taxonID	type	description	language	source
039D87D5802E0E4E83E5FE1202B66F26.taxon	type_taxon	Type species. Spathiomorpha varinervis Tobias, 1976, by original designation. Re-description (Figs 1, 2). Head transverse. Ocelli arranged in obtuse triangle with base larger than ts sides. Frons not or only weakly concave. Eyes glabrous. Occipital carina complete, lower often fused with hypostomal carina upper base of mandible. Malar suture absent. Clypeal suture rather distinct and almost complete. Clypeus with distinct protruding flange along its lower margin. Hypoclypeal depression round. Postgenal bridge present but rather narrow. Palps usually long; maxillary palps 6 - segmented, labial palps 4 - segmented. Scape of antenna rather thick and short, without apical lobe and basal constriction. Flagellum weakly thickened, first flagellomere more or less longer than second. Mesosoma. Neck of prothorax short; pronotal keel usually distinct and situated near middle. Mesonotum usually smooth or sparsely punctate, often highly and almost vertically elevated above pronotum. Median lobe of mesoscutum without anterolateral corners and usually without complete medial longitudinal furrow. Notauli complete and sculptured. Prescutellar depression relatively long, sculptured or almost smooth, always with median carina. Metanotum with short, subpointed or obtuse median tooth (lateral view). Precoxal sulcus distinct, medium length, almost straight, oblique, mostly crenulate. Prepectal carina distinct and complete. Propodeum with basolateral areas and areola deliniated by distinct carinae; lateral tubercles and propodeal bridge absent or sometimes present short and thick tubercles. Propodeal spiracles small and round. Rarely mesosoma partly reduced (micropterous species). Wings. Pterostigma of fore wing wide. Radial vein (r) arising from or behind middle of pterostigma. Radial (marginal) cell not shortened. Both radiomedial veins (2 - SR, r-m) present. Recurrent vein (m-cu) always antefurcal. Nervulus (cu-a) postfurcal. Discoidal (discal) cell petiolate, petiole (1 - SR) short. Parallel vein (CU 1 a) not interstitial, arising from posterior third of distal margin of brachial (subdiscal) cell. Brachial (subdiscal) cell wide. Transverse anal veins (2 A, a) absent. In hind wing, radial (marginal) cell almost parallel-sided, without additional transverse vein (r). Medial (basal) cell wide. Submedial (subbasal) cell long; first abscissa of mediocubital vein (M + CU) about as long as second abscissa (1 - M). Recurrent vein (m-cu) present and sclerotised. Legs. Fore and middle tibiae with numerous and thick spines arranged in almost single vertical line. Hind coxa with distinct basoventral corner and tubercle. Hind femur relatively narrow. Hind basitarsus 0.7 – 0.9 × as long as second – fifth segments combined. Metasoma. First tergite usually relatively long and narrow, petiolate; dorsope small; spiracular tubercles distinct and placed usually in basal third of tergite. Acrosternite of first segment 0.5 – 0.6 × as long as first tergite, its apical margin situated on or usually behind spiracles. Second suture usually absent or rarely fine; second and third tergites without furrows and areas. Only second tergite with separate laterotergites, usually completely smooth or sometimes shortly striate basally. Ovipositor shorter or longer than metasoma.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802E0E4E83E5FE1202B66F26.taxon	diagnosis	Diagnosis. This genus is morphologically and molecularly similar to the widely distributed genus Ontsira Cameron, 1900 from the tribe Doryctini (Zaldivar-Riverón et al., 2008; Belokobylskij and Austin, 2013). It can be distinguished from the latter by having a petiolate first metasomal segment with distinctly elongated acrosternite (longer than half of tergite). Also Spathiomorpha is similar by petiolate first segment with the genus Spathius Nees, 1819 from the tribe Spathiini, but distinctly differs from later in fore wing by the antefurcal position of recurrent vein (m-cu) to the first radiomedial vein (2 - SR) (always postfurcal in Spathius), the arising of parallel vein (CU 1 a) from the posterior fourth of the vein (3 - CU 1) closed brachial (subdiscal) cell distally (from or before middle of this vein in Spathius), and in hind wing the long submedial (subbasal) cell with first abscissa of mediocubital vein (M + CU) longer than second abscissa (1 - M) (cell and abscissa short in Spathius). Composition. Spathiomorpha enderleini Belokobylskij, 1996 (China: Taiwan); S. japonica Belokobylskij et Maeto, 2006 (Japan); S. jenningsi Belokobylskij et Austin, 2013 (Australia: New South Wales); S. longipalpis Belokobylskij, 1985 (Russia: Far East, Korea), S. mexicana Belokobylskij et Zaldívar-Riverón, sp. nov. (Mexico), S. tasmanica Belokobylskij, Iqbal et Austin, 2004 (Australia: Tasmania), S. varinervis Tobias, 1976 (Azerbaijan, Georgia, Macedonia, Russia: North Caucasus, Serbia, Turkey) (Tobias, 1976; Belokobylskij, 1985, Papp, 1987; Belokobylskij, 1996; Belokobylskij et al., 2004; Belokobylskij and Maeto, 2006; Yu et al., 2016).	