taxonID	type	description	language	source
03FD87925833FFC7FF56FADD82F642A1.taxon	description	Figs 1 – 16, 82 – 85, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925833FFC7FF56FADD82F642A1.taxon	type_taxon	Type species. Borniella parva spec. nov.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925833FFC7FF56FADD82F642A1.taxon	etymology	Etymology. The generic name is derived from the name of the island Borneo, where the type species was recorded. The gender is feminine.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925833FFC7FF56FADD82F642A1.taxon	diagnosis	Diagnosis. Males can be distinguished from other genera of the Heteropodinae by the RTA proximally arising from palpal tibia in combination with the presence of three tegular apophyses and the absence of an embolic apophysis (present in most Sinopoda species) (Figs 1 – 3). Females are similar to those of Sinopoda and Menarik gen. nov. in having a medially fused internal duct system with anteriorly situated glandular appendages (Figs 4 – 7) but can be distinguished from Menarik gen. nov. by lobal septum present with two separate copulatory openings (septum absent and with copulatory atrium in Menarik gen. nov., Figs 23 – 26); from Sinopoda by 1. Copulatory openings distinctly separated from epigynal pockets, those latter developed as shallow pockets only in anterior half, but as rims only in posterior half, and 2. Connecting ducts between anterior and posterior part of internal duct system not freely visible in dorsal view, but running ventrally and hidden behind internal structures (copulatory openings always connected directly or indirectly with epigynal pockets or connecting rims and connecting part of internal duct system distinctly visible in dorsal view, i. e. such internal structures absent in Sinopoda species). Notes. Within the genus Sinopoda there is a huge range of variation in the female copulatory organ (for example in S. chiangmaiensis Grall & Jäger, 2020 or S. scurion Jäger, 2012). In contrast, the male palps show only little variation of the ground pattern. In males of all Sinopoda species described so far, the RTA arises from tibia distally and it is divided into a dorsal and ventral branch (exceptions are few species of the okinawana - group). This is true even for species with uncommon female copulatory organs like S. arboricola Grall & Jäger, 2020 or S. scurion Jäger, 2012. Additionally, the embolus possesses a distinct apophysis in almost all Sinopoda spp. In those species of the okinawana - group with emboli without such an apophysis, the RTA arises distally from the tibia, and the embolus arises from tegulum in a much more proximal position, and the tegulum lacks any additional apophysis. Females are in fact superficially similar to those of the genus Sinopoda, but functional epigynal pockets are situated in this new genus exclusively in the anterior half. In Sinopoda spp., in contrast, functional epigynal pockets are longer, running from postero-lateral to antero-medial (in most of the species). In any case the copulatory opening is situated in the pockets and not visible in ventral view as in the new genus.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925833FFC7FF56FADD82F642A1.taxon	description	Description. See description of type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925833FFC7FF56FADD82F642A1.taxon	distribution	Distribution. Northern Borneo (Malaysia: Sarawak; Brunei) (Fig. 98: green circles). Species included. Only the type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	description	Figs 1 – 16, 82 – 85, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	materials_examined	Type material. MALAYSIA: Sarawak Province: Holotype male (PJ 1227): Mulu Expedition, Camp 1, 4 ° 3 ’ 2.88 ” N, 114 ° 52 ’ 1.20 ” E, ca. 330 m, MDF, leaf litter near waterfall, 24 May 1978, F. Wanless (NHM). Paratypes (4 females): 2 females (PJ 1228 – 1229) with same data as for holotype (NHM; 1 female: SMF). 2 females (PJ 1204 – 1205) with same data as for holotype, except: shrub layer, 20 June 1978 (NHM). Additional material examined (14 females). MALAYSIA: Sarawak Province: 9 females with same data as for holotype, but: 1 female (PJ 1221), Tapin river, 4 ° 3 ’ 35.25 ” N, 114 ° 51 ’ 47.83 ” E, ca. 100 m, waterside, vegetation, 5 June 1978 (NHM); 3 females (PJ 1218 – 1220), 150 m, Plot 2, forest leaf litter (NHM); 5 females (SD 1449, PJ 1222 – 1226) 2 March 1978, H. W. Vallame (NHM). BRUNEI: Temburong: 2 females, Ulu Temburong National Park, Kuala Belalong Field study Centre, ca. 4 ° 32 ’ 49.31 ” N, 115 ° 9 ’ 20.36 ” E, 147 m, 1 – 25 February 2013, O. Machac (SMF).