taxonID	type	description	language	source
D82B87B5FFD4B15CFF33FEC2FAF5FADD.taxon	description	(Figs 1 – 3, 22, 32)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD4B15CFF33FEC2FAF5FADD.taxon	materials_examined	Material examined. Mozambique. 1 male, Maputo Special Reserve, West Gate, Sand Forest, 22 m, 26 ° 30 ’ 14.2 ” S, 32 ° 42 ’ 59.6 ” E, 30. v. – 9. vi. 2017, Actinic Light Trap, Aristophanous, M., László, G., Miles, W., Vetina, A. leg., ANHRT: 2017.26, gen. slide No.: LGNA 374, BOLD process / DNA barcode id.: ANLMN 2794 - 21 / ANHRTUK 00013214; 2 males, 1 female, same site, 21 – 22. ii. 2018, Actinic Light Trap, László, G., Mulvaney, J., Smith, L. leg; 1 female, same site, method and collectors, 13 – 15. ii. 2018; 3 males, 1 female, Maputo Special Reserve, Futi Corridor (Sand Forest Woodland Mosaic), 17 m, 26 ° 32 ’ 10.1 ” S, 32 ° 43 ’ 09.7 ” E, 23 – 24. ii. 2018, Actinic Light Trap, László, G., Mulvaney, J., Smith, L. leg., gen. slide Nos: LGNA 440 (male), LGNA 441 (female), BOLD process / DNA barcode ids: ANLMN 2795 / ANHRTUK 00019572, ANLMN 2796 - 21 / ANHRTUK 00019566 (ANHRT). Zambia. 1 male, Kalungu, North of Isoka, 1280 m, S 9 ° 40 ’ 52 ”, E 32 ° 42 ’ 50 ”, 30. iv. – 1. v. 2019, Actinic Light Trap, Dérozier, V., László, G., Miles, W. leg., ANHRT: 2019.12, gen. slide No.: LGNA 1062, BOLD process / DNA barcode id.: ANLMN 2797 - 21 / ANHRTUK 00136186; 1 female, Kalambo Falls, 30 km N of Mbala, Northern Province, 1189 m, 08 ° 35 ’ 50 ” S, 31 ° 14 ’ 26 ” E, 9 – 11. v. 2019, Actinic Light Trap, Dérozier, V., László, G., Miles, W. leg., ANHRT: 2019.12, gen. slide No.: LGNA 1063, BOLD process / DNA barcode id.: ANLMN 2798 - 21 / ANHRTUK 00083347 (ANHRT). Genetic information. Hampsonola elachistomorpha has been assigned the BIN URI: BOLD: AEJ 8170. The intraspecific divergence ranges between 0.00 – 0.31 %. According to BOLD, the nearest neighbour of the species is a probably undescribed species collected in coastal Kenya misidentified as Nola minima (BIN URI: BOLD: ABW 4977). The genetic pairwise distances between H. elachistomorpha and its congeners H. argyropasta and H. nanographa are rather large, in the range of 7.4 – 8.07 % and 8.08 – 8.41 %, respectively, which may refer to a genuslevel separation between H. elachistomorpha and the H. argyropasta-nanographa species-pair. Nevertheless, as no autapomorphies in the genital morphology could be observed, the three species are treated here as congeneric.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD4B15DFF33F920FD90FBB1.taxon	description	(Figs 4 – 5, 23, 33)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD4B15DFF33F920FD90FBB1.taxon	materials_examined	Material examined. Kenya. 1 male, Central Province, Thika District, 8 km SW Thika, Karamaini Estate, 1550 m, S: 01 ° 02 ’ 746 ”, E: 36 ° 59 ’ 472 ”, 19. x. 2010, A. J. Kingston, gen. slide No.: LGNA 1298; 1 female, same site, 19. vii. 2011, gen. slide No.: LGNA 1299 (coll. A. J. Kingston, Albrighton, UK). Taxonomic note. The remarkably distinctive species was described from a single female holotype in the genus Nola. Subsequently, Hacker (2014) figured two male specimens from Yemen without illustrating the male genitalia. The examination of the genital apparatuses of a male and a female specimen recently collected in Kenya sharing an identical habitus with those illustrated in Hacker et al. (2012) and Hacker (2014) has revealed close affinities with the genus Hampsonola, although displaying unique characters such as the two apical processes of the valva and the large lobe-like harpe. To clarify whether or not these characters represent generic autapomorphies, genetic studies involving a wide range of Oriental and Afrotropical Hampsonola taxa would be required, which was beyond the scope of this paper. The species is, therefore, tentatively treated here as a member of the genus Hampsonola: Hampsonola aulombardiella (Hacker, 2012) comb. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD4B15DFF33F920FD90FBB1.taxon	description	Description of the male adult and genitalia. Adult (Figs 4 – 5). Forewing length 6 – 7.3 mm. Head small, antenna bipectinate with very short, finely ciliate rami in male, filiform in female; labial palp short and relatively narrow, porrect, dark brown; frons and vertex brownish-grey. Thorax dark brown in cranial third, creamy-white in caudal two-thirds, collar dark brown, tegula dark brown cranially, creamy-white caudally. Abdomen creamy-white. Wing narrow, distal third of costal margin evenly arcuate, apex rounded. Costal third dark brown, subapically with pale, diffuse, creamy patches, ventro-basal two-thirds creamy-white, tornus and subterminal area light rusty brown; transverse lines absent. Cilia long, dark greyish-brown. Hindwing off-white, costal and apical area somewhat greyish; cilia long, pale grey. Underside of wings dark greyish-brown, ventro-basal area pale grey. Male genitalia (Fig. 23). Uncus very short, broad, evenly tapered with rounded margins, apex truncate. Tegumen moderately long and broad, margins gently arched. Subscaphium short, narrow, weakly sclerotized. Valva moderately broad, more or less quadrangular with straight dorsal and slightly undulate ventral margin, distally with long, narrow, membranous, slightly setose dorsal and ventral process. Dorsal membranous part of valva with a long, arched crest serving as a base of short androconial scales. Sacculus elongate-quadrangular. Harpe situated distally, proximal section narrow, ridge-like, connected to base of costa, distal section dilated, triangular, with finely dentate ventral margin, curved ventrad, apically pointed. Transtilla very broad, ribbon-like, arms medially fused and dilated. Juxta large, gently curved V-shaped; vinculum short, apically rounded V-shaped. Aedeagus short, relatively thick medially, gradually tapered distally, coecum very narrow, apically slightly dilated and truncate; vesica with a narrow, elongate scobinated area.