identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
C822F213FFCC2470CB88A20E7003229A.text	C822F213FFCC2470CB88A20E7003229A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisini Reitter 1882	<div><p>Tribe Batrisini Reitter, 1882</p> <p>Chinese common name: 毛唇à甲⁂</p> <p>Diagnostic characters, an introduction, and a discussion of the tribe can be found in Nomura (1991), Chandler (2001), and Löbl &amp; Kurbatov (2001). Following Jeannel (1954a), Leleup (1970, 1971, 1973) and Chandler (2001), all taxa treated in this paper belong to Batrisina. The other three subtribes, e. g., Ambicocerina Leleup, Leupeliina Jeannel, and Stilipalpina Jeannel, are confined to Africa. Kurbatov (2007) recently found that a broad range of batrisine genera uniquely have four large specialized setae at the middle of the anterior margin of the labrum, in contrast to the presence of zero to two such setae for the other pselaphine supertribes. This is by far the only unambiguous evidence suggesting a monophyletic Batrisitae. In total there are 80 species classified in 13 genera found in Tibet (including the new taxa described in this paper). The genera may be distinguished by using the following key.</p> </div>	http://treatment.plazi.org/id/C822F213FFCC2470CB88A20E7003229A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFCC2470CB88A3FE7721254B.text	C822F213FFCC2470CB88A3FE7721254B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pselaphinae Latreille 1802	<div><p>Subfamily Pselaphinae Latreille, 1802</p> <p>Supertribe Batrisitae Reitter, 1882</p></div> 	http://treatment.plazi.org/id/C822F213FFCC2470CB88A3FE7721254B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFCC247FCB88A45B70FB2417.text	C822F213FFCC247FCB88A45B70FB2417.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisini Reitter 1882	<div><p>Key to Tibetan genera of Batrisini</p> <p>1 Whole dorsal surface of body bears coarse punctation; each elytron with four basal foveae; aedeagus simple, plate-like (Figs 52–61)............................................................................... Sathytes Westwood</p> <p>- Dorsal surface of body finely punctate to glabrous, or only parts of body coarsely punctate; each elytron with one to three basal foveae; aedeagus usually composed of median and dorsal lobes................................................. 2</p> <p>2 Each elytron with two basal foveae; pronotum with transverse antebasal sulcus connecting lateral antebasal foveae; aedeagus with articulated dorsal lobe............................................................................. 3</p> <p>- Each elytron with one or three basal foveae; pronotum lacking transverse antebasal sulcus; aedeagus with partially fixed dorsal lobe................................................................................................ 6</p> <p>3 Tergite 1 (IV) with pair of oblique inner marginal carinae (Fig. 76).................................. Trisinus Raffray</p> <p>- Tergite 1 (IV) lacking inner marginal carinae............................................................... 4</p> <p>4 Aedeagus with small, constricted basal capsule (Fig. 29)..................................... Batriscenaulax Jeannel</p> <p>- Aedeagus with large or extended basal capsule.............................................................. 5</p> <p>5 Scape lacking inner glandular structure at apicolateral margin; dorsal lobe of aedeagus erect (Figs 2–28)................................................................................................... Arthromelodes Jeannel</p> <p>- Scape with inner glandular structure at apicolateral margin; dorsal lobe of aedeagus recumbent or strongly bent at base (Figs 30–37)................................................................................ Batrisiella Raffray</p> <p>6 Head and pronotum strongly transverse; each elytron with one basal fovea (Yin &amp; Li 2015a: figs 1, 3)..................................................................................................... Hingstoniella Jeannel</p> <p>- Head at most moderately transverse; pronotum usually with length and width subequal; each elytron with three basal foveae 7</p> <p>7 Head lacking carina or sulcus; antennomeres moniliform; pronotum lacking discal or lateral longitudinal sulcus/carina..... 8</p> <p>- Head carinate or sulcate; antennomeres non-moniliform; pronotum with discal or lateral longitudinal sulcus/carina........ 9</p> <p>8 Body glabrous; head sub-trapezoidal, with trichome at posterolateral margin (Yin et al. 2012b: fig. 1; Yin &amp; Li 2013a: fig. 1)....................................................................................... Tangius Yin &amp; Li</p> <p>- Body pubescent; head sub-rectangular, lacking trichome at posterolateral margin (Yin 2021: fig. 1)....................................................................................................... Myrmicophila Yin &amp; Li</p> <p>9 Pronotum with pairs of sinuate discal and lateral longitudinal sulci; metatibia of male with long pencil of setae at apex (Figs 38, 39)................................................................................... Batrisodes Reitter</p> <p>- Pronotum at most with one pair of longitudinal sulci, often with pair of discal longitudinal carinae; metatibia lacking or with short tuft of setae at apex.............................................................................. 10</p> <p>10 Pronotum with pairs of anterolateral and antero-hypomeral foveae (Fig. 40E).................. Coryphomimus gen. nov.</p> <p>- Pronotum lacking anterolateral or antero-hypomeral foveae................................................... 11</p> <p>11 Elytra with inner two basal foveae close, disc coarsely punctate (Figs 43–45).................... Coryphomodes Jeannel</p> <p>- Elytra with inner two basal foveae clearly separated, disc finely punctate........................................ 12</p> <p>12 Body length less than 1.7 mm; vertex strongly convex; lacking lateral frontal fovea; pronotum with short discal longitudinal carina present only at basal 1/2; lacking external male sexual characters (Figs 46–48)................. Gyrongita gen. nov.</p> <p>- Body length usually no less than 2.0 mm; lateral frontal fovea present; vertex weakly to moderately convex; pronotum often with complete discal carina; male sexual characters usually present on antennae or legs, rarely on head (Figs 62–74)............................................................................................ Tribasodites Jeannel</p></div> 	http://treatment.plazi.org/id/C822F213FFCC247FCB88A45B70FB2417	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFC3247ECB88A3D670FB2130.text	C822F213FFC3247ECB88A3D670FB2130.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes Jeannel 1954	<div><p>Arthromelodes Jeannel, 1954</p> <p>Chinese common name: njà甲¤</p> <p>Arthromelodes Jeannel, 1954b: 247; Jeannel 1957 (correction of the locality of the type species); Nomura 1991 (description, new generic synonym, key to Japanese species); Yin 2018c (first country record in China). Type species: Arthromelodes cariei Jeannel, 1954b: 248 (original designation).</p> <p>= Pseudobatriscenus Jeannel, 1958: 66. Type species: Batrisocenus dilatatus Raffray, 1909: 25 (original designation). Synonymized by Nomura 1991: 337.</p> <p>= Plocamarthrus Jeannel, 1960: 423. Type species: Plocamarthrus championi Jeannel, 1960: 423 (original designation). syn. nov.</p> <p>The genus Arthromelodes previously comprised 21 species distributed in Japan (Nomura 1991), China (Yin 2018c), Vietnam (Nomura &amp; Pham 2019), Laos (Jeannel 1957), and Myanmar (Nomura &amp; Aung 2020), and is strongly diversified in East to Southeast Asia and along the Himalaya. There is no determined synapomorphy that links all included species, but members share a large basal capsule and erect dorsal lobe of the aedeagus. Plocamarthrus with four species distributed in northern India has a more elongate ventral stalk of the median lobe, and an elongate and apically strongly curved dorsal lobe of the aedeagus. Otherwise, the genus cannot be effectively separated from Arthromelodes by external morphology. Here, Arthromelodes is proposed as a junior synonym of Plocamarthrus, with the former being the older name, syn. nov. The synonymy creates the following new combinations: Arthromelodes championi (Jeannel, 1960), comb. nov., A. cameroni (Jeannel, 1960), comb. nov., A. planiceps (Jeannel, 1960), comb. nov., and A. abdominalis (Jeannel, 1960), comb. nov. Nomura (1991: 338) was the first to point out the morphological similarity between these two genera, and one of the new species described here is named in his honor.</p> <p>Twenty-six new species from Tibet are described in this paper, and A. championi (Jeannel) is recorded from China (in Tibet) for the first time. The 27 Tibetan species are keyed below.</p> <p>Key to Tibetan species of Arthromelodes (male)</p> <p>1 Sexual characters present on antenna or tergites............................................................. 2</p> <p>- Antenna or tergites lacking sexual characters.............................................................. 23</p> <p>2 Sexual characters present on antenna...................................................................... 3</p> <p>- Sexual characters present on tergites...................................................................... 7</p> <p>3 Antennomeres 9 and 10 with dense, short setae.............................................................. 4</p> <p>- Antennomeres 9 and 10 normally setose................................................................... 5</p> <p>4 Vertex with distinct median longitudinal sulcus (Fig. 7B); antennomere 7 small, unmodified; metaventrite densely setose (Fig. 9D).............................................................................. A. crassicornis sp. nov.</p> <p>- Vertex lacking median longitudinal sulcus (Fig. 18B); antennomere 7 enlarged (Fig. 18C); metaventrite lacking dense setae......................................................................................... A. lebus sp. nov.</p> <p>5 Antennal modification presents on antennomere 11 (Fig. 8C); mesotibia with markedly large apical spine (Fig. 8E)................................................................................................ A. cona sp. nov.</p> <p>- Antennal modification present on antennomere 9; mesotibia lacking or with small apical spine........................ 6</p> <p>6 Antennomere 7 mesally expanded (Fig. 6C); elytra truncate at bases (Fig. 6A); mesotibia with triangular apical spine (Fig. 6D)............................................................................... A. championi (Jeannel)</p> <p>- Antennomere 7 apically projecting on anterolateral margin (Fig. 25C); elytra constricted at bases (Fig. 25A); mesotibia lacking apical spine.......................................................................... A. speciosus sp. nov.</p> <p>7 Mesofemur with protuberance or spine on ventral margin...................................................... 8</p> <p>- Mesofemur lacking protuberance or spine on ventral margin................................................... 9</p> <p>8 Elytra constricted at bases (Fig. 13A); ventral spine of mesotrochanter asetose (Fig. 13C); mesotibia with small apical tubercle; tergites 1–4 (IV–VII) modified (Fig. 13D)................................................... A. gyamda sp. nov.</p> <p>- Elytra truncate at bases (Fig. 21A); ventral spine of mesotrochanter setose (Fig. 21C); mesotibia with large, curved apical spine (Fig. 21D); only tergite 1 (IV) modified (Fig. 21E)............................................. A. monba sp. nov.</p> <p>9 Metafemur distinctly thickened distally................................................................... 10</p> <p>- Metafemur normally formed, not distinctly thickened distally................................................. 11</p> <p>10 Vertex with long, distinct median carina (Fig. 11B); abdomen slightly longer than wide due to elongate tergite 1 (IV) (Fig. 11A); tergite 1 (IV) lacking setose patches lateral to central cavity (Fig. 11D).......................... A. cylindricus sp. nov.</p> <p>- Vertex with short, indistinct median carina (Fig. 20B); abdomen much wider than long (Fig. 20A); tergite 1 (IV) with setose patches lateral to central cavity (Fig. 20D)................................................... A. markam sp. nov.</p> <p>11 Tergite 1 (IV) lacking central cavity or impression.......................................................... 12</p> <p>- Tergite 1 (IV) with central cavity........................................................................ 17</p> <p>12 Body bicolored, with black head, pronotum and abdomen black, elytra reddish-brown (Fig. 5A); tergite 1 (IV) with median projection at posterior margin (Fig. 5D)...................................................... A. bicolor sp. nov.</p> <p>- Body unicolorous, reddish-brown; tergite 1 (IV) lacking median projection at posterior margin....................... 13</p> <p>13 Tergite 5 (VIII) modified, with broad and deep basal cavity (Fig. 24D)........................ A. songxiaobini sp. nov.</p> <p>- Tergite 5 (VIII) simple, unmodified...................................................................... 14</p> <p>14 Protibia with angulate protuberance at lateral margin near apex (Fig. 28C)..................... A. zhentangensis sp. nov.</p> <p>- Protibia lacking protuberance at lateral margin.............................................................. 15</p> <p>15 Mesotrochanter with long bunch of setae on ventral margin (Fig. 16C); mesotibia with tiny tubercle at apex (Fig. 16D)......................................................................................... A. langjicuo sp. nov.</p> <p>- Mesotrochanter with short setose protuberance on ventral margin; mesotibia with moderately large tubercle at apex...... 16</p> <p>16 Median projection of tergite 1 (IV) with two widely separated posterolateral angles (Fig. 2E)........... A. alpitorus sp. nov.</p> <p>- Median projection of tergite 1 (IV) with single posterior or narrowly separated posterolateral angles (Fig. 26E)................................................................................................... A. torus sp. nov.</p> <p>17 Lateral margin of tergite 1 (IV) angulate (Fig. 3D)............................................ A. angulatus sp. nov.</p> <p>- Lateral margin of tergite 1 (IV) smooth................................................................... 18</p> <p>18 Pronotum with exceptionally elongate and erect setae at lateral margin.......................................... 19</p> <p>- Pronotum normally setose, lacking exceptionally elongate setae at lateral margin.................................. 20</p> <p>19 Body length over 2.6 mm; vertex with complete reversed U-shaped sulcus connecting foveae (Fig. 12B); antennomeres distinctly elongate (Fig. 12A).............................................................. A. flosculus sp. nov.</p> <p>- Body length less than 1.8 mm; vertexal sulcus incomplete (Fig. 19B); antennomeres slightly elongate (Fig. 19A).................................................................................................. A. lotus sp. nov.</p> <p>20 Protibia distinctly expanded (Fig. 15A)........................................................ A. lage sp. nov.</p> <p>- Protibia slender, not expanded.......................................................................... 21</p> <p>21 Lateral and posterior margins of elytra densely setose (Fig. 14A); antennomeres distinctly elongate (Fig. 14A); central cavity of tergite 1 (IV) exceptionally large (Fig. 14E)............................................... A. intricatus sp. nov.</p> <p>- Lateral and posterior margins of elytra lacking dense setae; antennomeres moderately elongate; central cavity of tergite 1 (IV) relatively much smaller............................................................................... 22</p> <p>22 Mesotibia with tiny apical tubercle; tergite 1 (IV) with single central cavity, lacking tubercle or projection (Fig. 4D)............................................................................................. A. aniqiao sp. nov.</p> <p>- Mesotibia with distinct apical spur; tergite 1 (IV) with median projection anterior to and large tubercle posterior to central cavity (Fig. 7F)......................................................................... A. complexus sp. nov.</p> <p>23 Vertex with dense, exceptionally long setae (Fig. 10B)......................................... A. criniger sp. nov.</p> <p>- Vertex normally setose, lacking dense, long setae........................................................... 24</p> <p>24 Body bicolored (Fig. 22A); protibia strongly sinuate (Fig. 22D)............................... A. nepaeformis sp. nov.</p> <p>- Body unicolorous; protibia slightly curved or nearly straight.................................................. 25</p> <p>25 Lateral margin of pronotum normally rounded, not expanded (Fig. 23A); mesotibia moderately expanded mesally at basal fourth, with small apical projection (Fig. 23C)................................................ A. nomurai sp. nov.</p> <p>- Lateral margin of pronotum moderately to strongly expanded; mesotibia not expanded mesally at basal fourth, with large apical spine.............................................................................................. 26</p> <p>26 Lateral margin of pronotum strongly expanded (Fig. 17B); apical spine of mesotibia curved (Fig. 17D); ventral stalk of median lobe moderately broad at apex in dorsoventral view (Fig. 17F, H)................................ A. latithorax sp. nov.</p> <p>- Lateral margin of pronotum moderately expanded (Fig. 27B); apical spine of mesotibia straight (Fig. 27D); ventral stalk of median lobe strongly broad at apex in dorsoventral view (Fig. 27G).............................. A. zhangmu sp. nov.</p></div> 	http://treatment.plazi.org/id/C822F213FFC3247ECB88A3D670FB2130	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFC1247BCB88A1DB719725DE.text	C822F213FFC1247BCB88A1DB719725DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes alpitorus Yin 2022	<div><p>Arthromelodes alpitorus sp. nov.</p> <p>(Fig. 2)</p> <p>Chinese common name: Ṉ瘤únjà甲</p> <p>Type material (60 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, pass to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.49638&amp;materialsCitation.latitude=27.916666" title="Search Plazi for locations around (long 87.49638/lat 27.916666)">Zhêntang Town</a>, 27°55’0”N, 87°29’47”E, 3500–3650 m, 26.vi.2021, Peng, Yin &amp; Zhang, ƱẪU喀NJŤĠĿƋ沟Ḃ山杜ḁff ’ (SNUC). PARATYPES: CHINA: 21 ♂♂, 20 ♀♀, same data as that of holotype; 11 ♂♂, 7 ♀♀, same data as that of holotype except ‘3650-4020, 27.vi.2021 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.8–2.9 mm. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 smallest, 11 as long as 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to more than apical 1/3 of elytral length. Fore and hind legs simple, mesotrochanter with setose tuft on ventral margin, mesotibia with large, curved apical spine. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 with broad, bifid protuberance at middle. Aedeagus strongly asymmetric, median lobe with large basal capsule and elongately oval foramen, ventral stalk broad and dilated at apex, dorsal lobe elongate, forked at apex, parameres reduced and forming single membranous structure. Female. Body length approximately 2.7–2.8 mm, legs and abdomen lacking modifications, genitalia as in Fig. 2I.</p> <p>Description. Male. Body (Fig. 2A) length 2.80–2.88 mm; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 2B) subtruncate at base, approximately as long as wide, length 0.58–0.59 mm, width across eyes 0.56–0.58 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with distinct, curved transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercle moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina present, additional incomplete carina present below posteroventral articulation of mandible. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, median carina extending before fovea anteriorly to mouthpart, gradually thickened anteriorly. Eyes moderately prominent, composed of approximately 26 ommatidia. Antenna 1.46 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 3 and 8 of same length, shortest, 8 much shorter and slightly narrower than 7, 9 longer but narrower than 10, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 2B) slightly longer than wide, length 0.65–0.66 mm, width 0.62 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus much shorter than lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.85–0.87 mm, width 0.94–0.96 mm, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea posteriorly to more than apical 1/3 of elytral length; subhumeral fovea absent, thin marginal stria extending posteriorly from posterior to middle to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with long, narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 2C) with short setose tuft on ventral margin, mesotibia (Fig. 2D) with thick, curved spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.81–0.88 mm, width 0.85–0.86 mm; with modified tergite 1 (IV). Tergite 1 (Fig. 2E) longer than tergites 2–4 (V–VII) combined, with broad, strongly raised projection at middle, projection with acute posterolateral angles in fully-developed individuals, less so in few other specimens, area posterior to projection moderately raised along middle, basal impression broken by mediobasal and basolateral foveae, with pair of short discal carinae, with incomplete outer marginal carina; tergites 2 and 3 (V and VI) subequal in length and 4 (VII) slightly longer than 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of short lateral carinae; midlength of sternite 2 (IV) as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin roundly emarginate at middle, sternite 7 (IX) (Fig. 2F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 2G, H) 0.45 mm long, strongly asymmetric; median lobe with large basal capsule and elongately oval foramen, basoventral projection short, ventral stalk broad and protruding, in lateral view strongly dilated at apex; dorsal lobe obliquely erect, apically forked into two lobes; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking spines/projections or modifications; each compound eye composed of approximately 22 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.66–2.80 mm; length/width of head 0.55–0.57/ 0.56–0.57 mm, pronotum 0.62–0.63/ 0.58–0.60 mm, elytra 0.77–0.79/ 0.93–0.94 mm; abdomen 0.80–0.83/ 0.87–0.88 mm; length of antenna 1.31–1.35 mm; maximum width of genitalia (Fig. 2I) 0.29 mm.</p> <p>Comparative notes. Arthromelodes alpitorus is morphologically most similar to A. torus by sharing most external characters, as well as a similar male tergal modification and form of the aedeagus. The new species can be separated by the larger body size (2.80–2.88 mm vs. 2.64 mm in male; 2.66–2.80 mm vs 2.53–2.61 mm in female), a much shorter median longitudinal sulcus of the pronotum, the male having thicker and more strongly curved spine at the apex of mesotibia, the median projection of tergite 1 (IV) with acute posterolateral angles (with single, acute posterior corner in A. torus), and area posterior to the projection raised along middle (flat in A. torus), and a more expanded apex of the median lobe of the aedeagus. It is worth noting that A. alpitorus inhabits Rhododendron forests experiencing low temperatures at an altitude of 3500–3650 m, which is much colder than the mixed forests inhabited by A. torus at 3000 m. It appears that Arthromelodes alpitorus is well-isolated from A. torus by high altitude geographical barriers.</p> <p>On the other hand, this species is also close to A. angulatus, A. aniqiao, A. complexus, A. lage and A. langjicuo, sharing with them a setose male mesotrochanter, as well as a similar general form of the aedeagus, but A. alpitorus can be otherwise readily separated by the unique form of the male tergal modification and the aedeagus.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 77A, 93A).</p> <p>Etymology. The species name is combined from ‘ Alpînus (alpine)’ and ‘ torus (swelling, protuberance)’, indicating a high-altitude distribution as well as the morphological similarity of this species to A. torus.</p> </div>	http://treatment.plazi.org/id/C822F213FFC1247BCB88A1DB719725DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFC72479CB88A29E76572206.text	C822F213FFC72479CB88A29E76572206.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes angulatus Yin 2022	<div><p>Arthromelodes angulatus sp. nov.</p> <p>(Figs 3, 77A, 97C)</p> <p>Chinese common name: Pṅnjà甲</p> <p>Type material (13 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, 2190 m, 2019.vii.15–viii.01, FIT, Z.-W. Yin leg., [ƱẪffżḦËƖǎ]’ (SNUC). PARATYPES: CHINA: 1 ♂, same data as that of holotype; 1 ♂, 1 ♀, same locality and collector, except ‘vii.17, leaf litter’; 4 ♂♂, 3 ♀♀, same locality, except ‘ 30°0’35”N, 94°57’58”E, 2180 m, 10.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg. ’; 2 ♂♂, same data as precedent, except collected by ‘FIT’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.5 mm. Head sub-rounded at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae large and asetose; antenna long, antennomeres each elongate, lacking modifications. Discal stria of elytron extending to apical 1/5 of elytral length. Fore and hind legs simple, mesotrochanter with small tubercle on ventral margin, mesotibia with tiny apical spur. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) posterolaterally expanded, area mesal to expansion impressed, with transverse central cavity. Aedeagus strongly asymmetric, ventral stalk strongly protruding apically, apex in dorsal view expanded, dorsal lobe strongly curved at apex, parameres reduced and forming single membranous structure. Female. Body length 2.45 mm; mesotibia lacking apical spur, tergite 1 (IV) lacking modifications, genitalia as in Fig. 1H.</p> <p>Description. Male. Body (Fig. 3A) length 2.52–2.53 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 3B) sub-rounded at base, slightly broader than long, length 0.50–0.52 mm, width across eyes 0.54 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles weakly raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina fusing with ventral margin of antennal socket. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 32 large ommatidia. Antenna lacking distinct club and modification, length 1.17–1.21 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 3B) slightly longer than wide, length 0.53–0.56 mm, width 0.49–0.51 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.72–0.78 mm, width 0.86–0.90 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; humerus slightly prominent; discal stria extending from outer basal fovea to apical 1/5 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from middle to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 3C) with setose ventral tubercle thin at base and dilated apically, mesotibia with tiny tubercle at apex.</p> <p>Abdomen narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.64–0.79 mm, width 0.80–0.82 mm. Tergite 1 (IV) (Fig. 3D) strongly modified, longer than tergites 2–4 (V–VII) combined; basolateral margin angularly expanded laterally, area mesal to expansion broadly impressed, with broad, transverse median cavity, anterior margin of cavity projecting at middle, densely setose lateral to projection, posterior margin broadly projecting, with X-shaped protrusion at middle; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with very short triangular discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of short lateral carinae; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 3E) with weakly sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 3F, G) 0.33 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk strongly protruding and dilated at apex in dorsal view; dorsal lobe elongate and strongly curved, narrowing apically; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and tergite 1 (IV) lacking modifications; each compound eye composed of approximately 22 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.45 mm; length/width of head 0.52–0.53/ 0.52–0.54 mm, pronotum 0.51–0.54/ 0.49 mm, elytra 0.59–0.68/ 0.82–0.90 mm; abdomen 0.70–0.72/ 0.82–0.84 mm; length of antenna 1.08–1.13 mm; maximum width of genitalia (Fig. 3H) 0.28 mm.</p> <p>Comparative notes. This species is most similar to A. aniqiao by sharing a similar structure of the male abdominal modifications and the aedeagus, but can be readily separated from the latter by the distinct lateral angulations of the abdomen and a much longer median lobe of the aedeagus. Arthromelodes lage also has the abdominal modifications at a similar location, but lacks lateral angulations of the abdomen, the protibiae are expanded, and the ventral stalk of the aedeagus is relatively much shorter.</p> <p>Distribution. Chagyib District (巴Ù区) (Nyingchi), Xizang, SW China (Figs 77A, 97C).</p> <p>Etymology. The specific name ‘ angulatus, - a, - um ’ is a Latin adjective meaning ‘cornered, angular’, referring to the laterally expanded male tergite 1 (IV) of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FFC72479CB88A29E76572206	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFC52467CB88A5E971DB25FA.text	C822F213FFC52467CB88A5E971DB25FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes aniqiao Yin 2022	<div><p>Arthromelodes aniqiao sp. nov.</p> <p>(Figs 4, 77A, 98E, F)</p> <p>Chinese common name: Nj尼桥njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang A. R., Nyingchi City, Mêdog County, A’niqiao, alt 1100 m, 16–18.viii.2005, L. Tang leg. [ƱẪffżẸṘNj尼桥]’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.5 mm. Head sub-rounded at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae large and asetose; antenna long, antennomeres each elongate, lacking modifications. Discal stria of elytron extending to apical 2/5 of elytral length. Fore and hind legs simple, mesotrochanter with small trichome on ventral margin, mesotibia with tiny apical spur. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) with transverse central cavity and impressed lateral areas. Aedeagus strongly asymmetric, ventral stalk protruding apically, apex in dorsal view truncate, dorsal lobe strongly curved at apex, parameres reduced and forming single semi-sclerotized structure.</p> <p>Description. Male. Body (Fig. 4A) length 2.33 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 4B) sub-rounded at base, slightly broader than long, length 0.52 mm, width across eyes 0.55 m; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles weakly raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 32 large ommatidia. Antenna with loosely formed club, lacking modifications, length 1.20 m; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, much shorter than 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 4B) approximately as long as wide, length 0.54 mm, width 0.52 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.71 mm, width 0.80 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; humerus barely prominent; discal stria extending from outer basal fovea to apical 2/5 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from subbase to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 4C) with small trichome on ventral margin, mesotibia with tiny tubercle at apex.</p> <p>Abdomen narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.68 mm, width 0.76 mm. Tergite 1 (IV) (Fig. 4D) strongly modified, longer than tergites 2–4 (V–VII) combined; with broad, transverse central cavity, anterior margin of cavity broadly projecting at middle, densely setose lateral to projection, posterior margin with X-shaped protrusion at middle and two lateral projections, areas lateral to cavity broadly impressed, with smooth surface; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with very short triangular discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of short lateral carinae; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 4E) with moderately sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 4F, G) 0.31 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk protruding and in dorsal view truncate at apex; dorsal lobe elongate and strongly curved, narrowing both basally and apically; parameres reduced to single semi-membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species is most similar to A. angulatus by sharing a similar structure of the male abdominal modifications and the aedeagus. These two species can be separated primarily by the lack of distinct lateral expansions of the abdomen, and the relatively much shorter ventral stalk of the aedeagus of A. aniqiao. Arthromelodes aniqiao is also similar to A. lage, but differs by the slender male protibiae, and the much shorter and less protruding ventral stalk of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Fig. 77A, 98E, F).</p> <p>Etymology. The specific name is taken from the type locality of this species, i. e., Aniqiao.</p></div> 	http://treatment.plazi.org/id/C822F213FFC52467CB88A5E971DB25FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFDB2465CB88A2BA70E2243E.text	C822F213FFDB2465CB88A2BA70E2243E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes bicolor Yin 2022	<div><p>Arthromelodes bicolor sp. nov.</p> <p>(Figs 5, 77B, 98C, D)</p> <p>Chinese common name: ųDznjà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang A. R., Mêdog Co., Hanmi (汗ṁ), alt. 2200 m, 19.viii.2005, TANG Liang leg.’ (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.6 mm, head, pronotum and abdomen much darker in color than elytra. Head sub-rounded at base; vertex with transverse sinuate sulcus between antennal tubercles and long mediobasal carina, vertexal foveae large and asetose; antenna long, antennomeres each elongate, lacking modifications. Discal stria of elytron extending to apical 4/5 of elytral length. Fore and hind legs simple, mesotibia with large apical tubercle. Abdomen with tergite 1 (IV) longer than tergites 2–5 (V–VIII) combined; tergite 1 (IV) with distinct median projections near posterior margin. Aedeagus strongly asymmetric, ventral stalk broad in dorsal view, dorsal lobe strongly curved and pointed at apex, parameres reduced and forming single membranous structure.</p> <p>Description. [The holotype is a squashed specimen with part of the elytra and abdomen broken] Male. Body (Fig. 5A) length 2.64 mm; head dark reddish-brown, pronotum and abdomen blackish-brown, elytra reddish-brown, tarsi and mouthparts light reddish-brown. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 5B) sub-rounded at base, approximately as broad as long, length 0.56 mm, width across eyes 0.57 mm; vertex finely punctate at center, roughly punctate posterior to antennal tubercles, with large, asetose vertexal foveae (dorsal tentorial pits), with sinuate transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles weakly raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with sparse moderately large punctures, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 large ommatidia. Antenna lacking distinct club and modification, length 1.29 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, slighter longer than 9 and 10 combined, fusiform.</p> <p>Pronotum (Fig. 5B) slightly broader than long, length 0.56 mm, width 0.59 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.81 mm, width 0.94 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; humerus slightly prominent; discal stria extending from outer basal fovea to apical 4/5 of elytral length; subhumeral fovea absent, marginal stria extending from middle to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotibia (Fig. 5C) with large tubercle at apex.</p> <p>Abdomen narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.58 mm, width 0.86 mm. Tergite 1 (IV) (Fig. 5D) longest, longer than tergites 2–5 (V–VIII) combined; with broad median tubercle and one much longer but thinner projection just posterior to tubercle at posterior margin; setose basal sulcus separated by mediobasal and one pair of basolateral sockets, with pair of short discal carinae; tergites 2–5 (V–VIII) tightly compressed, tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of long lateral carinae; midlength of sternite 2 (IV) longer than sternites 3–5 (V–VII) combined, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 5E) with weakly sclerotized, round apical part and membranous basal part.</p> <p>Aedeagus (Fig. 5F, G) 0.43 mm long, strongly asymmetric; median lobe with large basal capsule and triangular foramen, basoventral projection short, ventral stalk broad in dorsal view and expanded at apex; dorsal lobe elongate and strongly curved, narrowing apically and pointed at apex; parameres reduced to single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species can be readily recognized and separated from all other members of the genus by the large and bicolored body, as well as the tergite 1 (IV) with a median tubercle and a projection at the posterior margin and the form of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 77B, 98C, D).</p> <p>Etymology. The specific name ‘ bicolor (having two colors)’ refers to the bicolored body of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFDB2465CB88A2BA70E2243E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFD92463CB88A3FE768C229B.text	C822F213FFD92463CB88A3FE768C229B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes championi (Jeannel 1960) Yin 2022	<div><p>Arthromelodes championi (Jeannel, 1960), comb. nov.</p> <p>(Figs 6, 77B, 88A, B, 89, 90, 93D–G)</p> <p>Chinese common name: K氏njà甲</p> <p>Plocamarthrus championi Jeannel, 1960: 423; figs 29, 30. Type locality: United Provinces. Kumaon: Chakrata div., Deoban, Dodoro Khud, Khedar Khud, alt. 2,500 m.</p> <p>Additional material examined (56 exx.). 5 ♂♂, 26 ♀♀, ‘ China: Xizang, Gyirong County, Gyirong Valley, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.346115&amp;materialsCitation.latitude=28.359722" title="Search Plazi for locations around (long 85.346115/lat 28.359722)">Jifu Vill.</a>, 28°21’35”N, 85°20’46”E, 2600–2700 m, 19.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉 PW吉ė村ṉữ山ě; 1 ♂, same data as precedents, except ‘slope nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32806&amp;materialsCitation.latitude=28.376945" title="Search Plazi for locations around (long 85.32806/lat 28.376945)">Jifu Vill.</a>, 28°22’37”N, 85°19’41”E, 2400- 2700 m, 22.vi.2021, ƱẪU喀NJ吉PW吉ė村ṉữ山坡 ’; 7 ♂♂, 11 ♀♀, same collectors and also from Gyirong Valley, except ‘pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.263885&amp;materialsCitation.latitude=28.429722" title="Search Plazi for locations around (long 85.263885/lat 28.429722)">Ru Kupuqiong</a>, 28°25’47”N, 85°15’50”E, 3000 m, 20.vi.2021, ƱẪU喀NJ吉PWả库 âAEṉữ ’; 1 ♂, ‘ China: Xizang, Dinggyê County, Zhêntang Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Xiuxiongma Vill.</a>, 27°54’11”N, 87°22’42”E, 2700-3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJŤĠ县ĿƋWšŏ玛村 ’; 1 ♂, 3 ♀♀, same data as precedent, except ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.44972&amp;materialsCitation.latitude=27.857224" title="Search Plazi for locations around (long 87.44972/lat 27.857224)">Nadang Vill.</a>, 27°51’26”N, 87°26’59”E, 2500 m, 28.vi.2021, Yin &amp; Zhang, ƱẪŤ Ġ县ĿƋWḄ当村 ’; 1 ♂, same data as precedent, except ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.1 mm. Head rounded at base; vertex with deep transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna moderately elongate, antennomeres each sub-moniliform to slightly elongate, antennomeres 7 and 8 moderately enlarged, 9 strongly expanded mesally. Discal stria of elytron extending posteriorly to more than apical 1/3 of elytral length. Fore and hind legs simple, mesotrochanter with acute spine on ventral margin, mesotibia with large, subtriangular apical spine. Abdomen with large Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, simple. Aedeagus strongly asymmetric, ventral stalk in dorsal view broad through length, strongly curved leftwards in ventral view, dorsal lobe thin and elongate, strongly curved downwards at apical part, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.0 mm, antenna and legs lacking modifications, genitalia as in Fig. 6H.</p> <p>Redescription (based on populations of China). Male. Body (Fig. 6A) length 2.17–2.19 mm; color reddishbrown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 6B) rounded at base, approximately as long as wide, length 0.45–0.46 mm, width across eyes 0.52 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), with deep, slightly curved transverse sulcus between antennal tubercles, distinct mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 moderately large ommatidia. Antenna 1.03 mm long, with moderately enlarged antennomeres 7–8 and modified antennomere 9 (Fig. 6C); antennomere 1 thick, subcylindrical, 2–6 each elongate, successively shorter, 7 much larger than 6 and slightly larger than 8, 9 strongly expanded at mesal margin, 10 as long as wide, sub-trapezoidal, 11 largest, simple, as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 6B) approximately as long as wide, length 0.51–0.52 mm, width 0.51 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus slightly shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.67–0.68 mm, width 0.84–0.85 mm; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humerus angularly prominent; discal stria extending from outer basal fovea to more than apical 1/3 of elytral length; subhumeral fovea small, marginal stria extending posteriorly from fovea to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae in transverse opening, broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle. Longitudinal marginal ridge of meso- and metaventrite complete.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter with acute ventral spine, mesotibia (Fig. 6D) with large, sub-triangular, blade-like spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.60–0.61 mm, width 0.76 mm; lacking modifications. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) shorter than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral foveae, lacking lateral carinae; midlength of sternite 2 (IV) slightly longer than sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 6E) slightly oval, moderately sclerotized, with scattered long setae along apical margin.</p> <p>Aedeagus (Fig. 6F, G) 0.42 mm long, strongly asymmetric; median lobe with large basal capsule and elongate foramen, basoventral projection moderately long, ventral stalk in dorsal view broad though entire length, in ventral view strongly curved towards left; dorsal lobe extremely elongate and strongly curved downwards at apex, gradually narrowing from base towards apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter; antenna and legs lacking modifications; each compound eye composed of approximately 21 ommatidia; elytra constricted at bases, humeral angle rounded. Measurements (as for male): body length 2.09–2.12 mm; length/width of head 0.44–0.46/ 0.48–0.49 mm, pronotum 0.49/ 0.49 mm, elytra 0.59–0.60/ 0.79–0.82 mm; abdomen 0.69–0.71/ 0.78–0.79 mm; length of antenna 0.95–0.97 mm; maximum width of genitalia (Fig. 6H) 0.35 mm.</p> <p>Comparative notes. This is the type species of Jeannel’s Plocamarthrus and center of the P. championi group. The males can be readily separated from all congeners of the group by the enlarged antennomeres 7–8 and mesally expanded antennomere 9, in combination with the distinct apical spine of the mesotibia and a dorsally broad, strongly curved median lobe of the aedeagus.</p> <p>Distribution. Himachal Pradesh and Uttarakhand, India; Gyirong County and Dinggyê County, Tibet, SW China (Figs 77B, 88A, B, 89, 90, 93D–G). New country record for China.</p></div> 	http://treatment.plazi.org/id/C822F213FFD92463CB88A3FE768C229B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFDF2461CB88A45A748D2112.text	C822F213FFDF2461CB88A45A748D2112.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes complexus Yin 2022	<div><p>Arthromelodes complexus sp. nov.</p> <p>(Figs 7, 77B, 93A–C)</p> <p>Chinese common name: Ắúnjà甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, pass to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.53917&amp;materialsCitation.latitude=27.904167" title="Search Plazi for locations around (long 87.53917/lat 27.904167)">Zhêntang Town</a>, 27°54’15”N, 87°32’21”E, 3650–4020 m, 27.vi.2021, Peng, Yin &amp; Zhang, ƱẪŤĠ县ĿƋ沟Ḃ山矮杜ḁff ’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 1 ♀, same data as that of holotype; 1 ♂, 4 ♀♀, same data as that of holotype except ‘ 27°53’58”N, 87°27’21”E, 3060 m, 28.vi.2021, Yin &amp; Zhang leg., ƱẪŤĠ县ĿƋ沟公ḡṅðff ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.5 mm. Head subtruncate at base; vertex with V-shaped sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 5–7 markedly long, 8 smallest, 11 approximately as long as 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to apical 2/5 of elytral length. Hind legs simple, protrochanter with small ventral spine, protibia slightly emarginate at apical 1/3, mesotrochanter with apically setose protuberance on ventral margin, mesotibia with thin, curved apical spine.Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 with broad, bifid protuberance at middle, followed by central cavity and large tubercle at base. Aedeagus strongly asymmetric, median lobe with large basal capsule and elongately oval foramen, ventral stalk broad and dilated at apex, dorsal lobe elongate, forked at apex, parameres reduced and forming single membranous structure. Female. Body length over 2.5 mm, legs and abdomen lacking modifications, genitalia as in Fig. 7J.</p> <p>Description. Male. Body (Fig. 7A) length 2.55–2.58 mm; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 7B) subtruncate at base, approximately as long as wide, length 0.55–0.56 mm, width across eyes 0.54 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with distinct, V-shaped transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, median carina extending before fovea anteriorly to mouthpart, gradually thickened anteriorly. Eyes moderately prominent, composed of approximately 27 ommatidia. Antenna 1.41–1.43 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 5–7 markedly longer than neighboring ones, 3 and 8 of same length, shortest, 9 slightly longer but narrower than 10, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 7B) slightly longer than wide, length 0.62–0.64 mm, width 0.60–0.61 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus slightly shorter than lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.86–0.87 mm, width 0.87–0.89 mm, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea posteriorly to apical 2/5 of elytral length; subhumeral fovea absent, thin marginal stria extending posteriorly from approximately basal 1/3 to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with long, narrow split at middle.</p> <p>Legs elongate, hind legs simple. Protibia (Fig. 7C) broadly emarginate on mesal margin for apical 1/3, protrochanter with small ventral spine; mesotrochanter (Fig. 7D) with short, apically setose protuberance on ventral margin, mesotibia (Fig. 7E) with thin, curved spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.57–0.64 mm, width 0.71–0.74 mm; with modified tergite 1 (IV). Tergite 1 (Fig. 7F) dorsally longer than tergites 2–4 (V–VII) combined, with broad, strongly raised projection at middle, projection with acute posterolateral angles, with transverse cavity posterior to projection, followed by large posteromedial tubercle, posterolateral areas devoid of setae, glabrous, basal impression broken by mediobasal and two pairs of basolateral foveae, with pair of short, thick, triangular discal carinae, with incomplete outer marginal carina; tergites 2 and 3 (V and VI) subequal in length and 4 (VII) slightly longer than 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, lateral carina of tergites 2 and 3 short, 4 complete, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of short lateral carinae; midlength of sternite 2 (IV) slightly longer than 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 7G) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 7H–I) 0.36 mm long, strongly asymmetric; median lobe with large basal capsule and elongately oval foramen, basoventral projection short, ventral stalk protruding, in lateral view strongly dilated at apex; dorsal lobe obliquely erect, apically forked into two lobes; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking spines/projections or modifications; each compound eye composed of approximately 20 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.51–2.54 mm; length/width of head 0.53/ 0.50–0.53 mm, pronotum 0.59–0.60/ 0.57–0.58 mm, elytra 0.80–0.82/ 0.86–0.87 mm; abdomen 0.64–0.70/ 0.75–0.76 mm; length of antenna 1.23–1.30 mm; maximum width of genitalia (Fig. 7J) 0.26 mm.</p> <p>Comparative notes. Arthromelodes complexus is closest to A. alpitorus, A. langjicuo and A. torus by sharing a setose mesotrochanter, similar placement of the male tergal modification and form of the aedeagus. Arthromelodes complexus can be separated from these congeners by the unique emargination of the male protibia, and the structure of the modified male tergite 1 (IV).</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 77B, 93A–C).</p> <p>Etymology. The new species epithet ‘ complexus (complex)’ is a Latin noun referring to the complicated modification of male tergite 1 (IV).</p> </div>	http://treatment.plazi.org/id/C822F213FFDF2461CB88A45A748D2112	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFDC246ECB88A1DB71FB278F.text	C822F213FFDC246ECB88A1DB71FB278F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes cona Yin 2022	<div><p>Arthromelodes cona sp. nov.</p> <p>(Figs 8, 77B, 95A, B)</p> <p>Chinese common name: DZḄnjà甲</p> <p>Type material (60 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.7525&amp;materialsCitation.latitude=27.819445" title="Search Plazi for locations around (long 91.7525/lat 27.819445)">Lebu</a> (勒布), 27°49’10”N, 91°45’09”E, 2600 m, 19.viii.2010, leaf litter, sifted, Jian-Qing Zhu leg.’ (SNUC). PARATYPES: CHINA: 12 ♀♀, same data as that of holotype; 2 ♂♂, 2 ♀♀, same locality and collector, except ‘ 27°49’32”N, 91°45’35”E, 2500 m, 17.viii.2010 ’; 18 ♂♂, 25 ♀♀, same locality, except ‘pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400-2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛乡 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.2 mm. Head subtruncate at base; vertex with deep transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, antennomeres each slightly elongate except for sub-trapezoidal antennomere 10, antennomere 11 modified with impression and ridge on ventral side. Discal stria of elytron extending to approximately apical 4/5 of elytral length. Fore and hind legs simple, mesotrochanter with long, deeply forked spine on ventral margin, mesotibia with exceptionally long apical spine. Abdomen with large Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, simple. Aedeagus strongly asymmetric, median lobe protruding apically, apex pointed, dorsal lobe strongly curved downwards at apical part, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.0 mm, antenna and legs lacking modifications, genitalia as in Fig. 8I.