taxonID	type	description	language	source
54519917FF93FFD6087AF8FA0E5DE456.taxon	materials_examined	ISRAEL • 1 ♀; near Shoham, Hadom Shomeron Nature Reserve, Tinshemet cave; 31.99938 ° N, 34.96813 ° E; 122 m. a. s. l.; 20 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev and S. Aharon leg.; Mediterranean scrubland, under stones, outside the cave; MACN-Ar 41797.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF95FFD0085BFEB00E2DEE7F.taxon	materials_examined	ISRAEL • 2 ♂♂, 1 imm.; central Negev, Sede Boqer, Haluqim Ridge; [30.87801 ° N, 34.78559 ° E]; 2 Jul. 1992; Y. Lubin leg.; pitfall; HUJ-INVAr 21011 – 21013 • 2 ♂♂, holotype and paratype; same locality as for preceding; 4 May 1994; Y. Lubin leg.; HUJ-INVAr 21016 – 21017 • 1 ♂, paratype; same collection data as for preceding; SMNH • 2 ♂♂, paratypes; western Negev, Nizzana; [30.91666 ° N, 34.4 ° E]; 6 May 1992; Y. Lubin leg.; pitfall; HUJ-INVAr 21014 – 21015.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF95FFD008CCFB0109CBE97C.taxon	materials_examined	PALESTINE • 1 ♀, possibly subadult; Dead Sea Valley, En Fashkha (= ‘ Enot Zuqim); [31.72358 ° N, 35.4575 ° E]; 18 Feb. 1943; A. Shulov leg.; HUJ-INVAr 20277.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF95FFD008D4FAD30E55E9A0.taxon	materials_examined	ISRAEL • 1 ♀; Golan Heights, near Odem, Ya’ar Mas’ada Nature Reserve, Odem forest; 33.20351 ° N, 35.74009 ° E; 888 m. a. s. l.; 19 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev, Z. Ganem, S. Aharon and M. Arnedo leg.; Mediterranean woodland; under stones; MACN-Ar 41263.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF95FFD00824FDCE0ECEE82E.taxon	materials_examined	ISRAEL • 1 ♂, holotype; Negev, near Be’er Mash’abbim, sand dunes; pitfall; [31.0019 ° N, 34.7538 ° E]; 14 Mar. 1990; Y. Ayal leg.; HUJ-INVAr 21018 • 2 ♂♂, paratypes; same locality as for preceding; 21 Jul. 1990; Y. Ayal leg.; sand dunes; pitfall; HUJ-INVAr 21019 – 21020 • 1 ♂, paratype; same collection data as for preceding; SMNH • 3 ♂♂, paratypes; same locality as for preceding; 15 May 1991; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21021 – 21023 • 5 ♂♂, paratypes; same locality as for preceding; 17 May 1992; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21024 – 21028 • 1 ♂, paratype; same locality as for preceding; 27 May 1992; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21029 • 3 ♂♂, paratypes; same locality as for preceding; 3 Jul. 1991; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21030 – 21032 • 4 ♂♂, paratypes; same locality as for preceding; 30 Apr. 1993; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21033 – 21036 • 1 ♂, paratype; same collection data as for preceding; SMNH • 11 ♂♂, paratypes; same locality as for preceding; 8 Jul. 1992; Y. Lubin leg.; sand dunes; pitfall; HUJ-INVAr 21038 – 21048 • 1 ♂, paratype; same locality as for preceding; 13 May 1991; Y. Lubin leg.; HUJ-INVAr 21037 • 1 ♂; same locality as for preceding; Jun. 1991; Y. D. Lubin leg.; MCZ 39913.