identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7FEFCA6AC53D5FEEAF4B7F011B9C410F.text	7FEFCA6AC53D5FEEAF4B7F011B9C410F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abia brevicornis Leach 1817	<div><p>Abia brevicornis Leach, 1817 nom. rev.</p><p>Abia brevicornis Leach, 1817: 114. Sex not stated [but probably female, because conspicuous dark dorsal patches on abdomen are not mentioned]. Syntypes (assumed). Type locality: not stated. Type material probably lost or destroyed.</p><p>Cimbex splendida Klug, 1820 [incorrect original spelling]: 98-99. ♀, ♂. Syntypes. Type locality: Germany; rare in this area [around Berlin]. Syntype ♂ [examined]:  “GBIF-GISHym2903”,  “13567”,  “Germania”, "nitens L. Soldanski det.". ZMHB.</p><p>Abia rossica Semenov, 1896: 159, 167-168. ♀. Holotype [not examined]. Type locality: Ukraine, Kamjanez-Podilskyj.</p><p>Abia nitens auct. nec Linnaeus. Misidentification of  Tenthredo nitens by, for example: de Dalla Torre (1894), Konow (1905b), Taeger (1998), Liston and  Späth (2006).</p><p>Notes.</p><p>Taeger et al. (2010) also listed  Abia nitens var. vernetensis Pic, 1928 (type locality: France, Allier) as a junior synonym of  Abia nitens auct. However, no type specimen has been examined subsequent to its description, as far as we are aware. Pic wrote [translated from French] "differs from the typical form in the black-marked base of the posterior femora". This suggests that it possibly does not belong to  A. brevicornis .</p></div>	https://treatment.plazi.org/id/7FEFCA6AC53D5FEEAF4B7F011B9C410F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
46786109F6CB53C790750E4FF34E9B93.text	46786109F6CB53C790750E4FF34E9B93.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Abia nitens (Linnaeus 1758)	<div><p>Abia nitens (Linnaeus, 1758)</p><p>Tenthredo nitens Linnaeus, 1758: 556. ♂ [because the conspicuous dark dorsal patches on abdomen are mentioned]. Syntypes (assumed). Type locality: Europe. Lectotype designated by Malaise and Benson (1934). LSUK. Images of the lectotype (LINN 2402) in dorsal and lateral view are available (The Linnean Society of London 2022).</p><p>Tenthredo sericea Linnaeus, 1767: 921. Sex not stated [but probably female, because dark dorsal patches on abdomen are not mentioned]. Syntypes (assumed). Type locality: Leipzig. syn. nov.</p><p>Abia sericea: de Dalla Torre (1894), Konow (1905b), Enslin (1917), Liston and  Späth (2006), Taeger et al. (2010).</p><p>Abia dorsalis Costa, 1859: 5-6. ♀. Holotype [not examined]. Type locality: Italy, Cape Miseno near Naples. syn. nov.</p><p>Notes.</p><p>Abia nitens was successively mentioned as occurring in Sweden by  Fallén (1807), Dahlbom (1836), and Thomson (1871), who all placed  Tenthredo sericea as its synonym.  Thomson’s description of leg color indicates that he had before him specimens of the species called by Taeger (1998)  A. sericea, or perhaps  A. candens [not distinguished from the former in  Thomson’s time]. Influenced by the continued, widespread use of the name  Abia nitens in Scandinavian literature, Taeger et al. (2006) and ArtDatabanken (2015) included Sweden within the range of  A. brevicornis [as  A. nitens auct.]. In fact, no Swedish specimens of  A. brevicornis have been located in the MZLU or NHRS collections, and there are no published records from other Fennoscandian countries.  Abia brevicornis is restricted to extremely dry, summer-warm sites, where its larval hosts occur ( Scabiosa spp.) (Liston and  Späth 2006). Probably it has a strongly continental distribution: its most north-westerly known localities in Europe are in Central Germany, whereas it is not definitely known in France (Noblecourt 2020), and a single old record from Spain needs confirmation. Although a presence of  A. brevicornis in southern Sweden cannot be ruled out, we consider it likely that all references to  A. nitens auct. in Sweden relate to either  A. nitens [=  Abia sericea] or  A. candens .</p></div>	https://treatment.plazi.org/id/46786109F6CB53C790750E4FF34E9B93	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B23797F1FBA45A4FACFAD4F70C2B6C95.text	B23797F1FBA45A4FACFAD4F70C2B6C95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calameuta punctata (Klug 1803)	<div><p>Calameuta punctata (Klug, 1803)</p><p>Figs 3, 4</p><p>Astatus punctatus Klug, 1803: 55, plate VII figs 2a, b. ♀. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of  Klug’s work]. Neotype designated below.</p><p>Calameuta punctata: Zombori (1978).</p><p>Astatus floralis Klug, 1803: 53-54, plate VI figs 5a, b. ♂. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of  Klug’s work]. syn. nov.</p><p>Astatus analis Klug, 1803: 54-55, plate VII fig. 1. ♀. Syntypes (assumed). The type material is considered to be lost or destroyed. Published type locality: Germany [implicit from title of  Klug’s work]. syn. nov.</p><p>Cephus filum Gussakovskij, 1935: 112, 358, 361. ♀, ♂. Syntypes. Type locality: Sarepta, Caucasus, and southern Siberia. ZIN. syn. nov.</p><p>Calameuta filum: Benson (1946).</p><p>Type material examined and taxonomic notes.</p><p>To help resolve the taxonomic disagreements in the interpretation of these nominal taxa, and promote the future stability of nomenclature, a neotype is designated for  Astatus punctatus:</p><p>Astatus punctatus Klug, 1803. Neotype ♀ (DEI-GISHym21255, Fig. 4A-D), hereby designated. Germany, Brandenburg, Landkreis  Märkisch-Oderland,  Müncheberg, Trebnitz, 52.535°N, 14.204°E, damp meadow, swept from  Alopecurus pratensis, 16.05.2015, leg. A. Liston (deposited in the SDEI). Labelling [printed on pale paper if not stated otherwise]: "Germany: Brandenburg; Landkreis  Märkisch-Oderland,  Müncheberg Trebnitz 16.05.2015 leg. A. D. Liston",  “21255” [handwritten] with part of a leg gummed to card],  “DEI-GISHym21255”, "♀  Calameuta punctata (Klug) [handwritten] det. A. Liston 2018", "NEOTYPE ♀  Astatus punctatus Klug, 1803 designated A. Liston 2022" [red].  Klug’s description states that abdominal segment 4 has obscure, paired dorsal spots; segment 5 four separate yellow spots, one pair laterally, the other dorsally; segments 6 and 7 with spots on their lower posterior margins; segment 8 immaculate; segment 9 completely yellow. Accordingly, we selected as neotype a specimen with small pale markings on terga 4-7 as well as 8-10 (Fig. 4A-D). The abdomen of a second female (DEI-GISHym21260) collected at the same place and time has fewer and less extensive pale markings and is thus intermediate in this respect to other female  C. punctata specimens collected in Germany and all known Finnish and Estonian specimens, which have a completely black abdomen apart from terga 8-10 (Fig. 3A, C).</p><p>Astatus floralis and  A. analis have in the past generally been treated as synonyms of  Calameuta haemorrhoidalis auct. [our  Calameuta variabilis], e.g. by Konow (1905a). The opinion that  A. floralis is a synonym of  Cephus pygmeus (Linnaeus, 1767), as in de Dalla Torre (1894), cannot be accepted:  Klug’s description of leg colour does not fit  C. pygmeus . Our reason for placing  A. floralis and  A. analis as synonyms of  Calameuta punctata rather than of  C. variabilis is based primarily on one of the main characters which distinguishes  C. punctata from  C. variabilis: the structure of the maxillary palps. In the description of  Astatus which precedes the descriptions of  A. floralis,  A. analis and  A. punctatus, Klug characterized the genus thus: "Palpi[...]anteriores[...]sexarticulati, articulis duobus baseos cylindricis, aequalibus, tertio crassiori, longiori, subcylindrico, quarto longissimo, graciliori, quinto brevissimo, ultimo longitudine fere tertii subulato[...]". The described proportions of maxillary palpomeres 5 and 6 therefore fit  C. punctata (Fig. 3D), not  C. variabilis (Fig. 5B).  Calameuta variabilis is unique in  Calameuta in having maxillary palpomeres 5 and 6 of almost equal length (Gussakovskij 1935; Benson 1968; Zombori 1978). Zombori (1978) correctly identified  Calameuta variabilis [which he called  C. haemorrhoidalis] as a taxon distinct from  C. punctata, and summarized the characters that distinguish them, but interpreted some of the names wrongly. Notably, Zombori (1978) did not mention the major contradiction in the morphology of the maxillary palps, as described by Klug, when he tentatively suggested that  A. floralis and  A. analis might be synonyms of  Astatus haemorrhoidalis auct. ["the description of the latter two [ Astatus floralis,  Astatus analis] rather corresponds to the one given by Fabricius for  C. haemorrhoidalis, accordingly, they are considered as synonyms of the latter name."].  Zombori’s main reason for doubting that  Astatus analis was synonymous with  Calameuta punctata, seems to have been the wording of  Klug’s descriptions, which suggested that the thorax of  Astatus analis is shinier than that of  Calameuta punctata . Apart from this,  Klug’s description of  A. analis fits the darker forms within the rather wide range of variability in the female sex of  C. punctata . Strangely, in his discussion of these names, Zombori (1978) does not mention  Calameuta filum at all.</p><p>The explicit collection data given by Klug (1803) for  Astatus floralis ("Locus in editioribus argillosis; in floribus"),  A. analis ("Locus in editioribus; in floribus"), and  A. punctatus ("Locus in floribus") are, in part, not easy to interpret. Clearly, "in floribus" means that the specimens were collected from flowers. We think that "in editioribus argillosis" refers simply to the type of locality, i.e. an elevated place on clayey ground. This fits well with the type of sites at which  C. punctata has recently been collected in Germany (see below).</p><p>The synonymy of  Calameuta filum with  C. punctata can be proposed with a high degree of confidence. The characters described by Gussakovskij (1935) for the former are precisely those used by Zombori (1978) to characterize the latter. The same characters are also given by Viitasaari (1975) in his description of Finnish specimens identified as  Calameuta filum, and which he compared with a syntype of that species. Viitasaari (1984) subsequently noted that  Calameuta punctata sensu Zombori (1978) and  C. filum are probably conspecific.</p><p>Biology and distribution.</p><p>The only recorded host plant of  Calameuta punctata is  Alopecurus pratensis L. (Vikberg 1978; Liston 2015), on which it is possibly monophagous. Accordingly,  C. punctata occurs mostly in rather moist places. Its wider geographic range is not entirely clear, particularly because the identity of  Calameuta pravei (Dovnar-Zapolskij, 1926) remains unresolved. This has been considered to be a valid species (e.g. Llorente and Gayubo 1990), or a synonym of  C. punctata (e.g. Taeger et al. 2010).  Calameuta pravei was recorded by Gussakovskij (1935) from Transcaucasia, Crimea, and the western Kopet-Dagh (Turkmenia). Llorente and Gayubo (1990) added records from Spain. Excluding these records,  C. punctata is known from south-east Russia, Transcaucasia and south Siberia to Irkutsk (Gussakovskij 1935, as  Cephus filum), and central and northern Europe (specimens examined by us). The distribution of  C. punctata is therefore rather different from that of  C. variabilis (see below), but their ranges overlap at least in parts of south-central Europe.</p><p>Calameuta variabilis [as  C. haemorrhoidalis auct.] has been stated to occur in Germany based on the mention of Germany as the type locality in the original descriptions by Fabricius (1781) of  Tenthredo haemorrhoidalis and by Klug (1803) of  Astatus punctatus,  A. floralis and  A. analis, coupled to an apparently faulty understanding of which taxa are represented by these names. Blank et al. (1998, 2001) listed  C. haemorrhoidalis from Germany, only for Berlin-Brandenburg, dating the record respectively as "vor 1803" and  “1802” . This refers to the type material of  Klug’s species. Later, Liston et al. (2012) treated  C. haemorrhoidalis auct. [ C. variabilis] as extinct in Germany, and added  C. punctata to the German list, based on recently collected specimens. Although it cannot be ruled out that  C. variabilis once occurred in Germany, but has since disappeared, we think it more likely that in historical times only  C. punctata ever occurred there, and propose in future to include only it in the list of German  Symphyta . Fennoscandian and Estonian specimens which were previously identified as  C. filum also belong to  C. punctata .</p><p>Based on COI sequences,  C. punctata is split into two barcode clusters. Three specimens from Finland and one from Estonia are identical (BOLD:ACQ7596), but differ from two German specimens by 5.0-5.5% (no BIN assigned yet, GenBank accessions MW353981 and MW353982). The BOLD:ACQ7596 is closer to  C. pallipes, differing by a minimum of 4.1%.</p></div>	https://treatment.plazi.org/id/B23797F1FBA45A4FACFAD4F70C2B6C95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A24CA63B1BF65A26B698AB9EEDE40F68.text	A24CA63B1BF65A26B698AB9EEDE40F68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calameuta variabilis (Mocsary 1886)	<div><p>Calameuta variabilis (Mocsary, 1886)
 comb. nov.</p><p>Fig. 5</p><p>Cephus quadriguttatus Costa, 1882: 198. ♂. Syntypes. Type locality: Sardinia. Synonymy with  C. haemorrhoidalis auct. by Ghigi (1905). Primary homonym of  Cephus quadriguttatus Westwood, 1874. syn. nov.</p><p>Cephus variabilis Mocsáry, 1886a: 101, 103. ♀, ♂. Syntypes. Type locality:  Kecskemétnél,  Herkulesfürdõ,  Nagyváradnál,  Szilágymegyében Tasnádon,  Tasnád-Szántó, and S.-A.-  Ujhelynél ( Mocsáry 1886b). Lectotype, ♀, designated below.</p><p>Cephus quadriguttulatus Dalla Torre, 1894: 412. Replacement name for C.  Cephus quadriguttatus Costa. syn. nov.</p><p>Cephus quadrisignatus Costa, 1894: 252. Replacement name for C.  Cephus quadriguttatus Costa. syn. nov.</p><p>Cephus haemorrhoidalis var. signifer Konow, 1896b: 317-318. ♀. Holotype. Type locality: Syria, Akbes. syn. nov.</p><p>Cephus pseudotabidus Kokujev, 1910: 136-137. ♀, ♂. Syntypes. Type locality: Transcaspia. Synonymy with  C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov.</p><p>Cephus diversipes Ghigi, 1915: 308-309. ♂. Holotype. Type locality: Greece, Rhodos. Synonymised with  C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov.</p><p>Trachelus syriacus Pic, 1917: 1-2. ♀. Syntypes. Type locality: Syria, Alexandrette. Synonymised with  C. haemorrhoidalis auct. by Gussakovskij (1935). syn. nov.</p><p>Calameuta haemorrhoidalis: Gussakovskij (1935), Benson (1946), and most subsequent authors.</p><p>Type material examined and taxonomic notes.</p><p>Lectotype of  Cephus variabilis Mocsáry, 1886, hereby designated: ♀, id nr.017651 HNHM Hym.coll. (Fig. 5A); labels (Fig. 5A). Type locality: Romania,  Herkulesfürdõ (HNHM). Paralectotypes (all HNHM): 5♂ and 2♀ also belong to the type series and have been labelled as paralectotypes: details of their sexes and localities were given by Zombori (1978). Note that the type series is heterogeneous, and contains specimens of  Calameuta punctata as well as  C. variabilis . Zombori noted the heterogeneity of the type series, but did not publish a lectotype designation, although the specimens were labelled by him as lectotype and paralectotypes. The types were found by Z. Vas in the HNHM, grouped as stated by Zombori. We do not follow  Zombori’s intention according to his labels, but designate the female from  Herculesfürdö (= Baile Herculane, Romania) (id nr. 017651 HNHM Hym. coll., labelled by Z. Vas) as lectotype (Fig. 5A). Thus,  Calameuta variabilis ( Mocsáry, 1886) comb. nov. can be used as a valid name for  Calameuta haemorrhoidalis sensu Gussakovskij et auct.</p><p>Cephus atripes Stephens, 1835 has sometimes been listed as a synonym of  C. variabilis, e,g. by Taeger et al. (2010, under  Calameuta haemorrhoidalis). The type specimen, or specimens, is probably lost. The description is short, and does not state the sex of the described specimens(s). de Dalla Torre (1894) treated  C. atripes as a synonym of  C. pygmeus (Linnaeus, 1767) and Konow (1905a) as a synonym of  C. haemorrhoidalis . The color pattern described by Stephens does not fit very well with either of these, in either sex, nor with any other known north-west European cephid species.  Stephens’ name is best treated as a species inquirendae.</p><p>Biology and distribution.</p><p>According to Macek et al. (2020, under  Calameuta haemorrhoidalis) the host plants of  C. variabilis are various  Poaceae, including cereals such as rye ( Secale cereale) and wheat ( Triticum). According to our personal experiences, adults occur in dry places, mostly on or near wild grasses. The taxon to which the name  C. variabilis is now applied has so far been found only in the West Palaearctic, and has an essentially Mediterranean distribution, summarized by Gussakovskij (1935) as comprising southern Europe, Crimea, Caucasus, western Turkmenia (Kopet-Dagh) and Syria. In Central Europe, it reaches at least as far north as Hungary (Zombori 1978), but specimens recorded from Austria and the Czech Republic (see Taeger et al. 2006) should be checked.</p></div>	https://treatment.plazi.org/id/A24CA63B1BF65A26B698AB9EEDE40F68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
