identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038E04381F73FFAFFF25AD99FEEAA395.text	038E04381F73FFAFFF25AD99FEEAA395.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus affinis (H. Milne Edwards 1837)	<div><p>Metapenaeus affinis (H. Milne Edwards, 1837)</p> <p>(Figs. 1, 8a)</p> <p>Penaeus affinis H. Milne Edwards 1837: 416 (type locality: India).</p> <p>Metapenaeus affinis — Maki &amp; Tsuchiya 1923: 38, pl. 2-fig. 3.— Lee &amp; Yu 1977: 91, figs. 62-63.— Miquel 1982: 47, figs. 11- 12.— Yu &amp; Chan 1986: 143, fig. 16B, 2 unnumbered figs.— Lee et al. 1999: 445.</p> <p>Material examined. Taipei City, Taipei market, 7 May 1985, 1 male cl 18.5 mm (NTOU M02305). Yunlin County, Mailiao, Station 1, 1 female cl 14.4 mm (NTOU M02428). Chiayi County, Budai fishing port, 2 Jul 2002, 9 males cl 18.9-21.7 mm, 15 females cl 21.0- 28.4 mm (NTOU M02306); 25 Feb 2021, 1 female cl 21.6 mm (NTOU M02412).— Dongshi fishing port, 12 Dec 2021, 2 males cl 22.2, 23.8 mm (NTOU M02403). Kaohsiung City, Kaohsiung fishing port, Mar 1975, 4 males cl 16.0- 22.3 mm, 4 females cl 15.8-22.8 mm (NTOUM02312).— Neiwei market, 17 Oct 2019, 1 female cl 29.2 mm (NTOU M02313).— Singda fishing port, 24 Jul 1984, 1 male cl 17.9 mm (NTOU M02314).— Kaohsiung Port, 5 Jan 1995, 9 males cl 12.8-19.2 mm, 9 females cl 16.3-33.2 mm (NTOU M02311); Station 1, 1 Mar 1994, 6 males cl 10.9-20.8 mm, 9 females cl 14.6-21.5 mm (NTOU M02308); Station 2, 1 Mar 1994, 1 male cl 19.5 mm, 2 juveniles cl 8.6, 13.0 mm (NTOU M02309); Station 3, 1 Mar 1994, 1 male cl 16.11 mm (NTOU M02413); Station 6, 1 Mar 1994, 1 male cl 19.2 mm, 3 females cl 20.2-31.6 mm (NTOU M02310). Pingtung County, Donggang fishing port, 27 Dec 1974, 2 males cl 19.4, 22.8 mm, 1 female cl 23.3 mm (NTOU M02315); 21 Oct 1995, 1 male cl 22.15 mm, 1 female cl 18.6 mm (NTOU M02414); 14 Aug 2015, 1 male cl 23.5 mm (NTOU M02316); 9 May 2019, 1 male cl 22.2 mm, 4 females cl 23.9-28.3 mm (NTOU M02317); 5 Mar 2021, 1 male cl 23.1 mm (NTOU M02318). No Specific data, 1 female cl 34.4 mm (NTOU M02429)</p> <p>Diagnosis. Rostrum with 6–9 (excluding epigastric tooth) teeth along entire dorsal border, generally straight and with tip curved upwards, more or less extending to tip of antennular peduncle. Pereiopod I with ischial spine absent to distinct. In males, basial spine of pereiopod III not particular elongated, merus of pereiopod V bearing basal notch accompanied with carinated and twisted tubercle. Telson with minute movable lateral spinules. Male petasma with distomedian projections very low and crescent shaped, reaching behind tips of distolateral projections. Female thelycum with anterior plate having posterior part much wider than anterior part; coxal projections of pereiopod IV round plate like and often concealing anterior plate but not extending to lateral plates, lateral plates as two overlapped rounded flaps without any sharp ridge and median longitudinal fissure.</p> <p>Coloration. Body generally greyish green and densely covered with dark dots. Antennal flagella, distal parts of pereiopods and pleopods reddish. Uropods red margined with distal parts somewhat greenish yellow.</p> <p>Distribution. Widely distributed in the Indo-West Pacific from the Persian Gulf to Japan and Papua New Guinea, intertidal to 92 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997; Chan 1998).</p> <p>Remarks. Metpenaeus affinis mainly occurs in southern Taiwan and rather common there. Amongst the species of this genus in Taiwan, M. affinis is most similar to M. ensis (De Haan, 1844). These two species can be distinguished by the following characters. The petasma has the distomedian projections very low in M. affinis (Fig. 1b) but as huge protuberances in M. ensis (Fig. 3b). The lateral plate of the thelycum has the posterolateral margins strongly folded anteriorly in M. ensis (Fig. 3c) but not folded up at all in M. affinis (Fig. 1b). In mature females the coxal projection of pereiopod IV is normal and not expanded in M. ensis, but round plate like and often concealing the anterior plate of thelycum in M. affinis. In mature males, the basal protuberance at the merus of pereiopod V is rounded in M. affinis (Fig. 1d) but