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD5B15AFF33FB4FFE3DFB95.taxon	description	(Figs 6 – 8, 24, 34 – 35)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD5B15AFF33FB4FFE3DFB95.taxon	materials_examined	Type material examined. Holotype, female. Red circular label “ Type ” / “ Gold Coast. Bibianaha. 23. X – 2. XI. 1911. H. G. F. Spurrell. 1911 - 431. ” / with handwritten “ Celama argyropasta type ♀. Hmpsn. ” / QR code label with unique number “ NHMUK 010598788 ” / genitalia slide No. “ SLIDE NHMUK 010317591 ” (prepared by László) (NHMUK). Additional material examined. Liberia. 1 male, Liberia, 140 m, Krahn-Bassa Reserve, Sinoe County, Juboe River, 7.5 km South West Pellokon Town, 5 ° 39 ’ 4 ” N, 8 ° 39 ’ 4 ” W, 14 – 21. i. 2018, Cold Cathode UV Bucket Light Trap, Geiser, M., Sáfián, Sz., Simonics, G. leg. ANHRT: 2017.33, gen. slide No.: LGNA 1229, BOLD process / DNA barcode id.: ANLMN 2800 - 21 / ANHRTUK 00101064; 1 female, 611 m, Lofa County, Wologizi Mts, base camp forest, 8 ° 07 ’ 17 ” N, 9 ° 57 ’ 42 ” W, 20. xi. – 01. xii. 2017, MV Light Trap (125 W) Aristophanous, M., Sáfián, Sz., Simonics, G. & Smith, L. leg. ANHRT: 2017.33. gen. slide No.: LGNA 599, BOLD process / DNA barcode id.: ANLMN 2801 - 21 / ANHRTUK 00026556; 1 male, 15 m, Grand Cape Mount County, Lake Piso area (Bomi forest — savannah mosaic), 6 ° 39 ’ 19.19 ” N, 11 ° 7 ’ 71.35 (sic!) ” W, 2 – 9. I. 2018, MV Light Trap, Sáfián, Sz. Simonics, G. leg., ANHRT: 2017.33 (ANHRT). Taxonomic note. Hacker et al. (2012) treated the species as a Meganola without studying the genitalia of the female holotype. As the facies of the species is highly reminiscent of Meganola nanographa Hacker, 2012, the examination of the holotype genitalia of Celama argyropasta was crucial in order to elucidate its generic position and test the validity of Hacker’s taxon. The dissections of the C. argyropasta holotype collected in Ghana, and a series of DNA barcoded West African and Gabonese specimens of both sexes have revealed that the West African argyropasta and M. nanographa described from Uganda belong to the genus Hampsonola rather than to Meganola. In the male genitalia, the two species are distinguished by the less dilated dorsal arm of the valva and the considerably more curved aedeagus of H. nanographa. The females of the two taxa differ in the somewhat longer apophyses and considerably larger signa bursae of argyropasta compared to the Gabonese specimens. Analyses of DNA barcodes have confirmed the conspecifity of the Gabonese specimens and the holotype of M. nanographa (BOLD process / DNA barcode id.: GWOSG 784 - 10 / BC ZSM Lep 41781), and furthermore, a pairwise distance of 3.31 % was calculated between the West African H. argyropasta and Central African H. nanographa, corroborating the observed morphological divergence. Consequently, H. argyropasta and H. nanographa are confirmed here as distinct species and transferred to the genus Hampsonola: Hampsonola argyropasta (Hampson, 1914) comb. n., Hampsonola nanographa (Hacker, 2012) comb. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD5B15AFF33FB4FFE3DFB95.taxon	description	Description of the male genitalia (Fig. 24). Uncus short, narrow, gradually tapered, apically pointed. Tegumen long and narrow. Subscaphium short and narrow, weakly sclerotized, proximally finely scobinate. Valva broad at base, costal lobe long and narrow, apically slightly clubbed, ventral lobe short, ventral margin evenly rounded, dorsal margin produced into a short point. Medio-distal part of valva with a short rugged crest bearing long androconial scales. Sacculus short and broad, trapezoidal; harpe very long, slightly S-curved, apically dilated and rounded, twice as long as length of ventral valval lobe. Juxta short, simple, overturned C-shaped. Vinculum moderately long, very narrow, apically pointed. Aedeagus short and thin, slightly S-curved, postmedially tapered, apically slightly dilated bearing a rounded subapical plate; coecum penis short, rounded; vesica without cornuti or scobination. Description of the female genitalia (Figs 34 – 35). Ovipositor rather short and broad, papilla analis trapezoidal, posterior apophysis rather long and thick. Eighth tergite very short, medially constricted, anterior apophysis almost as long as posterior one, rather thin. Ostium bursae moderately sclerotized, cup-shaped, ductus bursae tubular, relatively long, moderately thick; cervix bursae unmodified, ductus seminalis relatively thick. Distal tubular part of corpus bursae short, ca. one-fifth of length of bursa, dilated proximal part of corpus bursae ovoid, with two welldeveloped, apically acute, more or less thorn-shaped signum situated wide apart from one another. Genetic information. The species has been assigned the BIN URI: BOLD: AEK 6843. The intraspecific divergence based on two samples is 1.08 %. The nearest neighbour of H. argyropasta is H. nanographa with a pairwise distance of 3.31 %. The genetic divergence between H. argyropasta and H. elachistomorpha is discussed under the latter species.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD2B15AFF33FB2BFD72F83A.taxon	description	(Figs 9 – 10, 25, 36)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD2B15AFF33FB2BFD72F83A.taxon	materials_examined	Material examined. Gabon. 