9495D1C86C0C56AF873890DF3FDD581E.text	9495D1C86C0C56AF873890DF3FDD581E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalcia arvensis Panzer 1802	<div><p>Cephalcia arvensis Panzer, 1802</p><p>Fig. 6</p><p>Cephalcia arvensis Panzer, 1802: vol. 86 pl. 9. ♀. Syntypes. Type locality: Germany [according to title of work]. Blank et al. (2009); Nomen protectum with respect to  Psen lucorum Schrank, 1802, nomen oblitum.</p><p>Cephalcia abietis var. intermedia Hellén, 1948: 40-41. ♀. Syntypes. Type locality: Russia, Karelia,  Paanajärvi .</p><p>Cephalcia intermedia: Vikberg (1982); raised to species rank. Van Achterberg and van Aartsen (1986); synonymy with  C. lariciphila (Wachtl, 1898). syn. nov.</p><p>Notes.</p><p>Cephalcia intermedia has in recent years mostly been understood as a distinct species close to  C. arvensis (e.g. Viitasaari 2002b, Taeger et al. 2006), but with more extensively dark-patterned adults, occurring in northern Europe and the central European mountains. Because of its dark coloration, specimens of  C. intermedia can also be mixed up with  C. lariciphila (Wachtl, 1898), but the host plant of  C. intermedia is  Picea, as in  C. arvensis, not  Larix as in  C. lariciphila . The status of  C. intermedia and the supposed differences to  C. arvensis were discussed by Vikberg (1982), Shinohara (1985), Midtgaard (1987) and Viitasaari (2002b). The only differences between  C. arvensis and  C. intermedia are in coloration: mainly of the abdomen, and less so of the antennae and legs. Shinohara (1985) pointed out that an unbroken range of color variability occurs between pale  C. arvensis and the dark specimens identified as  C. intermedia. This is in accordance with our observations, as the amount of dark color on the male abdomen varies considerably even in a single locality (Fig. 6). Small differences in colouration of larvae are possibly not constant, as the larvae of  C. intermedia used for comparison were offspring of a single female. At present, there seems to be no convincing evidence for treating  C. intermedia as a species distinct from  C. arvensis, and we therefore synonymize them.</p></div>	https://treatment.plazi.org/id/9495D1C86C0C56AF873890DF3FDD581E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5FD5309B01BD5F0EB98A5474CED58AEB.text	5FD5309B01BD5F0EB98A5474CED58AEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Claremontia brevicornis (Brischke 1883)	<div><p>Claremontia brevicornis (Brischke, 1883)</p><p>Fig. 7B, C</p><p>Blennocampa brevicornis Brischke, 1883: 282-283. ♀. Syntypes (assumed). Type locality: Poland [ “Westpreußen” = western Prussia]. Types probably lost (Blank and Taeger 1998).</p><p>Blennocampa puncticeps Konow, 1886a: 215-216. ♀, ♂. Syntypes. Type locality: Switzerland,  Zürich . Lectotype ♀ (SDEI) designated by Koch (1988). Treated by Koch (1988) as a synonym of  Claremontia confusa .</p><p>Monophadnoides puncticeps: Benson, 1952; treated as species distinct from  M. confusa, and  Poterium sanguisorba [currently  Sanguisorba minor] recorded as host of larva. Chambers (1961):  Potentilla reptans recorded as host of larva.</p><p>Biology.</p><p>ZMUO specimens, from Finland, are not reared;  Sanguisorba, recorded as a host plant by Benson (1952), can be excluded as a possible host plant at these localities, but  Potentilla erecta is present in abundance at one of the localities in Eastern Finland.</p><p>Taxonomic notes.</p><p>Konow (1886a), in a key, described the tibiae of  Claremontia confusa (as  Blennocampa confusa) as mainly black with only the knees very narrowly whitish and the protibia only whitish on the anterior face, and wrote that the tibiae of  C. puncticeps were mainly pale. Benson (1952) and Lacourt (2020), among others, distinguished  Claremontia brevicornis (as  Claremontia confusa) from  C. puncticeps using several characters, also including the color of the metatibia of females. However, Brischke (1883) described the metatibia of  Claremontia brevicornis, a nominal species not known to Konow (1886a), as extensively yellow-white. Numerous barcoded females (SDEI, ZMUO) belong to two separate COI sequence clusters (distance 3.1-5.8%), which correlate fully with their leg color. Accordingly, we think that the original description of  C. brevicornis refers to what has more recently come to be known as  C. puncticeps, and that the correct name for the other species is  C. confusa . Unlike some previous authors, we did not detect a clear difference in the sculpture of the head of the two forms. Distinction of the males is problematic, because of the lack of reliably identified male specimens of  C. confusa . Benson (1952) stated that  C. confusa "is entirely parthenogenetic, at least in Britain". On the other hand, several male specimens of  C. brevicornis (Fig. 7C) have been barcoded, and can thus definitely be associated with that species. Based mainly on barcode-sequenced specimens in ZMUO, females of  Claremontia confusa and  C. brevicornis may be separated as follows.</p><table><tr><td colspan="1" rowspan="1">1</td><td colspan="1" rowspan="1">a Length of antenna subequal to length of costa (Fig. 7A) b Metatibia completely black, or with base narrowly pale (Fig. 7A) [More robust body shape and slightly darker wings]</td><td colspan="1" rowspan="1">C. confusa (Konow, 1886)</td></tr><tr><td colspan="1" rowspan="1">-</td><td colspan="1" rowspan="1">aa Antenna approximately 0.65-0.80 as long as costa (Fig. 7B) bb Metatibia usually extensively pale, with at least base whitish (Fig. 7B) [More slender body shape and slightly paler wings]</td><td colspan="1" rowspan="1">C. brevicornis (Brischke, 1883)</td></tr></table></div>	https://treatment.plazi.org/id/5FD5309B01BD5F0EB98A5474CED58AEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
742BFB48E2B15B8B8EED1F77953D42A2.text	742BFB48E2B15B8B8EED1F77953D42A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Claremontia confusa (Konow 1886)	<div><p>Claremontia confusa (Konow, 1886) sp. rev.</p><p>Fig. 7A</p><p>Blennocampa confusa Konow, 1886b: 82. ♀. Type locality: Germany. Neotype ♀ (ZMHB) designated by Koch (1988). Synonymy with  Claremontia brevicornis proposed by Koch (1988), wherein he overlooked the priority of the name  Claremontia brevicornis (Blank and Taeger 1998).  Monophadnoides confusa: Benson (1952); treated as species distinct from  M. puncticeps, cultivated  Fragaria species recorded as hosts of larva.</p><p>Biology.</p><p>Substantiating the statement about the host plant by Benson (1952), the ZMUO specimens, collected in the Helsinki area, were reared from larvae feeding on cultivated  Fragaria .</p></div>	https://treatment.plazi.org/id/742BFB48E2B15B8B8EED1F77953D42A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
84F088DC0D7D54A0B628FC52A6CF4357.text	84F088DC0D7D54A0B628FC52A6CF4357.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus aericeps Thomson 1871	<div><p>Dolerus aericeps Thomson, 1871</p><p>Dolerus aericeps Thomson, 1871: 285. ♀, ♂. Syntypes. Published type locality:  “södra Sverige" [southern Sweden]. Lectotype designated below.</p><p>Tenthredo Eglanteriae [sic!] Fabricius, 1793: 109-110. syn. nov., nomen oblitum. ♀. Syntypes (assumed). Published type locality: "Habitat in Germania. Dom. Smidt." [Germany]. Lectotype designated below.</p><p>Tenthredo (Dolerus) palustris Klug, 1818: 296-297. ♀, ♂. Syntypes. Published type locality:  “Deutschland” . Lectotype designated below. Primary homonym of  Tenthredo (Allantus) palustris Klug, 1818. syn. nov.</p><p>Dolerus bajulus Serville, 1823: 58. ♀. Syntypes (assumed). Type locality:  “Paris” . Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus aericeps Thomson, 1871. Lectotype ♀ hereby designated, labelled:  “Båst[ad]” [Type locality: Sweden,  Skåne,  Båstad], "  Dolerus aericeps " [blue line along upper margin], " MZLU 2013 416" [pale green, printed, loan record], "  Lectotypus 2014  Dolerus aericeps ♀ Thomson, 1871 M.Heidemaa design." [printed, red], "  Dolerus aericeps Thomson, 1871 M.Heidemaa det.". In excellent condition. MZLU. Paralectotypes: 2♀, 2♂. MZLU.</p><p>Tenthredo eglanteriae Fabricius, 1793. Lectotype ♀ hereby designated, labelled: " eglan [...] riae ", "LECTOTYPE  Tenthredo eglanteriae Fabricius, 1793 des. SMBlank 2008" [red, printed], "  Dolerus eglanteriae (F.) =  Dolerus bajulus Serv. [handwritten] det. S.M.Blank 2008 ✓" [printed], " ZMUC-GISHym 1014", "  Dolerus aericeps Thomson, 1871 M.Heidemaa det." [printed]. Left hind tarsus and some other tarsomeres missing. ZMUC.</p><p>Among 2♀ and 1♂ specimens preserved in the ZMUC collection under the name  T. eglanteriae, only the above female can be regarded as a syntype. The scutellum of the other female is red, and the tip of the abdomen of the male black. Both characters conflict with  Fabricius’ original description. Compared with the other two specimens, the pale parts of the legs of the lectotype are darkened. The lectotype agrees with the species which has for a long time been known as  Dolerus aericeps Thomson, 1871 (Zhelochovtsev 1994), characterized for example by the laterally directed and distally curved setae of the ovipositor sheath. Lacourt (2000) used the name  D. bajulus Serville, 1823 for this species, but Blank et al. (2009) presented arguments for the use of the name  D. aericeps (nomen protectum) versus  D. bajulus (nomen oblitum).  Tenthredo eglanteriae was synonymized with  T. germanica by Klug (1819) and with  Athalia glabricollis Thomson, 1870 by Konow (1897a). Since  Brullé (1846),  T. eglanteriae (nomen oblitum) has never again been used as valid, while between 1971-2021 the name  Dolerus aericeps (nomen protectum) was used as valid by more than 120 authors in over 150 publications (Article 23.9.1, ICZN 1999). One paralectotype male of  D. cothurnatus Serville belongs to this species (see under  D. germanicus).</p><p>Tenthredo (Dolerus) palustris Klug, 1818. Lectotype ♂ hereby designated, labelled:  “14190” [collection catalog no.], "cotypus von  Dol. palustris Klg. =  Dolerus aericeps Thoms. O.Conde det 1941" [handwritten by Otto Conde], "GBIF-GISHym 2342" [printed], "  Lectotypus [printed] ♂  Tenthredo (Dolerus) palustris Klug, 1818 [handwritten in block letters] M.Heidemaa des. [printed]" [red], "  Dolerus aericeps Thomson, 1871 M.Heidemaa det.", "Zool. Mus. Berlin" [printed]. Left forewing and metatarsi missing, half of the genital capsule glued on a paper card and pinned with the lectotype. Paralectotype ♂ ("GBIF-GISHym 2341") with same labelling (except "  Paralectotypus "). All in ZMHB.</p><p>Formerly placed as a synonym of  D. cothurnatus auct. (=  D. junci (Stephens, 1835)), e.g. by Taeger et al. (2010).</p><p>Dolerus bajulus Serville, 1823. Lectotype ♀ hereby designated, labelled: "  Dolerus bajulus, Lep.", "Lectotype [red, printed in block letters]", "Lectotype  D. bajulus LEP. det.: H.Haris 1996" [white], "  D. aericeps THOMS. ♀ det.: A.Haris", "  Lectotypus [printed] ♀[handwritten]  Dolerus bajulus Serville, 1823 [handwritten in block letters] M.Heidemaa design." [printed, red]. "  Dolerus aericeps Thomson, 1871 M.Heidemaa det" [printed]. MNHN.</p></div>	https://treatment.plazi.org/id/84F088DC0D7D54A0B628FC52A6CF4357	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5821C75CF43A5312BEC6D962BB6CEBDE.text	5821C75CF43A5312BEC6D962BB6CEBDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus aericepsellus Heidemaa & Mutanen 2022	<div><p>Dolerus aericepsellus Heidemaa &amp; Mutanen sp. nov.</p><p>Figs 8, 9</p><p>Description.</p><p>Holotype ♀. Figs 8A, B, 9B, C.</p><p>Colour. Fig. 8A, B. Body black with terga 2-7 and sterna 2-7 orange. Terga 8-9 brownish black, 10 brownish basally but yellowish brown apically, apical third of protibiae brownish. Ventral margin of valvula 3 and cerci brownish. Wings clear.</p><p>Measurements (mm). Body length: 5.0, distance from tegula to base of pterostigma: 3.1, head breadths: 1.46 (max. at eyes), 1.28 (max. behind eyes), 1.07 (min. behind eyes), head length (behind eyes): 0.21, max diameter of eye: 0.62, breadth of postocellar field: 0.36, OC: 0.96, OOCL: 0.35, OOL: 0.20, POL: 0.19. Length of: metafemur: 1.18, metatibia: 1.76, ovipositor sheath (from the base of valvifer 2 to the apex of valvula 3): 1.01. Length of flagellomeres: 1: 0.48, 2: 0.41, 3: 0.37, 4: 0.31, 5: 0.29, 6: 0.26, 7: 0.25.</p><p>Large structures. Clypeal emargination almost half as deep as clypeal median length. Clypeus asymmetrical, its antero-lateral lobes round at apex, the left lobe longer than the right. Distance between antennal sockets 1.6  × as long as the malar space. Length of antennomere 3 about 1.24  × that of 4. Oblique furrow not outlined. Distance between cenchri about 1.1  × as long as width of cenchrus. Metatarsomere 1 about 1.75  × as long as 2. Mesoscutellar appendage with weakly outlined ridge medially.</p><p>Setae. Metascutellum with few setae (about 5). Abdomen dorsally mostly pubescent from segment 7 to last one (laterally and ventrally from 2 to last one). Longest setae of valvula 3 in dorsal view markedly curved in apical third, and forming an angle of about 80-90° (Fig. 9B).</p><p>Macrosculpture. Pits on head rather irregular in size and distribution, more distinct and separated on postocular and postocellar area. Density and size of pits on thorax vary, the largest (partly fused) on lateral sides of median mesoscutal lobes and on mesoscutellum. Pits on lateral lobes of mesoscutellum and near the median mesoscutal groove sparse and small. Pits on mesepisternum largest, partly isolated and round, partly polygonal and fused (Fig. 9C). Distinct pits on pectus nearly absent (Fig. 9C).</p><p>Microsculpture. Meshes on mesoscutellar appendage extensive, on tergum 1 absent, on metepimeron (Fig. 9C) and on ventral surface of metacoxa absent. Sculpticells on anterior half of katepimeron of mesopleuron rib-like, irregular and of uneven height, and absent on metepimeron, pectus, and terga (all glossy).</p><p>Male. Figs 8C, D, 9A, D-G. Closely resembles the female, but the anterior margin of tergum 2 blackish, basal 2/3 of hind tibia (with apical spurs), and base of metatarsomere 1 reddish yellow (Fig. 8C, D). Penis valve valviceps long and narrow (Fig. 9G).</p><p>Measurements (mm). Body length (paratype male ZMUO.035740): 5.3, distance from tegula to base of pterostigma: 3.2, max. diameter of eye: 0.60, head breadth: 1.47 (max. at eyes), breath of postocellar field: 0.38, head length (behind eyes): 0.24, OC: 1.01, OOCL: 0.18, OOL: 0.29, POL: 0.21, metafemur: 1.25, metatibia: 1.64, metatarsus: 0.50. Length of flagellomeres: 1: 0.59, 2: 0.51, 3: 0.50, 4: 0.44, 5: 0.41, 6: 0.39, 7: 0.41.</p><p>Genetic data.</p><p>Three specimens with DNA barcode sequence data are available. They comprise a distinct cluster BIN BOLD:ABV8002, with 0.3% intraspecific variability. Based on sequences longer than 600 bp, the minimum distance to the closest relative  D. aericeps (n=26) is 3.9%, but a few specimens identified as  D. yukonensis from North America show a minimum distance of even less, 3.6%.</p><p>Differential diagnosis.</p><p>Adults of  D. aericepsellus are most similar to  D. aericeps Thomson, but are distinguished by the following characters: smaller body size (similar to  D. elderi Kincaid, 1900, or smaller); very short, strongly narrowing postocular area; glossy metascutellum with few distinct pits and setae (3-6); pectus without distinct pits; and thorax black, including tegulae (based on the barcoded specimens: 1♀ 2♂). Lateral postocellar furrows very distinct in both sexes (Fig. 9F), like in  D. aericeps and  D. incisus . The thorax of female  D. aericeps is usually largely red, whereas in the  D. aericepsellus paratype female it is almost entirely black.  Dolerus aericeps ab. nigricollis was described by Lindqvist (1943) from northern Finland close to the localities where  D. aericepsellus was collected. We examined the type specimen of  Dolerus aericeps ab. nigricollis deposited in the MZH and found that it is not conspecific with  D. aericepsellus but with  D. aericeps, although it has a nearly completely black thorax, with two obscure reddish flecks. Some other examined specimens of  D. aericeps from northern Finland show reduced red markings on the thorax, suggesting that this character varies in  D. aericeps .</p><p>Type material.</p><p>Holotype ♀. Type locality: Finland,  Enontekiö, Pousu. Labelled [white, printed if not given otherwise]: "FIN Le  Enontekiö Pousu 7651:3266 [68.846°N, 21.197°E] 27.6.2020 Marko Mutanen leg.", "Sawfly tissue 2020-1079 M.Mutanen" [greenish],  “http://id.zmuo.oulu/fi/ZMUO.044566”, "Holotypus [block letters] ♀  Dolerus aericepsellus Heidemaa &amp; Mutanen, 2022 [handwritten in block letters] des. M. Heidemaa" [red]. Good condition: wings spread, left fore and mid legs removed as tissue samples. ZMUO.</p><p>Paratypes. Total: 1 ♀, 3 ♂: Finland: Lapponia inarensis, Utsjoki  Pulmankijärvi, YKJ grid coordinates: 7761:3539 [69.928°N, 28.031°E], 1♂ (ZMUO.033245) 07.07.2017, leg. M. Mutanen, N. Mutanen, A. Mutanen, SDEI. 1♂ (ZMUO.035740) (Fig. 9A, D-G), same collection data as preceding, but 05.07.2018, ZMUO. Lapin  Lääni, Nuorgam 17 km SSE, 69.944°N, 28.041°E, 1♀ (BC ZSM HYM 11450), leg. M. Kraus, ZSM. 1♂ (BC ZSM HYM 11451), same collection data as preceding, ZSM.</p><p>Etymology.</p><p>The name is an adjective derived from the species name of its closely similar sister species,  D. aericeps Thomson, using the Latin diminutive suffix - ellus.</p></div>	https://treatment.plazi.org/id/5821C75CF43A5312BEC6D962BB6CEBDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