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	etymology	Etymology. The species name is derived from the Latin word “ parvus ” meaning “ small ” and refers to the small body size of this species; adjective.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	diagnosis	Diagnosis. Small Sparassidae with TL <4.5 (Figs 82 – 85). Borniella parva spec. nov. may be recognised by the following combination of characters: Males (Figs 1 – 3): 1. Single RTA long, reaching distally to retrolateral cymbial bulge, 2. Embolus arising from tegulum at 9 - o’clock-position, running roughly a semicircle and 3. Distal TA largest with small tooth ventrally, median TA mid-range in size, triangular in prolateral view, proximal TA smallest, acute triangular, situated close to spermophor winding. Females (Figs 4 – 7): 1. Posterior part of vulva with spermathecae distinctly wider than anterior part with glandular appendages and 2. Copulatory openings with roughly right-angled rims.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	description	Description. Male (holotype): TL 3.3; PL 1.7, PW 1.7, AW 0.95; OL 1.6, OW 1.1. Eye measurements (Fig. 8): AME 0.11; ALE 0.16; PME 0.10; PLE 0.22; AME – AME 0.06; AME – ALE 0.01; PME – PME 0.15; PME – PLE 0.18; AME – PME 0.15; ALE – PLE 0.16; clypeus AME 0.14; clypeus ALE 0.13. PLE on small humps. Measurements of palp and legs: palp: 2.55 (0.75, 0.40, 0.50, 0.90); I: 6.15 (1.65, 0.80, 1.60, 1.50, 0.60); II: 7.85 (2.20, 0.90, 2.10, 1.95, 0.70); III: 6.60 (1.90, 0.80, 1.70, 1.60, 0.60); IV: 6.70 (1.90, 0.70, 1.70, 1.75, 0.65). Leg formula: 2431. Spination: palp: 131, 101, 2110; legs: Fe I 322, Fe II & III 222, Fe IV 231; Pa I – IV 000; Ti I & II 1016, Ti III & IV 2226; Mt I & II 0006, Mt III 2036, Mt IV 3036. Chelicerae with 3 promarginal, 5 retromarginal teeth, 13 – 14 denticles in patch close to promarginal teeth and 1 escort seta (Fig. 13). Serrula with 24 denticles. Prolateral claw of leg II with 17 teeth (Fig. 10). Trilobate membrane with narrow median hook, slightly shorter than lateral projections (Fig. 15). Palp as in diagnosis (Figs 1 – 3). RTA long and tapering, almost straight in ventral view, moderately curved in retrolateral view. Conductor short and slender, arising at 12.30 - o’clock-position from tegulum. Retrolateral part of tegulum kidney-shaped. Spermophor running retrolaterally submarginally along tegular margin, with distinct prolaterad U-turn proximally and another retrolaterad U-turn following directly after that. Embolus slender and semicircular. Distal tegular apophysis sail-shaped, connected directly with its prolateral rim to the embolus base. Colouration in ethanol (Figs 82 – 83): Prosoma yellowish-white, posterior and lateral margins with brown band, medio-anteriorly light brown semi-circular pattern, medially with brown longitudinal stripe along fovea. Sternum yellowish-white. Chelicerae yellow-brown. Coxa I – III medio-dorsally with brown stripe. Palps and legs yellowish-white. Opisthosoma yellowish-white, laterally and posteriorly with brown spots, ventral part white, posteriorly brown. Female (paratype): TL 3.7; PL 1.8, PW 1.7, AW 1.0; OL 1.9, OW 1.2. Eye measurements (Fig. 9): AME 0.10; ALE 0.16; PME 0.09; PLE 0.19; AME – AME 0.10; AME – ALE 0.03; PME – PME 0.15; PME – PLE 0.21; AME – PME 0.15; ALE – PLE 0.19; clypeus AME 0.15; clypeus ALE 0.13. PLE on small humps. Leg formula: 2431; measurements of palp and legs: palp: 2.90 (0.80, 0.50, 0.70, 0.90); I: 5.85 (1.60, 0.70, 1.55, 1.45, 0.55); II: 7.25 (2.10, 0.80, 2.00, 1.75, 0.60); III: 6.20 (1.80, 0.70, 1.65, 1.50, 0.55); IV: 6.45 (1.80, 0.65, 1.70, 1.70, 0.60). Leg formula: 2431. Spination: palp: 131, 101, 2221, 1004; legs: Fe I 322, Fe II & III 222, Fe IV 221; Pa I – IV 000; Ti I & II 1016, Ti III 2006, Ti IV 2226; Mt I & II 0006, Mt III 1016, MT IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 14 – 15 denticles in patch close to promarginal teeth and 1 escort seta (Fig. 14). Serrula with 26 denticles. Palpal claw with 5 long and 2 – 3 short teeth (Fig. 11). Retrolateral claw of leg III with 13 teeth (Fig. 12). Trilobate membrane with narrow median hook, almost as long as lateral projections (Fig. 16). Copulatory organ as in diagnosis (Figs 4 – 7). Epigynal field oval, wider than long, without slit sensilla or anterior bands. Internal duct system wider than long. Posterior part of internal duct system wider than anterior part. Fertilization ducts short and slender, arising postero-laterally. Membranous structure covers posterior part of internal duct system. Colouration in ethanol (Figs 84 – 85): Prosoma yellowish-white, medio-anteriorly with semi-circular light brown pattern, medially with brown longitudinal stripe, posterior and lateral margin brown. Sternum white. Opisthosoma greyish-white with brown spots, posteriorly brown, ventral part greyish-white. Chelicerae yellowish-brown. Coxa I – III medio-dorsally with brown stripe. Palps yellowish-white. Legs yellowish-white; Fe proximally light brown; Pa ventrally light brown. Variation female (n = 14). TL 3.25 – 4.1; PL 1.65 – 1.9, PW 1.6 – 1.8, AW 1.0 – 1.1; OL 1.6 – 2.3, OW 0.9 – 1.7. Measurements leg I: total length 5.62 – 6.20, Fe 1.50 – 1.70, Pa 0.70 – 0.80, Ti 1.45 – 1.60, Mt 1.35 – 1.50, Ta 0.55 – 0.60. Spination: palps: Ti 2121; Mt 1014; legs: Fe II & III 322; Ti III 1016, 2026; Mt III 1006, 2006, 2016, 2026, 3026.