taxonID	type	description	language	source
B23797F1FBA45A4FACFAD4F70C2B6C95.taxon	description	Figs 3, 4	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B23797F1FBA45A4FACFAD4F70C2B6C95.taxon	biology_ecology	Biology and distribution. The only recorded host plant of Calameuta punctata is Alopecurus pratensis L. (Vikberg 1978; Liston 2015), on which it is possibly monophagous. Accordingly, C. punctata occurs mostly in rather moist places. Its wider geographic range is not entirely clear, particularly because the identity of Calameuta pravei (Dovnar-Zapolskij, 1926) remains unresolved. This has been considered to be a valid species (e. g. Llorente and Gayubo 1990), or a synonym of C. punctata (e. g. Taeger et al. 2010). Calameuta pravei was recorded by Gussakovskij (1935) from Transcaucasia, Crimea, and the western Kopet-Dagh (Turkmenia). Llorente and Gayubo (1990) added records from Spain. Excluding these records, C. punctata is known from south-east Russia, Transcaucasia and south Siberia to Irkutsk (Gussakovskij 1935, as Cephus filum), and central and northern Europe (specimens examined by us). The distribution of C. punctata is therefore rather different from that of C. variabilis (see below), but their ranges overlap at least in parts of south-central Europe. Calameuta variabilis [as C. haemorrhoidalis auct.] has been stated to occur in Germany based on the mention of Germany as the type locality in the original descriptions by Fabricius (1781) of Tenthredo haemorrhoidalis and by Klug (1803) of Astatus punctatus, A. floralis and A. analis, coupled to an apparently faulty understanding of which taxa are represented by these names. Blank et al. (1998, 2001) listed C. haemorrhoidalis from Germany, only for Berlin-Brandenburg, dating the record respectively as " vor 1803 " and " 1802 ". This refers to the type material of Klug's species. Later, Liston et al. (2012) treated C. haemorrhoidalis auct. [C. variabilis] as extinct in Germany, and added C. punctata to the German list, based on recently collected specimens. Although it cannot be ruled out that C. variabilis once occurred in Germany, but has since disappeared, we think it more likely that in historical times only C. punctata ever occurred there, and propose in future to include only it in the list of German Symphyta. Fennoscandian and Estonian specimens which were previously identified as C. filum also belong to C. punctata. Based on COI sequences, C. punctata is split into two barcode clusters. Three specimens from Finland and one from Estonia are identical (BOLD: ACQ 7596), but differ from two German specimens by 5.0 - 5.5 % (no BIN assigned yet, GenBank accessions MW 353981 and MW 353982). The BOLD: ACQ 7596 is closer to C. pallipes, differing by a minimum of 4.1 %.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A24CA63B1BF65A26B698AB9EEDE40F68.taxon	description	Fig. 5	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A24CA63B1BF65A26B698AB9EEDE40F68.taxon	biology_ecology	Biology and distribution. According to Macek et al. (2020, under Calameuta haemorrhoidalis) the host plants of C. variabilis are various Poaceae, including cereals such as rye (Secale cereale) and wheat (Triticum). According to our personal experiences, adults occur in dry places, mostly on or near wild grasses. The taxon to which the name C. variabilis is now applied has so far been found only in the West Palaearctic, and has an essentially Mediterranean distribution, summarized by Gussakovskij (1935) as comprising southern Europe, Crimea, Caucasus, western Turkmenia (Kopet-Dagh) and Syria. In Central Europe, it reaches at least as far north as Hungary (Zombori 1978), but specimens recorded from Austria and the Czech Republic (see Taeger et al. 2006) should be checked.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
9495D1C86C0C56AF873890DF3FDD581E.taxon	description	Fig. 6	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5FD5309B01BD5F0EB98A5474CED58AEB.taxon	description	Fig. 7 B, C	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5FD5309B01BD5F0EB98A5474CED58AEB.taxon	biology_ecology	Biology. ZMUO specimens, from Finland, are not reared; Sanguisorba, recorded as a host plant by Benson (1952), can be excluded as a possible host plant at these localities, but Potentilla erecta is present in abundance at one of the localities in Eastern Finland.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