</p> <p>Description. Male. Body (Fig. 8A) length 2.29–2.35 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 8B) subtruncate at base, approximately as long as wide, length 0.49–0.51 mm, width across eyes 0.51–0.52 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with curved deep, slightly sinuate transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocularmandibular carina complete. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 moderately large ommatidia. Antenna 1.26–1.32 mm long, with modified antennomeres 10 and 11 (Fig. 8C); antennomere 1 thick, subcylindrical, 2–9 each elongate, 10 transverse, sub-trapezoidal, 11 large, slightly longer than 9 and 10 combined, ventral surface with oblique carina demarcating broadly impressed area, dorsal surface with mesal margin round-angularly expanded mesally.</p> <p>Pronotum (Fig. 8B) approximately as long as wide, length 0.50–0.53 mm, width 0.53 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.73–0.76 mm, width 0.88–0.91 mm; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humerus weakly prominent; discal stria extending from outer basal fovea to apical 4/5 of elytral length; subhumeral fovea small, marginal stria extending posteriorly from middle to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, forked internally. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 8D) with long, deeply forked spine at base, mesotibia (Fig. 8E) with large blade-like spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.56–0.58 mm, width 0.78–0.79 mm; lacking modifications. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with pair of short discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral sockets in setose basal impression, with pair of short, triangular lateral carinae; midlength of sternite 2 (IV) longer than sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 8F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 8G, H) 0.47 mm long, strongly asymmetric; median lobe with large basal capsule and sub-triangular foramen, basoventral projection moderately long, ventral stalk protruding and pointed at apex; dorsal lobe elongate and strongly curved downwards apically, gradually narrowing towards apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; antenna and legs lacking modifications; each compound eye composed of approximately 20 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.20–2.22 mm; length/width of head 0.47–0.48/ 0.47–0.49 mm, pronotum 0.50–0.51/ 0.51–0.52 mm, elytra 0.62–0.67/ 0.84–0.86 mm; abdomen 0.58–0.59/ 0.77–0.79 mm; length of antenna 0.98–0.99 mm; maximum width of genitalia (Fig. 8I) 0.27 mm.</p> <p>Comparative notes. Arthromelodes cona is placed as a member of the A. championi group based on the aedeagal form. The new species can be readily separated from all members of the group, except for A. abdominalis, by the male’s unique antennal modifications, the large, blade-like spine of the mesotibia, as well as the form of the aedeagus. Arthromelodes abdominalis was described based on a single female, and its identity remains elusive until an associated is made available. However, it appears to be unlikely that A. abdominalis and A. cona represent the same species given the larger body size (2.6 mm vs. 2.20–2.22 mm) and V-shaped frontal sulcus (slightly sinuate in A. cona) of the former species, as well as their widely separated distributions.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 77B, 95A, B).</p> <p>Etymology. The species is named after Cona County, where its type locality is situated.</p></div> 	http://treatment.plazi.org/id/C822F213FFDC246ECB88A1DB71FB278F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFD2246CCB88A36E71B6229A.text	C822F213FFD2246CCB88A36E71B6229A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes crassicornis Yin 2022	<div><p>Arthromelodes crassicornis sp. nov.</p> <p>(Figs 9, 77C, 97C, 100A)</p> <p>Chinese common name: ÝOnjà甲</p> <p>Type material (59 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.50278&amp;materialsCitation.latitude=29.685833" title="Search Plazi for locations around (long 95.50278/lat 29.685833)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, 2190 m, 2019.vii.17, leaf litter, Z.-W. Yin leg., [ƱẪffżḦË 318 Ǟdzƛḡ]’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 4 ♀♀, same data as that of holotype; 9 ♂♂, same locality and collector, except vii.15– viii.1, FIT; 13 ♂♂, 12 ♀♀, same locality, except ‘ 2069 m, 30°1’11.1”N, 95°0’0.46”E, 15.viii.2017, X.-B. Song leg.’; 4 ♂♂, 9 ♀♀, same locality, except ‘ 30°0’35”N, 94°57’58”E, 2180 m, 10.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg.’; 4 ♂♂, 1 ♀, ‘China: Xizang, Mêdog County, near 80K, 29°41’09”N, 95°30’10”E, alt. 2330 m, mixed leaf litter, sifted, 09.vii.2018, Cheng, Peng &amp; Shen leg.’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length 1.92–1.94 mm. Head sub-rectangular at base; vertexal foveae large and asetose, lacking sulcus connecting them; antenna short; antennomeres moniliform, 9–11 enlarged and with dense short setae on dorsal surface. Discal stria of elytron extending to apical 2/5 of elytral length. Legs almost simple, only mesotibia with small apical spur. Metaventrite with dense setose area admesally. Sternite 2 (IV) with pair of trichomes lateral to midline. Aedeagus strongly asymmetric; median lobe with large basal capsule and triangular foramen, dilated at apex; dorsal lobe curved, apical part strongly bent downwards, with pointed apex. Female. Body length 1.85–1.92 mm; antennal club and metaventrite lacking modifications, mesotibia lacking apical spur, genitalia as in Fig. 5I.</p> <p>Description. Male. Body (Fig. 9A) length 1.92–1.94 mm; color dark reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 9B) sub-rectangular at base, approximately as long as wide, length 0.40–0.43 mm, width across eyes 0.41–0.44 mm; vertex finely punctate, with two exceptionally long setae near base and normal ones, with large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae or mediobasal carina, with T-shaped sulcus at middle and few large punctures anterior and lateral to anterior margin of sulcus; antennal tubercles hardly raised; frons slightly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina present. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 35 ommatidia. Antenna relatively short, length 0.86 mm, apical three antennomeres forming distinct club; antennomere 1 thick, subcylindrical, 2–8 each moniliform, 9 much larger than 8, 10 slightly longer and broader than 9, 11 largest, as long as 9 and 10 combined, truncate at base and narrowed apically, dorsal surface of antennomeres 9–11 (Fig. 9C) covered with short, dense setae.</p> <p>Pronotum (Fig. 9B) approximately as long as broad, length 0.42–0.46 mm, width 0.44 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with few exceptionally long setae at base and lateral areas of disc and normal ones, median longitudinal sulcus slightly shorter than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with large lateral procoxal foveae; hypomeral ridge present at basal 2/3, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.57–0.64 mm, width 0.69–0.75 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to apical 2/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, sulcate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite (Fig. 9D) broadly impressed at middle, lateral areas of impression with long, dense setae; with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate, lacking distinct modifications. Mesotibia (Fig. 9E) with tiny apical tubercle.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.46–0.49 mm, width 0.65–0.67 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking discal or marginal carinae; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 longer than 2 and 3 combined along middle, tergite 5 (VIII) broad and slightly transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and two pairs of basolateral foveae in broad setose basal impression; midlength of sternite 2 (IV) approximately as long as 3–6 (V–VIII) combined, 3–5 each with one pair of indistinct basolateral foveae. Sternite 7 (IX) (Fig. 9F) transverse, posterior margin with scattered setae.</p> <p>Aedeagus (Fig. 5G, H) 0.37 mm long, strongly asymmetric; median lobe with large basal capsule and sub-triangular foramen, ventral stalk in ventral view with broadened apex; dorsal lobe elongate and strongly curved, apically strongly bent downwards and pointed at apex; parameres absent.</p> <p>Female. Similar to male in external morphology; antenna shorter; antennal club (Fig. 5I), metaventrite and sternite 2 (IV) lacking modifications; each compound eye composed of approximately 20 ommatidia; humeral angle rounded; mesotibia lacking apical spur, tergite 5 (VIII) not emarginate medially. Measurements (as for male): body length 1.85–1.92 mm; length/width of head 0.40–0.42/ 0.41 mm, pronotum 0.43/ 0.43–0.44 mm, elytra 0.51– 0.55/ 0.70 mm; abdomen 0.51–0.53/ 0.65–0.68 mm; length of antenna 0.77–0.79 mm; maximum width of genitalia (Fig. 5J) 0.25 mm.</p> <p>Comparative notes. The new species can be readily identified by the unique modifications of the male antennal clubs and the form of the aedeagus. The setose posterior area of the metaventrite of the male is similarly shared with A. nepaeformis and A. lebus, which may indicate close relationships between these species.</p> <p>Distribution. Chagyib District (Nyingchi) and Mêdog County, Tibet, SW China (Figs 77C, 97C, 100A).</p> <p>Etymology. The specific epithet is formed by the Latin words ‘ crassus -a -um (dense, thick, solid)’ and ‘ cornis (horned)’, referring to the relatively short antennae and stout male antennal clubs of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FFD2246CCB88A36E71B6229A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFD0246BCB88A45A774721AC.text	C822F213FFD0246BCB88A45A774721AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes criniger Yin 2022	<div><p>Arthromelodes criniger sp. nov.</p> <p>(Figs 10, 77C, 93A, B)</p> <p>Chinese common name: Ợbnjà甲</p> <p>Type material (8 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, pass to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.53917&amp;materialsCitation.latitude=27.904167" title="Search Plazi for locations around (long 87.53917/lat 27.904167)">Zhêntang Town</a>, 27°54’15”N, 87°32’21”E, 3650–4020 m, 27.vi.2021, Peng, Yin &amp; Zhang, ƱẪŤĠ县ĿƋ沟Ḃ山矮杜ḁff ’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 5 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length over 2.4 mm. Head subtruncate at base; median area of vertex with dense, markedly long setae oriented posteriorly, vertex with thin V-shaped sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 smallest, 11 approximately as long as 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to slightly more than half of elytral length. Fore legs simple, mesotrochanter with apically setose protuberance on ventral margin, mesotibia with short apical spine, metatrochanter with setose tuft on ventral margin. Abdomen lacking modifications, tergite 1 (IV) with long, moderately dense setae at posteromedian area, longer than tergites 2–4 (V–VII) combined. Aedeagus strongly asymmetric, median lobe with large basal capsule and elongately oval foramen, ventral stalk broad and gradually dilated at apex, dorsal lobe elongate, forked at apex, parameres reduced and forming single membranous structure. Female. Body length over 2.3 mm, legs and abdomen lacking modifications, genitalia as in Fig. 10J.</p> <p>Description. Male. Body (Fig. 10A) length 2.43–2.46 mm; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with moderately elongate pubescence.</p> <p>Head (Fig. 10B) subtruncate at base, slightly longer than wide, length 0.52–0.53 mm, width across eyes 0.50– 0.51 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with thin, V-shaped transverse sulcus between antennal tubercles, median area with dense, markedly long setae oriented posteriorly and passing posterior margin of head, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, median carina extending from fovea anteriorly to mouthpart, gradually thickened anteriorly. Eyes moderately prominent, composed of approximately 25 ommatidia. Antenna 1.27–1.32 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 3 and 8 of same length, shortest, 9 as long as but slightly narrower than 10, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 10B) as long as wide, length 0.54–0.56 mm, width 0.54–0.56 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus slightly shorter than lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.76–0.81 mm, width 0.84–0.87 mm, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea posteriorly to slightly more than half of elytral length; subhumeral fovea absent, thin marginal stria extending posteriorly from posterior middle to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with long, narrow split at middle.</p> <p>Legs elongate, fore legs simple. Mesotrochanter (Fig. 10C) with short, apically setose protuberance on ventral margin, mesotibia (Fig. 10D) with short spine at apex; metatrochanter (Fig. 10E) with short setose tuft on ventral margin.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.68 mm, width 0.75– 0.77 mm; lacking modifications. Tergite 1 (Fig. 10F) dorsally longer than tergites 2–4 (V–VII) combined, basal impression broken by broadly separated mediobasal and one pair of basolateral foveae, with pair of short discal carinae, with incomplete outer marginal carina; tergites 2 and 3 (V and VI) subequal in length and 4 (VII) slightly longer than 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, lateral carina of tergites 2 and 3 short, 4 complete, 5 (VIII) semicircular, posterior margin roundly and deeply emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of short lateral carinae; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 10G) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 10H–I) 0.35 mm long, strongly asymmetric; median lobe with large basal capsule and elongately oval foramen, basoventral projection short, ventral stalk protruding, in lateral view gradually dilated at apex; dorsal lobe obliquely erect, apically forked into two lobes; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking spines/projections or modifications; each compound eye composed of approximately 20 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.33–2.48 mm; length/width of head 0.49–0.52/ 0.48–0.52 mm, pronotum 0.55–0.58/ 0.52–0.55 mm, elytra 0.69–0.75/ 0.82–0.88 mm; abdomen 0.68–0.75/ 0.74–0.83 mm; length of antenna 1.16–1.21 mm; maximum width of genitalia (Fig. 10J) 0.27 mm.</p> <p>Comparative notes. Arthromelodes complexus is allied with A. torus and a number of species described in this paper by the similar form of the aedeagus, but can be readily separated from all of them by the dense, elongate setae that cover the central part of the vertex in both sexes.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 77C, 93A, B).</p> <p>Etymology. The new species name ‘ criniger (having long hair)’ is a Latin adjective suggested by long setae on the male vertex of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFD0246BCB88A45A774721AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFD62468CB88A773773D2446.text	C822F213FFD62468CB88A773773D2446.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes cylindricus Yin 2022	<div><p>Arthromelodes cylindricus sp. nov.</p> <p>(Figs 11, 78A, 96E, F)</p> <p>Chinese common name: Ñ型njà甲</p> <p>Type material (19 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, Shejila, nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.69278&amp;materialsCitation.latitude=29.61389" title="Search Plazi for locations around (long 95.69278/lat 29.61389)">Sheji’ema</a>, 29°36’50”N, 95°41’34”E, 4340 m, 05.vii.2018, leaf litter, sifted, Cheng, Peng &amp; Shen leg. [ƱẪffż市DzĻŃ]’ (SNUC). PARATYPES: CHINA: 9 ♂♂, 4 ♀♀, same data as that of holotype; 2 ♂♂, 3 ♀♀, also from Shejila, except ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.69861&amp;materialsCitation.latitude=29.616945" title="Search Plazi for locations around (long 94.69861/lat 29.616945)">Lulang Tourist Center</a>, 29°37’01”N, 94°41’55”E, 4300 m, 14.vii.2021, Peng, Yin &amp; Zhang, DzĻŃ游Β中 心北坡 ’ (SNUC).</p> <p>Diagnosis. Male. Body length over 2.60–2.67 mm. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna long, antennomeres each slightly elongate, lacking modifications. Discal stria of elytron extending to approximately apical 1/4 of elytral length. Fore legs simple, mesotrochanter with small tuft of setae on ventral margin, mesotibia with distinct apical spur, metafemur with dorsal setose patch at apex. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) with deep central cavity. Aedeagus strongly asymmetric, median lobe protruding apically, ventral stalk in dorsal view broad, dorsal lobe laterally strongly curved at middle, parameres reduced and forming single semi-sclerotized structure.</p> <p>Description. Male. Body (Fig. 11A) length 2.60–2.67 mm, habitus generally cylindrical; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 11B) subtruncate at base, as long as broad, length and width across eyes 0.50–0.51 mm; vertex, frons and clypeus roughly punctate; vertex with moderately large, asetose vertexal foveae (dorsal tentorial pits), with curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus, with short frontal-clypeal carinae at sides; clypeus with anterior margin carinate and moderately raised; ocular-mandibular carina thick and complete. Venter with gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 19 large ommatidia. Antenna with loosely formed club, lacking modifications, length 1.23–1.24 mm; antennomere 1 thick, subcylindrical, anterolateral margin slightly impressed and filled with short setae (but not to form prominent trichome), 2–11 each elongate, 8 smallest, 11 largest, much shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 11B) approximately as long as wide, length and width 0.55–0.56 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae, with additional pair of marginal antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.73–0.74 mm, width 0.82 mm, constricted at bases; each elytron with two large, asetose basal foveae, lacking subbasal fovea; humerus lacking prominence or denticle; discal stria extending from outer basal fovea to approximately apical 1/4 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from basal 1/3 to apex. Hind wings absent.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore simple. Mesotrochanter with small setose tuft on ventral margin, mesotibia with distinct apical spur; metatrochanter setose on ventral margin, metafemur (Fig. 11C) with dorsal sensory area covered with short, dense setae at apex.</p> <p>Abdomen narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.75–0.78 mm, width 0.73–0.74 mm. Tergite 1 (IV) (Fig. 11D) strongly modified, much longer than tergites 2–4 (V–VII) combined; with deep central cavity, inside cavity with ‘ ± ’-shaped thick ridge, with broad, truncate projection posterior to cavity, areas lateral to projection markedly concave; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) slightly longer than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of short lateral carinae, with thin marginal carinae much longer than lateral ones; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 11E) with moderately sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 11F, G) 0.51 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk protruding and in dorsal view broad, apex narrowly and shortly split; dorsal lobe in lateral view elongate and bent at middle, in dorsal view strongly curved rightwards (position in figure) at apical 1/5, gradually narrowing apically and with pointed apex; parameres reduced to single semi-membranous structure.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter; legs and abdomen lacking modifications; each compound eye composed of approximately 19 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.56–2.57 mm; length/width of head 0.51–0.52/ 0.50–0.51 mm, pronotum 0.55–0.58/ 0.55–0.58 mm, elytra 0.70–0.72/ 0.85–0.87 mm; abdomen 0.75–0.80/ 0.80 mm; length of antenna 1.17 mm; maximum width of genitalia (Fig. 11H) 0.36 mm.</p> <p>Comparative notes. The new species can be readily separated from all known congeners by the relatively large body size and a generally cylindrical habitus, combined with the unique sensory patch of the male metafemur and the structure of the abdominal modifications.</p> <p>Distribution. Chagyib District (Nyingchi), Xizang, SW China (Figs 78A, 96E, F).</p> <p>Etymology. The specific epithet ‘ cylindricus (- a, - um)’ is a Latin adjective (of Ancient Greek origin) referring to the cylindrical habitus of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FFD62468CB88A773773D2446	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFEB2456CB88A1DB77D8278E.text	C822F213FFEB2456CB88A1DB77D8278E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes flosculus Yin 2022	<div><p>Arthromelodes flosculus sp. nov.</p> <p>(Figs 12, 78A, 99D, G)</p> <p>Chinese common name: 小ǩnjà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.27383&amp;materialsCitation.latitude=29.31093" title="Search Plazi for locations around (long 95.27383/lat 29.31093)">Guoguotang Great Ben</a>, 29°18’39.35”N, 95°16’25.75”E, 1180 m, 2019.vii.22, leaf litter, Z.-W. Yin leg., [ƱẪẸṘ果果ḙ]’ (SNUC).</p> <p>Diagnosis. Male. Body length over 2.6 mm. Head sub-rectangular at base; vertex with complete reversed Ushaped sulcus connecting vertexal foveae. Head, pronotum, elytra and abdomen with few long thickened, erect setae and normal ones. Legs simple, lacking modifications. Tergite 1 (IV) strongly modified, composed of central cavity filled with specialized trichomes, mediobasal tubercle and lateral setose patches. Aedeagus strongly asymmetric; median lobe with large basal capsule and foramen, dorsal lobe elongate and strongly twisted at apical 1/2.</p> <p>Description. Male. Body (Fig. 12A) length 2.62 mm; color dark reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with long pubescence.</p> <p>Head (Fig. 12B) sub-rectangular at base, approximately as long as wide, length 0.58 mm, width across eyes 0.56 mm; vertex finely punctate, with small, asetose vertexal foveae (dorsal tentorial pits), with complete reversed U-shaped sulcus connecting foveae, mediobasal carina extending from near head base anteriorly to sulcus, with few long thickened setae at antennal tubercles and near base; antennal tubercles slightly raised; frons impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and slightly raised; ocularmandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 large ommatidia. Antenna lacking distinct club and modification, length 1.43 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, as long as 9 and 10 combined, fusiform.</p> <p>Pronotum (Fig. 12B) slightly broader than long, length 0.60 mm, width 0.56 mm, widest at middle; lateral margins rounded, with few long thickened, erect setae pointed anteriorly; disc slightly convex, finely punctate, with scattered long setae laterally, with one median and one pair of semicircular lateral longitudinal sulci; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with large lateral procoxal foveae; hypomeral ridge present at basal halves; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.82 mm, width 0.98 mm; each elytron with two large, asetose basal foveae; humerus slightly prominent; discal stria extending from outer basal fovea to apical 3/5 of elytral length; subhumeral fovea absent, with carinate marginal stria extending from subbase to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, lacking distinct modifications. Mesotrochanter with small ventral tubercle, mesotibia with tiny tubercle at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.90 mm, width 0.70 mm. Tergite 1 (IV) (Fig. 12C) strongly modified, longer than tergites 2–4 (V–VII) combined; structure of modification complex, composed of central cavity filled with specialized trichomes at middle and sides, with large mediobasal tubercle covered with dense setae at apex and broad lateral setose patches with coarse punctation; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, short marginal carinae extending for basal 1/3 of tergal length; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 longer than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and basolateral foveae in broad setose basal impression, with pair of short lateral carinae; midlength of sternite 2 (IV) approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 12D) with weakly sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 12E, F) 0.47 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk narrowing apically, with flat apex in ventral view; dorsal lobe elongate and strongly twisted, with pointed apex; parameres reduced and forming single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species can be readily separated from all known congeners by the unique structure of the male modifications on tergite 1 (IV), as well as by the form of the aedeagus. The exceptionally long setae along the posterolateral margins of the head, lateral margins of the pronotum and elytral disc is shared with A. lotus. But otherwise these two species can be readily separated by the larger body size, much more elongate antennae, and a totally different form and structure of the male tergal modification and the aedeagus of A. flosculus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 78A, 99D, G).</p> <p>Etymology. The specific epithet ‘ flosculus ’ (a small flower, flowery ornament) refers to the flower-like modification of male tergite 1 (IV) of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFEB2456CB88A1DB77D8278E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFEA2454CB88A7BF759724F6.text	C822F213FFEA2454CB88A7BF759724F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes gyamda Yin 2022	<div><p>Arthromelodes gyamda sp. nov.</p> <p>(Figs 13, 78A, 96A, D)</p> <p>Chinese common name: ȕḭnjà甲</p> <p>Type material (10 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gongbo'gyamda, near <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.23944&amp;materialsCitation.latitude=29.9" title="Search Plazi for locations around (long 93.23944/lat 29.9)">Gaduogang</a> vil., 29°53’60”N, 93°14’22”E, 3620 m, mixed leaf litter, sifted, 18.vii.2018, Cheng, Peng &amp; Shen leg. [工布ȕḭ]’ (SNUC). PARATYPES: CHINA: 4 ♂♂, 4 ♀♀, same data as that of holotype; 1 ♂, China: Xizang, Nyingchi City, valley nr. Jiare Vill., Chongge, 30°03’19”N, 93°46’46”E, 3500–3750 m, 08.vii.2021, Peng, Yin, Zhang, ƱẪffż ẘ热村ṗẋ ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.1 mm. Head subtruncate at base; vertex with shallow, slightly curved transverse sulcus between antennal tubercles and moderately long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres 1–7 each slightly elongate, 8 smallest, 9 and 10 approximately as long as wide, 11 as long as 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately half of elytral length. Fore and hind legs simple, mesotrochanter with long ventral protuberance, mesofemur with short ventral spine, mesotibia with small apical denticle. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergites 1–4 each modified (IV–VII). Aedeagus strongly asymmetric, median lobe relatively short, ventral stalk in dorsal view broad, dorsal lobe elongate, apex strongly twisted, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.0 mm, legs and abdomen lacking modifications, genitalia as in Fig. 13I.</p> <p>Description. Male. Body (Fig. 13A) length 2.17–2.28 mm; color light reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 13B) subtruncate at base, approximately as long as wide, length 0.43–0.44 mm, width across eyes 0.44 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with shallow and thin, slightly curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles slightly raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two tiny gular foveae (posterior tentorial pits) in single small pit, with distinct median carina extending from fovea anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 18–20 ommatidia. Antenna 1.01–1.02 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, much shorter and narrower than 7, 9–11 successively larger, 11 largest, slightly shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 13B) slightly longer than wide, length 0.48–0.49 mm, width 0.46 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.61–0.63 mm, width 0.76–0.78 mm, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea to apical 1/2 of elytral length; subhumeral fovea lacking, thin marginal stria extending posteriorly from basal third to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite slightly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 13C) with long protuberance at base, mesofemur (Fig. 13C) with short ventral spine, mesotibia with small denticle at apex.</p> <p>Abdomen approximately as broad as elytra, widest at lateral margins of tergite 1 (IV), length 0.64–0.71 mm, width 0.75–0.77 mm; with modified tergites 1–4 (IV–VII) (Fig. 13D). Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, with small, median protuberance near posterior margin, which is decorated with dense golden setae admesally; setose basal sulcus separated by mediobasal and one pair of large basolateral foveae, with very short discal carinae; tergites 2 and 3 (V and VI) each with median protuberance and dense golden setae along posterior margin, tergite 4 (VII) much longer than 2 and 3 combined along middle, with large median projection, area anterior projection strongly impressed, tergites 2–4 (V–VII) each with one pair of small basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral sockets, lacking obvious basal impression or sulcus, with pair of moderately long lateral carinae; midlength of sternite 2 (IV) approximately as long as sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 13E) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 13F–H) 0.40 mm long, strongly asymmetric; median lobe with large basal capsule and slightly oval foramen, basoventral projection short, ventral stalk short and in dorsal view dilated at apex; dorsal lobe elongate and gradually narrowing from middle towards apex, strongly twisted at apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking modifications; each compound eye composed of approximately 18 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.13–2.16 mm; length/width of head 0.44/ 0.43–0.44 mm, pronotum 0.46–0.50/ 0.45–0.46 mm, elytra 0.59–0.60/ 0.78 mm; abdomen 0.66–0.67/ 0.76–0.79 mm; length of antenna 0.96–1.00 mm; maximum width of genitalia (Fig. 13I) 0.27 mm.</p> <p>Comparative notes. The males of the new species can be readily separated from those of all congeners by the relatively small head and short antennae, the unique modifications of tergites 1–4 (IV–VII), as well as the form of the aedeagus.</p> <p>Distribution. Gongbo'gyamda County, Tibet, SW China (78A, 96A, D).</p> <p>Etymology. The specific epithet ‘ gyamda ’ is taken from Gongbo'gyamda County, the type locality of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFEA2454CB88A7BF759724F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFEF2452CB88A1DB772E2492.text	C822F213FFEF2452CB88A1DB772E2492.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes intricatus Yin 2022	<div><p>Arthromelodes intricatus sp. nov.</p> <p>(Figs 14, 78B, 93E)</p> <p>Chinese common name: Ḕúnjà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Zhêntang Town</a>, Xiuxiongma Vill., 27°54’11”N, 87°22’42”E, 2700–3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJŤĠ县Ŀ ƋWšŏ玛村 ’ (SNUC).</p> <p>Diagnosis. Male. Body length over 2.4 mm; dorsal surface covered with long pubescence. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles. Pronotum distinctly broader than head. Posterolateral areas of elytra with exceptionally long pubescence. Mesotibia with long golden setae on ventral margin; ventral surface of metatrochanter covered by dense, short setae. Tergite 1 (IV) strongly modified, composed of large central cavity of complex structures and lateral patches with dense, short setae.Aedeagus strongly asymmetric; median lobe with large basal capsule and triangular foramen, dorsal lobe protruding and forked apically.</p> <p>Description. Male. Body (Fig. 14A) length 2.46 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with long pubescence.</p> <p>Head (Fig. 14B) subtruncate at base, approximately as long as wide, length 0.53 mm, width across eyes 0.55 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), with slightly curved transverse sulcus between antennal tubercles, long mediobasal carina extending from head base anteriorly to sulcus, lateral carina short, extending from near head base anteriorly to level of anterior margin of eyes; antennal tubercles moderately raised, postantennal area and frons roughly punctate; frons broadly and weakly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and slightly raised; ocular-mandibular carina complete, branched twice below eye, with middle carina shorter than upper and lower ones, lower carina extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 23 large ommatidia. Antenna with long pubescence, lacking distinct club and modification, length 1.48 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, shorter than 9 and 10 combined, fusiform.</p> <p>Pronotum (Fig. 14B) approximately as long as wide, length 0.61 mm, width 0.63 mm, widest at middle; lateral margins rounded, with dense setae at apical halves; disc slightly convex, finely punctate, with scattered long setae, median longitudinal sulcus shorter than semicircular lateral ones; lacking median antebasal fovea, mediobasal carina short, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part longer than coxal part, with small lateral procoxal foveae; thick hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra strongly convex, much wider than long, length 0.75 mm, width 0.89 mm, moderately constricted at bases; each elytron with two large, asetose basal foveae; humerus slightly prominent; discal stria extending from outer basal fovea to apical 1/5 of elytral length; small subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed medially, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, with meso- and metatrochanter and mesotibia weakly modified. Mesotrochanter (Fig. 14C) with small ventral tubercle and long golden setae on ventral margin, mesotibia with tiny tubercle at apex; metatrochanter (Fig. 14D) with ventral surface densely covered by short setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.56 mm, width 0.72 mm. Tergite 1 (IV) (Fig. 14E) strongly modified, dorsally longer than tergites 2–4 (V–VII) combined; structure of modification complex, large central cavity with middle and basal projections, median row of dense golden setae and clustered setae at middle of raised lateral margins, area lateral to cavity sloping laterally, densely covered with short setae; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking distinct marginal carina; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 longer than 2 and 3 combined along middle, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal foveae and large basolateral sockets, with pair of short lateral carinae; midlength of sternite 2 (IV) longer than 3–5 (V–VII) combined, sternites 3–5 each with pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate and emarginate at middle, sternite 7 (IX) (Fig. 14F) with sclerotized rounded apical half and few setae along apical margin.</p> <p>Aedeagus (Fig. 14G, H) 0.36 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk narrowing apically, apex pointed on right side and with acute subapical projection; dorsal lobe elongate and strongly bent, deeply forked at apex; parameres reduced and forming single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. Arthromelodes intricatus resembles A. flosculus by the elongate antennae, and a similar pattern of tergal modifications. The new species can be readily separated by the lack of a reversed U-shaped sulcus on vertex, the more strongly expanded lateral margins of the pronotum, the more elongate elytra with dense setae along the sides and posterolateral margins, and a different structure of the tergal modification and the aedeagus.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 78B, 93E).</p> <p>Etymology. The specific epithet ‘ intricatus ’ (intricate, complex) refers to the complicated structure of the male tergal modification of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFEF2452CB88A1DB772E2492	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFED2450CB88A1DB71F424DA.text	C822F213FFED2450CB88A1DB71F424DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes lage Yin 2022	<div><p>Arthromelodes lage sp. nov.</p> <p>(Fig. 15, 78B, 98A, B)</p> <p>Chinese common name: Ń格njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang A. R., Mêdog County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.98944&amp;materialsCitation.latitude=29.470001" title="Search Plazi for locations around (long 94.98944/lat 29.470001)">Lage</a> (ca 5.5 km E Dochula Pass, 29°28’12”N, 94°59’22”E), 3000–3500 m, 11.viii.2005, Liang Tang leg., [ƱẪẸṘ县Ń格]’ (SNUC).</p> <p>Diagnosis. Male. Body length 2.6 mm. Head rounded at base; vertex with transverse V-shaped sulcus between antennal tubercles, vertexal foveae asetose and small; antenna long and slender, antennomeres each slightly elongate, lacking modifications. Discal stria of elytron extending to approximately half of elytral length. Protibia expanded at middle; mesotrochanter with tufted golden setae on ventral margin. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) with broad and deep central cavity and laterally impressed. Aedeagus strongly asymmetric, ventral stalk of median lobe protruding, apex membranous, dorsal lobe strongly curved at apex, parameres reduced and forming single membranous structure.</p> <p>Description. Male. Body (Fig. 15A) length 2.60 mm; color reddish-brown, tarsi and mouthparts paler. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 15B) rounded at base, slightly broader than long, length 0.51 mm, width across eyes 0.57 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with V-shaped sulcus between antennal tubercles, mediobasal carina relatively long, distinct, extended from posterior margin of vertexal sulcus towards near head base, antennal tubercles strongly raised; frons broadly impressed medially, confluent with clypeus at middle; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two small gular foveae (posterior tentorial pits) in single pit, with thin median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 small ommatidia. Antenna lacking modification, length 1.38 mm, club loosely formed by antennomeres 9–11; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 15B) slightly longer than wide, length 0.58 mm, width 0.53 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae small. Prosternum with anterior part approximately as long as coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with indistinct lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.75 mm, width 0.88 mm; each elytron with two medium-sized, asetose basal foveae, lacking subbasal fovea; humerus slightly prominent; discal stria extending from outer basal fovea to approximately half of elytral length; lacking subhumeral fovea, marginal stria extending posteriorly from basal 1/4 to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite weakly impressed at middle, with pair of setose lateral metaventral foveae at middle; posterior margin with small, narrow split at middle.</p> <p>Legs elongate. Protibia expanded from basal 1/3 to near apex; mesotrochanter (Fig. 15C) with long tufted golden setae on ventral margin.</p> <p>Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.69 mm, width 0.85 mm. Tergite 1 (IV) (Fig. 15D) strongly modified, with broad, deep central cavity, anterior margin of cavity strongly carinate, posterior margin with narrow projection at middle, inside central cavity another ‘∞’-shaped cavity with densely setose anterior margin, areas lateral to cavity strongly impressed and smooth; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with thick, triangular discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) lacking basal sulcus, with large mediobasal and two pairs of basolateral foveae, with pair of distinct lateral carinae; midlength of sternite 2 (IV) as long as 3–5 (V–VII) combined, sternites 3–5 approximately subequal in length along midline, each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 15E) with weakly sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 15F, G) 0.35 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, with short basoventral projection, ventral stalk strongly protruding, apex in dorsal view expanded; dorsal lobe in lateral view broader than apical part of median lobe, strongly curved in dorso-ventral view; parameres reduced to single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. Arthromelodes lage is similar to A. angulatus in the location of the central cavity and lateral impressed areas of male tergite 1 (IV), as well as the form of the aedeagus in ventral view. Arthromelodes lage can be readily separated from the latter species by the expanded male protibiae (slender in A. angulatus), the presence of tufted setae at the ventral margin of the mesotrochanter (lacking in A. angulatus), the different structures of the abdominal modifications, and a relatively much less protruding median lobe of the aedeagus. The new species is also similar to A. aniqiao, but can be separated by the expanded male protibiae and a different shape of the median lobe of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 78B, 98A, B).</p> <p>Etymology. The specific epithet is taken from ‘Lage’, type locality of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FFED2450CB88A1DB71F424DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFE3245DCB88A1DB713C27D6.text	C822F213FFE3245DCB88A1DB713C27D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes langjicuo Yin 2022	<div><p>Arthromelodes langjicuo sp. nov.</p> <p>(Figs 16, 78B, 91)</p> <p>Chinese common name: 朗吉śnjà甲</p> <p>Type material (60 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, Gyirong Valley, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.399445&amp;materialsCitation.latitude=28.402777" title="Search Plazi for locations around (long 85.399445/lat 28.402777)">Langjicuo Lake</a>, 28°24’10”N, 85°23’58”E, 4000–4100 m, 21.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW 朗吉śéae ’ (SNUC). PARATYPES: CHINA: 30 ♂♂, 29 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length over 2.3 mm. Head subtruncate at base; vertex with V-shaped transverse sulcus between antennal tubercles and moderately long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 smallest, 11 as long as 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately half of elytral length. Fore and hind legs simple, mesotrochanter with long setose tuft on ventral margin, mesotibia with small apical denticle. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 weakly modified, with pair of small admesal tubercles at posterior 1/4. Aedeagus strongly asymmetric, median lobe with large basal capsule and elongate foramen, ventral stalk narrowed and protruding, dilated at apex, dorsal lobe elongate, apical half thin and pointing towards apex, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.3 mm, legs and abdomen lacking modifications, genitalia as in Fig. 16I.</p> <p>Description. Male. Body (Fig. 16A) length 2.37–2.41 mm; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 16B) subtruncate at base, approximately as long as wide, length 0.52–0.53 mm, width across eyes 0.51–0.52 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with distinct V-shaped transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles slightly raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, with distinct median carina extending from fovea anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 30 ommatidia. Antenna 1.33–1.37 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 8 smallest, much shorter and narrower than 7, 9 and 10 of same length, 10 slightly broader than 9, 11 largest, as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 16B) slightly longer than wide, length 0.54–0.56 mm, width 0.51–0.52 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus slightly shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.76–0.77 mm, width 0.87–0.91 mm, truncate at base; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humerus weakly raised; discal stria extending from outer basal fovea to approximately half of elytral length; subhumeral fovea absent, thin marginal stria extending posteriorly from basal third to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite slightly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 16C) with long setose tuft on ventral margin, mesotibia (Fig. 16D) with small denticle at apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.62–0.64 mm, width 0.77–0.82 mm; with modified tergite 1 (IV). Tergite 1 (Fig. 16E) longer than tergites 2–4 (V–VII) combined, degree of modification development varying among individuals, with two small admesal tubercles at middle, posterior half longitudinally raised at middle, with pair of admesal carinae or tubercles, area surrounding tubercle weakly to strongly impressed (surface with round, small cavity), basal impression broken by mediobasal and basolateral foveae, with pair of short discal carinae; tergites 2 and 3 (V and VI) subequal in length along midline, 4 (VII) as long as 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of moderately long lateral carinae; midlength of sternite 2 (IV) longer than 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 16F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 16G, H) 0.37 mm long, strongly asymmetric; median lobe with large basal capsule and elongate foramen, basoventral projection short, ventral stalk protruding and dilated at apex; dorsal lobe elongate and in dorsal view flattened at middle, apical part strongly bent rightwards and narrowing towards apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology, antenna shorter, legs and abdomen lacking spines/projections or modifications, each compound eye composed of approximately 25 ommatidia and humeral angle rounded. Measurements (as for male): body length 2.33–2.37 mm; length/width of head 0.50–0.51/ 0.51 mm, pronotum 0.53–0.54/ 0.50–0.51 mm, elytra 0.70–0.74/ 0.87–0.89 mm; abdomen 0.70–0.71/ 0.85–0.86 mm; length of antenna 1.21–1.22 mm; maximum width of genitalia (Fig. 16I) 0.26 mm.</p> <p>Comparative notes. Arthromelodes langjicuo inhabits leaf litter of a Rhododendron forest at an altitude of 4,100 m, being one of the highest-dwelling species within the genus. It is close to previously described A. angulatus, A. aniqiao, and A. lage, and A. torus, sharing with them a setose tuft on the ventral margin of the male mesotrochanter, as well as the generally similar form of the aedeagus. The unique form of the male tergal modification and the aedeagus readily leads to a separation of the new species from these similar congeners.</p> <p>Distribution. Gyirong County, Tibet, SW China (Figs 78B, 91).</p> <p>Etymology. The new species is named after its type locality, i.e., Langjicuo Lake.</p></div> 	http://treatment.plazi.org/id/C822F213FFE3245DCB88A1DB713C27D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFE1245BCB88A09677B82492.text	C822F213FFE1245BCB88A09677B82492.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes latithorax Yin 2022	<div><p>Arthromelodes latithorax sp. nov.</p> <p>(Figs 17, 78C, 92C)</p> <p>Chinese common name: Pḃnjà甲</p> <p>Type material (6 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyalam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.983055&amp;materialsCitation.latitude=27.987501" title="Search Plazi for locations around (long 85.983055/lat 27.987501)">Zhangmu Town</a> (樟木W), 27°59’15”N, 85°58’59”E, 2250 m, leaf litter, sifted, 22.vii.2010, Jian-Qing Zhu leg.’ (SNUC). PARATYPES: CHINA: 3 ♀♀, same data as that of holotype; 2 ♀♀, same data as that of holotype except ‘ 27°57’21”N, 85°58’41”E, 27.vii.2010 ’ (all in SNUC).</p> <p>Diagnosis. Male. Body length over 2.1 mm. Head sub-rectangular at base; vertex with deep transverse sulcus between antennal tubercles and moderately long mediobasal carina, vertexal foveae moderately large and asetose; antenna elongate, lacking modifications; antennomeres 1–7 each slightly elongate, 8 smallest, 9 and 10 approximately subequal in width, 9 longer than 10, 11 slightly shorter than 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately apical 3/5 of elytral length. Fore and hind legs simple, mesotrochanter with short ventral spine, mesofemur strongly thickened, mesotibia with thickened, with large curved spine at apex. Abdomen lacking modifications, with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined. Aedeagus strongly asymmetric, median lobe relatively broad in dorsal view, dorsal lobe strongly elongate, ventral stalk strongly curved and with pointed apex, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.0 mm, legs lacking modifications, genitalia as in Fig. 17I.</p> <p>Description. Male. Body (Fig. 17A) length 2.16 mm; color reddish-brown, head and pronotum slightly darker than elytra and abdomen, tarsi and mouthparts lighter. Dorsal surface of body covered with dense long pubescence.</p> <p>Head (Fig. 17B) sub-rectangular at base, wider than long, length 0.47 mm, width across eyes 0.52 mm; vertex finely punctate, with relatively large, asetose vertexal foveae (dorsal tentorial pits), with deep, curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles weakly prominent; frons almost flat, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 large ommatidia. Antenna 1.16 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, much shorter and narrower than 7, 9–10 successively shorter, subequal in width, 11 largest, slightly shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 17B) slightly wider than long, length 0.58 mm, width 0.62 mm, widest at middle; lateral margins laterally expanded and rounded; disc weakly convex, finely punctate, with median longitudinal sulcus longer than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, anteriorly confluent with additional setose marginal sulcus, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.76 mm, width 0.90 mm, narrowed at base; each elytron with two large basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea to apical 3/5 of elytral length; subhumeral fovea present, thin marginal stria extending posteriorly from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite slightly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 17C) with short, setose spine at base, mesofemur (Fig. 17C) strongly thickened, mesotibia (Fig. 17D) with large, curved spine at apex.</p> <p>Abdomen as broad as elytra, widest at lateral margins of tergite 1 (IV), length 0.43 mm, width 0.90 mm, lacking modifications. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; lacking basal sulcus\impression, with single mediobasal fovea (likely formed by medially moved ones) and one pair of basolateral foveae, lacking discal carinae; tergites 2 and 3 (V and VI) each short, tergite 4 (VII) approximately as long as 2 and 3 combined along middle, tergites 2–4 (V–VII) each with one pair of small basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with single, large mediobasal and large, round basolateral foveae, with setose basal impression laterally, lacking carinae; midlength of sternite 2 (IV) approximately as long as sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 17E) slightly oval, moderately sclerotized, apical margin rounded.</p> <p>Aedeagus (Fig. 17F–H) 0.53 mm long, strongly asymmetric; median lobe with moderately large basal capsule and slightly oval foramen, basoventral projection relatively long, ventral stalk long and in dorsal view broad and moderately curved; dorsal lobe elongate and gradually narrowing from base towards apex, strongly curved at apical portion, with pointed apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; legs lacking modifications; each compound eye composed of approximately 18 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.05–2.23 mm; length/width of head 0.47–0.48/ 0.52–0.54 mm, pronotum 0.57–0.58/ 0.61–0.62 mm, elytra 0.73–0.75/ 0.91–0.96 mm; abdomen 0.45/ 0.87–0.94 mm; length of antenna 1.10–1.15 mm; maximum width of genitalia (Fig. 17I) 0.47 mm.</p> <p>Comparative notes. Arthromelodes latithorax is a member of the A. championi group based on the unique form of the aedeagus. It is most similar to the sympatric A. zhangmu by sharing similar pronotal expansions, thickened male mesofemur, and mesotibia with a large apical spine. They can be differentiated by the more strongly expanded lateral margins of the pronotum, the more strongly curved mesotibia and apical spine, as well as by the dorsally much narrower ventral stalk of aedeagal median lobe of Arthromelodes latithorax. The new species also shares with the northern Indian A. planiceps the more or less laterally expanded tempora and pronotum, but can be readily separated by the male lacking a prominent humerus, the thickened mesofemur, and the mesotibia with a large, curved apical spine. Arthromelodes planiceps has a denticulate elytral humerus in the male, and lacks obvious sexual characters on the middle legs.</p> <p>Distribution. Nyalam County, Tibet, SW China (Figs 78C, 92C).</p> <p>Etymology. The species name is a combination of Latin ‘ latus, - a, - um ’ (wide, broad) and ‘ thorax ’, referring to the laterally expanded prothorax of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FFE1245BCB88A09677B82492	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFE72459CB88A252777925F9.text	C822F213FFE72459CB88A252777925F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes lebus Yin 2022	<div><p>Arthromelodes lebus sp. nov.</p> <p>(Figs 18, 78C, 95A, B)</p> <p>Chinese common name: 勒布njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 1.8 mm. Head sub-rectangular at base; vertexal foveae large and asetose, lacking sulcus connecting them; antenna short; antennomeres moniliform, 7 enlarged, 9 and 10 with dense short setae on mesal surface. Discal stria of elytron extending posteriorly to more than 1/5 of elytral length. Legs almost simple, only mesotibia with small apical spur. Metaventrite with dense setose area admesally. Aedeagus strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, in lateral view narrowing apically; dorsal lobe narrowing apically, with pointed apex.</p> <p>Description. Male. Body (Fig. 18A) length 1.84 mm; color reddish-brown, head, pronotum and abdomen slightly darker than elytra, antennae and most parts of legs yellowish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 18B) sub-rectangular at base, slightly wider than long, length 0.39 mm, width across eyes 0.43 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae or mediobasal longitudinal sulcus, with short transverse sulcus between antennal tubercles, with few large punctures lateral sulcus; antennal tubercles weakly raised; frons slightly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two small gular foveae (posterior tentorial pits) in large oval opening, with distinct median carina extending from foveae anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 32 ommatidia. Antenna moderately short, length 0.97 mm, antennomere 7 and apical three antennomeres (Fig. 18C) forming distinct club; antennomere 1 thick, subcylindrical, 2–6 and 8 each moniliform, 7 strongly enlarged, 9 much larger than 8, 10 slightly short than 9, 9 and 10 with short, dense setae on mesal margin, 11 largest, slightly shorter than 9 and 10 combined, truncate at base and narrowed apically.</p> <p>Pronotum (Fig. 18B) approximately as long as broad, length 0.47 mm, width 0.44 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, median longitudinal sulcus slightly shorter than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea or mediobasal carina, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 2/3, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.64 mm, width 0.76 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to more than apical 1/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, sulcate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite broadly impressed at middle, admesal areas with long, dense setae; with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate, lacking distinct modifications; mesotibia (Fig. 18D) with tiny apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.43 mm, width 0.69 mm. Tergite 1 (IV) in dorsal view longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and one pair of transverse basolateral sockets, with pair of thick discal carinae; tergites 2 and 3 (V and VI) lacking foveae, 4 (VII) as long as 2 and 3 combined along middle, with one pair of basolateral foveae, 2–4 each with pair of thin lateral carinae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral sockets, with pair of long, thick lateral carinae; midlength of sternite 2 (IV) as long as sternites 3–5 (V–VII) combined, 3 and 4 short, 5 slightly longer than 4, 3–5 lacking foveae, sternite 6 (VIII) transverse, posterior margin emarginate at middle, sternite 7 (IX) (Fig. 18E) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 18F, G) 0.28 mm long, strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, ventral stalk in lateral view broad along basal 2/3 and narrowing apically; dorsal lobe elongate, broadest near middle, with pointed apex; parameres reduced to single broad membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species is allied with A. crassicornis by the similar modifications of antennomeres 9 and 10, setose metaventrite, and general form of the aedeagus. These two species can be readily separated by the lack of a median longitudinal sulcus of the vertex, the strongly enlarged antennomeres 7, and a much stouter form of the aedeagus of A. lebus. The setose metaventrite is also shared with A. nepaeformis, but the male of the latter species possesses strongly sinuate protibiae and modified protarsomeres 1, which are characteristic for that species.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 78C, 95A, B).</p> <p>Etymology. The new species epithet is a Latin adjective combined from ‘Lebu’ (after Lebu Valley) and suffix ‘- us (- a, - um)’ (of or from a place).</p></div> 	http://treatment.plazi.org/id/C822F213FFE72459CB88A252777925F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFE52447CB88A2BA761B25B2.text	C822F213FFE52447CB88A2BA761B25B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes lotus Yin 2022	<div><p>Arthromelodes lotus sp. nov.</p> <p>(Figs 19, 78C, 99F)</p> <p>Chinese common name: Ṑ+njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.3575&amp;materialsCitation.latitude=29.3125" title="Search Plazi for locations around (long 95.3575/lat 29.3125)">Renqingbeng Temple</a> (ca. 29°18’45”N, 95°21’27”E), 1760–2200 m, 19.iii.2017, Wen-Xuan Bi leg., [ƱẪẸṘ仁ů崩寺]’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.8 mm. Head subtruncate at base; vertexal foveae large and asetose, lacking sulcus connecting them; antenna elongate; antennomeres slightly elongate, lacking modifications. Discal stria of elytron extending posteriorly to approximately apical 1/3 of elytral length. Mesotrochanter with small ventral spine, mesotibia with triangular apical spur. Tergite 1 (IV) heavily modified, with large central cavity and projections inside. Aedeagus strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, ventral stalk dilated at apex; dorsal lobe broad at middle, narrowing towards base and apex.</p> <p>Description. Male. Body (Fig. 19A) length 1.78 mm; color dark reddish-brown, antennae, tibiae, tarsi and mouthparts yellowish-red. Dorsal surface of body covered with scattered long pubescence and normal ones.</p> <p>Head (Fig. 19B) subtruncate at base, slightly wider than long, length 0.37 mm, width across eyes 0.42 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae, mediobasal carina extending from head base anteriorly to transverse sulcus between antennal tubercles, post ocular areas and frons coarsely punctate; antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus; clypeus with scattered large punctures, its anterior margin carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with gular foveae (posterior tentorial pits) fused in single oval opening, with distinct median carina extending from middle anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 19 large ommatidia. Antenna moderately elongate, length 0.94 mm, simple, club loosely formed by moderately enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each slightly elongate, 3 and 8 smallest, 9 much larger than 8, 10 as long and as wide as 9, 11 largest, longer than 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 19B) approximately as long as broad, length and width 0.44 mm, widest at middle; lateral margins rounded, with few exceptionally long setae; disc slightly convex, finely punctate, median longitudinal sulcus as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea or mediobasal carina, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with small lateral procoxal foveae; hypomeral ridge extending from base anteriorly towards half of apical part, with lateral antebasal hypomeral impression; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.50 mm, width 0.61 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to approximately apical 3/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, sulcate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite broadly impressed at middle, admesal areas weakly prominent; with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate; protrochanter with tiny ventral spine; mesotrochanter (Fig. 19C) with small, acute spine on ventral margin, mesotibia (Fig. 19D) with distinct triangular apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.51 mm, width 0.56 mm. Tergite 1 (IV) (Fig. 19E) in dorsal view longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with pair of relatively long discal carinae, large central cavity occupying most surface of tergite, inside with large tubercles at apex of and two small laminae at posterolateral margins of cavity, with broad longitudinal sulcus lateral to each side of cavity; tergites 2 and 3 (V and VI) lacking foveae, 4 (VII) as long as 2 and 3 combined along middle, with one pair of basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral sockets, with pair of long, thick lateral carinae; midlength of sternite 2 (IV) as long as sternites 3–5 (V–VII) combined, 3 and 4 short, 5 slightly longer than 4, 3–5 lacking foveae, sternite 6 (VIII) transverse, posterior margin emarginate at middle, sternite 7 (IX) (Fig. 19F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 19G, H) 0.30 mm long, strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, ventral stalk dilated at apex and with pointed expansion before apex; dorsal lobe elongate, broadest near middle, narrowing towards base and apex; parameres reduced to single broad membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species resembles A. flosculus by the presence of exceptionally long setae along the posterolateral margin of the head, lateral margin of the pronotum, and elytral disc. Arthromelodes lotus can be readily separated from A. flosculus by the much smaller body size (1.78 mm vs. 2.62 mm), the more robust antennae, the presence of a large apical spur of the mesotibiae, and a different structure and form of the abdominal modification and the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 78C, 99F).</p> <p>Etymology. The specific epithet is a Latin adjective meaning ‘washed, bathed’, referring to Mêdog (Pemako) being regarded as the ‘Lotus Holy Land’ to the Tibetan Buddhists.</p></div> 	http://treatment.plazi.org/id/C822F213FFE52447CB88A2BA761B25B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFFB2445CB88A572719925B2.text	C822F213FFFB2445CB88A572719925B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes markam Yin 2022	<div><p>Arthromelodes markam sp. nov.</p> <p>(Figs 20, 79A, 100F)</p> <p>Chinese common name: Ẽ康njà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang (Chamdo City), Markam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.31528&amp;materialsCitation.latitude=29.552221" title="Search Plazi for locations around (long 98.31528/lat 29.552221)">Juebashan Pass</a> (29°33’8”N, 98°18’55”E), ca. 3900 m, 27.vi.2011, Wen-Xuan Bi leg., [ƱẪẼ康ẅ巴山垭口]’ (SNUC). PARATYPES: CHINA: 1 ♂, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.2 mm. Head subtruncate at base; vertex with complete, reversed U-shaped sulcus connecting relatively small, asetose vertexal foveae; antenna long, antennomeres each slightly elongate, lacking modifications. Discal stria of elytron extending to approximately apical 1/5 of elytral length. Mesotrochanter with small ventral tubercle, mesotibia distinctly constricted before apex and with large apical spur; metafemur strongly clavate, basal third narrowed and then expanded for apical two-thirds. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) with small central impression near posterior margin and pair of setose patches laterally. Aedeagus strongly asymmetric, ventral stalk of median lobe protruding and slightly curved, apex in dorsal view rounded, dorsal lobe broader than ventral stalk, parameres reduced and forming single membranous structure.</p> <p>Description. Male. Body (Fig. 20A) length 2.22–2.27 mm; color light reddish-brown, tarsi and mouthparts paler. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 20B) subtruncate at base, slightly broader than long, length 0.42–0.43 mm, width across eyes 0.45 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with complete, reversed U-shaped sulcus connecting foveae, mediobasal carina short, indistinct, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse opening, with thin median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 30 relatively large ommatidia. Antenna lacking distinct club and modification, length 1.08–1.13 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, distinctly shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 20B) approximately as long wide, length 0.45–0.47 mm, width 0.45 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, with median longitudinal sulcus slightly longer than lateral sulci in dorsal view; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part approximately as long as coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with indistinct lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.64–0.66 mm, width 0.73–0.76 mm; each elytron with two medium-sized, asetose basal foveae, lacking subbasal fovea; humerus slightly prominent; discal stria extending from outer basal fovea to apical 1/5 of elytral length; subhumeral fovea present, marginal stria extending posteriorly from fovea to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite weakly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore legs simple. Mesotrochanter with small ventral tubercle, mesotibia (Fig. 20C) distinctly constricted before apex and with large apical spur; metafemur strongly clavate, basal third narrowed and then expanded for apical two-thirds.</p> <p>Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.65–0.67 mm, width 0.69–0.72 mm. Tergite 1 (IV) (Fig. 20D) strongly modified, with small central impression near posterior margin, posterior margin of cavity with small half-round tubercle, with several modified setae along anterior margin of impression, areas lateral to impression with large setose patches; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with very thin discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) lacking basal sulcus, with mediobasal and basolateral foveae and pair of short lateral carinae; midlength of sternite 2 (IV) shorter than 3–5 (V–VII) combined, sternites 3–5 approximately subequal in length along midline, each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 20E) with weakly sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 20F, G) 0.34 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, lacking distinct basoventral projection, ventral stalk strongly protruding and slightly curved ventrally, apex in dorsal view rounded; dorsal lobe elongate and broader than ventral stalk, narrowing apically; parameres reduced to single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. This is a quite distinct species of which the males can be readily separated from those of all known congeners by the unique modification of the abdomen, the forms of the mesotibiae and metafemora, as well as the form of the aedeagus.</p> <p>Distribution. Markam County, Tibet, SW China (Figs 79A, 100F).</p> <p>Etymology. The new species is named after Markam County, where the type locality is situated.</p></div> 	http://treatment.plazi.org/id/C822F213FFFB2445CB88A572719925B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFF92443CB88A5727151229A.text	C822F213FFF92443CB88A5727151229A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes monba Yin 2022	<div><p>Arthromelodes monba sp. nov.</p> <p>(Figs 21, 79A, 95A, B)</p> <p>Chinese common name: Ḣ巴njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.2 mm. Head subtruncate at base; vertex with deep transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna moderately elongate, antennomeres slightly elongate or as long as wide, lacking modifications. Discal stria of elytron extending to apical 1/4 of elytral length. Fore and hind legs simple, mesotrochanter with apically setose protuberance on ventral margin, mesofemur with short spine at distal 1/3, mesotibia with tufted setae at apical 1/3 and long apical spine. Abdomen with large Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, tergites 1 (IV) prominent at middle. Aedeagus strongly asymmetric, median lobe in dorsal view markedly broad through length, dorsal lobe extremely elongate, strongly curved downwards apically, parameres reduced and forming single membranous structure.</p> <p>Description. Male. Body (Fig. 21A) length 2.21 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 21B) subtruncate at base, approximately as long as wide, length 0.45 mm, width across eyes 0.44 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), with deep, slightly curved transverse sulcus between antennal tubercles, distinct mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with foveae (posterior tentorial pits) in single small round opening, with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 30 moderately large ommatidia. Antenna 1.10 mm long, club formed by moderately enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each elongate, with 5–7 much longer than neighboring antennomeres, 8 smallest, 9 much larger than 8, 10 as wide as and slightly shorter than 9, 11 largest, simple, approximately as long as 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 21B) approximately as long as wide, length 0.50 mm, width 0.48 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus slightly longer than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with lateral antebasal impression; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.71 mm, width 0.80 mm; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humerus roundly prominent; discal stria extending from outer basal fovea to apical 1/4 of elytral length; subhumeral fovea small, marginal stria extending posteriorly from fovea to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae in transverse opening, broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle. Longitudinal marginal ridge of meso- and metaventrite complete.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 21C) with short, apically setose ventral spine, mesofemur (Fig. 21C) with short, apically curved spine at distal 1/3, mesotibia (Fig. 21D) with acute tuft of setae at apical 1/3, then strongly narrowing apically and with long, curved blade-like spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.66 mm, width 0.67 mm; with first dorsally visible tergite modified. Tergite 1 (IV) (Fig. 21E) longer than tergites 2–4 (V–VII) combined; with moderately large, round projection at middle of posterior margin, transverse area lateral to projection covered with dense, short setae, setose basal sulcus separated by broadly separated mediobasal and one pair of basolateral foveae, lacking discal carinae; tergites 2–3 (V–VII) lacking foveae or lateral carinae, tergite 4 (VII) as long as 2 and 3 combined along middle, with pair of complete lateral carinae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral foveae, lacking lateral carinae; midlength of sternite 2 (IV) slightly longer than sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 21F) slightly oval, moderately sclerotized, with scattered long setae along apical margin.</p> <p>Aedeagus (Fig. 21G, H) 0.39 mm long, strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, basoventral projection moderately long, ventral stalk in dorsal view markedly broad though entire length, in ventral view narrowing from base toward apex; dorsal lobe extremely elongate and strongly curved downwards at apex, gradually narrowing from base towards apex; parameres reduced to single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species belongs to a group of species centered on A. championi that bear a long apical spine of the mesotibia, and a strongly elongate and strongly curved dorsal lobe of the aedeagus. The male of Arthromelodes monba can be readily separated by the presence of an acute setose tuft of mesotibia, a spinose mesofemur, a unique form of tergal modification, as well as a dorsally markedly broad median lobe of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 79A, 95A, B).</p> <p>Etymology. The new species is named after the local Monba people (Ḣ巴⁂).</p></div> 	http://treatment.plazi.org/id/C822F213FFF92443CB88A5727151229A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFFF2441CB88A45A774821D8.text	C822F213FFFF2441CB88A45A774821D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes nepaeformis Yin 2022	<div><p>Arthromelodes nepaeformis sp. nov.</p> <p>(Figs 22, 79A, 95, 96D)</p> <p>Chinese common name: OE形njà甲</p> <p>Type material (55 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, Lebu Valley, pass to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Mama Vill.</a>, 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟 Þ玛乡 ’ (SNUC). PARATYPES: CHINA: 13 ♂♂, 6 ♀♀, same data as that of holotype; 25 ♂♂, 9 ♀♀, also from <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.85445&amp;materialsCitation.latitude=27.920277" title="Search Plazi for locations around (long 91.85445/lat 27.920277)">Lebu Valley</a>, except ‘ 27°55’13”N, 91°51’16”E, 3650-4050 m, 05.vii.2021 ’; 1 ♂ ‘ China: Xizang, Nyingchi City, valley nr. Jiare Vill., <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.77945&amp;materialsCitation.latitude=30.055277" title="Search Plazi for locations around (long 93.77945/lat 30.055277)">Chongge</a>, 30°03’19”N, 93°46’46”E, 3500-3750 m, 08.vii.2021, Peng, Yin, Zhang Ʊ Ẫffżẘ 热村ṗẋ ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.7–1.8 mm. Head subtruncate at base; vertex with deep transverse sulcus between antennal tubercles, lacking mediobasal carina, vertexal foveae moderately large and asetose; antenna relatively short, antennomeres each slightly elongate or moniliform, lacking modifications. Discal stria of elytron extending posteriorly to more than apical 1/3 of elytral length. Protibia strongly sinuate, protarsomere 1 with tuft of umbellate setae; mesotibia with long apical spine. Posteromedial area of metaventrite covered with dense, thickened setae. Abdomen with large Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, simple. Aedeagus strongly asymmetric, median lobe protruding apically, ventral stalk strongly curved in lateral view, dorsal lobe strongly curved at apex, parameres broad, weakly sclerotized. Female. Body length slightly over 1.7 mm, legs and metaventrite lacking modifications, genitalia as in Fig. 22I.</p> <p>Description. Male. Body (Fig. 22A) length 1.73–1.82 mm; bicolored, head, maxillary palpomere 4, pronotum, most parts of protibia and abdomen dark brown, antennae, elytra and legs (other than protibia) reddish-brown, tarsi and mouthparts (other than maxillary palpomere 4) lighter in color. Dorsal surface of body covered with relatively short pubescence.</p> <p>Head (Fig. 22B) subtruncate at base, approximately as long as wide, length 0.37–0.38 mm, width across eyes 0.38 mm; vertex finely punctate, with relatively large, asetose vertexal foveae (dorsal tentorial pits), with slightly curved, deep and short transverse sulcus between antennal tubercles, mediobasal carina absent, antennal tubercles moderately raised; frons broadly and weakly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 32 moderately large ommatidia. Antenna 0.86–0.87 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much larger than 8, approximately as long as wide, 10 as long as and slightly broader than 9, 11 large, as long as 9 and 10 combined.</p> <p>Pronotum (Fig. 22B) approximately as long as wide, length 0.41–0.42 mm, width 0.40–0.41 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus much shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pits; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.60 mm, width 0.71 mm; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humerus weakly prominent; discal stria extending from outer basal fovea to more than apical 1/3 of elytral length; subhumeral fovea small, marginal stria extending posteriorly from middle to apex.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, forked internally. Metaventrite (Fig. 22C) moderately impressed at middle, with pair of setose lateral metaventral foveae, posteromedial area covered with dense, thickened setae, with pair of angulate admesal projections, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, hind leg simple. Protibia (Fig. 22D) strongly sinuate, surface with linear microsculpture and erect setae, protarsomere 1 apically extended and with tuft of umbellate setae at apex; mesotibia (Fig. 22E) with long, slightly curved spine at apex.</p> <p>Abdomen much narrower than elytra, widest at basolateral margins of tergite 1 (IV), length 0.46–0.50 mm, width 0.60–0.61 mm; lacking modifications. Tergite 1 (IV) in dorsal view longer than tergites 2–4 (V–VII) combined; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with pair of thick, triangular discal carinae; tergites 2 and 3 (V and VI) lacking foveae, 4 (VII) as long as 2 and 3 combined along middle, with one pair of basolateral foveae, 2–4 each with pair of thin lateral carinae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral sockets, with pair of long, thick lateral carinae; midlength of sternite 2 (IV) as long as sternites 3–5 (V–VII) combined, 3 and 4 short, 5 slightly longer than 4, 3–5 lacking foveae, sternite 6 (VIII) transverse, posterior margin emarginate at middle, sternite 7 (IX) (Fig. 22F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 22G, H) 0.33 mm long, strongly asymmetric; median lobe with large basal capsule and subtriangular foramen, lacking basoventral projection, ventral stalk protruding, in lateral view strongly curved, narrowing towards apex; dorsal lobe elongate and strongly curved at apex; parameres fused to form broad plate, weakly sclerotized.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and metaventrite lacking modifications; each compound eye composed of approximately 22 ommatidia; humeral angle rounded; tergite 4 (VII) with thickened lateral ridges. Measurements (as for male): body length 1.72–1.78 mm; length/width of head 0.35–0.37/ 0.36–0.37 mm, pronotum 0.38/ 0.38–0.39 mm, elytra 0.51–0.52/ 0.65–0.67 mm; abdomen 0.50–0.58/ 0.61–0.62 mm; length of antenna 0.73–0.77 mm; maximum width of genitalia (Fig. 22I) 0.21 mm.</p> <p>Comparative notes. This species appears to be well isolated in the genus due to the unique bicolored body, and the oddly modified protibiae and tarsomeres 1 of the male. The setose metaventrite is shared with A. crassicornis and A. lebus, but the setae are of a different density and length.</p> <p>Distribution. Cona County and Chagyib District (Nyingchi), Xizang, SW China (Figs 79A, 95, 96D).</p> <p>Etymology. The new species name is a combination of ‘ nepa (a crab)’ and ‘ fôrma (shape)’, referring to the crab-like habitus of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FFFF2441CB88A45A774821D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFFC244FCB88A1DB741120CA.text	C822F213FFFC244FCB88A1DB741120CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes nomurai Yin 2022	<div><p>Arthromelodes nomurai sp. nov.</p> <p>(Figs 23, 79B, 97C, D)</p> <p>Chinese common name: Ǚ村氏njà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, 2190 m, 2019.vii.15–viii.01, FIT, Z.-W. Yin leg., [ƱẪffżḦËƖǎ]’ (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.0 mm. Head sub-rectangular at base; vertexal foveae large and asetose, with trapezoidal sulcus connecting foveae; lacking mediobasal carina; antenna moderately long, antennomeres each elongate, lacking modifications. Pronotum with median longitudinal sulcus much shorter than semicircular lateral ones. Discal stria of elytron extending to apical 1/3 of elytral length. Legs more or less robust; pro- and mesofemur thickened, mesotibia expanded mesally at basal 1/3 and with small apical protuberance, mesotrochanter with small, thin ventral spine. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 (IV) simple; sternite 2 (IV) with short, transverse median projection. Aedeagus strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, basoventral projection moderately long, ventral stalk relatively short, narrowing apically in lateral view, dorsal lobe extremely elongate, apical half strongly curved ventrally, parameres reduced and forming single membranous structure.</p> <p>Description. Male. Body (Fig. 23A) length 2.05 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with scattered exceptionally long pubescence and normal ones.</p> <p>Head (Fig. 23B) sub-rectangular at base, approximately as long as wide, length 0.38 mm, width across eyes 0.40 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with complete reversed trapezoidal sulcus connecting foveae, with pairs of long setae just posterior to anterior margin of sulcus and near head base, mediobasal carina absent; antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and slightly raised; ocular-mandibular carina complete. Venter with single small gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 30 ommatidia. Antenna distinctly clubbed and lacking modifications, length 0.90 mm; antennomere 1 thick, subcylindrical, 2–11 each elongate, 11 largest, slightly shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 23B) slightly longer than wide, length 0.44 mm, width 0.41 mm, widest at middle; lateral margins rounded, with long thickened, erect setae scattered on lateral margins and disc; disc slightly convex, finely punctate, median longitudinal sulcus much shorter than semicircular lateral longitudinal sulci; lacking median antebasal fovea, with complete transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal halves, with lateral hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra slightly wider than long, length 0.60 mm, width 0.69 mm; each elytron with two large, asetose basal foveae; humerus weakly prominent; discal stria extending from outer basal fovea to apical 1/3 of elytral length; small subhumeral fovea present, marginal stria extending from subbase to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae moderately separated, lateral mesoventral foveae large and setose. Metaventrite moderately impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate; pro- and mesofemur thickened; mesotrochanter with small ventral spine, mesotibia (Fig. 23C) expanded at basal 1/3, with small apical protuberance.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.59 mm, width 0.61 mm, with scattered exceptionally long setae on dorsal surface. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, lacking modifications, setose basal sulcus separated by mediobasal and one pair of broad basolateral sockets, lacking marginal or discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 4 (VII) much shorter than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) (Fig. 23D) with large mediobasal foveae and transverse basolateral sockets, with pair of short lateral carinae, with short, transverse median projection near posterior margin, midlength approximately as long as 3–5 (V–VII) combined, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 23E) with weakly sclerotized, rounded apex and few long setae along apical margin.</p> <p>Aedeagus (Fig. 23F–H) 0.30 mm long, strongly asymmetric; median lobe short, with large basal capsule and sub-triangular foramen, basoventral projection relatively long, ventral stalk narrowing apically in lateral view, with broad apex in dorso-ventral view; dorsal lobe strongly elongate and strongly curved ventrally, with pointed apex; parameres reduced and forming single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. Arthromelodes nomurai belongs to the A. championi group based on the aedeagal form. The new species can be readily separated from congeners by the unique modification of the mesotibia, as well as by the relatively much shorter ventral stalk of the aedeagus.</p> <p>Distribution. Chagyib District (Nyingchi), Xizang, SW China (Figs 79B, 97C, D).</p> <p>Etymology. The new species is named after the Japanese staphylinidologist Shûhei Nomura (Ǚ村周平) who described most of the species of Arthromelodes, and was the first to point out the similarity between Arthromelodes and Plocamarthrus.</p> </div>	http://treatment.plazi.org/id/C822F213FFFC244FCB88A1DB741120CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFF3244DCB88A7AF71D3216A.text	C822F213FFF3244DCB88A7AF71D3216A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes songxiaobini Yin 2022	<div><p>Arthromelodes songxiaobini sp. nov.</p> <p>(Fig. 24, 79B, 97C, D)</p> <p>Chinese common name: Ṫ氏njà甲</p> <p>Type material (7 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, nr. Bomê County, 29°52’22”N, 95°42’15”E, ca. 2735 m, 12.iii.2019, X.-B. Song leg., [ƱẪ波ṁ县ṉữ]’ (SNUC). PARATYPES: CHINA: 1 ♂, 3 ♀♀, same data as that of holotype’; 2 ♂♂, ‘ China: Xizang, Nyingchi City, Pailong Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.87583&amp;materialsCitation.latitude=29.983889" title="Search Plazi for locations around (long 94.87583/lat 29.983889)">Layue Bridge</a>, 29°59’02”N, 94°52’33”E, 2550 m, 13.vii.2021, Peng leg., Ʊ Ẫffż 市ḦË乡Ń月ss桥 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length 2.01–2.24 mm. Head subtruncate at base; vertexal foveae small, asetose, lacking sulcus connecting them; antenna moderately elongate; antennomeres slightly elongate to moniliform, antennomeres 9–11 enlarged to form indistinct club. Discal stria of elytron extending to apical 1/5 of elytral length. Mesotrochanter with thin, strongly curved protuberance, mesotibia expanded mesally at basal 1/3, with curved projection at apex. Tergite 5 (VIII) with broad and deep cavity at basal 1/2, projecting at middle apically. Aedeagus strongly asymmetric; median lobe with large basal capsule and basoventral projection, ventral stalk broad in dorsal view; dorsal lobe broad at middle and narrowing both basally and apically; parameres reduced to single membranous structure. Female. Body length 1.90–1.97 mm; antenna shorter, middle legs and abdomen lacking modifications, genitalia as in Fig. 24H.</p> <p>Description. Male. Body (Fig. 24A) length 2.01–2.24 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 24B) roundly-rectangular at base, approximately as long as wide, length 0.41 mm, width across eyes 0.42 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking Ushaped sulcus connecting foveae or mediobasal carina, with transverse, sinuate sulcus between antennal tubercles; antennal tubercles weakly raised; frons almost flat medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two small gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 28 ommatidia. Antenna moderately elongate, length 0.85–0.88 mm, apical three antennomeres forming indistinct club; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much larger than 8, 10 slightly longer and broader than 9, 11 largest, as long as 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 24B) approximately as long as broad, length 0.44–0.45 mm, width 0.45 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with small lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.64–0.65 mm, width 0.76–0.77 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to apical 1/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, sulcate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite broadly impressed at middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate; protrochanter with small ventral tubercle; mesotrochanter with thin, long and strongly curved protuberance, mesotibia (Fig. 24C) expanded mesally at basal 1/3, with distinct curved projection at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.55–0.70 mm, width 0.70–0.71 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, lacking discal carinae; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 5 (VIII) (Fig. 24D) semicircle, transverse, with large central cavity at basal 1/2, inside cavity four tubercles each with one seta, posterior margin of cavity with short, dense setae, apical half of tergite projecting medially. Sternite 2 (IV) with large mediobasal foveae and pair of broad, setose basolateral sockets, midlength longer than 3–5 (V–VII) combined, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, sternite 7 (IX) (Fig. 24E) with rounded apex, moderately sclerotized on apical half, membranous on basal half.</p> <p>Aedeagus (Fig. 24F, G) 0.30 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, ventral stalk short and broad in dorso-ventral view, round at apex; dorsal lobe broad at middle and narrowing both basally and apically, with pointed apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 18 ommatidia; humeral angle rounded; legs and abdomen lacking modifications. Measurements (as for male): body length 1.90–1.97 mm; length/width of head 0.39–0.41/ 0.41 mm, pronotum 0.42–0.43/ 0.43 mm, elytra 0.59/ 0.72–0.73 mm; abdomen 0.53–0.62/ 0.68–0.69 mm; length of antenna 0.80–0.81 mm; maximum width of genitalia (Fig. 24H) 0.26 mm.</p> <p>Comparative notes. The mesally expanded mesotibiae of the male of the new species is shared only with A. nomurai. These two species can be readily separated by the medially concave male tergite 5 (VIII) and the characteristic form of the aedeagus of A. songxiaobini.</p> <p>Distribution (Fig. 96). Bomê County and Chagyib District (Nyingchi), Tibet, SW China (Figs 79B, 97C, D).</p> <p>Etymology. The new species is named after Xiao-Bin Song, the collector of the holotype.</p></div> 	http://treatment.plazi.org/id/C822F213FFF3244DCB88A7AF71D3216A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFF0244ACB88A1DB749F27F2.text	C822F213FFF0244ACB88A1DB749F27F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes speciosus Yin 2022	<div><p>Arthromelodes speciosus sp. nov.</p> <p>(Figs 25, 79B, 90)</p> <p>Chinese common name: ŁƟnjà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32806&amp;materialsCitation.latitude=28.376945" title="Search Plazi for locations around (long 85.32806/lat 28.376945)">Gyirong Valley</a>, slope nr. Jifu Vill., 28°22’37”N, 85°19’41”E, 2400–2700 m, 22.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW 吉ė村ṉữ山坡 ’ (SNUC).</p> <p>Diagnosis. Male. Body length over 2.5 mm. Head sub-rectangular at base; vertex with deep transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna elongate, lacking modifications; antennomeres 1–7 each slightly elongate, 8 shortest sub-moniliform, 9 strongly modified, 10 asymmetric, 11 much shorter than 9 and 10 combined. Pronotum enlarged, distinctly broader than head. Discal stria of elytron thin and shallow, extending posteriorly to approximately apical 1/6 of elytral length. Metaventrite densely setose admesally. Fore and hind legs simple, mesotrochanter with small ventral tubercle, mesotibia moderately thickened, with tiny apical protuberance. Abdomen lacking modifications, with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined. Aedeagus strongly asymmetric, ventral stalk elongate and dilated at apically, dorsal lobe extremely elongate and strongly bent downwards apically, parameres reduced and forming single membranous structure.</p> <p>Description. Male. Body (Fig. 25A) length 2.57 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with dense, moderately long pubescence.</p> <p>Head (Fig. 25B) sub-rectangular at base, slightly wider than long, length 0.54 mm, width across eyes 0.57 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), with deep, slightly curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles weakly prominent; frons almost flat, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with small gular foveae (posterior tentorial pits) in single pit, with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 large ommatidia. Antenna 1.36 mm long, with modified antennomeres 9 and 10 (Fig. 25C); antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, sub-moniliform, much shorter and narrower than 7, 9 strongly dilated, with large apicomesal projection, 10 asymmetric, with oblique nasal margin, 11 largest, much shorter than 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 25B) enlarged, much wider than head (width of pronotum / head) and slightly wider than long, length 0.61 mm, width 0.66 mm, widest at middle; lateral margins moderately expanded and rounded; disc weakly convex, finely punctate, with median longitudinal sulcus approximately as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.90 mm, width 0.69 mm, constricted at bases; each elytron with two large basal foveae, lacking subbasal fovea; humeral prominence barely visible; discal stria extending from outer basal fovea to apical 1/6 of elytral length; small subhumeral fovea present, thin marginal stria extending posteriorly from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite slightly impressed at middle, with pair of setose lateral metaventral foveae, admesal areas strongly convex, densely setose; posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Procoxa with exceptionally elongate seta at base. Mesotrochanter with small ventral tubercle, mesotibia with tiny, indistinct tubercle at apex.</p> <p>Abdomen as broad as elytra, widest at lateral margins of tergite 1 (IV), length 0.82 mm, width 0.87 mm, lacking modifications. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; broadly separated mediobasal fovea and one pair of basolateral foveae separating basal sulcus, with pair of short discal carinae; tergites 2 and 3 (V and VI) each short, tergite 4 (VII) approximately as long as 2 and 3 combined along middle, tergites 2–4 (V–VII) each with one pair of small basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal foveae and large basolateral sockets, with setose basal impression laterally, with pair of basolateral carinae; midlength of sternite 2 (IV) approximately as long as sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin almost straight, sternite 7 (IX) (Fig. 25D) slightly oval, moderately sclerotized, apical margin rounded, with long setae scattered along apical margin.</p> <p>Aedeagus (Fig. 25E, F) 0.62 mm long, strongly asymmetric; median lobe with enlarged basal capsule and elongate foramen, lacking basoventral projection, ventral stalk long and laterally broad; dorsal lobe extremely elongate and gradually narrowing from middle towards apex, apical portion strongly curved downwards, with pointed apex; parameres reduced to single membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species belongs to the A. championi group by sharing an extremely elongate endophallus of the aedeagus that is strongly curved apically. It is close to A. championi by the similar placement of the antennal modifications. The relatively enlarged pronotum and the uniquely modified male antennomeres 9 and 10 lead to a quick separation of the new species from A. championi.</p> <p>Distribution. Gyirong County, Tibet, SW China (Figs 79B, 90).</p> <p>Etymology. The specific epithet ‘ speciosus (good-looking, beautiful)’ is a Latin adjective referring to the modification of male antennal clubs.</p> </div>	http://treatment.plazi.org/id/C822F213FFF0244ACB88A1DB749F27F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFF62448CB88A0B274082206.text	C822F213FFF62448CB88A0B274082206.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes torus Yin 2022	<div><p>Arthromelodes torus sp. nov.</p> <p>(Figs 26, 79C, 93C, E)</p> <p>Chinese common name: 瘤únjà甲</p> <p>Type material (20 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Dinggyê County, pass to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Zhêntang Town</a>, 27°53’58”N, 87°27’21”E, 3060 m, 28.vi.2021, Yin &amp; Zhang leg., ƱẪŤĠ县ĿƋ沟公ḡṅðff ’ (SNUC). PARATYPES: CHINA: 6 ♂♂, 8 ♀♀, same data as that of holotype; 5 ♂♂, also from Zhêntang, except ‘ Xiuxiongma Vill., 27°54’11”N, 87°22’42”E, 2700-3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJŤĠ县Ŀ ƋWšŏ玛村 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.6 mm. Head subtruncate at base; vertex with transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae relatively small and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 smallest, 11 slightly shorter than 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately apical 2/5 of elytral length. Fore and hind legs simple, mesotrochanter with setose tuft on ventral margin, mesotibia with large apical spine. Abdomen with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; tergite 1 with strongly raised protuberance at middle. Aedeagus strongly asymmetric, median lobe with large basal capsule and oval foramen, ventral stalk narrowed and protruding, dilated at apex, dorsal lobe elongate, forked at apex, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.5 mm, legs and abdomen lacking modifications, genitalia as in Fig. 26I.</p> <p>Description. Male. Body (Fig. 26A) length 2.64 mm; color reddish-brown, tarsi and mouthparts slightly lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 26B) subtruncate at base, approximately as long as wide, length 0.52–0.55 mm, width across eyes 0.54–0.55 mm; vertex finely punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with distinct transverse sulcus between antennal tubercles, mediobasal carina extending from head base anteriorly to sulcus, antennal tubercles moderately raised; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina present, thin carina present below posteroventral articulation of mandible. Venter with tiny gular foveae (posterior tentorial pits) in single small pit, median carina extending from fovea anteriorly to mouthpart, with thin basal half and thick apical half. Eyes moderately prominent, composed of approximately 25 ommatidia. Antenna 1.34–1.38 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–8 each elongate, 8 smallest, much shorter and slightly narrower than 7, 9 longer but narrower than 10, 11 largest, shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 26B) slightly longer than wide, length 0.60–0.62 mm, width 0.58–0.59 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus as long as semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.81–0.82 mm, width 0.91–0.94 mm, constricted at bases; each elytron with two moderately-sized, widely separated basal foveae, lacking subbasal fovea; humeral angle rounded; discal stria extending from outer basal fovea to approximately apical 2/5 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from posterior to middle to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter (Fig. 26C) with moderately long setose tuft on ventral margin, mesotibia (Fig. 26D) with long spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.77–0.79 mm, width 0.85–0.86 mm; with modified tergite 1 (IV). Tergite 1 (Fig. 26E) longer than tergites 2–4 (V–VII) combined, with single, strongly raised projection at middle, degree of development of projection slightly varying among individuals, basal impression broken by mediobasal and basolateral foveae, with pair of short discal carinae, with incomplete outer marginal carina; tergites 2 and 3 (V and VI) subequal in length along midline, 4 (VII) slightly longer than 2 and 3 combined along middle, 2–4 (V–VII) each with one pair of small basolateral foveae, 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with two mediobasal foveae and large basolateral sockets, lacking basal impression or sulcus, with pair of short lateral carinae; midlength of sternite 2 (IV) as long as 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 26F) slightly oval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 26G, H) 0.42 mm long, strongly asymmetric; median lobe with large basal capsule and oval foramen, basoventral projection short, ventral stalk protruding and in lateral view dilated at apex; dorsal lobe recumbent, apically forked into two lobes; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking spines/projections or modifications; each compound eye composed of approximately 23 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.53–2.61 mm; length/width of head 0.53–0.54/ 0.53 mm, pronotum 0.56–0.57/ 0.56 mm, elytra 0.72–0.76/ 0.89–0.90 mm; abdomen 0.77–0.80/ 0.85–0.87 mm; length of antenna 1.26–1.27 mm; maximum width of genitalia (Fig. 26I) 0.30 mm.</p> <p>Comparative notes. Arthromelodes torus is morphologically most similar to A. alpitorus by sharing most external characters, as well as a similar male tergal modification and the form of the aedeagus. The new species can be separated by the smaller body size (2.64 mm vs. 2.80–2.88 mm for male; 2.53–2.61 mm vs 2.66–2.80 mm for female), a much longer median longitudinal sulcus of the pronotum, the male having a thinner and slightly curved spine at the apex of the mesotibia, the median projection of tergite 1 (IV) with single posteromedial corner (with acute posterolateral angles in A. alpitorus), the area posterior to the projection flat along the midline (raised in A. torus), and a less expanded apex of the median lobe of the aedeagus. These two species appear to be well isolated by altitudinal barriers. Arthromelodes torus is also close to A. angulatus, A. aniqiao, A. complexus, A. lage, and A. langjicuo, sharing with them a setose male mesotrochanter, as well as a similar general form of the aedeagus. The male of the new species can be readily separated by the unique presence of a large central projection on tergite 1 (IV), as well as the long apical spine of the mesotibia.