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925830FFC1FF56FD5E84764269.taxon	distribution	Distribution. Borneo (Malaysia: Sarawak; Brunei) (Fig. 98: green circles).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	description	Figs 17 – 46, 86 – 89, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	type_taxon	Type species. Menarik kecil spec. nov.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	etymology	Etymology. “ Menarik ” is the Malayan word for “ attractive, fascinating, interesting ” and refers to the specific tegular apophysis and retrolateral tibial apophysis of the male, the unique epigynal atrium of the female as well as to the enigmatic scars around the female epigyne. The gender is masculine.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	diagnosis	Diagnosis. Males may be recognised by the following combination of characters (Figs 17 – 22): 1. Massive TA extending distinctly beyond prolateral cymbial margin, 2. Massive RTA, proximally arising from palpal tibia and bulging distinctly retrolaterally and 3. Complete loss of conductor. Females are similar to those of Sinopoda and Borniella gen. nov. in having a fused internal duct system divided into an anterior and a posterior part with glandular appendages situated anteriorly but can be easily distinguished from both latter genera by the anteriorly situated atrium, the absence of epigynal pockets and lobal septum, and the semicircular first windings of the internal duct system (Figs 23 – 37; both other genera without atrium, epigynal pockets and lobal septum present and copulatory openings separated, i. e. not situated together in an atrium, and first winding different).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	description	Description. See description of type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925837FFC3FF56F99E82F64181.taxon	distribution	Distribution. Borneo (Malaysia: West Sarawak) (Fig. 98: blue squares). Species included. Only the type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	description	Figs 17 – 46, 86 – 89, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	materials_examined	Type material. MALAYSIA: Sarawak: Holotype male (SD 1366): West Sarawak, Semengoh arboretum, 4 – 10 January 1984, C. L. & P. R. Deeleman (RMNH). Paratypes (9 females): 2 females with same data as for holotype (RMNH); 4 females (SD 1365 & 1561) with same data as for holotype (RMNH); 1 female with same data as for holotype except: 23 March 1985 (RMNH); 2 females (SD 1368) with same data as for holotype except: 24 & 27 March 1985 (SMF). Additional material examined (1 male, 3 females, 1 subadult male, 17 juveniles, 2 egg-sacs with 15 nymphs). MALAYSIA: Sarawak Province: 7 juveniles with same data as for holotype (RMNH); 4 juveniles, 1 egg-sac with 11 nymphs with same data as for holotype (RMNH); 1 male (SD 1370), 3 females (SD 1372 & 1560), 1 subadult male (SD 1371), 3 juveniles (SD 1373), 1 egg-sac with 4 nymphs, Borneo, West Sarawak, Bako National Park, moeras [= swampy] bog, 29 & 30 March 1985, C. L. & P. R. Deeleman [+ 1 juvenile Sinopoda sp., + 1 juvenile Heteropodinae indet.] (RMNH; 1 male, 1 female: SMF); 3 juveniles (SD 1369) with same data as for holotype except: 24 & 27 March 1985 (SMF).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	etymology	Etymology. The species name is derived from the Malayan word “ kecil ”, meaning “ little ”, and refers to the small body size of this species; term in apposition.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	diagnosis	Diagnosis. Small Sparassidae with TL <5.0 (Figs 86 – 89). Males of Menarik kecil spec. nov. may be recognised by the following combination of characters (Figs 17 – 22): 1. Apical part of embolus forming almost a circle in a plane oblique to cymbial length axis, 2. Tegular apophysis arising from a 7 - to 8 - o’clock-position, curved, its tip distad and 3. RTA with basal part bulgy and with various lobes, apical branch acutely tapering. Females are unique in the subfamily Heteropodinae having an oval to slit-like copulatory atrium at anterior half of epigyne (Figs 23 – 24, 28 – 36). Moreover, posterior part with large spherical spermathecae and long antero-laterad fertilisation ducts (Fig. 25).