taxonID	type	description	language	source
6BE9A183A5D3576BAFCD9B2782EB3610.taxon	description	New Japanese name: Himejimukade-zoku Fig. 5	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6BE9A183A5D3576BAFCD9B2782EB3610.taxon	diagnosis	Diagnosis. Partly modified from Foddai et al. (2003). Adult body length ca 10 mm (Fig. 5 A). Cephalic plate only slightly longer than wide, with frontal line absent or replaced by areolation. Two small clypeal plagulae covering ca one-sixth of the clypeus. Bucca without setae. Stilus present, relatively short. Spiculum absent. Side-pieces of labrum only incompletely subdivided into anterior and posterior alae by fragmented line very poorly marked. Mandible provided with four well-developed pectinate lamellae. Coxosternite of first maxillae medially divided. Coxosternite of second maxillae undivided, without suture or membranous isthmus. Metameric pore close to posterior margin of coxosternum of second maxillae, not to lateral ones. Claw of second maxillae only represented by terminal spine. Forcipular telopodites far behind anterior margin of head in the closed position. Forcipular trochanteroprefemur with pigmented single distal denticle; femur without teeth; tarsungulum with basal, well-developed denticle. Forcipular tergite without median sulcus. Sternal sulcus not anteriorly furcate. Last metasternite subtriangular. Ventral surface of each coxopleuron with numerous pores. Anal pore present. Forty-one pairs of legs.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6440CA979C1E52D8B3882368A02A34F9.taxon	description	New Japanese name: Kaguya-himejimukade Figs 5, 6, 7, 8, 9, 10, 11	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6440CA979C1E52D8B3882368A02A34F9.taxon	etymology	Etymology. The species name is derived from unmarried in Latin. In the Japanese population, males of this species have not been discovered despite the wide collection range in Japan.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6440CA979C1E52D8B3882368A02A34F9.taxon	diagnosis	Diagnosis. Clypeus with a pair of smooth or weakly areolate areas along the posterior part of the paraclypeal sutures; forcipular trochantroprefemur with a large denticle (longer than wide); tarsungulum with a well-pigmented denticle; metasternite of ultimate leg-bearing segment wider than long.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6440CA979C1E52D8B3882368A02A34F9.taxon	description	Description. General features (Fig. 5 A, B): Body 7.0 - 12.0 mm long (holotype 12.0 mm), gradually attenuated posteriorly, almost uniformly very pale yellow, with head and forcipular segment pale ocher. Cephalic capsule (Fig. 6 A-D): Cephalic plate ca 1.4 - 1.6 x as long as wide; lateral margins more distinctly converging anteriorly than posteriorly; posterior margin straight; scutes approximately isometric and up to 15 μm wide; transverse suture absent but areolate line present in some individuals (Fig. 6 A, C); setae up to ca 37.5 μm long. Clypeus ca 1.4 - 1.5 x as wide as long, with lateral margins complete, almost uniformly areolate, with scutes ca 15 μm wide, clypeal areas absent; clypeus with 11 - 17 setae, 1 + 1 postantennal, 1 - 2 + 2 median, 3 - 6 + 3 - 5 prelabral; clypeal ratio ca 1: 6 - 1: 7; clypeal plagulae with additional smooth or weak areolation area along posterior part of paraclypeal sutures. Anterior and distolateral parts of pleurites areolate, without setae. Side-pieces of labrum medially in contact, only incompletely divided into anterior and posterior alae by weak chitinous line, without longitudinal stripes on posterior alae, with slightly visible short fringe on posterior margin of side-pieces; mid-piece as long as wide, converging anteriorly and posteriorly. Antenna (Fig. 7 A-D): Antenna with 14 articles, when stretched, ca 2.1 - 2.6 x as long as head length. Intermediate articles slightly longer than wide. Article XIV ca 2.5 x as long as wide, ca 1.9 - 2.4 x as long as article XIII, and 1.8 - 2.4 x as long as intermediate articles. Setae on articles VIII-XVI denser than articles I-VII. Setae gradually shorter from article VIII to XIV, up to 65 μm long on article I, up to 25 µm long on article VIII and <15 μm long on article XIV. Article XIV with two types of sensilla; apical sensilla (arrows in Fig. 7 C, D) ca 10 μm long, with wide flat ring at mid-length; club-like (arrowheads in Fig. 7 C, D) sensilla ca 15 μm long, clustered in the distal part of the internal and the external sides of the article. Three longitudinal rows, each consists of ca 9 proprioceptive spine-like sensilla, at bases of antennal articles III-V, VII-IX, approximately dorsal, ventro-internal and ventro-external; the rows each consisting of 1 - 3 spine-like sensilla on articles I and VI, and 6 on the article II; 0 - 1 spine-like sensilla on articles X-XIV. A few pointed sensilla, up to 2.5 μm long, on both dorso-external and ventro-internal position, close to distal margin of articles II, V, IX and XIII. Mandible (Fig. 8 A): At least four pectinate lamellae present; first and second lamellae with ca 5 elongated teeth. Each tooth ca 2 x as long as wide. Ventral surface hairy. First maxillae (Fig. 8 B): Coxosternite medially divided but slightly, without setae, faintly areolate. Coxal projections well developed and hyaline distally, with 1 - 2 + 1 - 2 setae and 3 - 4 + 3 - 4 small sensilla. Telopodite uni-articulated and hyaline distally, with one (two) seta (e). No lobes on either coxosternite or telopodites. Second maxillae (Fig. 8 C, D): Coxosternite medially undivided, without suture, with 2 + 2 setae along anterior margin, with 6 - 7 setae behind anterior margin, with 1 - 2 + 1 - 2 sensilla on posterior lateral margin in some individuals. with anterior margin slightly concave, with metameric pores on posterior part. Telopodites tri-articulate, reaching medial projections and telopodites of first maxillae in some individuals. Claw of telopodite virtually absent, represented by short spine only. Forcipular segment (Fig. 9 A-F): Tergite trapezoidal, ca 1.4 - 1.9 x as wide as long, with lateral margins converging anteriorly, approximately as wide as cephalic plate and ca 0.7 x as wide as following tergites; 1 + 1 setae of similar length arranged in an anterior row, and 3 + 3 setae of similar length arranged in a posterior row; one pair of longitudinal rows of three tiny setae located between middle and distal setae in posterior row. Mid-longitudinal sulcus of tergite not visible. Exposed part of coxosternite ca 1.1 x as wide as long; anterior margin with shallow medial concavity and with one pair of denticles; coxopleural sutures complete in entire ventrum, sinuous and diverging anteriorly; chitin-lines absent. Trochanteroprefemur ca 1.3 - 1.4 x as long as wide; with a well-developed and strong pigmented denticle at distal internal margin, ca 1.3 - 1.6 x as long as wide. Intermediate articles distinct, with a tubercle on tibia (arrowed in Fig. 9 E, some individuals not visible). Tarsungulum with well-pigmented basal denticle; both external and internal margins uniformly curved, except for moderate mesal basal bulge; ungulum not distinctly flattened. Elongated poison calyx (circle in Fig. 9 F), ca 9 x as long as wide, lodged inside intermediate forcipular articles. Leg-bearing segments (Figs 9 A, B, 10 A-C): Forty-one pairs of legs present. Metatergite 1 slightly wider than subsequent one, with two paramedian sulci visible on tergites of anterior half of body, with pretergite. No paratergites. Walking legs shorter than width of trunk; legs of first pair much smaller than following ones; claws simple, uniformly bent, with 2 accessory spines; anterior spine reaching at most 10 % of length of claw; posterior spine equal in length of the anterior spine. Metasternites slightly longer than wide. Sternal sulcus visible on a few anterior sternites, represented by very shallow mid-longitudinal thickening, anteriorly not furcate. No ventral glandular pores on each metasternite. Ultimate leg-bearing segment (Fig. 11 A-D): Pretergite not accompanied by pleurites but incomplete traces of sutures present at both sides. Metatergite subtrapezoidal, ca 1.1 - 1.3 x as wide as long; lateral margins convex and converging posteriorly; posterior margin slightly curved. Coxopleuron ca 1.2 - 1.7 x as long as metasternite; coxal organs of each coxopleuron opening through 5 - 10 independent pores, placed ventrally. Metasternite trapezoidal, ca 1.3 - 1.6 x as wide as long, anteriorly ca 1.7 - 2.2 x as wide as posteriorly; lateral margins slightly convex and converging backward; setae almost arranged symmetrically, dense on posterior margin. Telopodite ca 9 - 11 x as long as wide, ca 1.9 - 2.1 x as long and ca 1.5 - 1.6 x as wide as penultimate telopodite, with six articles; tarsus 2 ca 3.6 - 4.2 x as long as wide and ca 1.2 - 1.6 x as long as tarsus 1; setae arranged uniformly, <70 μm long; pretarsus represented by small tubercle. Female postpedal segments (Fig. 11 A, B): Two gonopods basally touching, subtriangular, without traces of articulation, covered with setae. Anal pore present. Male postpedal segments unknown (male unknown).	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
