identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C187DA1A40FFACC7CDFC2C0B9D5D95.text	03C187DA1A40FFACC7CDFC2C0B9D5D95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marchantiopsida Doweld	<div><p>Key to the families of sterile complex thalloid liverworts of Sri Lanka</p> <p>1. Thallus well differentiated into layers with air chambers..................................................................................................................2</p> <p>- Thallus weakly differentiated into layers without distinct air chambers.................................................................. Dumortieraceae</p> <p>2. Thallus with compound air pores in the epidermis, gemmae present cupules cup-shaped..................................... Marchantiaceae</p> <p>- Thallus with simple air pores in the epidermis, gemmae absent or present, cupules not cup-shaped...............................................3</p> <p>3. Gemmae present in crescent-shaped cupulues............................................................................................................ Lunulariaceae</p> <p>- Gemmae absent..................................................................................................................................................................................4</p> <p>4. Thallus very thin, 2-layered, translucent..................................................................................................................... Cyathodiaceae</p> <p>- Thallus thick, several layered, not translucent...................................................................................................................................5</p> <p>5. Air chambers with chlorophyllose filaments......................................................................................................................................6</p> <p>- Air chambers without chlorophyllose filaments.................................................................................................................................7</p> <p>6. Thallus green with purplish pigmentation................................................................................................................... Targioniaceae</p> <p>- Thallus green, without purplish pigmentation.......................................................................................................... Exormotheceae</p> <p>7. Ventral scales in 2 or several rows, appendages absent..................................................................................................... Ricciaceae</p> <p>- Ventral scales in 2 rows, with 1–4 appendages............................................................................................................... Aytoniaceae</p> <p>Key to families of fertile complex thalloid liverworts of Sri Lanka</p> <p>1. Perigonial chambers on stalked receptacles.......................................................................................................................................2</p> <p>- Perigonial chambers aggregated in cushions or embedded in groups on dorsal surface of thallus or on short branches..................3</p> <p>2. Perigonial receptacles bearing compound air pores, stalk with 2 to 4 rhizoid furrows; involucre bivalved or cup-shaped; pseudoperianths present............................................................................................................................................ Marchantiaceae</p> <p>- Perigonial receptacles bearing few open-air chambers, stalk with 2 rhizoid furrows; involucres tubular, bristly, opening by a slit; pseudoperianths absent............................................................................................................................................. Dumortieraceae</p> <p>3. Sporophyte on stalked receptacles.....................................................................................................................................................4</p> <p>- Sporophytes lacking receptacles.........................................................................................................................................................5</p> <p>4. Perigonial chambers aggregated in terminal cushions on thallus; seta massive, elongating prior to spore dispersal............................................................................................................................................................................................................... Lunulariaceae</p> <p>- Perigonial chambers embedded in dorsal groove of thallus or in irregular rows on thallus; seta remaining short............................6</p> <p>5. Sporophytes embedded singly in the thallus; involucres absent; seta absent; elaters absent; capsules cleistocarpous..... Ricciaceae</p> <p>- Sporophytes ventral at the thallus apex; involucres bivalved; seta remaining short; elaters few per capsule; capsule dehiscence by a lid and irregular valves....................................................................................................................................................................7</p> <p>6. Receptacles bearing compound air pores; involucres bivalved, cup-shaped or