</p> <p>Distribution (Fig. 92). Dinggyê County, Tibet, SW China (Figs 79C, 93C, E).</p> <p>Etymology. The new species name ‘ torus (swelling, protuberance)’ is a Latin noun referring to the protuberant male tergite 1 (IV).</p> </div>	http://treatment.plazi.org/id/C822F213FFF62448CB88A0B274082206	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFF42437CB88A5E6713B21E6.text	C822F213FFF42437CB88A5E6713B21E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes zhangmu Yin 2022	<div><p>Arthromelodes zhangmu sp. nov.</p> <p>(Figs 27, 79C, 92C)</p> <p>Chinese common name: 樟木njà甲</p> <p>Type material (7 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyalam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.983055&amp;materialsCitation.latitude=27.987501" title="Search Plazi for locations around (long 85.983055/lat 27.987501)">Zhangmu Town</a> (樟木W), 27°59’15”N, 85°58’59”E, 2200 m, leaf litter, sifted, 17.vii.2010, Jian-Qing Zhu leg.’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 2 ♀♀, same locality as holotype, except ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.97806&amp;materialsCitation.latitude=27.955833" title="Search Plazi for locations around (long 85.97806/lat 27.955833)">Lixin Village</a> (ĒẾ村), 27°57’21”N, 85°58’41”E, 2400– 2600 m, 27.vii.2010 ’; 1 ♂, same locality, except ‘ 2600 m, 18.vii.2010 ’; 1 ♀, same locality, except ‘ 27°59’15”N, 85°58’59”E, 2250 m, 22.vii.2010 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length over 2.47–2.64 mm. Head sub-rectangular at base; vertex with deep transverse sulcus between antennal tubercles and long mediobasal carina, vertexal foveae moderately large and asetose; antenna elongate, lacking modifications; antennomeres each slightly elongate, 8 shortest, 9 and 10 approximately subequal in width, 9 longer than 10, 11 much shorter than 9 and 10 combined. Discal stria of elytron thin and shallow, extending posteriorly to approximately apical 1/5 of elytral length. Fore legs simple, mesotrochanter with short ventral spine, mesofemur moderately thickened, mesotibia with large apical, metatrochanter with lamellar projection. Abdomen lacking modifications, with large tergite 1 (IV) longer than tergites 2–4 (V–VII) combined. Aedeagus strongly asymmetric, median lobe in dorsal view broad, dorsal lobe strongly elongate, ventral stalk strongly curved and with pointed apex, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.24–2.39 mm, legs lacking modifications, genitalia as in Fig. 27I.</p> <p>Description. Male. Body (Fig. 27A) length 2.47–2.64 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with dense, moderately long pubescence.</p> <p>Head (Fig. 27B) sub-rectangular at base, wider than long, length 0.50–0.52 mm, width across eyes 0.51–0.54 mm; vertex finely punctate, with relatively large, asetose vertexal foveae (dorsal tentorial pits), with deep, curved transverse sulcus between antennal tubercles, mediobasal carina extending from near head base anteriorly to sulcus, antennal tubercles weakly prominent; frons almost flat, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single gular fovea (posterior tentorial pit), with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 large ommatidia. Antenna 1.20–1.26 mm long, lacking modifications; antennomere 1 thick, subcylindrical, 2–11 each elongate, 8 smallest, slightly shorter and narrower than 7, 9–10 successively shorter and broader, 11 largest, much shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 27B) slightly wider than long, length 0.55–0.60 mm, width 0.55–0.59 mm, widest at middle; lateral margins moderately expanded and rounded; disc weakly convex, finely punctate, with median longitudinal sulcus slightly longer than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge present at basal 1/2, extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.76–0.80 mm, width 0.90–0.98 mm, narrowed at base; each elytron with two large basal foveae, lacking subbasal fovea; humerus weakly prominent; discal stria extending from outer basal fovea to apical 1/5 of elytral length; small subhumeral fovea present, thin marginal stria extending posteriorly from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite slightly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate, fore legs simple. Mesotrochanter (Fig. 27C) with short ventral spine at base, mesofemur moderately thickened, mesotibia (Fig. 27D) with large spine at apex, metatrochanter (Fig. 27E) with truncate, lamellar projection on ventral margin.</p> <p>Abdomen as broad as elytra, widest at lateral margins of tergite 1 (IV), length 0.67–0.73 mm, width 0.84–0.89 mm, lacking modifications. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined; broadly separated mediobasal fovea and one pair of basolateral foveae separating basal sulcus, with pair of short discal carinae; tergites 2 and 3 (V and VI) each short, tergite 4 (VII) approximately as long as 2 and 3 combined along middle, tergites 2–4 (V–VII) each with one pair of small basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal foveae and large basolateral sockets, with setose basal impression laterally, with pair of basolateral carinae; midlength of sternite 2 (IV) approximately as long as sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 27F) slightly oval, moderately sclerotized, apical margin rounded.</p> <p>Aedeagus (Fig. 27G, H) 0.42 mm long, strongly asymmetric; median lobe with moderately large basal capsule and elongate foramen, basoventral projection relatively large, ventral stalk long and broad in dorsal view; dorsal lobe elongate and gradually narrowing from middle towards apex, apical part strongly curved ventrally, with pointed apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; legs lacking modifications; each compound eye composed of approximately 18 ommatidia; humeral angle rounded. Measurements (as for male): body length 2.24–2.39 mm; length/width of head 0.48–0.49/ 0.49–0.50 mm, pronotum 0.52–0.53/ 0.50–0.51 mm, elytra 0.60–0.66/ 0.83–0.88 mm; abdomen 0.63–0.64/ 0.75–0.81 mm; length of antenna 1.17–1.19 mm; maximum width of genitalia (Fig. 27I) 0.29 mm.</p> <p>Comparative notes. The new species is morphologically most similar to the sympatrically distributed A. latithorax, sharing the laterally expanded pronotal margins, the thickened mesofemur, and a large apical spine of the mesotibia. These two species can be readily separated by the lesser expansion of the pronotal lateral margins, the less curved mesotibia and apical spine, a dorsally much broader ventral lobe of the aedeagus, and the different structure of the female genitalia.</p> <p>Distribution. Nyalam County, Tibet, SW China (Figs 79C, 92C).</p> <p>Etymology. The new species is named after its type locality, i. e., Zhangmu Town.</p></div> 	http://treatment.plazi.org/id/C822F213FFF42437CB88A5E6713B21E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF8A2434CB88A75E713D273A.text	C822F213FF8A2434CB88A75E713D273A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arthromelodes zhentangensis Yin 2022	<div><p>Arthromelodes zhentangensis sp. nov.</p> <p>(Figs 28, 79C, 93E)</p> <p>Chinese common name: ĿƋnjà甲</p> <p>Type material (3 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Zhêntang Town</a>, Xiuxiongma Vill., 27°54’11”N, 87°22’42”E, 2700–3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJŤĠ 县ĿƋWšŏ玛村 ’ (SNUC). PARATYPES: CHINA: 1 ♂, 1 ♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.0 mm. Head subtruncate at base; vertex lacking distinct sulcus, with long median carina, vertexal foveae moderately large and asetose; antenna relatively short, antennomeres each sub-moniliform to slightly elongate, lacking modifications. Discal stria of elytron extending to approximately apical 1/3 of elytral length. Protibia with angulate projection before apex. Abdomen with large Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, with central impression at posterior half. Aedeagus strongly asymmetric, ventral stalk and dorsal lobe strongly twisted or curved dorsoventrally, parameres reduced and forming single membranous structure. Female. Body length slightly over 2.0 mm, antenna and legs lacking modifications, genitalia as in Fig. 28H.</p> <p>Description. Male. Body (Fig. 28A) length 2.06–2.08 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 28B) subtruncate at base, transverse, length 0.44 mm, width across eyes 0.50–0.51 mm; vertex mostly finely punctate, area posterior to antennal tubercle with coarse punctation, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking distinct sulcus between antennal tubercles, long median carina extending from head base anteriorly to apex of frons, antennal tubercles weakly raised; frons convex medially, confluent with clypeus; clypeus with coarse surface, its anterior margin carinate and moderately raised; thick ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter strongly convex at apical 1/2, with gular foveae (posterior tentorial pits) in small, shared opening, with distinct median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 28 moderately large ommatidia. Antenna 1.07–1.08 mm long, antennomeres 9–11 forming indistinct club; antennomere 1 thick, subcylindrical, 2 distinctly elongate, 3–8 each slightly elongate or sub-moniliform, with 7 much longer and broader than 6 and 8, 9 and 10 of similar size, moderately enlarged, 11 largest, as long as 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 28B) approximately as long as wide, length 0.49–0.50 mm, width 0.50–0.51 mm, widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with median longitudinal sulcus much shorter than semicircular lateral sulci in dorsal view; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae; outer and inner pair of basolateral foveae distinct, mediobasal carina short and indistinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.67 mm, width 0.81–0.83 mm; each elytron with two large, widely separated basal foveae, lacking subbasal fovea; humerus roundly prominent; discal stria extending from outer basal fovea to more than apical 1/3 of elytral length; subhumeral fovea absent, marginal stria extending posteriorly from basal 1/3 to apex of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae in transverse opening, broadly separated, lateral mesoventral foveae large and setose, not forked internally. Metaventrite broadly and shallowly impressed at middle, with pair of setose lateral metaventral foveae, posterior margin with small and narrow split at middle. Longitudinal marginal stria of meso- and metaventrite complete.</p> <p>Legs moderately elongate, middle and hind legs simple. Protibia (Fig. 28C) with roundly angulate protuberance near apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.54–0.55 mm, width 0.70–0.73 mm; lacking modifications. Tergite 1 (IV) (Fig. 28D) longer than tergites 2–4 (V–VII) combined; posterior half with oval, transverse and moderately deep impression at middle, anterior margin of impression with dense setae, forming setose edge; setose basal sulcus separated by mediobasal and one pair of basolateral foveae, discal carinae distinct; tergites 2–4 (V–VII) each with one pair of small basolateral foveae, tergite 4 (VII) approximately as long as 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and broad basolateral foveae, with pair of distinct lateral carinae; midlength of sternite 2 (IV) shorter than sternites 3–5 (V–VII) combined, 3–5 each with one pair of tiny basolateral foveae, sternite 6 (VIII) transverse, posterior margin evenly curved, sternite 7 (IX) (Fig. 28E) roundly quadrate, weakly sclerotized, with scattered a few setae along apical margin.</p> <p>Aedeagus (Fig. 28F, G) 0.31 mm long, strongly asymmetric; median lobe with large, expanded basal capsule and elongately oval foramen, basoventral projection short, ventral stalk in dorsal view broad and strongly twisted apically; dorsal lobe broad at base and narrowing apically, apical 1/2 strongly bent in dorsal view; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter; protibia and abdomen lacking modifications; each compound eye composed of approximately 19 ommatidia; elytra shortened, constricted at bases, humeral angle rounded. Measurements (as for male): body length 2.08 mm; length/width of head 0.43/ 0.48 mm, pronotum 0.47/ 0.48 mm, elytra 0.58/ 0.77 mm; abdomen 0.64/ 0.77 mm; length of antenna 1.01 mm; maximum width of genitalia (Fig. 28H) 0.31 mm.</p> <p>Comparative notes. Arthromelodes zhentangensis has a transverse head that lacks a distinct sulcus on the vertex, and possesses a long median carinae on the medially convex frons. When combined with the angulate protibiae, posteriorly placed oval impression of tergite IV, and the unique form of the aedeagus of the male, the new species can be readily separated from all other members of the genus.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 79C, 93E).</p> <p>Etymology. The new species is named after its type locality, i. e., Zhêntang Town.</p></div> 	http://treatment.plazi.org/id/C822F213FF8A2434CB88A75E713D273A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF882433CB88A7F575A227D6.text	C822F213FF882433CB88A7F575A227D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batriscenaulax Jeannel 1958	<div><p>Batriscenaulax Jeannel, 1958</p> <p>Chinese common name: ậḇà甲¤</p> <p>Batriscenaulax Jeannel, 1958: 57; Newton &amp; Chandler 1989 (catalog); Nomura 1991 (description, key to Japanese species). Type species: Batrisus modestus Sharp, 1874: 116 (original designation).</p> <p>The genus Batriscenaulax includes five species distributed in Japan, all share modified protibiae and tergite 1 (IV) in the male and a constricted basal capsule of the aedeagus (Nomura 1991). The three Japanese species of the genus Physomerinus Jeannel having the similar forms of the aedeagi and exhibiting male sexual characters on metafemora may be congeneric with Batriscenaulax. The major problem is the type species of Physomerinus, Batrisus septemfoveolatus Schaufuss, L. W., 1877 distributed in Thailand and Vietnam, has an enlarged basal capsule of the aedeagus and apicolateral trichomes on first antennomeres (Jeannel 1952; Nomura pers. comm.), a condition congruent with members of Batrisiella Raffray. Reexamination of the type and all included species of Physomerinus would help to clarify the identities of its included members.</p> <p>One new species of Batriscenaulax is described here, which appears to fit well the current concept of the genus.</p> </div>	http://treatment.plazi.org/id/C822F213FF882433CB88A7F575A227D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF8F2431CB88A09974B8243E.text	C822F213FF8F2431CB88A09974B8243E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batriscenaulax campestris Yin 2022	<div><p>Batriscenaulax campestris sp. nov.</p> <p>(Fig. 29, 80A, 99A)</p> <p>Chinese common name: 低地ậḇà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Tibet, Mêdog County, Beibeng Vill. (Ü崩村, ca. 29°14’36”N, 95°10’22”E), 800-1000 m, 01.viii.2014, Wen-Xuan Bi leg.’ (SNUC). PARATYPE: CHINA: 1 ♂, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.80–1.85 mm. Head sub-rectangular at base; vertexal foveae large and asetose, lacking sulcus connecting them; antenna elongate; antennomeres slightly elongate, lacking modifications. Discal stria of elytron long, extending posteriorly to approximately apical 15.5% of elytral length. Protibia with triangular setose brush at apicomesal margin; mesotibia with relatively long apical spine. Tergite 1 (IV) modified, with semi-cinquefoil central cavity and trichomes inside or along margins, with small setose patches lateral to cavity. Aedeagus strongly asymmetric; median lobe with constricted basal capsule and small foramen, ventral stalk broad and bi-sinuate, pointed towards apex; dorsal lobe elongate, flat in lateral view, narrowing towards apex.</p> <p>Description. Male. Body (Fig. 29A) length 1.81–1.86 mm; color reddish-brown, antennae, tibiae, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 29B) rounded triangular, sub-rectangular at base, slightly wider than long, length 0.35–0.36 mm, width across eyes 0.42–0.43 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae, mediobasal carina short, faint; antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus; clypeus smooth, its anterior margin carinate and moderately raised; ocular-mandibular carina complete, weakly presented. Venter with small gular foveae (posterior tentorial pits) narrowly separated, with distinct median carina extending from foveae anteriorly to mouthparts. Eyes strongly prominent, composed of approximately 30 large ommatidia. Antenna moderately elongate, length 0.96–1.09 mm, simple, club formed by moderately enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each elongate, 8 shortest, 9 much larger than 8, 10 as long and as wide as 9, 11 largest, slightly shorter than 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 29B) approximately as long as broad, length 0.45–0.46 mm, width 0.47–0.48 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, median longitudinal sulcus broad, approximately as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea or mediobasal carina, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with small lateral procoxal foveae; hypomeral ridge short, present only at basal part, with lateral antebasal hypomeral impression; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.65–0.67 mm, width 0.77–0.80 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria long, carinate, extending from outer basal fovea to approximately apical 15.5% of elytral length; humerus angulate, with small subhumeral fovea at middle of elytral flank, sulcate marginal stria extending from fovea to posterior margin of elytron, with thin, curved carinate sulcus extending anteriorly from fovea to humerus.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite broadly impressed at middle, admesal areas moderately prominent; with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin broadly emarginate, lacking split at middle.</p> <p>Legs moderately elongate; protrochanter with tiny ventral protuberance, protibia (Fig. 29C) with small triangular setose brush at apicomesal margin; mesotibia (Fig. 29D) with relatively long spine at apex.</p> <p>Abdomen compressed, widest at lateral margins of tergite 1 (IV) (Fig. 29E), length 0.47–0.50 mm, width 0.71–0.72 mm. Tergite 1 (IV) in dorsal view longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and two pairs of basolateral foveae, lacking discal carinae, semi-cinquefoil central cavity deep, with pair of trichomes along anterior margin, inside cavity one transverse projection and two long trichomes, with small setose patch lateral to each side of cavity, with thin, complete marginal stria; tergites 2 and 3 (V and VI) lacking foveae, 4 (VII) twice as long as 2 and 3 combined along middle, with one pair of basolateral foveae, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle. Sternite 2 (IV) with mediobasal and two basolateral foveae, with pair of long, thick lateral carinae; midlength of sternite 2 (IV) as long as sternites 3–5 (V–VII) combined, 3 and 4 short, 5 slightly longer than 4, 3–5 lacking foveae, sternite 6 (VIII) transverse, posterior margin emarginate at middle, sternite 7 (IX) (Fig. 29F) weakly sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 29G, H) 0.27 mm long, strongly asymmetric; median lobe with constricted basal capsule and small, round foramen, ventral stalk broad at base, bi-sinuate, narrowing towards apex; dorsal lobe strongly bent at middle, flat in lateral view, with narrowed apex; parameres reduced to single broad membranous structure.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species resembles its five Japanese congeners that have the male protibiae with an apical setose brush and a modified tergite 1 (IV). Batriscenaulax campestris can be readily separated by the larger elytra in relation to the head and pronotum, the apical brushes located on the mesal margin of the protibiae, and the more strongly sinuate ventral stalk of the aedeagus. All the Japanese species of Batriscenaulax have the male protibiae moderately to strongly swollen near their apices, and have the preapical setose brush located on the lateral margins of the tibiae.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 80A, 99A).</p> <p>Etymology. The specific epithet is a Latin adjective meaning ‘field, plain’, indicating the species was collected from a lowland area of Tibet.</p></div> 	http://treatment.plazi.org/id/C822F213FF8F2431CB88A09974B8243E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF8D2430CB88A3FE70FB277D.text	C822F213FF8D2430CB88A3FE70FB277D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella Raffray. Reexamination 1904	<div><p>Batrisiella Raffray, 1904</p> <p>Chinese common name: 小毛唇à甲¤</p> <p>Batrisiella Raffray, 1904: 59; Jeannel 1960: 426 (key to Indian and Pakistani species), 1961: 436 (key to Sri Lankan species); Löbl &amp; Kurbatov 2001 (redescription, key to Sri Lankan species). Type species: Eubatrisus caviventris Raffray, 1894a: 52 (monotypy).</p> <p>= Arthromelinus Jeannel, 1952: 98. Type species: Batrisus angulatus Raffray, 1893: 487 (original designation). Synonymized by Jeannel 1960: 426.</p> <p>The genera Batrisiella and Batriscenellus Jeannel from the Oriental Region and the African Arthromelus Jeannel share a similar external morphology and a large basal capsule with a basally recumbent dorsal lobe of the aedeagus. The first two genera were separated by the presence/absence of a trichome at the apicolateral margin of the scape (e.g., Nomura 1991; Yin et al. 2011b; Wang &amp; Yin 2015; Jiang &amp; Yin 2017), with Batrisiella lacking such a structure nor a glandular orifice (Löbl &amp; Kurbatov 2001: 671). In this paper, eight species are added to the Tibetan fauna, all possessing an inner glandular structure of varying size (only visible in cleared specimens put in liquid) at the apicolateral or lateral margin of the scape, and two species (B. ganma, B. zhujianqingi) have a tiny apicolateral trichome composed of short, thickened setae. It is most likely that such a glandular structure further develops into large, distinct trichomes and is widely represented in Asian species. Thus, there will be no discreet character separating Batrisiella from Batriscenellus. No formal synonymy is proposed here until the types and a wide range of known species of these groups can be examined, but all new species described here are placed in the oldest genus, i. e., Batrisiella. The eight species can be separated in the following key.</p> <p>Key to Tibetan species of Batrisiella (male)</p> <p>1 Sexual character present on profemur..................................................................... 2</p> <p>- Profemur unmodified, sexual character present on other body parts.............................................. 4</p> <p>2 Metatibia slender, unmodified; vertex with thin median carina (Fig. 30B); ventral stalk of median lobe in lateral view broad, recumbent (Fig. 30H).................................................................... B. acclivis sp. nov.</p> <p>- Metatibia broadened at distal 1/2; vertex lacking median carina (Fig. 31B); ventral stalk of median lobe in lateral view slender, erect (Fig. 31H)........................................................................ B. beibeng sp. nov.</p> <p>3 Metafemur with angulate tubercle at distal 1/3 (Fig. 34E), metatibia sinuate at basal 1/3 (Fig. 30F)... B. curvitibialis sp. nov.</p> <p>- Metafemur and metatibia simple, unmodified............................................................... 4</p> <p>4 Antennomeres 8 and 9 with short, thickened setae, 10 constricted at base and dilated at apex (Fig. 32C)....................................................................................................... B. bomea sp. nov.</p> <p>- Antennomeres 8–10 simple, unmodified................................................................... 5</p> <p>5 Tergite 4 (VII) with disc-like lateral protuberances, 5 (VIII) with broad and deep central cavity (Fig. 36C); sternites 2 and 3 (IV and V) projecting and 4 (VI) with long, thick setae (Fig. 36E, F).................................. B. guttata sp. nov.</p> <p>- Tergites 4 and 5 (VII and VIII) and sternites 2–4 (IV–VI) unmodified............................................ 6</p> <p>6 Vertex with distinct median sulcus (Fig. 33B)................................................. B. concisa sp. nov.</p> <p>- Vertex lacking median sulcus............................................................................ 7</p> <p>7 Scape with small, indistinct trichome on apicolateral margin; anterior margin of central cavity of tergite 1 (IV) densely setose (Fig. 35C); ventral stalk of aedeagus in dorsoventral view narrowing apically, with round, slightly broadened apex (Fig. 35F)...................................................................................... B. ganma sp. nov.</p> <p>- Scape with distinct trichome on apicolateral margin (Fig. 37C); anterior margin of central cavity of tergite 1 (IV) smooth (Fig. 37D); ventral stalk of aedeagus in dorsoventral view obliquely dilated (Fig. 37F)................ B. zhujianqingi sp. nov.</p></div> 	http://treatment.plazi.org/id/C822F213FF8D2430CB88A3FE70FB277D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF8C243ECB88A039748F24F6.text	C822F213FF8C243ECB88A039748F24F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella acclivis Yin 2022	<div><p>Batrisiella acclivis sp. nov.</p> <p>(Figs 30, 80B, 95A, B)</p> <p>Chinese common name: ṖḎ小毛唇à甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC). PARATYPES: CHINA: 6 ♂♂, 2 ♀♀, same data as that of holotype (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.8 mm. Head sub-rectangular at base; vertexal foveae moderately large and asetose, surrounding areas strongly impressed, lacking sulcus connecting foveae; antenna moderately elongate; antennomeres each slightly elongate, 9–11 enlarged to form moderately distinct club. Discal stria of elytron extending to approximately apical 1/5 of elytral length. Legs with profemur strongly swollen distally, with dense trichomes at apex, mesotibia with tiny tubercle at apex. Aedeagus strongly asymmetric; median lobe with large basal capsule, basoventral projection short, ventral stalk broad in lateral view; dorsal lobe strongly curved, apically split into two elongate sclerites; parameres reduced to single, small membranous structure. Female. Body length approximately 1.80 mm; antenna shorter, legs lacking modifications, genitalia as in Fig. 30I.</p> <p>Description. Male. Body (Fig. 30A) length 1.80–1.85 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 30B) sub-rectangular at base, approximately as long as broad, length 0.38–0.39 mm, width across eyes 0.40–0.42 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae, surrounding areas of foveae strongly impressed, median carina thin and long, with broad transverse sulcus between weakly raised antennal tubercles; frons broadly impressed medially, smooth, anteriorly confluent with clypeus; clypeus coarsely punctate, anterior margin carinate and moderately raised; ocular-mandibular ridge (with upper and lower carinae) broad. Venter with two tiny gular foveae (posterior tentorial pits) originating from shared impression, with thin median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna moderately elongate, length 0.91–0.92 mm, apical three antennomeres forming moderately distinct club; antennomere 1 thick, subcylindrical, laterally with internal glandular structure (Fig. 30C), 2–8 each elongate, 8 smallest, 9 much larger than 8, 10 as long as and broader than 9, 11 largest, as long as 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 30B) approximately as long as broad, length 0.43–0.44 mm, width 0.42–0.43 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus broad, slightly shorter than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with round lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.58–0.60 mm, width 0.71–0.76 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to approximately apical 1/5 of elytral length; humerus angulate, small subhumeral fovea present, with carinate marginal stria extending posteriorly from fovea to elytral posterior margin and anteriorly to dorsal surface of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, not forked internally; intercoxal process short. Metaventrite projecting admesally, inclined toward middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin emarginate at middle, lacking split.</p> <p>Legs elongate; profemur (Fig. 30D) strongly swollen at distal 1/2, dorsal surface near apex deeply incised, with dense trichomes inside incision; mesotibia (Fig. 30E) with tiny, truncate protuberance at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.52–0.54 mm, width 0.64–0.66 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by two mediobasal and one pair of basolateral foveae, with pair of long discal carinae, marginal carina present only at basal 1/2; tergites 2–4 (V–VII) tightly compressed, 2–3 lacking and 4 with one pair of basolateral foveae, tergite 4 shorter than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, roundly emarginate at middle of posterior margin. Sternite 2 (IV) with large mediobasal and pair of broad, setose basolateral foveae, with long lateral carinae, midlength slightly longer than 3–4 (V–VI) combined, sternites 3–5 (V–VII) lacking foveae, sternite 6 (VIII) with posterior margin slightly emarginate, sternite 7 (IX) (Fig. 30F) slightly oval, weakly sclerotized at apical part and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 30G, H) 0.21 mm long, strongly asymmetric; median lobe with large basal capsule and subtriangular foramen, basoventral projection short, ventral stalk broad in lateral view, apex shortly forked; dorsal lobe elongate, recumbent, strongly curved and C-shaped in anterior view, apically split into two elongate lobes, upper one gradually narrowing apically, lower one forked at apex; parameres reduced to single, small membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 21 ommatidia; humerus also angulate; legs lacking modifications. Measurements (as for male): body length 1.78–1.82 mm; length/width of head 0.36–0.37/ 0.39–0.40 mm, pronotum 0.41–0.42/ 0.42 mm, elytra 0.52/ 0.70–0.71 mm; abdomen 0.58–0.61/ 0.66–0.67 mm; length of antenna 0.88 mm; maximum width of genitalia (Fig. 30I) 0.22 mm.</p> <p>Comparative notes. The new species is most similar to B. beibeng by the strongly swollen profemora of the male, but can be readily distinguished by the presence of a thin median carina of the vertex (lacking in B. beibeng), the more strongly setose swollen part of the profemora, as well as by the recumbent and broader ventral stalk and the deeply split dorsal lobe of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 80B, 95A, B).</p> <p>Etymology. The specific epithet ‘ acclîvis (rising, sloping)’ is a Latin adjective referring to the swollen male profemora of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FF8C243ECB88A039748F24F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF82243CCB88A3A3705A2352.text	C822F213FF82243CCB88A3A3705A2352.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella beibeng Yin 2022	<div><p>Batrisiella beibeng sp. nov.</p> <p>(Figs 31, 80B, 99A)</p> <p>Chinese common name: Ü崩小毛唇à甲</p> <p>Type material (23 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Mêdog County, nr. Beibeng, 29°14’11”N, 95°09.54”E, 825 m, 11.vii.2018; mixed leaf litter, sifted, Cheng, Peng Shen leg., [Ʊ ẪẸṘÜ 崩乡]’ (SNUC). PARATYPES: CHINA: 7 ♂♂, 15 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length 1.82–1.85 mm. Head sub-rectangular at base; vertexal foveae moderately large and asetose, lacking sulcus connecting them; antenna moderately elongate; antennomeres each elongate, 9–11 enlarged to form indistinct club. Discal stria of elytron extending to approximately apical 1/8 of elytral length. Legs with profemur markedly thickened at distal 1/2, with disc-like protuberance at apical 1/4 on dorsal surface, metatibia broadened at distal 1/2. Aedeagus strongly asymmetric; median lobe with large basal capsule, lacking basoventral projection, ventral stalk curved rightwards in ventral view (position in figure); dorsal lobe strongly curved for whole length, forked apically; parameres reduced to single, weakly membranous structure. Female. Body length 1.80–1.86 mm; antenna shorter, legs lacking modifications, genitalia as in Fig. 31K.</p> <p>Description. Male. Body (Fig. 31A) length 1.82–1.85 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 31B) sub-rectangular at base, slightly broader than long, length 0.38–0.39 mm, width across eyes 0.41–0.42 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae or mediobasal carina, with transverse, medially emarginate sulcus between antennal tubercles; antennal tubercles moderately raised; frons broadly impressed medially, demarcated from clypeus by nearly complete frontal-clypeal ridge; clypeus with rough surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with two small gular foveae (posterior tentorial pits) close originating from shared opening, with thin median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 30 ommatidia. Antenna moderately elongate, length 0.96–0.98 mm, apical three antennomeres forming indistinct club; antennomere 1 thick, subcylindrical, apicolateral portion with large internal glandular structure, indistinct, 2–8 each elongate, 8 smallest, 9 much larger than 8, 10 as long as and slightly broader than 9, 11 largest, slightly shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 31B) approximately as long as broad, length 0.43–0.45 mm, width 0.42 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus, with small antebasal spines; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.53–0.55 mm, width 0.67–0.69 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to approximately apical 1/8 of elytral length; humerus angulate, small subhumeral fovea present, sulcate marginal stria extending posteriorly from fovea to elytral posterior margin and anteriorly to dorsal surface of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite longitudinally sulcate along midline, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin emarginate at middle, lacking split.</p> <p>Legs elongate; profemur (Fig. 31C) strongly thickened at distal 1/2, dorsal surface (Fig. 31D) near apex with disc-like protuberance, metatibia (Fig. 31E) expanded at distal 1/2, surface with dense short setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.48–0.53 mm, width 0.63–0.65 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by two mediobasal and one pair of basolateral foveae, with pair of short discal carinae, marginal carina present only at basal 1/2; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 (VII) shorter than 2 and 3 combined along middle, tergite 5 (VIII) broad and slightly transverse. Sternite 2 (IV) with large mediobasal and pair of broad, setose basolateral sockets, with long lateral carinae, midlength slightly longer than 3–4 (V–VI) combined, sternites 3–5 (V–VII) each with one pair of indistinct basolateral foveae, sternite 6 (VIII) with posterior margin slightly emarginate, sternite 7 (IX) (Fig. 31F) asymmetric, transversely slightly oval, weakly sclerotized at apical part.</p> <p>Aedeagus (Fig. 31G–J) 0.25 mm long, strongly asymmetric; median lobe with large basal capsule and sub-triangular foramen, lacking basoventral projection, ventral stalk narrowing towards apex, curved rightwards in ventral view (position of figure); dorsal lobe elongate, strongly curved, apex split into two unequal thin lobes, dorsal part with additional spine-like lobe; parameres reduced to single weakly membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 20 ommatidia; humerus also prominent; legs lacking modifications. Measurements (as for male): body length 1.80–1.86 mm; length/width of head 0.36–0.38/ 0.40 mm, pronotum 0.43–0.44/ 0.42 mm, elytra 0.48–0.50/ 0.64–0.69 mm; abdomen 0.52–0.56/ 0.62–0.65 mm; length of antenna 0.85–0.90 mm; maximum width of genitalia (Fig. 31K) 0.19 mm.</p> <p>Comparative notes. The new species shares with B. acclivis the broad male profemora, but differs by the femora lacking dense setae and with a disc-like protuberance, the thickened male metatibiae, as well as a generally more elongate aedeagus with a slenderer ventral stalk of the median lobe. The modified profemora are also shared by B. strictus Jiang &amp; Yin which is distributed in Guangxi, southern China. Apart from this, the two species can be readily separated by the lack of a mediobasal carina of the vertex, the much longer discal carinae of the elytra, and the distally broadened metatibiae (normally slender in B. strictus) of the new species.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 80B, 99A).</p> <p>Etymology. The specific epithet refers to the type locality of the new species, i. e., Beibeng Township.</p></div> 	http://treatment.plazi.org/id/C822F213FF82243CCB88A3A3705A2352	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF80243BCB88A41271BC2182.text	C822F213FF80243BCB88A41271BC2182.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella bomea Yin 2022	<div><p>Batrisiella bomea sp. nov.</p> <p>(Figs 32, 80B, 97A, B, D)</p> <p>Chinese common name: 波ṁ小毛唇à甲</p> <p>Type material (64 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200–2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’ (SNUC). PARATYPES: CHINA: 5 ♂♂, 3 ♀♀, same data as that of holotype; 23 ♂♂, 32 ♀♀, China: Xizang, Nyingchi City, Tongmai Town, pass nr. 102 <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.13528&amp;materialsCitation.latitude=30.070835" title="Search Plazi for locations around (long 95.13528/lat 30.070835)">Tunnel</a>, 30°04’15”N, 95°08’07”E, 2150-2400 m, 12.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫȃ 9 102 Ṁdz (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.7–1.8 mm. Head subtruncate at base; vertexal foveae moderately large and asetose, with transverse sulcus between antennal tubercles; antenna moderately elongate; antennomeres each slightly elongate, 9–11 enlarged to form moderately distinct club, with modified antennomeres 8–10. Discal stria of elytron extending to approximately apical 1/5 of elytral length. Legs simple. Aedeagus strongly asymmetric; median lobe with large basal capsule, basoventral projection short, ventral stalk narrowing towards apex; dorsal lobe strongly curved, with broad, rounded apex; parameres reduced to single membranous structure. Female. Body length approximately 1.7–1.8 mm; antenna shorter, antennae lacking modifications, genitalia as in Fig. 32G.</p> <p>Description. Male. Body (Fig. 32A) length 1.69–1.78 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 32B) subtruncate at base, slightly broader than long, length 0.36–0.38 mm, width across eyes 0.40– 0.43 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking median carina, with transverse, sinuate sulcus between antennal tubercles; frons coarsely punctate, weakly impressed medially, anteriorly confluent with clypeus; clypeus smooth, anterior margin carinate and moderately raised; ocularmandibular ridge (with upper and lower carinae) broad. Venter with tiny gular foveae (posterior tentorial pits) originating from shared small pit, with thin median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 23 ommatidia. Antenna moderately short, length 0.85–0.90 mm, apical three antennomeres forming moderately distinct club, with modified antennomeres 8–10 (Fig. 32C); antennomere 1 thick, subcylindrical, laterally with internal glandular structure, 2–7 each slightly elongate, 8 broader than 7, 9 much larger than 8, 8 and 9 each with short, thickened setae, 10 larger than 9, constricted at base and dilated at apex, 11 longest, sub-fusiform.</p> <p>Pronotum (Fig. 32B) approximately as long as broad, length 0.40–0.41 mm, width 0.42–0.43 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus broad, as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with round lateral antebasal hypomeral impression; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.53–0.57 mm, width 0.68–0.74 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to slightly more than apical 1/5 of elytral length; humerus angulate, small subhumeral fovea present, with carinate marginal stria extending posteriorly from fovea to elytral posterior margin and anteriorly to dorsal surface of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite moderately projecting admesally, inclined toward middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin broadly and weakly emarginate at middle, lacking split.</p> <p>Legs simple, lacking spines or tubercles.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.46–0.51 mm, width 0.62–0.67 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by two mediobasal and one pair of basolateral foveae, with pair of long discal carinae, marginal carina present only at basal third; tergites 2–4 (V–VII) tightly compressed, 2–3 lacking and 4 with one pair of basolateral foveae, tergite 4 shorter than 2 and 3 combined along middle, tergite 5 (VIII) semicircular, posterior margin roundly and shallowly emarginate at middle. Sternite 2 (IV) with large mediobasal and pair of broad, setose basolateral sockets, with long lateral carinae, midlength slightly longer than 3–4 (V–VI) combined, sternites 3–5 (V–VII) lacking foveae, sternite 6 (VIII) with posterior margin slightly emarginate, sternite 7 (IX) (Fig. 32D) slightly oval, weakly sclerotized at apical part and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 32E–F) 0.23 mm long, strongly asymmetric; median lobe with large basal capsule and subtriangular foramen, basoventral projection short, ventral stalk elongate and protruding, curved, narrowing toward apex; dorsal lobe elongate, recumbent, strongly curved and C-shaped in anterior view, in lateral view apically broadened and with rounded apex; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 18 ommatidia; humerus weakly angulate; legs simple. Measurements (as for male): body length 1.71–1.81 mm; length/width of head 0.36–0.38/ 0.41–0.42 mm, pronotum 0.40/ 0.41–0.42 mm, elytra 0.52–0.53/ 0.69–0.70 mm; abdomen 0.56–0.57/ 0.66 mm; length of antenna 0.84–0.86 mm; maximum width of genitalia (Fig. 32G) 0.25 mm.</p> <p>Comparative notes. The new species can be readily separated from all congeners in Tibet and northern India by the location of the male sexual characters on the antennal clubs and their unique forms, as well as by the characteristic form of the aedeagus.</p> <p>Distribution. Bomê County, Tibet, SW China (Figs 80B, 97A, B, D).</p> <p>Etymology. The new species is named after Bomê County, where the type locality is located.</p></div> 	http://treatment.plazi.org/id/C822F213FF80243BCB88A41271BC2182	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF862438CB88A791762E2446.text	C822F213FF862438CB88A791762E2446.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella concisa Yin 2022	<div><p>Batrisiella concisa sp. nov.</p> <p>(Figs 33, 80B, 99B)</p> <p>Chinese common name: Ṋú小毛唇à甲</p> <p>Type material (3 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.18413&amp;materialsCitation.latitude=29.246986" title="Search Plazi for locations around (long 95.18413/lat 29.246986)">Beibeng</a>, road to Gelin Vill., 29°14’49.15”N, 95°11’2.86”E, 1060 m, 2019.vii.23, leaf litter, Z.-W. Yin leg., [ƱẪÜ崩乡格ff村]’ (SNUC). PARATYPES: CHINA: 1 ♂, 1 ♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length 1.76–1.78 mm. Head sub-rectangular at base; vertexal foveae small and asetose, lacking sulcus connecting them; antenna long, antennomeres each elongate, lacking modifications. Discal stria of elytron extending to apical 1/10 of elytral length. Legs almost simple, only mesotibia with tiny apical spur. Abdomen with large tergite 1 (IV) and vertical tergites 2–5 (V–VIII); tergite 1 with large posteromedial cavity and setose lateral patches. Aedeagus strongly asymmetric; ventral stalk narrowing apically and with truncate apex; dorsal lobe composed of one broad and one long, thin and curved structures; parameres reduced and forming single membranous structure. Female. Body length 1.65 mm; mesotibia lacking apical spur, tergite 1 lacking modifications, genitalia as in Fig. 4H.</p> <p>Description. Male. Body (Fig. 33A) length 1.76–1.78 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 33B) sub-rectangular at base, approximately as long as wide, length 0.37–0.38 mm, width across eyes 0.39 mm; vertex finely punctate, sparsely pubescent, with small, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae or mediobasal carina, with T-shaped sulcus at middle and few large punctures anterior and lateral to anterior margin of sulcus; antennal tubercles slightly raised; frons slightly impressed between antennal tubercles, anterolateral margins carinate and with small tufts of setae; clypeus with smooth surface, its anterior margin weakly carinate; ocular-mandibular carina present. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 25 large ommatidia. Antenna elongate, length 0.93 mm, apical three antennomeres forming moderately distinct club; antennomere 1 thick, subcylindrical, apicolateral portion with small inner glandular structure (Fig. 33C), lacking trichome, 2–7 each elongate, 8 moniliform, 9 much larger than 8, 10 slightly broader than and as long as 9, 11 largest, as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 33B) approximately as long as broad, length 0.40–0.43 mm, width 0.41 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate and sparsely pubescent; median longitudinal sulcus as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by thin antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with large lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.58 mm, width 0.73–0.74 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria carinate, extending from outer basal fovea to apical 1/10 of elytral length; humerus moderately prominent, with small subhumeral fovea, with marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, not forked internally; intercoxal process short. Metaventrite broadly impressed at middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate, lacking distinct modifications; mesotibia with tiny apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.46–0.47 mm, width 0.65 mm. Tergite 1 (IV) (Fig. 33D) much longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with short discal and outer marginal carinae; with large posteromedial cavity, anterior margin of cavity carinate, inside cavity there is one small anteromedial process and row of dense median setae, area lateral to cavity with large setose patches covered with dense, short setae; tergites 2–5 (V–VIII) tightly compressed, vertical, 2–4 each with one pair of basolateral foveae, tergite 4 (VII) longer than 2 and 3 combined along middle, tergite 5 (VIII) broad and slightly transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and two pairs of basolateral foveae in broad setose basal impression; midlength of sternite 2 (IV) approximately as long as 3–4 (V–VI) combined, sternites 3–5 (V–VII) each with one pair of small basolateral foveae, sternite 6 (IX) (Fig. 33E) rounded, apical 1/3 moderately sclerotized and with a few setae along anterior margin, basal 2/3 membranous.</p> <p>Aedeagus (Fig. 33F–H) 0.33 mm long, strongly asymmetric; median lobe with large basal capsule and sub-triangular foramen, ventral stalk narrowing apically and with truncate apex in dorso-ventral view; dorsal lobe transversely protruding and then strongly curved towards apex, with one thin, long and curved sclerite; parameres reduced and forming single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; frons and tergite 1 (IV) lacking modifications; each compound eye composed of approximately 20 ommatidia; humeral angle rounded; mesotibia lacking apical spur. Measurements (as for male): body length 1.65 mm; length/width of head 0.36/ 0.37 mm, pronotum 0.40/ 0.38 mm, elytra 0.46/ 0.64 mm; abdomen 0.47/ 0.58 mm; length of antenna 0.84 mm; maximum width of genitalia (Fig. 33I) 0.21 mm.</p> <p>Comparative notes. The new species is close to B. ganma by sharing the similar placement and form of the male tergal modification and the aedeagus. Batrisiella concisa can be readily separated from B. ganma by the presence of a broad, deep median longitudinal sulcus on the vertex of the head (sulcus absent in B. ganma).</p> <p>Distribution (Fig. 98). Mêdog County, Tibet, SW China (Figs 80B, 99B).</p> <p>Etymology. The specific epithet ‘ concîsus, - um, - a ’ is a Latin adjective meaning ‘cut, broken up’, referring to the vertical plane of tergites 2–5 (V–VIII) of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FF862438CB88A791762E2446	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF842425CB88A650767827AA.text	C822F213FF842425CB88A650767827AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella curvitibialis Yin 2022	<div><p>Batrisiella curvitibialis sp. nov.</p> <p>(Figs 34, 80C, 97C, 99B, C)</p> <p>Chinese common name: DZḇ小毛唇à甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.18413&amp;materialsCitation.latitude=29.246986" title="Search Plazi for locations around (long 95.18413/lat 29.246986)">Beibeng</a>, road to Gelin Vill., 29°14’49.15”N, 95°11’2.86”E, 1060 m, 2019.vii.23; leaf litter, Z.-W. Yin leg., [ƱẪÜ崩乡格ff村]’ (SNUC).</p> <p>PARATYPES: CHINA: 1 ♂, 2 ♀♀, same data as that of holotype; 1 ♂, China: Xizang, Mêdog County, nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.24573&amp;materialsCitation.latitude=29.2623" title="Search Plazi for locations around (long 95.24573/lat 29.2623)">Yarang Power Station</a>, 29°15’44.28”N, 95°14’44.63”E, 885 m, 2019.vii.24, leaf litter, Z.-W. Yin leg., [ƱẪẸṘ亚üΨ ‖]; 3 ♂♂, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.48917&amp;materialsCitation.latitude=29.656666" title="Search Plazi for locations around (long 95.48917/lat 29.656666)">China</a>: Tibet, Mêdog County, 80K (ca. 29°39’24”N, 95°29’21”E), 2111 m, 11.viii.2016, Chen Zhi-Lin leg.; 1 ♂, China: Nyingchi County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, 2190 m, 2019.vii.15–viii.04, FIT, Z.-W. Yin leg., [ƱẪffżḦËƖǎ]’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length 2.02–2.06 mm. Head sub-rectangular at base; vertexal foveae moderately large and asetose, lacking sulcus connecting them; antenna moderately elongate; antennomeres each elongate, 9–11 enlarged to form indistinct club. Discal stria of elytron extending to apical 5/6 of elytral length. Legs with mesotibia with small apical tubercle, metafemur protuberant on ventral margin and at apex, metatibia sinuate at base. Admesal area of posterior part of metaventrite densely setose. Surface of tergite 4 (VII) broadly and transversely impressed, with pair of small, oval cavities at middle. Aedeagus strongly asymmetric; median lobe with large basal capsule and short basoventral projection, ventral stalk curved ventrally; dorsal lobe transverse at base and strongly curved, apex split into two lobes; parameres reduced to round, membranous structure. Female. Body length 1.92 mm; antenna shorter, metaventrite, legs and abdomen lacking modifications, genitalia as in Fig. 6K.