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	description	Description. Male (holotype): TL 3.4; PL 1.8, PW 1.8, AW 1.1; OL 1.6, OL 1.1. Eye measurements (Fig. 40): AME 0.10; ALE 0.18; PME 0.10; PLE 0.20; AME – AME 0.10; AME – ALE 0.02; PME – PME 0.15; PME – PLE 0.21; AME – PME 0.18; ALE – PLE 0.19; clypeus AME 0.14; clypeus ALE 0.12. Leg formula: 2431. Measurements of palp and legs: palp: 2.80 (0.80, 0.45, 0.55, 1.00); I: 6.60 (1.80, 0.80, 1.80, 1.60, 0.60); II: 8.60 (2.40, 0.90, 2.30, 2.20, 0.80); III: 7.20 (2.10, 0.80, 1.90, 1.80, 0.60); IV: 7.40 (2.10, 0.80, 1.85, 1.95, 0.70). Spination: palp: 131, 101, 2001; legs: Fe I 323, Fe II & III 222, Fe IV 221; Pa I – IV 000; Ti I & II 1026, Ti III & IV 2226; Mt I & II 0006, Mt III 2036, Mt IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 17 intermarginal denticles and 1 escort seta (Fig. 45). Retrolateral claw of right leg I with 16 teeth (Fig. 42). Trilobate membrane of right leg II with narrow median hook, slightly shorter than lateral projections (Fig. 27). Palp as in diagnosis (Figs 17 – 22). Embolus basally wide, apical part filiform. Spermophor curved distinctly at distal part of tegulum, slightly in proximal half (best seen in ventral view). Tegular apophysis wide, curved and tapering. Distal part of tegulum bulging ventrally. Apical part of RTA strongly acute, beak-shaped in retrolateral view with dorsal branch; basal part broad and wide, roughly kidney-shaped in ventral view. Colouration in ethanol (Figs 86 – 87): Prosoma yellowish-brown, median part with brown semi-circular pattern, posterior and lateral margin dark brown. Sternum yellowish-white. Opisthosoma brown with yellow spots, posteriorly dark brown, ventrally yellow, laterally with brown stripes. Chelicerae yellowish-brown. Coxa I – III medio-dorsally with brown stripe. Palps yellowish-brown. Fe and Pa distally darker brown. Legs yellowish-brown; Fe lateral and distal part darker brown; Pa and Ti laterally with brown spots. Female (paratype): TL 4.4; PL 2.0, PW 1.9, AW 1.2; OL 2.4, OW 1.7. Eye measurements (Fig. 41): AME 0.10; ALE 0.18; PME 0.11; PLE 0.25; AME – AME 0.12; AME – ALE 0.04; PME – PME 0.16; PME – PLE 0.22; AME – PME 0.21; ALE – PLE 0.18; clypeus AME 0.19; clypeus ALE 0.17. Leg formula: 2431; measurements of palp and legs: palp: 3.00 (0.80, 0.50, 0.70, 1.00); I: 6.80 (1.80, 0.90, 1.80, 1.70, 0.60); II: 8.50 (2.40, 1.00, 2.30, 2.10, 0.70); III: 7.30 (2.10, 0.90, 1.90, 1.80, 0.60); IV: 7.60 (2.10, 0.80, 1.90, 2.10, 0.70). Spination: palp: 131, 101, 2221, 1004; legs: Fe I 222 / 322, Fe II & III 222, Fe IV 221; Pa I – IV 000; Ti I & II 1016, Ti III 2126, Ti IV 2226; Mt I & II 0006, Mt III 2026, MT IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 16 intermarginal denticles and 1 escort seta (Fig. 46). Palpal claw with 6 long teeth and 1 short tooth (Fig. 43), retrolateral claw of right leg I with 16 teeth (Fig. 44). Copulatory organ as in diagnosis (Figs 23 – 26, 28 – 37). Epigynal field wider than long with two long anterior “ horns ” and two indistinct lateral bulges. Lateral lobes fused, with few “ fusion bubbles ” in posterior half and with slight median indentation at posterior margin, extending very slightly beyond epigastric furrow. Internal duct system wider than long. Glandular appendages short and mainly visible in ventral view. Fertilization ducts arising posterolaterally, long and slender, tips antero-laterad. Colouration in ethanol (Figs 88 – 89): Prosoma yellowish-brown, medially with semi-circular brown pattern, posterior and lateral margin dark brown. Sternum yellowish-white. Opisthosoma brown with yellow spots, medially with yellow transversal stripes, posterior part darker brown, ventral part medially yellow, laterally with brown stripes. Chelicerae yellowish-brown. Coxa I – III medio-dorsally with brown stripe. Palps yellowish-brown; Fe distally brown. Legs yellowish-brown; Fe distally and medially brown; Pa laterally brown. Variation. Male (SD 1370). TL 4.45; PL 2.05, PW 1.95; OL 2.4, OW 1.65. Measurements of leg II: 9.60 (2.70, 1.00, 2.70, 2.40, 0.80); Spination: palp: Mt 1110; legs: Fe II 223, Fe III 222 / 322; Ti II 2026; Mt II 006, Mt III 2026. Chelicerae denticles anterior 3, posterior 5 teeth. In the male from Bako National Park the disto-dorsal apophysis of the male palpal tibia and the retrolateral outline of the basal RTA is differently shaped (Fig. 22). Additionally, the dorsal branch of the RTA was longer and more pronounced (Fig. 21). Females (n = 11). TL 3.7 – 4.75; PL 1.8 – 2.15, PW 1.75 – 2.05, AW 1.1 – 1.15; OL 1.8 – 2.6, OW 1.0 – 1.9. Measurements leg VI: total length 6.90 – 7.90, Fe 1.95 – 2.20, Pa 0.70 – 0.80, Ti 1.80 – 2.00, Mt 1.80 – 2.10, Ta 0.65 – 0.80. Spination: palp: Ti 2121; legs: Fe I 323, Fe II 322, 232, Fe III 232, Fe IV 321, 231; Ti III 1006, 2026, 2226, Ti IV 2126; Mt III 1016, 2006, 2016, 3036. Chelicerae with 3 prolateral, and 5 retrolateral teeth. One female with 6 (left) and 7 (right) retromarginal teeth. Female epigynes with various shapes of atrium (Figs 28 – 36). Spermathecae in some females antero-laterad, ducts leading to spermathecae were narrower (Figs 29, 31, 33), in some female laterad (Figs 28, 30), in some females in between (Figs 32, 34). Notes. The majority of females examined showed scars on the surface of the epigyne. In most females with scars present they are most distinct in the right half (Figs 23, 28 – 29, 32 – 33). Only one female showed a larger scar on the left side (Fig. 30). Egg-sacs contained nymphs with TL 1.2, PL 0.65 – 0.75, the egg-sac with 11 nymphs had a diameter of 3.5 mm and was 2.0 mm thick.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925834FFCEFF56FE7E84E34181.taxon	distribution	Distribution and habitat. Known only from the type locality (Fig. 98: blue squares).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	description	Figs 47 – 65, 90 – 93, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	type_taxon	Type species. Micropoda daviesae spec. nov.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	etymology	Etymology. The generic name is a combination of the prefix “ micro ” (Greek, means “ small / little ”) and the second part of the genus name Heteropoda. It was chosen in reference to the small body size of the only known species of the new genus. The gender is feminine.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	diagnosis	Diagnosis. Small Sparassidae with TL <7.0 (Figs 90 – 93). Micropoda gen. nov. may be recognised by the following combination of characters: Males (Figs 47 – 52): 1. Tegulum with a narrow, but distinct ridge, running a semi-circle from proximal to distal, 2. Alveolus proximally with indistinct pocket in which embolus base is partly hidden, 3. Conductor (homologue) reduced to indistinct bulge at distal tegulum and 4. Embolus with apophysis. Other Heteropodinae with an embolic apophysis, such as most Sinopoda, few Heteropoda or all Yiinthi species (sub “ flagellum ”: Davies 1994) possess a well-developed conductor and lack the alveolus pocket as well as the tegular ridge. Females (Figs 53 – 57): 1. First winding dorsad, both sides fused, running a distinct semi-circle, leaving a window between outer cuticle and internal duct, 2. Lateral lobes ear-shaped and with narrow and long epigynal pockets, situated laterally and 3. Internal duct system with only short glandular appendages anteriorly (but functionally after the first winding) and distinctly convoluted ducts in the posterior part. Few Sinopoda spp. exhibit a similar window between outer cuticle and duct system (e. g., S. derivata Jäger & Ono, 2002 or S. ogatai Jäger & Ono, 2002), but it is formed by the part functionally behind the glandular appendages. Moreover, epigynal pockets in Sinopoda are never exclusively situated laterally, and the internal duct system is much simpler and shows no coils.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	description	Description. See description of type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCEFF56FE7E82F64523.taxon	distribution	Distribution. Papua New Guinea (New Britain) (Fig. 98: orange squares). Species included. Only the type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	description	Figs 47 – 65, 90 – 93, 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	materials_examined	Type material. PAPUA NEW GUINEA: East New Britain: Holotype male (PJ 936): S. E. Bay, Dörpen Peak, South Seas expedition, 14 September 1909, Dr. G. Duncker (ZMH). Paratype: 1 female (PJ 937) with same data as for holotype (ZMH). Additional material examined (4 females and 1 subadult male): PAPUA NEW GUINEA: Morobe: 3 females (PJ 927 – 929, SD 1598) and 1 subadult male (SD 1361), Siassi archipelago, Malabaya, South Seas expedition, 22 October 1909, Dr. G. Duncker (ZMH; 1 female: SMF). West New Britain: 1 female (PJ 926, SD 1597; epigyne not fully sclerotised), S. W. Coast, Lieblich Islands, South Seas expedition, 1909, Dr. G. Duncker (ZMH).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	etymology	Etymology. The species is dedicated to the late Valerie Todd Davies, a pioneering Australian arachnologist; name in genitive case.