D6B0BCA16A4C5615B22176AF98FFE5EC.text	D6B0BCA16A4C5615B22176AF98FFE5EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus anthracinus (Klug 1818)	<div><p>Dolerus anthracinus (Klug, 1818)</p><p>Fig. 10A</p><p>Notes.</p><p>Tenthredo (Dolerus) anthracina Klug, 1818: 302. ♀, ♂. Syntypes. Type locality: Germany, Gartz in Pommern. Lectotype ♂ hereby designated, labelled:  “14205” [collection catalog no., printed], "  Anthracina Kl." [ Klug’s handwriting],  “Kl.” [ Klug’s handwriting], "GBIF-GISHym 2308" [printed], "Zool. Mus. Berlin" [printed], "  Lectotypus ♀  Tenthredo (Dolerus) anthracina Klug, 1818 M.Heidemaa des. '22 [handwritten in block letters]" [red], "  Dolerus anthracinus (Klug, 1818) M.Heidemaa det. [printed] not nitens Zadd." [handwritten in block letters]. ZMHB.</p></div>	https://treatment.plazi.org/id/D6B0BCA16A4C5615B22176AF98FFE5EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
29598963BCC5524EAB34094F8FA0A166.text	29598963BCC5524EAB34094F8FA0A166.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus bimaculatus (Geoffroy 1785)	<div><p>Dolerus bimaculatus (Geoffroy, 1785)</p><p>Tenthredo bimaculata Geoffroy in Fourcroy, 1785: 368, not 386 as given by Taeger et al. (2010). Sex not given. Syntypes (assumed). Type locality: France. The earlier description in French by Geoffroy (1762: 279, no. 16).</p><p>Tenthredo tristis Fabricius, 1804: 39. Sex not given. Syntypes (assumed). Type locality: Germany ("Habitat in Germania Dr. Panzer."). Lectotype designated below. Primary homonym of  Tenthredo tristis Fabricius, 1779.</p><p>Type material examined and taxonomic notes.</p><p>Tenthredo tristis Fabricius, 1804. Lectotype ♂ hereby designated, labelled: "  Tenthredo tristis ", "LECTOTYPE  Tenthredo tristis Fabricius, 1775 [sic!] des. SMBlank 2008" [red, printed], "  Dolerus bimaculatus (Geoff.) [handwritten] det. S.M.Blank 2008 ✓" [printed], "  Dolerus bimaculatus (Geoffroy) M.Heidemaa det." [printed], " ZMUC-GISHym 1067". Left flagellum, right flagellomeres 6-7, left fore tarsomeres 4-5 missing. ZMUC.</p><p>Contrary to  Fabricius’ original description, the posterior half of tergum 2 is red in addition to terga 3-4.</p><p>The application of the name  Tenthredo bimaculata to this particular  Dolerus species remains uncertain until the type material (syntype(s) in coll. E. L. Geoffroy, MNHN or Museum  d’Historié Naturelle  d’Autun, France?) should be located, or a neotype designated. The diagnosis by Geoffroy (1785) for  Tenthredo bimaculata is uninformative: "deux taches blanches au corcelet", but based on the description by Geoffroy (1762) it was regarded as a possible senior synonym of  D. tristis (F.) by de Dalla Torre (1894). Konow (1897a, 1905b) subsequently affirmed this synonymy.</p></div>	https://treatment.plazi.org/id/29598963BCC5524EAB34094F8FA0A166	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
8F59D6608D2B56D386653723F6213B7A.text	8F59D6608D2B56D386653723F6213B7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus coracinus (Klug 1818)	<div><p>Dolerus coracinus (Klug, 1818)</p><p>Tenthredo (Dolerus) coracina Klug, 1818: 302-303. ♀. Syntypes (assumed). Published type locality:  “Kärnthen” (Austria, Carinthia). Lectotype designated below.</p><p>Dolerus anthracinus auct. nec Klug. Misidentifications of  D. anthracinus by, for example: Konow (1885, 1886b), Enslin (1913), Benson (1952), Zhelochovtsev (1994), Lacourt (2020).</p><p>Dolerus atricapillus Hartig, 1837: 239. Lectotype ♂ [examined] designated by Blank and Taeger (1992). ZSM. Type locality: Germany. syn. nov.</p><p>Type material examined.</p><p>Tenthredo (Dolerus) coracina Klug, 1818. Lectotype ♀ hereby designated, labelled:  “14207” [collection catalog no., printed], "Carinth. Kl." [type locality: Austria, Carinthia, probably Klagenfurt], "  Coracina Kl." [ Klug’s handwriting], "GBIF-GISHym 2311" [printed], "Zool. Mus. Berlin" [printed], "  Lectotypus ♀  Tenthredo (Dolerus) coracina Klug, 1818 M.Heidemaa des.  ‘22” [block letters, red], "  Dolerus coracinus (Klug, 1818) M.Heidemaa det. [printed] =  D. anthracinus auct." [handwritten]. ZMHB.</p></div>	https://treatment.plazi.org/id/8F59D6608D2B56D386653723F6213B7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6FED188EF92E549296338AB6A1DCB807.text	6FED188EF92E549296338AB6A1DCB807.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus coruscans Konow 1890	<div><p>Dolerus coruscans Konow, 1890 sp. rev.</p><p>Fig. 17A-H</p><p>Dolerus varispinus sensu Konow, 1884: 351.</p><p>Dolerus coruscans Konow, 1890a: 10. Described by indication on the description of  Dolerus varispinus sensu Konow, 1884 (above). ♀, ♂. Syntypes. Published type locality: not given. Lectotype designated below.</p><p>Dolerus nigrominutus Haris, 1998: 131-132. ♂. Holotype. Type locality: Budakeszi  Hârsbokorhegy (Hungary). Apex of abdomen missing, genitalia dissected and pinned with the specimen. Paratypes: 3♂. All in HNHM. syn. nov.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus coruscans Konow, 1890.  Fürstenberg /Mecklenburg [Germany]. Lectotype ♂ hereby designated, labelled: "  Dolerus varispinus Htg fbg. [ Fürstenberg] 4.84." [Type locality: Germany, Brandenburg,  Fürstenberg], "  Dolerus coruscan s Knw.",  “Type” [red, printed], "  Syntypus " [red, printed], "Coll. Konow" [printed], "GBIF-GISHym 3762" [printed], "  Lectotypus [printed] ♂  Dolerus coruscans Konow, 1884 M.Heidemaa design.  ‘22”, "  Dolerus [printed] ♂ coruscans Konow, 1890 [handwritten in block letters] M.Heidemaa det. [printed]". Antennomeres 5-9 missing, genitalia dissected, pinned with the specimen. SDEI. Paralectotype: 1♀ (GBIF-GISHym 3761), Germany,  Schönberg [probably Brandenburg]. Here determined as  D. anthracinus. SDEI.</p><p>Oehlke and Wudowenz (1984) correctly linked  D. coruscans Konow, 1890 to the description of  D. varispinus Hartig sensu Konow, 1884, but following the definitions used in the Code (ICZN 1999), they were wrong in categorizing it as a nomen novum proposed as a replacement for a previously existing name.  Dolerus coruscans was, in fact, described as a new species, by indication on the description by Konow (1884). Konow (1890a) altered his opinion on his 1884 identification of  D. varispinus, and stated that this taxon represented  D. anthracinus sensu Thomson. Goulet (1986) followed  Konow’s 1890 opinion, but this cannot be accepted, because Konow clearly based his 1884 description on material in his own collection, not in  Thomson’s collection. Although  D. coruscans was later synonymized (Enslin 1909) with  D. nitens Zaddach, it has been used in published works more than once as a valid name since 1899 (e.g. Dittrich 1905; Fedchenko 1905; Dalglish 1914) and cannot be treated as a nomen oblitum. However, it was listed as a synonym of  D. anthracinus by Liston (1981), while Konow (1890b, 1905b) and Cameron (1893) regarded it as possibly conspecific with  D. possilensis Cameron. A recent study of the syntypes of  D. coruscans (♀, ♂) revealed that the penis valve of the lectotype male (Fig. 17A) is identical to  D. nigrominutus Haris, 1998 (the holotype and two paratype males studied) but the penis valve drawing by Haris (1998) was apparently based on a deformed valve because the other valves examined (including those of the holotype) have no such distinct depression on the ventroapical margin near the valvispina. The paralectotype female of  D. coruscans (see above) is probably a female (the only one known at present) of  D. anthracinus Klug (nec auct.).</p><p>Characters of the female.</p><p>The female of  D. coruscans, based on two specimens, resembles  D. picipes in many characters (the males are even more similar) and the melanic form of  D. liogaster with black legs, but differs by its mostly smaller body size of 6-7 mm and the rather inconspicuous setation of the ovipositor sheath (valvula 3) (Fig. 17G). Due to their similarity, the diagnostic characters of  Dolerus coruscans are compared to their states in  Dolerus picipes .</p><p>Head. (Fig. 17B, C) Closely resembles  D. picipes, but the minimal distance between the antennal sockets is about 2  × the length of the malar space (clearly shorter in  Dolerus picipes). Pits on face and vertex more unevenly distributed and vary more in their size. Postocellar field less distinctly defined than in  Dolerus picipes and lateral postocellar furrows less distinct: short and pit-like, rather than long and line-like in  Dolerus picipes . Glossy patches beside the lateral postocellar furrows less distinct (almost no difference between the males).</p><p>Thorax. (Fig. 17D, E) The katepimeron of the mesopleuron (Fig. 17D) with more numerous and distinct pits than in  Dolerus picipes . The lateral mesoscutal lobes (Fig. 17E) less distinctly and rather sparsely pitted.</p><p>Abdomen. (Fig. 17F-H) The ovipositor sheath in dorsal view broadens slightly towards the apex (Fig. 17G). Cerci (Fig. 17H) yellowish (black in  Dolerus picipes, Fig. 17I). Setae on valvula 3 rather short and delicate compared to most other  Poodolerus species (Fig. 17G, H). Setae on the abdominal terga shorter and sparser, except on the apical terga. The species can be easily differentiated from  D. picipes as follows.</p><table><tr><td colspan="1" rowspan="1">1 a Lateral postocellar furrows distinct: long and line-like b Minimal distance between the antennal sockets about 1.5  × the length of malar space c Ovipositor sheath in dorsal view strongly broadening towards apex, the apical setae well developed and clearly curved (Fig. 17I)</td><td colspan="1" rowspan="1">D. picipes (Klug, 1818)</td></tr><tr><td colspan="1" rowspan="1">- aa Lateral postocellar furrows indistinct: short and pit-like (Fig. 17C) bb Minimal distance between the antennal sockets about 2  × the length of malar space (Fig. 17B) cc Ovipositor sheath in dorsal view slightly broadening towards the apex, the apical setae delicate, nearly straight or slightly curved (Fig. 17G)</td><td colspan="1" rowspan="1">D. coruscans Konow, 1890</td></tr></table><p>Distribution.</p><p>Dolerus coruscans is recorded from Austria (N-Burgenland, Ungerberg, Weiden am See, 1♀, 24.03.1957, leg. F. Kasy, SDEI), Germany (lectotype: see above), Hungary ( “Budapest”, 1♀, SDEI. Haris 1998, 2003: "Budakeszi, Csepel,  Látrány,  Rákospalota”; Roller and Haris 2008: "Simontornya,  Csévharaszt,  Szécsény”, in total 14♂), and Greece (Nom.  Grevená,  Kónitsa E 28 km  Vasilítsa Ski Resort, 40.050°N, 21.050°E, 1780 m, 2♂, 11.05.2007, leg. S. M. Blank et al. / leg. M. Wei, SDEI. The records of  D. coruscans from Austria and Greece are the first for these countries. The species apparently has a wide distribution, but is probably seldom collected because of its rarity and early flight period. Probably it has also sometimes remained unidentified, or has been misidentified. Possibly, the males of this species in collections could have been placed under  D. picipes and its rather small females have perhaps been mistaken for dwarf females of  D. liogaster /  D. puncticollis (both species also have melanic color forms).</p><p>Remarks.</p><p>Haris (2003) speculated that the female of this species might be wingless, or have reduced wings. Present data exclude this possibility. Nor does the female remain unknown, as Haris (1998) suggested it would, because of the destruction of the very small type locality of  D. nigrominutus . In Central Europe  D. coruscans has been collected early in the season (March-April).</p></div>	https://treatment.plazi.org/id/6FED188EF92E549296338AB6A1DCB807	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
5E0E8E524CA85ADB94813CBB5607D40B.text	5E0E8E524CA85ADB94813CBB5607D40B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus ferrugatus Serville 1823	<div><p>Dolerus ferrugatus Serville, 1823</p><p>Dolerus ferrugatus Serville, 1823: 59. ♀. Syntypes. Type locality: Paris. Lectotype designated below.</p><p>Dolerus brevicornis Thomson, 1871: 288. ♀, ♂. Syntypes. Published type locality:  “södra Sverige" [southern Sweden]. Lectotype designated below. Primary homonym of  D. brevicornis Zaddach, 1859.</p><p>Dolerus thomsoni var. miricolor Konow, 1887a: 283. ♂. Syntypes (assumed). Type locality:  Fürstenberg / Mecklenburg. Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus ferrugatus Serville, 1823. Lectotype ♀ hereby designated, labelled:  “Saint-Fargeau” [round],  “Lectotype” [red, printed in block letters], "Lectotype  D. ferrugatus LEP. ♀ det. A.Haris 1996", "  D. ferrugatus LEP. ♀ det.: A.Haris", "  Lectotypus [printed]♀  Dolerus ferrugatus Serville, 1823 [handwritten in block letters] M.Heidemaa design. [printed]" [red], "  Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. In good condition (the right flagellum missing). MNHN.</p><p>Dolerus brevicornis Thomson, 1871. Lectotype ♀ hereby designated (type locality: Sweden,  Öland), labelled:  “Ö.” [ Öland], " MZLU 2013 414" [green, printed (a loan record label)], "  Lectotypus [printed] ♀ 2014  Dolerus brevicornis Thomson, 1871 [handwritten in block letters] M.Heidemaa des. [printed]" [red], "  Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. 2♀ (from  Öland and Scania), 1♂ (the melanic form, from Lund) labelled as paralectotypes. All in MZLU.</p><p>Dolerus thomsoni var. miricolor Konow, 1887. Lectotype ♂ hereby designated, labelled: "f. [ Fürstenberg] 12/4. [18]84.", "  Dolerus v. [var.]  Dolerus miricolor Knw. [Konow] fbg. 12/4. 84. Typus" [with double black frame], "  Dolerus ferrugatus Lep. f. miricolor Knw. Typus. ♂ det. OConde [Otto Conde] 1943",  “Typus” [red, printed],  “Holotypus” [red, printed], "Coll. Konow" [printed], "GBIF-GISHym 4215" [printed], "  Lectotypus [printed] ♂  Dolerus thomsoni var. miricolor [handwritten in block letters] M.Heidemaa des. [printed]" [red], "  Dolerus ferrugatus Serville, 1823 M.Heidemaa det." [printed]. Right flagellum missing, penis valves dissected (a microscope slide:  Symphyta coll. Nr: 159). All in SDEI.</p><p>The melanic colour form of  D. ferrugatus, described by Konow as  D. thomsoni var. miricolor, has sometimes been mixed up with  D. pachycerus Hartig, 1837. Apart from their different penis valves, they can be separated by the shape and the structure of the mesoscutellar appendage: long, concave at the sides, and with a distinct longitudinal keel in  D. pachycerus compared with  D. ferrugatus .</p></div>	https://treatment.plazi.org/id/5E0E8E524CA85ADB94813CBB5607D40B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