spiniform in M. ensis (Fig. 3d). Moreover, the ischial spine of the pereiopod I in males is often (but not always) indistinct or absent in M. affinis but distinct in M. ensis. The tip of the rostrum is somewhat curved upwards in M. affinis (Fig. 1a) but generally straight in M. ensis (Fig. 3a). The posterior parts of the uropods are often greenish yellow in M. affinis (Fig. 8a) instead of bluish purple in M. ensis (Fig. 8c). Moreover, in large specimens the pereiopods are somewhat banded with red and while in M. ensis (Fig. 8c) but non-banded in M. affinis (Fig. 8a). Nevertheless, juveniles of these two species with gentalia not fully developed are rather difficult to separate.</p> </div>	http://treatment.plazi.org/id/038E04381F73FFAFFF25AD99FEEAA395	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F72FFAAFF25AD1BFB74A468.text	038E04381F72FFAAFF25AD1BFB74A468.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus elegans De Man 1907	<div><p>Metapenaeus elegans De Man, 1907</p> <p>(Figs. 2, 8b)</p> <p>Metapenaeus elegans De Man 1907: 130 (type locality: Indonesia).— Miquel 1982: 83, figs. 31-32.— Lee et al. 1999: 445.</p> <p>Penaeopsis elegans —De Man 1911: 58, pl. 6-fig. 16.</p> <p>Material examined. Yilan County, Dasi fishing port, 14 Oct 1988, 1 male cl 19.1 mm (NTOU M02319).</p> <p>Diagnosis. Rostrum with 8 (excluding epigastric tooth) teeth along entire dorsal border, generally straight and extending to tip of antennular peduncle. Pereiopod I with small ischial spine. In males, basial spine of pereiopod III not particular elongated, merus of pereiopod V with basal notch accompanied by carinated tubercle, which slightly bent outward. Telson with minute movable lateral spinules. Male petasma with distomedian projections semi-circular and flap-like, extending to tips of distolateral projections.</p> <p>Coloration. Body somewhat pinkish grey and covered with dense dark dots. Antennal flagella and distal half of uropods reddish.</p> <p>Distribution. Indo-West Pacific from Sri Lanka to Taiwan and Wallis and Futuna Islands, intertidal to 55 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997; Chan 1998).</p> <p>Remarks. Although the list of Penaeoidea from Taiwan and adjacent areas by Lee et al. (1999) contained M. elegans as a new record, their sampling areas included many localities far from Taiwan in the South China Sea, East China Sea and northwest of the Philippines (Lee et al. 1999: fig.1). Moreover, Lee at al. (1999) only listed M. elegans with no further information how such record was based (e.g., no voucher specimen information provided). This work formally reports M. elegans from Taiwan for the first time and extends the northern geographical range of this species from the Philippines to Taiwan. This species is very rare in Taiwan and at present only one male has been collected from the northeastern coast of the island despite of the recent extensive surveys on the decapod crustacean fauna in Taiwan. Amongst the species of this genus in Taiwan, M. elegans is most similar to M. moyebi (Kishinouye, 1896). Nevertheless, their petasma are very different with the distomedian projections extending far beyond the distolateral projections in M. moyebi (Fig. 6b) but reaching as far as the tips of distolateral projections in M. elegans (Fig. 2b). According to Miquel (1982: fig. 31c), the thylcum of M. elegans has the anterior end of anterior plate smooth while the lateral margins of the lateral plates distinctly ridged (v.s. anterior end of anterior plate bearing three tubercles while lateral plates with margins not ridged in M. moyebi, Fig. 7c).</p> </div>	http://treatment.plazi.org/id/038E04381F72FFAAFF25AD1BFB74A468	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F77FFA8FF25A98CFEB9A44D.text	038E04381F77FFA8FF25A98CFEB9A44D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus ensis (De Haan 1844)	<div><p>Metapenaeus ensis (De Haan, 1844)</p> <p>(Figs. 3, 8c)</p> <p>Penoeus Ensis De Haan 1844: pl. 46-fîg. 2 (type locality: Japan).</p> <p>Metapenaeus monoceros — Maki &amp; Tsuchiya 1923: 39, pl. 3-fig. 4.— Chang 1965: 6, 3 unnumbered figs.— Lee &amp; Yu 1977: 94, figs. 64-65. [not Fabricius, 1798]</p> <p>Metapenaeus ensis — Miquel 1982: 89, figs. 35-36.