742BFB48E2B15B8B8EED1F77953D42A2.taxon	description	Fig. 7 A	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
742BFB48E2B15B8B8EED1F77953D42A2.taxon	biology_ecology	Biology. Substantiating the statement about the host plant by Benson (1952), the ZMUO specimens, collected in the Helsinki area, were reared from larvae feeding on cultivated Fragaria.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5821C75CF43A5312BEC6D962BB6CEBDE.taxon	description	Figs 8, 9	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5821C75CF43A5312BEC6D962BB6CEBDE.taxon	description	Description. Holotype ♀. Figs 8 A, B, 9 B, C. Colour. Fig. 8 A, B. Body black with terga 2 - 7 and sterna 2 - 7 orange. Terga 8 - 9 brownish black, 10 brownish basally but yellowish brown apically, apical third of protibiae brownish. Ventral margin of valvula 3 and cerci brownish. Wings clear. Measurements (mm). Body length: 5.0, distance from tegula to base of pterostigma: 3.1, head breadths: 1.46 (max. at eyes), 1.28 (max. behind eyes), 1.07 (min. behind eyes), head length (behind eyes): 0.21, max diameter of eye: 0.62, breadth of postocellar field: 0.36, OC: 0.96, OOCL: 0.35, OOL: 0.20, POL: 0.19. Length of: metafemur: 1.18, metatibia: 1.76, ovipositor sheath (from the base of valvifer 2 to the apex of valvula 3): 1.01. Length of flagellomeres: 1: 0.48, 2: 0.41, 3: 0.37, 4: 0.31, 5: 0.29, 6: 0.26, 7: 0.25. Large structures. Clypeal emargination almost half as deep as clypeal median length. Clypeus asymmetrical, its antero-lateral lobes round at apex, the left lobe longer than the right. Distance between antennal sockets 1.6 x as long as the malar space. Length of antennomere 3 about 1.24 x that of 4. Oblique furrow not outlined. Distance between cenchri about 1.1 x as long as width of cenchrus. Metatarsomere 1 about 1.75 x as long as 2. Mesoscutellar appendage with weakly outlined ridge medially. Setae. Metascutellum with few setae (about 5). Abdomen dorsally mostly pubescent from segment 7 to last one (laterally and ventrally from 2 to last one). Longest setae of valvula 3 in dorsal view markedly curved in apical third, and forming an angle of about 80 - 90 ° (Fig. 9 B). Macrosculpture. Pits on head rather irregular in size and distribution, more distinct and separated on postocular and postocellar area. Density and size of pits on thorax vary, the largest (partly fused) on lateral sides of median mesoscutal lobes and on mesoscutellum. Pits on lateral lobes of mesoscutellum and near the median mesoscutal groove sparse and small. Pits on mesepisternum largest, partly isolated and round, partly polygonal and fused (Fig. 9 C). Distinct pits on pectus nearly absent (Fig. 9 C). Microsculpture. Meshes on mesoscutellar appendage extensive, on tergum 1 absent, on metepimeron (Fig. 9 C) and on ventral surface of metacoxa absent. Sculpticells on anterior half of katepimeron of mesopleuron rib-like, irregular and of uneven height, and absent on metepimeron, pectus, and terga (all glossy). Male. Figs 8 C, D, 9 A, D-G. Closely resembles the female, but the anterior margin of tergum 2 blackish, basal 2 / 3 of hind tibia (with apical spurs), and base of metatarsomere 1 reddish yellow (Fig. 8 C, D). Penis valve valviceps long and narrow (Fig. 9 G). Measurements (mm). Body length (paratype male ZMUO. 035740): 5.3, distance from tegula to base of pterostigma: 3.2, max. diameter of eye: 0.60, head breadth: 1.47 (max. at eyes), breath of postocellar field: 0.38, head length (behind eyes): 0.24, OC: 1.01, OOCL: 0.18, OOL: 0.29, POL: 0.21, metafemur: 1.25, metatibia: 1.64, metatarsus: 0.50. Length of flagellomeres: 1: 0.59, 2: 0.51, 3: 0.50, 4: 0.44, 5: 0.41, 6: 0.39, 7: 0.41.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5821C75CF43A5312BEC6D962BB6CEBDE.taxon	etymology	Etymology. The name is an adjective derived from the species name of its closely similar sister species, D. aericeps Thomson, using the Latin diminutive suffix - ellus.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
