taxonID	type	description	language	source
594D037701061223FF656B96FC8DF9D5.taxon	type_taxon	Type species: Calosega kamiteta Terayama, 1999: 102.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061223FF656B96FC8DF9D5.taxon	diagnosis	Diagnosis. Scapal basin shallow and cross-ridged, dorsal margin not carinate; posterior margin of vertex almost straight; occipital carina complete; malar sulcus developed; flagellum compact, intermediate flagellomeres shorter (♀) or at most slightly longer (♂) than wide; pronotum thick, about as long as or slightly longer than mesoscutum along mid-line, with posteromedial and lateral pits; mesoscutum with notauli complete; short track of parapsidal signum present, sometimes indicated as small depression; mesopleuron with omaulus and posterior oblique sulcus (= scrobal sulcus by Kimsey & Bohart (1991 )), sometimes weakly differentiated from other surface sculptures; metanotum as long as mesoscutellum; median enclosure of metanotum present, sculptured as mesoscutum; metapectal-propodeal complex with spine-like posterior propodeal projection (female) or posterolateral corner angulate (male); both sexes fully winged; narrow R 1 2 v indicated or not differentiated from pterostigma; distal part of pterostigma + R 1 2 v 0.8 × 2 r-rs 2 v; 2 r-rs 2 v almost straight or weakly curving; dark streak of Rs 2 v almost straight or weakly curving; Rs & M 2 v arising at Cu-a 2 v; tarsal claw with submedian tooth; metacoxa with dorsal carina; metasoma sculptured by small and sparsely located punctures. General characteristics of Calosega are similar to those of Isegama and Cladobethylus Kieffer, 1922. However, Isegama has a median pronotal groove (not pit-like), and a rounded posterolateral corner of metapectal-propodeal complex (the posteromedian pit is present, and the metapectal-propodeal complex is angulate or spinose in Calosega); and Cladobethylus has an impunctate welt on vertex (welt absent).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061223FF656B96FC8DF9D5.taxon	biology_ecology	Host. Unknown.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061223FF656B96FC8DF9D5.taxon	distribution	Distribution. Known from Palaearctic Japan (Honshu), Oriental Japan (Ryukyus) (Terayama, 1999; Terayama & Suda, 2014; Katayama, 2021) and Thailand (Kimsey, 1995).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061223FF656B96FC8DF9D5.taxon	discussion	Remarks. Two species Calosega kamiteta Terayama, 1999, C. flavipes (Kimsey, 1995) are known. Calosega kamiteta was rarely found in temperate area (Honshu, Japan) (Terayama & Suda, 2014; Katayama, 2021), but the center of abundance of the genus is considered to be in subtropics.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061220FF656FB9FC02FE6B.taxon	description	(Figs 1 – 4)	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061220FF656FB9FC02FE6B.taxon	materials_examined	Holotype ♀, “ THAILAND: 180 km N. E. Bangkok, Khao Yai N. P., 780 m, 11 - 18 - IV- 1990, M. T., Moist Semi Evergreen Forest, B. V. Brown ”, “ HOLOTYPE, Perissosega flavipes Kimsey ”, “ HOLOTYPE Perissosega flavipes Kimsey, CNC No. 22205 ”, “ CNC, 957622 ” (CNC, photos examined).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701061220FF656FB9FC02FE6B.taxon	discussion	Remarks. 2 r-rs 2 v is almost straight and the dark streak of Rs 2 v is also almost straight, but the connection is strongly arced (Fig. 1). It appears similar to those of Perissosega and other related genera. However, as indicated by Kimsey (1995), other important characters do not match the diagnosis of Perissosega (Krombein, 1983; Kimsey & Bohart 1991). The absence of a smooth strip on vertex (Fig. 3), the well-developed and not-excavated vertex, the presence of a complete omaulus and posterior oblique sulcus (Fig. 2), and also the presence of a dorsal carina on metacoxa (Fig. 4) are common with Isegama among the genera known in the Indo-China region. However, the presence of postero-medial pit on pronotum and the developed posterior propodeal tooth places this species in the genus Calosega. The general habitus, and the smaller and sparsely located punctures of the body are similar to Calosega kamiteta Terayama, 1999; however, C. flavipes is distinguished from C. kamiteta by the wider flagellum, F 2 2.0 × wider than long (F 2 1.5 × wider than long) and the unique condition of Rs 2 v (Rs 2 v is normal in C. kamiteta, the 2 r-rs 2 v and dark streak of Rs 2 v is weakly curving, the connection does not from a strong arc).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701051221FF656C15FC28FF4B.taxon	type_taxon	Type species: Perissosega venablei Krombein, 1983: 27.