EW 1.44, EL/PL 2.65, EW/PWmax 1.24), widest just behind middle; sides rounded apically and basally, almost straightly diverging in medial portion; subapical sinuation shallow, but distinct. Humerus angulate, without denticle at apex. Sutural angle almost right, slightly blunted at tip. Basal edge moderately arched, forming an obtuse angle with lateral margin. Striae impressed along entire length, reaching anteriorly basal elytral edge, slightly widened apically. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Intervals narrowed apically, slightly convex on disc and more strongly convex before apex. Intervals 3 and 5 with a series of small discal setigerous pores adjoining striae 2 and 5 respectively along entire length; interval 7 with one discal pore adjoining stria 6 in its basal quarter on left elytron and without pore on right elytron. All intervals densely and somewhat coarsely punctate, with two or three punctures across each interval in its middle portion; two lateral intervals with scattered very fine and short setae. Marginal umbilicate series consisting of 18 and 22 setigerous pores, concentrated apically and basally. Lateral groove slightly widened apically, with short elongate convexity along the lateral groove at apex. Microsculpture on intervals recognized only preapically (meshes obliterated, transverse); lateral groove and bottom of striae with distinct isodiametric microsculpture. Wings fully developed.</p> <p>Ventral side of thorax finely punctate and with very fine and short scattered setae. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</p> <p>Metacoxae finely punctate and pubescent, each only with two obligatory setigerous pores. Profemur with moderately deep longitudinal excavation on inner side distally. Metafemur ventrally with two long setae at posterior margin and with two very short setae at anterior margin laterally. Protibia without longitudinal sulcus on dorsal side, with (in male) two or three preapical spines on outer margin. Tarsi very sparsely setose dorsally. Pro- and mesotarsomeres 2–4 (in male) weakly widened, with biseriate adhesive scales ventrally (only with a few scales on mesotarsomeres); pro- and mesotarsomere 1 elongate, without scales, about as long as pro- and mesotarsomeres 2 and 3, respectively, combined. Metatarsus slender, as long as HWmax; metatarsomeres 1 and 2 only slightly and metatarsomere 3 and 4 markedly widened distally; metatarsomere 1 elongate, as long as metatarsomeres 2 and 3 combined. Tarsomere 5 with two or three pairs of ventro-lateral setae.</p> <p>Abdominal sternites with fine and short setae; apex of last visible sternite (VII) (in male) widely rounded, with two pairs of marginal setae; these setae not distant from margin.</p> <p>Aedeagus (Figs. 46–49) with comparatively large parameres (Figs. 48, 49). Median lobe of aedeagus in lateral view (Fig. 46) arcuate, with ventral margin convex in middle portion; in dorsal view (Fig. 47) almost straight, with large oval lamella protruded on the left side; terminal lamella wide, much longer than wide, rounded at apex. Apical orifice in dorsal position. Internal sac three large groups of long spines.</p> <p>Etymology. The species is named after my good friend and colleague Hongbin Liang (Beijing), a talented carabidologist and tireless researcher of the Chinese, particularly of Yunnan, fauna.</p> <p>Comparison. Except for the exclusively Oriental taxon Harpaliscus Bates, 1892, treated either as a separate genus (e. g. Kataev &amp; Wrase, 2017), or as a subgenus of Siopelus (e. g. Facchini, 2021), or even as a synonym of the latter genus (Noonan, 1985a), only one species of Siopelus in narrow sense, S. tamilnadensis Kataev, 2002, has been known from the Oriental region so far, with all other members distributed in the Ethiopian region. Siopelus tamilnadensis, known from India, Nepal and Myanmar (Kataev, 2002; Kataev &amp; Wrase, 2016), is isolated from other Siopelus not only geographically but also taxonomically due to an unique combination of morphological characters somewhat intermediate between Siopelus s. str. and Laparhetes Jeannel, 1946 (Kataev, 2002). In having ciliate basal edge of pronotum and distinct pubescence on lateral intervals of elytra, S. tamilnadensis is also similar to the members of the Ethiopian genus Afromizonus Basilewsky, 1947 and the Oriental genus Prakasha Andrewes, 1919. In contrast, S. liangi sp. n. has all distinctive features of the subgenus Neosiopelus Basilewsky, 1946; it distinctly differs from S. tamilnadensis in having flatter body lacking bluish green lustre on elytra, pronotum and elytra more coarsely and densely punctate on disc, pronotal basal edge without fringe of short setae, elytral interval 7 without series of discal setigerous pores (only one such pore present on left elytron in the holotype), lateral intervals more scarcely (rather indistinctly) pubescent, protibia not sulcate dorsally and male pro- and mesotarsi narrower. The new species also differs from S. tamilnadensis and all other congeners in very unusual asymmetrical median lobe of aedeagus with lateral lamella on the left side.</p> <p>Among the Ethiopian species of Neosiopelus, S. liangi sp. n. is similar in habitus to S. babaulti (Basilewsky, 1946) from Central and East Africa, which also has pronotum and elytra densely and coarsely punctate, elytral interval 7 without row of setigerous pores, and mentum edentate, but well differs from the new species in greater body size (7.5–8.5 mm) and in having pronotum less transverse, with basal angles more widely rounded and elytral striae wider. Additionally, discal pores of S. babaulti are larger, its male pro- and mesotarsi are more strongly widened and median lobe of aedeagus is without lateral lamella.</p> <p>Distribution. Known from southern Yunnan, China. The first record of Siopelus (subgenus Neosiopelus) from China.</p> </div>	http://treatment.plazi.org/id/E04B4F00FFF3FFD4FF4FFAB57CB95134	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kataev, Boris M.	Kataev, Boris M. (2022): Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China. Zootaxa 5168 (3): 285-305, DOI: https://doi.org/10.11646/zootaxa.5168.3.2