— Yu &amp; Chan 1986: 139, figs. 15B, 16A., 4 unnumbered figs.— Lee et al., 1999: 445.</p> <p>Material examined. Yilan County, Dasi fishing port, 5 Aug 1982, 3 males cl 20.3-33.1 mm (NTOU M02320); 17 Jul 1984, 1 male cl 24.2 mm, 1 female cl 21.0 mm (NTOU M02430); 29 Jul 1984, 1 female cl 20.3 mm (NTOU M02431); 10 Mar 1985, 1 female cl 31.1 mm (NTOU M02321); 16 Apr 1988, 1 male cl 20.5 mm (NTOU M02432); 8 Jan 2008, 1 male cl 20.4 mm, 1 female cl 32.2 mm (NTOU M02322); 19 Oct 2008, 1 male cl 28.1 mm, 1 female cl 41.4 mm (NTOU M01205).— Nanfang-ao fishing port, 20 Apr 1985, 1 male cl 30 mm (NTOU M02324).— Luodong market, 3 May 1988, 1 male cl 17.5 mm (NTOU M02433).— Lanyang River estuary, 18 Jan 2005, 2 females cl 13.9, 18.4 mm (NTOU M02348). Taipei City, Taipei market, 25 Oct 1984, 2 females cl 20.4, 22.3 mm (NTOU M02415); 7 May 1985, 3 cl 16.9-19.5 mm, 4 females cl 18.5-23.0 mm (NTOU M02424). Taichung City, Wuci, 16 Jan 1995, 2 males cl 17.9, 20.1 mm, 2 females cl 24.0, 29.4 mm (NTOU M02434). Changhua County, Fangyuan, 26 Nov 1992, 1 female cl 15.7 mm (NTOU M02435).— Shengang, 30 Jul 2009, 1 female cl 15.4 mm (NTOU M02436). Yunlin County, Mailiao, Station 1, 30 Nov 2010, 1 female cl 16.2 mm (NTOU M02437); Station 2, 30 Nov 2021, 1female cl 13.6 mm (NTOU M02438). Chiayi County, Budai fishing port, 2 Jul 2002, 8 males cl 21.6-25.2 mm, 6 females cl 22.2-29.9 mm (NTOU M02325). Tainan City, Anping fishing port, 24 Jun 1984, 1 male cl 20.1 mm, 1 female cl 17.5 mm (NTOU M02425). Kaohsiung City, Kaohsiung Port, Station 4, 1 Mar 1994, 2 males cl 25.6, 27.9 mm (NTOU M02439).— Neiwei market, 17 Oct 2019, 2 males cl 23.9, 24.4 mm, 2 females cl 26.2, 28.8 mm (NTOU M02326). Pingtung County, Donggang fishing port, 3 Nov 2011, 1 female cl 32.7 mm (NTOU M02327); 14 Aug 2015, 1 male cl 23.9 mm (NTOU M02328); 9 May 2018, 2 males cl 19.6, 21.3 mm (NTOU M02323); 9 May 2018, 1 male cl 20 mm (NTOU M02426). No specific locality, 1 Feb 1974, 1 male cl 24.8 mm, 1 female cl 23.2 mm (NTOU M02440); 1 Apr 1993, 1 female cl 14.7 mm (NTOU M02441); Oct 2012, 1 male cl 24.7 mm (NTOU M02442). No specific data, 3 males cl 16.1-21.8 mm, 1 female cl 19.7 mm (NTOU M02443); 3 males cl 20.7-21.6 mm, 2 females cl 20.5, 21.8 mm, 1 juvenile cl 12.8 mm (NTOU M02444); 1 male cl 20.3 mm, 2 females cl 22.5, 25.5 mm (NTOU M02445); 1 female cl 29.5 mm (NTOU M02446); 2 males cl 14.7, 15.8 mm, 4 females cl 15.1- 18.3 mm (NTOU M02447).</p> <p>Diagnosis. Rostrum with 6–10 (excluding epigastric tooth) teeth along entire dorsal border; generally straight and more or less extending to middle of distal segment of antennular peduncle. Pereiopod I generally with ischial spine distinct. In males, basial spine of pereiopod III not particular elongated, merus of pereiopod V with basal notch accompanied with long inwardly curved spiniform process and row of tubercles. Telson with minute movable lateral spinules. Male petasma with distomedian projections greatly swollen and as a huge triangular or subquadrate protuberances extending far beyond distolateral projections. Female thelycum with anterior plate bearing pair of small rounded lateral processes; coxa of pereiopod IV normal, not expanded or elongated; lateral plates with posterolateral margins highly elevated and strongly recurved inwards.</p> <p>Coloration. Body greenish greyish to dark grey and covered with dense dark dots. Antennal flagella and distal parts of uropods reddish. Large adults greyish pink and with pereiopods more or less banded with red and white, uropods red margined and with distal parts bluish to purplish.</p> <p>Distribution. Widely distributed in the Indo-West Pacific from India to Japan and Willis and Funtuna Islands, intertidal to 95 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997; Chan 1998).</p> <p>Remarks. Metapenaeus ensis is the commonest species of the genus in Taiwan and occurs all around Taiwan. This species was reported under the name M. monoceros (Fabricius, 1798) in local literatures (Maki &amp; Tsuchiya 1923; Chang 1965; Lee &amp; Yu 1977) before Miquel’s (1982) revision of the genus Metapenaeus. Adults of M. ensis can be easily distinguished by its very different shapes of gentalia and banded pereiopods. Nevertheless, its juveniles and immature specimens with undeveloped gentalia are very difficult to separate from M. affinis, M. elegans and M. moyebi.