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701051221FF656C15FC28FF4B.taxon	diagnosis	Diagnosis. Scapal basin cross-ridged, sometimes dorsal margin surrounded by strong carina; occipital carina variable, but incomplete; malar sulcus developed; posterior margin of vertex weakly excavated; flagellum compact, intermediate flagellomeres shorter than (♀) or as long as (♂) wide; pronotum thick, about as long as mesoscutum along mid-line, with posteromedial pit and pit before lateral lobe; mesoscutum with notauli and short track of parapsidal signum; mesopleuron without omaulus and posterior oblique sulcus; metanotum more than half as long as mesoscutellum; median enclosure of metanotum present, sculptured as mesoscutum; metapectal-propodeal complex with posterior propodeal projection; both sexes fully winged; R 1 2 v not discriminated from pterostigma; distal part of pterostigma + R 1 2 v 0.4 – 0.7 × 2 r-rs 2 v; 2 r-rs 2 v weakly curving; dark streak of Rs 2 v extended at abrupt angle from 2 r-rs 2 v; Rs & M 2 v variable, usually arising before Cu-a 2 v; tarsal claw with submedian tooth; metacoxa with dorsal carina; metasoma more or less densely punctate by large punctures. Perissosega is probably most closely related to Atoposega. This genus is distinguished from Atoposega by following characteristics: the body is largely black (the ground color of the body is red and black in Atoposega); the metanotum is punctate as mesoscutellum (differently sculptured from the mesoscutellum); and the metasoma is sculptured by large and densely located punctures (usually more or less finely scratched) (Krombein, 1983; Kimsey & Bohart, 1991; Kimsey, 2014).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701051221FF656C15FC28FF4B.taxon	biology_ecology	Host. Unknown.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701051221FF656C15FC28FF4B.taxon	distribution	Distribution. Oriental genus known from Sri Lanka (Krombein, 1983) and Japan, new record.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701051221FF656C15FC28FF4B.taxon	discussion	Remarks. Two species, P. venablei and P. sulcata sp. nov. are known.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701041221FF656803FEC1FBD0.taxon	description	(Figs 5 – 10)	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701041221FF656803FEC1FBD0.taxon	diagnosis	Diagnosis. Body 2.9 – 3.5 mm in female, 3.6 – 2.9 mm in male; black with dark brown mandibles, antennae and legs, articulation of legs paler; wings strongly infuscate with two clear bands, legs sometimes brown; head punctate, almost contiguous; scapal basin with dorsal margin surrounded by strong carina (Fig. 7); occipital carina posterior to ocellar region absent or effaced (Fig. 9), laterally always complete, extending ventral to hypostoma; ocellar triangle narrow, POL 0.8 × OL (Fig. 8); mesosoma (Fig. 10) punctate, punctures almost contiguous; mesopleuron with punctures, dorsally denser, almost punctate-reticulate (Fig. 5); median enclosure of metanotum 0.75 × mesoscutellum; metapectal-propodeal complex bearing spine-like posterior propodeal projection; distal part of pterostigma + R 1 2 v 0.7 × 2 r-rs 2 v (Fig. 6); 2 r-rs 2 v weakly curving; Rs & M 2 v arising before (holotype) or almost at (one paratype ♀) Cua 2 v; metasoma punctate, punctures shallow, somewhat larger and denser than those on mesosoma; median polished line more or less present on T 2 (♀) or absent (♂). About the recognition of P. venablei from the closely allied species, see the diagnosis of P. sulcata sp. nov. Specimens examined. Holotype ♀, “ SRI LANKA: Gal. Dist., Kanneliya Jungle ”, “ 14 - VII- 1978, K. V. Krombein, P. B. Karunaratne ”, “ HOLOTYPE, PERISSOSEGA VENABLEI ♀, Karl V. Krombein ”, “ Type No. 100444, U. S. N. M. ” (USNM). Paratypes 1 ♀, Sri Lanka, Mat. Dist., Kibissa, 0.5 miwest, Sigiriya, Jungle, 3. VI. 1978, K. V. Krombein, P. B. Karunaratne leg. (USNM); 1 ♂ (allotype), Sri Lanka, Rat. Dist., Gilimale, Induruwa, Jungle, 2. II. 1979, Malaise trap, K. V. Krombein, P. B. Karunaratne, T. Wijesinhe, S. Siriwardane, T. Gunawardane leg. (USNM).