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	diagnosis	Diagnosis. Micropoda daviesae spec. nov. may be recognised by the following combination of characters: Males (Figs 47 – 52): 1. RTA proximally arising from tibia, with 2 lateral apices, a large medial tooth and a small additional tooth (Fig. 51), 2. Embolus arising from tegulum at 5 - o’clock-position, filiform and apically coiled and 3. Embolic apophysis arising from embolus in 10 - o’clock-position, running parallel to and with similar width of embolus up to their tip. Females (Figs 53 – 57): 1. Lateral lobes extending postero-laterally and running latero-anteriorly in anterior half, 2. Glandular appendages situated at the end of the first winding of the internal duct system and 3. Internal duct system with 1 big loop posteriorly and 4 subsequent turns.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	description	Description. Male (holotype): TL 6.6; PL 3.3, PW 3.1, AW 1.7; OL 3.3, OW 2.1. Eye measurements (Fig. 58): AME 0.19; ALE 0.26; PME 0.18; PLE 0.27; AME – AME 0.18; AME – ALE 0.06; PME – PME 0.18; PME – PLE 0.31; AME – PME 0.18; ALE – PLE 0.21; clypeus AME 0.18; clypeus ALE 0.16. Leg formula: 2431. Measurements of palp and legs: palp: 4.80 (1.60, 0.70, 0.80, 1.70); I: 12.20 (3.30, 1.50, 3.30, 3.00, 1.10); II: 14.30 (3.90, 1.70, 4.00, 3.50, 1.20); III: 12.40 (3.40, 1.40, 3.40, 3.10, 1.10); IV: 12.60 (3.40, 1.20, 3.40, 3.40, 1.20). Spination: palp: 131, 101, 2101; legs: Fe I – III 323, Fe IV 321; Pa I – IV 001; Ti I – III 2026, Ti IV 2126; Mt I 0014, Mt II 1004, Mt III 2016, Mt IV 3036. Chelicerae with 3 promarginal, 4 retromarginal teeth, 15 – 20 denticles near promarginal teeth and 1 escort seta (Fig. 63). Retrolateral claw of right leg I with 12 teeth (Fig. 60). Trilobate membrane of right leg II with triangular median hook, with about same length as lateral projections (Fig. 65). Palp as in diagnosis (Figs 47 – 52). Tibia half as long as cymbium. RTA reaching the margin of the alveolus, with 2 lateral apices, a medial tooth and an additional small tooth, dorso-lateral apex only visible in retrolateral or ventro-distal view (Fig. 51). Apices generally rounded in ventral view, pointed in retrolateral view. Alveolus pocket extending from retrolateral margin to postero-prolateral margin. Tegulum at least as wide as cymbium, retrolaterally extending slightly beyond cymbial margin in ventral view, widest in distal half; spermophor double-curved distinctly in ventral view, with one large and one small U-shaped curve. Embolic apophysis filiform with narrow tip. Embolus tip ventrad, with subapical opening of spermophor. Conductor homologue situated in a 2 - o’clock-position at distal part of tegulum. Colouration in ethanol (Figs 90 – 91): Prosoma yellowish, anterior part with two wide brown stripes running medially to ALE, medially reddish-brown with brown stripes, posterior part medially with semi-circular yellow pattern, lateral and latero-posterior margin brown. Fovea with longitudinal reddish-brown stripe. Sternum pale yellow. Opisthosoma whitish-grey with brown spots, posteriorly darker, laterally with brown stripes, ventral part without pattern. Chelicerae reddish-brown with two dark longitudinal brown stripes, retrolaterally with brown spot. Palps yellowish-brown. Legs yellowish-brown; Fe distally, medially and proximally with brown band; Ti proximally with brown band. Female (paratype): TL 6.5; PL 3.1, PW 2.9, AW 1.8; OL 3.4, OW 2.3. Eye measurements (Fig. 59): AME 0.18; ALE 0.25; PME 0.17; PLE 0.24; AME – AME 0.18; AME – ALE 0.07; PME – PME 0.20; PME – PLE 0.36; AME – PME 0.18; ALE – PLE 0.26; clypeus AME 0.17; clypeus ALE 0.18. Leg formula: 2431. Measurements of palp and legs: palp: 4.40 (1.30, 0.70, 1.00, 1.40); I: 10.30 (2.90, 1.40, 2.70, 2.40, 0.90); II: 12.10 (3.40, 1.60, 3.30, 2.80, 1.00); III: 10.70 (3.00, 1.40, 2.90, 2.50, 0.90); IV: 10.90 (3.00, 1.30, 2.80, 2.80, 1.00). Spination: palp: 131, 101, 2121, 1014; legs: Fe I – III 323, Fe IV 321; Pa I – IV 001; Ti I – III 2026, Ti IV 2126; Mt I & II 0004, Mt III 2006, Mt IV 3036. Chelicerae with 3 promarginal, 4 retromarginal teeth, 15 – 20 denticles near promarginal teeth and 1 escort seta (Fig. 64). Copulatory organ as in diagnosis (Figs 53 – 57). Epigynal field wider than long with fusion bubbles situated antero-medially. Lateral lobes kidney-shaped and distinct. Copulatory orifices situated medio-anteriorly. Internal duct system longer than wide. Glandular appendages short and straight, extending only in anterior half of internal duct system. Fertilization ducts situated medio-posteriorly, short. Colouration in ethanol (Figs 92 – 93): Prosoma yellowish, anterior part with two wide, brown stripes running medially to ALE, medially brown