D6B0BCA16A4C5615B22176AF98FFE5EC.taxon	description	Fig. 10 A	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6FED188EF92E549296338AB6A1DCB807.taxon	description	Fig. 17 A-H	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6FED188EF92E549296338AB6A1DCB807.taxon	description	Characters of the female. The female of D. coruscans, based on two specimens, resembles D. picipes in many characters (the males are even more similar) and the melanic form of D. liogaster with black legs, but differs by its mostly smaller body size of 6 - 7 mm and the rather inconspicuous setation of the ovipositor sheath (valvula 3) (Fig. 17 G). Due to their similarity, the diagnostic characters of Dolerus coruscans are compared to their states in Dolerus picipes. Head. (Fig. 17 B, C) Closely resembles D. picipes, but the minimal distance between the antennal sockets is about 2 x the length of the malar space (clearly shorter in Dolerus picipes). Pits on face and vertex more unevenly distributed and vary more in their size. Postocellar field less distinctly defined than in Dolerus picipes and lateral postocellar furrows less distinct: short and pit-like, rather than long and line-like in Dolerus picipes. Glossy patches beside the lateral postocellar furrows less distinct (almost no difference between the males). Thorax. (Fig. 17 D, E) The katepimeron of the mesopleuron (Fig. 17 D) with more numerous and distinct pits than in Dolerus picipes. The lateral mesoscutal lobes (Fig. 17 E) less distinctly and rather sparsely pitted. Abdomen. (Fig. 17 F-H) The ovipositor sheath in dorsal view broadens slightly towards the apex (Fig. 17 G). Cerci (Fig. 17 H) yellowish (black in Dolerus picipes, Fig. 17 I). Setae on valvula 3 rather short and delicate compared to most other Poodolerus species (Fig. 17 G, H). Setae on the abdominal terga shorter and sparser, except on the apical terga. The species can be easily differentiated from D. picipes as follows. 1 a Lateral postocellar furrows distinct: long and line-like b Minimal distance between the antennal sockets about 1.5 x the length of malar space c Ovipositor sheath in dorsal view strongly broadening towards apex, the apical setae well developed and clearly curved (Fig. 17 I) D. picipes (Klug, 1818) - aa Lateral postocellar furrows indistinct: short and pit-like (Fig. 17 C) bb Minimal distance between the antennal sockets about 2 x the length of malar space (Fig. 17 B) cc Ovipositor sheath in dorsal view slightly broadening towards the apex, the apical setae delicate, nearly straight or slightly curved (Fig. 17 G) D. coruscans Konow, 1890	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6FED188EF92E549296338AB6A1DCB807.taxon	distribution	Distribution. Dolerus coruscans is recorded from Austria (N-Burgenland, Ungerberg, Weiden am See, 1 ♀, 24.03.1957, leg. F. Kasy, SDEI), Germany (lectotype: see above), Hungary (" Budapest ", 1 ♀, SDEI. Haris 1998, 2003: " Budakeszi, Csepel, Latrany, Rakospalota "; Roller and Haris 2008: " Simontornya, Csevharaszt, Szecseny ", in total 14 ♂), and Greece (Nom. Grevena, Konitsa E 28 km Vasilitsa Ski Resort, 40.050 ° N, 21.050 ° E, 1780 m, 2 ♂, 11.05.2007, leg. S. M. Blank et al. / leg. M. Wei, SDEI. The records of D. coruscans from Austria and Greece are the first for these countries. The species apparently has a wide distribution, but is probably seldom collected because of its rarity and early flight period. Probably it has also sometimes remained unidentified, or has been misidentified. Possibly, the males of this species in collections could have been placed under D. picipes and its rather small females have perhaps been mistaken for dwarf females of D. liogaster / D. puncticollis (both species also have melanic color forms).	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
DB6A6BBD7A3C50A3B4A61B950A7F7BA8.taxon	description	Figs 11 A-D, 12 A, C, E, G-I	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
31AA3E3992E65E95BC92589CF688FD1B.taxon	description	Fig. 19	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
BCFC3E42126051FFBAD8C863C00CF548.taxon	description	Fig. 20 A-C	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6AD49FE0E2485E8996BBF91345E39D18.taxon	description	Fig. 16 A, C, E, H, K, N, Q	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
592652E4A2765C02B5A3C92AEEDBDBA9.taxon	description	Figs 13 A, B, 14 C, 14 F, 15	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A5AE91BDF997598D81218B845B442C3F.taxon	description	Fig. 21	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
59D2DF44FD3956259223EDD0DF6DF0B1.taxon	description	Figs 11 E, F, 12 B-D, F, H	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F2688D367F7C520AA61D5D2417D58EAB.taxon	description	Fig. 16 B, D, F, I, L, O, R	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