6 males, 3 females, 430 m, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 0 ° 29 ’ 47 ” N, 11 ° 10 ’ 42 ” E, 28. vii. – 12. viii. 2019, Actinic, MV and LepiLED Light Trap, Albert, J-L., Aristophanous, M., Bie Mba, J., Derozier, V., Moretto, P. leg., ANHRT: 2019.17, gen. slide Nos: LGNA 1227 (male), LGNA 1228 (female), BOLD process / DNA barcode ids: ANLMN 2802 - 21 / ANHRTUK 00137825, ANLMN 2803 - 21 / ANHRTUK 00138677, ANLMN 2804 - 21 / ANHRTUK 00162579 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD2B15AFF33FB2BFD72F83A.taxon	description	Description of the female genitalia (Fig. 36). Ovipositor rather short and broad, papilla analis trapezoidal, posterior apophysis moderately long and thick. Eighth tergite very short, medially constricted, anterior apophysis 20 % shorter than posterior one, rather thin. Ostium bursae moderately sclerotized, cup-shaped, ductus bursae tubular, relatively long, moderately thick; cervix bursae unmodified, ductus seminalis relatively thick. Distal tubular part of corpus bursae short, ca. one-fifth of bursa length, dilated proximal part of corpus bursae ovoid, with two small, sclerotized, amorphous plates of signa, each bearing very short, acute process; signa situated wide apart from one another. Genetic information. The species has been assigned the BIN URI: BOLD: AAQ 1292. The average intraspecific pairwise distance according to BOLD is 0.40 %. The nearest neighbour of H. nanographa is H. argyropasta with 3.31 % pairwise distance between the two species. The genetic divergence between H. nanographa and H. elachistomorpha is discussed under the latter species.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD0B158FF33FF72FCD9FB52.taxon	description	(Figs 11 – 12, 26)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD0B158FF33FF72FCD9FB52.taxon	materials_examined	Type material examined. Syntype, male. Red circular label “ Type ” / “ Sierra Leone. W. G. Clements. 99 - 116. July 1895 ” / with handwritten “ Nola transecta. type ♂. Hmpsn. ” / QR code label with unique number “ NHMUK 010598812 ” (NHMUK). Syntype male. “ Sierra Leone. W. G. Clements. 99 - 116 ” / QR code label with unique number “ NHMUK 014173187 ” / genitalia slide No. “ SLIDE NHMUK 010317594 ” (prepared by László) (NHMUK). Taxonomic note. Hacker et al. (2012) illustrated male and female syntypes of Nola transecta Hampson, 1901, following the original generic combination without attempting to revise the generic assignment of the species. The author of this paper had the opportunity to dissect a male syntype in order to obtain morphological information to clarify the taxonomy of this peculiar species. The configuration of the male genitalia has revealed a close affinity to Hampsonola characterized by the bilobate valva comprised of a narrow, long dorsal lobe and a short ventral lobe and the conspicuously elongate harpe. This species, however, does not have androconial scales on the valva, additionally possessing a fully developed uncus, reminiscent of Meganola and an extensive ventro-distal sclerotized plate of the valva. It is difficult to establish whether these characters represent generic autapomorphies without conducting in depth genetic studies, therefore the species is tentatively placed in the genus Hampsonola: Hampsonola transecta (Hampson, 1901) comb. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD0B158FF33FF72FCD9FB52.taxon	description	Description of the male genitalia (Fig. 26). Uncus rather long, narrow, medially slightly dilated, distally tapered, apically pointed. Tegumen elongate, very narrow; subscaphium short, narrow, weakly sclerotized. Valva narrow basally, dorsal lobe long, slightly curved, distally gradually dilated with apex rounded, costal margin rather wide at base, then abruptly narrowed and gradually dilated towards apex. Ventral lobe of valva moderately long, more or less quadrangular, distal margin depressed, ventro-distally bearing a conspicuous, heavily sclerotized, broad L-shaped plate. Sacculus short and narrow, weakly sclerotized; harpe relatively small, bulb-shaped basally continued in a very thin, gently curved needle-like distal process. Juxta small, shield-like; vinculum moderately long, pointed V-shaped. Aedeagus short, dilated medially, gradually tapered to pointed apex, coecum moderately long, narrow, rounded; vesica without cornuti or scobination.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FF3AFB1BFB74.taxon	description	(Figs 13, 27)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FF3AFB1BFB74.taxon	type_taxon	Type species: Gabonola smithi sp. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FF3AFB1BFB74.taxon	diagnosis	Diagnosis. The male genitalia of Gabonola gen. n. is characterized by the fully sclerotized dorsal lobe of the valva bearing a rounded subapical lobe and a long, acute apical process in contrast to the ventrally membranous, apically rounded dorsal valval lobe of Hampsonola lacking any lobes or processes. While the harpe is well-developed in Hampsonola, it is fully reduced in Gabonola, substituted by a curved, sclerotized band lacking any prominence or process. The configuration of the ventral lobe of the valva is highly characteristic in its dense distal scobination. Compared to Vandamia van Son, the new genus lacks the conspicuous hair tuft of the uncus and the dorsal valval lobe which are diagnostic features in the former genus, while Vandamia always has an apically rounded dorsal valval lobe and a well-developed harpe.