</p> <p>Description. Male. Body (Fig. 34A) length 2.02–2.06 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 34B) sub-rectangular at base, slightly broader than long, length 0.41–0.42 mm, width across eyes 0.44–0.45 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae, with thin mediobasal carina extending from subbase to level of anterior margins of vertexal foveae, with V-shaped sulcus between antennal tubercles; antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus at middle and carinate anterolaterally; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina complete. Venter with single, small gular fovea (posterior tentorial pit), with thin median carina extending from fovea anteriorly to mouthparts. Compound eyes prominent, composed of approximately 30 ommatidia. Antenna moderately elongate, length 1.08 mm, apical three antennomeres forming indistinct club; antennomere 1 thick, subcylindrical, apicolateral portion with small inner glandular structure, lacking trichome, 2–8 each elongate, 8 smallest, 9 much larger than 8, 10 as long as and slightly broader than 9, 11 largest, as long as 9 and 10 combined, slightly oval.</p> <p>Pronotum (Fig. 34B) approximately as long as broad, length 0.43–0.44 mm, width 0.45–0.46 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with distinct lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.61–0.62 mm, width 0.76–0.78 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to apical 5/6 of elytral length; humerus angulate, subhumeral fovea absent, sulcate marginal stria extending from where fovea supposed to be to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, not forked internally; intercoxal process short. Metaventrite broadly impressed at middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae; admesal area (Fig. 34C) densely setose at posterior half; posterior margin with small and narrow split at middle.</p> <p>Legs elongate; mesotibia (Fig. 34D) with small apical tubercle; metafemur (Fig. 34E) projecting on ventral margin at apical 1/3, apex strongly extended on ventral side such that femoral-trochanteral joint is covered, metatibia (Fig. 34F) thin and sinuate at basal 1/3.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.69–0.71 mm, width 0.58–0.61 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by two mediobasal and one pair of basolateral foveae, with pair of short discal carinae; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 (VII) (Fig. 34G) slightly shorter than 2 and 3 combined along middle, with pair of small, oval cavities at middle, surface with broad and transverse median impression, tergite 5 (VIII) broad and slightly transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and pair of broad, setose basolateral sockets, with short lateral and long, complete marginal carinae, midlength approximately as long as 3–4 (V–VI) combined, sternites 3–5 (V–VII) each with one pair of indistinct basolateral foveae, sternite 6 (VIII) with posterior margin slightly emarginate, sternite 7 (IX) (Fig. 34H) asymmetric, transversely slightly oval, weakly sclerotized.</p> <p>Aedeagus (Fig. 34I, J) 0.33 mm long, strongly asymmetric; median lobe with large basal capsule and sub-triangular foramen, ventral stalk narrowing towards apex; dorsal lobe transverse, strongly curved, broad throughout whole length, apex split into two unequal lobes; parameres reduced to single round membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 22 ommatidia; humerus weakly prominent; metaventrite lacking densely setose area, legs lacking tubercles or projections; abdomen lacking modifications. Measurements (as for male): body length 1.96 mm; length/width of head 0.40–0.41/ 0.41 mm, pronotum 0.42–0.43/ 0.42–0.45 mm, elytra 0.52–0.54/ 0.69–0.74 mm; abdomen 0.61– 0.66/ 0.66–0.68 mm; length of antenna 0.95–0.98 mm; maximum width of genitalia (Fig. 34K) 0.25 mm.</p> <p>Comparative notes. The new species is morphologically distinct and can be readily separated from all known congeners by the conspicuously modified hindlegs, as well as the form of the modification of male tergite VII. Females from the same locality may be identified by the shape of the genitalia without association with the male.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 80C, 97C, 99B, C).</p> <p>Etymology. The specific epithet is combined from ‘ curvus, - a, - um ’ (bent, curved) and ‘ tibalis, - le ’ (tibial), referring to the sinuate male metatibia of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FF842425CB88A650767827AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF992423CB88A30A70E325FA.text	C822F213FF992423CB88A30A70E325FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella ganma Yin 2022	<div><p>Batrisiella ganma sp. nov.</p> <p>(Figs 35, 80C, 93E–G)</p> <p>Chinese common name: HḼ小毛唇à甲</p> <p>Type material (11 exx.). HOLOTYPE: CHINA: ♂, China: Xizang, Dinggyê County, Zhêntang Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (SNUC). PARATYPES: CHINA: 6 ♂♂, same data as that of holotype; 1 ♂, 3 ♀♀, also from <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Zhêntang</a>, except ‘Xiuxiongma Vill., 27°54’11”N, 87°22’42”E, 2700-3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU 喀NJŤĠ县ĿƋWšŏ玛村 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length slightly over 1.65 mm. Head sub-rectangular at base; vertexal foveae moderately small and asetose, with transverse sulcus between antennal tubercles, lacking mediobasal carina; antenna moderately long, antennomeres slightly elongate, lacking modifications. Discal stria of elytron extending to apical 1/5 of elytral length. Legs almost simple, only mesotibia with tiny apical spur. Abdomen with large tergite 1 (IV) and vertical tergites 2–5 (V–VIII); tergite 1 with large posteromedial cavity and setose lateral patches. Aedeagus strongly asymmetric; ventral stalk narrowing apically and with rounded apex; dorsal lobe strongly twisted, in dorsal view expanded at apex; parameres reduced and forming single membranous structure. Female. Body length over 1.7 mm; mesotibia lacking apical spur, tergite 1 lacking modifications, genitalia as in Fig. 35G.</p> <p>Description. Male. Body (Fig. 35A) length 1.66–1.67 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 35B) sub-rectangular at base, approximately as long as wide, length 0.36–0.38 mm, width across eyes 0.39–0.41 mm; vertex finely punctate, sparsely pubescent, with moderately small, asetose vertexal foveae (dorsal tentorial pits), with short, transverse sulcus between antennal tubercles, lacking median longitudinal sulcus; antennal tubercles moderately raised; frons with roughened surface slightly impressed between antennal tubercles, anterolateral margins carinate, medially confluent with clypeus; clypeus with roughened surface, its anterior margin moderately carinate and raised; ocular-mandibular carina present. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 21 large ommatidia. Antenna elongate, length 0.88–0.92 mm, apical three antennomeres enlarged and forming moderately distinct club; antennomere 1 thick, subcylindrical, apicolateral portion with elongate inner glandular structure and indistinct trichome, 2–7 each elongate, 8 sub-moniliform, 9 much larger than 8, 10 slightly broader than and as long as 9, 11 largest, as long as 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 35B) approximately as long as broad, length 0.42–0.43 mm, width 0.41–0.43 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate and sparsely pubescent; median longitudinal sulcus as long as semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by deep antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with large lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.49–0.50 mm, width 0.66–0.70 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria carinate, extending from outer basal fovea to apical 1/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, with marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, not forked internally; intercoxal process short. Metaventrite weakly impressed at middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin broadly emarginate at middle, lacking split.</p> <p>Legs moderately elongate, lacking distinct modifications; mesotibia with tiny apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.49–0.50 mm, width 0.65–0.70 mm. Tergite 1 (IV) (Fig. 35C) much longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, discal carina long, marginal carina absent; posteromedial cavity large, with fringed anterior margin projecting posteriorly at middle, inside cavity there is one mediobasal process and longitudinal row of dense setae, area lateral to cavity with large setose patch covered with dense, short setae; tergites 2–5 (V–VIII) tightly compressed, vertical, 2–4 each with one pair of basolateral foveae, 4 longer than 2 and 3 combined along middle, 5 broad and slightly transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal foveae and large basolateral sockets in broad setose basal impression; midlength of sternite 2 (IV) approximately as long as 3–4 (V–VI) combined, sternites 3–5 (V–VII) each with one pair of small basolateral foveae, sternite 6 (IX) (Fig. 35D) rounded at apex, apical 1/3 weakly sclerotized and with a few setae along anterior margin, basal 2/3 membranous.</p> <p>Aedeagus (Fig. 35E, F) 0.31 mm long, strongly asymmetric; median lobe with large basal capsule and rounded triangular foramen, ventral stalk narrowing apicad and with rounded apex in dorso-ventral view; dorsal lobe transversely protruding and then strongly curved and expanded apically; parameres reduced and forming single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; tergite 1 (IV) lacking modifications; each compound eye composed of approximately 15 ommatidia; humeral angle rounded; mesotibia lacking apical spur. Measurements (as for male): body length 1.70–1.71 mm; length/width of head 0.37–0.38/ 0.39–0.40 mm, pronotum 0.42–0.43/ 0.41–0.42 mm, elytra 0.47–0.48/ 0.68–0.70 mm; abdomen 0.57–0.58/ 0.67–0.70 mm; length of antenna 0.84–0.87 mm; maximum width of genitalia (Fig. 35G) 0.19 mm.</p> <p>Comparative notes. The new species is close to B. concisa by sharing a similar placement and the form of the male tergal modification and the aedeagus. Batrisiella ganma can be readily separated from B. concisa by the lack of a median longitudinal sulcus on the vertex of the head (sulcus broad and deep in B. concisa).</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 80C, 93E–G).</p> <p>Etymology. The new species is name after Ganma (= Gama) Zangbo, a river lying south of Zhêntang Town.</p></div> 	http://treatment.plazi.org/id/C822F213FF992423CB88A30A70E325FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF9F2421CB88A2BA76AC239E.text	C822F213FF9F2421CB88A2BA76AC239E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella guttata Yin 2022	<div><p>Batrisiella guttata sp. nov.</p> <p>(Figs 36, 80C, 100C)</p> <p>Chinese common name: ųê小毛唇à甲</p> <p>Type material (8 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Zayü County, valley nr. Zhala Vill., 28°37’23.74”N, 97°21’8.10”E, 2190 m, 2019.vii.28, leaf litter / decomposing log, Z.-W. Yin leg., [ƱẪ察ø县扎 Ń村ṉữ]’ (SNUC). PARATYPES: CHINA: 5 ♂♂, 2 ♀♀, same data as that of holotype’ (SNUC).</p> <p>Diagnosis. Male. Body length 1.85–1.91 mm. Head sub-rectangular at base; vertexal foveae moderately large and asetose, lacking sulcus connecting them; antenna moderately elongate; antennomeres slightly elongate or moniliform, antennomeres 9–11 enlarged to form club. Discal stria of elytron extending to apical 3/5 of elytral length. Legs with pro- and mesotrochanter with small ventral tubercle, mesotibia with distinct apical spine. Tergite 4 (VII) with two large and round, disc-like lateral protuberances, tergite 5 (VIII) with broad and deep central cavity, posteromedian portion angularly projecting. Sternite 2 (IV) with pair of admesal setose projections, sternite 3 (V) with two similar, but much smaller projections, sternite 4 (IV) lacking projections but with long, thick setae at middle. Aedeagus strongly asymmetric; median lobe with large basal capsule and basoventral projection, ventral stalk dilated; dorsal lobe curved and transverse, apex pointed; parameres reduced to moderately sclerotized structure. Female. Body length 2.0 mm; antenna shorter, legs lacking tubercle or spine, abdomen lacking modifications, genitalia as in Fig. 7K.</p> <p>Description. Male. Body (Fig. 36A) length 1.85–1.91 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 36B) sub-rectangular at base, approximately as long as wide, length 0.41–0.42 mm, width across eyes 0.42–0.44 mm; vertex finely punctate, with moderately large, asetose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae or mediobasal carina, with transverse sulcus between antennal tubercles; antennal tubercles moderately raised; frons slightly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and moderately raised; ocular-mandibular carina present. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna moderately elongate, length 0.90–0.91 mm, apical three antennomeres forming distinct club; antennomere 1 thick, subcylindrical, apicolateral portion with small inner glandular structure, lacking trichome, indistinct, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much larger than 8, 10 slightly longer and broader than 9, 11 largest, slightly shorter than 9 and 10 combined, slightly oval.</p> <p>Pronotum (Fig. 36B) approximately as long as broad, length 0.45 mm, width 0.44–0.45 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, pubescent, median longitudinal sulcus slightly shorter than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by transverse antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with distinct lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.63 mm, width 0.74 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to apical 3/5 of elytral length; humerus weakly prominent, with small subhumeral fovea, sulcate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of thin admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite broadly impressed at middle, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs moderately elongate; pro- and mesotrochanter each with ventral tubercle, mesotibia with distinct apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.50–0.55 mm, width 0.67 mm. Tergite 1 (IV) longer than tergites 2–4 (V–VII) combined in dorsal view, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, with pair of triangular discal carinae; tergites 2–4 (V–VII) tightly compressed, each with one pair of basolateral foveae, tergite 4 (VII) (Fig. 36C) slightly shorter than 2 and 3 combined along middle, each side with one large and round projection, tergite 5 (VIII) (Fig. 36C, D) broad and slightly transverse, with large central cavity, posteromedian part angularly projecting and with dense setae at apex of projection, with pair of much smaller tubercles lateral to protuberance, each with few thickened setae, anterior margin of cavity with pair of small median protuberances, anterior margin of tergite with pair of round admesal tubercles. Sternite 2 (IV) with large mediobasal and pair of broad, setose basolateral sockets, midlength approximately as long as 3–5 (V–VII) combined, with pair of large, setose admesal projections (Fig. 36E, F), 3–5 each with one pair of indistinct basolateral foveae, sternite 3 (V) with pair of small, setose admesal projections (Fig. 36E, F), sternite 4 and 5 (VI and VII) lacking projections, but with thickened setae at middle, sternite 6 (VIII) as long as 5 (VII) at middle, posterior margin slightly emarginate, sternite 7 (IX) (Fig. 36G) transverse, rounded triangular, weakly sclerotized.</p> <p>Aedeagus (Fig. 36H–J) 0.33 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, ventral stalk strongly dilated; dorsal lobe transverse, strongly curved, narrowing from middle apically, with pointed apex; parameres reduced to single moderately sclerotized structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; each compound eye composed of approximately 22 ommatidia; humeral angle rounded; legs lacking tubercles or spines; abdomen lacking modifications. Measurements (as for male): body length 2.0 mm; length/width of head 0.40/ 0.41–0.43 mm, pronotum 0.42– 0.45/ 0.43–0.44 mm, elytra 0.53–0.59/ 0.70–0.74 mm; abdomen 0.68/ 0.66–0.69 mm; length of antenna 0.82–0.87 mm; maximum width of genitalia (Fig. 36K) 0.24 mm.</p> <p>Comparative notes. Batrisiella guttata can be readily separated from all congeners by the unique structure of the modifications on tergites 4 and 5 (VII and VIII), sternites 2 and 3 (IV and V), as well as the form of the aedeagus.</p> <p>Distribution. Zayü County, Tibet, SW China (Figs 80C, 100C).</p> <p>Etymology. The specific epithet ‘ guttâtus, - a, - um ’ is a Latin adjective meaning ‘spotted, speckled’, referring to the round lateral projections of male tergite 4 (VII) of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FF9F2421CB88A2BA76AC239E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF9D242FCB88A75E70FA27D6.text	C822F213FF9D242FCB88A75E70FA27D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisiella zhujianqingi Yin 2022	<div><p>Batrisiella zhujianqingi sp. nov.</p> <p>(Figs 37, 80C, 92C)</p> <p>Chinese common name: 朱氏小毛唇à甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Nyalam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.97806&amp;materialsCitation.latitude=27.955833" title="Search Plazi for locations around (long 85.97806/lat 27.955833)">Zhangmu Town</a>, 27°59’15”N, 85°58’59”E, 2200 m, 17.vii.2010, leaf litter, sifted, J.-Q. Zhu leg., [ƱẪỄŃ木县樟木W]’ (SNUC). PARATYPES: CHINA: 1 ♂, 6 ♀♀, same data as that of holotype; 1 ♀, also from Zhangmu, except ‘ Lixin Vill. (ĒẾ村), 27°57’21”N, 85°58’41”E, 2400–2600 m, 27.vii.2010 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length 1.81–1.87 mm. Head sub-rectangular at base; vertexal foveae small and asetose, with transverse sulcus disappearing just before foveae; antenna long, antennomeres each slightly elongate, lacking modifications. Discal stria of elytron extending to apical 1/7 of elytral length. Legs almost simple, only mesotibia with small apical spur. Abdomen with large tergite 1 (IV) and vertical tergites 2–5 (V–VIII); tergite 1 with large posteromedial cavity and setose lateral patches. Aedeagus strongly asymmetric; ventral stalk narrowing apically and with rounded apex; dorsal lobe strongly curved at base, broad apically; parameres reduced and forming single semi-membranous structure. Female. Body length 1.73–1.75 mm; mesotibia lacking apical spur, tergite 1 lacking modifications, genitalia as in Fig. 37H.</p> <p>Description. Male. Body (Fig. 37A) length 1.81–1.87 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with moderately long pubescence.</p> <p>Head (Fig. 37B) sub-rectangular at base, much broader than long, length 0.37–0.38 mm, width across eyes 0.44–0.45 mm; vertex smooth, sparsely pubescent, with large, asetose vertexal foveae (dorsal tentorial pits), with transverse sulcus posterior to antennal tubercles, disappearing before vertexal foveae, mediobasal carina absent; antennal tubercles weakly raised; frons slightly impressed between antennal tubercles, roughly punctate, confluent with clypeus; clypeus with roughly punctate surface, its anterior margin moderately carinate and raised; ocular-mandibular carina present. Venter with two small gular foveae (posterior tentorial pits) originating from shared transverse slit, with distinct median carina extending from foveae anteriorly to mouthparts. Compound eyes prominent, composed of approximately 20 ommatidia. Antenna elongate, length 0.90–0.95 mm, apical three antennomeres loosely forming club; antennomere 1 thick, subcylindrical, apicolateral portion (Fig. 37C) with inner glandular structure and small trichome composed of short, thickened setae, 2–8 each elongate, 8 smallest, 9 much longer and broader than 8, 10 slightly broad than and as long as 9, 11 largest, much shorter than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 37B) approximately as long as broad, length 0.44–0.47 mm, width 0.46–0.47 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate and sparsely pubescent; median longitudinal sulcus slightly longer than semicircular lateral sulci in dorsal view, the latter extending from dorsal surface laterally and posteriorly and then fusing with lateral ends of antebasal sulcus; lacking median antebasal fovea, lateral antebasal foveae connected by deep antebasal sulcus; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, with large lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with two pairs of small hypomeral foveae; margin of coxal cavity weakly carinate.</p> <p>Elytra much wider than long, length 0.50–0.52 mm, width 0.71–0.72 mm; each elytron with two large, asetose basal foveae, lacking subbasal fovea; discal stria extending from outer basal fovea to apical 1/7 of elytral length; humerus denticulate, with small subhumeral fovea, with marginal stria extending from fovea to posterior margin of elytron, with thin, curved carina extending from fovea to humerus.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where lateral mesocoxal foveae situated at mesal margins, with pair of strongly raised admesal carinae; setose median mesoventral foveae widely separated, lateral mesoventral foveae large and setose, broadly forked internally; intercoxal process short. Metaventrite weakly impressed medially, with large, setose lateral mesocoxal foveae and pair of smaller, setose lateral metaventral foveae, posterior margin broadly and shallowly emarginate at middle, lacking split.</p> <p>Legs moderately elongate, lacking distinct modifications; mesotibia with small apical spur.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.53–0.55 mm, width 0.70 mm. Tergite 1 (IV) (Fig. 37D) strongly longer than tergites 2–4 (V–VII) combined, setose basal sulcus separated by mediobasal and one pair of basolateral foveae, discal carinae long, outer marginal carinae present at basal 1/3; with large, transverse central cavity at posterior end of tergite, anterior margin of cavity with one middle and two admesal tubercles, posterior margin with broad projection and setose tuft at middle, areas lateral cavity covered with dense, short setae; tergites 2–5 (V–VIII) tightly compressed, vertical, 2–4 each with one pair of basolateral foveae, tergite 4 (VII) as long as 2 and 3 combined along middle, tergite 5 (VIII) broad and slightly transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with large mediobasal and two pairs of basolateral foveae in setose basal impression, with pair of moderately long lateral carinae; midlength of sternite 2 (IV) shorter than 3–4 (V–VI) combined, sternites 3–5 (V–VII) each with one pair of small basolateral foveae, sternite 6 (IX) (Fig. 37E) sub-rounded, apical 1/3 moderately sclerotized and with a few setae along anterior margin, basal 2/3 membranous.</p> <p>Aedeagus (Fig. 37F, G) 0.31 mm long, strongly asymmetric; median lobe with large basal capsule and small, rounded triangular foramen, ventral stalk dorsoventrally broad at base, strongly narrowed for short distance at middle, and then dilated apically, with rounded apex; dorsal lobe in dorso-ventral view strongly curved at base, broad at middle, strongly broad at apical portion; parameres reduced and forming single semi-membranous structure.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, apicolateral trichome of scape same as that of male; tergite 1 (IV) lacking modifications; each compound eye composed of approximately 18 ommatidia; humerus also denticulate; mesotibia lacking apical spur. Measurements (as for male): body length 1.73–1.75 mm; length/width of head 0.34–0.35/ 0.40–0.41 mm, pronotum 0.40–0.42/ 0.43 mm, elytra 0.46–0.47/ 0.65 mm; abdomen 0.53–0.54/ 0.64–0.66 mm; length of antenna 0.81–0.82 mm; maximum width of genitalia (Fig. 37H) 0.23 mm.</p> <p>Comparative notes. The new species shares with B. concisa and B. ganma a posteriorly positioned central cavity of male tergite 1 (IV), and vertical tergites 2–5 (V–VIII). It differs from both of these species by the posterior margin of the tergal cavity with broad projection and setose tuft at middle (projection absent in B. concisa and of a different form in B. ganma), as well as a broad, apically obliquely dilated dorsal lobe of the aedeagus.</p> <p>Distribution. Nyalam County, Tibet, SW China (Figs 80C, 92C).</p> <p>Etymology. The new species is named after Jian-Qing Zhu, a Lepidopterist and the collector of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FF9D242FCB88A75E70FA27D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF93242ECB88A65970FB2528.text	C822F213FF93242ECB88A65970FB2528.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisodes Reitter 1882	<div><p>Batrisodes Reitter, 1882</p> <p>Chinese common name: qà甲¤</p> <p>Batrisodes Reitter, 1882: 134; Park 1947 (North American species), 1948 (checklist of world species); Chandler 1997 (catalog of North American species); Besuchet 1981 (Palaearctic species); Newton &amp; Chandler 1989 (discussion of type species, catalog); Nomura 2007 (Japanese species). Type species: Batrisus delaporti Aubé, 1833: 509 (subsequent designation by Lucas, 1920: 135).</p> <p>= Alytus Hampe, 1863: 285. Type species: Trichonyx adnexus Hampe, 1863: 285 (monotypy). Nomen oblatum, see discussion in Newton &amp; Chandler 1989: 8.</p> <p>= Batrisodellus Jeannel, 1958: 37. Type species: Batrisodes nipponensis Raffray, 1909: 23 (original designation). Synonymized by Yin et al. 2015a: 53.</p> <p>In China, the genus is represented by 21 species (e. g., Nomura 2007; Yin &amp; Nomura 2011; Yin et al. 2015a; Jiang &amp; Yin 2016, 2017; Zhang &amp; Yin 2021), with most of these described only recently. Some of the Chinese species are known inquilines of a wide range of ant genera including Aphaenogaster Mayr, Brachyponera Emery, Formica Linné, Lasius Fabricius, Myrmica Latreille, Nylanderia Emery, Odontomachus Latreille, Pheidole Westwood, and Pseudolasius Emery (Yin &amp; Li 2013b; Yin &amp; Shen 2015; Jiang &amp; Yin 2016, 2017). Batrisodes guoguotang was collected by sifting leaf litter, and there is no information on the biology of B. hanmi. The type material of Batrisodes pruinosus Reitter (a female) was collected by Russian ethnographer Grigory Potanin from Amdo (ȐƷ) (Besuchet 1981), an area historically in a broader Tibet, but now belonging to Qinghai Province. This species is here excluded from the Tibetan fauna. The following key is provided to help with the separation of the two Tibetan species.</p> <p>Key to Tibetan species of Batrisodes (male)</p> <p>1 Central part of vertex finely punctate (Fig. 38B); metatrochanter with strongly curved ventral projection (Fig. 38C); dorsal lobe of aedeagus split into three lobes near apex (Fig. 38F)....................................... B. guoguotang sp. nov.</p> <p>- Central part of vertex coarsely punctate (Fig. 39B); metatrochanter with slightly curved ventral spine (Fig. 38C); dorsal lobe of aedeagus split into two lobes near apex (Fig. 39H)............................................... B. hanmi sp. nov.</p></div> 	http://treatment.plazi.org/id/C822F213FF93242ECB88A65970FB2528	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF92242CCB88A28674B22502.text	C822F213FF92242CCB88A28674B22502.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisodes guoguotang Yin 2022	<div><p>Batrisodes guoguotang sp. nov.</p> <p>(Figs 38, 81A, 99D, G)</p> <p>Chinese common name: 果果Ƌqà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, forest nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.27383&amp;materialsCitation.latitude=29.31093" title="Search Plazi for locations around (long 95.27383/lat 29.31093)">Guoguotang Great Bend</a>, 29°18’39.35”N, 95°16’25.75”E, 1180 m; 2019.vii.22, leaf litter, Z.-W. Yin leg. [ƱẪẸṘ果果ḙ]’ (SNUC).</p> <p>Diagnosis. Male. Body length 2.37 mm. Head sub-rectangular at base, approximately as broad as pronotum; head and antenna lacking modifications. Pronotum with five dorsally visible longitudinal sulci, discal sulci strongly curved at middle, one pair of marginal sulci present ventral to lateral expansion. Elytron with short discal stria. Mesofemur and mesotibia simple; metatrochanter with large ventral projection, metatibia with blunt apical projection and long bunch of setae at apex. Aedeagus strongly asymmetric; median lobe with large basal capsule and foramen, dorsal lobe well-developed, with narrowed apex deeply forked, split to three lobes.</p> <p>Description. Male. Body (Fig. 38A) length 2.37 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 38B) sub-rectangular at base, slight wider than long, length 0.52 mm, width across eyes 0.55 mm; vertex finely punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with U-shaped sulcus connecting foveae, with sparse large punctures along outer margin of sulcus, lateral vertexal carina faint, extending from occipital constriction to level of middle of eyes, mediobasal carina as long as lateral ones; antennal tubercles moderately raised; frons impressed at middle, confluent with clypeus; clypeus with rough surface, its entire anterior margin carinate and raised; ocular-mandibular carina indistinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 20 large ommatidia. Antenna lacking distinct club and modification, length 1.24 mm; antennomere 1 thick, subcylindrical, 2–7 each elongate, gradually shorter, 8 smallest, 9 much larger than 8, 10 slightly broader than 9, 11 largest, as long as 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 38B) approximately as long as wide, length 0.50 mm, width 0.53 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, antebasal tubercles small, lacking discal spines, with one median, one pair of discal and one pair of lateral longitudinal sulci, discal sulci strongly curved at middle; with one pair of marginal sulci beneath lateral expansions; with small asetose median pit and large, setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, well-demarcated from hypomera by pair of lateral carinae, which extend posterolaterally to form complete hypomeral ridges; lateral procoxal foveae small; anterior and lateral margin of coxal cavity carinate.</p> <p>Elytra wider than long, length 0.69 mm, width 0.86 mm; each elytron with three large, asetose basal foveae, inner two close; discal stria only broadly and shallowly present for short distance posterior to outer basal fovea; small subhumeral fovea present, with carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by thin carinae; median mesoventral foveae narrowly separated, lateral mesoventral foveae deeply forked internally. Metaventrite moderately impressed at middle, with welldeveloped lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs more or less robust, lacking distinct modifications except for metatrochanter. Pro- and mesotrochanter each with tiny tubercle on ventral margin, pro- and mesotibia with small apical denticle; metatrochanter (Fig. 38C) with large, curved ventral projection, metatibia (Fig. 38D) with dense golden setae along mesal margin of apical 1/2, apex with small, blunt projection and elongate bunch of long setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.68 mm, width 0.79 mm. Tergite 1 (IV) approximately twice as long as 2 (V), with pair of short, triangular discal carinae, basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with thick marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) as long as 2 and 3 combined, with pair of small tubercles at lateral margins; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly impressed, sternite 7 (IX) (Fig. 38E) with asymmetric, membranous basal part and semicircular, well-sclerotized apical part.</p> <p>Aedeagus (Fig. 38F, G) 0.52 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection short, ventral stalk with two long preapical setae; dorsal lobe well-developed, apically narrowed and deeply forked.</p> <p>Female. Unknown.</p> <p>Comparative notes. Batrisodes guoguotang is apparently allied with B. hanmi by sharing most external characters, as well as a similar form of the aedeagus. The new species may be separated from B. hanmi by the following characters: 1) more strongly curved discal longitudinal sulci of the pronotum, 2) a smaller ventral denticle of the mesotrochanter (much larger ventral spine in B. hanmi), 3) a large projection on the ventral margin of the metatrochanter (small tubercle in B. hanmi), and 4) the apical portion of the aedeagal dorsal lobe split into with three lobes (split into two lobes in B. hanmi).</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 81A, 99D, G).</p> <p>Etymology. The new species is named after the Guoguotang Great Bend of Yarlung Tsangpo Grand Canyon located near Mêdog County.</p></div> 	http://treatment.plazi.org/id/C822F213FF92242CCB88A28674B22502	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF90242ACB88A2E571C022FE.text	C822F213FF90242ACB88A2E571C022FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Batrisodes hanmi Yin 2022	<div><p>Batrisodes hanmi sp. nov.</p> <p>(Figs 39, 81A, 98C, D)</p> <p>Chinese common name: 汗ṁqà甲</p> <p>Type material (4 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Mêdog County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.12945&amp;materialsCitation.latitude=29.364723" title="Search Plazi for locations around (long 95.12945/lat 29.364723)">Hanmi</a> (ca. 29°21’53”N, 95°07’46”E), alt. 2200 m; 19.viii.2005, Liang Tang leg. [ƱẪẸṘ汗ṁ]’ (SNUC). PARATYPE: CHINA: 2 ♂♂, 1 ♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length 2.50–2.58 mm. Head sub-rectangular at base, approximately as broad as pronotum; head and antenna lacking modifications. Pronotum with five dorsally visible longitudinal sulci, discal sulci slightly curved at middle, one pair of marginal sulci present ventral to lateral expansion. Elytron with short discal stria. Protibia with tiny apical denticle, mesotrochanter with distinct ventral spine, metatrochanter with small ventral tooth, metatibia with blunt apical projection and long bunch of setae at apex. Aedeagus strongly asymmetric; median lobe with large basal capsule and foramen, dorsal lobe well-developed, with apex deeply forked, split to two lobes. Female. Body length approximately 2.5 mm; legs lacking spine or projection, genitalia as in Fig. 39I.</p> <p>Description. Male. Body (Fig. 39A) length 2.50–2.58 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 39B) sub-rectangular at base, slight wider than long, length 0.50–0.51 mm, width across eyes 0.55– 0.57 mm; vertex roughly punctate, with large, asetose vertexal foveae (dorsal tentorial pits), with U-shaped sulcus connecting foveae, lateral vertexal carina faint, extending from occipital constriction to level of anterior margin of eyes, mediobasal carina short, extending anteriorly to level of middle of eyes; antennal tubercles moderately raised; frons impressed at middle, confluent with clypeus; clypeus with rough surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, distinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared opening, with distinct median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna lacking distinct club and modification, length 1.16–1.27 mm; antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, 9 much larger than 8, 10 slightly broader but shorter than 9, 11 largest, longer than 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 39B) approximately as long as wide, length 0.57 mm, width 0.58 mm, widest at middle; lateral margins rounded; disc slightly convex, finely punctate, antebasal tubercles small, lacking discal spines, with one median, one pair of discal and one pair of lateral longitudinal sulci, discal sulci curved at middle; with one pair of marginal sulci beneath lateral expansions; with small asetose median pit and large, setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, well-demarcated from hypomera by pair of lateral carinae, which extend posterolaterally to form complete hypomeral ridges; lateral procoxal foveae distinct; anterior and lateral margin of coxal cavity carinate.</p> <p>Elytra wider than long, length 0.77–0.78 mm, width 0.89–0.96 mm; each elytron with three large, asetose basal foveae, inner two close; discal stria only broadly and shallowly present for short distance posterior to outer basal fovea; small subhumeral fovea present, with carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by thin carinae; median mesoventral foveae narrowly separated, lateral mesoventral foveae deeply forked internally. Metaventrite moderately impressed at middle, with welldeveloped lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs more or less robust. Protrochanter simple, protibia with tiny apical spur; mesotrochanter with distinct, slightly curved ventral spine (Fig. 39C); metatrochanter with tiny ventral tooth (Fig. 39D; note: the male paratype has ventral margin smooth and lacks this tooth, which was treated as an intraspecific variation), metatibia (Fig. 39E) with dense golden setae along mesal margin of apical 1/2, apex with small, blunt projection and elongate bunch of long setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.63–0.67 mm, width 0.85–0.89 mm. Tergite 1 (IV) approximately twice as long as 2 (V), with pair of short, triangular discal carinae, basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with thick marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) as long as tergites 2 and 3 combined, with pair of small tubercles at lateral margins; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly impressed, sternite 7 (IX) (Fig. 39F) with asymmetric, membranous basal part and semicircular, well-sclerotized apical part.</p> <p>Aedeagus (Fig. 39G, H) 0.67 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, basoventral projection relatively long, middle and apical portion of ventral stalk each with one long seta; dorsal lobe well-developed, apically narrowed and deeply forked, split into two lobes.</p> <p>Female. Similar to male in external morphology; antenna approximately as long as that of male; legs lacking spines or projections; each compound eye composed of approximately 12 ommatidia. Measurements (as for male): body length 2.47 mm; length/width of head 0.51/ 0.59 mm, pronotum 0.56/ 0.56 mm, elytra 0.77/ 0.93 mm; abdomen 0.74/ 0.87 mm; length of antenna 1.19 mm; genitalia (Fig. 39I) moderately sclerotized, rounded triangular, maximum width 0.38 mm.</p> <p>Comparative notes. The new species is morphologically similar to B. guoguotang by sharing most external characters and may be separated from the latter by the following characters: 1) slightly curved discal longitudinal sulci of the pronotum (strongly curved in B. guoguotang), 2) a large ventral spine of mesotrochanter (with small ventral denticle in B. guoguotang), 3) a small ventral tooth of the metatrochanter (large ventral projection in B. guoguotang), and 4) the apical portion of aedeagal dorsal lobe split into two lobes (split into three lobes in B. guoguotang).</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 81A, 98C, D).</p> <p>Etymology. The specific name refers to the type locality of the new species, i. e., Hanmi.</p></div> 	http://treatment.plazi.org/id/C822F213FF90242ACB88A2E571C022FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF962416CB88A5BE712724F6.text	C822F213FF962416CB88A5BE712724F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomimus Yin 2022	<div><p>Coryphomimus gen. nov.</p> <p>(Figs 40–42)</p> <p>Chinese common name: ṈỀȁà甲¤</p> <p>Type species. Coryphomimus levigatus sp. nov. (by present designation).</p> <p>Diagnosis. Head roundly-triangular; lacking sulcus connecting vertexal foveae, lacking distinct frontal rostrum. Pronotum moderately expanded laterally, broad median longitudinal impression with carinate margins, with discal and lateral longitudinal carinae, disc lacking spines or denticles, with large, setose lateral antebasal, anterolateral and antero-hypomeral foveae, both inner and outer pair of small basolateral foveae present. Elytra with three basal foveae, inner two close, with long discal stria, subhumeral fovea and marginal stria present. Abdomen with tergite 1 (IV) longer than 2–3 (V–VI) combined, constricted near base, with complete inner and outer marginal carinae. Aedeagus stout, with long basoventral projection.</p> <p>Description. Body length 1.9–2.2 mm; habitus (Fig. 42A) elongate; dorsal surface with scattered exceptionally long setae and normal pubescence; antenna relatively elongate, extending to more than half elytral length when extended posteriorly.</p> <p>Head (Fig. 40A–C) roundly-triangular; lacking distinct frontal rostrum, antennal tubercles strongly raised; vertexal foveae (Fig. 40A; vf) relatively small, asetose, lacking sulcus connecting them, mediobasal and lateral carinae present; eyes small, ocular-mandibular carina (Fig. 40B; omc) complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with small gular foveae (Fig. 40C; gf) originating from shared round opening, gular carina thin; antenna 11-segmented, club formed by three apical antennomeres; maxillary palpus with small palpomere 1, 2 pedunculate basally and broad at apex, 3 triangular, 4 sub-fusiform, with small apical cone.</p> <p>Pronotum (Fig. 40D–F) transverse, lateral margins roundly expanded anterior to middle, convergent apically and constricted at basal 1/2, strongly incised posterior to lateral expansion, anterior margin slightly emarginate, posterior margin slightly curved; lateral antebasal foveae (Fig. 40D; laf) large, covered by dense setae, median antebasal fovea reduced to oval impression, with additional pairs of distinct, setose anterolateral and antero-hypomeral foveae (Fig. 40E; alf, ahf), two pairs of basolateral foveae present, lacking marginal, discal or antebasal spines/tubercles, median longitudinal (Fig. 40D; mls) sulcus broad, with carinate margins, with long discal and short lateral longitudinal carina (Fig. 40D; dlc, llc), lacking transverse antebasal sulcus; hypomeral ridge (Fig. 40E; hr) complete; prosternum with small lateral procoxal foveae (Fig. 40F; lpf).</p> <p>Elytra truncate at bases, each elytron with three basal foveae (Fig. 40G; bef), inner two close, lacking subbasal fovea, with long discal stria (Fig. 40G; ds) and complete sutural stria, with subhumeral fovea and carinate marginal sulcus (Fig. 40G, H; shf, ms).</p> <p>Mesoventrite with median foveae (Fig. 41A; mmsf) narrowly separated, in transverse opening, with large lateral mesoventral foveae not forked (straight) internally (Fig. 41A; lmsf); metaventrite with large, setose lateral coxal foveae (Fig. 41A; lmcf), lateral metaventral foveae (Fig. 41A; lmtf) in broad, shared opening, posterior margin with small, narrow split at middle; coxae broadly separated.</p> <p>Abdomen constricted near base; tergite 1 (IV) slightly longer than 2–3 (V–VI) combined, with mediobasal and one pair of basolateral foveae (Fig. 41B; mbf, blf)), lacking discal carinae, tergites 2–4 (V–VII) (Fig. 41C) each with one pair of small basolateral foveae and thin lateral carinae; sternite 2 (IV) with one pair of mediobasal (Fig. 41D; mbf) and two to three pairs of basolateral foveae (Fig. 41C, D; blf), sternites 3–5 (V–VII) each with two pairs of basolateral foveae.</p> <p>Legs moderately elongate, tarsomeres 2 and 3 subequal in length, with one major and one setiform claw.</p> <p>Males have relatively longer antenna than females, antennomeres 10 and 11 modified; trochanters and mesofemur with spines or projections on ventral margins; sternite 2 (IV) with three pairs of basolateral foveae; aedeagus (Fig. 42H, I) asymmetric, stout, with long basoventral projection, median lobe with large basal capsule, dorsal lobe embracing median lobe and with elongate sclerite, parameres reduced to single membranous structure. Female with two pairs of basolateral foveae on sternite 2 (IV); genitalia (Fig. 42J) weakly sclerotized.</p> <p>Comparative notes. Coryphomimus shares with a few species of Coryphomodes Jeannel the lack of marginal spines on the pronotum, and close inner two pairs of basal foveae of the elytra. The new genus differs from Coryphomodes and all other similar genera centered on Tribasodites Jeannel primarily by the presence of large, setose anterolateral and antero-hypomeral foveae of the prothorax; other characters that may bear generic importance are: moderately expanded lateral margins and presence of lateral longitudinal carinae (instead of sulci) of the pronotum, long discal striae of the elytra, and the abdomen is constricted near the base, with tergite 1 (IV) longer than 2 and 3 (V and VI) combined.</p> <p>Etymology. The generic name is a combination of Greek ‘ koryphç (head, top)’ and ‘ mîmos (imitator, actor)’, indicating the morphological similarity to Coryphomodes. The gender is masculine.</p> </div>	http://treatment.plazi.org/id/C822F213FF962416CB88A5BE712724F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFAA2415CB88A3B6768E2112.text	C822F213FFAA2415CB88A3B6768E2112.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomimus levigatus Yin 2022	<div><p>Coryphomimus levigatus sp. nov.</p> <p>(Figs 42, 81B, 98C, D, 99F, 100A)</p> <p>Chinese common name: 光ẚṈỀȁà甲</p> <p>Type material (14 ex.). HOLOTYPE: CHINA: ♂, ‘ China: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.50278&amp;materialsCitation.latitude=29.685833" title="Search Plazi for locations around (long 95.50278/lat 29.685833)">Xiang</a>, Mêdog County, nr. 80K, 29°41’09”N, 95°30’10”E, alt. 2330 m, mixed leaf litter, sifted, 09.vii.2018, Cheng, Peng &amp; Shen leg. (中ǞƱẪẸṘ县 80K ṉ ữ)’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 7 ♀♀, same data as that of holotype; 1 ♂, 2 ♀♀, ‘ China: Mêdog County, pass from Bari Village to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.3575&amp;materialsCitation.latitude=29.3125" title="Search Plazi for locations around (long 95.3575/lat 29.3125)">Renqingbeng Temple</a>, 29°18’45”N, 95°21’27”E, ca. 1830 m, 19.iii.2017, X.-B. Song leg. [ƱẪẸṘ县巴U村 -仁ů崩寺]’; 1 ♀ ‘ China: Xizang A. R., Motuo Co., Hanmi, alt. 2200 m, 19.viii.2006, Tang Liang leg.’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length 2.16–2.20 mm; dorsal surface of body with scattered exceptionally long setae and normal pubescence. Head subtruncate at base, narrower than pronotum, tempus moderately short, rounded at posterolateral angle, vertex lacking sulcus, with relatively small, setose foveae, with long mediobasal carina extending from head base anteriorly to frons, with thin lateral carinae from subbase to posterior margin of strongly raised antennal tubercles; antenna elongate, with modified antennomeres 10–11, antennomere 10 with moderately large basal cavity on ventral surface, 11 with short basal projection. Pronotum with laterally carinate median longitudinal sulcus, with pairs of discal and short lateral carinae, lacking antebasal or marginal spines. Discal stria of elytron long, extending posteriorly to more than apical 1/5 of elytral length; disc finely punctate. Pro- and mesotrochanter and mesofemur with small, blunt ventral spine; metatrochanter with short, apically curved projection on ventral margin. Aedeagus strongly asymmetric, stout; median lobe with large basal capsule and slightly oval foramen, ventral stalk broad at middle and then abruptly narrowing apically on left side, dorsal lobe embracing median lobe, non-articulated, with elongate sclerite forked at apex. Female. Body length 1.89–2.04 mm; antenna lacking modifications, legs lacking spine or projection, genitalia as in Fig. 42J.</p> <p>Description. Male. Body (Fig. 42A) length 2.16–2.20 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence, with scattered exceptionally long setae present on vertex, lateral margins and discs of pronotum and elytra, meso- and metatibia and tergites 1–4 (IV–VII).</p> <p>Head (Fig. 42B) subtruncate at base, much wider than long, length 0.43 mm, width across eyes 0.51–0.52 mm; vertex smooth, markedly raised at middle, lacking sulcus, vertexal foveae (dorsal tentorial pits) setose, relatively small, mediobasal carina distinct and long, extending from head base anteriorly to level of bases of antennal tubercles, lateral carina thin, extending from subbase anteriorly to base of antennal tubercle; posterolateral angle rounded; frons anteriorly demarcated from clypeus by frontal-clypeal ridge, weakly impressed between large, strongly raised antennal tubercles; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with median carina faint from opening and more distinct near mouthparts. Compound eyes prominent, composed of approximately 40 ommatidia. Antenna elongate, length 1.25 mm, distinct club (Fig. 42C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much wider and longer than 8, sub-moniliform, 10 (Fig. 42D) much broader and longer than 9, asymmetric, ventral surface with moderately large cavity at base, 11 largest, much longer than 9 and 10 combined, sub-fusiform, with short and broad projection at base of mesal margin.</p> <p>Pronotum (Fig. 42B) wider than long, length 0.49–0.50 mm, width 0.58 mm, widest anterior middle; lateral margins rounded, strongly incised posterior to lateral humps, convergent basally and sub-parallel at basal 1/4; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of discal and short lateral longitudinal carinae, lacking antebasal, discal, or marginal spines/tubercles; with large, setose anterolateral and lateral antebasal foveae; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit and large, setose antero-hypomeral fovea; margin of coxal cavity broadly carinate.</p> <p>Elytra much wider than long, length 0.63 mm, width 0.75 mm; each elytron with three moderately large, asetose basal foveae, inner two close; long discal stria extending posteriorly from outer basal fovea to more than apical 1/5 of elytral length; humerus moderately prominent, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae moving medially into large, setose shared impression; posterior margin weakly emarginate at middle, with thin split.</p> <p>Legs elongate, femora; procoxa with exceptionally long seta at base, protrochanter with blunt ventral spine; mesotrochanter (Fig. 42E) with distinct blunt spine on ventral margin, mesofemur (Fig. 42E) with small ventral denticle near base; metatrochanter (Fig. 42F) with broad, apically curved projection on ventral margin, metatibia with short pencil-like apical tuft of setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.64–0.69 mm, width 0.66–0.68 mm. Tergite 1 (IV) more than twice as long as 2 (V), with large mediobasal foveae at lateral margins of short median sulcus, with one pair of large basolateral foveae, lacking discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of small basolateral foveae and pair of thin lateral carinae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, sternites 3–4 with tiny tubercle at middle, 3–5 each with two pairs of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 42G) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 42H, I) 0.41 mm long, in dorsal view asymmetric, stout; median lobe with broad basal capsule and slightly oval foramen, with long basoventral projection, ventral stalk in dorsal view broad at middle and abruptly narrowing apically at left side, with rounded, membranous apex; dorsal lobe broad, embracing median lobe, with elongate sclerite apically forked; parameres membranous, apically split into two lobes.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter; antenna lacking modifications, legs lacking spines or projections; each compound eye composed of approximately 13 ommatidia; sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae. Measurements (as for male): body length 1.89–2.04 mm; length/width of head 0.41/ 0.49 mm, pronotum 0.44–0.46/ 0.54–0.56 mm, elytra 0.49–0.52/ 0.67–0.71 mm; abdomen 0.61–0.66/ 0.67–0.70 mm; length of antenna 1.00– 1.10 mm; genitalia (Fig. 42J) weakly sclerotized, transverse, maximum width 0.36 mm.