</p> </div>	http://treatment.plazi.org/id/038E04381F77FFA8FF25A98CFEB9A44D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F75FFA8FF25A963FE64A3AA.text	038E04381F75FFA8FF25A963FE64A3AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus intermedius (Kishinouye 1900)	<div><p>Metapenaeus intermedius (Kishinouye, 1900)</p> <p>(Figs. 4, 8d)</p> <p>Penaeus intermedius Kishinouye 1900: 21, 1 unnumbered textfig. (type locality: Japan).</p> <p>Metapenaeus intermedius — Miquel 1982: 96, figs. 39-40.— Yu &amp; Chan 1986: 133, 1 unnumbered fig.— Lee et al. 1999: 445.</p> <p>Material examined. Kaohsiung City, Neiwei market, 2019 Jun, 1 female cl 25.6 mm (NTOU M02329). Pingtung County, Donggang fishing port, 6 Aug 1996, 1 female cl 35.1 mm (NTOU M02330); 9 May 2018, 3 females cl 27.3-35.8 mm (NTOU M02331). Penghu County, 10 Oct 1984, 2 females cl 23.4, 25.3 mm (NTOU M02333).— Magong fishing port, 1 Apr 2013, 2 females cl 33.6, 36.5 mm (NTOU M02332).— Third fishing port, 8 Jun 2013, 1 male cl 26.9 mm, 1 female cl 30.4 mm (NTOU M02334).</p> <p>Diagnosis. Rostrum with 8–11 (excluding epigastric tooth) teeth along entire dorsal border, generally straight and more or less extending to middle of distal segment of antennular peduncle. Pereiopod I with distinct ischial spine. In males, basial spine of pereiopod III not particular elongated, merus of pereiopod V with basal notch accompanied by a carinated tubercle. Telson armed with 3 pairs of large movable lateral spines. Male petasma with distomedian projection as triangular flap and slightly overreaching tip of spiniform distolateral projection. Female thelycum with anterior plate bearing 2 elongated subquadrate protuberances; coxal projections of pereiopod IV extremely elongate, flat and extending onto lateral plates; lateral plates with posteromedian ogival boss and posterolateral margins elevated into ridges.</p> <p>Coloration. Body yellowish grey and distributed with dense dark dots. Rostrum dark brown. Antennal flagella reddish. Uropods red margined and with distal parts bluish to dark blue.</p> <p>Distribution. Indo-West Pacific from the Andaman Sea to Japan, Malaysia and Indonesia, intertidal to 130 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997; Chan 1998).</p> <p>Remarks. Metapenaeus intermedius is mainly found in southern Taiwan and Penghu. This species can be readily distinguished from the other species of the genus in Taiwan by the telson bearing three pairs of large movable lateral spines (Fig. 4e).</p> </div>	http://treatment.plazi.org/id/038E04381F75FFA8FF25A963FE64A3AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F75FFA4FF25AD58FDADA509.text	038E04381F75FFA4FF25AD58FDADA509.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus joyneri (Miers 1880)	<div><p>Metapenaeus joyneri (Miers, 1880)</p> <p>(Figs. 5, 8e)</p> <p>Penaeus joyneri Miers 1880: 458, pl. 15-figs. 8-10 (type locality: Japan).</p> <p>Metapenaeus joyneri — Maki &amp; Tsuchiya 1923: 41, pl. 6-fig. 2.— Chang 1965: 7, 3 unnumbered figs.— Lee &amp; Yu 1977: 100, fig. 70.— Miquel 1982: 99, figs. 41-42— Yu &amp; Chan 1986: 135, fig. 15C, 1 unnumbered fig.— Lee et al. 1999: 445.</p> <p>Metapenaeus joyneri formosus Lee &amp; Yu 1977: 101, fig. 71 (type locality: Taiwan).</p> <p>Material examined. Yilan County, Luodong market, 5 Mar 1985, 2 females cl 21.6, 31.1 mm (NTOU M02335). Miaoli County, 7 Nov 2005, 1 male cl 14.8 mm, 1 female cl 19.2 mm (NTOU M02336). Changhua County, Station 2, 19 Mar 2018, 1 female cl 24.3 mm (NTOU M02337). Yunlin County, Mailiao, 29 Oct 2009, 30 males cl 13.5-18.2 mm, 18 females cl 11.5-18.4 mm (NTOU M02338); Station 1, 2 Sep 2009, 3 females cl 14.2-16.9 mm (NTOU M02343); Station 2C, 29 Oct 2009, 1 female cl 24.3 mm (NTOU M02341); Station 4A, 29 Oct 2009, 1 female cl 17.9 mm (NTOU M02342); Station 4B, 22 Jan 2009, 1 female cl 8.2 mm (NTOU M02340); Station X6 17 males cl 14.6-18.6 mm, 18 females cl 14.3-20.4 mm (NTOU M02339). Chiayi County, Budai fishing port, 5 Mar 2021, 14 males cl 19.1-21.3 mm, 6 females cl 21.8-24.0 (NTOU M02416), 1 male cl 20.7 mm (NTOU M02448, neotype of Metapenaeus joyneri formosus), 2 males cl 19.4, 20.3 mm (NTOU M02449).