6440CA979C1E52D8B3882368A02A34F9.taxon	distribution	Distribution. Honshu, Shikoku and Kyushu.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
1B1A7490A2C854008BEFD6BC3F5DAE32.taxon	description	New Japanese name: Amefuri-himejimukade Figs 12, 13, 14, 15, 16, 17	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
1B1A7490A2C854008BEFD6BC3F5DAE32.taxon	etymology	Etymology. The species name is derived from the name of Japanese mountain, namely Mt. Oyama. The word was further Latinized by adding the Latin masculine adjective suffix - ensis, to form Nannarrup oyamensis oyamensis. The last " a " of Oyama and the first " e " of - ensis are merged into " e. " Examined specimens were collected from Mt. Oyama, an object of the mountain worship.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
1B1A7490A2C854008BEFD6BC3F5DAE32.taxon	diagnosis	Diagnosis. Clypeus without smooth or weakly areolate areas along the posterior part of the paraclypeal sutures; forcipular trochantroprefemur with a large denticle (longer than wide); tarsungulum with a well-pigmented denticle; metasternite of ultimate leg-bearing segment wider than long.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
1B1A7490A2C854008BEFD6BC3F5DAE32.taxon	description	Description. General features: Body 8.6 mm long (holotype), gradually attenuate posterior, almost uniformly very pale yellow, with head and forcipular segment pale ocher. Cephalic capsule (Fig. 12 A, B): Cephalic plate ca 1.5 x as long as wide, lateral margins more distinctly converging anteriorly than posteriorly, posterior margin straight; scutes approximately isometric and up to ca 15 μm wide; transverse suture absent; setae up to ca 50 μm long. Clypeus ca 1.5 x as wide as long, with lateral margins complete, almost uniformly areolate, with scutes ca 10 μm wide, a pair of clypeal areas absent; 13 setae in holotype, 1 + 1 postantennal, 1 + 1 median, 5 + 4 prelabral; clypeal ratio ca 1: 7; clypeal plagulae without additional smooth area along posterior part of paraclypeal sutures; 17 pore-like organs on entire part of clypeus. Anterior and distolateral parts of pleurites areolate, without setae. Side-pieces of labrum medially in contact, only incompletely divided into anterior and posterior alae by weak chitinous line, without longitudinal stripes on posterior alae; with slightly visible short fringe on posterior margin of side-pieces; mid-piece as long as wide, converging anteriorly and posteriorly. Antenna (Fig. 13 A-D): Antenna with 14 articles, when stretched, ca 2.3 x as long as head length. Intermediate articles slightly longer than wide. Article XIV ca 2.0 x as long as wide, ca 1.9 x as long as article XIII, and 1.9 - 2.1 x as long as intermediate articles. Setae on articles VIII-XVI denser than articles I-VII. Setae gradually shorter from article VIII to XIV, up to 50 μm long on article I, up to 33 µm long on article VIII and <18 μm long on article XIV. Article XIV with two types of sensilla, apical sensilla (arrows in Fig. 13 C, D) ca 5 μm long, with wide flat ring at mid-length; club-like sensilla (arrowheads in Fig. 13 C, D) ca 10 μm long, clustered in the distal parts of the internal and external sides of the article. Three longitudinal rows consisted of ca 9 proprioceptive spine-like sensilla at bases of antennal articles II-V, VII-IX, approximately dorsal, ventro-internal and ventro-external; rows reduced to 1 - 3 spines on antennal articles I and VI, and 0 - 1 spine on antennal articles X-XIV. A few pointed sensilla, up to 2.5 μm long, on both dorso-external and ventro-internal position, close to distal margin of articles II, V, IX and XIII. Mandible (Fig. 14 A). At least four pectinate lamellae, with elongated teeth. Each tooth ca 2 x as long as wide. First maxillae (Fig. 14 B): Coxosternite medially divided but slightly, without