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 81B, 98C, D, 99F, 100A).</p> <p>Etymology. The specific epithet ‘ levigatus (- a, - um)’ is a Latin adjective meaning ‘smoothed, polished’, referring to the smooth, aspinose lateral margins of the pronotum of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FFAA2415CB88A3B6768E2112	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFA82414CB88A1DB70FB2582.text	C822F213FFA82414CB88A1DB70FB2582.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomodes Jeannel 1960	<div><p>Coryphomodes Jeannel, 1960</p> <p>Chinese common name: Ềȁà甲¤</p> <p>Coryphomodes Jeannel, 1960: 416; Newton &amp; Chandler 1989 (catalog); Chandler 2001 (redescription); Löbl &amp; Kurbatov 2001 (discussion of relationship). Type species: Coryphomodes cristatus Jeannel, 1960: 418 (original designation).</p> <p>Members of Coryphomodes have the inner two basal foveae of the elytra close, or to a greater degree, fused, and have marginal spines of the pronotum small or absent. Further characterization of this genus appears to be difficult, as the included species may represent polyphyletic and geographically widely separated groups. For example, C. cristatus (type species), C. calcaratus Jeannel and C. humeralis Jeannel from India share long carinate discal striae of the elytra and short first visible tergites (IV) (Löbl &amp; Kurbatov 2001), while C. dionysius (Schaufuss, L. W.) from Japan and the diverse Australian members have short or lack elytral discal striae, and the tergites 1 (IV) are much longer than tergites 2 (V) (Jeannel 1958; Chandler 2001). The three new species described here all have three basal elytral foveae, as does the type species, but have long tergites 1 more than twice the length of tergites 2. The Tibetan species may be linked by the coarsely punctate elytral disc, and may be separated by using the following key.</p> <p>Key to Tibetan species of Coryphomodes (male)</p> <p>1 Body length over 2.0 mm; vertex lacking lateral impressions (Fig. 45B)......................... C. chenzhilini sp. nov.</p> <p>- Body length less than 2.0 mm; vertex with round lateral impressions............................................. 2</p> <p>2 Vertex lacking mediobasal carina (Fig. 43B); areas lateral to median tubercle on frons smooth (Fig. 43B); marginal tubercles of pronotum more distinct (Fig. 43B); punctures of elytral disc sparser (Fig. 43C)........................ C. budda sp. nov.</p> <p>- Vertex with short mediobasal carina (Fig. 44B); areas lateral to median tubercle on frons with setose tufts (Fig. 44B); marginal tubercles of pronotum tiny (Fig. 44B); punctures of elytral disc coarser (Fig. 44C)................. C. cephalicus sp. nov.</p></div> 	http://treatment.plazi.org/id/C822F213FFA82414CB88A1DB70FB2582	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFA82412CB88A57C773225B2.text	C822F213FFA82412CB88A57C773225B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomodes budda Yin 2022	<div><p>Coryphomodes budda sp. nov.</p> <p>(Figs 43, 81C, 99F, 100A)</p> <p>Chinese common name: ṞĽỀȁà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘China: Tibet (Mêdog County), pass nr. Renqingbeng Temple (29°18’16”N, 95°21’13”E), alt. 1700 m; 17.viii.2016, Zhi-Lin Chen leg. [ƱẪẸṘ仁ů崩寺]’ (SNUC). PARATYPE: CHINA: 1 ♂, ‘China: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.488884&amp;materialsCitation.latitude=29.65611" title="Search Plazi for locations around (long 95.488884/lat 29.65611)">Mêdog County</a>, 80K (ca. 29°39’22”N, 95°29’20”E), 2110 m, 11.viii.2016, Z.-L. Chen leg. [ƱẪẸṘ县 80K]’ (SNUC).</p> <p>Diagnosis. Male. Body length 1.75 mm. Head rounded triangular, as broad as pronotum, tempus short, vertex and frons modified, vertex with lateral impressed areas, frons with median tubercle; antenna lacking modifications. Pronotum with laterally carinate median sulcus and lateral longitudinal sulci, with large antebasal spines and small marginal spines. Elytron with inner two basal foveae close, discal stria shallow and short, extending posteriorly to basal 1/4 of elytral length; disc with sparse, large punctures at posterior 3/4. Fore and middle legs simple, metatrochanter with large ventral projection. Aedeagus strongly asymmetric, stout; ventral stalk protruding apically, dorsal lobe plate-like, parameres reduced.</p> <p>Description. Male. Body (Fig. 43A) length 1.75 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 43B) rounded triangular, wider than long, length 0.34–0.35 mm, width across eyes 0.44 mm; vertex smooth, lacking foveae, mediobasal area strongly concave, laterally with round impressed areas, anteriorly preceded by short, transverse ridge; frons anteriorly confluent with clypeus, with antero-lateral ridges, with strongly raised tubercle at middle, sides of tubercle with short setae, posterior margin with tuft of thickened setae; clypeus with smooth surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, distinct. Venter with granulate surface, mediobasal area broadly impressed; small gular foveae (posterior tentorial pits) originating from shared transverse opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 35 ommatidia. Antenna lacking modification, length 0.78 mm, distinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each moniliform, 2 as long as wide, 3–7 short, each more or less transverse, 8 smallest, 9 much larger than 8, transverse, 10 broader but shorter than 9, 11 largest, longer than 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 43B) wider than long, length 0.39–0.40 mm, width 0.44 mm, widest posterior middle; lateral margins rounded at apical 1/2, basally subparallel; disc slightly convex, with sparse but distinct punctures, median longitudinal sulcus with carinate sides, with pair of shallow lateral longitudinal sulci, antebasal tubercles large, lacking discal spines; with small marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra wider than long, length 0.52–0.54 mm, width 0.67–0.68 mm; each elytron (Fig. 43C) with three large, asetose basal foveae, inner two close; discal stria shallow and short, extending posteriorly from outer basal fovea to basal 1/4 of elytral length; humerus weakly and broadly prominent, lacking subhumeral fovea, carinate marginal stria extending from basal 2/5 to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated in shared oval opening, large lateral mesoventral foveae forked internally, with short mesoventral process. Metaventrite prominent admesally, moderately impressed at middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina complete, extending from anterior margin of mesoventrite to posterior margin of metaventrite.</p> <p>Legs more or less robust; trochanters each with exceptionally long seta on ventral margin. Mesotrochanter with tiny tubercle on ventral margin; metatrochanter (Fig. 43D) with large ventral projection curved at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.46–0.50 mm, width 0.62–0.64 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking mediobasal foveae or discal carinae, basal sulcus thin, with large basolateral foveae at lateral margins of sulcus, inner marginal carina slightly oblique, complete, outer one thicker, complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and two pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 43E) membranous, with weakly sclerotized apex.</p> <p>Aedeagus (Fig. 43F, G) 0.24 mm long, strongly asymmetric, stout; median lobe with broad basal capsule and slightly oval foramen, basoventral projection short, ventral stalk with one long, thickened seta at middle, protruding and subtruncately rounded at apex; non-articulated dorsal plate short and flattened, roundly rectangular; parameres reduced.</p> <p>Female. Unknown.</p> <p>Comparative notes. The new species is morphologically similar to C. cephalicus and C. chenzhilini, and can be readily separated from both species by the smaller body size, a stouter habitus, and the characteristic cephalic modifications and form of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 81C, 99F, 100A).</p> <p>Etymology. The species name refers to the tantric Buddhist Guru Rimpoche (Äǩffik士), who is enshrined and worshiped at Renqingbeng Temple.</p></div> 	http://treatment.plazi.org/id/C822F213FFA82412CB88A57C773225B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFAE2410CB88A57271852222.text	C822F213FFAE2410CB88A57271852222.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomodes cephalicus Yin 2022	<div><p>Coryphomodes cephalicus sp. nov.</p> <p>(Figs 44, 81C, 99E, 100A)</p> <p>Chinese common name: 焰ĦỀȁà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘ China: nr. Mêdog County, alt. 1280 m, 29°18’48”N, 95°19’17”E; 18.iii.2017, X.-B. Song leg. [ƱẪẸṘ县]’ (SNUC). PARATYPE: CHINA: 1 ♀, ‘ China: Mêdog County, 80K (ca. 29°39’22”N, 95°29’20”E), 2110 m, 11.viii.2016, Z.-L. Chen leg. [ƱẪẸṘ县 80K]’ (SNUC).</p> <p>Diagnosis. Male. Body length 1.87 mm. Head rounded triangular, as broad as pronotum, tempus moderately long, vertex and frons modified, vertex with lateral impressed areas and large central cavity, frons with median projection covered with dense setae; antenna lacking modifications. Pronotum with laterally carinate median sulcus and lateral longitudinal sulci, with small antebasal and marginal spines. Elytron with inner two basal foveae close, discal stria broad and shallow, short, extending posteriorly from outer basal fovea to basal 1/4 of elytral length; disc with moderately sparse, large punctures. Fore legs simple, mesotrochanter with tiny ventral tubercle, metatrochanter with blunt triangular ventral spine. Aedeagus strongly asymmetric, stout; ventral stalk composed two plates apically, dorsal lobe plate-like, parameres reduced. Female genitalia as in Fig. 44G.</p> <p>Description. Male. Body (Fig. 44A) length 1.87 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 44B) rounded triangular, as wide as long, length 0.42 mm, width across eyes 0.43 mm; vertex smooth, large mediobasal cavity with ridged posterior margin, small foveae at anterolateral margin of cavity, areas lateral to cavity with round impressions filled with short setae, anterior margin of cavity formed by thick, transverse ridge; frons anteriorly confluent with clypeus, with antero-lateral ridges, with large central tubercle covering with dense setae at middle, with short longitudinal ridge and tuft of setae lateral to tubercle; clypeus with smooth surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, distinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared transverse opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 45 ommatidia. Antenna lacking modification, length 0.93 mm, indistinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each moniliform, 2 elongate, 3–8 short, each slightly elongate or as long as wide, 8 smallest, 9 much larger than 8, subconical, 10 slightly broader but shorter than 9, 11 largest, slightly shorter than 9 and 10 combined, subconical.</p> <p>Pronotum (Fig. 44B) approximately as long as wide, length 0.43 mm, width 0.42 mm, widest posterior middle; lateral margins rounded at apical 1/2, basally evenly convergent; disc slightly convex, with sparse but distinct punctures, median longitudinal sulcus with carinate sides, with pair of shallow lateral longitudinal sulci; with small antebasal and marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra wider than long, length 0.61 mm, width 0.77 mm; each elytron (Fig. 44C) with three large, asetose basal foveae, inner two close; discal stria broad and shallow, short, extending posteriorly from outer basal fovea to basal 1/4 of elytral length; humerus weakly and broadly prominent, lacking subhumeral fovea, carinate marginal stria extending from basal 1/4 to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared oval opening, large lateral mesoventral foveae forked internally, with short mesoventral process. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina complete, extending from anterior margin of mesoventrite to posterior margin of metaventrite.</p> <p>Legs more or less robust; protrochanter with exceptionally long seta on ventral margin, mesotrochanter with tiny tubercle on ventral margin; metatrochanter (Fig. 44D) with distinct rounded triangular spine on ventral margin.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.50 mm, width 0.70 mm. Tergite 1 (IV) approximately twice as long as 2 (V), lacking mediobasal foveae or discal carinae, basal sulcus thin, with large basolateral foveae at lateral margins of sulcus, inner marginal carina slightly oblique, complete, outer one thicker, complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and two pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) entirely membranous, flattened.</p> <p>Aedeagus (Fig. 44E, F) 0.23 mm long, strongly asymmetric, stout; median lobe with broad basal capsule and subtriangular foramen, basoventral projection short, ventral stalk apically expanded, short, truncate at apex; nonarticulated dorsal plate short and flattened, subtruncate; parameres reduced.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter; head and legs lacking modifications, vertex with reversed U-shaped sulcus connecting small foveae; each compound eye composed of approximately 28 ommatidia. Measurements (as for male): body length 1.83 mm; length/width of head 0.41/ 0.40 mm, pronotum 0.42/ 0.44 mm, elytra 0.60/ 0.75 mm; abdomen 0.49/ 0.70 mm; length of antenna 0.90 mm; genitalia (Fig. 44G) weakly sclerotized, transverse, maximum width 0.19 mm.</p> <p>Comparative notes. This species is morphologically most similar to the Tibetan C. budda and C. chenzhilini, and can be readily separated from both species by the large punctures that cover the dorsal surface of the elytra, and the characteristic cephalic modifications and form of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 81C, 99E, 100A).</p> <p>Etymology. The specific epithet refers to the strong cephalic modification of the new species.</p></div> 	http://treatment.plazi.org/id/C822F213FFAE2410CB88A57271852222	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFAC241ECB88A582749F230A.text	C822F213FFAC241ECB88A582749F230A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coryphomodes chenzhilini Yin 2022	<div><p>Coryphomodes chenzhilini sp. nov.</p> <p>(Figs 45, 81C, 100A)</p> <p>Chinese common name: Ŀ氏Ềȁà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.488884&amp;materialsCitation.latitude=29.65611" title="Search Plazi for locations around (long 95.488884/lat 29.65611)">China</a>: Mêdog County, 80K (ca. 29°39’22”N, 95°29’20”E), 2110 m, 11.viii.2016, Z.-L. Chen leg. [ƱẪẸṘ县 80K]’ (SNUC).</p> <p>Diagnosis. Male. Body length 2.20 mm. Head rounded triangular, approximately as broad as pronotum, tempus moderately long, vertex and frons lacking modifications, lacking distinct sulcus or carina, vertexal foveae asetose, relatively large, with thin, short mediobasal carina; antenna lacking modifications. Pronotum with laterally carinate median sulcus and lateral longitudinal sulci, with small antebasal and marginal spines. Elytron with inner two basal foveae close, discal stria broad and shallow, short, extending posteriorly to apical 2/3 of elytral length; disc with moderately dense, large punctures. Fore legs simple, mesotrochanter with large ventral spine, metatrochanter with large, curved ventral projection.Aedeagus strongly asymmetric, stout; ventral stalk apically divided into ventral and dorsal sclerites, dorsal lobe large, parameres reduced.</p> <p>Description. Male. Body (Fig. 45A) length 2.20 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 45B) rounded triangular, slightly wider than long, length 0.44 mm, width across eyes 0.49 mm; vertex smooth, lacking modifications, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina thin, extending from head base for short distance anteriorly; frons anteriorly confluent with clypeus, with curved antero-lateral ridges; clypeus with smooth surface, its entire anterior margin carinate and moderately raised; ocular-mandibular carina complete, distinct. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared transverse slit, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 45 ommatidia. Antenna lacking modification, length 1.06 mm, distinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each slightly elongate, 8 smallest, 9 much larger than 8, subconical, 10 broader and longer than 9, subconical, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 45B) approximately as long as wide, length 0.45 mm, width 0.46 mm, widest at middle; lateral margins rounded at apical 1/2, evenly convergent basally; disc slightly convex, with sparse but distinct punctures, median longitudinal sulcus with carinate sides, with pair of lateral longitudinal sulci; with small antebasal and marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra wider than long, length 0.67 mm, width 0.80 mm; each elytron (Fig. 45C) with three large, asetose basal foveae, inner two close; discal stria broad and shallow, short, extending posteriorly from outer basal fovea to apical 2/3 of elytral length; humerus weakly and broadly prominent, lacking subhumeral fovea, carinate marginal stria extending from basal 1/4 to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, large lateral mesoventral foveae forked internally, with short mesoventral process. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina complete, extending from anterior margin of mesoventrite to posterior margin of metaventrite.</p> <p>Legs elongate; foreleg simple, mesotrochanter (Fig. 45D) with large, triangular spine on ventral margin; metatrochanter (Fig. 45E) with large, apically curved ventral projection.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.61 mm, width 0.75 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking mediobasal foveae or discal carinae, basal sulcus thin, with large basolateral foveae at lateral margins of sulcus, inner marginal carina slightly oblique, complete, outer one thicker, complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and two pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 45F) membranous, weakly sclerotized laterally.</p> <p>Aedeagus (Fig. 45G, H) 0.36 mm long, strongly asymmetric, stout; median lobe with broad basal capsule and rounded triangular foramen, basoventral projection short, ventral stalk apically expanded, with narrowed dorsal sclerite truncate at apex; non-articulated dorsal lobe long and flattened, apically membranous; parameres reduced.</p> <p>Female. Unknown.</p> <p>Comparative notes. This new species is morphologically similar to the two previously described species and shares with them a similar general form of the aedeagus. It can be readily separated from both species by the lack of modifications on the male head, markedly denser large punctures of the elytra, presence of a large ventral spine on the male mesotrochanter, as well as the characteristic form of the aedeagus.</p> <p>Distribution. Mêdog County, Tibet, SW China (Fig. 81C, 100A).</p> <p>Etymology. The new species is named after myrmecologist Zhi-Lin Chen, who collected the holotype and provided the material for study.</p></div> 	http://treatment.plazi.org/id/C822F213FFAC241ECB88A582749F230A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFA2241ACB88A4EB763C243E.text	C822F213FFA2241ACB88A4EB763C243E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gyrongita Yin 2022	<div><p>Gyrongita gen. nov.</p> <p>(Figs 46–48)</p> <p>Chinese common name: 小吉Pà甲¤</p> <p>Type species. Gyrongita uniformis sp. nov. (by present designation)</p> <p>Diagnosis. Head subtriangular; vertex strongly convex, with reversed U-shaped sulcus connecting foveae, lacking distinct frontal rostrum, tempus angulate; eyes prominent. Pronotum with thin median and lateral longitudinal sulci and short discal carinae, small antebasal and marginal denticles present, with small, nude lateral antebasal and inner and outer pair of basolateral foveae. Elytra with three small basal foveae, lacking discal stria, subhumeral fovea and marginal stria present. Abdomen with tergite 1 (IV) longer than 2–3 (V–VI) combined, constricted near base and narrowing apically; tergite 1 with complete inner marginal carinae, lacking outer marginal carinae; sternites 3–5 (V–VII) lacking foveae. Aedeagus simple, flattened.</p> <p>Description. Body small, length less than 1.7 mm; habitus (Fig. 48A) elongate; dorsal surface with dense, relatively long setae; antenna relatively short, surpassing pronotal base when extended posteriorly.</p> <p>Head (Fig. 46A–C) subtriangular; lacking distinct frontal rostrum, antennal tubercles weakly raised; vertexal foveae (Fig. 46A; vf) relatively small, asetose, connected by reversed U-shaped sulcus, with short mediobasal and two pairs of lateral carinae; eyes prominent, ocular-mandibular carina (Fig. 46B; omc) complete, extending to posteroventral articulation of mandible. Venter with small gular foveae (Fig. 46C; gf) originating from shared opening, gular carina present for short distance behind mouthparts; antenna 11-segmented, club loosely formed by three apical antennomeres; maxillary palpus with small palpomere 1, 2 basally pedunculate and broad at apex, 3 subquadrate, 4 fusiform, with small apical cone.</p> <p>Pronotum (Fig. 46D–F) moderately transverse, lateral margins rounded anterior to middle, convergent apically and constricted at basal 2/5, anterior margin slightly emarginate, posterior margin slightly curved; with thin median and lateral longitudinal sulci (Fig. 46D; mls, lls) and short mediobasal and discal carinae (Fig. 46D; mbc, dc), lateral antebasal foveae (Fig. 46E; laf) small and asetose, two pairs of basolateral foveae (Fig. 46D; blf) present, with small marginal and two pairs of antebasal tubercles, lacking antebasal sulcus; constricted basal portion with rough surface; hypomeral ridge (Fig. 46E; hr) complete; prosternum with small lateral procoxal foveae (Fig. 46F; lpf).</p> <p>Elytra truncate at bases, each elytron with three widely separated basal foveae (Fig. 46G; bef), lacking subbasal fovea, discal stria (Fig. 46G; ds) presented but short, represented by indistinct impression, sutural stria complete, with subhumeral fovea and marginal sulcus (Fig. 46G, H; shf, ms).</p> <p>Mesoventrite with median foveae (Fig. 47A; mmsf) moderately broadly separated in transverse opening, with large lateral mesoventral foveae deeply forked internally (Fig. 47A; lmsf); metaventrite with large, setose lateral coxal foveae (Fig. 47A; lmcf), lateral metaventral foveae (Fig. 47A; lmtf) separated, posterior margin with small, narrow split at middle; coxae broadly separated.</p> <p>Abdomen constricted near base; tergite 1 (IV) slightly longer than 2–3 (V–VI) combined, with mediobasal and two pairs of basolateral foveae (Fig. 47B; mbf, blf), lacking discal carinae, tergites 2–4 (V–VII) each with one pair of tiny basolateral foveae and thin margin carinae; sternite 2 (IV) with one pair of mediobasal (Fig. 47D; mbf) and three pairs of basolateral foveae (Fig. 47C, D; blf), sternites 3–5 (V–VII) lacking foveae.</p> <p>Legs moderately elongate, tarsomeres 2 and 3 subequal in length, with one major and one setiform claw.</p> <p>Males lacking observable sexual characters; aedeagus (Fig. 48E, F) slightly asymmetric in dorsal view, flattened, with curved basoventral projection, median lobe with large basal capsule, dorsal lobe plate-like, embracing median lobe, parameres reduced to single membranous structure. Female unknown.</p> <p>Comparative notes. Gyrongita is defined here by a unique combination of characters that occur in various genera rather than by a definitive synapomorphy. It belongs to a group including Tribasodites, Coryphomodes, and those genera that possess spinose pronotal lateral margins, lack a transverse antebasal sulcus of the pronotum, and have three basal foveae on each elytron (= Nomura &amp; Idris (2003) ’s Tribasodes -group). Gyrongita differs from all known Asian genera of this group by the small body size, a subtriangular head with a strongly convex vertex, the angulate tempora and prominent eyes, the lack of a distinct gular carina, the pronotum with small marginal and antebasal tubercles, a roughened constricted base of the pronotum, the lack of foveae on sternites 3–5 (V–VII), as well as the lack of obvious sexual characters on the male.</p> <p>Etymology. The new generic name is combined from ‘Gyirong (= Gyrong)’, the type locality of the type species, and the suffix ‘- ita (small, little)’. The gender is feminine.</p></div> 	http://treatment.plazi.org/id/C822F213FFA2241ACB88A4EB763C243E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFA62419CB88A3FF760D22FE.text	C822F213FFA62419CB88A3FF760D22FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gyrongita uniformis Yin 2022	<div><p>Gyrongita uniformis sp. nov.</p> <p>(Figs 48, 82A, 90)</p> <p>Chinese common name: 同型小吉Pà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32806&amp;materialsCitation.latitude=28.376945" title="Search Plazi for locations around (long 85.32806/lat 28.376945)">Gyirong Valley</a>, slope nr. Jifu Vill., 28°22’37”N, 85°19’41”E, 2400–2700 m, 22.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW吉 ė村ṉữ山坡 ’ (SNUC). PARATYPE: CHINA: 1 ♂, same data as that of holotype’ (SNUC).</p> <p>Diagnosis. Male. Body length less than 1.7 mm; dorsal surface of body with dense, moderately long pubescence. Head subtriangular, transverse, slightly broader than pronotum, tempus moderately long, angulate at posterolateral angle, vertex with reversed U-shaped sulcus connecting small, asetose foveae, with short mediobasal carina extending from near head base anteriorly to point posterior to middle of eyes, with two pairs of distinct lateral carinae from base to posterior margin of weakly raised antennal tubercles; antenna moderately short, lacking modifications, club loosely formed by apical three moderately enlarged antennomeres. Pronotum with thin median and lateral longitudinal sulci, with pair of short discal carinae, with small antebasal and marginal tubercles. Discal stria of elytron indistinct, short; disc finely punctate. Legs lacking modifications. Aedeagus in dorsal view symmetric, flattened; median lobe with large, rounded triangular foramen, ventral stalk laterally narrowed, dorsal lobe embracing ventral stalk, lobe plate-like, parameres reduced to single membranous structure.</p> <p>Description. Male. Body (Fig. 48A) length 1.59–1.66 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 48B) subtriangular, wider than long, length 0.30–0.32 mm, width across eyes 0.39–0.40 mm; vertex smooth, markedly raised at middle, with reversed U-shaped sulcus connecting asetose, widely separated vertexal foveae (dorsal tentorial pits), mediobasal carina short, extending from near head base anteriorly to extend past posterior margin of eyes, with two pairs of distinct lateral carinae, both extending from near head base anteriorly to base of antennal tubercle; posterolateral angle angulate; frons anteriorly demarcated from clypeus by oblique frontalclypeal ridge, weakly impressed between weakly raised antennal tubercles; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with faint median carina present for short distance posterior to mouthparts. Compound eyes prominent, conical, each composed of approximately 45 ommatidia. Antenna relatively short, length 0.79 mm, club (Fig. 48C) loosely formed by moderately enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each moniliform, 9 much larger than 8, 10 much longer than 9, 11 largest, as long as 9 and 10 combined, sub-conical.</p> <p>Pronotum (Fig. 48B) approximately as long as wide, length 0.34–0.35 mm, width 0.35–0.37 mm, widest at middle; lateral margins rounded, convergent apically and basally; disc slightly convex, finely punctate, with thin median longitudinal sulcus and short mediobasal carina, with pair of short, indistinct discal carinae and lateral longitudinal impressions, with small antebasal, discal and marginal tubercles; with small, asetose lateral antebasal and outer and inner pair of basolateral foveae; collar region roughened. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity slightly carinate.</p> <p>Elytra much wider than long, length 0.44–0.46 mm, width 0.55–0.59 mm; each elytron with three moderately small, widely separated asetose basal foveae; indistinct discal stria extending posteriorly from outer basal fovea to slightly less than half of elytral length; humerus denticulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae moderately widely separated, originating from shared opening, large lateral mesoventral foveae deeply forked internally, with broad anterior mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae widely separated; posterior margin weakly emarginate at middle, with narrow split.</p> <p>Legs moderately elongate, lacking modifications.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.52–0.53 mm, width 0.51–0.55 mm. Tergite 1 (IV) more than twice as long as 2 (V), with one pair of mediobasal and two pairs of basolateral foveae, lacking discal carinae, inner marginal carina oblique, complete, outer marginal carina lacking; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of small basolateral foveae and pair of thin lateral carinae, tergite 5 (VIII) semicircular, posterior margin weakly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, with pair of lateral carinae; midlength of sternites 2–5 (IV–VVI) gradually shorter, sternites 3–5 lacking foveae, sternite 6 (VIII) short, strongly transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 48D) membranous, consisting of pair of subtriangular lateral plates and one elongately oval central plate.</p> <p>Aedeagus (Fig. 48E, F) 0.25 mm long, in dorsal view flattened; median lobe with broad basal capsule and large foramen, with curved basoventral projection, ventral stalk laterally narrowed and with rounded apex, with two apical setae; dorsal lobe broad, plate-like, embracing median lobe; parameres membranous, reduced to single membranous structure.</p> <p>Female. Unknown</p> <p>Distribution. Gyirong County, Tibet, SW China (Figs 82A, 90).</p> <p>Etymology. The specific epithet ‘ uniformis, - e ’ is a Latin adjective meaning ‘uniform’, referring to the lack of observable male secondary sexual characters of the new species.</p> </div>	http://treatment.plazi.org/id/C822F213FFA62419CB88A3FF760D22FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFA42406CB88A76A70FB271F.text	C822F213FFA42406CB88A76A70FB271F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes Westwood 1870	<div><p>Sathytes Westwood, 1870</p> <p>Chinese common name: ḛà甲¤</p> <p>Sathytes Westwood, 1870: 128; Löbl 1979 (revision of world species); Yin et al. 2012a (redescription); Yin &amp; Shen 2020 (checklist of world species). Type species: Sathytes punctiger Westwood, 1870: 128 (monotypy).</p> <p>= Batoxylina Jeannel, 1957: 8. Type species: Batoxylina clavalis Jeannel, 1957: 9 (original designation). Synonymized by Löbl 1979: 285.</p> <p>This is a rather easily recognizable genus with all included members possessing a coarsely punctate dorsal surface of the body, and a great number of foveae at the elytral bases and the bases of the abdominal segments. The males of all species, except S. simplex Löbl, possess strongly modified and enlarged antennal clubs of characteristic forms, thus making identifications less difficult in most cases with enough careful comparisons. In Tibet, it is common to find several species at the same location, making it sometimes impossible to associate females with males of species that have a similar body size with 100% assurance. Sathytes is diverse in China, and seven species have been previously reported from Tibet (Yin et al. 2012a; Shen &amp; Yin 2020; Yin &amp; Shen 2020), with another 12 species being described here. Additionally, Sathytes simplex is newly reported to occur in Tibet.</p> <p>The 20 Tibetan species are keyed below. It should be kept in mind that in some cases a direct and careful comparison between the specimens is the only means for separating morphologically similar species due to often different angles showing the antennal modifications in published literatures.</p> <p>Key to Tibetan species of Sathytes (male)</p> <p>1 Antenna simple, lacking modifications (Fig. 58C)............................................... S. simplex Löbl</p> <p>- Antenna with antennomeres 9 and 11 modified (Figs 49–57, 59–61)............................................. 2</p> <p>2 Antennomere 11 markedly elongate, ratio (length/width) greater than 1.85 (Figs 49C, 60C; Yin &amp; Shen 2020: fig. 3B)..... 3</p> <p>- Antennomere 11 weakly to moderately elongate, ratio (length/width) less than 1.80................................. 5</p> <p>3 Body length less than 2.7 mm (2.64 mm); ratio (length/width) of antennomere 11 less than 1.9 (1.89) (Fig. 60C); aedeagus expanded apically and with broadly emarginate apical margin (Fig. 60F).......................... S. virupaksa sp. nov.</p> <p>- Body length 2.8 mm or more; ratio (length/width) of antennomere 11 greater than 2.0; aedeagus apically as broad as basal half................................................................................................ 4</p> <p>4 Setae of head relatively much longer and sparser (Fig. 49A); pronotum more elongate, ratio (length/width) 80/71 (Fig. 49A, B); distributed in Cona..................................................................... S. aequalis sp. nov.</p> <p>- Setae of head relatively much shorter and denser (Yin &amp; Shen 2020: fig. 3A); pronotum less elongate, ratio (length/width) 80/78 (Yin &amp; Shen 2020: fig. 3A); distributed in Zayü.......................................... S. chayuensis Yin &amp; Shen</p> <p>5 Antennomere 11 with two basal protuberances (Yin &amp; Shen 2020: fig. 6B, D).................. S. linzhiensis Yin &amp; Shen</p> <p>- Antennomere 11 with single basal protuberance............................................................. 6</p> <p>6 Hyaline apophysis located on apical 1/2 of antennomere 9 (Figs 55D, 56D, 61D; Yin &amp; Shen 2020: figs 10C, 13C)........ 7</p> <p>- Hyaline apophysis located at middle or on basal 1/2 of antennomere 9 (e.g., D in Figs 50–54, 57, 59).................. 11</p> <p>7 Body length less than 2.0 mm (1.94–1.95 mm); antennomere 10 of unique form (Fig. 55D).......... S. pengzhongi sp. nov.</p> <p>- Body length greater than 2.0 mm; antennomere 10 not formed as above.......................................... 8</p> <p>8 Body length greater than 2.5 mm (2.61–2.72 mm); antennomere 9 strongly expanded mesally (Yin &amp; Shen 2020: fig. 13C); basal protuberance of antennomere 11 conspicuously extended posteriorly to reach antennomere 10 (Yin &amp; Shen 2020: fig. 13B)............................................................................ S. transversus Yin &amp; Shen</p> <p>- Body length less than 2.3 mm; antennomere 9 moderately expanded mesally or obliquely expanded at anteromesal portion; basal protuberance of antennomere 11 not short and not reach antennomere 10… 9</p> <p>9 Anterior margin of antennomere 9 distinctly oblique (Fig. 56D); antennomere 11 with relatively shorter basal protuberance, apical margin subtruncate (Fig. 56C)....................................................... S. proclivis sp. nov.</p> <p>- Anterior margin of antennomere 9 even; antennomere 11 with relatively longer basal protuberance and angulate apical margin.................................................................................................. 10</p> <p>10 Antennomere 11 relatively longer, ratio (length/width) 1.72 (Fig. 61C); hyaline apophysis of antennomere 9 more anteriorly positioned (Fig. 61D)................................................................. S. yigongensis sp. nov.</p> <p>- Antennomere 11 relatively shorter, ratio (length/width) 1.6 (Yin &amp; Shen 2020: fig. 10B); hyaline apophysis of antennomere 9 more posteriorly positioned (Yin &amp; Shen 2020: fig. 10C)................................. S. panzhaohuii Yin &amp; Shen</p> <p>11 Body length greater than 3.0 mm........................................................................ 12</p> <p>- Body length less than 3.0 mm.......................................................................... 14</p> <p>12 Antennomere 9 strongly inclined at mesal margin (Yin et al. 2012a: fig. 7F); aedeagus (Yin et al. 2012a: fig. 8F)............................................................................................... S. magnus Yin &amp; Li</p> <p>- Antennomere 9 oblique, but not as inclined as above; aedeagus of a different form................................. 13</p> <p>13 Body length approximately 3.5 mm; tempus relatively longer, length of eye/tempus 0.78 (Fig. 51B); aedeagus at middle weakly narrowed (Fig. 51F)................................................................. S. caichenyangi sp. nov.</p> <p>- Body length approximately 3.1 mm; tempus relatively shorter, length of eye/tempus 0.81 (Fig. 53B); aedeagus at middle strongly narrowed (Fig. 53F)................................................................ S. duojii sp. nov.</p> <p>14 Body length 2.1 mm or slightly less (Fig. 54A).............................................. S. laticornis sp. nov.</p> <p>- Body length greater than 2.3 mm........................................................................ 15</p> <p>15 Antennomere 9 relatively more elongate (Fig. 52D), ratio (length/width) 1.18–1.23.................... S. dawai sp. nov.</p> <p>- Antennomere 9 relatively less elongate, ratio (length/width) less than 1.05....................................... 16</p> <p>16 Hyaline apophysis located at approximately middle of antennomere 9 (Yin &amp; Shen 2020: fig. 16C).................................................................................................... S. xizangensis Yin &amp; Shen</p> <p>- Hyaline apophysis located on basal 1/2 of antennomere 9..................................................... 17</p> <p>17 Aedeagus broadened apically (Fig. 57F)............................................... S. pseudograndis sp. nov.</p> <p>- Apical part of aedeagus as broad as basal half.............................................................. 18</p> <p>18 Hyaline apophysis of antennomere 9 relatively longer (Fig. 59D); basal protuberance of antennomere 11 relatively short (Fig. 59C).............................................................................. S. sympatricus sp. nov.</p> <p>- Hyaline apophysis of antennomere 9 relatively shorter; basal protuberance of antennomere 11 relatively long........... 19</p> <p>19 Body smaller in size (length 2.7 mm or slightly more), antenna shorter (1.39–1.43 mm), antennomeres 9 slightly more elongate (Fig. 50D) and antennomere 11 stouter (Fig. 50C), apical margin of aedeagus not expanded on left side (Fig. 50F)................................................................................................ S. asura sp. nov.</p> <p>- Body larger in size (length 2.9 mm or slightly more), antenna longer (1.51 mm), antennomeres 9 slightly stouter (Yin &amp; Shen 2020: fig. 1C) and antennomere 11 more elongate (Yin &amp; Shen 2020: fig. 1B), apical margin of aedeagus expanded on left side (Yin &amp; Shen 2020: fig. 1E)............................................................. S. alpicola Yin &amp; Shen</p></div> 	http://treatment.plazi.org/id/C822F213FFA42406CB88A76A70FB271F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFBA2404CB88A0DE76B924DA.text	C822F213FFBA2404CB88A0DE76B924DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes aequalis Yin 2022	<div><p>Sathytes aequalis sp. nov.</p> <p>(Figs 49, 83A, 95A, B)</p> <p>Chinese common name: ẈƟḛà甲</p> <p>Type material (16 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟 Þ玛乡 ’ (SNUC). PARATYPE: CHINA: 2 ♂♂, 13 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.9 mm; ratio (length/width) of head 1.12, pronotum 1.05, elytra 0.82–0.86, length of eye/tempus 0.82. Antennomere 9 as long as wide (length/width excluding apophysis 1.03), with hyaline apophysis at middle of strongly oblique mesal margin, antennomere 11 elongate (length/width 2.07), with distinct protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad throughout length, with slightly emarginate apical margin. Female. Body length approximately 3.0 mm; antenna lacking modifications, genitalia as in Fig. 49G.</p> <p>Description. Male. Body (Fig. 49A) length 2.87–2.95 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 49B) sub-rectangular at base, slightly longer than wide, length 0.63–0.64 mm, width across eyes 0.56–0.57 mm, length/width 1.12; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, shallowly impressed medially; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking distinct median carina or sulcus. Compound eyes prominent, composed of approximately 19 large ommatidia, length of eye/tempus 0.82. Antenna elongate, length 1.49–1.54 mm, distinct club (Fig. 49C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 slightly shorter than and as broad as 7, 9 (Fig. 49D) strongly enlarged, approximately as long as wide, length/width excluding apophysis 1.03, mesal margin expanded at middle, anterior half strongly oblique, hyaline apophysis located at approximately middle of mesal margin, 10 transverse, length/width 0.71, 11 largest, approximately as long as 9 and 10 combined, length/width 2.07, with distinct protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 49B) approximately as long as wide, length 0.63–0.68 mm, width 0.60–0.63 mm, length/width 1.05, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.92–0.98 mm, width 1.12–1.14 mm, length/width 0.82–0.86; each elytron with four large, asetose basal foveae; discal stria shallow and short, present only near base; humerus weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, large lateral mesoventral foveae forked internally, mesoventral process short. Metaventrite moderately prominent admesally, weakly inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, lacking median pit anterior to posterior margin, posterior margin broadly emarginate at middle, with thin marginal carina.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.80–0.85 mm, width 0.99–1.00 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal and marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 49E) moderately sclerotized, elongate, apex lacking setae.</p> <p>Aedeagus (Fig. 49F) 0.32 mm long, slightly asymmetric, plate-like, with large foramen, median lobe broadened throughout entire length, with apex approximately as wide as base, apical margin slightly emarginate.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus faintly prominent; metaventrite with round median pit shortly anterior to posterior margin. Measurements (as for male): body length 2.95–3.13 mm; length/width of head 0.63/ 0.56–0.59 mm, pronotum 0.67–0.70/ 0.62–0.68 mm, elytra 0.85–0.94/ 1.11–1.16 mm; abdomen 0.91– 1.0/ 1.02–1.06 mm; length of antenna 1.34–1.37 mm; maximum width of genitalia (Fig. 49G) 0.31 mm.</p> <p>Comparative notes. The male of this species resembles that of S. chayuensis present in Zayü County (察ø 县) in the similar body size, relatively long antennomere 11, and similar form and structure of antennomere 9. They can be separated by the following subtle differences: 1) the more convergent postocular margins and less rounded posterolateral angles, 2) the setae of the head are much longer and less dense, 3) a relatively more elongate (less rounded) pronotum (relative length/width: 80/71 vs. 80/78), and 4) a different fine structure of the hyaline apophyses of antennomeres 9 and the different form of the aedeagus of S. aequalis (left portion of apical margin of the median lobe distinctly convex in S. chayuensis). Note these differences can be reliably determined only through direct comparisons between the specimens of the two species, not the figures. Sathytes aequalis is also similar to S. virupaksa in the form of male antennomeres 9, but can be readily separated by the large body size, the longer postocular margins in relation to eye length, the larger basal protuberances of male antennomeres 11, a much more elongate male sternite 7 (IX), and a non-dilated apex of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 83A, 95A, B).</p> <p>Etymology. The new species name ‘ aequâlis (similar, resembling in size or form)’ is a Latin adjective suggested by the morphological similarity of the new species to S. chayuensis.</p> </div>	http://treatment.plazi.org/id/C822F213FFBA2404CB88A0DE76B924DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB82404CB88A39A74982212.text	C822F213FFB82404CB88A39A74982212.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes alpicola Yin & Shen 2020	<div><p>Sathytes alpicola Yin &amp; Shen, 2020</p> <p>(Figs 83A, 100A)</p> <p>Chinese common name: Ḃëḛà甲</p> <p>Sathytes alpicola Yin &amp; Shen, 2020: 39. Type locality: Xizang, Motuo County, near 80 K; 29°41’09”N, 95°30’10”E; alt. 2330 m.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 83A, 100A).</p> <p>Remarks. Sathytes alpicola was described based on one male and three female specimens collected at 80K, Mêdog, which has been the only available material so far. This species is most similar to S. asura, and can be separated only by subtle differences.</p> </div>	http://treatment.plazi.org/id/C822F213FFB82404CB88A39A74982212	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB82402CB88A5D3719E229A.text	C822F213FFB82402CB88A5D3719E229A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes asura Yin 2022	<div><p>Sathytes asura sp. nov.</p> <p>(Figs 50, 83A, 97A, B, D)</p> <p>Chinese common name: Nj修Ɓḛà甲</p> <p>Type material (18 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200–2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’ (SNUC). PARATYPES: CHINA: 1 ♂, 3 ♀♀, same data as that of holotype; 5 ♂♂, 8 ♀♀, ‘ China: Xizang, Nyingchi City, Tongmai Town, pass nr. 102 <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.13528&amp;materialsCitation.latitude=30.070835" title="Search Plazi for locations around (long 95.13528/lat 30.070835)">Tunnel</a>, 30°04’15”N, 95°08’07”E, 2150-2400 m, 12.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫȃ 9 102 Ṁdz ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.70–2.8 mm; ratio (length/width) of head 1.11–1.18, pronotum 1.00–1.07, elytra 0.78, length of eye/tempus 0.78. Antennomere 9 approximately as long as wide (length/width excluding apophysis 1.04), with hyaline apophysis at basal 1/3 of mesal margin, antennomere 11 elongate (length/ width 1.70), with large protuberance at base, apex of protuberance with thin tuft of curved setae. Aedeagus broad throughout length, apical margin slightly emarginate. Female. Body length approximately 2.7–2.9 mm; antenna lacking modifications, genitalia as in Fig. 50G.</p> <p>Description. Male. Body (Fig. 50A) length 2.70–2.76 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 50B) sub-rectangular at base, slightly longer than wide, length 0.61–0.63 mm, width across eyes 0.55 mm, length/width 1.11–1.18; vertex weakly raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), median carina shortly present near foveae. Compound eyes prominent, composed of approximately 19 large ommatidia, length of eye/tempus 0.78. Antenna elongate, length 1.39–1.43 mm, distinct club (Fig. 50C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 50D) enlarged, approximately as long as wide, length/width excluding apophysis 1.04, mesal margin moderately oblique, hyaline apophysis located approximately basal 1/3 of mesal margin, 10 transverse, length/width 0.68, 11 largest, as long as 9 and 10 combined, length/width 1.70, with large protuberance at base, apex of protuberance with curved tuft of thin golden setae.</p> <p>Pronotum (Fig. 50B) approximately as long as wide, length 0.62 mm, width 0.58–0.62 mm, length/width 1.00– 1.07, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.82–0.83 mm, width 1.05–1.06 mm, length/width 0.78; each elytron with four large, asetose basal foveae; lacking discal stria; humerus weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused medially, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite slightly raised admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with small, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate, slightly convex at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.80–0.81 mm, width 0.91–0.94 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites depressed along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 50E) moderately sclerotized, elongate, apex with three long setae.</p> <p>Aedeagus (Fig. 50F) 0.30 mm long, slightly asymmetric, plate-like; median lobe with large foramen, slightly constricted at apical 2/5 and then slightly broadened, apical margin slightly emarginate at middle.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus angulate. Measurements (as for male): body length 2.72–2.87 mm; length/width of head 0.60–0.61/ 0.55–0.56 mm, pronotum 0.60–0.63/ 0.60–0.65 mm, elytra 0.71– 0.87/ 0.99–1.07 mm; abdomen 0.88–0.92/ 0.93–0.99 mm; length of antenna 1.27–1.33 mm; maximum width of genitalia (Fig. 50G) 0.27 mm.</p> <p>Comparative notes. This species morphologically resembles S. alpicola found in Mêdog, in the similar form of antennomere 9 and location of the hypophysis on it (at limit of approximately basal 1/3). These two species may be separated only by subtle differences: Sathytes asura is slightly smaller in all body parts and has shorter antennae, antennomeres 9 are relatively slightly more elongate and antennomeres 11 stouter, and the apical margin of the aedeagus is not expanded on left side.</p> <p>Distribution. Bomê County, Tibet, SW China (Figs 83A, 97A, B, D).</p> <p>Etymology. The new species is named after Asuras, demigods of the desire realm of Buddhism.</p></div> 	http://treatment.plazi.org/id/C822F213FFB82402CB88A5D3719E229A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFBE2400CB88A45A77232326.text	C822F213FFBE2400CB88A45A77232326.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes caichenyangi Yin 2022	<div><p>Sathytes caichenyangi sp. nov.</p> <p>(Figs 51, 83B, 95A, B)</p> <p>Chinese common name: Ḛ氏ḛà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC). PARATYPE: CHINA: 1 ♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 3.5 mm; ratio (length/width) of head 1.13, pronotum 1.07, elytra 0.84, length of eye/tempus 0.78. Antennomere 9 as long as wide (length/width excluding apophysis 1.0), with hyaline apophysis posterior to middle of mesal margin, antennomere 11 elongate (length/width 1.70), with moderately long protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad throughout length, with rounded apical margin. Female. Body length approximately 3.6 mm; antenna lacking modifications, genitalia as in Fig. 51G.