— Dongshi fishing port, 8 May 2012, 1 juvenile cl 6.9 mm (NTOU M02450). Kaohsiung City, Singda fishing port, 24 Jul 1984, 1 male cl 13 mm, 4 females cl 16.1-18.9 mm (NTOU M02344). Pingtung County, Donggang fishing port, 5 Mar 2021, 10 females cl 22.0- 24.8 mm (NTOU M02345), 1 female cl 23.1 mm (NTOU M02451). Lianjiang County, Nangan, 26 Nov 2000, 3 females cl 21.4-22.0 mm (NTOU M02346). Kinmen County, 1 Feb 2001, 3 females cl 20.6-21.7 mm (NTOUM02347). No specific data, 1 male cl 18.6 mm, 2 females cl 19.0, 21.3 mm (NTOU M02452).</p> <p>Diagnosis. Rostrum armed with 6–8 (excluding epigastric tooth) teeth on proximal part and dorsal margin, distal 1/3 unarmed, slightly sinuous and more or less extending to tip of antennular peduncle. Pereiopod I with small ischial spine. In males, basial spine of pereiopod III very long and barbed, merus of pereiopod V without basal notch and only with minute to distinct basal tubercle. Telson with minute movable lateral spinules. Male petasma with distomedian projections very long and ribbon-like, far exceeding distolateral projections and with distal parts recurved dorsally. Female thelycum with anterior plate styliform and shaprly pointed anteriorly; coxa of pereiopod IV normal, not expanded or elongated; lateral plates semicircular, extended anteriorly and enveloping anterior plate except for elongated tip of anterior plate.</p> <p>Coloration. Body greyish white to pale green, densely covered with dark dots. Antennal flagella reddish. Uropods with margins reddish, distal parts proximally somewhat yellowish green and distally reddish. Pleopods somewhat yellowish.</p> <p>Distribution. Restricted to Japan, Korea, China and Taiwan, intertidal to about 76 m deep (Miquel 1982; Liu &amp; Zhong 1988; Pérez Farfante &amp; Kensley 1997; Kim 2012)</p> <p>Remarks. Metapenaeus joyneri can be readily distinguished from the other species of the genus in Taiwan by the rostrum extending to the tip of antennular peduncle and with the distal 1/3 unarmed (Fig. 5a). Moreover, both the pestama and thelycum have very different shapes in this species (Fig. 5b, c), and the basial spine at the pereiopod III in males are very long and barbed (Fig. 5e).</p> <p>Lee &amp; Yu (1977) noted that some specimens of this species from Donggang (= Tongkang) in southern Taiwan had the carapace relatively more glabrous and robust (i.e., the integument was harder) than the typical form, and recognized it as new subspecies, M. joyneri formosus. One holotype and 12 paratypes were identified but the morphological differences were not illustrated by Lee &amp; Yu (1977). The material reported in Lee &amp; Yu (1977) was deposited either in the Taiwan Fisheries Research Institute or NTOU, the resident institutions of Lee and Yu. An extensive search in the collections of the Taiwan Fisheries Research Institute and NTOU failed to find any specimen labeled as material of this subspecies. In the NTOU are numerous specimens of M. joyneri from Taiwan, including Donggang but almost all are of the typical carapace form, possessing many pubescent sunken areas. There are three males (NTOU M02448, NTOU M02449) and a juvenile (NTOU M02450) from Chiayi County (southwestern Taiwan just north of Donggang) that have the carapace relatively more glabrous and with pubescence mainly restricted to the grooves and carinae; characters that fit the description of M. joyneri formosus. They are, however, not from the type locality and are not types.