with brown stripes, posterior part medially with semi-circular yellow pattern, lateral and latero-posterior margin brown. Fovea with longitudinal reddish-brown stripe. Sternum pale yellow. Opisthosoma yellowish-grey with brown spots, medially with four brown dots, in front of spinnerets white spot, ventral part yellowish-grey, medially grey, laterally brown. Chelicerae reddish-brown with two dark longitudinal brown stripes, retrolaterally with brown spot. Palps yellowish-brown. Legs yellowish-brown; Fe medially and proximally with brown band; Ti proximally with brown band. Variation. Females (n = 4). TL 5.5 – 6.7; PL 2.6 – 3.3, PW 2.4 – 3.2, AW 1.4 – 1.9; OL 2.9 – 3.4, OW 2.0 – 2.3. Measurements leg I: total length 8.20 – 10.80, Fe 2.20 – 3.00, Pa 1.10 – 1.50, Ti 2.20 – 2.90, Mt 1.90 – 2.50, Ta 0.80 – 1.00. Spination: legs: Ti I – III 1016.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD87925839FFCAFF56FAD9809843E4.taxon	distribution	Distribution. Known only from New Britain and Siassi Islands in Papua New Guinea (Fig. 98: orange squares).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	description	Figs 66 – 81, 94 – 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	type_taxon	Type species. Tiomaniella ladam spec. nov.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	etymology	Etymology. The generic name is derived from the name of Tioman Island. The gender is feminine.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	diagnosis	Diagnosis. Small Sparassidae with TL <7 (Figs 94 – 97) belonging to the subfamily Heteropodinae. Males of Tiomaniella gen. nov. can be distinguished from all other Heteropodinae by the subtegular functional conductor (Figs 66 – 67). Females may be recognised by the following combination of characters: 1. Copulatory organs situated laterally with epigynal pockets located posteriorly, with first winding of internal duct system postero-mediad, 2. Median septum not recognisable, “ fusion bubbles ” indicating fused seams between lateral lobes and median septum and 3. Internal duct system simple with very broad ducts leading to large spermathecae (Figs 70 – 72).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	description	Description. See description of type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFCAFF56FB9A82F64643.taxon	distribution	Distribution. Malaysia: Tioman Island (Fig. 98: red triangle). Species included. Only the type species.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	description	Figs 66 – 81, 94 – 98	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	materials_examined	Type material. MALAYSIA: Pahang: Holotype male (SD 1471): Pulau Tioman, pied du Gunung, rain forest, N 02 ° 47.181 ’, E 104 ° 07.892 ’, 160 m, 2 October 2001, L. Monod, field number TM- 17 (MHNG). Paratypes (1 male, 3 females): 1 female (SD 1472), with same data as for holotype (MHNG). MALAYSIA: Pahang: 1 male (SD 1474), Tioman Island, Westside, 2 km SE of Kg. Genting, N 02 ° 47 ’, E 104 ° 08 ’, 100 m, 28 June 2001, A. Schulz & K. Vock leg., field number M 01 - 71 (SMF); 2 females with same data as for previous specimen, except: 1 female 26 June 2001 with field number M 01 – 39 (MHNG); 1 female (SD 1473) 26 June 2001 with field number M 01 – 40 (SMF).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	etymology	Etymology. The species name refers to the Malayan word “ ladam ” meaning “ horse shoe ” and refers to the brown U-shaped patterning of the prosoma; term in apposition.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	diagnosis	Diagnosis. Tiomaniella ladam spec. nov. may be recognised by the following combination of characters: Males (Figs 66 – 69): 1. Distinct, apically fanned conductor homologue arising in 1 - o’clock-position from tegulum, 2. TA arising centrally, with broad base and retrolatero-distad acute tip, 3. Dorsal branch of RTA as long as ventral branch (best seen in retrolateral view). Females (Figs 70 – 72): 1. Epigyne with epigynal pockets running from laterally to postero-medially, 2. Lateral lobes extending not in anterior half of epigyne and 3. Internal duct system only fused anteriorly at ventral part.