</p> <p>Description. Male. Body (Fig. 51A) length 3.48 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 51B) sub-rectangular at base, slightly longer than wide, length 0.71 mm, width across eyes 0.63 mm, length/width 1.13; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking distinct median carina or sulcus. Compound eyes prominent, composed of approximately 28 large ommatidia, length of eye/tempus 0.78. Antenna elongate, length 1.71 mm, distinct club (Fig. 51C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 slightly narrower and shorter than 7, 9 (Fig. 51D) strongly enlarged, as long as wide, length/width excluding apophysis 1.0, mesal margin moderately expanded at middle, hyaline apophysis located slightly posterior to middle of mesal margin, 10 transverse, length/ width 0.77, 11 largest, approximately as long as 9 and 10 combined, length/width 1.70, with small protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 51B) approximately as long as wide, length 0.76 mm, width 0.71 mm, length/width 1.07, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 1.07 mm, width 1.27 mm, length/width 0.84; each elytron with four large, asetose basal foveae; discal stria shallow and short; humerus weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, large lateral mesoventral foveae forked internally, mesoventral process short. Metaventrite moderately prominent admesally, weakly inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate at middle, lacking split, with thin marginal carina.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.95 mm, width 1.15 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal and marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 51E) moderately sclerotized, elongate, apex with few long setae.</p> <p>Aedeagus (Fig. 51F) 0.36 mm long, slightly asymmetric and bent leftwards, plate-like; median lobe with large foramen, broadened throughout entire length, with apex approximately as wide as base, apical margin evenly rounded.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus faintly prominent. Measurements (as for male): body length 3.62 mm; length/width of head 0.77/ 0.63 mm, pronotum 0.81/ 0.74 mm, elytra 1.04/ 1.28 mm; abdomen 1.10/ 1.23 mm; length of antenna 1.74 mm; maximum width of genitalia (Fig. 51G) 0.32 mm.</p> <p>Comparative notes. This species is allied with S. grandis Löbl distributed in central Nepal and S. pseudograndis by sharing similar forms of male antennomeres 9 and 11, and a large body size. The male of Sathytes caichenyangi differs from S. grandis in the much larger body size (3.5 mm vs. 3.0 mm), the relatively much smaller antennomeres 9 in relation to the head size, and a much longer basal protuberance of antennomeres 11; from S. pseudograndis by the much larger size (3.5 mm vs. 2.80–3.0 mm), much longer postocular margins in relation to eye length (0.78 vs 0.95), the more posterior position of the hyaline apophysis of antennomeres 9, the much longer basal protuberance of antennomeres 11, and the narrower apex of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 83B, 95A, B).</p> <p>Etymology. This species is named after Chenyang Cai (ḚṚŖ) for his various support and help during my work on the Tibetan pselaphine fauna.</p></div> 	http://treatment.plazi.org/id/C822F213FFBE2400CB88A45A77232326	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFBC2400CB88A48676D621D4.text	C822F213FFBC2400CB88A48676D621D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes chayuensis Yin & Shen 2020	<div><p>Sathytes chayuensis Yin &amp; Shen, 2020</p> <p>(Figs 83B, 100D, E)</p> <p>Chinese common name: 察øḛà甲</p> <p>Sathytes chayuensis Yin &amp; Shen, 2020: 42. Type locality: China, Xizang, Chayu (= Zayü) Co., Xiachayu To., Talin Bridge; 28.53340539° N, 96.97800188° E; alt. 1541 m.</p> <p>Distribution. Zayü County, Tibet, SW China (Figs 83B, 100D, E).</p> <p>Remarks. Sathytes chayuensis was described based on three males and one female collected at Xiachayu, Zayü. This species morphologically resembles S. aequalis and S. virupaksa described in this paper in the distinctly elongate antennomeres 11, similar forms of antennomeres 9 and placements of the apophyses on them. It differs from both species by the shorter and denser setae of the head, and the apical margin of the aedeagus is convex on the left side (also see ‘Comparative notes’ under S. aequalis and S. virupaksa).</p> </div>	http://treatment.plazi.org/id/C822F213FFBC2400CB88A48676D621D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB3240DCB88A1DB7728271E.text	C822F213FFB3240DCB88A1DB7728271E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes dawai Yin 2022	<div><p>Sathytes dawai sp. nov.</p> <p>(Figs 52, 83B, 90, 93F, G)</p> <p>Chinese common name: ḭǟḛà甲</p> <p>Type material (17 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, Zhêntang Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (SNUC). PARATYPES: CHINA: 11 ♂♂, 4 ♀♀, same data as that of holotype; 1 ♂, ‘ China: Xizang, Gyirong County, Gyirong Valley, slope nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.32806&amp;materialsCitation.latitude=28.376945" title="Search Plazi for locations around (long 85.32806/lat 28.376945)">Jifu Vill.</a>, 28°22’37”N, 85°19’41”E, 2400-2700 m, 22.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW吉ė村ṉữ山坡 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length greater than 2.4 mm; ratio (length/width) of head 1.10–1.15, pronotum 0.96– 1.02, elytra 0.81–0.83, length of eye/tempus 0.95–1.13. Antennomere 9 slightly elongate (length/width excluding apophysis 1.18–1.23), with hyaline apophysis at middle of mesal margin, antennomere 11 elongate (length/ width 1.74–1.77), with single protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad throughout length, with apex as broad as base. Female. Body length over 2.3 mm; antenna lacking modifications, genitalia as in Fig. 52G.</p> <p>Description. Male. Body (Fig. 52A) length 2.42–2.51 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 52B) sub-rectangular at base, slightly longer than wide, length 0.52–0.53 mm, width across eyes 0.46–0.48 mm, length/width 1.10–1.13; vertex moderately raised, with distinct asetose foveae (dorsal tentorial pits); frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina present; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking median carina or sulcus. Compound eyes prominent, composed of approximately 20–22 large ommatidia, length of eye/tempus 0.95–1.13. Antenna elongate, length 1.25–1.32 mm, distinct club (Fig. 52C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 52D) strongly enlarged, slightly elongate, length/width excluding apophysis 1.18–1.23, mesal margin moderately expanded at middle, hyaline apophysis located anterior to middle of mesal margin, 10 transverse, length/width 0.75–0.76, 11 largest, as long as 9 and 10 combined, length/width 1.74–1.77, with truncate protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 52B) approximately as long as wide, length 0.53–0.56 mm, width 0.55 mm, length/width 0.96– 1.02, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.73 mm, width 0.88–0.90 mm, length/width 0.81–0.83; each elytron with four large, asetose basal foveae; lacking discal stria; humerus roundly and weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae contiguous, originating from shared opening, large lateral mesoventral foveae forked internally, anteromesal margin of coxal cavity with small tubercle, mesoventral process short, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin with indistinct, short split at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.69–0.78 mm, width 0.77–0.78 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, discal and marginal carina thin and short, indistinct; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites depressed along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 52E) moderately sclerotized, elongate, apex asetose.</p> <p>Aedeagus (Fig. 52F) 0.31–0.33 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broadened throughout entire length, with apex as broad as base.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus weakly prominent. Measurements (as for male): body length 2.32–2.46 mm; length/width of head 0.50–0.52/ 0.45–0.47 mm, pronotum 0.51–0.55/ 0.53–0.55 mm, elytra 0.65/ 0.84 mm; abdomen 0.75–0.79/ 0.79–0.81 mm; length of antenna 1.08–1.12 mm; maximum width of genitalia (Fig. 52G) 0.27 mm.</p> <p>Comparative notes. Sathytes dawai is morphological similar to S. montanus Löbl and S. grandis Löbl that are distributed in northern India and a number of Chinese species including S. alpicola Yin &amp; Shen, S. panzhaohuii Yin &amp; Shen, and S. xizangensis Yin &amp; Shen from Tibet that share a hyaline apophysis on the more or less expanded mesal margin of antennomere 9, and a single basal projection of antennomeres 11. The new species, however, can be separated from all these congeners by a combination of characters listed in the ‘Diagnosis’ section, and especially by the elongate antennomere 9 with its more apical placement of the hyaline apophysis on it.</p> <p>Distribution. Dinggyê and Gyirong County, Tibet, SW China (Figs 83B, 90, 93F, G).</p> <p>Etymology. The new species is named after Wa Da (ḭǟ), who provided logistical and other kinds of support during our field investigations in Tibet.</p></div> 	http://treatment.plazi.org/id/C822F213FFB3240DCB88A1DB7728271E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB1240CCB88A0DE774B21D7.text	C822F213FFB1240CCB88A0DE774B21D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes duojii Yin 2022	<div><p>Sathytes duojii sp. nov.</p> <p>(Figs 53, 83C, 97A, B)</p> <p>Chinese common name: Ʒ吉ḛà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200–2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’.</p> <p>Diagnosis. Male. Body length approximately 3.1 mm; ratio (length/width) of head 1.08, pronotum 1.03, elytra 0.82, length of eye/tempus 0.81. Antennomere 9 strongly enlarged, broad (length/width excluding apophysis 0.94), with hyaline apophysis slightly above middle of mesal margin, antennomere 11 elongate (length/width 1.53), with broad, moderately long protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad at base and apex, constricted at middle.</p> <p>Description. Male. Body (Fig. 53A) length 3.09 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 53B) sub-rectangular at base, slightly longer than wide, length 0.63 mm, width across eyes 0.62 mm, length/width 1.02; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, weakly impressed; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina distinct; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), with faint median carina. Compound eyes prominent, composed of approximately 18 large ommatidia, length of eye/tempus 0.81. Antenna elongate, length 1.54 mm, distinct club (Fig. 53C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 53D) strongly enlarged, slightly wider than long, length/width excluding apophysis 0.94, mesal margin oblique, hyaline apophysis located slightly above middle of mesal margin, 10 transverse, length/width 0.61, 11 largest, approximately as long as 9 and 10 combined, length/width 1.53, with broad protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 53B) approximately as long as wide, length 0.71 mm, width 0.69 mm, length/width 1.03, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.98 mm, width 1.19 mm, length/width 0.82; each elytron with four large, asetose basal foveae; discal stria represented by shallow and short impression near base; humerus weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, large lateral mesoventral foveae forked internally, mesoventral process short. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with small, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate at middle, lacking split, with thin marginal carina.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.88 mm, width 1.06 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal and marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 53E) moderately sclerotized, elongate, apical margin with three long setae.</p> <p>Aedeagus (Fig. 53F) 0.30 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broad at apex and base, moderately constricted at middle, apical margin slightly emarginate at middle.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species shares with S. caichenyangi the similar forms of male antennomeres 9, but can be readily separated by the smaller body size (3.1 mm vs. 3.5 mm), a less elongate head (ratio: 1.02 vs. 1.13), the shorter basal protuberances of antennomeres 11, and a more strongly constricted middle part of the generally stouter aedeagus.</p> <p>Distribution. Bomê County, Tibet, SW China (Figs 83C, 97A, B).</p> <p>Etymology. The new species is dedicated to Duoji Ciren, the driver and guide of our 2021 field trip in Tibet, who provided various kinds of help.</p></div> 	http://treatment.plazi.org/id/C822F213FFB1240CCB88A0DE774B21D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB7240ACB88A1DB7162210E.text	C822F213FFB7240ACB88A1DB7162210E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes laticornis Yin 2022	<div><p>Sathytes laticornis sp. nov.</p> <p>(Figs 54, 83C, 95A, B)</p> <p>Chinese common name: ÝƟḛà甲</p> <p>Type material (6 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC). PARATYPE: CHINA: 5 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.1 mm; ratio (length/width) of head 1.07, pronotum 1.02, elytra 0.77, length of eye/tempus 0.81. Antennomere 9 moderately transverse (length/width excluding apophysis 1.0), with hyaline apophysis at approximately middle of mesal margin, antennomere 11 elongate (length/width 1.71), with short protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broadly rounded at base, broadening through length, apical margin moderately emarginate. Female. Body length 2.1–2.2 mm; antenna lacking modifications, genitalia as in Fig. 54G.</p> <p>Description. Male. Body (Fig. 54A) length 2.08 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short, dense pubescence.</p> <p>Head (Fig. 54B) sub-rectangular at base, slightly longer than wide, length 0.48 mm, width across eyes 0.45 mm, length/width 1.07; vertex moderately raised, with small asetose foveae (dorsal tentorial pits); frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking median carina. Compound eyes moderately prominent, composed of approximately 18 large ommatidia, length of eye/tempus 0.81. Antenna elongate, length 1.04 mm, distinct club (Fig. 54C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 54D) strongly enlarged, as long as wide, length/width excluding 1.0, mesal margin strongly expanded at middle, moderately inclined apically and basally, hyaline apophysis located at approximately middle of mesal margin, 10 transverse, length/width 0.75, 11 largest, as long as 9 and 10 combined, length/width 1.71, with short protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 54B) approximately as long as wide, length 0.50 mm, width 0.49 mm, length/width 1.02, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae moderately small. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.67 mm, width 0.87 mm, length/width 0.77; each elytron with four small, asetose basal foveae; lacking discal stria; humerus weakly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, originating from shared opening, large lateral mesoventral foveae shallowly forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two broadly separated lateral metaventral foveae, with shallow, round median pit shortly anterior to posterior margin, posterior margin narrowly and shortly emarginate at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.57 mm, width 0.75 mm. Tergite 1 (IV) 2.5 times as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal carinae, with indistinct, thin and short marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 54E) moderately sclerotized, elongate, apex with three long setae.</p> <p>Aedeagus (Fig. 54F) 0.22 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broadest at apical 1/3, apical margin emarginate at middle and moderately expanded at right side.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 12 ommatidia; humerus weakly prominent. Measurements (as for male): body length 2.11–2.20 mm; length/width of head 0.48–0.49/ 0.45–0.46 mm, pronotum 0.49–0.51/ 0.46–0.49 mm, elytra 0.60–0.64/ 0.84–0.85 mm; abdomen 0.64–0.67/ 0.81–0.82 mm; length of antenna 1.01–1.02 mm; maximum width of genitalia (Fig. 54G) 0.22 mm.</p> <p>Comparative notes. The male of Sathytes laticornis differs from those of all congeners by the relatively broad antennomeres 9 (mesal margin less oblique), combined with the small body size (2.08 mm), the short basal protuberances of antennomeres 11, and the unique form of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 83C, 95A, B).</p> <p>Etymology. The specific epithet is combined from the Latin adjective ‘ lâtus, - a, - um (wide, broad)’ and ‘ cornis (horned)’, referring to the broad shape of male antennomeres 9 of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FFB7240ACB88A1DB7162210E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB52409CB88A1DB70672450.text	C822F213FFB52409CB88A1DB70672450.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes linzhiensis Yin & Li 2020	<div><p>Sathytes linzhiensis Yin &amp; Li, 2020</p> <p>(Figs 83C, 99A, F)</p> <p>Chinese common name: ffżḛà甲</p> <p>Sathytes linzhiensis Yin &amp; Shen, 2020: 47. Type locality: China, Xizang, Motuo County, path from Bari village to <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.35755&amp;materialsCitation.latitude=29.312449" title="Search Plazi for locations around (long 95.35755/lat 29.312449)">Renqingbeng Temple</a>, 29°18’44.82”N, 95°21’27.18”E, alt. 1833 m.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 83C, 99A, F).</p> <p>Remarks. Sathytes linzhiensis was described from four males found at Bari Village and Beibeng Township of Mêdog. This species is distinct in possessing a small body size (1.7–1.9 mm), the markedly elongate and straight hyaline apophyses on antennomeres 9, and there are two small basal projections of antennomeres 11.</p> </div>	http://treatment.plazi.org/id/C822F213FFB52409CB88A1DB70672450	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB52409CB88A32C748D227A.text	C822F213FFB52409CB88A32C748D227A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes magnus Yin & Li 2012	<div><p>Sathytes magnus Yin &amp; Li, 2012</p> <p>(Figs 84A, 98C, D)</p> <p>Chinese common name: 巨型ḛà甲</p> <p>Sathytes magnus Yin &amp; Li, 2012a: 845. Type locality: Xizang A. R., Motuo County, Hanmi, alt. 2200 m.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 84A, 98C, D).</p> <p>Remarks. Sathytes magnus was described based on one male and one female collected at Hanmi of Mêdog County (Yin &amp; Li 2012). This species morphologically resembles a number of congeners whose males share a single basal protuberance of antennomere 11, and markedly oblique mesal margins of antennomeres 9. It differs in the large body size (3.36 mm), the relatively broad antennomeres 11 (Yin &amp; Li 2012: fig. 4F), the unique forms of antennomeres 9 and their apophyses (Yin &amp; Li 2012: fig. 7F), an apically expanded aedeagus (Yin &amp; Li 2012: fig. 8F), as well as its distribution.</p> </div>	http://treatment.plazi.org/id/C822F213FFB52409CB88A32C748D227A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB52409CB88A53B77E12021.text	C822F213FFB52409CB88A53B77E12021.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes panzhaohuii Yin & Shen 2020	<div><p>Sathytes panzhaohuii Yin &amp; Shen, 2020</p> <p>(Figs 84A, 97A–C, 100D, E)</p> <p>Chinese common name: Ğ氏ḛà甲</p> <p>Sathytes panzhaohuii Yin &amp; Shen, 2020: 53; fig. 10. Type locality: China, Xizang, Nyingchi, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, alt. 2190 m.</p> <p>Additional material examined (26 exx.). 2 ♂♂, 3 ♀♀, ‘ China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96611&amp;materialsCitation.latitude=30.009722" title="Search Plazi for locations around (long 94.96611/lat 30.009722)">Pailong Township</a>, 30°0’35”N, 94°57’58”E, 2180 m, 10.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., Ʊ Ẫh 治区ffż市ḦË乡 ’; 14 ♂♂, 7 ♀♀, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200-2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’ (SNUC).</p> <p>Distribution. Bomê and Zayü (= Chayu) County, Tibet, SW China (Figs 84A, 97A–C, 100D, E).</p> <p>Remarks. Sathytes panzhaohuii was described based on material from Pailong, Yigong, and surprisingly, Xiazayü (T察ø) of Zayü County. The new material confirms that it is a common species in Nyingchi that is sympatric with S. yigongensis in Yigong Township.</p> </div>	http://treatment.plazi.org/id/C822F213FFB52409CB88A53B77E12021	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FFB524F7CB88A79C71EF2056.text	C822F213FFB524F7CB88A79C71EF2056.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes pengzhongi Yin 2022	<div><p>Sathytes pengzhongi sp. nov.</p> <p>(Figs 55, 84A, 93F, G)</p> <p>Chinese common name: 彭氏ḛà甲</p> <p>Type material (11 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, Zhêntang Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 8 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.0 mm; ratio (length/width) of head 1.05–1.08, pronotum 1.09–1.11, elytra 0.75–0.78, length of eye/tempus 0.89. Antennomere 9 moderately transverse (length/width excluding apophysis 0.87), with hyaline apophysis at anterior 1/4 of mesal margin, antennomere 11 elongate (length/width 1.59), with single protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broadly rounded at base, narrowing apically. Female. Body length approximately 2.0 mm; antenna lacking modifications, genitalia as in Fig. 55G.</p> <p>Description. Male. Body (Fig. 55A) length 1.94–1.95 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short, dense pubescence.</p> <p>Head (Fig. 55B) sub-rectangular at base, slightly longer than wide, length 0.46–0.49 mm, width across eyes 0.44–0.45 mm, length/width 1.05–1.08; vertex moderately raised, with small asetose foveae (dorsal tentorial pits); frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), short median carina present posterior mouthpart. Compound eyes moderately prominent, composed of approximately 18 large ommatidia, length of eye/tempus 0.89. Antenna elongate, length 1.06–1.07 mm, distinct club (Fig. 55C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 55D) strongly enlarged, moderately transverse, length/width excluding apophysis 0.87, mesal margin strongly expanded, hyaline apophysis located at apical fourth of mesal margin, 10 transverse, length/width 0.67, 11 largest, slightly longer than 9 and 10 combined, length/width 1.59, with small protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 55B) slightly longer than wide, length 0.49–0.51 mm, width 0.45–0.46 mm, length/width 1.09– 1.11, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae small. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.59–0.62 mm, width 0.79 mm, length/width 0.75–0.78; each elytron with four moderately large, asetose basal foveae; lacking discal stria; humerus moderately prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae contiguous, originating from shared opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two broadly separated lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin narrowly and shortly emarginate at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.51–0.53 mm, width 0.67–0.71 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal carinae, with indistinct, thin and short marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 55E) moderately sclerotized, elongate, apex with two long setae.</p> <p>Aedeagus (Fig. 55F) 0.21 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broad at base and narrowing from middle towards apex.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 12 ommatidia; humerus weakly prominent. Measurements (as for male): body length 1.98–2.02 mm; length/width of head 0.45–0.48/ 0.44–0.45 mm, pronotum 0.47–0.48/ 0.46–0.48 mm, elytra 0.53–0.58/ 0.76–0.83 mm; abdomen 0.63–0.65/ 0.76–0.78 mm; length of antenna 0.96–0.99 mm; maximum width of genitalia (Fig. 55G) 0.20 mm.</p> <p>Comparative notes. The male of Sathytes pengzhongi can be easily separated from that of all congeners from the Himalayan region by the unique shape of antennomeres 9 and the anteriorly placed apophysis on them, combined with the small body size of 1.9–2.0 mm.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 84A, 93F, G).</p> <p>Etymology. The new species is named after Zhong Peng, the collector of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FFB524F7CB88A79C71EF2056	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4B24F6CB88A71677ED2182.text	C822F213FF4B24F6CB88A71677ED2182.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes proclivis Yin 2022	<div><p>Sathytes proclivis sp. nov.</p> <p>(Figs 56, 84B, 95A, B)</p> <p>Chinese common name: ṆƟḛà甲</p> <p>Type material (13 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟 Þ玛乡 ’ (SNUC). PARATYPE: CHINA: 4 ♂♂, 8 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.2 mm; ratio (length/width) of head 0.92–0.98, pronotum 1.02– 1.04, elytra 0.77–0.80, length of eye/tempus 0.90. Antennomere 9 slightly wider than long (length/width excluding apophysis 0.96), with hyaline apophysis near apex of mesal margin, antennomere 11 elongate (length/width 1.72), with weak protuberance at base, apex of protuberance with tuft of slightly curved setae. Aedeagus broadly rounded at base, slightly narrowed at middle and then broadening apically. Female. Body length approximately 2.2–2.3 mm; antenna lacking modifications, genitalia as in Fig. 56G.</p> <p>Description. Male. Body (Fig. 56A) length 2.17–2.19 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short, dense pubescence.</p> <p>Head (Fig. 56B) sub-rectangular at base, approximately as long as wide, length 0.47 mm, width across eyes 0.48–0.51 mm, length/width 0.92–0.98; vertex slightly raised, with small asetose foveae (dorsal tentorial pits); frons anteriorly confluent with strongly declining clypeus, shallowly impressed medially; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), median carina present for anterior half. Compound eyes moderately prominent, composed of approximately 20 large ommatidia, length of eye/tempus 0.90. Antenna elongate, length 1.23–1.26 mm, distinct club (Fig. 56C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 shorter and slightly narrower than 7, 9 (Fig. 56D) strongly enlarged, obliquely trapezoidal, slightly wider than long, length/width excluding apophysis 0.96, anteromesal region strongly enlarged, hyaline apophysis located near apex of mesal margin, 10 transverse, length/width 0.63, 11 largest, longer than 9 and 10 combined, length/width 1.72, with weak protuberance at base, apex of protuberance with slightly curved tuft of golden setae.</p> <p>Pronotum (Fig. 56B) approximately as long as wide, length 0.51–0.52 mm, width 0.50 mm, length/width 1.02– 1.04, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae moderately small. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.69–0.71 mm, width 0.89–0.90 mm, length/width 0.77–0.80; each elytron with four small, asetose basal foveae; lacking discal stria; humerus weakly prominent, acute, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused at middle, originating from shared opening, large lateral mesoventral foveae shallowly forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two broadly separated lateral metaventral foveae, with shallow, round median pit shortly anterior to posterior margin, posterior margin narrowly and shortly emarginate at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.62–0.63 mm, width 0.77–0.78 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 56E) moderately sclerotized, elongate, apex with two long setae.</p> <p>Aedeagus (Fig. 56F) 0.23 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broadest at approximately apical 2/5, apical margin moderately convex at right side, with one short seta at left side.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus faintly prominent. Measurements (as for male): body length 2.25–2.28 mm; length/width of head 0.49/ 0.47 mm, pronotum 0.51–0.53/ 0.51 mm, elytra 0.63–0.64/ 0.86– 0.87 mm; abdomen 0.73–0.75/ 0.82–0.84 mm; length of antenna 1.06–1.08 mm; maximum width of genitalia (Fig. 56G) 0.22 mm.</p> <p>Comparative notes. The new species morphologically resembles S. longitrabis Yin &amp; Li found in Shaanxi, central China by the obliquely expanded male antennomeres 9, but clearly differs from the latter by the more elongate antennomeres 11 which have smaller basal projections, the more strongly oblique anterior margins of antennomeres 9, and the different form of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 84B, 95A, B).</p> <p>Etymology. The specific epithet ‘ prôclîvis (prone, steep)’ is a Latin adjective referring to the obliquely expanded male antennomeres 9 of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FF4B24F6CB88A71677ED2182	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4924F3CB88A71776E820BE.text	C822F213FF4924F3CB88A71776E820BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes pseudograndis Yin 2022	<div><p>Sathytes pseudograndis sp. nov.</p> <p>(Figs 57, 84B, 93F, G)</p> <p>Chinese common name: 伪ũḛà甲</p> <p>Type material (8 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, Zhêntang Town, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (SNUC). PARATYPES: CHINA: 2 ♂♂, 5 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length greater than 2.8 mm; ratio (length/width) of head 1.09–1.14, pronotum 1.03– 1.08, elytra 0.84–0.86, length of eye/tempus 0.95. Antennomere 9 approximately as long as wide (length/width excluding apophysis 1.04), with hyaline apophysis at middle of mesal margin, antennomere 11 elongate (length/width 1.56), with short protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad throughout length, slightly bent leftwards, with apical margin slightly emarginate. Female. Body length approximately 3.0 mm; antenna lacking modifications, genitalia as in Fig. 57G.</p> <p>Description. Male. Body (Fig. 57A) length 2.80–2.97 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 57B) sub-rectangular at base, slightly longer than wide, length 0.61–0.63 mm, width across eyes 0.55–0.56 mm, length/width 1.11–1.12; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking distinct median carina or sulcus. Compound eyes prominent, composed of approximately 20 large ommatidia, length of eye/tempus 0.95. Antenna elongate, length 1.39–1.46 mm, distinct club (Fig. 57C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 slightly narrower and shorter than 7, 9 (Fig. 57D) strongly enlarged, approximately as long as wide, length/width excluding apophysis 1.04, mesal margin moderately expanded at middle, hyaline apophysis located at middle of mesal margin, 10 transverse, length/width 0.67, 11 largest, as long as 9 and 10 combined, length/width 1.56, with small protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 57B) approximately as long as wide, length 0.66–0.69 mm, width 0.64 mm, length/width 1.03– 1.08, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.89–0.90 mm, width 1.04–1.06 mm, length/width 0.85; each elytron with four large, asetose basal foveae; lacking discal stria; humerus slightly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae contiguous, originating from shared oval opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite moderately prominent admesally, weakly inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate at middle, lacking split.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.80–0.91 mm, width 0.94–0.95 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal carina, marginal carina thin and short, faint; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites depressed along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 57E) moderately sclerotized, elongate, apex asetose.</p> <p>Aedeagus (Fig. 57F) 0.37 mm long, slightly asymmetric and bent leftwards, plate-like; median lobe with large foramen, broadened throughout entire length, with apex slightly wider than base, apical margin slightly emarginate.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 14 ommatidia; humerus weakly prominent. Measurements (as for male): body length 2.91–3.05 mm; length/width of head 0.61–0.63/ 0.57–0.58 mm, pronotum 0.66–0.67/ 0.65 mm, elytra 0.84–0.86/ 1.07–1.08 mm; abdomen 0.93–0.95/0.99–1.00 mm; length of antenna 1.30–1.41 mm; maximum width of genitalia (Fig. 57G) 0.29 mm.</p> <p>Comparative notes. This species is closely related to S. grandis Löbl, 1979 by sharing a similar body size, the position and form of the apophyses of antennomeres 9 and basal projection of antennomeres 11, and the form of the aedeagus. It differs from S. grandis by the relatively smaller eyes (ratio of eye/tempus 0.95 vs.&gt; 1.1 in S. grandis), and relatively markedly smaller antennomeres 9 in relation to 8 and 10. Sathytes pseudograndis is also similar to S. xizangensis by the similar male antennal structures, but can be separated from it by the much larger body size (2.80–2.97 mm vs. 2.53–2.73 mm), the more strongly inclined anteromesal margins of antennomere 9, and the much smaller basal projections of antennomeres 11.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 84B, 93F, G).</p> <p>Etymology. The specific epithet is a combination of the prefix pseudo - (false, fake)’ and ‘ grandis (large, great)’, indicating the morphological similarity of this species to S. grandis.</p> </div>	http://treatment.plazi.org/id/C822F213FF4924F3CB88A71776E820BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4E24F0CB88A1DB76C326A6.text	C822F213FF4E24F0CB88A1DB76C326A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes simplex Lobl 1979	<div><p>Sathytes simplex Löbl, 1979</p> <p>(Figs 58, 84B, 97A–C)</p> <p>Chinese common name: ÊƟḛà甲</p> <p>Sathytes simplex Löbl, 1979: 298; figs 19, 35. Type locality: Inde, Darjeeling distr., Tiger Hill, 2150 m.</p> <p>Additional material examined (6 exx.). 5 ♂♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200–2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’; 1 ♂ ‘China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96611&amp;materialsCitation.latitude=30.009722" title="Search Plazi for locations around (long 94.96611/lat 30.009722)">Pailong Township</a>, 30°0’35”N, 94°57’58”E, 2180 m, 10.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg. Ʊ Ẫh 治区ffż市ḦË乡 ’ (all in SNUC).</p> <p>Diagnosis. Male. Body length 2.45–2.6 mm; ratio (length/width) of head 1.04–1.16, pronotum 1.00–1.06, elytra 0.76–0.86, length of eye/tempus 0.86–0.91. Antennae simple, antennomere 9 slightly longer than wide (length/ width 1.06–1.15), lacking hyaline apophysis on mesal margin, antennomere 11 elongate (length/width 1.76–1.95), lacking protuberance at base. Aedeagus broad, slightly constricted at middle, apical margin slightly protuberant at left side. Female. See Löbl 1979.</p> <p>Redescription (based on populations of China, measurements of original description included in parentheses). Male. Body (Fig. 58A) length 2.45–2.56 (2.55–2.60) mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 58B) sub-rectangular at base, slightly longer than wide, length 0.53–0.58 mm, width across eyes 0.48–0.50 mm, length/width 1.10–1.16 (1.04); vertex weakly raised, with small asetose foveae (dorsal tentorial pits) located slightly posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, flat; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina thin and short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking median carina or sulcus. Compound eyes prominent, composed of approximately 18 large ommatidia, length of eye/tempus 0.86–0.91. Antenna elongate, length 1.27–1.28 mm, distinct club (Fig. 58C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly broader and 8 slightly narrower than 3–7, 9 enlarged, slightly longer than wide, length/width 1.06–1.15, lacking hyaline apophysis on mesal margin, 10 transverse, length/width 0.82–0.87, 11 largest, slightly longer than 9 and 10 combined (36/33), length/width 1.76–1.95 (1.9), lacking protuberance at base.</p> <p>Pronotum (Fig. 58B) approximately as long as wide, both length and width 0.56–0.57 mm, length/width 1.00 (1.06), widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.72–0.74 mm, width 0.84–0.92 mm, length/width 0.80–0.86 (0.76); each elytron with four large, asetose basal foveae; lacking discal stria; humerus slightly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae contiguous, originating from shared oval opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with moderately large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate and shallowly incised at middle.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.64–0.72 mm, width 0.77–0.83 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites depressed along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 58D) moderately sclerotized, elongate, apex with three elongate setae.</p> <p>Aedeagus (Fig. 58E) 0.29 (0.26) mm long, slightly asymmetric, broad, plate-like; median lobe with large foramen, slightly constricted at middle, apical margin slightly protuberant at left side.</p> <p>Female. No material has been collected in China, see Löbl 1979 for description.</p> <p>Comparative notes. Sathytes simplex is the only species of the genus known to possess simple antennae in the male.</p> <p>Distribution. Darjeeling District, N India; Bomê County and Chagyib District (Nyingchi), Xizang, SW China (Figs 84B, 97A–C). By discovering the new populations in China, the known range of this species extends considerably for some 750 km to the northeast. New country record for China.</p> <p>Remarks. Even when adjusted to a same angle, the aedeagi of the populations from China appear to be less protuberant at the apical margin than is shown in Löbl (1979: fig. 35). However, this difference could be reasonably fall into the range of intraspecific variation. The measurements of the body segments show no significant difference from those of specimens from northern India. The populations of China are temporarily identified as conspecific with S. simplex. A direct comparison with the holotype in the future is desired, since in many cases this is the only means to distinguish morphologically similar species within Sathytes.</p> </div>	http://treatment.plazi.org/id/C822F213FF4E24F0CB88A1DB76C326A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4C24FECB88A00674B327F2.text	C822F213FF4C24FECB88A00674B327F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes sympatricus Yin 2022	<div><p>Sathytes sympatricus sp. nov.</p> <p>(Figs 59, 84C, 95A, B)</p> <p>Chinese common name: ḲRḛà甲</p> <p>Type material (16 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟 Þ玛乡 ’ (SNUC). PARATYPE: CHINA: 10 ♂♂, 5 ♀♀, same data as that of holotype (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.7 mm; ratio (length/width) of head 1.00–1.08, pronotum 1.05, elytra 0.83–0.84, length of eye/tempus 0.95. Antennomere 9 as long as wide (length/width excluding apophysis 1.0), with hyaline apophysis at middle of mesal margin, antennomere 11 elongate (length/width 1.64), with short protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broad, slightly constricted at middle, apical margin slightly emarginate and convex on right side. Female. Body length approximately 2.6 mm; antenna lacking modifications, genitalia as in Fig. 59G.</p> <p>Description. Male. Body (Fig. 59A) length 2.66–2.68 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 59B) sub-rectangular at base, approximately as long as wide, length 0.52–0.53 mm, width across eyes 0.53–0.56 mm, length/width 1.00–1.08; vertex moderately raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, shallowly impressed at middle; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking distinct median carina or sulcus. Compound eyes prominent, composed of approximately 17 large ommatidia, length of eye/tempus 0.95. Antenna elongate, length 1.33 mm, distinct club (Fig. 59C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each transverse, subequal in width, 8 as wide as and much shorter than 7, 9 (Fig. 59D) strongly enlarged, as long as wide, length/width excluding apophysis 1.0, mesal margin moderately expanded at middle and strongly oblique anteriorly and posteriorly, hyaline apophysis located at middle of mesal margin, 10 transverse, length/width 0.71, 11 largest, as long as 9 and 10 combined, length/width 1.64, with short, apically truncate protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 59B) slightly longer than wide, length 0.58–0.61 mm, width 0.55–0.58 mm, length/width 1.05, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; large asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.82–0.86 mm, width 0.99–1.02 mm, length/width 0.83–0.84; each elytron with four large, asetose basal foveae; discal stria shallow and short; humerus weakly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, in oval opening, large lateral mesoventral foveae forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate at middle, lacking split.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.77–0.78 mm, width 0.87–0.88 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of large mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternites flat along middle; sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 59E) moderately sclerotized, elongate, with few long setae along apical margin.</p> <p>Aedeagus (Fig. 59F) 0.29 mm long, slightly asymmetric, plate-like; median lobe with large foramen, broad at base and slightly constricted at middle, apical margin moderately emarginate at middle and convex on right side.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 14 ommatidia; humerus weakly prominent. Measurements (as for male): body length 2.56–2.59 mm; length/width of head 0.56/ 0.52–0.56 mm, pronotum 0.58–0.60/ 0.56–0.59 mm, elytra 0.76–0.77/ 0.97–0.99 mm; abdomen 0.77–0.80/ 0.93–0.94 mm; length of antenna 1.19–1.23 mm; maximum width of genitalia (Fig. 59G) 0.26 mm.</p> <p>Comparative notes. The new species is related to a number of species centered on S. grandis, with all sharing a hyaline apophysis at the oblique mesal margin of antennomeres 9, and with a single basal protuberance consisting of a curved apical setose tuft on antennomere 11. Sathytes sympatricus differs particularly in the relatively small body size (2.7 mm), the long hyaline apophyses of antennomeres 9, the short basal protuberances of antennomeres 11, and the form of the aedeagus.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 84C, 95A, B)</p> <p>Etymology. The specific epithet ‘ sympatricus, - a, - um ’ is a Latin adjective indicating that this species is sympatric with a few congeners.</p> </div>	http://treatment.plazi.org/id/C822F213FF4C24FECB88A00674B327F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4224FECB88A0B2768125BE.text	C822F213FF4224FECB88A0B2768125BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes transversus Yin & Shen 2020	<div><p>Sathytes transversus Yin &amp; Shen, 2020</p> <p>(Figs 84C, 97A–C, 100A)</p> <p>Chinese common name: ẄƟḛà甲</p> <p>Sathytes transversus Yin &amp; Shen, 2020: 58; fig. 13. Type locality: China, Xizang, Nyingchi, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, alt. 2190 m.</p> <p>Additional material examined (7 exx.). 1 ♂, 3 ♀♀, ‘ China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96611&amp;materialsCitation.latitude=30.009722" title="Search Plazi for locations around (long 94.96611/lat 30.009722)">Pailong Town</a>, 30°0’35”N, 94°57’58”E, 2180 m, 10.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., Ʊ Ẫh 治区ffż市ḦË乡 ’; 1 ♂, 2 ♀♀, ‘China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200-2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’. (SNUC).</p> <p>Distribution. Chagyib District (Nyingchi) and Mêdog County, Tibet, SW China (Figs 84C, 97A–C, 100A).</p> <p>Remarks. Sathytes panzhaohuii was described based on material from Pailong and 80K near Mêdog County (ẸṘ县) seat, the new material extends the range of this species to Yigong.</p> </div>	http://treatment.plazi.org/id/C822F213FF4224FECB88A0B2768125BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4224FCCB88A57F745725B2.text	C822F213FF4224FCCB88A57F745725B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes virupaksa Yin 2022	<div><p>Sathytes virupaksa sp. nov.</p> <p>(Figs 60, 84C, 97D)</p> <p>Chinese common name: 广ƃḛà甲</p> <p>Type material (3 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, Tongmai Town, pass nr. 102 <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.13528&amp;materialsCitation.latitude=30.070835" title="Search Plazi for locations around (long 95.13528/lat 30.070835)">Tunnel</a>, 30°04’15”N, 95°08’07”E, 2150–2400 m, 12.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫȃ 9 102 Ṁdz ’ (SNUC). PARATYPES: CHINA: 2 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.6 mm; ratio (length/width) of head 1.07, pronotum 1.03, elytra 0.78, length of eye/tempus 0.88. Antennomere 9 as long as wide (length/width excluding apophysis 0.90), with hyaline apophysis at basal 2/5 of strongly oblique mesal margin, antennomere 11 elongate (length/width 1.89), with distinct protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus dilated at apex, apical margin moderately emarginate at middle. Female. Body length approximately 2.7–2.9 mm; antenna lacking modifications, genitalia as in Fig. 60G.</p> <p>Description. Male. Body (Fig. 60A) length 2.64 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short pubescence.</p> <p>Head (Fig. 60B) sub-rectangular at base, slightly longer than wide, length 0.59 mm, width across eyes 0.55 mm, length/width 1.07; vertex weakly raised, with small asetose foveae (dorsal tentorial pits) located posterior to level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, shallowly impressed medially; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), with faint carina posterior mouthparts. Compound eyes prominent, composed of approximately 20 large ommatidia, length of eye/tempus 0.88. Antenna elongate, length 1.31 mm, distinct club (Fig. 60C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, subequal in width, 9 (Fig. 60D) strongly enlarged, approximately as long as wide, length/width excluding apophysis 0.90, mesal margin expanded mesally, anteriorly strongly oblique, hyaline apophysis located at approximately basal 2/5 of mesal margin, 10 transverse, length/width 0.90, 11 largest, much longer than 9 and 10 combined, length/width 1.89, with short, distinct protuberance at base, apex of protuberance with curved tuft of golden setae.</p> <p>Pronotum (Fig. 60B) approximately as long as wide, length 0.62 mm, width 0.60 mm, length/width 1.03, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.82 mm, width 1.05 mm, length/width 0.78; each elytron with four large, asetose basal foveae; lacking discal stria; humerus weakly angulate, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused, large lateral mesoventral foveae forked internally, mesoventral process short. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, lacking median pit anterior to posterior margin, posterior margin broadly emarginate, slightly convex at middle, with thin marginal carina.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.77 mm, width 0.92 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal and marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 60E) moderately sclerotized, stout, apex with three long setae.</p> <p>Aedeagus (Fig. 60F) 0.32 mm long, slightly asymmetric, plate-like; median lobe with large foramen, apex dilated, much broader than base, apical margin moderately emarginate at middle.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 13 ommatidia; humerus weakly angulate. Measurements (as for male): body length 2.67–2.91 mm; length/width of head 0.62/ 0.53–0.55 mm, pronotum 0.62–0.63/ 0.61–0.62 mm, elytra 0.80/ 1.01–1.04 mm; abdomen 0.85–0.91/ 0.94–0.98 mm; length of antenna 1.28–1.37 mm; maximum width of genitalia (Fig. 60G) 0.28 mm.</p> <p>Comparative notes. This species resembles S. chayuensis which is found in Zayü County and S. aequalis by the relatively long antennomeres 11 and he markedly oblique mesal margins of antennomeres 9. Sathytes virupaksa can be distinguished from both of these species by the smaller body size (2.64 mm vs. 2.80–2.99 mm for S. chayuensis, 2.87–2.95 mm for S. aequalis), the relatively much shorter basal protuberances of antennomeres 11, the more posteriorly placed apophyses of antennomeres 9, as well as the dilated apex of the aedeagus.</p> <p>Distribution. Bomê County, Tibet, SW China (Figs 84C, 97D).</p> <p>Etymology. This species is named after Virűpâkṣa (广ƃƙƚ), who is one of the Four Great Heavenly Kings of Buddhism.</p></div> 	http://treatment.plazi.org/id/C822F213FF4224FCCB88A57F745725B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4024FCCB88A57271DF207D.text	C822F213FF4024FCCB88A57271DF207D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes xizangensis Yin & Shen 2020	<div><p>Sathytes xizangensis Yin &amp; Shen, 2020</p> <p>(Figs 85A, 97C, D)</p> <p>Chinese common name: ƱẪḛà甲</p> <p>Sathytes xizangensis Yin &amp; Shen, 2020: 63; fig. 16. Type locality: China, Xizang, Nyingchi, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.96601&amp;materialsCitation.latitude=30.009642" title="Search Plazi for locations around (long 94.96601/lat 30.009642)">Pailong Country</a>, 30°0’34.71”N, 94°57’57.64”E, alt. 2190 m.</p> <p>Additional material examined (8 exx.). 1 ♂, ‘ China: Xizang, Nyingchi City, Pailong Township, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.87583&amp;materialsCitation.latitude=29.983889" title="Search Plazi for locations around (long 94.87583/lat 29.983889)">Layue Bridge</a>, 29°59’02”N, 94°52’33”E, 2550 m, 13.vii.2021, Peng leg., Ʊ Ẫffż 市ḦË乡Ń月ss桥 ’; 2 ♂♂, 5 ♀♀, ‘ China: Xizang, Nyingchi City, Tongmai Town, pass nr. 102 <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.13528&amp;materialsCitation.latitude=30.070835" title="Search Plazi for locations around (long 95.13528/lat 30.070835)">Tunnel</a>, 30°04’15”N, 95°08’07”E, 2150–2400 m, 12.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫȃ 9 102 Ṁdz ’ (SNUC).</p> <p>Distribution. Chagyib District (Nyingchi) and Bomê County, Tibet, SW China (Figs 97C, D).</p> <p>Remarks. Sathytes xizangensis was described based on a large series from areas near Pailong Township. The new material confirms that it is a common species found from Pailong to Tongmai Townships.</p> </div>	http://treatment.plazi.org/id/C822F213FF4024FCCB88A57271DF207D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4024FACB88A73F7127271E.text	C822F213FF4024FACB88A73F7127271E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sathytes yigongensis Yin 2022	<div><p>Sathytes yigongensis sp. nov.</p> <p>(Figs 61, 85A, 97A–C)</p> <p>Chinese common name: ƌŭḛà甲</p> <p>Type material (13 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.93083&amp;materialsCitation.latitude=30.170555" title="Search Plazi for locations around (long 94.93083/lat 30.170555)">Yigong Tea Farm</a>, 30°10’14”N, 94°55’51”E, 2200–2400 m, 11.vii.2021, Peng, Yin &amp; Zhang, Ʊ Ẫffż 波ṁ县ƌŭē厂 ’ (SNUC). PARATYPES: CHINA: 3 ♂♂, 4 ♀♀, same data as that of holotype; 1 ♂, 4 ♀♀, ‘ China: Xizang, Nyingchi City, Pailong Township, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.87583&amp;materialsCitation.latitude=29.983889" title="Search Plazi for locations around (long 94.87583/lat 29.983889)">Layue Bridge</a>, 29°59’02”N, 94°52’33”E, 2550 m, 13.vii.2021, Peng leg. Ʊ Ẫffż 市ḦË乡Ń 月ss桥 ’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.2 mm; ratio (length/width) of head 1.06–1.08, pronotum 1.02– 1.06, elytra 0.79, length of eye/tempus 1.0. Antennomere 9 much wider than long (length/width excluding apophysis 0.80), with hyaline apophysis near apex of mesal margin, antennomere 11 elongate (length/width 1.72), with short protuberance at base, apex of protuberance with tuft of curved setae. Aedeagus broadly rounded at base, slightly bent towards right, apical margin convex at left side. Female. Body length approximately 2.3 mm; antenna lacking modifications, genitalia as in Fig. 61G.