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802E0E4E83E5FE1202B66F26.taxon	biology_ecology	Hosts. Unknown.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802E0E4E83E5FE1202B66F26.taxon	distribution	Distribution. Australasian, Oriental, Neotropical (first record) and Palaearctic regions.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	description	(Figs 1, 2)	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	materials_examined	Type material. Holotype: female, “ Mexico: Chiapas, 10 km ESE of San Cristobal, 2400 m, pine forest, 16.668022, - 92.550506, 6 – 7. viii. 2021, A. I. Khalaim ” (IB-UNAM).	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	description	Description. Female. Body length 6.5 mm; fore wing length 5.2 mm. Head width (dorsal view) 1.4 × its median length, 1.2 × width of mesoscutum. Head behind eyes (dorsal view) weakly convex in anterior half, evenly roundly narrowed in posterior half. Transverse diameter of eye 1.1 × longer than temple. Frons without carina, with distinct and rather wide complete median furrow. Ocelli mediumsized, arranged in triangle with base 1.15 × its sides. POL 1.3 × Od, 0.4 × OOL. Eye glabrous, practically without emargination opposite antennal socket, 1.3 × as high as broad. Malar space 0.4 × height of eye, 0.9 × basal width of mandible. Face along eyes without carinae, with distinct shallow elongate oblique depressions above clypeus; width of face 1.3 × height of eye and 1.4 × height of face and clypeus combined. Diameter of antennal socket 0.7 × distance between sockets and 1.4 × distance between socket and eye. Suture connected margin of antennal socket with inner margin of eye present and distinct. Clypeal suture shallow and complete. Hypoclypeal depression round, its width 1.1 × distance from edge of depression to eye, about 0.5 × width of face. Occipital carina coarse, below joined with hypostomal carina upper base of mandible. Palpi long, length of maxillary palpus almost equal to head height. Antenna relatively thick, weakly setiform, with 41 antennomeres, 1.3 × longer than body. Scape 1.8 × longer than its maximum width. First flagellomere 4.3 × longer than its apical width, 1.1 × longer than second. Penultimate flagellomere 2.5 × longer than wide, 0.5 × as long as first flagellomere, 0.8 × as long as apical flagellomere; the latter pointed apically and without spine. Mesosoma. Length 1.8 × its height. Pronotum (lateral view) convex dorsally, with median pronotal carina. Median lobe of mesoscutum (dorsal view) distinctly protruding forwards, with fine to very fine median furrow. Notauli deep, wide and coarsely crenulate. Prescutellar depression rather deep, with distinct median and two fine lateral carinae, almost smooth between carinae, 0.4 × as long as scutellum. Scutellum convex and without lateral carinae. Metanotal tooth short, but distinct and subpointed. Subalar depression relatively deep, more or less wide, coarsely rugose-striate. Precoxal sulcus wide, almost straight, oblique, coarsely and sparsely crenulate, running along anterior half of lower part of mesopleuron. Propodeum with distinct, thick lateral tubercles. Wings. Fore wing 3.4 × longer than its maximum width. Radial vein (r) arising behind middle of pterostigma, from its basal 0.7. Radial (marginal) cell not shortened; metacarp (1 - R 1) almost 1.5 × longer than pterostigma. First radial abscissa (r) 0.8 × as long as maximum width of pterostigma. Second radial abscissa (3 - SR) 3.6 × longer than first abscissa (r), 0.65 × as long as straight third abscissa (SR 1), 1.3 × longer than first radiomedial vein (2 - SR). Second radiomedial (submarginal) cell 2.7 × longer than its maximum width, 1.3 × longer than brachial (subdiscal) cell. First medial abscissa (1 - SR + M) distinctly sinuate. Mediocubital vein (M + CU 1) almost straight. Recurrent vein (m-cu) weakly curved, almost equal to