26DD1E9333C1525DB50EF40AF306C797.taxon	description	Fig. 20 D	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A169625F1E8A5801822B7CD36645130C.taxon	description	Figs 13 C, D, 14 A, 14 D, 14 E	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B3F952C45A855FEA97BFC72C85DFE1B5.taxon	description	Fig. 16 G, J, M, P, S	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F5EBCE8408025632806CB53D79897721.taxon	description	Fig. 22 A-E	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B9F8FBAE747D593F83B52006244A6DBE.taxon	description	Figs 36, 37	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.taxon	description	Description. Female. Figs 23, 24 A-C, 26 A, B Colour. Fig. 26 A, B. Black. Pale are: palpi, tegula, legs (except for arioli and extreme inner tip of metatibia); in fore wing base and apex of costa, apex of subcostal, base of stigma (Fig. 23 A), R 1, 1 A, 2 A + 3 A [color differences between veins of hind wing not clearly definable]; margins of median excision of abdominal tergum 1, medio-distal part of terga 2 - 4 (- 6) [pale area on tergum 3 is the largest of these], all sterna, more or less the downturned parts of terga 2 - 8, more or less hypopygium and ovipositor valvifer 2. Wing membranes hyaline. Head. Pedicel about as long as scape. Head in lateral view with widest point on temple about equal to length of pedicel (Fig. 23 E). Malar space slightly shorter than diameter of anterior ocellus. Lowest part of gena densely sculptured, without shining interspaces between the ill-defined pits (Fig. 23 E). Clypeus densely pitted, weakly shiny; anterior margin widely emarginate, to about 0.4 of its length (Fig. 23 D). Setae on upper head as long, or longer than, diameter of anterior ocellus (Fig. 23 E). Thorax. Whole pronotum except for small antero-ventral area dull, with sculpture (Fig. 23 H). Pits on upper mesepisternum large and well-defined (Fig. 23 H). Mesoscutellum with large, scattered pits anteriorly and laterally, interspaces shiny; medially and posteriorly nearly without pits and entirely unsculptured (Fig. 23 F). Anterior of mesoscutellar appendage densely pitted, with numerous setae, only small posterior area unpitted (Fig. 23 F). Abdomen. Anterior of tergum 1 densely setose. Terga 2 and 3 glabrous. Terga increasingly setose from tergum 4 to apex of abdomen. Weak surface sculpture on tergum 2 becoming stronger on more distal terga. Sawsheath in dorsal view broad, with blunt tip (Fig. 24 C). In lateral view exposed length of cercus about 0.5 x as long as exposed upper length of valvula 3 (Fig. 23 G). Lancet (Fig. 24 B, C): 13 - 14 serrulae; apical serrulae short and high. Body length: 4.5 - 8.0 mm Male. Figs 24 D, 26 C Similar to female, except for: red-brown antennal flagellum with basal two antennomeres more or less black; metatibiae completely pale; abdomen completely dark with small, obscure pale markings medially on terga 3 - 5. Penis valve: Fig. 24 D. Body length: 4.5 - 5.5 mm Variability: 5 - 6 flagellomeres, depending on whether or not the distal one is subdivided; the proportions of this antennomere are thus highly variable. Number of pits on upper mesepisternum variable. The scape and pedicel of females may be more or less pale.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.taxon	materials_examined	Other material. Finland: Nylandia, Vantaa, YKJ grid coordinates: 669: 37, 17.06.2021, 1 ♀, leg. Miikka Friman.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.taxon	etymology	Etymology. Named after Matti Viitasaari, who first recognised a very large female of this species as probably not belonging to Heptamelus ochroleucus or H. dahlbomi.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.taxon	biology_ecology	Biology. Many female specimens have been reared from larvae in Athyrium filix-femina (L.) Roth. (Fig. 25). Larvae found abundantly in 2021 in Matteucia struthiopteris (L.) Tod. in Kitee, Finland, were provisionally identified as H. viitasaarii by genetic comparison (identical to adults), as later confirmed when females started to emerge in 2022. The larvae feed inside the rachis, eating alternate portions nearly completely, or only partly (Fig. 25 B, C). This results internally in a " ladder-like " appearance, which is externally clearly visible, especially against the light (Fig. 25 A). This feeding habit is, however, possibly widespread in Heptamelus: at least H. dahlbomi larvae feed in the same way. Larvae of H. viitasaarii in M. struthiopteris regularly overwinter in a chamber made within the fertile fronds at the base of the stem. Strangely, no males have so far been reared, and the few males so far netted are all from a single locality (within an area of less than 1 km 2). It was observed that adults reared from plants of different size varied very much in body size, and that the smallest individuals had been feeding on the smallest plant, with correspondingly thin stalks.