</p> <p>There is one lot from the old collections in NTOU (NTOU M02452) without any location or collection label with three specimens (1 male and 2 females, each attached with a label “3”) that all have the carapace rather glabrous and very similar to the condition of the four naked specimens from Chiayi (NTOU M02448 - M02450). While we are confident that NTOU M02452 was collected from Taiwan, there is no precise data even that they may be the types - we just cannot be sure. No size information was given for the types of M. joyneri formosus in Lee &amp; Yu (1977) so we cannot match the present specimens with the type series. The petasma and thelycum illustrated for M. joyneri formosus by Lee &amp; Yu (1977: fig. 71A-C, G) have maximum lengths of 7.5 and 4.8 mm, respectively, according to the scales given for the figures. In the unlabeled lot NTOU M02452, the petasma of the male is 9.8 mm long and the thelycum of the two females are 2.8 and 3.8 mm long; and as such, the data sets do not agree. Moreover, the telsons of the male and one female have the tips broken while that of the other female does not possess distinct spinules, unlike the telson illustrated by Lee and Yu (1977: fig. 71J). Although we are sure that the specimens of NTOU M02452 are not those used for the illustrations of M. joyneri formosus in Lee &amp; Yu (1977), we cannot ascertain if they are the types. We do know, however, that some Taiwanese penaeid specimens identified by Lee and Yu before 1977 were not included in their 1977 report (e.g., M. ensis: NTOU M 02440 collected on 1 Feb 1974). In addition, both Lee and Yu were always very careful with the labelling of type material when the third author TYC worked with them. The registration numbers indicated in Lee &amp; Yu (1977) for the material of M. joyneri formosus are M 701, M 704 (paratypes), M 705 (holotype) and M 706 (paratype). The label “3” attached to the specimens of the NTOU M02452 lot does not match the registration number of Lee &amp; Yu’s (1977) M. joyneri formosus material. As such, it is unlikely that unlabeled NTOU M02452 are the type specimens of M. joyneri formosus.</p> <p>An attempt to sequence the barcoding gene COI of the three specimens of NTOU M02542 had failed. The COI (615 bp) of the most glabrous specimen from Chiayi (NTOU M02448, GenBank ON329768), however, is almost identical (sequence similarity 99.7%) with a densely pubescent specimen (NTOU M02451, GenBank ON329774). As none of the types of M. joyneri formosus is found or can be determined, they are now considered as lost. In order to fix the identity of M. joyneri formosus, the sequenced male with nearly glabrous carapace fitting the original description of the subspecies given by Lee &amp; Yu (1977) from Budai, Chiayi (NTOU M02448), a locality in southern Taiwan close to Donggang (type locality of M. joyneri formosus) is now selected as the neotype of M. joyneri formosus. Through this action, M. joyneri formosus, is now synonymized with the nominotypical form.</p> <p>It may also need to point out that parts of the specimens in the two lots of M. joyneri materials reported in Lee &amp; Yu (1977) are found in NTOU (Nanfang-ao, 5 Apr 1976, 5 specimens, NTOU M02453; Keelung fishing port, 9 Mar 1976, 10 specimens, NTOU M02454). However, these two lots with labels clearly mentioned that the specimens were caught by fishing boats off the Zhoushan Island at the river mouth of Changjiang River (or Yangtze River) and far from the northern Taiwan (also see Lee &amp; Yu 1977: 100). Therefore, these two lots with specimens not originated from Taiwan are excluded in this work.</p> </div>	http://treatment.plazi.org/id/038E04381F75FFA4FF25AD58FDADA509	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F79FFA4FF25A8ACFBF3A1AE.text	038E04381F79FFA4FF25A8ACFBF3A1AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus moyebi (Kishinouye 1896)	<div><p>Metapenaeus moyebi (Kishinouye, 1896)</p> <p>(Figs. 6, 8f)</p> <p>Penaeus moyebi Kishinouye 1896: 373 (type locality: Japan).</p> <p>Metapenaeus mastersii — Lee &amp; Yu 1977: 96, figs. 66-67. (non Haswell, 1879)</p> <p>Metapenaeus moyebi — Miquel 1982: 114, figs. 50-51.— Yu &amp; Chan 1986: 137.— Lee et al. 1999: 445.</p> <p>not Metapenaeus moyebi — Yu &amp; Chan 1986: 1 unnumbered fig. [= Parapenaeus sp.]</p> <p>Material examined. Yunlin County, Mailiao, 18 May 2010, 1 females cl 17.9 mm (NTOU M02349); 29 Jul 2011, 1 male cl 12.3 mm (NTOU M02350). Tainan City, Taijiang National Park, Liukong Pier, Aug-Sep 2017, 14 males cl 10.9-14.0 mm, 6 female cl 9.8-16.5 mm (NTOU M02351). Kaohsiung City, Kaohsiung port, Station 4, 1 Mar 1994, 1 female cl 21.4 mm (NTOU M02352). Pingtung County, Donggang fishing port, 7 Jan 1984, 1 female cl 22.3 mm (NTOU M02353); 9 May 2019, 1 female cl 16.8 mm (NTOU M02354).