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF95FFD00839F9BC0B39EAFF.taxon	materials_examined	SPAIN • 4 ♀♀; Almeria, Boca de los Frailes; [36.80564 ° N, 2.1322 ° W]; 8 Dec. 2019; J. Moya leg.; CRBA.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF96FFD90870FACA09C1EFBF.taxon	description	Figs 2, 3 C – D, I, 5 B, D, F, 6, 8 H, 10 A, E, Q	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF96FFD90870FACA09C1EFBF.taxon	discussion	Remarks Please see WSC (2022) for a full synonymic list of this species, and Zonstein & Marusik (2019) for description, diagnosis, natural history, and additional distribution records.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF96FFD90870FACA09C1EFBF.taxon	diagnosis	Updated diagnosis Males are similar to those of F. betarif sp. nov. and F. albens by having an elongate pedipalpal femur and tibia, and by a subconical copulatory bulb with a terminally hooked embolus (Fig. 6; see also Zonstein & Marusik 2019: fig. 27). Males differ from those of F. albens by their longer copulatory bulb and from those of F. betarif sp. nov. by their inconspicuous embolic keel (Fig. 8 H (arrow), 5 B, k) (conspicuous keel in F. betarif sp. nov., Fig. 5 A). Females have a pair of large spermathecae that may be partially divided or undivided (Fig. 3 C – D) and their variation includes the observed morphology of F. betarif sp. nov.; thus females are not diagnosable through morphology alone, they can be diagnosed using DNA barcoding of the COI gene.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF96FFD90870FACA09C1EFBF.taxon	materials_examined	Material examined ANGOLA • 1 ♀; Luanda, Luanda; [8.83833 ° S, 13.23444 ° E]; N. Chatelain leg.; USNM 1656 • 1 ♀; Namibe, Baia das Pipas, in house; [14.95 ° S, 12.18333 ° E]; Jan. 1960; Brühl leg.; ZMB. EGYPT • 1 ♀; Cairo; [30.04442 ° N, 31.23571 ° E]; O. Manley leg.; ZMB 4375. FRANCE • 1 ♂, 1 ♀; Alpes-Côte d’Azur, Var, Brignoles, Gonfaron; 43.34158 ° N, 6.29443 ° E; 2017; M. Stockmann leg.; MACN-Ar 39461 • 2 ♂♂, 4 ♀♀; Languedoc-Roussillon, Banyuls-sur-Mer; [42.48343 ° N, 3.12887 ° E]; 21 Mar. 1968; H. W. Levi, F. Levi and L. R. Levi leg.; MCZ 39864 • 1 ♀; no further data; [42.44097 ° N, 3.16469 ° E]; Spatz leg.; ZMB 1053 • 2 ♀♀; southern France; AMNH • 1 ♀; southern France; ZMB 5021. GREECE • 1 imm.; Attica, Sounion; [37.65915 ° N, 24.01476 ° E]; 30 Nov. 1978; B. Malkin leg.; AMNH • 2 ♀♀; Crete, Akrotiri; [35.55 ° N, 24.13333 ° E]; Feb. – Mar. 1925; A. Schultz leg.; ZMB 6205 • 1 ♀; Crete, Agios Nikolaos; 35.31006 ° N, 25.57781 ° E; Jul. 2019; M. Stockmann leg.; MACN-Ar 41844 • 3 ♀♀, 1 imm.; NW Crete; [35.55 ° N, 24.13333 ° E]; A. Schultz leg.; ZMB 6208 • 1 ♀; Crete; [36.39316 ° N, 25.46151 ° E]; Oertzen leg.; ZMB 6951 • 1 ♀; Cyclades Islands, Santorini; 36.40922 ° N, 25.43221 ° E; May 1988; M. J. Ramírez leg.; MACN-Ar 20563 • 1 imm.; Kea; [37.60758 ° N, 24.31037 ° E]; 30 Sep. 1925; A. Schultz leg.; ZMB 6203 • 2 ♀♀; Kythira; [36.26327 ° N, 22.97737 ° E]; Aug. 1925; ZMB • 1 ♀; Rhodes, Kalavarda; [36.33998 ° N, 27.94949 ° E]; 7 Dec. 1978; B. Malkin leg.; AMNH • 1 ♀; Thera; [36.39316 ° N, 25.46151 ° E]; 1972; Hillas and Goerligen (?) leg.; ZMB. ISRAEL • 1 ♂; upper Galilee, Ba’al-Shem-Tov Forest, Meron Mountains; 32.9821 ° N, 35.47381 ° E; 13 – 20 Nov. 2007; T. Levanony leg.; SMNH • 1 ♂, 1 ♀, 1 imm.; upper Galilee, Dan; 33.24 ° N, 35.655 ° E; 14 May 2012; S. Zonstein leg.; SMNH • 1 ♂; upper Galilee, En Ya’aqov; 33.00764 ° N, 35.23889 ° E; 4 – 9 Feb. 2007; I. Shtirberg leg.; pitfall; SMNH • 1 ♂; upper Galilee, Nahal Keziv; [33.04234 ° N, 35.17882 ° E]; 31 Jul. 2010; C. Drees and L. Friedman leg.; SMNH • 1 ♂; same locality as for preceding; 13 Nov. 2010; L. Friedman and C. Drees leg.; SMNH • 1 ♂, 2 ♀♀; upper Galilee, near Elon, Nahal Bezet Nature Reserve, Sharakh cave, entrance of cave; 33.074 ° N, 35.2379 ° E; 233 m. a. s. l.; 17 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev, Z. Ganem, S. Aharon, N. Givon and M. Arnedo leg.; MACN- Ar 41218 • 1 ♂, 2 ♀♀; same collection data as for preceding; MACN-Ar 41226 • 1 ♂; same locality as for preceding; 17 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev, Z. Ganem, S. Aharon, N. Givon and M. Arnedo leg.; twilight zone; MACN-Ar 41261 • 1 ♀; same collection data as for preceding; MACN-Ar 41825 • 1 ♂; upper Galilee, Senir; 33.23333 ° N, 35.66667 ° E; 250 m. a. s. l.; 2 Nov. 2013; S. Zonstein leg.; SMNH • 1 ♀; upper Galilee, Yehiam, in cave; [32.99611 ° N, 35.22109 ° E]; 15 Nov. 1949; A. Shulov leg.; HUJ-INVAr 20290 • 1 ♀; upper Galilee, Yir’on, small cave; 33.0672 ° N, 35.4672 ° E; 28 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21049 • 1 ♀; upper Galilee, Yonim cave, in depth; 32.9236 ° N, 35.2168 ° E; 25 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ - INVAr 21050 • 1 ♂; upper Galilee, north slope of Mount Meron, Meron Field School; 33.01667 ° N, 35.39167 ° E; 1 Nov. 2013; S. Zonstein leg.; SMNH • 2 imm.; lower Galilee, near Teverya (Tiberias), Berniki, large cave; 32.7775 ° N, 35.5401 ° E; 27 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21051 • 1 ♀; lower Galilee, near Teverya (Tiberias), Berniki, medium cave; 32.7768 ° N, 35.5413 ° E; 27 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21052 • 1 ♂; Mount Karmel, HaKarmel Nature Reserve, Karmel Ridge, Nahal Me’arot; Gamal cave; [32.6705 ° N, 34.96598 ° E]; 4 Nov. 2013; D. Rotem leg.; SMNH • 1 ♀; Mount Karmel, near Hefa (Haifa), HaKarmel Nature Reserve, Karmel Ridge; Ornit cave, inside cave; 32.7567 ° N, 34.98975 ° E; 209 m. a. s. l.; 20 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev, Z. Ganem, S. Aharon and M. Arnedo leg.; MACN-Ar 41215 • 1 ♂, 1 ♀; same collection data as for preceding; MACN-Ar 41813 • 1 ♂; Mount Karmel, Yagur; [32.74153 ° N, 35.07683 ° E]; 4 Dec. 1939; HUJ-INVAr 20283 • 1 ♀; coast of Karmel, ‘ Atlit; [32.69088 ° N, 34.94291 ° E]; 8 Sep. 1944; HUJ-INVAr 20287 • 1 ♂; coast of Karmel, Ma'agan Mikha'el, malaise trap; [32.55871 ° N, 34.91782 ° E]; 2 – 28 Dec. 2010; W. Kuslitzky leg.; SMNH • 1 ♂; same locality as for preceding; 12 Oct. 1992; Y. Lubin leg.; HUJ • 1 ♀; coast of Karmel, route