first radiomedial vein (2 - SR). Distance from nervulus (cu-a) to basal vein (1 - M) 0.6 × nervulus (cu-a) length. Parallel vein (CU 1 a) arising from posterior 0.2 of distal margin (3 - CU 1) of brachial (subdiscal) cell. Hind wing 5.2 × longer than maximum width. First costal abscissa (C + SC + R) 0.7 × as long as second abscissa (1 - SC + R). First abscissa of mediocubital vein (M + CU) 0.8 × as long as second abscissa (1 - M). Mediocubital (basal) cell large, distinctly widened toward apex, 5.7 × longer than wide, 0.4 × as long as wing. Recurrent vein (m-cu) weakly curved, subinterstitial, distinctly pigmented. Legs. Fore tibia with numerous slender spines arranged in narrow line. Hind femur rather narrow, 4.5 × longer than wide. Hind tarsus as long as hind tibia. Hind basitarsus 0.9 × as long as second – fifth tarsal segments combined. Second segment of hind tarsus 0.4 × as long as basitarsus, 1.5 × longer than fifth segment (without pretarsus). Metasoma 1.1 × longer than head and mesosoma combined. First tergite rather thick, with small dorsope, with distinct spiracular tubercles in basal 0.3, distinctly and weakly curvedly widened from base to apex. Maximum width of first tergite 1.6 × its width at level of spiracles, 2.2 × its minimum width; length of tergite 1.7 × its apical width, 1.8 × length of propodeum. Length of second and third tergites combined 1.2 × basal width of second tergite, 0.9 × their maximum width. Ovipositor sheath 1.1 × longer than body, 2.3 × longer than metasoma, 3.3 × longer than mesosoma, 1.4 × longer than fore wing. Sculpture and pubescence. Vertex and temple smooth; frons smooth posteriorly, rugose anteriorly; face finely striate with rugulosity, smooth in lower third. Sides of pronotum entirely coarsely rugose-striate, with coarsely crenulate wide and oblique median depression. Mesoscutum rather densely and finely punctate, coarsely striate in narrow medioposterior area in posterior half. Scutellum almost smooth. Mesopleuron almost entirely smooth. Propodeum with areas distinctly delineated by high carinae; basolateral areas large, almost entirely finely rugulosereticulate; areola short and rather wide, 1.2 × longer than wide; basal carina in basal half of propodeum long, 2.2 × longer than anterior fork of areola; posterior part of propodeum coarsely sparsely and rather finely rugose. Hind coxae finely rugulose dorsally, smooth on most part. Hind femur smooth, partly with fine and sparse punctation. First tergite entirely densely and coarsely striate with rugulosity between striae, medially coarsely transversely striate with rugulosity in basal 0.3. Second tergite mostly smooth, with short basal striation. Remaining tergites smooth. Vertex with rather dense, relatively long and semi-erect yellow setae, medially partly almost glabrous. Mesoscutum entirely with dense, rather short and semi-erect yellow setae. Hind tibia dorsally with rather short, dense and almost erect setae, length of these setae 0.4 – 0.6 × maximum width of hind tibia. Colour. Head mainly yellow, face mainly, malar space and temple brown to dark brown. Mesosoma mainly black, mesoscutum and scutellum brownish yellow to reddish brown. Metasoma reddish brown to dark reddish brown, first tergite black. Antennae mainly dark brown to black, segments in basal fifth light brown to reddish brown. Palpi pale yellow. Legs mainly yellow, all femora medially or dorso-medially distinctly infuscate; tibiae and tarsi brownish, hind tibia basally black at short part. Ovipositor sheath brown to dark brown, apically black. Fore wing distinctly infuscate. Pterostigma dark brown to black, pale brown shortly basally. Male. Unknown.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	etymology	Etymology. This species is named after the country from where this new species was collected.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	distribution	Distribution. Mexico (Chiapas).	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
039D87D5802D0E4983E5FD8705DA68FE.taxon	diagnosis	Diagnosis. This new species is similar to S. longipalpis Belokobylskij and S. japonica Belokobylskij et Maeto. The differences between these species are shown in the below key to species.	en	Belokobylskij, Sergey A., Zaldívar-Riverón, Alejandro (2022): First record of the genus Spathiomorpha Tobias, 1976 (Hymenoptera: Braconidae Doryctinae) in the New World, with the description of a new species. Zootaxa 5162 (4): 439-445, DOI: https://doi.org/10.11646/zootaxa.5162.4.8
