taxonID	type	description	language	source
182B9F50FFECFFDDFD22FB66FDDAFB7B.taxon	description	(Figs. 1, 5, 7, 9, 11)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFECFFDDFD22FB66FDDAFB7B.taxon	type_taxon	Type species: Thinobatis ferruginea Eschscholtz, 1831, by monotypy.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFECFFDDFD22FB66FDDAFB7B.taxon	description	Description. In addition to the characters stated above: Mandibles thick, base 2 X wider than apex, both mandibles with dorsal cusp on basal half, shorter on right than on left mandible (Fig. 5), lacking ventral fossa; labrum exposed, entirely sclerotized; clypeus short, not covering labrum, anterior margin straight or concave, nearly at same level with antennal insertions, lacking row of teeth (Fig. 5); prementum partially sclerotized, base concealed beneath mentum; anterior margin of mentum narrowly notched (Fig. 7); eyes small, not protruding, upper margin lacking carina; length of epicanthus 1.5 X length of eye (Fig. 5); antennae reaching posterior margin of pronotum, antennomere 1 as long as 3, emerging from distal half of antennal insertion (Fig. 7), antennomere 3 longer than 2 (Fig. 5), antennomeres 9 and 10 triangular, strongly expanded outwards, wider than long, antennomere 11 equal to or longer than 10 (Fig. 9). Pronotum with anterior angles acute, directed forwards, or rounded sloping towards venter (Fig. 1). Metasternum short, as long as mesosternum. Metacoxae separated by less than half metacoxal width; metacoxal width 3 X metacoxal length; distance between meso – and metacoxae equal to metacoxal length (Fig. 11). First tarsomere of hind legs longer than 2 + 3 combined.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFECFFDDFD22FB66FDDAFB7B.taxon	discussion	Species Included. T h i n ob a t i s f e r r u g i ne a Eschscholtz, Thinobatis rufipes rufipes Solier, Thinobatis rufipes penai Freude, Thinobatis rotundicollis Waterhouse, Thinobatis intermedia Philippi and Philippi, Thinobatis kuscheli Kulzer, Thinobatis brevicollis Kulzer, Thinobatis calderana Kulzer, Thinobatis melcheri Freude, Thinobatis simplex Peña, Thinobatis punctata Peña, Thinobatis confusa Peña, and Thinobatis arenaria Peña (Peña 1974 b).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFECFFDDFD22FB66FDDAFB7B.taxon	materials_examined	Material Examined. Thinobatis intermedia, T. melcheri, T. rufipes rufipes, and T. rufipes penai (IADIZA). We also examined T. rotundicollis, but it should be classified in a distinct genus because of the shape and proportional length and width of the pronotum, rounded pronotal posterior angles, lack of a dorsal cusp on both mandibles, and clypeus and frons with protuberances.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFF2AFB4CFB58FB19.taxon	description	(Figs. 2 – 4, 6, 8, 10, 12, 21)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFF2AFB4CFB58FB19.taxon	type_taxon	Type species: Vaniosus paradoxus Kulzer, 1956, by monotypy.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFF2AFB4CFB58FB19.taxon	description	Description. In addition to the characters stated above: Mandibles finer, acute, base 1.5 X wider than the apex, both mandibles with a dorsal cusp on basal half longer on the right than on the left mandible (Fig. 6), both mandibles with a ventral fossa (Fig. 8); labrum exposed, partially sclerotized; clypeus short, not covering the labrum, anterior margin triangular nearly at same level with antennal insertions, armed with a row of teeth directed downwards, in the apex with a large tooth prolonged anterad (Fig. 6); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave (Fig. 8); eyes small, protruding out, upper margin lacking carina; length of epicanthus 2.5 x the length of eye (Fig. 6); antennae surpassing posterior margin of pronotum, antennomere 1 sticks out distal half of antennal insertion (Fig. 8), antennomeres 9 and 10 cylindrical, not expanded outwards, longer than wide, antennomere 11 equal in length to 10 (Fig. 10). Pronotum with anterior angles obtuse, slopeing towards venter (Figs. 2 – 4). Metasternum twice the length of mesosternum. Metacoxae separated by less than half metacoxal width; metacoxal width 3 X metacoxal length; distance between meso – and metacoxae twice the metacoxal length (Fig. 12). Female genitalia (Fig. 21). Spiculum without arms. Paraprocts long (2.0 <P / C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, with annulate duct. Spermatheca single, compact, locular, at the end of vagina, near the duct of spermathecal accessory gland.