to Zikhron-Y’aqov [32.57391 ° N, 34.95198 ° E]; 24 Mar. 1940; HUJ-INVAr 20286 • 1 ♀; Judean Hills, ‘ Adullam National Park, Midras; 31.655 ° N, 34.94167 ° E; 15 Nov. 2012; S. Zonstein leg.; SMNH IFM- 0272 • 1 ♂, 1 ♀; Judean Hills, ‘ Adullam National Park; [31.64474 ° N, 34.96089 ° E]; Apr. 2003; U. Columbus leg.; SMNH • 2 ♂♂; same locality as for preceding; 20 May 2002; Y. Mandelik and A. Landsman leg.; SMNH • 1 ♂; Judean Hills, Ramat Avishur; [31.65 ° N, 34.92 ° E]; 14 May 2002; Y. Mandelik leg.; pitfall trap; HUJ • 1 ♀; Judean Mountains, Jerusalem; [31.78333 ° N, 35.2 ° E]; 29 May 1944; A. Shulov leg.; HUJ-INVAr 20288 • 1 ♀, 1 imm.; same locality as for preceding; Oct. 1936; A. Shulov leg.; HUJ-INVAr 20284 • 1 ♀; Judean Mountains, Newe Ya’akov (= Kefar Ivri); [31.84177 ° N, 35.24276 °]; 30 Dec. 1939; A. Shulov leg.; HUJ-INVAr 20289 • 1 ♀; Judean Mountains, Qiryat'Anavim, under bark; [31.80991 ° N, 35.12084 ° E]; 20 Jan. 1940; HUJ-INVAr 20285 • 1 ♂; Samaria, Gilboa'; [32.48248 ° N, 35.42137 ° E]; 30 Jan. 2011; L. Friedman and C. Drees leg.; SMNH • 1 ♀; Samaria, Sal’it cave; 32.2454 ° N, 35.0456 ° E; 25 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21053 • 1 ♂; Sharon plain, Ramot Hashavim, in house; 32.16667 ° N, 34.88333 ° E; 9 Mar. 2013; T. Eshcoly leg.; SMNH • 1 ♂; Jordan Valley, Nahal HaGal; [32.6341 ° N, 35.534 ° E]; 16 Jul. 2010; C. Drees and L. Friedman leg.; SMNH • 2 imm.; Jordan Valley, Nir Dawid (= Tel’ Amal); [32.50406 ° N, 35.45762 ° E]; 29 Sep. 1939; A. Shulov leg.; HUJ-INVAr 21061. ITALY • 1 ♂, 1 ♀; Sardinia, Baunei; [40.03175 ° N, 9.66327 ° E]; J. Wunderlich leg.; SMF 59579 • 1 ♂; Sardinia, Porto Torres; [40.83134 ° N, 8.40587 ° E]; 7 Apr. 1952; Esk. Zool. Just. Ffn. Leg.; SMF • 1 ♀; Sorgono; [40.0272 ° N, 9.10196 ° E]; 1917; Häuser and Krausse leg.; ZMB • 7 ♀♀, 6 imm.; Toscania, Siena, 4 km S of San Giminiano, Fattoria Voltrona; [43.46728 ° N, 11.04348 ° E]; 12 Dec. 2001; M. J. Ramírez leg.; MACN-Ar 10378 • 1 ♀; same collection data as for preceding; MACN-Ar 10379. JORDAN • 1 ♂; Amman, Fuhais; [32.00408 ° N, 35.78451 ° E]; Dec. 1980; R. Kinzelbach leg.; SMF 57068. MOROCCO • 1 ♀; Saffi; [32.30082 ° N, 9.2272 ° W]; Quedenfeldt leg.; ZMB • 1 ♂, 3 ♀♀; Sefrou, Travertine Falls; 33.83333 ° N, 4.83333 ° W; 25 Dec. 1986; V. and B. Roth leg.; CAS 9060650 • 1 imm.; same locality as for preceding; 25 Dec. 1996; V. Roth leg.; CAS 9057617 • 1 ♀, 1 imm.; Souss-Massa- Draâ, Ouarzazate, Amerzgane, Telouet; 31.06659 ° N, 7.14178 ° W; Oct. 2016; M. Stockmann leg.; MACN-Ar 41845 • 1 ♂, 1 ♀; Tafilalt, Errachidia, Aoufous, Ziz Valley; 31.75176 ° N, 4.1968 ° W; Oct. 2016; M. Stockmann leg.; MACN-Ar 39463. PALESTINE • 1 ♀; Judean desert, 15 km E of Jerusalem, Fawar Spring, Wadi Qelt; [31.8401 ° N, 35.3489 ° E]; 7 Mar. 1975; H. W. Levi leg.; MCZ 39890 • 1 ♀, 1 imm.; Judean desert, Nahal Perat Nature Reserve, Inbal cave; 31.8332 ° N, 35.3019 ° E; 26 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21054 • 1 ♂, 1 ♀; Judean desert, Nahal Perat Nature Reserve, I’qul cave; [31.8318 ° N, 35.30842 ° E]; 26 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 20276. SPAIN • 4 ♀♀; Alicante, Benidorm; [38.54106 ° N, 0.12249 ° W]; 1961; K. W. Haller leg.; AMNH • 1 ♂, 1 ♀, 3 imm.; same collection data as for preceding; AMNH • 1 ♀; Almeria, 25 km from Almeria, Salinas de Cerillos; [36.70498 ° N, 2.66982 ° W]; 25 Jun. 1967; B. Malkin leg.; AMNH • 1 imm.; Balearic Islands, Mallorca, Colonia de Sant Jordi, 50 km SE of Palma, on house walls; 39.31727 ° N, 2.99381 ° E; 15 Sep. 2007; M. J. Ramírez leg.; MACN-Ar 20558 • 1 ♀; same collection data as for preceding; MACN-Ar 20560 • 1 ♀; same collection data as for preceding; MACN-Ar 20561 • 1 imm.; same collection data as for preceding; MACN-Ar 20564 • 1 ♀; Balearic Islands, Mallorca, Colonia Saint Jordi, 50 km SE of Palma, on walls of houses; [39.31727 ° N, 2.99381 ° E]; 15 Sep. 2007; M. J. Ramírez leg.; MACN- Ar 42348 • 1 ♂; same collection data as for preceding; MACN-Ar 42347 • 1 ♂; same collection data as for preceding; MACN-Ar 42346 • 1 ♀; Barcelona; [41.38504 ° N, 2.17347 ° E]; 13 Sep. 1986; J. A. Coddington leg.; caught in the middle of an imaginal molt, has two sets of spermathecae; USNM 1656 • 1 ♀; same collection data as for preceding; USNM 1656 • 2 ♂♂; Castellón, N of L’Alcora, stone walls; 40.08974 ° N, 0.19945 ° W; 300 m. a. s. l.; 30 May 2010; S. Huber and A. Schönhofer leg.; at night; SMF • 1 ♀; Cataluña, Barcelona, Barcelona; 41.40423 ° N, 21.54461 ° E; Jan. 1988; M. J. Ramírez leg.; MACN- Ar 20562 • 1 ♀, 1 imm.; Madrid; [40.41678 ° N, 3.70379 ° W]; 1961; K. W. Haller leg.; AMNH • 3 ♀♀, 1 imm.; Madrid, Navas del Rey; [40.38776 ° N, 4.25 ° W]; 1961; K. W. Haller leg.; AMNH • 9 ♀♀, 11 imm.; Madrid, Vaciamadrid; [40.35191 ° N, 3.53573 ° W]; 1961; K. W. Haller leg.; AMNH • 22 ♀♀, 6 imm.; Madrid, Vaciamadrid; 1961; K. W. Haller leg.; AMNH • 1 ♀; Murcia, Fortuna, Espacio Natural de Ajanque; [38.14949 ° N, 1.10128 ° W]; CRBA 861 • 1 ♂; same locality as for preceding; 19 Jun. 2004; CRBA 860 • 4 ♀♀, 1 imm.; Tarragona, Monte Poulet; 41.11903 ° N, 1.24451 ° E; 23 Dec. 1986; M. J. Ramírez leg.; MACN-Ar 20559 • 1 ♂; Teruel, Molinos; 40.81945 ° N, 0.45083 ° W; Apr. 1985; J. Moles leg.; MACN-Ar 20565 • 1 ♀, 1 imm.; no further data, on a slope; [39.66422 ° N, 0.22674 ° W]; 17 Jun. 1993; S. Lideese leg.; ZMB 30578. TUNISIA • 1 ♀; Djerba; [33.8076 ° N, 10.84515 ° E]; V. Bilgner leg.; ZMB K. 786. VENEZUELA • 1 ♀; Miranda, Caracas suburbs, caves near Cementerio del Este; [10.45038 ° N, 66.81118 ° W]; Jul. 1986; J. A. Coddington leg.; USNM 3222. NO DATA • 1 ♂, 2 ♀♀, 2 imm.; Mediterranean region; ZMB • 1 ♀; ZMB 533 • 1 ♀; ZMB 580 • 1 ♀; MNRJ 1434.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF96FFD90870FACA09C1EFBF.taxon	distribution	Distribution Widely distributed in the Mediterranean region and the Middle East (Fig. 4 A), with records (probably introduced) in the Azores, Socotra Island, Cabo Verde, Angola and Venezuela. In Israel, it is distributed in Mediterranean areas in the north (Fig. 4 B); in some localities only females are known, and thus it is possible these records could belong to F. betarif sp. nov., although it is more likely these records are of F. insidiatrix since they broadly overlap with records of males.