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	description	Description. Male (holotype): TL 5.3; PL 2.7, PW 2.5, AW 1.3; OL 2.6, OW 1.6. Eye measurements (Fig. 73): AME 0.13; ALE 0.20; PME 0.11; PLE 0.25; AME – AME 0.13; AME – ALE 0.05; PME – PME 0.21; PME – PLE 0.26; AME – PME 0.20; ALE – PLE 0.21; clypeus AME 0.16; clypeus ALE 0.14. Leg formula: 2431. Measurements of palp and legs: palp: 4.00 (1.10, 0.70, 0.80, 1.40); I: 8.65 (2.40, 1.10, 2.20, 2.10, 0.85); II: 10.70 (3.00, 1.30, 2.90, 2.60, 0.90); III: 8.95 (2.60, 1.15, 2.30, 2.10, 0.80); IV: 9.20 (2.60, 1.00, 2.40, 2.40, 0.80). Spination: palp: 131, 101, 2111; legs: Fe I 323, Fe II & III 222, Fe IV 221; Pa I – IV 000; Ti I & II 1026, Ti III & IV 2126; Mt I & II 1026, Mt III & Mt IV 3036. Cheliceral furrow with 3 promarginal, 5 retromarginal teeth, 16 intermarginal denticles and 1 escort seta (Fig. 78). Retrolateral claw of right leg I with 11 teeth (Fig. 75). Trilobate membrane of left leg I with short and narrow median hook, shorter than lateral projections (Fig. 80). Palp as in diagnosis (Figs 66 – 69). RTA arising subdistally from tibia, with two branches, both latter connected with a concave margin. Cymbium roughly twice as long as tibia, with enormous retrolateral bulge and small retroproximal projection. Subtegulum sail-shaped, covers half of the embolus length in ventral view, tip filiform elongated parallel to embolus tip. Median part of tegulum bulging prolaterally. Spermophor distinctly S-shaped. Embolus basally wide, then filiform, running a semicircle, its tip close to conductor homologue. Colouration in ethanol (Figs 94 – 95): Prosoma yellowish-brown, latero-posteriorly with brown U-shaped pattern, medially with brown longitudinal stripe, lateral margin with three dark brown dots. Sternum yellow. Opisthosoma grey with brown spots, medially with grey longitudinal stripe, posteriorly with grey dot, ventral part grey, laterally with brown stripes. Chelicerae yellowish-brown. Coxa dorsally with brown spots. Palps yellowish-brown. Legs yellowish-brown; Ti distally reddish-brow; Mt & Ta reddish-brown. Female (paratype): TL 6.0; PL 2.6, PW 2.5, AW 1.5; OL 3.4, OW 2.2. Eye measurements (Fig. 74): AME 0.16; ALE 0.23; PME 0.14; PLE 0.27; AME – AME 0.12; AME – ALE 0.04; PME – PME 0.19; PME – PLE 0.32; AME – PME 0.21; ALE – PLE 0.23; clypeus AME 0.15; clypeus ALE 0.13. Leg formula: 2431; measurements of palp and legs: palp: 4.10 (1.10, 0.70, 1.00, 1.30); I: 8.55 (2.30, 1.20, 2.25, 2.00, 0.80); II: 10.20 (2.80, 1.30, 2.70, 2.50, 0.90); III: 9.20 (2.60, 1.20, 2.40, 2.20, 0.80); IV: 9.30 (2.50, 1.10, 2.30, 2.50, 0.90). Spination: palp: 131, 101, 2121, 1004; legs: Fe I 323, Fe II & III 222, Fe IV 221; Pa I – V 000; Ti I 1004, Ti II 0004, Ti III 2026, Ti IV 2125; Mt I & II 0004, Mt III 1006, MT IV 3036. Chelicerae with 3 promarginal, 4 retromarginal teeth, 22 intermarginal denticles and 1 escort seta (Fig. 79). Palpal claw with 6 long and 1 – 2 short teeth (Fig. 76). Retrolateral claw of left leg I with 13 teeth (Fig. 77). Trilobate membrane of left leg I with short and narrow median hook, slightly shorter than lateral projections (Fig. 81). Copulatory organ as in diagnosis (Figs 70 – 72). Epigynal field slightly wider than long, with distinct anterior bands. Many “ fusion bubbles ” situated almost over entire epigyne. Lateral lobes posteriorly with median indentation. Internal duct system nearly as long as wide. Dorsal part of ducts S-shaped. Fertilization ducts arising posteromedially, long and curved. Colouration in ethanol (Figs 96 – 97): Prosoma yellowish-brown, latero-posteriorly with brown U-shaped pattern, medially with brown longitudinal stripe, lateral margin with three dark brown spots. Sternum yellow, margin light brown. Opisthosoma brown with yellowish-white spots, medially with white longitudinal stripe, ventral part yellowish-brown, laterally with brown spots. Chelicerae yellowish-brown, right chelicera with three brown dots. Coxa dorsally with brown spots. Palps yellowish-brown; Fe prolaterally, retrolaterally and distally with brown dots; Mt distal part reddish-brown. Legs yellowish-brown; Fe medially and distally with brown band; Pa prolaterally and retrolaterally brown; Ti I – III distally and proximally with brown band, Ti IV only distal part with brown band; Mt and Ta reddish-brown. Variation. Male (n = 1): TL 4.85; PL 2.45, PW 2.4; OL 2.4, OW 1.4. Measurements leg I: total length 8.20, Fe 2.20, Pa 1.10, Ti 2.40, Mt 2.00, Ta 0.80. Spination: palp: Ti 1121; legs: Fe I 322, Ti I 2026, Ti II 2026; Mt I 2026, Mt II 2026. Females (n = 2): TL 6.0 – 6.8; PL 2.6 – 3.1, PW 2.5 – 2.9, AW 1.5 – 1.7; OL 3.3 – 3.7, OW 2.2 – 2.5. Measurements leg I: total length 8.55 – 9.55, Fe 2.30 – 2.60, Pa 1.20 – 1.30, Ti 2.25 – 2.50, Mt 2.00 – 2.30, Ta 0.75 – 0.85. Spination: palp: Mt 1014; legs: Fe II 223; Ti I 1006, Ti II 006, Ti IV 2135; Mt III 2006, 2016.	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
03FD8792583DFFD1FF56F938838F46B9.taxon	distribution	Distribution. Known only from the type locality (Fig. 98: red triangle).	en	Grall, Elena, Jäger, Peter (2022): Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae). Zootaxa 5169 (1): 1-25, DOI: https://doi.org/10.11646/zootaxa.5169.1.1