DB6A6BBD7A3C50A3B4A61B950A7F7BA8.text	DB6A6BBD7A3C50A3B4A61B950A7F7BA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus fumosus Stephens 1835	<div><p>Dolerus fumosus Stephens, 1835</p><p>Figs 11A-D, 12A, C, E, G-I</p><p>Dolerus fumosus Stephens, 1835: 87. Sex not stated. Syntypes (assumed). Type locality: Hertford (UK). Lectotype designated below.</p><p>Dolerus Lucens [sic!]  André, 1880: 276-277. ♀, ♂. Syntypes. Published type locality: Hongrie (Hungary). Lectotype designated below. syn. nov.</p><p>Dolerus sanguinicollis var.  Dolerus sanguinicollis Reicherti [sic!] Konow, 1894b: 134. ♀. Holotype. Type locality: near Leipzig (Germany). SDEI. syn. nov.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus fumosus Stephens, 1835. Lectotype ♀ hereby designated, labelled: "Type H.T." [round with wide red margin, printed], "B.M. TYPE HYM. [printed] 1.572.", "B.M. TYPE HYM. [printed]  Dolerus fumosus (Stephens 1835)", "  Dolerus fumosus .", "Stephens Coll. 53-46" [printed], "  Lectotypus 2012  Dolerus fumosus ♀ [the symbol handwritten] Stephens, 1835 M.Heidemaa des." [red, printed], "  Dolerus sanguinicollis Klug M.Heidemaa det." [printed] [Identified before it became clear that two species were mixed up under this name.] Condition poor: mid- and hind tarsi incomplete, 2 apical flagellomeres of the right antenna missing, abdomen partly damaged. BMNH.</p><p>Kirby (1882) referred to the "Types (♂, ♀)". The number of syntypes of  D. fumosus was not given by Stephens, but only one syntype female is now present in the collection (BMNH).</p><p>Dolerus lucens André, 1880. Lectotype ♂ hereby designated, labelled:  “Jászó” [Type locality: Hungary,  Jászó (Jasov)],  “272”,  “Hongrie” [printed in block letters], "  D. lucens,  André.” [printed], "DEI GISHym 89371" [printed], "coll. Desbrochers [handwritten] k. [kollektsija = coll.] A. Jakovleva [nomin. A. Jakovlev]" [printed], "SYNTYPUS [printed in red]  Dolerus lucens ANDRÉ, 1880 [handwritten] teste A. Taeger 20[printed]18 [handwritten]" [white label with a red frame], "  Lectotypus [printed]  Dolerus lucens André, 1880 [handwritten] M.Heidemaa des. [printed]" [red label], "  Dolerus fumosus Stephens, 1835 M.Heidemaa det." [white, printed], ZIN. Paralectotype: 1♂ with same original collecting labels as the lectotype, but the no. is  “271” and "  Dolerus lucens André (typ.)" [handwritten, probably by Ed.  André]. Left forewing missing. HNHM.</p><p>The paralectotype of  D. lucens mentioned above bears a lectotype label by L. Zombori ("  Lectotypus ♂  Dolerus lucens [ …] des. Zombori 1980"), but Zombori never published a lectotype designation. Although the paralectotype male has a handwritten label, most probably by  André ("  Dolerus lucens André (typ.)", Fig. 12I) and its penis valve is dissected, another syntype male (the right forewing and flagellum missing) is selected as the lectotype because it is deposited in  André’s collection (in ZIN), with many other type specimens of species described by him.</p><p>The holotype female of  D. sanguinicollis var. reicherti has reddish lateral mesoscutal lobes and median mesoscutal lobes only slightly reddish on the anterior part, but according to the other diagnostic characters mentioned above, it belongs to  D. fumosus .</p></div>	https://treatment.plazi.org/id/DB6A6BBD7A3C50A3B4A61B950A7F7BA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F0CAE4C0333A523E922425B3294049B6.text	F0CAE4C0333A523E922425B3294049B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus germanicus (Fabricius 1775)	<div><p>Dolerus germanicus (Fabricius, 1775)</p><p>Tenthredo germanica Fabricius, 1775: 321. ♀. Syntypes (assumed). Type locality: Germany ("Habitat in Germania."). Lectotype designated below.</p><p>Dolerus arcticus Thomson, 1871: 284-285. ♀, ♂. Syntypes. Type locality: "Lapland, Dalarna,  Ångermanland” . Type locality: Lapland. Lectotype designated below.</p><p>Dolerus cothurnatus Serville, 1823: 60. ♂. Syntypes. Type locality: Paris. Lectotype designated below. syn. nov.</p><p>Dosytheus fuscipennis Stephens, 1835: 85. Sex not given. Syntypes (assumed). Type locality: south of Scotland. Lectotype ♀ designated by Kirby (1882).</p><p>Dosytheus hyalinalis Stephens, 1835: 83. ♀, ♂. Syntypes. Type locality: Hertford (UK). Lectotype designated below. syn. nov.</p><p>Dosytheus xanthopus Stephens, 1835: 83-84. Sex not given. Syntypes (assumed). Type locality: Hertford (UK).</p><p>Type material examined and taxonomic notes.</p><p>Tenthredo germanica Fabricius, 1775. Lectotype ♀ hereby designated, labelled: "  Tenthredo germanica ", "LECTOTYPE  Tenthredo germanica Fabricius, 1775 des. SMBlank 2008", [red, printed], " ZMUC-GISHym 1050", "  Dolerus germanicus (F.) [handwritten] det. S.M.Blank 2008 ✓" [printed], "  Dolerus germanicus (Fabricius) M.Heidemaa det." [printed], " ZMUC-GISHym 1043". Left flagellum, right fore tarsus, some additional distal tarsomeres missing. ZMUC.</p><p>Two specimens are present in the ZMUC collection under the name  Tenthredo germanica . Klug (1819) referred to one of them, which has black legs and an infuscated tip of the abdomen ("Das vorhandene Exemplar hatte schwarze Beine und einen an der Spitze  schwärzlichen Hinterleib, [...]"). This specimen is selected as the lectotype. It corresponds with the current concept of  Dolerus germanicus, e.g. in the shape of the sawsheath (valvula 3) setation. The species needs a thorough revision, because DNA barcoding divides the specimens into two BINs (BOLD:AAI9736 and BOLD:ABV8027) separated by a minimum of over 5% divergence, indicating the possible presence of two distinct species. The two BINs are associated with color differences, of which some appear stable within the cluster and others are variable. Based on about 80 barcoded specimens from Finland, specimens of BOLD:ABV8027 consistently have entirely black hind legs, while those of BOLD:AAI9736 usually have largely orange hind femora and tibiae, although sometimes they are completely black. Additionally, females with a red mesoscutellum always fall in BOLD:AAI9736 and always have red tegulae, pronotum and mesoscutum, whereas in BOLD:ABV8027 the mesoscutellum is always entirely or mostly black, but the tegulae, pronotum and mesoscutum vary from black to red. It is also worth mentioning that the division of specimens into two groups was not reflected in two nuclear genes (POL2 and NaK). Likewise, no useful structural characters have yet been found, but a more detailed systematic study of the male genitalia might be helpful.</p><p>Dolerus arcticus Thomson, 1871. Lectotype ♀ hereby designated, labelled:  “Lapp” [Lapland] [printed], "  Dolerus arcticus m[mihi]" [ Thomson’s handwriting], " MZLU 2013 421" [green, printed loan label], "  Lectotypus [printed] ♀ 2014  Dolerus arcticus Thomson, 1871 [handwritten in block letters] M.Heidemaa des. [printed]" [red], "  Dolerus germanicus (Fabricius) M.Heidemaa det." [printed]. Ovipositor dissected, glued on a piece of paper pinned below the specimen. Paralectotypes: 3♀ [MZLU 422, 424, 425], and 1♂ [MZLU 423]. All in MZLU.</p><p>Dolerus cothurnatus Serville, 1823. Lectotype ♂ hereby designated, labelled: [green, round, without text],  “Lectotype” [red, printed in block letters], "  D. cothurnatus LEP. Det.: A.Haris", "Lectotype  D. cothurnatus LEP. Det.: A.Haris", "  Lectotypus [printed] ♀[handwritten], 2014  Dolerus cothurnatus Serville, 1823 [handwritten in block letters] M.Heidemaa des. [printed]" [red label], "  Dolerus germanicus (Fabricius, 1775) M.Heidemaa det." [printed]. Two paralectotype males were also examined: one of them is  D. germanicus, the other is  D. aericeps . All in MNHN.</p><p>A study of the penis valves of the three syntypes of  D. cothurnatus revealed that two of them belong to  D. germanicus, including the specimen here designated as the lectotype, and one to  D. aericeps . These old specimens all have rather pale wings, but probably they have faded:  D. germanicus and  D. junci [=  Dolerus cothurnatus auct.] typically have darker wings. The specimen labelled by Haris as lectotype is designated here as a lectotype for  D. cothurnatus Serville and  D. cothurnatus is regarded as a synonym of  D. germanicus (see above, under  Dolerus cothurnatus auct. nec Serville, 1823).</p><p>Dosytheus fuscipennis Stephens, 1835. Lectotype ♀. Labelled: "Type H.T. [holotype]" [round with a red margin, printed], "B.M. TYPE HYM. [printed] 1.578.", "B.M. TYPE HYM. [printed]  Dosytheus fuscipennis Stephens 1835.", " fuscipennis.", "  Dolerus pratensis, L." [printed], "Stephens Coll. 53-46" [printed], "  Dolerus germanicus (Fabricius) M.Heidemaa det." [printed]. Condition good, legs partly destroyed. BMNH.</p><p>The number of specimens was not given by Stephens. The species description should therefore be assumed to have been based on syntypes. However, Kirby (1882) referred to the "♀. (Type of  D. fuscipennis .) S. Scotland. J. F. Stephens", and according to the labels and the collection catalogue this was the only specimen from the Stephens collection in the BMNH. The specimen fits the original description, and  Kirby’s statement qualifies as a valid lectotype designation, because there is no evidence in the original description that more than one type specimen existed (ICZN 1999 Article 74.6.).</p><p>Dosytheus hyalinalis Stephens, 1835. Lectotype ♀ hereby designated, labelled: "B.M. TYPE HYM [printed] 1.577.", "B.M. TYPE HYM [printed]  Dosytheus hyalinalis . (Stephens 1835)", "  Dosytheus hyalinalis Steph", "42 1b" [round], "  Dolerus pratensis, L." [printed], "  Dolerus germanicus (F.) M.Heidemaa det." [printed]. BMNH.</p><p>In the original description Stephens indicates that he had more than one specimen, because he gives color characters for both sexes.  Kirby’s statement "♀ Type of  D. hyalinalis " (Kirby 1882) refers to the only syntype from the Stephens collection that was present in the BMNH, but cannot be accepted as a lectotype designation, because it is clear that Stephens based his description on more than one specimen (ICZN 1999 Article 74.5.). The synonymy with  D. pratensis, based on a misidentification, was adopted for example by de Dalla Torre (1894) and most other subsequent works.</p><p>Dosytheus xanthopus Stephens, 1835. The only specimen found in the collection (BMNH) labelled as "  Dosytheus xanthopus " and "B.M. TYPE 1.257." belongs to  D. germanicus . It cannot be a syntype of  D. xanthopus, because the specimen disagrees with the original description: its mesoscutellum is orange, not black as described, and the locality label ("Kent, Darenth, J. F. Stephens BM 1853 - 42", probably added by Benson) does not match the locality given by Stephens. Interestingly, it was labelled by Benson (handwritten): "  Dolerus etruscus ?".</p></div>	https://treatment.plazi.org/id/F0CAE4C0333A523E922425B3294049B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
31AA3E3992E65E95BC92589CF688FD1B.text	31AA3E3992E65E95BC92589CF688FD1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus gessneri Andre 1880	<div><p>Dolerus gessneri 
Andre, 1880</p><p>Fig. 19</p><p>Notes.</p><p>Dolerus Gessneri [sic!]  André, 1880: 273. Sex not given. Syntypes. Type locality: Switzerland. Lectotype ♀ hereby designated, labelled:  “Suisse”, "DEI GISHym 89367" [printed], "coll. Desbrochers [handwritten] k. [kollektsija = coll.] A. Jakovleva [nomin. A. Jakovlev]" [printed], "SYNTYPUS [printed in red]  Dolerus gessneri ANDRÉ, 1880 [handwritten] teste A. Taeger 20[printed]18 [handwritten]" [white with a red frame]. "Lectotype  Dolerus gessneri André 1880 vide A. Taeger 2022 des." [red, printed]. Good condition, but slightly dusty. Paralectotypes: 2♂ (one without flagella) with the same data (except  “♂” [printed], "DEI-GISHym 89365" and  “D… 89366" and "  Paralectotypus " instead of "  Lectotypus "). All in ZIN.</p><p>The color form of  Dolerus gessneri with a red-banded abdomen was described as a distinct species,  D. labiosus Konow, 1897 (Konow 1897b), but was later mostly treated as a synonym (sometimes also as a subspecies) of  D. gessneri, because intermediate color forms with variable red markings on the abdomen occur. The specimens from the Russian Far East with completely black forelegs, which have sometimes been determined as  D. gessneri, probably belong to some other  Dolerus (subgen. Equidolerus) species.  Dolerus gessneri, a Holarctic species, needs a revision, because COI barcoding data suggest that it could include more than one species, and some morphological characters correlate with the genetic data. There are three BIN clusters (BOLD:AAL2317, BOLD:ACE7304, BOLD:ACE3617) diverging by 1.8-2.3%. BOLD:AAL2317 is closest to  D. timidus and BOLD:ACE7304 is closest to  D. pratensis (minimum divergence in both cases 1.2%). All three BIN clusters are represented in Fennoscandia, two of which (Fig. 19) have been found in Finland.</p></div>	https://treatment.plazi.org/id/31AA3E3992E65E95BC92589CF688FD1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B19286EDE8B85545B2EB8B3FB701D40D.text	B19286EDE8B85545B2EB8B3FB701D40D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus gonager (Fabricius 1781)	<div><p>Dolerus gonager (Fabricius, 1781)</p><p>Tenthredo gonagra Fabricius, 1781: 412. Sex not given. Syntypes (assumed). Type locality: "Habitat in Germaniae plantis." [Germany]. Lectotype designated below.</p><p>Dolerus femoratus Eversmann, 1847: 24-25. ♀. Syntypes. Type locality: Kasan (Russia, Taterstan). Lectotype designated below.</p><p>Dolerus magnicornis Eversmann, 1847: 25. ♂. Syntypes (assumed). Type locality: Russia, Orenburg Province. Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Tenthredo gonagra Fabricius, 1781. Lectotype ♀ hereby designated, labelled: "  Tenthredo gonagra "; "LECTOTYPE  Tenthredo gonagra Fabricius, 1781 des. SMBlank 2008" [red, printed: paralectotypes labelled in corresponding way], " ZMUC-GISHym 1050", "  Dolerus gonager (F.) [handwritten] det. S.M.Blank 2008 ✓" [printed], "  Dolerus gonager (Fabricius, 1781) M.Heidemaa det." [printed]. Right flagellomeres 6-7, right hind tarsus missing. Paralectotype 1♀: "  Tenthredo gonagra ", " ZMUC-GISHym 1051". ZMUC.</p><p>The type specimens correspond with  Zhelochovtsev’s definition of  Dolerus (Poodolerus) gonager, which has red knees, superficially pitted lateral mesonotal lobes and backwards directed setae of the parallel-sided ovipositor sheath (similar to Zhelochovtsev 1994: fig. 165.2, but with some of the longest hairs curved distally).</p><p>Dolerus femoratus Eversmann, 1847. Lectotype ♀ hereby designated, labelled: "Kas[an] 16.V.", "  Dolerus femoratus . Kl." [with black frames], "DEI-GISHym 30068" [white, printed], "Lectotype  Dolerus femoratus Eversmann 1847 vide A. Taeger 2014 des." [red, printed]. In good condition. Belongs to  D. gonager . Paralectotypes: 2♀ (DEI-GISHym 30069, 30070), are  D. puncticollis Thomson, 1871. All in ZIN.</p><p>Dolerus magnicornis Eversmann, 1847. Lectotype ♂ hereby designated, labelled:  “Orb” [Orenburg], "  Dolerus magnicornis mihi" [with black frames], "DEI-GISHym 30067" [white, printed], "Syntype  Dolerus magnicornis Eversmann 1847 vide A. Taeger 2014" [red, printed], Lectotype  Dolerus magnicornis Eversmann 1847 M.Heidemaa des." [red, printed]. In good condition. Belongs to  D. gonager . ZIN.</p></div>	https://treatment.plazi.org/id/B19286EDE8B85545B2EB8B3FB701D40D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