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	description	urn: lsid: zoobank. org: act: 86 B 4 AAD 0 - D 69 E- 4 C 79 - BF 3 F-DE 5 E 7 DC 0 EDB 1 Figs 1, 3 E – F, H, 5 A, C, E, 7 – 9	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	diagnosis	Diagnosis Males are similar to those of F. insidiatrix and F. albens by having a pedipalp with elongate femur and tibia, and by a subconical copulatory bulb with terminally hooked embolus (Fig. 8 A – B; see also Zonstein & Marusik 2019: fig. 27). They differ from both species by the conspicuous embolic keel (Fig. 8 F (arrow), 5 A, k) (inconspicuous keel in F. insidiatrix and F. albens; Figs 5 B, 10 O). Females have a pair of large spermathecae that appear undivided (Fig. 9); they fall within the morphological variation observed for F. insidiatrix (Fig. 3 C – D; Zonstein & Marusik 2019: fig. 17 d – h) and thus females are not diagnosable through morphology only; they can be diagnosed using DNA barcoding of the COI gene.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	etymology	Etymology The specific epithet is a noun in apposition and refers to Bet ‘ Arif, the name of the ‘ wadi ’ (dry riverbed) in which the two caves where the species was collected (Tinshemet and Oah) are located.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	materials_examined	Type material examined Holotype ISRAEL • 1 ♂; Judean hills, near Shoham, Hadom Shomeron Nature Reserve; Bet ‘ Arif wadi, Tinshemet cave, inside cave; 31.99938 ° N, 34.96813 ° E; 122 m. a. s. l.; 20 Feb. 2020; I. L. F. Magalhaes, E. Gavish- Regev and S. Aharon leg.; HUJ-INVAr 21058. Paratypes ISRAEL • 1 ♀; Judean hills, near Shoham, Hadom Shomeron Nature Reserve; Bet ‘ Arif wadi, Oah cave, inside cave; 32.0053 ° N, 34.9722 ° E; 25 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21055 • 2 ♀♀; same locality as for preceding; 9 Mar. 2014; S. Aharon and E. Gavish-Regev leg.; HUJ- INVAr 21059 • 1 ♂, 6 ♀♀; Judean hills, near Shoham, Hadom Shomeron Nature Reserve, Bet ‘ Arif wadi, Tinshemet cave, inside cave; 31.99938 ° N, 34.96813 ° E, 122 m. a. s. l.; 20 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev and S. Aharon leg.; MACN-Ar 41222 • 2 ♂♂, 3 ♀♀, 2 imm.; same collection data as for preceding; HUJ-INVAr 20275 • 4 ♀♀, 1 imm.; same collection data as for preceding; MACN-Ar 41791 • 1 ♂; same collection data as for preceding; MACN-Ar 41793 • 2 imm.; same locality as for preceding, Mediterranean scrubland, outside the cave, under stones; 20 Feb. 2020; I. L. F. Magalhaes, E. Gavish-Regev and S. Aharon leg.; MACN-Ar 41829 • 1 ♀; same locality as for preceding; Mediterranean scrubland, outside the cave; 31.9994 ° N, 34.9681 ° E; 25 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21056 • 7 ♀♀; same locality as for preceding; inside cave; 31.99938 ° N, 34.96813 ° E; 25 May 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 20301 • 1 ♀; same collection data as for preceding; HUJ-INVAr 21057 • 3 imm; same locality as for preceding; 9 Mar. 2014; S. Aharon and E. Gavish-Regev leg.; HUJ-INVAr 21060. Additional material examined None.	