</p> <p>Diagnosis. Rostrum with 7–9 (excluding epigastric tooth) teeth along entire dorsal border, generally straight and more or less extending to middle of distal segment of antennular peduncle. Pereiopod I with ischial spine minute to absence. In males, basial spine of pereiopod III not particular elongate, merus of pereiopod V with basal notch accompanied with carinated and twisted tubercle. Telson with minute movable lateral spines. Male petasma with distomedian projections enlarged and laminose or flap-like, diverging and far overreaching distolateral projections. Female thelycum with anterior plate having anterior margin convex and bearing 3 subequal tubercles, lateral margin distinctly concave; coxal projections of pereiopod IV round plate like and often concealing anterior plate, but not extended posteriorly to lateral plates; lateral plates flattened with lateral margins convex to subquadrate, bearing distinct median longitudinal furrow at least along anterior half.</p> <p>Coloration. Body greenish grey and densely covered with dark dots. Antennal flagella reddish. Uropods yellowish green and with distal parts reddish.</p> <p>Distribution. Indo-West Pacific from India to Japan, Indonesia and Thursday Island of northern Australia, intertidal to 45 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997; Chan 1998).</p> <p>Remarks. This species was firstly reported from Taiwan by Lee &amp; Yu &amp; (1977) under the name M. mastersii. Chan &amp; Yu (1986) followed Miquel (1982) in using M. moyebi for this species but their photograph provided (Chan &amp; Yu 1986: 137 -unnumbered photo) was a species of Parapenaeus Smith, 1885. Metapenaeus moyebi is uncommon in Taiwan and found only in the southwestern coasts, though quite a few young specimens were recently collected in a national park. Amongst the local species of Metapenaeus, M. moyebi is most similar to M. elegans and their distinguishing characters can be referred to the “Remarks” of the latter species.</p> </div>	http://treatment.plazi.org/id/038E04381F79FFA4FF25A8ACFBF3A1AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F7FFFA0FF25AB9FFDBAA435.text	038E04381F7FFFA0FF25AB9FFDBAA435.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus tenuipes Kubo 1949	<div><p>Metapenaeus tenuipes Kubo, 1949</p> <p>(Fig. 7)</p> <p>Metapenaeus tenuipes Kubo 1949: 348, figs. 7R, 22B, 31G, H, 62B, B’, 74N, T, 81H, 125G, 127 (type locality: Indonesia).— Racek &amp; Dall 1965: 82, pl.7-fig.1, pl. 12-fig. 2.— Miquel 1982: 125, figs. 58-59.— Yu &amp; Chan 1986: 131, 1 unnumbered fig.— Lee et al. 1999: 445.</p> <p>Metapenaeus spinulatus Kubo 1949: 355, figs. 7T, 47G, 74M, S, 81K, 129F, 130 [type locality: Thailand].— Lee &amp; Yu 1977: 98, figs. 68-69.</p> <p>Material examined. None.</p> <p>Diagnosis. Based on descriptions of the single female collected by Lee &amp; Yu (1977). Rostrum with distinct basal crest, bearing 5 (excluding epigastric tooth) dorsal teeth restricted to basal crest; distal 1/2 unarmed; short and straight, extending to middle of second segment of antennular peduncle. Pereiopod I with ischial spine. Telson with rows of minute and 2 larger movable lateral spines. Female thelycum with anterior plate small and rounded; coxal projections of pereiopod IV round plate like and not extending to lateral plates; lateral plates large, somewhat rhombus-shape and anteriorly with a “W”-like notch, lacking median longitudinal furrow.</p> <p>Coloration. According to Miquel (1982) body generally whitish to yellowish with green dots. Uropods red margined and with distal part greyish green.</p> <p>Distribution. Western Pacific from Thailand to Indonesia, Philippines and Taiwan, intertidal to 30 m deep (Miquel 1982; Pérez Farfante &amp; Kensley 1997).