BCFC3E42126051FFBAD8C863C00CF548.text	BCFC3E42126051FFBAD8C863C00CF548.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus incisus Goulet 1986	<div><p>Dolerus incisus Goulet, 1986</p><p>Fig. 20A-C</p><p>Notes.</p><p>Finland: Regio kuusamoensis, Kuusamo Siikauopaja, YKJ grid coordinates: 7362:3612 [66.334°N, 29.511°E], 2♀ (ZMUO.035155, ZMUO.035156), 12.06.2018. 1♀ (ZMUO.045265), 20.06.2020. All specimens leg. M. Mutanen. ZMUO.</p><p>First record in the Palaearctic. Recorded previously only in boreal North America: Alaska, Alberta, Yukon, and North West Territories (Goulet 1986). The genetic and morphological affinity of  D. incisus to  D. junci and  D. yukonensis suggests that the host plant of  D. incisus is also a species of  Equisetum (Goulet 1986). The habitat in Finland is a shallow oxbow lake (meander) of the Oulanka river with plenty of  Equisetum fluviatile, which is also the host of  D. junci . This habitat matches perfectly with that given for  D. incisus in North America by Goulet (1986). Several other  Dolerus species occur syntopically, including the closely related  D. junci Stephens (=  D. cothurnatus auct., see above). The melanic form of  D. incisus is recorded from Finland (all three are females, see above, Fig. 20A) and Norway (Heimdalsmunnen, alt 1150 m, leg. E. Heibo, CEH). Melanic females of  D. subarcticus (recorded from Norway and Kamtschatka, see under  D. subarcticus) can be distinguished by their postocellar furrows, the sawsheath (valvula 3) shape, and its apical setation (Fig. 20B). Melanic males can be separated most reliably by their penis valves (Fig. 20C, 20D). Only the melanic colour form of  D. incisus has so far been recognized from Fennoscandian countries, whereas in Canada and the USA both the black color form and a form with red-banded abdomen and partly reddish legs are known. We studied paratypes of  D. incisus: 2♀, 2♂, some of them dissected, CNC.</p></div>	https://treatment.plazi.org/id/BCFC3E42126051FFBAD8C863C00CF548	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F8AD7C874E7F5541A3ECD63CFC629AEA.text	F8AD7C874E7F5541A3ECD63CFC629AEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus junci (Stephens 1835)	<div><p>Dolerus junci (Stephens, 1835) nom. rev.</p><p>Dosytheus Junci [sic!] Stephens, 1835: 84-85. Sex not explicitly stated. Syntypes. Published type localty: "Devonshire, and near Windsor" (England). Lectotype designated below.</p><p>Dolerus Busaei [sic!] Snellen van Vollenhoven, 1858: 277. Sex not given. Lectotype ♂ designated by Thomas (1987). Type locality:  “Haarlem” (Netherlands). syn. nov.</p><p>Dolerus cothurnatus auct. nec Serville, 1823 (e.g. Benson 1952, Zhelochovtsev 1994, Lacourt 2020).</p><p>Dolerus thargitai Zombori, 1994: 185-187. ♀. Holotype. Type locality:  Csíkszentkirály,  Borsáros,  Kérújfürdő: Tolvajos patak (Romania). syn. nov.</p><p>Remarks.</p><p>Similarly to the melanic color form of  D. yukonensis (see below), which was described as a distinct species,  D. scoticus, by Cameron (1881a), the melanic form of  D. junci was also described as a distinct species,  D. thargitai, by Zombori (regarded as a synonym of  D. yukonensis /  D. scoticus until now). The melanic color form of  D. junci has also been recorded from central Europe (Germany) and southern Europe (Italy, Switzerland: see, e.g. Pesarini 2012), but is not known from Fennoscandia. The males of the melanic forms of  D. junci (01.06.1984) and  D. yukonensis (24.05.1984) have been collected at the same locality at nearly the same time (Germany: Federsee, Brackenhof, leg. E. Jansen, CEJ). In addition to their different penis valve structure and other characters used in the keys by Benson (1952) and Zhelochovtsev (1994),  D. junci and  D. yukonensis differ in the shape and sculpture of the mesoscutellar appendage: almost unsculptured and without a distinct longitudinal keel in the former, and clearly sculptured and with a distinct keel in the latter. Most likely, the melanic color form of  D. junci does not occur within the geographic range of  D. incisus, but they can also be distinguished by the shape and the sculpture of the mesoscutellar appendage: meshed (nearly matt) and long, with a distinct median longitudinal keel ( D. incisus) versus short and almost unsculptured (more or less glossy) and without a distinct median keel ( D. junci).</p><p>Type material examined and taxonomic notes.</p><p>Dosytheus junci Stephens, 1835. Lectotype ♀ hereby designated, labelled: "Type H.T. [holotype]" [round with wide red margin, printed; not a holotype but a syntype!], "B.M. TYPE HYM. [printed] 1.575.", "B.M. TYPE HYM. [printed]  Dosytheus Junci Stephens 1835.", "  Junci .", "Stephens Coll. 53-46" [printed], "  Dolerus palustris, Klug." [printed], "  Lectotypus 2012  Dosytheus junci [printed]♀ [symbol handwritten] Stephens, 1835 M.Heidemaa des. [printed]", "  Dolerus junci Stephens, 1835 M.Heidemaa det." [printed]. Condition: legs partly destroyed, left midleg missing, the apical flagellomeres missing, apex of left forewing broken. BMNH.</p><p>The original description of  D. junci by Stephens (1835) refers to syntypes ("Devonshire and near Windsor"). Although Kirby (1882) mentioned a "Type of  D. junci ", this cannot be regarded as a lectotype designation, even though he was apparently only able to locate a single specimen.</p><p>Dolerus busaei Snellen van Vollenhoven, 1858. Photos of the lectotype and of the penis valve were checked. Left midleg and two segments of the right antenna missing; the genital capsule is glued on a card. RMNH. This nominal taxon was previously treated as a synonym of  D. cothurnatus auct.</p><p>Dolerus cothurnatus auct. nec Serville, 1823. Haris and later Lacourt (2000) studied the syntypes (3 ♂) of  D. cothurnatus Serville, but not their genitalia. Based on their penis valve structure, none of the syntypes fits  D. cothurnatus auct.: the two syntypes (among them the one labelled as lectotype) represent  D. germanicus and the third one is a male of  D. aericeps . The name  D. cothurnatus by Serville is placed here as a junior synonym of  D. germanicus, by designating its lectotype (see also under  D. germanicus), and the name  Dolerus junci (Stephens, 1835) has to be used for  D. cothurnatus auct. nec Serville.</p><p>Dolerus thargitai Zombori, 1994. The holotype and some paratypes (2♀, 4♂) were studied. HNHM.</p></div>	https://treatment.plazi.org/id/F8AD7C874E7F5541A3ECD63CFC629AEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6AD49FE0E2485E8996BBF91345E39D18.text	6AD49FE0E2485E8996BBF91345E39D18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus liogaster Thomson 1871	<div><p>Dolerus liogaster Thomson, 1871</p><p>Fig. 16A, C, E, H, K, N, Q</p><p>Dolerus liogaster Thomson, 1871: 286. ♀, ♂. Syntypes. Type locality: Sweden. Lectotype designated below.</p><p>Dolerus schneideri Kiaer, 1898: 62-63. ♀, ♂. Syntypes. Type locality:  Tromsø . Lectotype designated below.</p><p>Dolerus truncatus ?  Dolerus truncatus Lacourt, 1988: 233-235. ♂. Holotype. Type locality: France, Alpes-Maritimes,  Vallée de la  Minière . CTN.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus liogaster Thomson, 1871. Lectotype ♂ hereby designated, labelled: "Lp. in." [?Lapland], "LECTOTYPUS [printed]  Dolerus liogaster ♂ Thomson, 1871 [handwritten in block letters] design. 1999 M. Heidemaa and M. Viitasaari [printed]" [red], "  Dolerus liogaster ♂ [handwritten] M.Heidemaa det. [printed]" [white]. Penis valves dissected, pinned with the specimen. Paralectotypes: 2♀, 1♂. All in MZLU.</p><p>Dolerus schneideri Kiaer, 1898. Lectotype ♀ hereby designated, labelled:  “Tromsö 22.6.77", "lget expl" [spelling doubtful], "Ex coll. H. Kiaer" [printed],  “261” [yellow], "LECTOTYPUS [printed]  Dolerus schneideri ♀ Kiaer, 1898 [handwritten in block letters] design. 1999 M. Heidemaa and Viitasaari" [printed (excl. year), red], "  Dolerus liogaster ♀M. Heidemaa det." [printed label]. Ovipositor dissected and pinned with the specimen, right antenna missing. Paralectotypes: 2♀. All in ZMUN.</p><p>The synonymy of  D. schneideri with  D. liogaster was first proposed by Lindqvist (1943). Treated here as a color form of  D. liogaster, but still separated in the key, because the melanic color form of  D. liogaster also has black legs, and  D. schmidti has a color form with a red-marked thorax. Specimens resembling  D. schneideri are also known from the mountain areas of Central Europe, e.g. from Switzerland at altitudes of 1500-1700 m (Benson 1961).</p><p>Dolerus truncatus Lacourt, 1988. The holotype (CTN) was studied. Its clypeus emargination looks as if it is abnormally developed. The penis valve (Fig. 18) appears to be somewhat distorted, perhaps during preparation, but resembles the valve of  D. liogaster . Tentatively, we treat  D. truncatus as a junior synonym of  D. liogaster .</p></div>	https://treatment.plazi.org/id/6AD49FE0E2485E8996BBF91345E39D18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
85AB248FCBC250D4A046640701CB40EB.text	85AB248FCBC250D4A046640701CB40EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus possilensis Cameron 1882	<div><p>Dolerus possilensis Cameron, 1882</p><p>Notes.</p><p>Dolerus possilensis Cameron, 1882a: 178-179. ♀. Syntypes (assumed). Type locality: Possil Marsh near Glasgow, Scotland. Lectotype ♀ hereby designated, labelled: "B.M. TYPE HYM. 1.573" [printed], "  Dolerus possilensis Cam.", "Cameron 96-76. [printed] Possil" [handwritten], "B.M. TYPE HYM. [printed]  Dolerus possilensis (Cameron 1882)", "  Lectotypus [printed]  Dolerus possilensis Cameron, 1882 [handwritten in block letters] M.Heidemaa design. [printed]" [red]. Most of the abdomen missing. Ovipositor preparation apparently lost, but probably figured in Cameron (1882a: plate XIX, fig. 8). BMNH.</p><p>Probably a parthenogenetic species: no male has ever been recorded. The only known Fennoscandian record is from Sweden: 1♀ [labelled as  Dolerus sp.]  “Resarö Waxholm / 6 1915", leg. R. Malaise, NHRS. New to the Swedish fauna.</p></div>	https://treatment.plazi.org/id/85AB248FCBC250D4A046640701CB40EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
592652E4A2765C02B5A3C92AEEDBDBA9.text	592652E4A2765C02B5A3C92AEEDBDBA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus pratensis (Linnaeus 1758)	<div><p>Dolerus pratensis (Linnaeus, 1758)</p><p>Figs 13A, B, 14C, 14F, 15</p><p>Tenthredo pratensis Linnaeus, 1758: 556. Sex not given. Syntypes (assumed). Lectotype ♀ designated by Malaise and Benson (1934). Type locality: Europe.</p><p>Tenthredo (Dolerus) deserta Klug, 1818: 300-301. ♂. Holotype. Published type locality: "In hiesiger Gegend gefunden" [meaning the environs of Berlin, Germany].</p><p>Type material examined and taxonomic notes.</p><p>Tenthredo pratensis Linnaeus, 1758. Lectotype. LSUK. Images of the lectotype (LINN 2501) in dorsal and lateral view and the labels are available (The Linnean Society of London 2022), but we did not examine the specimen itself.</p><p>Tenthredo (Dolerus) deserta Klug, 1818. Holotype (GBIF-GISHym 2316; Fig. 15). Good condition, genitalia dissected and pinned with the specimen. ZMHB.</p></div>	https://treatment.plazi.org/id/592652E4A2765C02B5A3C92AEEDBDBA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A5AE91BDF997598D81218B845B442C3F.text	A5AE91BDF997598D81218B845B442C3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus puncticollis Thomson 1871	<div><p>Dolerus puncticollis Thomson, 1871</p><p>Fig. 21</p><p>Notes.</p><p>Dolerus puncticollis Thomson, 1871: 286. ♀, ♂. Syntypes. Published type locality: [Sweden]. Lectotype ♂ hereby designated, labelled (printed on pale paper if not stated otherwise):  “Lund” [type locality Sweden, Lund], "  Lectotypus [printed] 1♂  Dolerus puncticollis Thomson, 1871 [handwritten in block letters] M.Heidemaa design. [printed]" [red]. Paralectotypes (3♀, 2♂,  Öland, Lund, Scania; the locality labels of 2♀ are illegible). All in MZLU.</p><p>Penis valve of the lectotype as in Fig. 21. The taxonomic status of a melanic colour form closely resembling  D. puncticollis (recorded from Portugal and Greece) is still under study.</p></div>	https://treatment.plazi.org/id/A5AE91BDF997598D81218B845B442C3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
59D2DF44FD3956259223EDD0DF6DF0B1.text	59D2DF44FD3956259223EDD0DF6DF0B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus sanguinicollis (Klug 1818)	<div><p>Dolerus sanguinicollis (Klug, 1818)</p><p>Figs 11E, F, 12B-D, F, H</p><p>Note.</p><p>Tenthredo (Dolerus) sanguinicollis Klug, 1818: 305. ♀ [not explicitly stated, but indicated by colour characters]. Syntypes. Published type locality: Austria. Lectotype ♀ hereby designated, labelled:  “14222” [printed], "Austr. Kl." [Type locality: Austria, Carinthia, Klagenfurt], "  Tenthredo sanguinicollis Kl.", "  Lectotypus " [printed in block letters] ♀ 2014 [handwritten]  Tenthredo (Dolerus) sanguinicollis Klug, 1818 [handwritten], "Des. M.Heidemaa" [red label], "GBIF-GISHym 2343", "Zool. Mus. Berlin" [printed], "  Dolerus sangunicollis (Klug, 1818) M.Heidemaa det.". Minor damage. 2♀ (same collecting data) labelled as paralectotypes (GBIF-GISHym: 2344, 2345). All in ZMHB.</p></div>	https://treatment.plazi.org/id/59D2DF44FD3956259223EDD0DF6DF0B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F2688D367F7C520AA61D5D2417D58EAB.text	F2688D367F7C520AA61D5D2417D58EAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus schmidti Konow 1884	<div><p>Dolerus schmidti Konow, 1884</p><p>Fig. 16B, D, F, I, L, O, R</p><p>Dolerus schmidti Konow, 1884: 350-351. ♀. Syntypes (assumed). Type locality: Hagen i. W. (Germany). Lectotype designated below.</p><p>Dolerus soniensis Dubois, 1920: 83-86, 94. ♀. Syntypes. Type locality: Uccle ( Forêt de Soignes, La Cambre), Belgium. Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus schmidti Konow, 1884. Lectotype ♀ hereby designated, labelled: "  Dolerus Schmidti m. Hagen i/W.",  “Typus” [red, printed],  “Holotypus” [red, printed],  “GBIF-GISHym4231” [printed], "  Dolerus 199[printed]7  Dolerus schmidti KONOW M. Heidemaa" [handwritten], "Eberswalde coll. DEI", "Coll. Konow", "LECTOTYPE ♀  Dolerus schmidti Konow, 1884 designated M. Heidemaa 2022" [red, printed]. Ovipositor dissected, gummed to a card and pinned with the specimen. SDEI.</p><p>Konow described  D. schmidti as a new species only because  D. liogaster Thomson was not known to him at the time. Later  D. schmidti was treated as a variety of  D. liogaster by Konow (1890b, 1898, 1905b) and synonymized with it by Enslin (1913). The lectotype female was wrongly interpreted as a holotype by Oehlke and Wudowenz (1984) and followed by Heidemaa (2004).</p><p>Dolerus soniensis Dubois, 1920. Lectotype ♀ hereby designated, labelled: "La Cambre Mai", "Collection Dubois", "  D. sonienis ♀ det [handwritten] Ed. Dubois 19 [printed]", "  Dolerus soniensis Dub ♀ [handwritten] Don et  dét . [a gift and the determination by] Ed. Dubois 1920 [printed]", " Type [printed with red ink]" [a pale label with a red frame], "  Dolerus 2000 [printed]  Dolerus schmidti KONOW [handwritten] M.Heidemaa det. [printed]", "  Lectotypus ♀  Dolerus soniensis Dubois, 1920 M.Heidemaa des." [printed white label with a red frame]. Ovipositor dissected and glued on a piece of card, pinned with the specimen. RBINS.</p></div>	https://treatment.plazi.org/id/F2688D367F7C520AA61D5D2417D58EAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
26DD1E9333C1525DB50EF40AF306C797.text	26DD1E9333C1525DB50EF40AF306C797.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus subarcticus Hellen 1956	<div><p>Dolerus subarcticus 
Hellen, 1956</p><p>Fig. 20D</p><p>Dolerus subarcticus Hellén, 1956: 99-100. ♀, ♂. Syntypes. Published type locality: 16 localities in Finnish Lapland, Russian Karelia and Kola Peninsula, and one locality in Siberia (Krasnojarsk oblast, Jenisseisk). Lectotype designated below.</p><p>Dolerus willoughbyi Benson, 1956: 55-58. ♀, ♂. Holotype. Type locality: Abisko (Sweden).</p><p>Type material examined and taxonomic notes.</p><p>Dolerus subarcticus Hellén, 1956. Lectotype ♀ hereby designated, labelled (printed on pale paper if not stated otherwise): "Fennia, Li. Utsjoki, Outakoski, 29.6.1947 [type locality], leg.  Hellén” [printed], "  Lectotypus 2014,  Dolerus subarcticus, ♀  Hellén, 1956, M.Heidemaa design." [red], "  Dolerus subarcticus Hellén, 1956, M.Heidemaa det." [white]. In good condition. 10♀, 8♂ paralectotypes also examined: Ivalo, Kantalaks,  Kilpisjärvi, Lutto, Muonio,  Paanajärvi, Petsamo, Umba, Utsjoki (Outakoski). All in MZH.</p><p>Dolerus willoughbyi Benson, 1956. 2♀, 2♂ paratypes studied. BMNH.</p><p>The melanic (black) form (♂) and a nearly black form, with obscure reddish patches on abdominal terga 2-4 (♀), are known from Norway (near Hovet and near Geitryggtunnelen, leg. E. Heibo, CEH), and the melanic form also from Kamtschatka, Russia (a syntype female of  D. pratensis var. totus determined by R. Malaise, in NHRS, see also under  D. yukonensis).</p></div>	https://treatment.plazi.org/id/26DD1E9333C1525DB50EF40AF306C797	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