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FF3AFB1BFB74.taxon	description	Description. Adult. Male (Fig. 13). Forewing length of type species 6.7 mm. Head small, antenna bipectinate with short, finely ciliate rami; labial palp short, porrect, cream; frons and vertex cream. Collar, tegula, mesothorax and abdomen brown. Forewing elongate-triangular, apically rounded, ground colour ochreous brown. Transverse lines diffuse, consisted of small darker brown patches. Cilia long, dark brown. Hindwing greyish-brown, veins and cilia darker than ground colour. Underside of wings greyish-brown. Male genitalia (Fig. 27). Uncus moderately long, narrow, apically rounded. Tegumen short and very narrow; subscaphium membranous, sparsely scobinate. Valva broad basally, bilobate distally, dorsal lobe relatively broad, elongate, uniformly sclerotized, postmedially with a semi-circular lobe, apically with a long, robust, acute process. Ventral lobe of valva short, relatively broad, rounded, distal half densely scobinate. Sacculus large, roundedtrapezoidal, weakly sclerotized, without processes. Juxta very narrow, V-shaped with long lateral arms. Vinculum robust, V-shaped, heavily sclerotized. Aedeagus short and narrow, dilated at ostium ducti ejaculatorii, coecum elongate, rounded, distal half of aedeagus digitiform, subapically slightly curved, apically broadly rounded; vesica without cornuti or scobination.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FF3AFB1BFB74.taxon	etymology	Etymology. The generic name refers to the country of origin of the new noline taxon.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FB08FC4EF884.taxon	description	(Figs 13, 27)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FB08FC4EF884.taxon	materials_examined	Holotype. Male. “ Gabon, 430 m, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 0 ° 29 ’ 47 ” N, 11 ° 10 ’ 42 ” E, 28. vii. – 12. viii. 2019, LepiLED Light Trap, Albert, J-L., Aristophanous, M., Bie Mba, J., Derozier, V., Moretto, P. leg., ANHRT: 2019.17 ”, unique id.: ANHRTUK 00111212, gen. slide No.: LGNA 1171, BOLD process / DNA barcode id.: ANLMN 2805 - 21 / ANHRTUK 00111212 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FB08FC4EF884.taxon	diagnosis	Diagnosis. The distinctive characters of the new species are discussed under the diagnosis of the new monotypic genus Gabonola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FB08FC4EF884.taxon	description	Description. The description of the external and genital morphology of Gabonola smithi sp. n. is given under the description of the new monotypic genus Gabonola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD1B159FF33FB08FC4EF884.taxon	etymology	Etymology. The new species is dedicated with great pleasure to Mr Richard Smith, Chairman of the Board of Trustees, African Natural History Research Trust, acknowledging his merits in advancing the entomological research in Sub-Saharan Africa. Genetic information. The new species has been assigned the BIN URI: BOLD: AEK 0327. As the species is known by its holotype only, the intraspecific genetic variability could not be calculated. The nearest neighbour of G. smithi in the BOLD database is an unidentified, probably undescribed Hampsonola (BOLD process / DNA barcode id.: GWOUA 461 - 19 / BC ZSM Lep 105017, BIN URI: BOLD: AEA 0093).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDEB157FF33F947FC4AF9E1.taxon	description	(14 – 15, 28)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDEB157FF33F947FC4AF9E1.taxon	type_taxon	Type species: Bellanola mikongo sp. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDEB157FF33F947FC4AF9E1.taxon	diagnosis	Diagnosis. The new genus is characterised externally by the extensive creamy-white basal area of the forewing which extends towards the undulate, interrupted postmedial line and the dark brownish-grey postmedial area with the fine, interrupted, creamy-white, dentate subterminal line and the continuous terminal line. The external habitus of Bellanola gen. n. is somewhat reminiscent of Laetonima László, 2020, however the latter is larger in size (forewing length 7.5 – 8.5 mm in males) and it has a more quadrangular, apically more broadly rounded forewing, a considerably smaller and brighter whitish area restricted to the ventro-basal third of the forewing, as well as a large, trapezoidal ventro-medial blotch followed by an oblique bright white tornal dash which is absent in Bellanola. The small size and the narrow-triangular, apically pointed forewing of the type species of the new genus are alike in Hampsonola, and the similar ground plan of the male genitalia, i. e. the similarly bilobate valva possessing a narrow, elongate dorsal and a broad, short ventral lobe suggests certain affinities between the two genera. Nevertheless, Bellanola is readily distinguished from Hampsonola by the ventral valval lobe having a broad sclerotized margin (it is always membranous in Hampsonola) bearing two short saccular processes and the absence of the harpe. The aedeagus in Bellanola is considerably shorter and broader than in Hampsonola, lacking the coecum and possessing a robust acute carina process, while the aedeagus of Hampsonola bears a coecum penis and lacks the carina process. Comparing the male genital morphology of Bellanola to Laetonima, the type species of the new genus possesses a considerably shorter and broader, rather quadrangular ventral valval lobe, a markedly smaller distal saccular process, and bears an additional, short subbasal saccular process, while Laetonima has only a single, large apical lobe of the sacculus. The configurations of the aedeagus are substantially different in Bellanola and Laetonima: the type species of the new genus has a remarkably short and thick, distally slightly dilated aedeagus lacking the coecum penis but bearing a medium long, thorn-shaped carina process, whereas the aedeagus is tubular, relatively short and narrow with a medium long coecum penis, and with a short, triangular, weakly sclerotized carina process in Laetonima. The female is unknown.