flap-like; capsule dehiscence by a lid................................................................................................................................................................................................................ Aytoniaceae</p> <p>- Receptacles bearing simple air pores, stalk with 1 rhizoid furrow; involucres tubular; capsule dehiscence by irregular valves or irregular lid and valves.......................................................................................................................................... Exormothecaceae</p> <p>7. Thallus thicker (more than 2 layers), not translucent; antheridia borne on special branches; sporophyte in a dark, swollen, mussellike bivalved involucre below the thallus apex foot and seta of sporophyte well developed...................................... Targioniaceae</p> <p>- Thallus very thin, 2-layered, translucent; antheridia scattered in clusters along the edge of the thallus; sporophyte in an apical notch on the ventral thallus surface; foot and seta of sporophyte represented by a single filament..................................... Cyathodiaceae</p> <p>Key to the genera of Complex Thalloid liverworts of Sri Lanka</p> <p>1. Thallus with cupules.......................................................................................................................................................................... 2</p> <p>- Thallus without cupules......................................................................................................................................................................3</p> <p>2. Gemma-receptacles cup-shaped; dorsal epidermis with compound pores....................................................................... Marchantia</p> <p>- Gemma-receptacles crescent-shaped; dorsal epidermis with simple pores......................................................................... Lunularia</p> <p>3. Photosynthetic tissue with narrow, vertical, channel-like air chambers or air chambers vestigial....................................................4</p> <p>- Photosynthetic tissue with air-chambers in one or more layers.........................................................................................................5</p> <p>4. Air chambers absent, thallus weakly differentiated into layers........................................................................................ Dumortiera</p> <p>- Air chambers in more than one layer or channel-like, thallus well differentiated into layers................................................... Riccia</p> <p>5. Thallus very thin, 2-layered, translucent......................................................................................................................... Cyathodium</p> <p>- Thallus thick, several layered.............................................................................................................................................................6</p> <p>6. Ventral scales hyaline, without appendages.................................................................................................................. Exormotheca</p> <p>- Ventral scales purplish, with appendages...........................................................................................................................................7</p> <p>7. Ventral scales with 1 appendage; sporophytes ventral at the thallus apex........................................................................... Targionia</p> <p>- Ventral scales with 1 to 4 appendages; sporophytes on stalked receptacles.......................................................................................8</p> <p>8. Ventral scales with ovate to lanceolate appendages; appendages purplish pink to hyaline, not constricted at base.................................................................................................................................................................................................................. Plagiochasma</p> <p>- Ventral scales with 2 or 3 filiform appendages..................................................................................................................... Reboulia</p></div> 	http://treatment.plazi.org/id/03C187DA1A40FFACC7CDFC2C0B9D5D95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruklani, Sumudu;Rubasinghe, Sumudu C. K.	Ruklani, Sumudu, Rubasinghe, Sumudu C. K. (2022): Thalloid Liverworts (Marchantiopsida) of Sri Lanka. Phytotaxa 551 (1): 1-82, DOI: 10.11646/phytotaxa.551.1.1