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.taxon	distribution	Distribution. Finland (North and South Karelia, Tavastia, and Helsinki area), Russia (Moscow). The record from Moscow was published by Vikberg (2017) as Heptamelus ochroleucus.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
AEF2BB47C0CE56318FD8399B02C100DD.taxon	description	Fig. 38	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
4A2976EA8ABB5F0791F772F7E338C1DF.taxon	description	Description. Antennal flagellomeres proportionately narrow; flagellomere 1 3.2 - 4.7 x as long as distal width; the basal flagellomeres not widening distally, and setae normal (not long and coarse as in Phymatocera). Outer orbit with pronounced groove behind nearly whole length of eye. Postgenal carina developed slightly below eye (clearly in P. sibiricola, but very weakly in P. gracilicornis). Posterior of mesoscutellum with some conspicuous pits. Epicnemium variably developed, e. g. present in P. sibiricola, absent in P. gracilicornis. Claws with a small to minute inner tooth. Stub of 2 A + 3 A (Astatus analis) of fore wings straight, curved towards anterior, or furcate at apex; hind wing with enclosed cell M.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
4A2976EA8ABB5F0791F772F7E338C1DF.taxon	diagnosis	Diagnosis. Phymatoceropsis can be distinguished from Rhadinoceraea by its proportionately narrower flagellomeres (flagellomere 1 3.2 - 4.7 x as long as distal width in Phymatoceropsis, 2.6 - 3.1 x as long as distal width in Rhadinoceraea); basal flagellomeres not distally widened (widened in Rhadinoceraea); outer orbit with pronounced groove behind whole length of eye (in Rhadinoceraea, if a groove is present, then this is behind only part of the eye. Phymatoceropsis differs from Lagonis in its mainly smooth mesepisternum (upper mesepisternum of Lagonis with numerous, large, crater-like pits). Phymatoceropsis can be separated from Paracharactus (based on North American species and the European P. hyalinus) by the mesoscutellum having at least a row of deep, well-defined pits on the posterior part (mesoscutellum entirely without pits in Paracharactus). Although in all examined specimens of Phymatoceropsis the stub of 2 A + 3 A of the fore wing is apically furcate (as is also usual in Rhadinoceraea), and in most specimens of various Nearctic Paracharactus and the European P. hyalinus (Konow, 1886) (Konow 1886 a) the apex of 2 A + 3 A is straight, in some individuals of Nearctic Paracharactus species, as already noted by Smith (1969), it is furcate / curved strongly upwards, e. g. in P. rudis (Norton, 1861).	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
7EC34ECDD8EF59C1A26469F2E7DF75C2.taxon	description	Figs 28, 29	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
7EC34ECDD8EF59C1A26469F2E7DF75C2.taxon	biology_ecology	Biology. Oviposition trial by Vikberg (3 / 2011 VV): three females captured in Hangastenmaeki in 2011 were used in this experiment. Adoxa, Sambucus racemosa, Lonicera xylosteum and Galium sp. were offered to them. They laid eggs only on Sambucus racemosa leaves. The egg is laid through the upperside of the leaf into the tissue of the underside, usually one to two eggs per leaflet, but in one small leaflet eight eggs were counted. Oviposition occurred on 21.05. - 24.05. The first larvae were observed on the morning of 28.05. Later, 28 larvae were counted. Five feeding instars were observed and after finishing feeding there was an " extra moult ". Prepupae were seen on 07.06 - 08.06. 1 st and 2 nd instar larvae are gray, with a brown head. 5 th instar larvae are 13 - 14 mm long; head width ca. 1.6 mm (Fig. 29). Head brownish, behind the eye a black fleck which is curved backwards and upwards. Body dorsally dark gray with greenish hue; ventrally whitish gray. Black suprastigmal flecks on thoracic segment 3 and abdominal segments 1 - 9.	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
855A00D5977850258AAB3FF0FBC60D02.taxon	description	Fig. 39	en	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