A169625F1E8A5801822B7CD36645130C.text	A169625F1E8A5801822B7CD36645130C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus timidus (Klug 1818)	<div><p>Dolerus timidus (Klug, 1818) sp. rev.</p><p>Figs 13C, D, 14A, 14D, 14E</p><p>Tenthredo (Dolerus) timida Klug, 1818: 300. ♀, ♂. Syntypes. Published type locality:  “Deutschland” [Germany]. Lectotype designated below.</p><p>Tenthredo (Dolerus) dubia Klug, 1818: 299-300. ♀, ♂. Syntypes. Type locality: Germany. Lectotype designated below. Primary homonym of  Tenthredo dubia Ström, 1768 [=  Tenthredo (Tenthredella) livida Linnaeus, 1758].</p><p>Dolerus dubius var. atratus Enslin, 1913: 159. ♂. Syntypes (assumed). Type locality: Switzerland. Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Tenthredo (Dolerus) timida Klug, 1818. Lectotype ♂ hereby designated, labelled:  “14199” [printed catalog no.], "Var.  T. timida Kl. [ Klug’s handwriting]", "GBIF-GISHym 2317", "Zool. Mus. Berlin" [printed], "  Lectotypus [printed] ♂  Tenthredo (Dolerus) timida Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]" [red], "  Dolerus timidus (Klug, 1818) M.Heidemaa det." [printed]. Good condition, penis valve dissected, pinned with the specimen (Fig. 14A). Paralectotypes: 4♂ 5♀ (GBIF-GISHym: 2312, 2318, 2320-2326). All in ZMHB.</p><p>Tenthredo (Dolerus) dubia Klug, 1818. Lectotype ♂ hereby designated, labelled:  “14198” [catalog no.], "  Dolerus pratensis (L.) ♂ [handwritten] det. F. Koch [printed]", "GBIF-GISHym 2315" [printed], "Zool. Mus. Berlin" [printed], "  Lectotypus [printed] ♂  Tenthredo (Dolerus) dubia Klug, 1818 [handwritten in block letters] M.Heidemaa design. [printed]" [red], "  Dolerus timidus (Klug, 1818) M.Heidemaa det." [printed], Condition satisfactory, genitalia dissected, penis valves on a slide pinned with the specimen (they fit  D. timidus). Paralectotypes: 1♂, 1♀ ("GBIF-GISHym 2313",  “… 2314"). All in ZMHB.</p><p>Dolerus dubius var. atratus Enslin, 1913. Lectotype ♂ hereby designated, labelled:  “Helvet.[ia]”,  “Type” [pink, printed], "  Dolerus ♂  Dolerus dubius Dolerus dubius var atratus [handwritten] Dr. Enslin det. [printed]", "Sammlung Dr. Enslin" [printed], "GBIF-GISHym 3102", "LECTOTYPE ♂  Dolerus dubius var. atratus Enslin, 1913 designated M. Heidemaa 2022" [red, printed]. Some damage: left flagellum missing, abdomen glued to a card together with the dissected penis valves and pinned with the specimen. ZSM.</p><p>This melanic color form was identified as  D. timidus based on its penis valve structure and the sculpture of the metepimeron.</p></div>	https://treatment.plazi.org/id/A169625F1E8A5801822B7CD36645130C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B3F952C45A855FEA97BFC72C85DFE1B5.text	B3F952C45A855FEA97BFC72C85DFE1B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus varispinus Hartig 1837	<div><p>Dolerus varispinus Hartig, 1837</p><p>Fig. 16G, J, M, P, S</p><p>Dolerus varispinus Hartig, 1837: 239. ♀, ♂. Syntypes. Lectotype ♂ designated by Blank and Taeger (1992). Type locality: Germany [implicit from title of  Hartig’s work].</p><p>Dolerus rugosus Konow, 1884: 281. ♀, ♂. Syntypes. Type locality:  Fürstenberg (Mecklenburg). Lectotype designated below. Junior primary homonym of  Dolerus rugosus Freymuth, 1870.</p><p>Dolerus rugosulus Dalla Torre, 1894: 16. A replacement name for  D. rugosus Konow.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus varispinus Hartig, 1837. Lectotype examined. ZSM.</p><p>Dolerus rugosus Konow, 1884. Lectotype ♂ hereby designated, labelled: "  Dolerus rugosus m [mihi]. fbg. 4.84" ( Fürstenberg / Mecklenburg, IV.1884) [with a double black frame], "  Dolerus rugosus D. T. [Dalla Torre]", "  Syntypus " [red, printed]; "  Lectotypus [printed] ♂  Dolerus rugosus Konow, 1884, design. M. Heidemaa and Viitasaari [printed] 1999 [handwritten]" [red label], "GBIF-GISHym 3770" [printed], "Coll. Konow" [printed], "Eberswalde coll. DEI" [printed], "  Dolerus 1997 [printed]  Dolerus varispinus Hartig [handwritten] M. Heidemaa det. [printed]". SDEI. The taxonomic identity of the lectotype is certain despite its poor condition, because the katepimeron of the mesopleuron is preserved.</p></div>	https://treatment.plazi.org/id/B3F952C45A855FEA97BFC72C85DFE1B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
9EC23B3EC5075F9CA506706C8E8513C1.text	9EC23B3EC5075F9CA506706C8E8513C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus vulneratus Mocsary 1878	<div><p>Dolerus vulneratus 
Mocsary, 1878</p><p>Notes.</p><p>Dolerus vulneratus Mocsáry, 1878: 199. ♀. Syntypes (assumed). Type locality: Siberia. Lectotype ♀ hereby designated, labelled:  “Siber[ia]”, "  Dolerus vulneratus ♀ Mocs.", "Holotypus [printed with red] ♀  Dolerus vulneratus Mocsáry, 1878 Zombori, 1977 [handwritten]", "  Lectotypus [printed] ♀  Dolerus vulneratus Mocsáry, 1878 M.Heidemaa des.  ‘22” [handwritten, red]. "  Dolerus vulneratus Mocsáry, 1878 M.Heidemaa det." [printed]. HNHM.</p><p>The  “holotype” label attached by Zombori has no nomenclatural significance, because this interpretation was never published. Moreover, the number of specimens was not given in the original description and the ♀ symbol does not necessarily indicate that  Mocsáry had only one specimen, even if only one specimen was found in the  author’s collection. Such assumptions about the status of specimens as holotypes should be avoided according to the Code (ICZN 1999: Articles 72.4.7, 73F). The larva is known through rearing and observations by Ponomarev (2022). Host plant: unidentified  Poaceae spec.</p></div>	https://treatment.plazi.org/id/9EC23B3EC5075F9CA506706C8E8513C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
21D834B7210157B29AD0682FAB3C0AD8.text	21D834B7210157B29AD0682FAB3C0AD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dolerus yukonensis Norton 1872	<div><p>Dolerus yukonensis Norton, 1872</p><p>Dolerus similis var. yukonensis Norton, 1872: 82. ♀, ♂. Syntypes. Lectotype ♀ designated by Ross (1931) [not examined]. Type locality: Alaska, USA.</p><p>Dolerus scoticus Cameron, 1881a: 206. ♀. Syntypes (assumed). Lectotype ♀ designated by Benson (1934a). Type locality: Braemar in Aberdeenshire, Scotland.</p><p>Dolerus pratensis var. totus Malaise, 1931b: 15-16. ♀, ♂. Syntypes. Type locality: Kamtschatka ("Klutchi, Petropawlowsk, Elisowo"). Lectotype designated below.</p><p>Type material examined and taxonomic notes.</p><p>Dolerus scoticus Cameron, 1881. Lectotype ♀. BMNH.</p><p>Dolerus pratensis var. totus Malaise, 1931. Lectotype ♀ hereby designated, labelled: "KAMTSCHATKA Malaise" [printed],  “586” [pink, printed],  “Typus” [red, printed], "  D. pratensis var totus n. sp. Type [handwritten] Malaise det. [printed]", "  Dolerus scoticus ♀ Cam [handwritten] A.Zhelochovtsev det 19[printed] 33 [handwritten]", "  Dolerus germanicus totus Mal." [printed and folded], "  Lectotypus [printed]  Dolerus pratensis var. totus Malaise, 1931 M.Heidemaa des. [handwritten in block letters]" [red label], "  Dolerus 2009 [printed]  Dolerus yukonensis [handwritten] M.Heidemaa det. [printed]". NHRS. Paralectotypes: 1♀,  “280” [printed], "  Dolerus 20 [printed] 11  Dolerus subarcticus [handwritten] M.Heidemaa det. [printed]" (a melanic female). NHRS. 1♀, "  Dolerus v. toto [sic!] n. var. Paratypus ♀ Malaise det. [written by R. Forsius?]", "  Dolerus yukonensis Norton M.Heidemaa det.", MZAT (coll. Forsius).</p><p>Dolerus yukonensis has a Holarctic distribution, and both red-banded/-marked as well as melanic color forms occur. According to Goulet (1986) the color forms developed independently in the coastal areas of both continents. This speaks against the subspecific status of the melanic color forms suggested by Benson (1959). According to Benson (1934a)  D. scoticus Cameron can be distinguished from melanic forms of  D. yukonensis by the strongly developed temporal furrows, like in  D. junci and  D. incisus, which are absent in  D. yukonensis . He also wrote that "  Dolerus totus Malaise, 1931, from Kamtchatka may be the same species as  D. scoticus Cam." Benson (1934b) synonymized them (and  D. arcticola Kiaer, 1898) with  D. scoticus and later all of them with  D. yukonensis (Benson 1958: corrigenda). Latterly, he regarded  D. scoticus as a subspecies of  D. yukonensis (Benson 1959) and reported it from Switzerland (Benson 1961) and southern Norway, Hallingskarvet District (Benson 1966). However, Benson was not aware of the presence of melanic forms of  D. incisus and  D. subarcticus in Norway, the males of which can most reliably be identified by their penis valves (Fig. 20C, D). The sculpture of the abdominal terga in  D. yukonensis, as in  D. subarcticus, is not as stable as is often assumed. The depth of the clypeus emargination and the lateral postocellar furrows also vary. However, no potential male of  D. scoticus, which could support its status as a distinct species as proposed by Haris (2000), has so far been recognized. The lectotype female of  D. scoticus probably belongs to  D. yukonensis, and certainly not to  D. junci or  D. incisus . Here we retain the synonymy of  D. scoticus with  D. yukonensis proposed by Benson (1958) and concurred with by Goulet (1986), but admit that more DNA sequence data are needed to test the taxonomic status of the melanic forms, which show at least some coinciding structural differences. We have studied melanic and red-marked color forms from the Palaearctic, but only red-marked forms from the Nearctic.</p></div>	https://treatment.plazi.org/id/21D834B7210157B29AD0682FAB3C0AD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
F5EBCE8408025632806CB53D79897721.text	F5EBCE8408025632806CB53D79897721.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Empria improba (Cresson 1880)	<div><p>Empria improba (Cresson, 1880)</p><p>Fig. 22A-E</p><p>Emphytus improbus Cresson, 1880: 11. ♀, ♂. Syntypes. Type locality: Nevada, USA. Lectotype ♂ (type No. 365) designated by Smith (1979). ANSP.</p><p>Tenthredo (Poecilostoma) hybrida Erichson in:  Ménétriés in: Middendorff, 1851: 60-61. ♀. Syntypes (assumed). Primary homonym of  Tenthredo (Tenthredo) hybrida Eversmann, 1847. Type locality: Udskoj Ostrog [Russia, Khabarovsk Krai, Udskoe]. Lectotype ♀ designated by Prous et al. (2011). ZIN. syn. nov.</p><p>Poecilosoma plana Jakowlew, 1891: 31. ♀. Type locality: Irkutsk, Russia. ♀. Holotype. ZIN. syn. nov.</p><p>Empria itelmena Malaise, 1931b: 23. ♀, ♂. Syntypes. Type locality: Kamtschatka, E[lisowo] [Russia, Kamchatka Krai]. Lectotype ♀ designated by Prous et al. (2011). NHRS. syn. nov.</p><p>Empria camtschatica Forsius, 1928: 46-47. ♀. Holotype. Type locality: Russia, Kamchatka Krai, Bolsheretsk [Bolscheretsk]. MZH. syn. nov.</p><p>Notes.</p><p>The species boundaries between willow-feeding taxa of the  Empria immersa group ( E. immersa,  E. camtschatica,  E. plana, and  E. improba) have proved to be difficult to elucidate (Prous et al. 2014, 2020). In Fennoscandia, two forms can commonly be found at the same time and place (Prous et al. 2014):  E. immersa with a dark pterostigma and short antenna, and  E. camtschatica with a pale pterostigma and long antenna. These two forms can also be distinguished by larval morphology (Fig. 22). Based on ex ovo rearings by M. Prous (two females from Sweden and Estonia) and ex larva rearings by Ponomarev (2022) of  E. immersa, and ex ovo rearings by M. Prous of  E. camtschatica (using two females from Sweden), the main difference seems to be in head coloration:  E. immersa with an occipital fleck or stripe (Fig. 22F-M) and  E. camtschatica with occipital and parietal stripes (Fig. 22A-E). An additional difference may be that glandubae (white conical warts) are more prominent in  E. immersa than in  E. camtschatica . Although based on limited specimen sampling, genome scale data (Prous et al. 2020) support  E. immersa as a distinct species most consistently compared to the other species in the  E. immersa group. In Fennoscandia, taxonomy is complicated by the presence of occasional specimens identifiable as  E. plana, somewhat intermediate in morphology between  E. immersa and  E. camtschatica (pterostigma like  E. camtschatica, saw intermediate). Genome scale data of one  E. plana female from Sweden do not indicate affinity with  E. immersa, but do show at least some affinity with  E. camtschatica (Sweden),  E. improba (Canada), and one other  E. plana (Hokkaido, Japan) (see fig. 5 in Prous et al. 2020). Given the above, we synonymize  E. plana and  E. camtschatica with  E. improba, because clear boundaries between these taxa cannot at present be drawn. Thus, in Europe, the specimens with dark pterostigma, short antennae and more prominent serrulae of the saw can be identified as  E. immersa, and those with pale pterostigma, usually longer antennae, and less prominent serrulae as  E. improba (see Prous et al. 2014). In North America, however, at least some  E. improba specimens look externally more like  E. immersa (dark pterostigma and metafemur), while the serrulae of the saw resemble  E. camtschatica . Lacourt (2020) suggested that  E. camtschatica could be a synonym of  E. improba, but genetically these taxa are not necessarily closer to each other than they are to  E. plana (Prous et al. 2020). If the circumscription of  E. improba as proposed here is considered incorrect, then it remains unclear how many additional species should be recognized, and how these should be delimited. For example, in Europe the morphological distinction between  E. camtschatica and  E. plana is not clear, although these forms can be more reliably distinguished from  E. immersa .</p></div>	https://treatment.plazi.org/id/F5EBCE8408025632806CB53D79897721	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
3352ED13D05652DDBD83037CBBF3F9BA.text	3352ED13D05652DDBD83037CBBF3F9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eutomostethus ephippium (Panzer 1797)	<div><p>Eutomostethus ephippium (Panzer, 1797)</p><p>Tenthredo ephippium Panzer, 1797: 52:5. ♀. Syntypes. Type locality: Germany [according to title of the publication].</p><p>Tomostethus ephippium var. nigrans Konow, 1887a: 275. Sex not stated. Syntypes. Type locality not stated.</p><p>Eutomostethus nigrans (Konow, 1887): Liston et al. (2006)</p><p>Eutomostethus nigrans Blank &amp; Taeger, 1998: 161-162. ♀. Holotype and paratypes. Type locality: Germany, Brandenburg, Luisenfelde, Langer Berg [Grumsiner Forst near  Groß Ziethen]. SDEI. Synonymy with  E. nigrans Konow by Liston et al. (2006).</p><p>Notes.</p><p>COI barcodes of a few specimens identified as  E. nigrans are indistinguishable from those of a large number of  E. ephippium (red-marked females). Vikberg et al. (2011) discussed the distribution of the two forms in Fennoscandia and more widely in Europe, noting that the patterns of occurrence are complex, and that they are regionally often sympatric, but that the dark form becomes more common towards the West, suggesting that climatic causes may play a role in maintaining this polymorphism. No other differences have been detected between the red-marked and black forms. We therefore revert to treating them as conspecific colour forms.</p></div>	https://treatment.plazi.org/id/3352ED13D05652DDBD83037CBBF3F9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