03C187DA1A68FF87C7CDFF080B9D5A1F.text	03C187DA1A68FF87C7CDFF080B9D5A1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marchantia L.	<div><p>Key to Sri Lankan Marchantia species</p> <p>1. Thallus margin lobulate or crenulate. Ventral scales in 4–6 rows extending over 75–100% of the thallus width. Laminal scales broader than long without apical papillae................................................................................................................. M. polymorpha</p> <p>- Thallus margin entire. Ventral scales in 4-rows extending over 16–66% of the thallus width. Laminal scales as long as or longer than wide, with apical papillae...........................................................................................................................................................2</p> <p>2. Inner opening of epidermal pore cruciate. Appendage of median scale entire or nearly so. Gemma cups with ciliate lobes and numerous papillae on outer surface. Male receptacles peltate......................................................................................... M. paleacea</p> <p>- Inner opening of epidermal pore with straight or convex wall, not cruciate. Appendage of median scales bluntly or sharply toothed. Gemma cups ciliate or nearly entire without papillae on outer surface. Male receptacle palmate...................................................3.</p> <p>3. Appendage of median scale yellowish. Male receptacles sessile. Female receptacles with involucres located underneath lobes.................................................................................................................................................................................................. M. acaulis</p> <p>- Appendage of median scale purplish or hyaline not yellowish. Male receptacles stalked. Female receptacles with involucres alternating with lobes.........................................................................................................................................................................4</p> <p>4. Appendage of median scale bluntly toothed. Female receptacle without shallow median projections, shallowly divided into 4–6 lobes, lobes not convex or broadened apically...................................................................................... M. pappeana subsp. robusta</p> <p>- Appendage of median scale sharply toothed. Female receptacle with a conspicuous median projection, deeply divided into 5–13 lobes. Lobes convex and broadened apically.....................................................................................................................................5</p> <p>5. Thallus with a dark median band on the dorsal surface. Mucilage cavities sometimes present in the ventral tissue. Appendage of median scales with a single apical cell or a row of 2–3 apical cells, marginal teeth oriented towards the apex of the appendage............................................................................................................................................................... M. papillata subsp. grossibarba</p> <p>- Thallus without a dark median band on the dorsal surface. Ventral tissue without mucilage cavities or rarely 1–2. Appendage of median scales with a row of 2–3 cells apically, marginal teeth, recurved towards the base of the appendage........... M. emarginata</p></div> 	http://treatment.plazi.org/id/03C187DA1A68FF87C7CDFF080B9D5A1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruklani, Sumudu;Rubasinghe, Sumudu C. K.	Ruklani, Sumudu, Rubasinghe, Sumudu C. K. (2022): Thalloid Liverworts (Marchantiopsida) of Sri Lanka. Phytotaxa 551 (1): 1-82, DOI: 10.11646/phytotaxa.551.1.1
03C187DA1A7BFF95C7CDF9D60B9C5BD1.text	03C187DA1A7BFF95C7CDF9D60B9C5BD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jungermanniopsida Stotler & Crand.	<div><p>Key to families of simple thalloid liverworts in Sri Lanka</p> <p>1. Plants with stem and leaves................................................................................................................................................................2</p> <p>- Plants thalloid.....................................................................................................................................................................................3</p> <p>2. Plants robust, reddish or purplish. Leaves deeply clasping the stem, undivided......................................................... Pleuroziaceae</p> <p>- Plants pale green, green or yellowish green, usually forming rosettes. Leaves succubously inserted to the stem............................................................................................................................................................................................................ Fossombroniaceae</p> <p>3. Thallus simple or pinnate, without midrib or with midrib only on branches. Gametangia on the thallus margin or on short lateral branches........................................................................................................................................................................... Aneuraceae</p> <p>- Thallus simple or dichotomous, with a midrib throughout. Gametangia on the midrib on dorsal or ventral side............................4</p> <p>4. Thallus usually 2–3 mm wide, margins with unicellular hairs or rhizoids. Midrib without central strand in cross section. Gametangia on the ventral side of the midrib.................................................................................................................................. Metzgeriaceae</p> <p>- Thallus usually more than 3 mm wide, margins without hairs. Midrib with central strand. Gametangia on the dorsal side of the midrib....................................................................................................................................................................... Pallaviciniaceae</p> <p>Key