setae, faintly areolate. Coxal projections well developed and hyaline distally, provided with 1 + 1 setae and 3 + 4 small sensilla. Telopodite uni-articulated and hyaline distally, with one (two) seta (e). No lobes on either coxosternite or telopodites. Second maxillae (Fig. 14 B): Coxosternite medially undivided, without suture, with 2 + 3 setae along the anterior margin, with 4 + 4 setae located behind anterior margin, with anterior margin slightly concave, with metameric pores on posterior part. Telopodites tri-articulate overreaching medial projections and telopodites of first maxillae. Claw of telopodite virtually absent, represented by short spine only. Forcipular segment (Fig. 15 A-D): Tergite trapezoidal, ca 1.9 x as wide as long, with lateral margins converging anteriorly, approximately as wide as cephalic plate and ca 0.7 x as wide as following tergite, 1 + 1 setae of similar length arranged in an anterior row, and 3 + 3 setae of similar length arranged in a posterior row, one pair of longitudinal rows of three tiny setae located between middle and distal setae in posterior row. Mid-longitudinal sulcus of tergite not visible. Exposed part of coxosternite ca 1.3 x as wide as long; anterior margin with shallow medial concavity and with one pair of denticles; coxopleural sutures complete in entirely ventrum, sinuous and diverging anteriorly; chitin-lines absent. Basal distance between forcipules ca 0.1 x of maximum width of coxosternite. Trochanteroprefemur ca 1.3 x as long as wide; with a well-developed and strong pigmented denticle at distal internal margin, ca 1.3 x as long as wide. Intermediate articles distinct, tubercle on tibia not visible. Tarsungulum with basal denticle well-pigmented; both external and internal margins uniformly curved, except for moderate mesal basal bulge; ungulum not distinctly flattened. Elongated poison calyx (circle in Fig. 15 D), ca 6 x as long as wide, lodged inside intermediate forcipular articles. Leg-bearing segments (Fig. 16 A-D): Forty-one pairs of legs present. Metatergite 1 slightly wider than subsequent one, with two paramedian sulci visible on tergites of anterior half of body, without pretergite. No paratergites. Walking legs shorter than width of trunk; legs of first pair much smaller than following ones; claws simple, uniformly bent, with two accessory spines; anterior spine reaching at most 10 % of length of claw; posterior spine shorter than anterior spine. Metasternites slightly longer than wide. Sternal sulcus visible on a few anterior sternites, represented by very shallow mid-longitudinal thickening, anterior not furcate. No ventral glandular pores on each metasternite. Ultimate leg-bearing segment (Fig. 17 A-D): Pretergite not accompanied by pleurites but incomplete traces of sutures present at both sides. Metatergite subtrapezoidal, almost as wide as long, lateral margins convex and converging posteriorly; posterior margin slightly curved. Coxopleuron ca 1.2 x as long as metasternite; coxal organs of each coxopleuron opening through five or six independent pores, placed ventrally. Metasternite subtriangular, ca 1.6 as wide as long, anteriorly ca 3.5 x as wide as posteriorly; lateral margins slightly convex and converging backward; setae almost arranged symmetrically, dense on posterior margin. Telopodite ca 11 x as long as wide, ca 1.9 x as long and ca 1.3 x as wide as penultimate telopodite, with 6 articles; tarsus 2 ca 2.7 x as long as wide and ca 1.5 x as long as tarsus 1; setae arranged uniformly, <50 μm long. Pretarsus represented by spines, up to 5 µm. Male postpedal segments (Fig. 17 A, B): Two gonopods, very widely separated from one another, conical in outline, uni-articulated without any sutures, covered with setae. Anal pore present. Female postpedal segments unknown (female unknown).	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
1B1A7490A2C854008BEFD6BC3F5DAE32.taxon	distribution	Distribution. Only known from Mt. Oyama, located in Isehara-shi, Kanagawa prefecture.	en	Tsukamoto, Sho, Shimano, Satoshi, Eguchi, Katsuyuki (2022): Two new species of the dwarf centipede genus Nannarrup Foddai, Bonato, Pereira & Minelli, 2003 (Chilopoda, Geophilomorpha, Mecistocephalidae) from Japan. ZooKeys 1115: 117-150, DOI: http://dx.doi.org/10.3897/zookeys.1115.83946, URL: http://dx.doi.org/10.3897/zookeys.1115.83946