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDEB157FF33F947FC4AF9E1.taxon	description	Description. Adult. Male (Figs 14 – 15). Forewing length of type species 6.7 – 7 mm. Head small, antenna ciliatefasciculate; labial palp very short, porrect, greyish-white; frons and vertex off-white. Collar, tegula, mesothorax and abdomen off-white. Forewing narrow, elongate-triangular, apically pointed, ground colour creamy-white in basal and medial area, with sparse greyish suffusion along costa, dark brown in postmedial and terminal area, off-white apically and subapically. Basal, subbasal, antemedial and medial transverse lines deleted, postmedial line undulate, interrupted, consisted of blackish dots. Subterminal line creamy-white, represented by short, interrupted dentate sections. Terminal line very fine, continuous, creamy-white. Cilia long, blackish brown. Hindwing off-white in proximal half, greyish-brown in distal half with darker veins; cilia long, brownish-grey. Underside of wings whitishgrey ventro-basally, dark grey elsewhere. Male genitalia (Fig. 28). Uncus moderately long, narrow, subapically curved, apically pointed. Tegumen elongate, narrow, arms thin, somewhat dilated medially. Subscaphium weakly sclerotized, short, rounded. Valva rather broad at base, transtilla wide basally, abruptly tapered subapically. Dorsal lobe of valva moderately long, narrow proximally, dilated distally, rounded apically; costal margin sclerotized in basal half. Ventral lobe of valva short, more or less quadrangular, proximal and distal margin broadly sclerotized. Sacculus relatively broad basally, tapered distally bearing a short, triangular distal and a longer, thorn-shaped subbasal process. Juxta very thin, broad U-shaped; vinculum moderately broad basally medium long, apically produced into a thin, pointed process. Aedeagus very short, thick, distally dilated, coecum penis absent, carina with moderately long and thick, straight, pointed process; vesica without cornuti or scobination. Female. Unknown.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDEB157FF33F947FC4AF9E1.taxon	etymology	Etymology. The prefix of the generic name is derived from the Latin adjective “ bella ” meaning “ beautiful ” referring to the attractive appearance of the type species of the new genus.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDFB154FF33F9BFFF74FE61.taxon	description	(Figs 14 – 15, 28)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDFB154FF33F9BFFF74FE61.taxon	materials_examined	Holotype. Male. “ Gabon, 430 m, Mikongo (Rougier), Monts de Cristal (Secondary Forest), 0 ° 29 ’ 47 ” N, 11 ° 10 ’ 42 ” E, 28. vii. – 12. viii. 2019, MV Light Trap, Albert, J-L., Aristophanous, M., Bie Mba, J., Derozier, V., Moretto, P. leg., ANHRT: 2019.17 ”, unique id.: ANHRTUK 00165526, gen. slide No.: LGNA 1294, BOLD process / DNA barcode id.: ANLMN 2856 - 21 / ANHRTUK 00165526 (ANHRT). Paratype. Male, with the same data as in the holotype, but collected by actinic light trap, unique id.: ANHRTUK 00162619, BOLD process / DNA barcode id.: ANLMN 2855 - 21 / ANHRTUK 00162619 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDFB154FF33F9BFFF74FE61.taxon	diagnosis	Diagnosis. The distinctive characters of the new species are discussed under the diagnosis of the new monotypic genus Bellanola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDFB154FF33F9BFFF74FE61.taxon	description	Description. The external and genital morphology of Bellanola mikongo sp. n. are discussed under the description of the new monotypic genus Bellanola. Genetic information. The species belongs to the BIN URI: BOLD: AEL 7152. The intraspecific genetic divergence between the two known specimens is 0.15 %. Within the species discussed in this study, the shortest pairwise genetic distance was observed between B. mikongo and H. argyropasta in the range of 8.58 – 9.10 %.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDFB154FF33F9BFFF74FE61.taxon	etymology	Etymology. The specific name refers to the type locality of the new species, the Mikongo Forest in central Gabon.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDCB154FF33FE3FFCC6F85C.taxon	description	(Figs 16 – 17, 29)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDCB154FF33FE3FFCC6F85C.taxon	type_taxon	Type species: Brunneonola nimba sp. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDCB154FF33FE3FFCC6F85C.taxon	diagnosis	Diagnosis. The facies of the type species of the new genus might suggest certain kinship with Hampsonola, however, the genital morphology of the two genera is fundamentally different as Brunneonola gen. n. has a unilobate valva, unlike the bilobate valva of Hampsonola. The external appearance of the new genus is characterised by the relatively short and broad, somewhat quadrangular forewing, and the extensive pale greyish-brown basal area extending towards the postmedial line which runs conspicuously close to the termen and parallel with the subterminal line. The costal area of the forewing bears a large, diffuse dark brown blotch postmedially, and the terminal area is bright red-brown, gradually narrowing towards the tornus. In the male genitalia, Brunneonola has a unilobate valva with a remarkably broad, sclerotized dorsal margin dilated distally and forming a rounded triangular process apically, a character which is unknown in any other Afrotropical Nolinae, although similar costal valval processes are observed in a several species of the Oriental genus Manoba Walker. The bifurcate harpe is also an uncommon feature in Nolinae, although it has also been found in Meganola. The most unique character of Brunneonola is the presence of a long and thin, apically dilated and dentate process arising from the valval base, a feature which is only known in the Oriental Maculonola László, Ronkay & Ronkay, 2015. The female is unknown.