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFF2AFB4CFB58FB19.taxon	discussion	Species included. Vaniosus paradoxus Kulzer and Vaniosus profana (Kulzer). KEY TO THE SPECIES OF VANIOSUS KULZER	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFD24FAB1FCA5F9C8.taxon	description	(Figs. 3 – 4)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFD24FAB1FCA5F9C8.taxon	diagnosis	Diagnosis. See Kulzer (1956).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFD24FAB1FCA5F9C8.taxon	materials_examined	Type Material. Holotype: Cerro Carambailloj, bei Cajamaros, Peru, 3000 m, II- 1942, leg. W. Weyrauch (NHMB) (Figs. 3 – 4).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDDFD24FAB1FCA5F9C8.taxon	distribution	Distribution. Known only from the type locality in Cajamarca, Peru.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	description	(Figs. 2, 6, 8, 10, 12, 21)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	diagnosis	Diagnosis. See Kulzer (1956).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	materials_examined	Type Material. Holotype: Santiago del Estero, Forres, XII- 1934 (NHMB) (Fig. 2). Paratype: Catamarca, Caspinchango, 12. III. 21 (NHMB). These and all localities mentioned in the original description correspond to Argentina and not to Peru, as stated by Kulzer (1956). Other Material Examined. 237 specimens in IADIZA. ARGENTINA: Salta: Dto. Chicoana: 2 km E Escoipe, 1856 m, 7 - XI- 2004, G. Flores - G. Zalazar, 70; Dto. Cafayate: Tolombón, 1812 m, 8 - XII- 2003, G. Flores, 4. Tucumán: Dto. Tafí del Valle: Ruinas de Quilmes, 17 - X- 1997, S. Roig, 16. Catamarca: 2 km N Villa Vil, 2358 m, 16 - XI- 2009, G. Flores, 1; Punta de Balasto, 3 - 5 - II- 2001, G. Arriagada, 1. La Rioja: Dto. Famatina: 2 km S Campanas, 10 - XII- 2003, G. Flores, 4, 10 km NE Pituil, 15 - X- 1997, S. Roig, 11; Dto. Gral. Belgrano: 20 km N Olta, 21 - X- 1997, S. Roig, 3. Córdoba: 15 km N Capilla del Monte, 11 - XII- 2000, C. Domínguez - S. Roig, 2. San Juan: Dto. Valle Fértil: Astica, 13 - V- 1979, S. Roig, 1, 10 km E Marayes, 23 - X- 1997, S. Roig, 1. Mendoza: Dto. La Paz: Desaguadero, 16 - XI- 1997, S. Roig, 2, 20 km E La Paz, 16 - XI- 1997, S. Roig, 1; Dto. Santa Rosa: Reserva Ñacuñán, 560 m, pitfall: 13 - IX- 1997, S. Lagos, 1, 20 - X- 1997 to 22 - XI- 1997, S. Lagos, 1, 7 - II- 1998, S. Lagos, 1, 15 - V- 1998, S. Claver, 1, 31 - V- 1998, S. Lagos, 1, 12 - XII- 1998, S. Lagos, 4, VI- 1999, S. Lagos, 2, 9 - V- 1999 to 11 - VI- 1999, S. Lagos, 4, 24 - III- 2010 to 25 - IV- 2010, G. Arriagada, 1, 22 - III- 2011, F. Ocampo, 1, Campo Quemado, 7 - II- 1998, S. Lagos, 1; Dto. Lavalle, Reserva Telteca, 548 m, pitfall: 9 - VIII- 1994 to 6 - IX- 1994, G. Flores, 1, 15 - XII- 1994 to 3 - II- 1995, G. Flores, 2, 15 - II- 1996 to 25 - III- 1996, G. Flores, 1, 15 - IV- 2008, L. Muñoz, 2, Campo Norte, 17 - XI- 1999, S. Roig, 2; Dto. Capital: CCT CONICET, 839 m, 91 specimens collected on pig carcasses (Sus scrofa L.): 15 - V- 2007, F. Aballay, 1, 17 - V- 2007, F. Aballay, 1, 18 - V- 2007, F. Aballay, 2, 19 - V- 2007, F. Aballay, 1, 24 - VII- 2007, F. Aballay, 1, 8 - IV- 2008, F. Aballay, 1, 12 - IV- 2008, F. Aballay, 1, 18 - IV- 2008, F. Aballay, 1, 21 - IV- 2008, F. Aballay, 1, 24 - IV- 2008, F. Aballay, 5, 28 - IV- 2008, F. Aballay, 12, 30 - IV- 2008, F. Aballay, 5, 8 - V- 2008, F. Aballay, 5, 15 - V- 2008, F. Aballay, 6, 30 - V- 2008, F. Aballay, 4, 30 - X- 2008, F. Aballay, 9, 31 - X- 2008, F. Aballay, 9, 1 - XI- 2008, F. Aballay, 3, 2 - XI- 2008, F. Aballay, 1, 3 - XI- 2008, F. Aballay, 1, 4 - XI- 2008, F. Aballay, 4, 7 - XI- 2008, F. Aballay, 2, 10 - XI- 2008, F. Aballay, 1, 11 - XI- 2008, F. Aballay, 4, 14 - XI- 2008, F. Aballay, 2, 15 - XI- 2008, F. Aballay, 3, 16 - XI- 2008, F. Aballay, 2, 17 - XI- 2008, F. Aballay, 1, 17 - XII- 2008, F. Aballay, 2.