B9F8FBAE747D593F83B52006244A6DBE.text	B9F8FBAE747D593F83B52006244A6DBE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fenusa ewaldi D. R. Smith 2011	<div><p>Fenusa ewaldi D. R. Smith, 2011 (Tenthredinidae, Blennocampinae)</p><p>Figs 36, 37</p><p>Notes.</p><p>A total of about 40♀ and numerous mines. All specimens leg. M. Mutanen, in ZMUO and SDEI:</p><p>Finland: Ostrobottnia ouluensis, Oulu Linnanmaa kasvipuutarha [Oulu Botanic Garden], YKJ grid coordinates: 7219:3427 [65.064N, 25.460E]: adults, 26.06.2018. Larvae in mines on  Rosa, 12.07.2019, Rearing 142/2019. Larvae in mines on  Rosa, 07.08.2019, Rearing 376/2019. Adults, 06.06.2020. Larvae in mines on  Rosa, 06.07.2020, Rearing: 30/2020.</p><p>Ostrobottnia kajanensis, Kuhmo Ulvinsalo, YKJ grid coordinates: 7103:3665 [63.992°N, 30.382°E]: about 20 mines on  Rubus chamaemorus, of which ca 8 inhabited, with one larva in alcohol and four females emerged, collected 10.08.2019, Rearing: 400/2019.</p><p>Fenusa ewaldi was described from two females reared from leaf-mines on  Rosa collected in Novosibirsk, West Siberia (Smith and Altenhofer 2011). Also reared from  Rosa by Ponomarev (2022): Russia, Moscow oblast, Voynovo-Gora, 55.846°N, 39.063°E, 10.07.2020. Since at least 2018, leaf-mines of  Fenusa ewaldi have been frequently found on  Rosa, e.g.  R. woodsii and  R. acicularis in Oulu Botanic Garden, and numerous females reared (Fig. 36). At these three localities, cultivated  Rosa species in parks and gardens were the hosts (Fig. 37A). Mines were found mostly on more sheltered rose bushes. The rearing of four females from leaf-mines on  Rubus chamaemorus (above) was therefore unexpected, considering the habitat was shady, moist, natural old-growth mixed forest dominated by spruce (Fig. 37B).</p></div>	https://treatment.plazi.org/id/B9F8FBAE747D593F83B52006244A6DBE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
3B6FBF67A5925845B575168A46CE3485.text	3B6FBF67A5925845B575168A46CE3485.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilpinia pallida (Klug 1812)	<div><p>Gilpinia pallida (Klug, 1812)</p><p>Lophyrus pallidus Klug, 1812: 54-55. ♀, ♂. Syntypes. Type locality: Sweden and northern Germany. ZMHB.</p><p>Lophyrus catocalus Snellen van Vollenhoven, 1858: 276-277. ♀, ♂. Syntypes. Lectotype ♀ designated by Thomas (1987). Type locality: Netherlands, Groesbeek near Nijmegen. syn. nov.</p><p>Gilpinia catocala: Thomas (1987) recombination.</p><p>Gilpinia verticalis Gussakovskij, 1947: 154-155, 227-228. ♀, ♂. Syntypes. Type locality: localities in Poland, Latvia and Russia. Synonymy with  G. catocala by Thomas (1987).</p><p>Notes.</p><p>Although we have not studied the lectotype of  Lophyrus catocalus, the interpretation of its taxonomic status is possible after referring to  Gussakovskij’s description of  G. verticalis and the other publications cited above. We agree with Pschorn-Walcher (1982) and Viitasaari and Varama (1987) that the two forms are an expression of individual variability, linked to population differences with a geographical component. Accordingly, we treat  Gilpinia catocala as a junior synonym of  G. pallida .</p></div>	https://treatment.plazi.org/id/3B6FBF67A5925845B575168A46CE3485	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
6E84B0EB412057629C5E020BB8A2BDD7.text	6E84B0EB412057629C5E020BB8A2BDD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptamelus viitasaarii Liston, Mutanen & Prous 2022	<div><p>Heptamelus viitasaarii Liston, Mutanen &amp; Prous sp. nov.</p><p>Description.</p><p>Female. Figs 23, 24A-C, 26A, B</p><p>Colour. Fig. 26A, B. Black. Pale are: palpi, tegula, legs (except for arioli and extreme inner tip of metatibia); in fore wing base and apex of costa, apex of subcostal, base of stigma (Fig. 23A), R1, 1A, 2A+3A [color differences between veins of hind wing not clearly definable]; margins of median excision of abdominal tergum 1, medio-distal part of terga 2-4 (-6) [pale area on tergum 3 is the largest of these], all sterna, more or less the downturned parts of terga 2-8, more or less hypopygium and ovipositor valvifer 2. Wing membranes hyaline.</p><p>Head. Pedicel about as long as scape. Head in lateral view with widest point on temple about equal to length of pedicel (Fig. 23E). Malar space slightly shorter than diameter of anterior ocellus. Lowest part of gena densely sculptured, without shining interspaces between the ill-defined pits (Fig. 23E). Clypeus densely pitted, weakly shiny; anterior margin widely emarginate, to about 0.4 of its length (Fig. 23D). Setae on upper head as long, or longer than, diameter of anterior ocellus (Fig. 23E).</p><p>Thorax. Whole pronotum except for small antero-ventral area dull, with sculpture (Fig. 23H). Pits on upper mesepisternum large and well-defined (Fig. 23H). Mesoscutellum with large, scattered pits anteriorly and laterally, interspaces shiny; medially and posteriorly nearly without pits and entirely unsculptured (Fig. 23F). Anterior of mesoscutellar appendage densely pitted, with numerous setae, only small posterior area unpitted (Fig. 23F).</p><p>Abdomen. Anterior of tergum 1 densely setose. Terga 2 and 3 glabrous. Terga increasingly setose from tergum 4 to apex of abdomen. Weak surface sculpture on tergum 2 becoming stronger on more distal terga. Sawsheath in dorsal view broad, with blunt tip (Fig. 24C). In lateral view exposed length of cercus about 0.5  × as long as exposed upper length of valvula 3 (Fig. 23G). Lancet (Fig. 24B, C): 13-14 serrulae; apical serrulae short and high.</p><p>Body length: 4.5-8.0 mm</p><p>Male. Figs 24D, 26C</p><p>Similar to female, except for: red-brown antennal flagellum with basal two antennomeres more or less black; metatibiae completely pale; abdomen completely dark with small, obscure pale markings medially on terga 3-5. Penis valve: Fig. 24D.</p><p>Body length: 4.5-5.5 mm</p><p>Variability: 5-6 flagellomeres, depending on whether or not the distal one is subdivided; the proportions of this antennomere are thus highly variable. Number of pits on upper mesepisternum variable. The scape and pedicel of females may be more or less pale.</p><p>Type material.</p><p>Holotype ♀ (ZMUO.044130). Type locality: Finland, Karelia borealis, Kitee Otravaara, YKJ grid coordinates: 6868:3663 [61.8877N 30.1096E]; 04.08.2019 [larva] in  Athyrium filix-femina (L.), Rearing 380/2019, leg. M. Mutanen. Labels white, printed, except for red holotype label. Deposited in the ZMUO.</p><p>Paratypes. Total: 92♀, 6♂. Finland [leg. M. Mutanen and in ZMUO unless otherwise stated]:</p><p>Karelia borealis, Kitee: Otravaara [Type locality]; 17♀ (including ZMUO.044126, ZMUO.044127, ZMUO.044128, ZMUO.044129, ZMUO.044131, ZMUO.044132, ZMUO.044133, ZMUO.044134, ZMUO.044135, ZMUO.044136), 04.08.2019 [larvae] in  Athyrium filix-femina (L.), Rearing 380/2019. 7♀ (including ZMUO.034957, ZMUO.034958, ZMUO.034959), Otravaara, YKJ grid coordinates: 6868:3663 [61.888°N, 30.110°E], 19.06.2018. 30♀ (including ZMUO.058509, ZMUO.058510, ZMUO.058511, ZMUO.058512, ZMUO.058513, ZMUO.058514, ZMUO.058515, ZMUO.058516, ZMUO.058517, ZMUO.058518, ZMUO.058519, ZMUO.058520), Otravaara, YKJ grid coordinates: 6868:3663 [61.888°N, 30.110°E], 24.08.2020, leg. M. Mutanen and M. Prous. Host:  Athyrium filix-femina, rearing 174/2020. 1♀ (ZMUO.046037), 15.06.2020. ZMUO and SDEI. Otravaara, YKJ grid coordinates: 6868:3664 [61.887°N, 30.129°E]; 8♀ (including ZMUO.034859, ZMUO.034860), 19.06.2018. 1♀ (ZMUO.034995), 18.06.2018.  Pajarinmäki, YKJ grid coordinates: 68899:36670 [62.078°N, 30.197°E]; 1♀ (ZMUO.032885), at light, 11.07.2017 [end date of about 5-day period]. 1♀ (ZMUO.033391), at light, 18.07.2017 [end date of about 7-day period]. 2♀ (including ZMUO.034899), 20.06.2018. 1♂ (ZMUO.061917), 14.06.2021, leg. A. Liston, M. Mutanen, M. Prous. 1♂ (ZMUO.061893), 15.06.2021, leg. N. Kiljunen, A. Liston, M. Mutanen, M. Prous. 4♂ 8 ♀♀ (including ZMUO.060946, ZMUO.060947, ZMUO.060948, ZMUO.060949, ZMUO.060950, ZMUO.060951, ZMUO.060952, ZMUO.060953, ZMUO.060954, ZMUO.060955, ZMUO.058165), 18.06.2021, leg. N. Kiljunen, A. Liston, M. Mutanen, M. Prous. 2 larvae from  Matteuccia struthiopteris (L.) (ZMUO.060557, ZMUO.060558) and 2 larvae from  Athyrium filix-femina (L.) (ZMUO.060561, ZMUO.060562), 26.07.2021, leg. M. Mutanen, M. Prous. Papinniemi, YKJ grid coordinates: 6883:3656 [62.025°N, 29.990°E]; 6♀ (including ZMUO.034761, ZMUO.034762), 18.06.2018, ZMUO and SDEI. 1♀ (ZMUO.040497), 12.06.2019. Puhos, YKJ grid coordinates: 6889:3653 [62.080°N, 29.938°E]; 2♀ (including ZMUO.034968), 20.06.2018. Hiidensaari, YKJ grid coordinates: 6891:3668 [62.092°N, 30.226°E]; 1♀ (ZMUO.034908), 20.06.2018. Potoskavaara, YKJ grid coordinates: 6893:3670 [62.109°N, 30.266°E]; 1♀ (ZMUO.046430), 28.06.2020, leg. Tupu Vuorinen. Satulavaara, YKJ grid coordinates: 6877:3672 [61.964°N, 30.289°E]; 3♀ (including ZMUO.061792, SDEI-GISHym14061), 14.06.2021, leg. A. Liston, M. Mutanen and M. Prous. ZMUO and SDEI.</p><p>Karelia australis: Imatra Kaikkallio, YKJ grid coordinates: 6778:3594 [61.104°N, 28.752°E]; 1♀ (ZMUO.058146), 16.06.2021, leg. A. Liston, M. Mutanen, N. Kiljunen, M. Prous. Joutseno Kuurmanpohja  Sunninmäki [61.071°N, 28.731°E]; 1♀ (ZMUO.016815), 06.06.2015, leg. Jussi Vilen, coll. Matti Viitasaari.</p><p>Tavastia australis: Kangasala Keisarinharju, YKJ grid coordinates: 6818:3347 [61.444°N, 24.141°E]; 1♀ (ZMUO.061964), 08.06.2021.</p><p>Other material.</p><p>Finland: Nylandia,  Vantaa, YKJ grid coordinates: 669:37, 17.06.2021, 1♀, leg. Miikka Friman.</p><p>Etymology.</p><p>Named after Matti Viitasaari, who first recognised a very large female of this species as probably not belonging to  Heptamelus ochroleucus or  H. dahlbomi .</p><p>Habitat.</p><p>Damp, shady places in woodland.</p><p>Biology.</p><p>Many female specimens have been reared from larvae in  Athyrium filix-femina (L.) Roth. (Fig. 25). Larvae found abundantly in 2021 in  Matteucia struthiopteris (L.) Tod. in Kitee, Finland, were provisionally identified as  H. viitasaarii by genetic comparison (identical to adults), as later confirmed when females started to emerge in 2022. The larvae feed inside the rachis, eating alternate portions nearly completely, or only partly (Fig. 25B, C). This results internally in a  “ladder-like” appearance, which is externally clearly visible, especially against the light (Fig. 25A). This feeding habit is, however, possibly widespread in  Heptamelus: at least  H. dahlbomi larvae feed in the same way. Larvae of  H. viitasaarii in  M. struthiopteris regularly overwinter in a chamber made within the fertile fronds at the base of the stem. Strangely, no males have so far been reared, and the few males so far netted are all from a single locality (within an area of less than 1km2). It was observed that adults reared from plants of different size varied very much in body size, and that the smallest individuals had been feeding on the smallest plant, with correspondingly thin stalks.</p><p>Distribution.</p><p>Finland (North and South Karelia, Tavastia, and Helsinki area), Russia (Moscow). The record from Moscow was published by Vikberg (2017) as  Heptamelus ochroleucus .</p><p>Differential diagnosis.</p><p>The characters which are most useful for the identification of European  Heptamelus species are presented below in a key. The extensively pitted and setose mesoscutellar appendage distinguishes  H. viitasaarii from the two other European species, as well as  H. magnocularis Malaise, 1931 (Malaise 1931b) from the Russian Far East (see Liston et al. 2018). Note that all body parts of  H. viitasaarii are conspicuously more densely setose, and pitted, than most other  Heptamelus species which we have so far examined.</p><p>From the females of  Heptamelus dahlbomi,  H. ochroleucus,  H. magnocularis,  H. montanus Togashi, 1961, and  H. takeuchii Togashi, 1961 (the latter two from Japan),  H. viitasaarii differs in its much shorter and higher apical serrulae of the lancet (Fig. 24A, B).  Heptamelus viitasaarii has a wider sawsheath in dorsal view than either  H. dahlbomi or  H. ochroleucus .  Heptamelus japonicus Togashi, 1961, only known in the male sex, differs from  H. viitasaarii in its entirely pale thorax, abdomen and fore wing pterostigma. The male of  H. takeuchii differs from  H. viitasaarii in its more shallowly emarginate and smoother clypeus, and the pale abdominal sternum 9. The male of  H. montanus resembles that of  H. viitasaarii in coloration, but as in females of these species, the clypeus of  H. montanus is much less deeply emarginate (approx. to about 0.2 of its length) and the setae on the upper head shorter (mostly shorter than the diameter of the anterior ocellus).</p></div>	https://treatment.plazi.org/id/6E84B0EB412057629C5E020BB8A2BDD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
AEF2BB47C0CE56318FD8399B02C100DD.text	AEF2BB47C0CE56318FD8399B02C100DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrophya infumata Rohwer 1925	<div><p>Macrophya infumata Rohwer, 1925 (Tenthredinidae, Tenthredininae)</p><p>Fig. 38</p><p>Notes.</p><p>Finland: Karelia borealis, Kitee: 1♂, Puhos [62.082°N, 29.935°E], 30.06.2017, leg. Juha Salokannel. 1♀,  Pajarinmäki, 16.06.2020, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. Jaakko  Pohjoismäki and M. Mutanen. 3♂,  Pajarinmäki, 18.06.2021, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. A. Liston, M. Mutanen, N. Kiljunen, M. Prous. Approximately 30 larvae, on  Sambucus racemosa,  Pajarinmäki, 26.07.2021, YKJ grid coordinates: 68898:36667 [62.078°N, 30.191°E], leg. M. Mutanen and M. Prous. All specimens in ZMUO (Fig. 38).</p><p>In Europe previously recorded from the Russian regions of Kirov and Perm (Zhelochovtsev and Zinovjev 1996), but in iNaturalist (2022) more western Russian records are to be found (Kursk, Moscow, and St Peterburg Regions). The species is widespread in the East Palaearctic, from W. Siberia to Sakhalin, Japan, and northern China (Mallach 1936; Yoshida 2017). Host plant is  Sambucus racemosa, including the closely related Japanese  S. sieboldiana (Sakurai et al. 2009). The larvae were beaten from lower branches of  S. racemosa in a shady habitat.</p></div>	https://treatment.plazi.org/id/AEF2BB47C0CE56318FD8399B02C100DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
C35D4AB78D9A5A8DBEC59393502A26D9.text	C35D4AB78D9A5A8DBEC59393502A26D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phymatoceropsis gracilicornis (Zaddach 1859)	<div><p>Phymatoceropsis gracilicornis (Zaddach, 1859) comb. nov.</p><p>Notes.</p><p>Selandria gracilicornis Zaddach, 1859: 34-35. ♀. Holotype. Type locality [see Introduction, p. 7]: probably  Königsberg [= Russia, Kaliningrad].  Blennocampa gracilicornis:  André (1881).  Monophadnus gracilicornis: Kirby (1882).  Rhadinoceraea gracilicornis: Konow (1886a).  Dicrostema gracilicornis: Benson (1952).  Paracharactus gracilicornis: Smith (1969).</p><p>The host plant of  P. gracilicornis is  Adoxa moschatellina L. ( Adoxaceae) (Chambers 1947).</p></div>	https://treatment.plazi.org/id/C35D4AB78D9A5A8DBEC59393502A26D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