to the genera of simple thalloid liverworts of Sri Lanka</p> <p>1. Plants thalloid.....................................................................................................................................................................................2</p> <p>- Plants leafy.........................................................................................................................................................................................6</p> <p>2. Thallus with a mid-rib, archegonia and antheridia on dorsal or ventral side of the midrib...............................................................3</p> <p>- Thallus mid-rib lacking or indistinct, archegonia and antheridia at thallus margin..........................................................................5</p> <p>3. Plants dendroid, thallus margin sharply toothed, thallus growing erect from a creeping rhizome....................................... Jensenia</p> <p>- Plants not dendroid; thallus margin entire or with hairs, not sharply toothed, thallus creeping/erect, rhizome lacking....................4</p> <p>4. Thallus more than 3 mm wide, with undulate wings, without hairs; antheridia in rows on midrib; archegonia aligned on midrib.............................................................................................................................................................................................. Pallavicinia</p> <p>- Thallus less than 3 mm wide, without undulate wings, with scarce hairs at margins or on ventral side; antheridia on ventral side of midrib; archegonia on ventral side of thallus..................................................................................................................... Metzgeria</p> <p>5. Thallus more than 3 mm wide, fleshy, irregularly branched, shortly stalked, multicellular; more than 6 oil-bodies in each epidermal cell; gemmae exogenous, shortly stalked, multicellular and bulbous..................................................................................... Aneura</p> <p>- Thallus less than 3 mm wide, pinnately branched or palmate; 0–2 oil-bodies in each epidermal cell; gemmae endogenous, formed within the walls of existing cells, 1–2 celled, at the thallus apex......................................................................................... Riccardia</p> <p>6. Plants green, to light green with two rows of lateral thallus lobes which are leaf-like; slime papillae present as ventral appendages or present on leaves; antheridia scattered along stem; mostly bright orange to yellow in colour; archegonia naked and dispersed dorsally on the stem; capsules spheroidal.................................................................................................................... Fossombronia</p> <p>- Plants reddish rarely green with succubous, complicatedly bilobed leaves; slime papillae lacking; antheridia solitary in the axils of reduced perigonial leaves; archegonia enclosed by a perianth; capsules ovoid to subspheroidal........................................ Pleurozia</p></div> 	http://treatment.plazi.org/id/03C187DA1A7BFF95C7CDF9D60B9C5BD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruklani, Sumudu;Rubasinghe, Sumudu C. K.	Ruklani, Sumudu, Rubasinghe, Sumudu C. K. (2022): Thalloid Liverworts (Marchantiopsida) of Sri Lanka. Phytotaxa 551 (1): 1-82, DOI: 10.11646/phytotaxa.551.1.1
03C187DA1A0AFFE5C7CDFBA50A5A5DF1.text	03C187DA1A0AFFE5C7CDFBA50A5A5DF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blasia pusilla L.	<div><p>Blasia pusilla L.</p> <p>Hattori (1968) in his “Hepaticae collected by F.Schmid in Ceylon and Pakistan ” included Blasia pusilla L. However, this specimen had been collected by F. Schmid in 1953 from Pakistan not from Sri Lanka. and was not recorded during the present study,confirming that Blasia pusilla L. does not occur in Sri Lanka. The species had been erroneously included in the latest checklist, by Long and Rubasinghe (2014) and should be corrected.</p> </div>	http://treatment.plazi.org/id/03C187DA1A0AFFE5C7CDFBA50A5A5DF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruklani, Sumudu;Rubasinghe, Sumudu C. K.	Ruklani, Sumudu, Rubasinghe, Sumudu C. K. (2022): Thalloid Liverworts (Marchantiopsida) of Sri Lanka. Phytotaxa 551 (1): 1-82, DOI: 10.11646/phytotaxa.551.1.1
03C187DA1A0AFFE5C7CDFA9A0A9E5CBC.text	03C187DA1A0AFFE5C7CDFA9A0A9E5CBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metzgeria latifrons Steph.	<div><p>Metzgeria latifrons Steph.</p> <p>According to So (2003), the collection of M. latifrons from Sri Lanka cited by Stephani (1917) (collected at Poondeloya, 4-8000 ft, J. Nietner 155 as M. latifrons, 1868, det. Müller, EGR, G, M, n.v.) belongs to M. crassipilis.Also, M. latifrons is not accepted in the world checklist of hornworts and liverworts by Söderström et al. (2016).</p> </div>	http://treatment.plazi.org/id/03C187DA1A0AFFE5C7CDFA9A0A9E5CBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Ruklani, Sumudu;Rubasinghe, Sumudu C. K.	Ruklani, Sumudu, Rubasinghe, Sumudu C. K. (2022): Thalloid Liverworts (Marchantiopsida) of Sri Lanka. Phytotaxa 551 (1): 1-82, DOI: 10.11646/phytotaxa.551.1.1