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	distribution	Distribution. Argentina: Salta, Tucumán, Catamarca, La Rioja, Santiago del Estero, Córdoba, San Juan, and Mendoza provinces.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	biology_ecology	Habitat. This species occurs in xerophilous warm steppes and dry forests east of the Andes, in the biogeographic provinces of Monte and Chaco (Morrone 2014) at altitudes of 150 to 2,400 m. Specimens were collected throughout the year, even in the coldest months.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEDFFDEFD71F9E2FC6CFBE5.taxon	biology_ecology	Biology. Specimens collected in Mendoza at the Science and Technology Center (91 individuals) were found feeding on decomposing pig carcasses, mainly on the carcass in direct sunlight (88 individuals) and very few on the carcass in the shade (3 individuals). These specimens were recorded in greatest abundance in autumn (46 individuals) and spring (44 individuals), with only one record in the winter and none in the summer. This species was associated with the Advanced Decay and Dry Remains stages of decomposition defined by Payne (1965). Because of these three behavioral characteristics, 1) preference for carcasses under direct sun, 2) seasonality (autumn and spring), and 3) association with specific stages of decomposition (Advanced Decay and Dry Remains), this species should be considered for potential forensic importance for estimating time of death (PMI = Post Mortem Interval) in bodies found in arid environments. Specimens that were collected using pitfall traps partly filled with water and propylene glycol in two reserves of Mendoza province, Telteca and Ñacuñán, could have been attracted by the smell of decomposition of propylene glycol, similar to the smell of a carcass.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEEFFDFFCCEFBEBFDB5FAFA.taxon	description	(Figs. 13 – 20, 22)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEEFFDFFCCEFBEBFDB5FAFA.taxon	type_taxon	Type species: Achanius anthicoides Erichson, 1847, by monotypy.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEEFFDFFCCEFBEBFDB5FAFA.taxon	description	Description. In addition to the characters stated above: Mandibles finer, acute, base 1.5 X wider than apex, with dorsal cusp only on right mandible on basal half, left mandible with sharp, raised dorsal edge (Fig. 17), both mandibles with ventral fossa (Fig. 18); labrum not exposed, entirely membranous; clypeus oval, produced anteriorad of antennal insertions, covering labrum, anterior margin triangular or concave and armed with row of teeth directed downwards, apically with a large tooth prolonged anteriorad, genoclypeal suture wellmarked (Fig. 17), epicanthus separated from clypeus (Fig. 19); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave (Fig. 18); eyes not protruding, upper margin with slightly or wellmarked carina always present, starting posterior to epicanthus and ending in posterior margin of eye; length of epicanthus equal to length of eye, head widest at eyes, postgenal margin inconspicuous (Fig. 19); antennae surpassing posterior margin of pronotum, all antennomeres longer than wide, antennomere 1 longer than 3, emerging from distal ¾ of antennal insertion (Fig. 19), antennomere 2 equal to or longer than 3 or shorter than 3 but almost equal (proportion 2 / 3 = 17 / 19) (Fig. 19); antennomeres 9 and 10 triangular, expanded outwards, longer than wide (Figs. 13, 14). Pronotum with anterior angles rounded, sloping towards venter. Metacoxae transverse, separated by less than third of metacoxal width (Fig. 20). First tarsomere of hind legs longer than 2 + 3 combined (Figs. 14, 16). Female genitalia as in Fig. 22. Spiculum without arms. Paraprocts long (2.0 <P / C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, duct not annulate. Spermatheca single, compact, locular, emptying at end of vagina, spermathecal accessory gland emptying inside spermatheca.