4A2976EA8ABB5F0791F772F7E338C1DF.text	4A2976EA8ABB5F0791F772F7E338C1DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phymatoceropsis Rohwer 1916	<div><p>Phymatoceropsis Rohwer, 1916</p><p>Phymatoceropsis Rohwer, 1916: 107-109. Type species:  Phymatoceropsis fulvocincta Rohwer, 1916, by original designation.</p><p>Dicrostema Benson, 1952: 97-98, 101. Type species:  Selandria gracilicornis Zaddach, 1859, by original designation. Smith (1969): synonymy with  Paracharactus. syn. nov.</p><p>Description.</p><p>Antennal flagellomeres proportionately narrow; flagellomere 1 3.2-4.7  × as long as distal width; the basal flagellomeres not widening distally, and setae normal (not long and coarse as in  Phymatocera). Outer orbit with pronounced groove behind nearly whole length of eye. Postgenal carina developed slightly below eye (clearly in  P. sibiricola, but very weakly in  P. gracilicornis). Posterior of mesoscutellum with some conspicuous pits. Epicnemium variably developed, e.g. present in  P. sibiricola, absent in  P. gracilicornis . Claws with a small to minute inner tooth. Stub of 2A + 3A ( Astatus analis) of fore wings straight, curved towards anterior, or furcate at apex; hind wing with enclosed cell M.</p><p>Diagnosis.</p><p>Phymatoceropsis can be distinguished from  Rhadinoceraea by its proportionately narrower flagellomeres (flagellomere 1 3.2-4.7  × as long as distal width in  Phymatoceropsis, 2.6-3.1  × as long as distal width in  Rhadinoceraea); basal flagellomeres not distally widened (widened in  Rhadinoceraea); outer orbit with pronounced groove behind whole length of eye (in  Rhadinoceraea, if a groove is present, then this is behind only part of the eye.  Phymatoceropsis differs from  Lagonis in its mainly smooth mesepisternum (upper mesepisternum of  Lagonis with numerous, large, crater-like pits).  Phymatoceropsis can be separated from  Paracharactus (based on North American species and the European  P. hyalinus) by the mesoscutellum having at least a row of deep, well-defined pits on the posterior part (mesoscutellum entirely without pits in  Paracharactus). Although in all examined specimens of  Phymatoceropsis the stub of 2A + 3A of the fore wing is apically furcate (as is also usual in  Rhadinoceraea), and in most specimens of various Nearctic  Paracharactus and the European  P. hyalinus (Konow, 1886) (Konow 1886a) the apex of 2A + 3A is straight, in some individuals of Nearctic  Paracharactus species, as already noted by Smith (1969), it is furcate / curved strongly upwards, e.g. in  P. rudis (Norton, 1861).</p><p>Comments.</p><p>The phylogeny of the  Blennocampinae (Fig. 27) requires additional study, including analysis of genetic data obtained from a larger number of taxa. The large number of genus names currently in use as valid (Taeger et al. 2010 listed over 100), and their often weak morphological characterization, lead us to suspect that significant  “oversplitting” may have occurred. However, at present it seems reasonable to retain  Phymatoceropsis as valid, and to place  P. sibiricola and  P. gracilicornis there. The synonymy of  Phymatoceropsis and  Dicrostema is based on the close genetic similarity of the type species of  Dicrostema to  Phymatoceropsis sibiricola, and because these two species possess the same combination of characters exhibited by other  Phymatoceropsis species, including its type species. The host plant ranges of lineages of the Phymatocerini may correlate to a certain degree with their phylogeny. As far as is known,  Rhadinoceraea species are attached to  Iridaceae and  Liliaceae (Smith 1969), and  Phymatoceropsis species to  Adoxaceae, i.e.  P. gracilicornis on  Adoxa, and  P. japonica and  P. sibiricola on  Sambucus . Interestingly,  Sambucus species are also the hosts of  Lagonis nevadensis, and  L. opacicollis, which genetically also group with  Phymatoceropsis, albeit with weak statistical support (Fig. 27). Unfortunately, the hosts of  Paracharactus species are not known for certain. Smith (1969) reasoned that North American species might be attached to  Carex, based on a tentative identification of a larva, whereas Lacourt (1985) speculated that the West Palaearctic  P. hyalinus feeds on  Ranunculus aconitifolius and  R. platanifolius, without presenting any supporting data or observations. On the other hand, Okutani (1967) recorded the East Palaearctic  Paracharactus leucopodus Rohwer, 1910 from  Smilax ( Liliales,  Smilacaceae), but confirmation of this would be desirable. Both European  Phymatoceropsis species are univoltine, as are probably nearly all Phymatocerini. Possible exceptions are  Eurhadinoceraea ventralis (Severin 1997) and  Phymatocera aterrima (Chevin and Silvestre de Sacy 2001), but the prolonged phenological periods of activity recorded for these species may be the result of polymodal emergence of adults, rather than true plurivoltinism.</p></div>	https://treatment.plazi.org/id/4A2976EA8ABB5F0791F772F7E338C1DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
7EC34ECDD8EF59C1A26469F2E7DF75C2.text	7EC34ECDD8EF59C1A26469F2E7DF75C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phymatoceropsis sibiricola (Zhelochovtsev 1939)	<div><p>Phymatoceropsis sibiricola (Zhelochovtsev, 1939) comb. nov.</p><p>Figs 28, 29</p><p>Notes.</p><p>Rhadinoceraea sibiricola Zhelochovtsev, 1939: 159-160. ♀. Holotype. Type locality: Russia, Krasnoyarsk Krai, Vostotschnoe (35 km SE of Minussinsk).</p><p>In the original description, Zhelochovtsev (1939) compared  R. sibiricola with "  Rh. japonica Malaise" [ Phymatoceropsis japonica (Malaise, 1931) (Malaise 1931a)], and stated that they are "closely allied". Comparison of female  P. sibiricola specimens from Finland (Fig. 28) with a Japanese female of  P. japonica in the SDEI collection confirmed their similarity. As mentioned by Zhelochovtsev,  P. japonica has much longer antennae, but the differences described in the structure of the frontal area and mesopleura seemed less clear. The host plant of  P. sibiricola, as discovered by Vikberg in Finland (see below), is  Sambucus racemosa ( Adoxaceae), since verified several times by Mutanen. The host plant of  P. japonica is  Sambucus sieboldiana Blume (Okutani 1956), which is sometimes treated as a synonym or subspecies of  S. racemosa L.</p><p>Summary of records in Finland</p><p>[not all more recent records are listed].</p><p>Uusimaa:  Mäntsälä, Saari, 67368:34139 [60.732°N, 25.423°E], 1♀ 18.05.2010, leg. Iiro Kakko (in his collection in  Hämeenlinna).</p><p>South  Häme: Janakkala,  Laurinmäki, 67526:3695 [60.862°N, 24.599°E], 1♀ 01.06.2009, 1♀ 26.05.2010, 1♀ 31.05.2010, leg. V.Vikberg (VVT). Janakkala,  Hangastenmäki, 67525:3692 [60.861°N, 24.594°E], 1♀ 21.05.2011, 2♀ 22.05.2011, 2♀ 24.05.2011, 1♀ 31.05.2011, 1♀ 16.05.2012, 1♂ 21.05.2013, 1♀ 29.05.2013; in 2014-2021 females were collected every year and a few males altogether, leg. V.Vikberg (VVT). One male and one female collected in 2013 were photographed by Pekka Malinen http://id.luomus.fi/GL.7655 (♂), http://id.luomus.fi/GL.7656 (♀).</p><p>North  Häme:  Jyväskylä . Vikberg identified two females in photographs taken in two different years by Raimo Pelkonen. The last year was 2019, but the first year much earlier.</p><p>South Karelia: Imatra,  Räikkölä, 61.121°N, 28.790°E, 1♀ 30.05.2021, 1♀ 01.06.2021, leg. M. Mutanen &amp; M. Prous.</p><p>Karelia borealis: Kitee,  Pajarinmäki, 62.075°N, 30.186°E, 1♀ 22.05.2016, leg. M. Mutanen; 1♀ 06.06.2016, leg. M. Mutanen.  Pajarinmäki, 62.078°N, 30.191°E, 1♀ 14.05.2018, leg. M. Mutanen; 1♀ 15.05.2018, leg. M. Mutanen; 1♀ 4♂ 20.06.2018, leg. M. Mutanen; 24♀ 1♂ 17.06.2020, leg. M. and I. Mutanen; 1♀ 18.5.2021, leg. M. Mutanen &amp; M. Prous. Papinniemi, 62.025°N, 29.990°E, 1♀ 16.05.2018, leg. M. Mutanen. Puuteniemi, 62.167°N, 29.970°E, 1♀ 1♂ 15.05.2018, leg. M. Mutanen. Potoskavaara, 62.109°N, 30.266°E, 1♀ 15.05.2018, leg. T. Vuorinen. Puhos, 62.089°N, 29.958°E, 2♀ 18.06.2018, leg. M. Mutanen. Puhos, 62.089°N, 29.939°E, 1♀ 05.06.2019, leg. M. Mutanen.  Tyynelä, 62.080°N, 30.321°E, 1♀ 13.05.2019, leg. M. Mutanen. Vanhahovi, 62.169°N, 29.974°E, 2♀ 15.05.2019, leg. M. Mutanen.</p><p>Tavastia australis:  Hämeenlinna, Hattelmalanharju, 60.971°N, 24.475°E, 1♀ 15.05.2013, leg. M. Raekunnas (ZMUO).</p><p>Biology.</p><p>Oviposition trial by Vikberg (3/2011 VV): three females captured in  Hangastenmäki in 2011 were used in this experiment.  Adoxa,  Sambucus racemosa,  Lonicera xylosteum and  Galium sp. were offered to them. They laid eggs only on  Sambucus racemosa leaves. The egg is laid through the upperside of the leaf into the tissue of the underside, usually one to two eggs per leaflet, but in one small leaflet eight eggs were counted. Oviposition occurred on 21.05.-24.05. The first larvae were observed on the morning of 28.05. Later, 28 larvae were counted. Five feeding instars were observed and after finishing feeding there was an "extra moult". Prepupae were seen on 07.06-08.06.</p><p>1st and 2nd instar larvae are gray, with a brown head. 5th instar larvae are 13-14 mm long; head width ca. 1.6 mm (Fig. 29). Head brownish, behind the eye a black fleck which is curved backwards and upwards. Body dorsally dark gray with greenish hue; ventrally whitish gray. Black suprastigmal flecks on thoracic segment 3 and abdominal segments 1-9.</p></div>	https://treatment.plazi.org/id/7EC34ECDD8EF59C1A26469F2E7DF75C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
4D49FDB220E1535699067E44863869D5.text	4D49FDB220E1535699067E44863869D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pristiphora coactula (Ruthe 1859)	<div><p>Pristiphora coactula (Ruthe, 1859)</p><p>Nematus coactulus Ruthe, 1859: 307-308. ♀. Holotype. Type locality: Iceland.</p><p>Lygaeonematus (Lygaeotus) trochantericus Lindqvist, 1952: 101-102. ♀. Holotype (http://id.luomus.fi/GL.7708) and paratypes (♀, ♂). Type locality: Finland, Utsjoki, Outakoski. syn. nov.</p><p>Notes.</p><p>The nuclear sequence data obtained for this study revealed three main clusters within the  Pristiphora carinata group:  P. carinata,  P. coactula, and  P. borea +  P. groenblomi +  P. albilabris (Fig. 30). No nuclear sequence data are yet available for  P. breadalbanensis (Cameron, 1882b) and  P. lativentris (Thomson, 1871). A nearly perfect match morphologically to the  L. trochantericus holotype is ZMUO.035514, which falls within the  P. coactula cluster based on nuclear DNA (Fig. 30). There are two main clusters based on COI sequences, one of which contains only  P. borea (Konow, 1904) and  P. groenblomi (Lindqvist, 1952) and the other one all species (Fig. 31). Within the COI cluster containing all species (Fig. 31),  P. borea,  P. groenblomi, and  P. albilabris (Boheman, 1852) ( Betula feeders) tend to separate from  P. coactula ( Salix) and  P. carinata (Hartig, 1837) ( Vaccinium). Based on the specimens having nuclear data, the species (mainly females) of the  Pristiphora carinata group may be separated by the following key, although it might not always work for all specimens, particularly  P. coactula and  P. borea . Excluded from the key are the (sub)arctic species  P. breadalbanensis and  P. lativentris .  Pristiphora Pristiphora lativentris may have somewhat different serrulae from the other species (almost papilliform, see fig. 215 in Prous et al. 2017). The identity of  Pristiphora breadalbanensis (most similar to  P. borea and  P. coactula) needs further research to confirm if characters (e.g. structure of median mesoscutal lobes) mentioned by Benson (1958) to separate this species are reliable.</p><table><tr><td colspan="1" rowspan="1">1</td><td colspan="1" rowspan="1">a Pterostigma distinctly darker than costa b Legs largely orange or reddish c In female, valvifer 2 and terga 9-10 black or slightly pale d In male, sternum 9 blac</td><td colspan="1" rowspan="1">P. albilabris (Boheman, 1852) ♂♀ and  P. groenblomi (Lindqvist, 1952) ♂♀ in part</td></tr><tr><td colspan="1" rowspan="1">-</td><td colspan="1" rowspan="1">aa Pterostigma similarly pale as costa or somewhat darker than costa bb Legs largely black to pale cc In female, valvifer 2 and terga 9-10 extensively pale dd In male, sternum 9 black to pale</td><td colspan="1" rowspan="1">2</td></tr><tr><td colspan="1" rowspan="1">2(1)</td><td colspan="1" rowspan="1">a Pterostigma somewhat darker than costa b Legs largely orange or reddish c Metafemur completely pale d In female, terga 2-8 and sterna black e In male, sternum 9 (always?) black</td><td colspan="1" rowspan="1">P. groenblomi ♂♀ in part</td></tr><tr><td colspan="1" rowspan="1">-</td><td colspan="1" rowspan="1">aa Pterostigma similarly pale as costa bb Legs largely black to yellowish cc Metafemur black to pale dd In female, terga 2-8 and sterna black or partly pale (starting from tergum 2 and sternum 2) ee In male, sternum 9 black to pale ... ♀ (males of the following species not separated)</td><td colspan="1" rowspan="1">3</td></tr><tr><td colspan="1" rowspan="1">3(2)</td><td colspan="1" rowspan="1">a Valvula 3 in dorsal view gradually narrowing, without invagination and with sharp tip (see figs 98-99 in Prous et al. 2017) b Usually only terga 8-10 or 9-10 extensively pale, but sometimes more (starting from tergum 5)</td><td colspan="1" rowspan="1">P. carinata (Hartig, 1837)</td></tr><tr><td colspan="1" rowspan="1">-</td><td colspan="1" rowspan="1">aa Valvula 3 in dorsal view more or less truncate, with or without indistinct invagination and with broader tip bb Usually terga 8-10 or more (starting from tergum 2) at least partly pale</td><td colspan="1" rowspan="1">4</td></tr><tr><td colspan="1" rowspan="1">4(3)</td><td colspan="1" rowspan="1">a Valvula 3 short, truncate and usually with indistinct invagination (Fig. 33C, D) b Abdomen usually becoming gradually paler from base to apex, dorsally usually starting from tergum 7, laterally and ventrally from tergum 2 and sternum 2 c Metafemur usually completely pale d Clypeus usually mostly pale</td><td colspan="1" rowspan="1">P. coactula (Ruthe, 1859)</td></tr><tr><td colspan="1" rowspan="1">-</td><td colspan="1" rowspan="1">aa Valvula 3 usually longer, slightly narrowed at apex and without invagination (Fig. 33A), but sometimes not distinguishable from  P. coactula (Fig. 33B) bb Abdomen usually slightly or extensively pale only at apex, dorsally usually terga 8-10, laterally usually terga 7-10, ventrally usually sternum 7 cc Metafemur black to completely pale dd Clypeus mostly black to mostly pale</td><td colspan="1" rowspan="1">P. borea (Konow, 1904)</td></tr></table><p>Examples of lancets of  P. borea,  P. carinata, and  P. coactula are shown in Fig. 32A-C, but more specimens need to be examined to check if there are any consistent differences between the species. Morphological differences between the males of  P. borea,  P. carinata, and  P. coactula are not clear. Externally, it seems that  P. coactula tends to be paler (clypeus, pronotal angles, tegula, metafemur, and sternum 9 completely or mostly pale) than  P. borea and  P. carinata . The dorso-apical margin of the paravalva of  P. borea (Fig. 34B) may be more strongly inclined basally compared to  P. carinata and  P. coactula, but differences between the penis valves of the latter two species are not clear (Fig. 34A, D, E). The most distinctive penis valve in the  P. carinata group seems to belong to  P. albilabris (Fig. 34F), which has the most distinctly inclined dorso-apical margin of paravalva. Overall shape of penis valve of  P. groenblomi (Fig. 34C) is most similar to  P. borea, but it may be larger.</p></div>	https://treatment.plazi.org/id/4D49FDB220E1535699067E44863869D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
855A00D5977850258AAB3FF0FBC60D02.text	855A00D5977850258AAB3FF0FBC60D02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stromboceros koebelei Rohwer 1910	<div><p>Stromboceros koebelei Rohwer, 1910 (Tenthredinidae, Selandriinae)</p><p>Fig. 39</p><p>Notes.</p><p>Finland: Regio kuusamoensis, Kuusamo Uopajanpuro, YKJ grid coordinates: 7363:3613 [66.342°N, 29.534°E]: 1♂, 2♀, 13.06.2018, leg. Marko Mutanen. 4♂, 17.06.2019, leg. Iina Eskelinen and Riikka Jarkko. 15♂ [approximately], 1♀, 20.06.2020, leg. Marko Mutanen. All specimens in ZMUO.</p><p>The only previous records in the West Palaearctic were from Russia: Zhelochovtsev (1951) mentioned Archangelsk oblast,  “Molotowsk” [since 1956 Severodvinsk], and Ermolenko (1975) wrote [translated] "distributed from Perm and the Urals to Sakhalin and Japan". Zhelochovtsev and Zinovjev (1996) mentioned only  “Ural” as the area of occurrence within European Russia. Without locating voucher specimens, it is therefore impossible to decide whether the published distributional information for European Russia is accurate.  Stromboceros koebelei occurs in the Russian Far East and Amur Region, but records are lacking from more western parts of the Russian East Palaearctic territories (Zhelochovtsev and Zinovjev 1996). It is also widespread in Japan, and occurs on the Korean Peninsula (Yoshida 2017). In Japan, species of  Athyrium (Isaka et al. 2015),  Dryopteris, and  Polystichum (Naito 1979) have been recorded as host plants. The only known Finnish site for the species is a shady creek valley with mixed trees and rich vegetation. All specimens were swept from  Dryopteris expansa (C. Presl) Fraser-Jenk. and Jermy, which is almost certainly the food plant at this site. Other fern species present on the site, including  Matteucia struthiopteris, did not yield any adults. It is noticeable that all specimens from several other localities in the same region have turned out to represent the common  S. delicatulus, suggesting that  S. koebelei is very local and possibly has a narrow host range.</p></div>	https://treatment.plazi.org/id/855A00D5977850258AAB3FF0FBC60D02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liston, Andrew;Mutanen, Marko;Heidemaa, Mikk;Blank, Stephan M.;Kiljunen, Niina;Taeger, Andreas;Viitasaari, Matti;Vikberg, Veli;Wutke, Saskia;Prous, Marko	Liston, Andrew, Mutanen, Marko, Heidemaa, Mikk, Blank, Stephan M., Kiljunen, Niina, Taeger, Andreas, Viitasaari, Matti, Vikberg, Veli, Wutke, Saskia, Prous, Marko (2022): Taxonomy and nomenclature of some Fennoscandian Sawflies, with descriptions of two new species (Hymenoptera, Symphyta). Deutsche Entomologische Zeitschrift 69 (2): 151-218, DOI: http://dx.doi.org/10.3897/dez.69.84080, URL: http://dx.doi.org/10.3897/dez.69.84080