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	description	Description Male (holotype, HUJ-INVAr 21058) Coloration yellowish cream except where noted. Carapace lined with black pigment, with marbled brown clypeal markings and brown, V-shaped median pattern. Legs I – III with three brown ventral spots. Abdomen brownish gray. Anterior margin of the carapace unmodified, eye apodemes absent. Sternum subrounded, sigilla not visible. Total length 5.25. Carapace length 2.38, width 1.88. Clypeus length 0.28. Eye diameters and interdistances: AME 0.14, PME 0.15, ALE 0.20, PLE 0.18, AME – AME 0.06, PME – PME 0.14. Sternum length 1.26, width 1.05. Pedipalp: femur length 2.34, height 0.30, tibia length 2.10, height 0.34. Leg I: 14.88 (3.82, 1.09, 3.94, 3.91, 2.12). II: 11.29 (3.05, 0.94, 2.78, 3.17, 1.35). III: 10.23 (2.70, 0.84, 2.52, 2.91, 1.26). IV: 12.52 (3.57, 0.90, 3.28, 3.44, 1.33). Abdomen: length 2.97, width 1.72. Pedipalp macrosetae on ventral surface of femur. Leg macrosetae: fe I d 1 - 0 - 0, dp 0 - 0 - 2, ti I v 0 - 2 - 2 - 2, mt I v 2 - 2 - 2 - 4, fe II d 1 - 1 - 0, dp 0 - 0 - 2, ti II v 0 - 2 - 2, mt II v 2 - 2 - 4, fe III d 1 - 0 - 0, dp 0 - 0 - 1, dr 0 - 0 - 2, ti III d 1 - 2 - 2, v 1 - 1, mt III dr 0 - 0 - 1, v 1 - 2 - 2 - 4, fe IV d 1 - 1 - 0, dr 0 - 0 - 2, ti IV v 1 - 1, mt IV v 1 - 1 - 2 - 4, all tarsi with ventral macrosetae. Pedipalp as in diagnosis (Figs 5 A, C, E, 8 A – G). State of the specimen: good, right leg I removed for molecular analysis, left pedipalp dissected. Female (paratype, MACN-Ar 41222) Coloration as in male. Eye apodemes absent. Sternum subrounded, with one pair of posterior sigilla. Total length 7.76. Carapace length 3.17, width 2.51. Clypeus length 0.40. Eye diameters and interdistances: AME 0.14, PME 0.19, ALE 0.27, PLE 0.23, AME – AME 0.04, PME – PME 0.23. Sternum length 1.64, width 1.50. Pedipalp: femur length 1.70, height 0.67, tibia length 1.06, height 0.65. Leg I: 13.60 (3.66, 1.37, 3.49, 3.23, 1.85). II: 9.66 (2.84, 1.13, 2.26, 2.16, 1.27). III: 7.82 (2.34, 1.03, 1.66, 1.64, 1.15). IV: 10.54 (3.30, 1.25, 2.64, 2.27, 1.08). Abdomen: length 4.67, width 3.08. Palpal macrosetae on ventral surface of tibia and tarsus. Leg macrosetae: fe I d 1 - 0 - 0, ti I v 0 - 2 - 0, mt I v 2 - 2 - 2 - 4, fe II d 1 - 0 - 0, ti II v 0 - 2 - 0, mt II v 2 - 2 - 2 - 4, fe III d 1 - 0 - 0, ti III d 0 - 2 - 0, v 0 - 1 - 2, mt III dr 1 - 1 - 1, v 1 - 2 - 2 - 4, fe IV d 1 - 0 - 0, ti IV v 1 - 1, mt IV v 1 - 1 - 1 - 4, all tarsi with ventral macrosetae. Calamistrum with three staggered rows with <6 setae; calamistrum gap present. Epigastric furrow unmodified. Endogyne of females from the same locality as in the diagnosis (Fig. 9). State of the specimen: good. Variation Males (N = 3): total length 4.89 – 5.67 (5.27), carapace length 2.25 – 2.78 (2.47), femur I length 3.62 – 4.04 (3.83), femur / carapace ratio 1.45 – 1.61 (1.56). Females (N = 5): total length 6.47 – 8.95 (7.79), carapace length 3.17 – 3.98 (3.57), femur I length 3.5 – 4.18 (3.86), femur / carapace ratio 1.01 – 1.15 (1.08). The shape of the spermathecae varies only slightly among the dissected females. Natural history Filistata betarif sp. nov. is a troglophilic species, inhabiting both epigean and hypogean (caves) habitats. It is known from the mesic Mediterranean region of Israel, in scrubland habitats, and usually hides in a refugium made of cribellate silk in crevices, holes or under stones and on rocky walls (Fig. 1 A – B). All specimens were collected from Bet ‘ Arif wadi near Shoham, Hadom Shomeron Nature Reserve, central Israel. The species was found in low abundances outside of caves (Fig. 1 D) and in very high abundances, year-around, in Tinshemet cave that is characterized by a high guano amount of the Egyptian fruit bat [Rousettus aegyptiacus (Geoffroy, 1810)]. Tinshemet (Arabic name: Mugharet al-Watwat) is a small to medium-sized Paleolithic cave, formed in a limestone rock of the Turonian Bina formation. It is located on the east bank of Bet ‘ Arif wadi (Arabic name: Wadi Adasiyeh), 95 meters a. s. l. and 15 meters above the wadi (Frumkin et al. 2016). It has a main opening on the side of the cliff, and an additional chimney opening, which the frugivorous bats frequently use (personal observation). Filistata betarif sp. nov. was also found in lower abundances in the nearby Oah cave that houses no bats (Cuff et al. 2021; Gavish- Regev et al. 2021). Oah is a small cave, situated on the same east bank of Bet ‘ Arif wadi, north to Tinshemet cave, and is formed in a similar rock and age (Boaz Langford, pers. comm.). In both caves, Filistata betarif sp. nov. was found mainly at the entrance zones, on the cave walls (Fig. 1 A – B). Other caves surveyed by us, at the mesic Mediterranean, semi-arid, and arid regions of Israel and Palestine, were not found to be occupied by F. betarif sp. nov., but instead by its sibling troglophilic species F. insidiatrix [Berniki, Yir’on, Yonim (Galilee); Ezba’, Oren (Karmel); Sal’it (northern Samaria); Andartat HaBiqa’ (central Jordan Valley); Perat (northern Judean desert); Qumeran (northern Dead-Sea area)] (Cuff et al. 2021; Gavish-Regev et al. 2021).	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
54519917FF9CFFC708A9FBBB08A5E88A.taxon	distribution	Distribution Known only from two caves and under stones in their surroundings in Bet ‘ Arif wadi, Hadom Shomeron Nature Reserve, central Israel (Fig. 4).	en	Magalhaes, Ivan L. F., Aharon, Shlomi, Ganem, Zeana, Gavish-Regev, Efrat (2022): A new semi-cryptic Filistata from caves in the Levant with comments on the limits of Filistata insidiatrix (Forsskål, 1775) (Arachnida: Araneae: Filistatidae). European Journal of Taxonomy 831: 149-174, DOI: https://doi.org/10.5852/ejt.2022.831.1875, URL: http://dx.doi.org/10.5852/ejt.2022.831.1875