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEEFFDFFCCEFBEBFDB5FAFA.taxon	materials_examined	Material Examined. Syntype of A. (Achanius) anthicoides (MNHUB) (Fig. 13); three specimens of Achanius (Achanius) bembidioides Fairmaire (1 NHMB, 2 IADIZA); six specimens of Achanius (Achanius) minutus Kulzer (IADIZA); holotype (MNHN) and 113 specimens (IADIZA) of Achanius (Ambigatus) rufonitens Fairmaire; holotype (NHMB) and three specimens (IADIZA) of A. (Ambigatus) stricticollis; holotype (NHMB) and 39 specimens (IADIZA) of Achanius (Ambigatus) bicolor Kulzer; holotype of Achanius (Ambigatus) rhinosomoides Kulzer (NHMB); paratype of Achanius (Ambigatus) piceus Kulzer (IADIZA); holotype of Achanius (Ambigatus) wittmeri Kulzer (NHMB) (Fig. 16); holotype, allotype, and nine paratypes of A. (Ambigatus) antofagastensis Flores and Aballay, new species.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFF45FA9CFB59FE46.taxon	description	(Fig. 13)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFF45FA9CFB59FE46.taxon	type_taxon	Type species: Achanius anthicoides Erichson, 1847, by monotypy.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFF45FA9CFB59FE46.taxon	diagnosis	Diagnosis. Kulzer (1950 in key) stated the humeri are not elevated and hind wings are absent. This study: metasternum 2 X length of mesosternum; metacoxal width twice metacoxal length; distance between meso – and metacoxae exceeding metacoxal length but not longer than 1.5 X metacoxal length.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFF45FA9CFB59FE46.taxon	biology_ecology	Biology. Species of Achanius (Achanius) are apterous and have been collected under stones or using pitfall traps partly filled with water and propylene glycol (Flores et al. 2004; Lagos 2004).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFF45FA9CFB59FE46.taxon	discussion	Species included. Achanius anthicoides, A. bembidioides, Achanius (Achanius) castanescens Fairmaire, Achanius (Achanius) piceofuscus Fairmaire, A. minutus, Achanius (Achanius) obscurus Kulzer, Achanius (Achanius) peruensis Kulzer, Achanius (Achanius) puncticollis Kulzer, and Achanius (Achanius) angusticollis Kulzer (Kulzer 1950, 1956).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFD71FE61FCE7FB71.taxon	description	(Figs. 14 – 20, 22)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFD71FE61FCE7FB71.taxon	type_taxon	Type species: Ambigatus stricticollis Fairmaire, 1892. Achanius (Ambigatus): Kulzer 1950: 31 (revision), 1956: 901 (revision).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFD71FE61FCE7FB71.taxon	diagnosis	Diagnosis. Kulzer (1950 in key) stated the humeri are not elevated and hind wings are present. This study: metasternum 3 X length of mesosternum; metacoxal width 3 X metacoxal length; distance between meso – and metacoxae 2.5 X metacoxal length (Fig. 20).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFD71FE61FCE7FB71.taxon	biology_ecology	Biology. Species of Achanius (Ambigatus) are winged, capable of flying, and of crepuscular and nocturnal habits. They were collected at white and UV lights, also using pitfall traps partly filled with water and propylene glycol (Flores et al. 2004; Lagos 2004), and cattle dung-baited pitfall traps.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFDFFD71FE61FCE7FB71.taxon	discussion	Species Included. Achanius rufonitens, A. stricticollis, A. bicolor, A. rhinosomoides, A. piceus, A. wittmeri (Kulzer 1950, 1956), and Achanius (Ambigatus) antofagastensis Flores and Aballay, new species.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	description	(Figs. 14 – 15)	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	diagnosis	Diagnosis. Antennomere 2 longer than 3, antennomeres 1 – 5 subcylindrical, antennomeres 6 – 10 triangular, longer than wide, antennomere 11 equal to 10, pear-shaped; apical maxillary palpomere ovoid, acuminate; prothorax longer than wide, anterior margin of pronotum exceeding width of posterior margin (Fig. 14). Achanius antofagastensis superficially resembles A. bicolor and A. wittmeri (Fig. 16) in color pattern. It differs from these species by having reduced wings, antennomere 2 longer than 3 and antennomere 11 equal to 10, whereas A. bicolor and A. wittmeri have well-developed wings, antennomere 2 shorter than 3, and antennomere 11 longer than 10. In addition, in A. wittmeri the prothorax is wider than long, and the elytra have an additional triangular dark brown spot pointed backwards between the humeri (Fig. 16).