taxonID	type	description	language	source
03F5312357788C7BFD6C674BDF0D74BA.taxon	materials_examined	Type species: Pelops bifurcatus EWING, 1909 (= Pelopsis nudiuscula HALL, 1911) Diagnosis. Adults areuniqueamong theMycobatidaein having thefollow - ing combination of character states: granular cerotegument extending ventrally in groove at edge of ventral plate; rostrum with lateral minitecta; enantiophysis present between base of seta in and bothridium; seta in borne on ridge extending between enantiophysis; seta in broad, heavily barbed dorsally and laterally, bifurcate distally; tutorium with large triangular cusp; pedotectum I with proximal third concave, expressed as narrow ridge, strongly convex medially; semicircular carina present close to margin of ventral plate, between circumpedal carina and porose area Ah; circumpedal carina not merging with discidium; octotaxic system of four pairs of saccules; posterior notogastral tectum without overlapping lobes; subcapitulum without mental tectum. Description. Poronotic, brachypyline oribatid mites, with character states of the Mycobatidae (GRANDJEAN 1954). Adult: Granular cerotegument restricted to region between pteromorphs, pedotectum I, tutorium, and lateral body wall, extending medially along dorsosejugal scissure and circumventrally in groove at edge of ventral plate (Fig. 14). Rostrum convex, margin medially concave, with or without lateral dens. Rostrum with pair of lateral minitecta extending from ventral of lamellar setae to edge of genal process (Fig. 12). Rostral seta barbed, directed anteromedially. Lamella present or reduced to ridge, with or without cusp; translamella absent. Lamellar setae borne anteriorly or anteroventrally on cusp, or borne directly on prodorsum. Interlamellar seta broad, strongly barbed dorsally and laterally, bifurcatedistally (Figs 7, 9), borneon ridge (Fig. 9). Enantiophysis welldeveloped between base of seta in and bothridium (Fig. 9). Bothridium cup-shaped (Fig. 9). Sensillus clavate (Figs 1, 8, 9). Porose area Ad present. Tutorium with well-developed triangular cusp. Pedotectum I with proximal third concave, expressed as narrow ridge, strongly convex medially; not covering base of exobothridial seta (Fig. 3). Porose areas Am and Ah present, porose area Al absent. Discidium large, triangular (Fig. 2). Posteriorly directed, semicircular carina present posterodorsal of acetabulum IV, between circumpedal carina and porose area Ah (Fig. 3; arrowhead). Postanal porose area present or absent. Dorsal apodemes (dorsophragmata) separate. Notogaster with 10 pairs of setae or their alveoli. Notogaster with medial process on anterior tectum, process flattened or slightly convex, often covering base of seta in (Fig. 8). Pair of thickened bands of integument bordering medial process, evident in transmitted light. Concave ridges extending from medial process almost to anterior edge of pteromorph (Figs 9, 10). Lenticulus present. Octotaxic system present as four pairs of saccules. Pteromorphs curved ventrally, with line of desclerotization clearly evident, extending four-fifths length of pteromorph (Fig. 7). Undivided posterior notogastral tectum present. Epimeral setal formula 3 – 1 – 3 – 3. Small custodium present. Circumpedal carina ending on ventral plate medial to discidium (Fig. 13). Spermapositor (male genital sclerite) normal for family, approximately half length of genital plate, when measured in ventral view on slide-mounted specimens. Genital papilla Va of maleand femalesubequal in sizeand shapeto Vm and Vp. Palp setal formula 0 – 2 – 1 – 3 – 9 (1). Subcapitulum without mental tectum. Gena without posteriorly directed tectum covering base of seta m. Seta l’ of palptibia setiform. Axillary saccule of subcapitulum present. Tarsi tridactylous. Dorsal integument of tibiae and tarsi II and IV thickened (Fig. 5). Femora II to IV with projection ventrally and abaxially. Seta s of tarsus I eupathidial; seta l ” on genua I and II setiform (Fig. 14). Tarsus II with or without dens abaxial to solenidia. Immatures – Unknown. Relationship. We have examined the syntypes of Pelopsis bidentatus (HAMMER 1961). The purported longitudinal groove and the “ sense papillae ” on the rostrum noted by HAMMER (1961) arethickenings of thebaseof therostral tectum. These, present in all species of Pelopsis, are visible in transmitted light, but being enclosed in the camerostome, are