</p> <p>Remarks. Metapenaeus tenuipes can be readily separated from the other species of the genus in Taiwan by having a distinct basal rostral crest and a short rostrum (Fig. 7a). No specimen of this species has been collected again from Taiwan since Lee &amp; Yu (1977) recorded a single female nearly half a century ago (1974). Although this female specimen (registration number M 750 in Lee &amp; Yu 1977) is now missing (not found in the Taiwan Fisheries Research Institute and NTOU, the affiliated institutions of Lee and Yu respectively), its identity is unmistaken from the figures provided by Lee &amp; Yu (1977: figs. 68-69). According to Lee &amp; Yu (1977), the single specimen of this species was collected from the Keelung fishing port in northern Taiwan with landing sometimes from fishing boats operated overseas. Nevertheless, M. tenuipes may just be rare in Taiwan like M. elegans which also only has one single specimen collected since 1988 despite of the recent extensive surveys of the decapod crustacean fauna in Taiwan.</p> <p>According to Racek &amp; Dall (1965) and Miquel (1982), there are marked sexual dimorphisms in M. tenuipes and with males having low basal rostral crest. This is also why Kubo (1949) named females of this species as M. tenuipes while males as M. spinulatus.</p> </div>	http://treatment.plazi.org/id/038E04381F7FFFA0FF25AB9FFDBAA435	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
038E04381F7DFFA0FF25A9BAFA61A36E.text	038E04381F7DFFA0FF25A9BAFA61A36E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metapenaeus Wood-Mason	<div><p>Key to species of Metapenaeus in Taiwan</p> <p>1. Rostrum with distinct basal crest and failed to reach distal segment of antennular peduncle.................. M. tenuipes</p> <p>-. Rostrum without distinct basal crest and more or less extending to tip of antennular peduncle......................... 2</p> <p>2. Telson with 3 pairs of large movable spines..................................................... M. intermedius</p> <p>-. Telson without distinct movable spines.................................................................... 3</p> <p>3. Rostrum unarmed at distal 1/3; males with basial spine of the pereiopod III extremely long and barbed......... M. joyneri</p> <p>-. Rostrum armed with teeth along entire dorsal border; males with basial spine of pereiopod III not elongated and simple.... 4</p> <p>4. Males with basal notch at merus of pereiopod V accompanied by a spiniform process and row of tubercles, petasma with distomedian projections very large and greatly swallow, triangular or subquadrate in shape; female thelycum with posterolateral edges of lateral plates highly elevated and strongly recurved inward....................................... M. ensis</p> <p>-. Males with basal notch at merus of pereiopod V only accompanied with a large tubercle, petasma with distomedian projections flap-like or reduced; female thelycum with margins of lateral plates flattened or if raised, not strongly curved inwards..... 5</p> <p>5. Male petasma with distomedian and distolateral projections more or less extending to same level; female thelycum with lateral plates having lateral margins raised as ridges....................................................... M. elegans</p> <p>-. Male petasma with distomedian projections considerably overreaching or falling short of distolateral projections; female thelycum with lateral plates having lateral margins not ridged...................................................... 6</p> <p>6. Male petasma with distomedian projections very large and extending far beyond distolateral projections; female thelycum with anterior margin of anterior plate convex and bearing 3 tubercles, lateral plates with median longitudinal furrow.. M. moyebi</p> <p>-. Male petasma with distomedian projections reduced, very low and far from tips of distolateral projections; female thelycum with anterior margin of anterior plate straight or concave, without tubercles, lateral plates lacking median longitudinal furrow............................................................................................ M. affinis</p></div> 	http://treatment.plazi.org/id/038E04381F7DFFA0FF25A9BAFA61A36E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsu, Yen-Cheng;Chen, Chien-Lin;Chan, Tin-Yam	Hsu, Yen-Cheng, Chen, Chien-Lin, Chan, Tin-Yam (2022): On the penaeid shrimps of the genus Metapenaeus Wood-Mason in Wood-Mason & Alcock, 1891 (Crustacea, Decapoda, Penaeidae) of Taiwan. Zootaxa 5169 (4): 331-346, DOI: https://doi.org/10.11646/zootaxa.5169.4.3