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	description	Description. Length 3.6 – 4.7 mm. Head and prothorax dark brown, elytra light brown on anterior half and posterior quarter, with dark brown spot on penultimate quarter extending along suture towards apex; antennae, legs, and mouthparts light brown as elytra. One specimen (Fig. 15) with same color pattern but elytra with spot outlined in dark brown. One specimen (Catamarca, 8 km W Antofagasta de la Sierra (16 km W of the holotype locality )) with similar color pattern but head, prothorax, legs, and elytral spot very dark brown. Humeri elevated, wings present, reduced, narrow, reaching 3 / 4 length of elytra, not folded (Fig. 15); metasternum 3 X length of mesosternum; metacoxal width 3 X metacoxal length (as in Fig. 20).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	etymology	Etymology. Named “ antofagastensis ” after the collection site of Antofagasta de la Sierra, Catamarca province, Argentina.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	materials_examined	Type Material. Holotype: Male (Fig. 14): [Argentina: Catamarca, 8 km NE / Antofagasta de la Sierra, Punta de la / Peña 3593 m 26 ° 01 ′ 39.54 ″ S, 67 ° / 20 ′ 38.79 ″ W, 17 - 20 - XI- 2009 / coll. G. Flores, F. Aballay] [Achanius (Ambigatus) / antofagastensis n. sp. / HOLOTYPUS male / Det. G. Flores and / F. Aballay 2015] (IADIZA). Allotype: Female (IADIZA) and four paratypes with the same data as holotype: one male (NHMB), one male (FMNH), one female (MACN), one not sexed (MACN). Paratypes: One male: [Argentina: Catamarca / (Dto.) Antofagasta de la Sierra, 8 km / NE Antofagasta de la Sierra / llama M- 4 P- 3, 3585 m / 26 ° 01 ′ 33.4 ″ S / 67 ° 20 ′ 42.5 ″ W, 21 - XI- 2009 / coll. F. Aballay] (MEKRB); one female with same label information except M- 11, 28 - XI- 2009 (IADIZA) (Fig. 15); one specimen: [Argentina: Catamarca / (Dto.) Antofagasta de la Sierra, 8 km / NE Antofagasta de la Sierra / cerdo al sol M- 11 P- 3, 3578 m 26 ° 01 ′ 32.3 ″ S, 67 ° / 20 ′ 36.5 ″ W, 27 - XI- 2009 / coll. F. Aballay] (MLPA); one male: [Argentina: Catamarca / (Dto.) Antofagasta de la Sierra, 8 km / NE Antofagasta de la Sierra / cerdo sombra M- 17 P- 1, 3595 m 26 ° 01 ′ 38.2 ″ S, 67 ° / 20 ′ 31.6 ″ W, 03 - XII- 2009 / coll. F. Aballay] (IADIZA), one specimen with same label information except M- 20, 06 - XII- 2009 without head, not labelled paratype (IADIZA); one specimen: [Argentina: Catamarca 8 km W / Antofagasta de la Sierra, Salar 3434 m / 26 ° 01 ′ 54.6 ″ S, 67 ° 27 ′ 07.7 ″ W / 11 - XII- 2009 / coll. G. Flores] (IADIZA).	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
182B9F50FFEFFFD2FD60FB11FCE6FE46.taxon	biology_ecology	Biology. Specimens collected in the spring at an altitude of over 3,500 m in Antofagasta de la Sierra, Catamarca province, were found feeding on decomposing pig carcasses (12 individuals) and on llama (Lama glama Linnaeus) carcasses (four individuals). These specimens showed a preference for pig carcasses, nine of which were recorded on the pig carcass in the shade and three under direct sun. This species was associated with the intermediate stages of decomposition (Active Decay and Advanced Decay) defined by Payne (1965). Because of these three behavioral characteristics, this species should be considered for potential forensic importance for estimating time of death (PMI) in arid high environments during the spring.	en	Flores, Gustavo E., Aballay, Fernando H. (2015): Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson. The Coleopterists Bulletin (mo 14) 69: 167-179, DOI: 10.1649/0010-065X-69.mo4.167