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDCB154FF33FE3FFCC6F85C.taxon	description	Description. Adult. Male (Figs 16 – 17). Forewing length of type species 5.8 – 6.7 mm. Head small, antenna short, bipectinate with short, finely ciliate rami, distal quarter filiform; labial palp relatively long, porrect, pale brown; frons and vertex pale brown. Collar rusty-brown, tegula and mesothorax off-white, and abdomen light brown. Forewing relatively broad, elongate-quadrangular, apically rounded, ground colour pale brownish-grey in basal and medial area, dark brown in postmedial and red-brown in terminal area. Basal and subbasal lines deleted, antemedial line gently arched, slightly undulate. Medial line deleted, postmedial line very fine, greyish-brown, evenly arched, interrupted. Subterminal line narrow, ochreous, slightly wavy; terminal line very fine, ochreous. Cilia moderately long, red-brown in dorsal, blackish brown in ventral half. Hindwing pale brownish-grey with slightly darker veins; cilia long, dark brown. Underside of wings dark brownish-grey, hindwing somewhat paler the forewing. Male genitalia (Fig. 29). Uncus medium long, relatively broad at base, gradually tapered, subapically curved, apically pointed. Tegumen elongate, narrow with very thin arms; subscaphium membranous, with a finely scobinate semi-circular area proximally. Valva relatively narrow in its entire length, dorsal margin concave, very broadly sclerotized, subapically dilated, apically produced into a short, rounded triangular process. Ventral margin membranous, distally slightly dilated, apex truncate. Harpe broad basally, bifurcate, consisting of two short, rounded triangular lobes arising relatively far apart, pointing caudad. Sacculus short and narrow, distally rounded. Valva base possessing a narrow, ribbon-like process, projecting caudad with dilated, roughly dentate ovoid apex curved inwards. Transtilla thin, weakly sclerotized, arms fused medially. Juxta short, simple, quadrangular; vinculum short, rounded. Aedeagus short and thin with long coecum and rounded apex bearing a short carinal dent subapically; vesica without cornuti or scobination. Female. Unknown.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDCB154FF33FE3FFCC6F85C.taxon	etymology	Etymology. The prefix in the generic name is derived from the Latin adjective “ brunneus ” meaning “ brown ” referring to the coloration of the type species of the new genus.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB155FF33FF72FABDFC69.taxon	description	(Figs 16 – 17, 29)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB155FF33FF72FABDFC69.taxon	materials_examined	Holotype. Male. “ Liberia, 750 m, Nimba County, Nimba Mts., ENNR, Cellcom road, 7 ° 33 ’ 3.78 ” N, 8 ° 31 ’ 46.49 ” W, 16 – 28. xii. 2018 Cold Cathode UV Light Trap (8 W), Sáfián, Sz., Simonics, G. leg. ANHRT: 2018.43 ”, unique id.: ANHRTUK 00073957, gen. slide No.: LGNA 1158, BOLD process / DNA barcode id.: ANLMN 2810 - 21 / ANHRTUK 00073957 (ANHRT). Paratypes. 4 males, with the same data as in the holotype, unique ids: ANHRTUK 00073955, 0073956,00073959, 00143276, gen. slide No.: LGNA 1303, BOLD process / DNA barcode ids: ANLMN 2809 - 21 / ANHRTUK 00073955, ANLMN 2808 - 21 / ANHRTUK 00073959 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB155FF33FF72FABDFC69.taxon	diagnosis	Diagnosis. The distinctive features of the new species are discussed under the diagnosis of the new monotypic genus Brunneonola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB155FF33FF72FABDFC69.taxon	description	Description. The external and genital morphology of Brunneonola nimba sp. n. are discussed under the description of the new monotypic genus. Genetic information. The new species has been assigned the BIN URI: BOLD: AEJ 3404. The intraspecific genetic divergence is 0.31 %. The nearest neighbour according to the BOLD database is a noline species identified as Callinola bifiliferata from South America, diverging at 6.41 %, however, the relatively short genetic distance based on COI barcodes only, does not necessarily refer to a close relationship between the species. Within the taxa discussed in this study, the shortest pairwise genetic distance was observed between B. nimba and H. nanographa in the range of 7.90 – 8.24 %.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB155FF33FF72FABDFC69.taxon	etymology	Etymology. The specific name refers to the type locality of the new species, the Nimba Mountains.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB152FF33FC36FE17FF11.taxon	description	(Figs 18, 30)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB152FF33FC36FE17FF11.taxon	type_taxon	Type species: Ivindonola ipassa sp. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB152FF33FC36FE17FF11.taxon	diagnosis	Diagnosis. The facies of the type species of the new genus is reminiscent of Brunneonola due to its similarly small size, brownish coloration and the red-brown terminal area of the forewing. Despite the similar habitus, Ivindonola displays a rather unique and distinct genital morphology expressed by the long, pointed and densely setose uncus, a character which is rather prevalent in Meganola, the remarkably narrow tegumen, the medially strongly constricted, unilobate valva, the lack of a harpe, and the presence of a short membranous lobe of the sacculus. The combination of these characters is unknown in any other noline genera and thus a new monotypic genus is erected here.