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701031227FF6569B3FEB3FC1F.taxon	diagnosis	Diagnosis. Body 2.1 – 2.3 mm; black, legs partly testaceous; fore wing strongly infuscate around 2 r-rs 2 v; head punctate, punctures almost contiguous on vertex; scapal basin with dorsal margin not carinate; occipital carina present only posterior to ocellar region; ocelli forming regular triangle; mesosoma densely punctate; dorsal part of mesopleuron densely punctate and rugose; posterolateral corner of metapectal-propodeal complex angulate, forming a blunt posterior propodeal projection; distal part of pterostigma + R 1 2 v 0.4 × 2 r-rs 2 v; 2 r-rs 2 v weakly curving; Rs & M 2 v arising almost at Cu-a 2 v; metasoma densely punctate, punctures shallow, somewhat smaller and sparser than those on mesosoma; median polished line present from T 1 to T 2. Male unknown. This new species is distinguished from P. venablei by the shallow scapal basin without a strong dorsal carina (scapal basin is deep and the dorsal margin is surrounded by a strong transverse carina in P. venablei); the incomplete occipital carina only present posterior to ocellar region (laterally complete, only absent or effaced posterior to ocellar region); the blunt posterior propodeal projection on the metapectal-propodeal complex (a long spine-like projection is present); the median smooth line on T 1 (smooth line absent on T 1).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701031227FF6569B3FEB3FC1F.taxon	description	Description. Holotype female. Body length 2.05 mm. Head (Figs 12, 14, 15) punctate except scapal basin, 1.6 × as wide as long in dorsal view, as wide as deep in frontal view; punctures 0.3 – 0.6 × MOD, 0.2 – 0.5 × PD apart with smooth interspaces, denser and contiguous on vertex; scapal basin (Fig. 14) shallow, cross-ridged with smooth median line; strong carina absent around dorsal margin; frons dorsally converging, shortest distance between eyes 0.33 × head width; MS 0.36 × maximum length of eye; lateral ocelli almost touching eye, OL 1.0, OPL 2.1, POL 1.0, OOL 0.1, MOD 0.6; occipital carina present only posterior to ocellar triangle; posterior margin of vertex excavated. Mandible cylindrical. Eye setose. Antenna compact, F 2 – F 8 wider than long; length (width) of F 1 – F 4 following ratio: 6.0 (2.0): 1.5 (2.5): 2.0 (3.0): 2.0 (3.0). Mesosoma (Fig. 16) covered with short decumbent setae. Pronotum punctate with smooth interspaces; punctures somewhat longitudinally contiguous, as large as those on frons, 0.3 – 1.0 × PD apart, sparser posteriorly; medial longitudinal line reaching 0.8 × length of pronotum; median length of pronotum 0.62 × longer than wide. Mesoscutum as long as pronotum, punctate with smooth interspaces; punctures smaller and denser on median part, larger and sparser on lateral part. Mesoscutellum 0.6 × mesoscutum, sculptured as median part of mesoscutum. Mesopleuron punctate with smooth interspaces; punctures 0.5 MOD, 0.2 – 0.6 × PD apart; dorsal part densely punctate and rugose. Metanotum with median triangular enclosure, median length 0.54 × mesoscutellum, densely punctate. Metapectalpropodeal complex rugose, with blunt posterior propodeal projection; dorsal surface slightly shorter than propodeal declivity, with median longitudinal carina; propodeal declivity weakly rugose, with median, sublateral and lateral carina. Fore wing tinged with brown, around 2 r-rs 2 v more strongly infuscate (Fig. 13). Metasoma (Fig. 14) punctate with smooth interspaces, surface covered with short decumbent setae; punctures on T 1 somewhat longitudinally contiguous, ca. 0.3 × MOD, 0.2 – 1.0 × PD apart, denser posteriorly, punctures on T 2 as large as T 1, anteriorly denser, posteriorly sparser; median smooth line with one fine groove present on T 1 and T 2. Color. Body black except mandible, antenna and tegula dark brown, joints of legs, protibia and tarsi testaceous, other part of legs dark brown, basal part of metasoma brownish. Specimens examined. Holotype ♀, “ JPN: Okinawa Pref., Iriomote-jima Isl., Haiminaka, yellow pan trap, 24.2897 N, 123.8775 E, 20 m alt., 28 - VIII- 2020, T. Mita leg. ” (ELKU). Paratype 1 ♀, same island as holotype, but Komi, Airagawa, 24.3261 N, 123.8914 E, 15. V. 2014, H. Handa leg. (ELKU).	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701031227FF6569B3FEB3FC1F.taxon	biology_ecology	Host. Unknown.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701031227FF6569B3FEB3FC1F.taxon	etymology	Etymology. The specific epithet sulcata, a Latin adjective, meaning bearing a groove. The species name refers to the median groove on metasoma.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
594D037701031227FF6569B3FEB3FC1F.taxon	discussion	Remarks. The paratype female shows the following variation: body length 2.25 mm; punctures on metasoma slightly denser.	en	Mita, Toshiharu (2022): Taxonomy of Perissosega Krombein (Hymenoptera: Chrysididae, Amiseginae) with description of a new species from Japan. Zootaxa 5169 (3): 279-285, DOI: https://doi.org/10.11646/zootaxa.5169.3.4