not evident in scanning electron micrographs (Figs 10, 12). The lamellae are lamellate proximally and ridge-like distally. The “ short ridges ” on the anterior of the notogaster are the thickenings of the notogastral tectum present in all species of Pelopsis. In Pelopsis the cerotegument extends as a circumventral band in the groove formed at the edge of the ventral plate. A similar development of the cerotegument is found in all members of the ceratozetoid family Zetomimidae (BEHAN- PELLETIER & EAMER 2003). It is possiblethat in Pelopsis this band of cerotegument forms a “ circumventral plastron ” as has been observed in the zetomimid genus Heterozetes (BEHAN- PELLETIER & EAMER 2003). The presence of a semicircular carina posterodorsal of acetabulum IV and close to the margin of the ventral plate, is a character state shared with the mycobatid genera Mycobates and Punctoribates. These latter genera also have a linear carina, anterior to the semicircular carina, which is also found in the mycobatid genus Zachvatkinibates (BEHAN- PELLETIER 1994). The pedotectum I with strongly concave dorsal margin is a character state also found in the genera Punctoribates and Minunthozetes and in some species of Zachvatkinibates (BEHAN- PELLETIER 1988). The presence of (1) minitecta on the rostrum; (2) bifurcate and barbed interlamellar setae, and (3) octotaxic system developed as saccules, are character states uniqueto Pelopsis among theMycobatidae (B EHAN- PELLETIER 1988). Pelopsis shares the following four character states with the mycobatid genera Zachvatkinibates, Punctoribates and Minunthozetes: interlamellar setae borne on a ridge; enantiophysis present between interlamellar setae and bothridium; notogaster with medial process; thickened bands bordering medial process. These character states are considered synapomorphic (BEHAN- PELLETIER 1988), and definethesubfamily Minunthozetinae GRANDJEAN (sensu SHALDYBINA (1975) and PAVLITSHENKO (1994 )) of the family Mycobatidae.	en	Behan-Pelletier, V., Eamer, B., K. W (2003): Redefinition Of Pelopsis (Acari: Oribatida: Mycobatidae), With Description Of Pelopsis Baloghi Sp. N. From Costa Rica. Acta Zoologica Academiae Scientiarum Hungaricae 49 (1): 5-15, DOI: 10.5281/zenodo.5731728
03F53123577C8C77FD636426DE5C73FA.taxon	description	(Figs 1 – 14) Material examined – Holotype: adult m. Costa Rica: Heredia, Estación Biologica La Selva, Swampo Experimentale, 10 ° 26 ’ N 84 ° 01 ’ W, 9 June 1997 (V. BEHAN- PELLETIER), aquatic vegetation and debris, deposited in the Acari collections of INBio, Insitituto Nacional de Biodiversidad, Santo Domingo, Costa Rica. Paratypes: 10 with same data as holotype; 2 with same data as holotype, except 17 May 1995, from saturated soil. Paratypes deposited in the Canadian National Collection of Insects and Arachnids, Agricultureand Agri-Food Canada, Ottawa, Canada, INBio, thecollections of R. A. N ORTON and the Hungarian Natural History Museum, Budapest. Etymology – This species is named in honor of Professor JÁNOS BALOGH for his outstanding contributions to knowledge of Oribatida worldwide. Diagnosis. Adult – Total length 420 – 508 µm; integument striate on notogaster, pteromorphs, epimeres, ventral, genital and anal plates and mentum; lamella developed as very strong ridge, 91 – 101 µm long; lamellar setae arising anteroventrally on lamellar cusps; seta in 98 – 127 µm long; sensillus 60 – 72 µm long, with heavily barbed, long clavate head; notogastral setae very short, thin, at most 6 µm long. Description. Adult: Measurements – Mean total length: females (n = 10) 454 µm (range 420 – 508); males (n = 3) 441 µm (range 422 – 499). Mean notogastral width: females (n = 10) 304 µm (range 288 – 336), males (n = 3), 303 µm (range 288 – 336). Integument. Microtuberculate on prodorsum, tutorium, genal process, pedotectum I, anterolaterally on epimere I, and on leg segments. Striate on notogaster, pteromorphs, epimeres, ventral, genital and anal plates, mentum, abaxially on femora III and IV and dorsodistally on trochanters III and IV. Prodorsum – Rostral margin strongly concave medially, with 2 lateral dens (Figs 11, 12). Seta ro 53 – 58 µm long, weakly barbed, acuminate (Figs 1, 12). Lamella developed as very strong ridge, 91 – 101 µm long, almost parallel (Figs 1, 9), ridge undulating in bothridial region (Fig. 9); lamellar cusp 17 – 19 µm long, parallel (Fig. 8). Seta le barbed, 