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB152FF33FC36FE17FF11.taxon	description	Description. Adult. Male (Fig. 18). Forewing length of type species 7 mm. Head small, antenna short, bipectinate with short, finely ciliate rami, distal quarter filiform; labial palp short, porrect, dark brown; frons and vertex grey. Collar and tegula dark grey, suffused by pale grey scales, mesothorax and abdomen pale greyish-brown. Forewing relatively broad, elongate-quadrangular, apically rounded, ground colour pale creamy-grey with sparse, very fine dark brown suffusion; costal margin with small, rounded patches; dorsal two-thirds of terminal area redbrown. Basal and subbasal lines deleted, antemedial line gently arched, diffuse. Medial line deleted, postmedial line very fine, greyish-brown, interrupted. Subterminal line rather broad, dark brown, interrupted, represented by dark brown patches in dorsal-, small dots in ventral area; terminal line very fine, ochreous. Cilia moderately long, red-brown in dorsal, cream in ventral half. Hindwing grey, somewhat darkened apically veins slightly darker; cilia long, proximal half dark grey, distal half pale greyish-brown. Underside of wings ochreous brown with well-defined discal spots. Male genitalia (Fig. 30). Uncus relatively long, distal quarter tapered and densely setose, apex pointed. Tegumen very narrow and elongate, arms thin; subscaphium membranous with elongate, densely scobinate plate. Valva long, relatively narrow, medially strongly constricted, dorsal margin concave basally, slightly convex submedially, straight distally, weakly sclerotized. Valva apex broadly rounded, densely scobinate, ventral margin deeply concave medially, gently convex in proximal half. Harpe absent; sacculus narrow, bearing a short, longitudinal lobe distally. Transtilla weakly sclerotized, ribbon-like arms medially fused. Juxta small, U-shaped, vinculum short, V-shaped. Aedeagus long and very thin, gently S-curved, coecum long, apically bifid, Y-shaped, carina absent; vesica without cornuti or scobination. Female. Unknown.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDDB152FF33FC36FE17FF11.taxon	etymology	Etymology. The prefix in the generic name refers to the type locality of the type species of the new genus, the Ivindo National Park.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB152FF33FEAFFE26FCB5.taxon	description	(Figs 18, 30)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB152FF33FEAFFE26FCB5.taxon	materials_examined	Holotype. Male. “ Gabon, Ogooue Ivindo P. N., Ivindo Station de Recherche d’Ipassa, 450 m, 0 ° 30 ’ 43 ” N, 12 ° 48 ’ 12 ” E, 14 – 26. vi. 2016, Light Trap Ruzzier, E., Tasane, T. leg. ANHRT: 2017.19 ”, unique id.: ANHRTUK 00042531, gen. slide No.: LGNA 590, BOLD process / DNA barcode id.: ANLMN 2811 - 21 / ANHRTUK 00042531 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB152FF33FEAFFE26FCB5.taxon	diagnosis	Diagnosis. The distinctive characters of the new species are discussed under the diagnosis of the new monotypic genus Ivindonola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB152FF33FEAFFE26FCB5.taxon	description	Description. The external and genital morphology of Ivindonola ipassa sp. n. are described under the description of the new monotypic genus Ivindonola. Genetic information. Ivindonola ipassa has been assigned the BIN URI: BOLD: AEK 9606. As the species is known only from its holotype, intraspecific genetic divergence could not be calculated. The pairwise distances between I. ipassa and all other taxa discussed in this paper exceed 9 %.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB152FF33FEAFFE26FCB5.taxon	etymology	Etymology. The specific name refers to the type locality of the new species, the Ipassa-Makokou Biosphere Reserve in central Gabon.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB153FF33FC4BFCBAFCD1.taxon	description	(Figs 19 – 21, 31, 37)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB153FF33FC4BFCBAFCD1.taxon	type_taxon	Type species: Cryptonola confundata sp. n.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB153FF33FC4BFCBAFCD1.taxon	diagnosis	Diagnosis. The facies of the type species of Cryptonola gen. n. could fit well in the mega-diverse genus Meganola based on the bipectinate antenna possessing long rami, the narrow, elongate, rather pointed forewing and the brownish-grey coloration with diffuse, oblique transverse lines. The configuration of the male genitalia, however, displays no close kinship between the two genera, but suggests closer affinities with the Evonima Walker lineage, in particular with Vansonima László, 2020 due to the presence of a seemingly homologous process arising rather distally from a medio-distal plate extending from the dorsal margin of the valva and projecting ventrad (referred to as harpe, probably incorrectly, in Hacker et al. (2012) and László (2020 )). Nevertheless, the following diagnostic characters undoubtedly support the distinctness of Cryptonola: the uncus is remarkably stout and short, covered in long, dense setae, and quite uniquely for the whole subfamily, associated with a pair of large, triangular socii basally. Compared to Vansonima, the valva of Cryptonola is much narrower and more elongate, bearing a much shorter and narrower, more pointed ventro-distal process and an additional saccular process similar in shape, projecting nearly from the base of the ventro-distal process.