19 – 24 µm long, arising anteroventrally on lamellar cusp, directed anterolaterally, extending beyond tip of rostrum (Figs 10, 11). Seta in 98 – 127 µm long, inserted on transverse ridge, dorsoventrally flattened, strongly barbed dorsally and laterally, narrow proximally and distally, broadening medially, with conspicuous medial and lateral dens about 24 µm long and small dens between them about 4 – 7 µm long (Figs 1, 10). Mutual distance of setal pairs ro-ro, le-le and in-in, about 60 µm, 22 – 24 µm and 29 – 31 µm, respectively. Seta ex barbed, about 35 µm long. Sensillus 60 – 72 µm long, with heavily barbed, long clavate head, rounded distally, curved anteromedially (Figs 1, 8). Dorsal apodemes (dorsophragmata) well separated basally (Fig. 1). Porose area Ad circular and medial to bothridium. Lateral aspect of prodorsum – Genal process triangular, tapered to point, with well-developed ridge (Fig. 12). Tutorium, including cusp, about 112 µm long, distinctly pointed distally, (Figs 3, 12), with longitudinal ridges dorsoproximally (Fig. 9); tutorial cusp about 36 µm long. Tutorium strongly curved medially forming concise fit with ridge on genal process (Figs 11, 12). Pedotectum I with strongly concave dorsal margin, with dorsal insertion posterior of seta ex (Fig. 3). Custodium about 12 µm long (Fig. 14). Circumpedal carina strongly developed, curving to level of setae 3 b (Figs 3, 13, 14). Notogaster – Longer than wide, ratio 1.2: 1. Lenticulus well-developed, subrectangular in shape (Fig. 1). Notogastral striae with thumb-print pattern, directed longitudinally on anterior third of notogaster and transversly posterior to level of saccule S 1 (Fig. 6). Notogastral setae very short, thin, at most 6 µm long (Fig. 1). Saccules small, clearly evident, Sa positioned anterior of seta lm, S 1 posterolateral of lp, S 2 midway between setae h 3 and h 2, S 3 lateral of seta h 1 (Fig. 1). Ventral region – Epimere I tuberculate anterolaterally, remainder of epimeres I to IV striate (Figs 11, 12). Epimeral setae weakly barbed, tapered, 3 b longest setae, about 34 µm, other epimeral setae 16 – 24 µm long. Genital plates with longitudinal striae, anal plates with predominantly transverse striae, striae on ventral plate predominantly longitudinal (Fig. 2). Genital setae about 17 µm, g 1 and g 2 weakly barbed, positioned on anterior margin of plate (Fig. 13). Aggenital, anal and adanal setae smooth, thin, about 6 – 10 µm long. Postanal porose area not evident. Gnathosoma – Mentum with predominantly longitudinal striae; gena with striae directed anteromedially (Figs 2, 11). Axillary saccule of subcapitulum about 10 µm long. Legs – Setation (I-IV): trochanters, 1 – 1 – 2 – 1; femora, 5 – 5 – 2 – 2; genua 3 (1) – 3 (1) – 1 (1) – 2; tibiae, 4 (2) – 4 (1) – 3 (1) – 3 (1); tarsi, 20 (2) – 15 (2) – 15 – 12 (Figs 4, 5). Tibia I with small anterodorsal dens rior of circumventral groove indicated by arrowhead (Fig. 4). Tibiae I and II indented abaxially (Fig. 10). Femur II with ventral ridge and ventral projection, with seta bv ” positioned abaxial to ridge (Fig. 5). Tibia and tarsus II with dorsal ridge; tarsus II with two tandem dorsal spines abaxial of solenidia (Fig. 5). Trochanters and femora III and IV with ventral carina and ridge; femora III and IV and trochanter IV with striae abaxally. Tibia and tarsus IV thickened dorsally; dorsal integument of tibia and tarsus III not thickened. Remarks – The presence of dens dorsally on tarsus II is one of the character states used by HAMMER (1961) to separate Pelopsis bidentatus from thetypespe - cies, Pelopsis bifurcata. Neither EWING (1909), in theoriginal description of the type species, or WOOLLEY (1958) who redescribed the species, noted dens dorsally on tarsus II. Dens are not evident on a cotype slide specimen of P. bifurcata that we have examined. However, variation in this character state in specimens from North America argues for caution in its use as a key character. Adults of Pelopsis baloghi though similar to the two other described species in the genus, can be easily distinguished on the basis of character states outlined in the following key.	en	Behan-Pelletier, V., Eamer, B., K. W (2003): Redefinition Of Pelopsis (Acari: Oribatida: Mycobatidae), With Description Of Pelopsis Baloghi Sp. N. From Costa Rica. Acta Zoologica Academiae Scientiarum Hungaricae 49 (1): 5-15, DOI: 10.5281/zenodo.5731728