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB153FF33FC4BFCBAFCD1.taxon	description	Description. Adult (Figs 19 – 21). Forewing length of type species 8 mm in male, 9.3 mm in female. Sexual dimorphism limited, expressed by the larger size and darker coloration in females. Head small, male antenna bipectinate with long, finely ciliate rami, distal third filiform, that of female filiform in entire length; labial palp relatively long and slim in male, somewhat broader in female, porrect, dark brown laterally, inner side off-white; frons and vertex bright white. Collar, tegula and mesothorax off white, abdomen brownish-grey. Forewing narrow, elongate-triangular, apically rounded, ground colour pale grey with sparse, very fine blackish-brown suffusion; costal margin dark brown, medially with a short, blackish dash; terminal area dark greyish-brown. Basal and subbasal lines deleted, antemedial line arched, diffuse. Medial- postmedial- and subterminal lines diffuse, shadowlike, barely traceable. Terminal line pale brown, chequered with black. Cilia moderately long, pale brown chequered with dark grey. Hindwing grey, somewhat darkened apically veins slightly darker; cilia long, colour as ground colour. Underside of wings uniformly dark grey. Male genitalia (Fig. 31). Uncus short, very broad basally, conical, lateral margins gently arched, distal half covered in dense, long, spine-like setae, apex blunt, dorso-apically bearing a moderately long and broad, slightly curved, apically pointed, knife-shaped process. Socius well-developed, long and stout, elongate-triangular, distally slightly curved, apex pointed, distal margin densely spinulose. Tegumen short and narrow, arms relatively wide, sparsely scobinate; subscaphium relatively broad, elongate, moderately sclerotized. Valva narrow, rather long, dorsal margin slightly concave, ventral margin gently convex, distal third slightly dilated, broadly rounded. Sclerotization of dorsal margin largely broadened postmedially covering two-thirds of the surface of distal dilated part of valva, produced into a narrow triangular, apically rounded process projecting ventrad, slightly overreaching ventral margin. Sacculus long and narrow, with dorsal margin gently sinuous, distally produced into a short, but robust, apically hooked process, overlapping with costal projection, overraching ventral valval margin. Transtilla very narrow, lace like, arms fused medially. Juxta relatively large, basally rounded with well-sclerotized, sinuous lateral arms. Vinculum short and broad, V-shaped. Aedeagus relatively short and thin, tubular, coecum penis short, broadly rounded, carina absent; vesica without cornuti or scobination. Female genitalia (Fig. 37). Ovipositor short, papilla analis narrow, apically pointed, moderately setose, posterior apophysis relatively long, thin. Eighth tergite short, ribbon-like with parallel margins, anterior apophysis very short, thin apically clubbed. Ostium bursae narrow, with gently arched, fine, sclerotized margin antero-laterally. Ductus bursae short, tubular, membranous, cervix bursae, slightly swollen, membranous. Posterior half of corpus bursae tubular, gradually dilating proximad, distal half of corpus bursae sack-like; signum bursae consisted of two large, thorn-like strongly sclerotized process of uniform shape, situated close to one another.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFDAB153FF33FC4BFCBAFCD1.taxon	etymology	Etymology. The generic name refers to the puzzling taxonomy of the genus where a rather unusual genital morphology is combined with a typical Meganola habitus.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD9B151FF33FF72FBB0FC22.taxon	description	(Figs 19 – 21, 31, 37)	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD9B151FF33FF72FBB0FC22.taxon	materials_examined	Holotype. Male. “ Ivory Coast, 40 m, Banco National park, 05 ° 23 ’ 3.8 ” N, 04 ° 03 ’ 11.2 ” W, 29. xi. – 5. xii. 2019, MV Light Trap, Aristophanous, M., Derozier, V., Moretto, P., Ouattara, S. leg., ANHRT: 2019.23 ”, unique id.: ANHRTUK 00134260, gen. slide No.: LGNA 1179, BOLD process / DNA barcode id.: ANLMN 2807 - 21 / ANHRTUK 00134260 (ANHRT). Paratypes. Ivory Coast. 1 female, Tai NP., Tai Research Station (SRET), 174 m, 05 ° 50 ’ 00 ” N, 07 ° 20 ’ 32.0 W, 25. iii. – 17. iv. 2017, MV light, Aristophanous, A., Aristophanous, M., Geiser, M., Moretto, P. leg., ANHRT: 2017.25, unique id.: ANHRTUK 00018629, gen. slide No.: LGNA, BOLD process / DNA barcode id.: ANLMN 2786 - 21 / ANHRTUK 00018629 (ANHRT). Sierra Leone. 1 male, Western Area Peninsula Forest Reserve, 180 m, N 08 ° 20 ’ 57 ”, W 13 ° 10 ’ 42 ” 24. x. 2015, Light Trap, R. Goff coll., leg. Smith, R & Takano, H., unique id.: ANHRTUK 00194157, gen. slide No.: LGNA 290, BOLD process / DNA barcode id.: ANLMN 2806 - 21 / ANHRTUK 00194157 (ANHRT).	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD9B151FF33FF72FBB0FC22.taxon	diagnosis	Diagnosis. The diagnostic features of the new species are discussed under the diagnosis of the new monotypic genus Cryptonola.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD9B151FF33FF72FBB0FC22.taxon	description	Description. The external and genital morphology of Cryptonola confundata sp. n. are discussed under the description of the new monotypic genus Cryptonola. Genetic information. The new species has been assigned the BIN URI: BOLD: AEJ 7184. The intraspecific divergence is 0.46 – 1.40 %. The nearest neighbour according to the BOLD database is a taxon identified as Meganola latifusca at 5.93 % pairwise distance.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
D82B87B5FFD9B151FF33FF72FBB0FC22.taxon	etymology	Etymology. The prefix “ confundatus ” in the specific name is a Latin adjective meaning “ confused ”, referring to the controversy between the external and genital morphology of the new taxon.	en	László, Gyula M. (2022): Taxonomic studies on Afrotropical Nolinae: review of the Afrotropical taxa of the genus Hampsonola László, Ronkay & Ronkay, 2015, descriptions of five new genera and five new species and introduction of a new synonymy (Lepidoptera, Nolidae). Zootaxa 5169 (2): 147-164, DOI: https://doi.org/10.11646/zootaxa.5169.2.3