</p> <p>Description. Male. Body (Fig. 61A) length 2.18 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with short, dense pubescence.</p> <p>Head (Fig. 61B) sub-rectangular at base, slightly longer than wide, length 0.50–0.52 mm, width across eyes 0.47–0.48 mm, length/width 1.06–1.08; vertex weakly raised, with small asetose foveae (dorsal tentorial pits) at level of posterior margin of eyes; frons anteriorly confluent with strongly declining clypeus, weakly impressed medially; clypeus with smooth surface, anterior margin carinate and moderately raised; ocular-mandibular carina short; postgenal region strongly projecting. Venter with small, broadly separated gular foveae (posterior tentorial pits), lacking median carina. Compound eyes moderately prominent, composed of approximately 18 large ommatidia, length of eye/tempus 1.0. Antenna elongate, length 1.13 mm, distinct club (Fig. 61C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each transverse, 2 slightly wider, 3–8 subequal in width, 9 (Fig. 61D) strongly expanded, slightly wider than long, length/width excluding apophysis 0.80, mesal margin moderately oblique, hyaline apophysis located near apex of mesal margin, 10 transverse, length/width 0.62, 11 largest, longer than 9 and 10 combined, length/width 1.72, with short, apically denticulate protuberance at base, apex of protuberance with short, curved tuft of golden setae.</p> <p>Pronotum (Fig. 61B) approximately as long as wide, length 0.51–0.52 mm, width 0.49–0.50 mm, length/width 1.02–1.06, widest at middle; lateral margins rounded, convergent apically and basally; disc weakly convex; asetose lateral antebasal foveae moderately small. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge absent; margin of coxal cavity moderately carinate.</p> <p>Elytra much wider than long, length 0.69–0.71 mm, width 0.87–0.90 mm, length/width 0.79; each elytron with four small, asetose basal foveae; lacking discal stria; humerus faintly prominent, subhumeral fovea present, marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by strongly curved carinae; median mesoventral foveae fused at middle, originating from shared opening, large lateral mesoventral foveae shallowly forked internally, mesoventral process short, with complete marginal stria. Metaventrite weakly prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two broadly separated lateral metaventral foveae, with large, round median pit shortly anterior to posterior margin, posterior margin broadly emarginate.</p> <p>Legs elongate, simple.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.57–0.62 mm, width 0.78–0.79 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking discal or marginal carina; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) longer than 3 but much shorter than 2 and 3 combined; tergites 2–3 (IV–VI) each with one pair of mediobasal and two pairs of basolateral foveae, 4 with one pair of basolateral foveae and marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 (IV–VI) each with three pairs and 5 (VII) with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 61E) moderately sclerotized, elongate, apex lacking setae.</p> <p>Aedeagus (Fig. 61F) 0.22 mm long, slightly bent rightwards, plate-like; median lobe with large foramen, apical margin slightly convex at left side.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications; each compound eye composed of approximately 15 ommatidia; humeral prominence faint. Measurements (as for male): body length 2.28–2.30 mm; length/width of head 0.50–0.52/ 0.47–0.48 mm, pronotum 0.51/ 0.51–0.52 mm, elytra 0.61–0.63/ 0.86 mm; abdomen 0.73–0.75/ 0.81 mm; length of antenna 1.00– 1.05mm; maximum width of genitalia (Fig. 61G) 0.25 mm.</p> <p>Comparative notes. This species is morphologically most similar to S. panzhaohuii which are sympatric in Yigong, sharing with it a similar body size and an acute apical tubercle of the basal protuberance of antennomeres 11. They can be best separated by the relatively more elongate antennomeres 11, a steeper declining angle of the apicomesal margin of antennomeres 9, and a more anteriorly placed location of the apophysis on this antennomere for S. yigongensis. The new species is also similar to S. pengzhongi and S. proclivis by the oblique mesal margins and apically located apophyses of male antennomeres 9. The male of S. yigongensis can be readily separated by the angulate tubercles at the apices of the basal protuberances of antennomeres 11, different forms of antennomeres 9 and the apophyses on them, as well as the different form of the aedeagus.</p> <p>Distribution. Bomê County, Tibet, SW China (Figs 85A, 97A–C).</p> <p>Etymology. The new species is named after its type locality, Yigong Township.</p></div> 	http://treatment.plazi.org/id/C822F213FF4024FACB88A73F7127271E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4524F8CB88A22D70FB2288.text	C822F213FF4524F8CB88A22D70FB2288.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites Jeannel 1960	<div><p>Tribasodites Jeannel, 1960</p> <p>Chinese common name: ûḃà甲¤</p> <p>Tribasodites Jeannel, 1960: 411; Newton &amp; Chandler 1989 (catalog); Sugaya &amp; Maruyama 2003 (biological notes); Yin et al. 2011d, 2015b (cave species); Nomura &amp; Aung 2020 (first record from Myanmar). Type species: Tribasodites antennalis Jeannel, 1960: 412 (original designation).</p> <p>This genus appears to be poorly defined by Jeannel (1960), since the presence of three basal elytra foveae and discal and marginal spines of the pronotum, as well as a relatively short tergite 1 (IV) of the abdomen is shared by many related genera. All species described here possess a lateral frontal fovea, which may potentially serve as a synapomorphy of the genus, or of a broader group that contains Tribasodites. Two major lineages occur in Tibet. The first contains six species that have modified antennomeres 11 (and often antennomeres 10), relatively short postocular margins, lack of a distinct row of discal spines on the pronotum, and the aedeagus with a plate-like dorsal lobe that partially surrounds the ventral stalk of the median lobe. The second lineage also includes six species, with all having elongate and simple antennae, the postocular margins of the head are relatively much longer, the pronotum usually bears two distinct rows of discal spines, and the aedeagus often has an elongate and slender ventral stalk of the median lobe and dorsal lobe. Tribasodites vertexalis, which possesses simple antennae, a modified vertex, and has a different aedeagal form, is also placed here. Numerous undescribed species possessing male sexual and aedeagal characters of varying forms have been seen from China and other areas of Asia.</p> <p>In Asia, Tribasodites contains many species that inhabit cave environments (Yin et al. 2011d, 2015b; Yin &amp; Zhou 2018; Yin &amp; He 2020), and others were commonly collected in decomposing logs or under bark of dead fallen trees, often with ants or termites (Nomura 1986; Zhao et al. 2010; Inoue &amp; Maruyama 2020). However, there is no direct evidence, both ecologically or morphologically, indicating that these beetles are oblique myrmecophiles or termitophiles (Sugaya &amp; Maruyama 2003), because many species of Tribasodites were often found without ants/ termites from the same habitats. Tribasodites hubeiensis Yin, Nomura &amp; Li, originally described from material collected in two caves in southern Hubei, was found in a leaf litter sample at Maoer Mountain (á儿山) of Guangxi (W.-X. Zhang &amp; Z.-W. Yin unpublished data), some 500 km south from the type locality. It is inferred that at least some of the cavernicolous Tribasodites are loosely associated with cave habitats. The fact that most of the described species have been taken only in caves likely reflects inadequate sampling efforts in the surrounding environments. Three species described here and that are sympatric in Gyirong were collected under barks on the same day: T, gyirong, T, kiypu, T, pugiunculus.</p> <p>The 13 Tibetan species may be distinguished by using the following key.</p> <p>Key to Tibetan species of Tribasodites (male)</p> <p>1 Vertex and frons each with large median tubercle (Fig. 73B).................................... T. vertexalis sp. nov.</p> <p>- Vertex and frons simple, lacking modifications (Figs 62–72, 74)................................................ 2</p> <p>2 Antennomere 10 and/or 11 modified; pronotum lacking row of spines along discal carina; aedeagus with broad dorsal lobe partially surrounding ventral stalk of median lobe (Figs 62–64, 66, 68, 72)........................................ 3</p> <p>- Antenna simple, lacking modifications; pronotum often with row of spines along discal carina (spines lacking in two species that have modified metafemur); aedeagus usually with long and slender median and dorsal lobe (one species has broad dorsal lobe, but not partially surrounding ventral stalk) (Figs 65, 67, 69–71, 74)......................................... 8</p> <p>3 Antennomere 10 lacking impression or cavity on ventral side.................................................. 4</p> <p>- Antennomere 10 with impression or cavity on ventral side (Figs 62C, 66D, 68C, 72D).............................. 5</p> <p>4 Pronotum with small marginal spines (Fig. 63B); mesofemur lacking spine on ventral margin........ T. cellulanus sp. nov.</p> <p>- Pronotum lacking marginal spines (Fig. 64B); mesofemur with long ventral spine (Fig. 64D) at base....................................................................................................... T. constrictus sp. nov.</p> <p>5 Head dark brown, much darker than pronotum and elytra which are reddish-brown (Fig. 66A, B); antennomere 11 with broad, short prominence at base (Fig. 66C)....................................................... T. elongatus sp. nov.</p> <p>- Head, pronotum and elytra reddish-brown, unicolorous; antennomere 11 with distinctly much longer projection or protuberance at base.............................................................................................. 6</p> <p>6 Antennomere 11 with markedly large and extended basal projection (Fig. 62C); mesotibia with indistinct apical tubercle............................................................................................. T. bari sp. nov.</p> <p>- Antennomere 11 with distinct conical basal protuberance; mesotibia with large, blade-like apical spine.................. 7</p> <p>7 Body length greater than 2.5 mm (2.68–2.78 mm); anterior portion of vertex with distinct transverse impression (Fig. 68B); aedeagus (Fig. 68I, J)..................................................................... T. gyirong sp. nov.</p> <p>- Body length smaller than 2.5 mm (2.30–2.41 mm); anterior portion of vertex lacking distinct impression (Fig. 72B); aedeagus (Fig. 72I, J)........................................................................ T. pugiunculus sp. nov.</p> <p>8 Body length greater than 2.5 mm (2.52–2.53 mm)............................................... T. kiypu sp. nov.</p> <p>- Body length smaller than 2.2 mm......................................................................... 9</p> <p>9 Metatibia expanded and with curved setose tuft on dorsal margin (Fig. 70F)........................ T. mirabilis sp. nov.</p> <p>- Metatibia simple..................................................................................... 10</p> <p>10 Pronotum lacking row of spines along discal carina; metafemur modified........................................ 11</p> <p>- Pronotum with row of spines along discal carina; metafemur simple............................................ 12</p> <p>11 Head relatively broader, ratio (length/width) 43:47, eyes weakly prominent (with 35 ommatidia); dorsal lobe of aedeagus strongly twisted at apex (Fig. 67H–J)..................................................... T. grandiceps sp. nov.</p> <p>- Head relatively longer, ratio (length/width) 49:48, eyes markedly prominent (with 45 ommatidia); dorsal lobe of aedeagus slightly curved at apex (Fig. 71H–J).................................................... T. prolixicornis sp. nov.</p> <p>12 Dorsal lobe of aedeagus conspicuously broad in dorsal view (Fig. 65G)............................ T. dilophus sp. nov.</p> <p>- Dorsal lobe of aedeagus long and slender in dorsal view (Fig. 74H, J).............................. T. yatung sp. nov.</p></div> 	http://treatment.plazi.org/id/C822F213FF4524F8CB88A22D70FB2288	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF4424E5CB88A44077DB27AA.text	C822F213FF4424E5CB88A44077DB27AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites bari Yin 2022	<div><p>Tribasodites bari sp. nov.</p> <p>(Figs 62, 85C, 99F)</p> <p>Chinese common name: 巴Uûḃà甲</p> <p>Type material (10 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Mêdog County, pass from <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.3575&amp;materialsCitation.latitude=29.3125" title="Search Plazi for locations around (long 95.3575/lat 29.3125)">Bari Village</a> to Renqingbeng Temple, 29°18’45”N, 95°21’27”E, ca. 1830 m, 19.iii.2017, X.-B. Song leg. [ƱẪẸṘ县巴U村 -仁ů崩寺]’ (SNUC). PARATYPES: CHINA: 4 ♂♂, 5 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.0 mm. Head sub-rectangular at base, narrower than pronotum, tempus moderately short, denticulate at posterolateral angle, vertex lacking sulcus, with relatively large, asetose foveae, with thin, short mediobasal carina extending from head base anteriorly beyond level of anterior margin of eyes, with distinct lateral carina from base to posterior margin of antennal tubercle; antenna elongate, with modified antennomeres 9–11, antennomere 11 with long basal projection extended posteriorly from mesal margin of basal cavity. Pronotum with laterally carinate median and lateral longitudinal sulci, with pair of discal carinae, with small antebasal and much larger marginal spines. Discal stria of elytron shallow, extending posteriorly to basal 3/5 of elytral length; disc with relatively large and shallow punctures. Protibia with small preapical denticle, protrochanter with tiny ventral tubercle; mesotibia with distinct apical spine; metatrochanter with large projection on ventral margin, mesal margin of metatibia with row of dense golden setae along apical 2/5. Aedeagus strongly asymmetric; median lobe with elongate basal capsule and foramen, ventral stalk broad at middle and then strongly narrowing apically, dorsal lobe large, plate-like, protruding apically and with numerous spine-like structures at apex. Female. Body length approximately 2.0 mm; antenna lacking modifications, legs lacking spine, tubercle, or projection, genitalia as in Fig. 62H.</p> <p>Description. Male. Body (Fig. 62A) length 1.96–1.98 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 62B) sub-rectangular at base, much wider than long, length 0.36–0.37 mm, width across eyes 0.45– 0.46 mm; vertex smooth, lacking sulcus, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina thin, extending from head base anteriorly to beyond level of anterior margin of eyes, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral angle angulate; frons anteriorly demarcated from clypeus by frontal-clypeal ridge, weakly impressed between large, moderately raised antennal tubercles; clypeus with smooth surface, its entire anterior margin carinate and moderately raised; ocular-mandibular carina complete, distinct. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared round opening, with faint median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 45 ommatidia. Antenna elongate, length 1.10–1.13 mm, distinct club (Fig. 62C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, 9 much wider than 8, asymmetric, 10 much broader and longer than 9, anterior margin inclined, ventral surface with broad impression at base, 11 largest, longer than 9 and 10 combined, constricted at basal 1/3, large basal projection extended posteriorly from mesal margin of deep basal cavity.</p> <p>Pronotum (Fig. 62B) wider than long, length 0.45 mm, width 0.50–0.51 mm, widest at middle; lateral margins rounded at apical 1/2, convergent basally and parallel at basal 1/3; disc slightly convex, finely punctate, median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of discal carinae and lateral longitudinal sulci; with small antebasal and much larger marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra wider than long, length 0.61–0.62 mm, width 0.72–0.74 mm; each elytron with three moderately large, asetose basal foveae; shallow discal stria extending posteriorly from outer basal fovea to basal 3/5 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared opening, large lateral mesoventral foveae forked internally, with short mesoventral process, marginal stria complete. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle.</p> <p>Legs elongate; protrochanter with small ventral spine, protibia with small preapical denticle at mesal margin; mesotibia with short but distinct spine at apex; metatrochanter (Fig. 62D) with large ventral projection, metatibia with dense golden setae along mesal margin from basal 3/5 to apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.54–0.55 mm, width 0.65–0.68 mm. Tergite 1 (IV) more than twice as long as 2 (V), thin basal sulcus separated by widely separated mediobasal and one pair of large basolateral foveae, inner marginal carina oblique, complete, outer one much thicker, complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, with short lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 62E) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 62F, G) 0.44 mm long, strongly asymmetric, elongate; median lobe with elongate basal capsule and foramen, ventral stalk in dorsal view broad at middle, narrowing towards apex; dorsal lobe large, plate-like, apical part protruding and curved, with numerous spine-like structures at apex; parameres membranous, apically split into two parts.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter, lacking modifications, legs lacking tubercle, spine or projection; each compound eye composed of approximately 40 ommatidia; humerus slightly denticulate. Measurements (as for male): body length 1.98–2.09 mm; length/width of head 0.38–0.42/ 0.47–0.51 mm, pronotum 0.45–0.47/ 0.53 mm, elytra 0.60–0.62/ 0.74–0.79 mm; abdomen 0.56–0.61/ 0.67–0.71 mm; length of antenna 1.00 mm; maximum width of genitalia (Fig. 62H) 0.19 mm.</p> <p>Comparative notes. Tribasodites bari is apparently close to T. antennalis Jeannel from northern India, which is the type species of the genus, by the similar form of the male antennal modifications and a similar general form of the aedeagus. They can be separated by the denticulate posterolateral angles of the head (rounded in T. antennalis), a much broader mesal margin of antennomere 10 (roundly truncate in T. bari, anterior and posterior margin inclined and mesally forming an acute apex in T. antennalis), and a much longer basal projection of antennomere 11. The new species also shares with T. pugiunculus a similar appearance of the head, pronotum and elytra, but can be separated from this species by the different structure of the antennal modifications and the aedeagus, and much smaller apical spines of the male mesotibiae. The denticulate posterolateral angles of the head of T. bari similarly occur in the Japanese T. picticornis Nomura (1986: fig. 1), but otherwise these two species can be distinguished by the different form of the male antennae and the aedeagus, as well as by their distributions.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 85C, 99F).</p> <p>Etymology. The new species is named after its type locality, i. e., Bari Village.</p> <p>Remarks. One male specimen was found with parasitic Laboulbeniales on the anterior margin of right femur (Fig. 62I) and on the lateral margin of right tibia.</p></div> 	http://treatment.plazi.org/id/C822F213FF4424E5CB88A44077DB27AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5924E3CB88A30A70FA229A.text	C822F213FF5924E3CB88A30A70FA229A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites cellulanus Yin 2022	<div><p>Tribasodites cellulanus sp. nov.</p> <p>(Figs 63, 85C, 92C)</p> <p>Chinese common name: Ḵ士ûḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyalam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.97806&amp;materialsCitation.latitude=27.955833" title="Search Plazi for locations around (long 85.97806/lat 27.955833)">Zhangmu</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.97806&amp;materialsCitation.latitude=27.955833" title="Search Plazi for locations around (long 85.97806/lat 27.955833)">Lixin Village</a> (ĒẾ 村), 27°57’21”N, 85°58’41”E, leaf litter, sifted, 2400–2600 m, 27.vii.2010, Jian-Qing Zhu leg.’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.2 mm. Head subtruncate at base, slightly narrower than pronotum, tempus moderately short, rounded at posterolateral angle, vertex with transverse impression anteriorly, with relatively large, setose foveae, with distinct mediobasal carina extending from near head base anteriorly slightly at level of middle length of eyes, with distinct oblique lateral carina from base to posterior margin of antennal tubercle; antenna elongate, antennomere 11 with short broad projection at base. Pronotum with laterally carinate median and lateral longitudinal sulci, with pair of discal carinae, with two pairs of antebasal and distinct marginal spines. Discal stria of elytron long, extending posteriorly to more than apical 3/4 of elytral length; disc finely punctate.All trochanters with ventral spine or projection, mesotibia simple. Aedeagus strongly asymmetric; median lobe with extended basal capsule and large foramen, ventral stalk in lateral view elongate and sinuate, dorsal lobe large, plate-like, with elongate sclerite broadly forked at apex.</p> <p>Description. Male. Body (Fig. 63A) length 2.19 mm; head, pronotum, abdomen dark reddish-brown, legs, antennae and elytra reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 63B) sub-rectangular at base, much wider than long, length 0.43 mm, width across eyes 0.52 mm; vertex smooth, with broad transverse impression demarcating from frons, vertexal foveae (dorsal tentorial pits) setose, relatively large, mediobasal carina distinct, extending from near head base anteriorly to slightly posterior to level of middle length of eyes, lateral carina distinct, curved, extending from head base to posterior margin of antennal tubercle; posterolateral margin rounded; frons weakly impressed between large, moderately raised antennal tubercles, anteriorly demarcated from clypeus by frontal-clypeal ridge; clypeus with smooth surface, short, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with smooth surface; small gular foveae (posterior tentorial pits) fused, originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes moderately prominent, composed of approximately 35 ommatidia. Antenna moderately elongate, length 1.13 mm, distinct club (Fig. 63C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate to as long as wide, 8 smallest, moniliform, 9 much wider than 8, transverse, 10 much broader and slightly longer than 9, 11 largest, approximately as long as than 8–10 combined, with short, subtriangular projection at base of mesal margin, apex of projection with small setose tuft.</p> <p>Pronotum (Fig. 63B) slightly wider than long, length 0.50 mm, width 0.56 mm, widest at middle; lateral margins rounded, smooth at apical 1/2, convergent basally; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of curved discal carinae and lateral longitudinal sulci; with two pairs of blunt antebasal and distinct marginal spines; lateral antebasal foveae small and setose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thick hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity broadly carinate.</p> <p>Elytra much wider than long, length 0.57 mm, width 0.76 mm; each elytron with three large, asetose basal foveae; long discal stria extending posteriorly from outer basal fovea to more than apical 1/3 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared median opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae moving medially and in large, setose shared impression; posterior margin roundly emarginate at middle, lacking split.</p> <p>Legs elongate; procoxa with exceptionally long seta at base, protrochanter (Fig. 63D) with small ventral spine, protibia with tiny preapical denticle at mesal margin; mesotrochanter (Fig. 63E) with long ventral spine, mesotibia simple; metatrochanter (Fig. 63F) with long, strongly curved ventral projection at apex, metatibia lacking tuft of long setae at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.71 mm, width 0.75 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with broadly separated mediobasal and one pair of basolateral foveae, with pair of short discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, 3 and 4 each with three and 5 with two pairs of small basolateral foveae and short lateral carinae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 63G) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 63H, I) 0.51 mm long, strongly asymmetric, elongate; median lobe with large extended basal capsule and large, elongate foramen, ventral stalk in dorsal view broad, laterally sinuate and narrowing towards apex; dorsal lobe large, plate-like, with elongate sclerite broadly forked at apex; parameres reduced to single broad membrane.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species shares with T. bari, T. constrictus, T. gyirong, and T. pugiunculus modified male antennomeres 11 and similar aedeagal forms, and is sympatric with T. pugiunculus near Zhangmu. The male of Tribasodites cellulanus can be distinguished from that of T. pugiunculus by the shorter elytra, the more robust antennomeres 11, the lack of a long apical spine on the mesotibia, as well as a different aedeagal form. This combination of characters also separates this species from all other similar congeners.</p> <p>Distribution. Nyalam County, Tibet, SW China (Figs 85C, 92C).</p> <p>Etymology. The species epithet ‘ cellulanus (hermit, recluse)’ is a Latin noun referring to the rarity of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FF5924E3CB88A30A70FA229A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5F24E1CB88A45A7720278E.text	C822F213FF5F24E1CB88A45A7720278E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites constrictus Yin 2022	<div><p>Tribasodites constrictus sp. nov.</p> <p>(Figs 64, 85C, 95A, B)</p> <p>Chinese common name: 束aûḃà甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Cona County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.763336&amp;materialsCitation.latitude=27.837778" title="Search Plazi for locations around (long 91.763336/lat 27.837778)">Lebu Valley</a>, pass to Mama Vill., 27°50’16”N, 91°45’48”E, 2400–2700 m, 04.vii.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪDZḄ县勒布沟Þ玛 乡 ’ (SNUC). PARATYPES: CHINA: 3 ♂♂, 5 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.2 mm. Head subtruncate at base, slightly narrower than pronotum, tempus moderately short, rounded at posterolateral angle, vertex with incomplete reversed U-shaped sulcus connecting foveae, with relatively large, setose foveae, with distinct mediobasal carina extending from head base anteriorly to level of eye midlength, lateral carina absent; antenna elongate, antennomere 11 with thin, long basal projection. Pronotum with laterally carinate median and lateral longitudinal sulci, with pair of discal carinae, with large antebasal protuberance, lacking marginal spine. Elytra constricted at bases, discal stria thin and shallow, extending posteriorly to approximately apical 1/3 of elytral length; disc finely punctate. Mesotibia with small apical spine. Aedeagus strongly asymmetric; median lobe with extended basal capsule and elongate foramen, ventral stalk in lateral view deeply forked at apical portion, dorsal lobe large, plate-like, embracing median lobe. Female. Body length approximately 2.1 mm, antenna shorter than male, antenna and legs lacking modifications, genitalia as in Fig. 64J.</p> <p>Description. Male. Body (Fig. 64A) length 2.22–2.27 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively long pubescence.</p> <p>Head (Fig. 64B) roundly at base, much wider than long, length 0.40–0.42 mm, width across eyes 0.48–0.50 mm; vertex smooth, with incomplete reversed U-shaped sulcus connecting relatively large, setose vertexal foveae (dorsal tentorial pits), mediobasal carina distinct, extending from head base anteriorly to level of eye midlength, lateral carina absent; posterolateral margin rounded; frons impressed between large, moderately raised antennal tubercles, anteriorly confluent with clypeus; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, median carina present only posterior mouthparts for short distance. Compound eyes prominent, composed of approximately 22 ommatidia. Antenna elongate, length 1.09–1.14 mm, indistinct club (Fig. 64C) formed by moderately enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each sub-moniliform, 7 distinctly enlarged, much longer than neighboring antennomeres, 9 slightly transverse, much wider than 8, 10 strongly transverse, much broader and approximately as long as 9, ventral surface lacking impression, 11 largest, longer than 9 and 10 combined, with distinct basoventral projection, apex of projection with elongate setae.</p> <p>Pronotum (Fig. 64B) slightly wider than long, length 0.50–0.51 mm, width 0.53 mm, widest at middle; lateral margins rounded, convergent anteriorly and evenly narrowing posteriorly for basal 1/3; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with large, oval antebasal impression and short mediobasal carina, with pair of discal carinae and lateral longitudinal sulci; with large pair of antebasal protuberances, lacking marginal spines; lateral antebasal foveae large and setose; with distinct outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thick hypomeral ridge extending from base to middle of anterior part, with anterior and lateral antebasal hypomeral impressions; margin of coxal cavity carinate.</p> <p>Elytra basally constricted, much wider than long, length 0.60–0.61 mm, width 0.74–0.75 mm; each elytron with three moderately large, asetose basal foveae; long discal stria extending posteriorly from outer basal fovea to approximately apical 1/3 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae and pair of lateral metaventral foveae; posterior margin roundly emarginate at middle, with narrow split.</p> <p>Legs elongate; procoxa with exceptionally long seta at base, protibia with tiny preapical denticle at mesal margin; mesotrochanter (Fig. 64D) with short ventral tubercle, mesofemur (Fig. 64D) with long ventral spine at base, mesotibia (Fig. 64E) with small apical protuberance; metatrochanter (Fig. 64F) with large, long ventral projection strongly curved at apex.</p> <p>Abdomen narrowed before base, widest at lateral margins of tergite 1 (IV), length 0.73–0.76 mm, width 0.75– 0.80 mm. Tergite 1 (IV) approximately 1.6 times as long as 2 (V), lacking basal sulcus, with large mediobasal and one pair of basolateral foveae, with pair of short discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin narrowly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, 3 and 4 each with three pairs and 5 with one pair of basolateral foveae, lacking lateral carinae, sternite 6 (VIII) longer than 3–5 combined, medially impressed, posterior margin broadly emarginate and slightly convex at middle, sternite 7 (IX) (Fig. 64G) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 64H, I) 0.58 mm long, strongly asymmetric, elongate; median lobe with large extended basal capsule and elongate foramen, ventral stalk in lateral view deeply forked into two elongate lobes; dorsal lobe large, plate-like, embracing median lobe, with rounded apex; parameres short, membranous.</p> <p>Female. Similar to male in external morphology; antenna slightly shorter than male; antenna and legs lacking modifications; each compound eye composed of approximately 15 ommatidia; tergite 1 (IV) with only one pair of basolateral foveae, lacking mediobasal foveae. Measurements (as for male): body length 2.08–2.10 mm; length/width of head 0.40/ 0.48–0.49 mm, pronotum 0.49/ 0.52–0.54 mm, elytra 0.53–0.54/ 0.70 mm; abdomen 0.71– 0.77/ 0.72 mm; length of antenna 0.96–0.97 mm; genitalia weakly sclerotized (Fig. 64J), with pair of membranous lateral plates, maximum width 0.32 mm.</p> <p>Comparative notes. The male of this species is close to those of T. bari, T. gyirong, and T. pugiunculus by antennomeres 11 having a basal projection, and a similar general form of the aedeagus. Tribasodites constrictus differs in the lack of marginal spines on the pronotum, the relatively small and basally constricted elytra, the presence of a long mesofemoral spine, and a different aedeagal form.</p> <p>Distribution. Cona County, Tibet, SW China (Figs 85C, 95A, B).</p> <p>Etymology. The specific epithet ‘ cônstrictus (bound together, tied up)’ is a Latin adjective suggested by the constricted elytral base of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FF5F24E1CB88A45A7720278E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5C24EECB88A1DB77032502.text	C822F213FF5C24EECB88A1DB77032502.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites dilophus Yin 2022	<div><p>Tribasodites dilophus sp. nov.</p> <p>(Figs 65, 86A, 96B, C)</p> <p>Chinese common name: ḦẆûḃà甲</p> <p>Type material (20 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.383606&amp;materialsCitation.latitude=29.57528" title="Search Plazi for locations around (long 94.383606/lat 29.57528)">Chongzhe</a>, 29°34’31”N, 94°23’01”E, 3230 m, 03.vii.2018, mixed leaf litter, sifted, Cheng, Peng &amp; Shen leg. [ƱẪffżΦĖṉữ]’ (SNUC). PARATYPES: CHINA: 2 ♂♂, same data as that of holotype; 5 ♂♂, 12 ♀♀, also from Nyingchi, except ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.16917&amp;materialsCitation.latitude=29.758888" title="Search Plazi for locations around (long 94.16917/lat 29.758888)">Kading Valley</a>, 29°45’32”N, 94°10’09”E, 3200 m, 08.vii.2021, Peng, Yin &amp; Zhang leg., Ʊ Ẫffż 市巴Ù区ϮŤ沟 Ḩā 区 ’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.1 mm. Head sub-rounded at base, approximately as wide as pronotum, tempus moderately long, with rounded posterolateral margin, vertex lacking sulcus, with relatively large, sparsely setose foveae, with distinct mediobasal carina extending from head base anteriorly slightly beyond level of anterior margin of vertexal foveae, with distinct lateral carina from base to posterior margin of antennal tubercle; antenna elongate, lacking modifications, antennomeres elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of longitudinal discal carinae composed of spines, with pair of large antebasal and small marginal spines. Discal stria of elytron extending posteriorly to approximately basal 1/3 of elytral length; disc finely punctate. Mesotrochanter with distinct ventral spine; metatrochanter with strongly curved projection on ventral margin. Aedeagus slightly asymmetric; median lobe with large basal capsule and elongate foramen, ventral stalk broad apically, lacking distinct dorsal lobe or parameres. Female. Body length approximately 2.0 mm, antenna shorter than male, legs lacking modifications, genitalia as in Fig. 65I.</p> <p>Description. Male. Body (Fig. 65A) length 2.08–2.15 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 65B) sub-rounded at base, approximately as wide as long, length 0.44–0.46 mm, width across eyes 0.46 mm; vertex smooth, lacking sulcus, vertexal foveae (dorsal tentorial pits) setose, relatively large, mediobasal carina distinct, extending from head base anteriorly to slightly beyond level of anterior margin of vertexal foveae, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral margin rounded; frons anteriorly demarcated from clypeus by thin frontal-clypeal ridge, area between moderately raised antennal tubercles weakly impressed; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 45–50 ommatidia. Antenna elongate, length 1.08–1.16 mm, indistinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each slightly elongate, 8 shortest, 9 much wider and longer than 8, elongate, 10 shorter than 9, subconical, 11 largest, slightly longer than 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 65B) approximately as long as wide, length 0.46–0.48 mm, width 0.44–0.45 mm, widest at middle; lateral margins rounded at middle, convergent apically and parallel at basal 1/4; disc slightly convex, finely punctate, thin median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and short mediobasal carina, pair of discal longitudinal carinae (Fig. 65C) each composed of five distinct spines, with shallow lateral longitudinal sulci; with pairs of large antebasal and small marginal spines; lateral antebasal foveae small and setose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity slightly carinate.</p> <p>Elytra much wider than long, length 0.66–0.69 mm, width 0.76–0.77 mm; each elytron with three moderately large, asetose basal foveae; discal stria short, extending posteriorly from outer basal fovea to approximately basal 1/3 elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long seta at base; mesotrochanter (Fig. 65D) with distinct ventral spine, mesotibia with small apical spur; metatrochanter (Fig. 65E) with long, strongly curved ventral projection, metatibia with short pencil-like apical tuft of setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.54–0.61 mm, width 0.65–0.67 mm. Tergite 1 (IV) approximately twice as long as 2 (V), lacking basal sulcus, with mediobasal and two pairs of basolateral foveae, with pair of moderately long, thick discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), with one pair of small mediobasal foveae, two pairs of small basolateral foveae and thin marginal carinae, 3 (VI) with small basolateral foveae and thin marginal carinae, 4 (VII) shorter than tergites 2 and 3 combined, with one pair of small basolateral foveae, lacking marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, sternites 3–4 each with three pairs and 5 with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, roundly convex at middle, sternite 7 (IX) (Fig. 65F) membranous, with weakly sclerotized central part.</p> <p>Aedeagus (Fig. 65G, H) 0.31 mm long, in dorsal view moderately asymmetric, elongate; median lobe with large laterally broad basal capsule and elongate foramen, ventral stalk in dorsal view broad, largely membranous, with weakly sclerotized, curved and apically expanded ventral lobe; lacking distinct articulated dorsal lobe or parameres.</p> <p>Female. Similar to male in external morphology; antenna as long as male; legs lacking modifications; each compound eye composed of approximately 35 ommatidia. Measurements (as for male): body length 2.0– 2.07 mm; length/width of head 0.43/ 0.43–0.45 mm, pronotum 0.45/ 0.44 mm, elytra 0.62–0.65/ 0.72–0.74 mm; abdomen 0.57– 0.58/ 0.64–0.65 mm; length of antenna 1.07–1.14 mm; genitalia weakly sclerotized (Fig. 65I), with elongate, forked lateral arms, maximum width 0.20 mm.</p> <p>Comparative notes. Tribasodites dilophus belongs to a group of species that share elongate and simple male antennae, relatively long postocular margins, often with longitudinal rows of discal spines on the pronotum, and the aedeagus usually with elongate median and dorsal lobes. This species differs by the broad ventral stalk and dorsal lobe of the aedeagus, and by its distribution.</p> <p>Distribution. Chagyib District (Nyingchi), Xizang, SW China (Figs 86A, 96B, C).</p> <p>Etymology. The specific epithet is taken from the Greek ‘ dilophos ’ (two-crested), referring to the two longitudinal rows of spines on the pronotal disc.</p></div> 	http://treatment.plazi.org/id/C822F213FF5C24EECB88A1DB77032502	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5224ECCB88A2E271772313.text	C822F213FF5224ECCB88A2E271772313.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites elongatus Yin 2022	<div><p>Tribasodites elongatus sp. nov.</p> <p>(Figs 66, 86A, 100C)</p> <p>Chinese common name: ḞẦẆḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.35225&amp;materialsCitation.latitude=28.62326" title="Search Plazi for locations around (long 97.35225/lat 28.62326)">China</a>: Xizang, Zayü County, valley nr. Zhala Vill., 28°37’23.74”N, 97°21’8.10”E, 2190 m, 2019.vii.28, leaf litter / decomposing log, Z.-W. Yin leg.; [ƱẪ察ø县扎 Ń村ṉữ]’ (SNUC).</p> <p>Diagnosis. Male. Habitus elongate; body length 2.61 mm. Head slightly narrower than pronotum, vertex with long mediobasal and complete lateral carinae; antennomere 10 concave on ventral surface, antennomere 11 with large tubercle at base. Pronotum with broad and laterally carinate median longitudinal sulcus, one pair of discal carinae and one pair of lateral sulci, with discal, antebasal and lateral denticles. Discal stria of elytron long, extending posterior to apical 5/6 of elytral length. Protibia with preapical denticle, trochanters each with ventral spine or projection. Aedeagus strongly asymmetric; median lobe strongly, basal capsule extended, strongly constricted at base, ventral stalk narrowing and strongly curved apically; with large, plate-like dorsal lobe; parameres reduced.</p> <p>Description. Male. Body (Fig. 66A) length 2.61 mm; color reddish-brown, head darkish brown, tarsi and mouthparts light reddish-brown. Dorsal surface of body covered with relatively short setae.</p> <p>Head (Fig. 66B) sub-rounded at base, slight wider than long, length 0.43 mm, width across eyes 0.48 mm; vertex finely punctate, with setose vertexal foveae (dorsal tentorial pits), lacking U-shaped sulcus connecting foveae, lateral vertexal carinae extending from head base to base of antennal tubercles, mediobasal carina extending from head base to level of middle of eyes; antennal tubercles moderately raised; frons impressed between antennal tubercles, demarcated from clypeus by transverse carina; clypeus with smooth surface, anterior margin carinate and raised; ocular-mandibular. carinae distinct. Venter with gular foveae (posterior tentorial pits) merged into single pit, gular carina present only for short distance near mouthpart. Eyes moderately prominent, composed of approximately 35 ommatidia. Antenna elongate, length 1.25 mm; club formed by apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each elongate, 8 smallest, 9 much larger than 8, subconical, 10 (Fig. 66C, D) much larger than 9, subconical, ventral surface broadly concave at base, 11 (Fig. 66C, D) largest, longer than 9 and 10 combined, truncate at base and narrowing apically, with large prominence at base, apex of prominence covered with short, thick setae.</p> <p>Pronotum (Fig. 66B) slightly broader than long, length 0.48 mm, width 0.51 mm, widest anterior middle; lateral margins rounded, with distinct lateral denticles; disc slightly convex, finely punctate, with small discal and antebasal denticles, broad median longitudinal impression with carinate edges extending posteriorly to round mediobasal impression, followed by short basal carina, with one pair of discal carinae evenly curved and one pair of lateral longitudinal sulci extending to large lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part as long as coxal part, hypomeral ridge extending from base to middle of anterior part; lateral procoxal foveae widely separated.</p> <p>Elytra wider than long, length 0.65 mm, width 0.77 mm, lateral margins rounded; each elytron with three large, asetose basal foveae, inner two close; humerus weakly denticulate; thin discal stria extending from outer fovea to apical 5/6 of elytral length; small subhumeral fovea present, with carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, confluent with metaventrite; median mesoventral foveae widely separated, lateral mesoventral foveae extending internally below mesoventral ones. Metaventrite moderately impressed at middle, with well-developed lateral mesocoxal foveae and two indistinct lateral metaventral foveae at middle; posterior margin with small and narrow split at middle.</p> <p>Legs elongate. Protrochanter (Fig. 66E) with short ventral spine, protibia (Fig. 66F) thickening apically, with triangular preapical denticle; mesotrochanter with (Fig. 66G) with ventral spine shorter than that of protrochanter; metatrochanter (Fig. 66H) with broad, curved ventral projection.</p> <p>Abdomen elongate, widest at lateral margins of tergite 1 (IV), length 0.98 mm, width 0.68 mm. Tergite 1 (IV) moderately longer than tergite 2 (V), lacking discal carinae, short basal sulcus embraced by small mediobasal foveae, with one pair of large basolateral foveae, with thin and short inner marginal carina and complete, thick outer marginal carinae; tergites 2–4 (V–VII) subequal in length, each with basal sulcus between small basolateral foveae, tergite 5 (VIII) (Fig. 66I) broad, semicircular. Sternite 2 (IV) as long as sternites 3–4 (V–VI) at middle, with one pair of mediobasal and three pairs of basolateral foveae, midlength of sternites 2–4 gradually shortened; sternites 3 and 4 (V and VI) each with three pairs of basolateral foveae, sternite 5 (VII) with one pair of basolateral foveae, sternite 6 (VIII) (Fig. 66J) transverse, posterior margin broadly impressed, sternite 7 (IX) (Fig. 66K) oval, membranous.</p> <p>Aedeagus (Fig. 66L–N) 0.58 mm long, strongly asymmetric; median lobe with large basal capsule and foramen, strongly constricted at base, ventral stalk strongly narrowed apically and strongly curved; dorsal lobe in dorsal view broad, plate-like; parameres reduced.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species differs from all known congeners by the elongate body, a darker color of the head, the characteristic structures of male antennomere 10 and 11, as well as the form of the aedeagus.</p> <p>Distribution. Zayü County, Tibet, SW China (Figs 86A, 100C).</p> <p>Etymology. The specific epithet refers to the elongate body of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FF5224ECCB88A2E271772313	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5024EACB88A4D177812183.text	C822F213FF5024EACB88A4D177812183.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites grandiceps Yin 2022	<div><p>Tribasodites grandiceps sp. nov.</p> <p>(Figs 67, 86A, 92A, B)</p> <p>Chinese common name: 巨Ħûḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, China: Xizang, Nyalam County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.96305&amp;materialsCitation.latitude=27.969723" title="Search Plazi for locations around (long 85.96305/lat 27.969723)">Zhangmu Town</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.96305&amp;materialsCitation.latitude=27.969723" title="Search Plazi for locations around (long 85.96305/lat 27.969723)">Youyi Bridge</a>, 27°58’11”N, 85°57’47”E, leaf litter, sifted, 1800 m, 29.vii.2010, Jian-Qing Zhu leg. (ƱẪỄŃ木县樟木W友谊 桥)’ (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.0 mm. Head sub-rounded at base, distinctly broader than pronotum, tempus moderately long, with rounded posterolateral margin, vertex with shallow reversed U-shaped impression, with moderately small foveae, with distinct mediobasal carina extending from head base anteriorly to level of anterior margin of eyes, with thin lateral carina from base to posterior margin of antennal tubercle; antenna moderately elongate, lacking modifications, antennomeres each distinctly elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of thin longitudinal discal carinae, with pair of small antebasal and tiny marginal spines. Elytra small, basally constricted; discal stria shallow, extending posteriorly to apical 1/2 of elytral length; disc finely punctate. Mesotibia with small preapical tubercle; metafemur expanded before base. Aedeagus elongate, asymmetric; median lobe with elongate basal capsule and round foramen, ventral stalk dilated at apex, dorsal lobe strongly curved at apex, parameres fused to broad membrane.</p> <p>Description. Male. Body (Fig. 67A) length 1.95 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 67B) sub-rounded at base, slightly broader than long, length 0.43 mm, width across eyes 0.47 mm; vertex smooth, with shallow reversed U-shaped impression, vertexal foveae (dorsal tentorial pits) asetose, moderately small, mediobasal carina distinct, extending from head base anteriorly to level of anterior margin of eyes, lateral carina thin; posterolateral margin rounded; frons anteriorly demarcated from clypeus by thin frontal-clypeal ridge, area between moderately raised antennal tubercles weakly impressed; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina thin, complete, branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with granulate surface; small gular foveae (posterior tentorial pits) in single oval pit, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes weakly prominent, composed of approximately 35 small ommatidia. Antenna moderately elongate, length 1.25 mm, indistinct club formed by slightly enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, antennomeres each markedly elongate, 8 shortest, 9 much wider and longer than 8, cylindrical, 10 wider but slightly shorter than 9, moderately broadened at apical 1/2, 11 largest, much longer than 9 and 10 combined, sub-conical, anterolateral margin slightly impressed.</p> <p>Pronotum (Fig. 67B) narrower than head, slightly longer than wide, length 0.45 mm, width 0.42 mm, widest approximately at middle; lateral margins rounded at middle, convergent apically and basally; disc slightly convex, finely punctate, thin median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and moderately short mediobasal carina, pair of thin discal longitudinal carinae (Fig. 67C) avoid additional spines, with thin lateral longitudinal sulci; with pairs of small antebasal and tiny marginal spines; lateral antebasal foveae small and asetose; with tiny outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, with small antebasal pit; margin of coxal cavity slightly carinate.</p> <p>Elytra slightly wider than long, length 0.56 mm, width 0.61 mm; each elytron with three moderately large, asetose basal foveae; discal stria extending posteriorly from outer basal fovea to half of elytral length; humerus weakly prominent, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae broadly forked internally, with short mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long seta at apex; mesotibia (Fig. 67D) with small preapical tubercle; metatrochanter (Fig. 67E) with long curved ventral protuberance, metafemur (Fig. 67F) broadly expanded at middle, with long setae along expansion.</p> <p>Abdomen constricted near base, widest at lateral margins of tergite 1 (IV), length 0.55 mm, width 0.64 mm. Tergite 1 (IV) longer than 2 and 3 (V and VI) combined, lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae and pair of thin discal carinae, complete, thin inner and thickened outer marginal carinae subparallel; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined, 2–4 each with one pair of small basolateral foveae and pair of thin marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin narrowly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral two pairs in large sockets, lateral carina thick; midlength of sternite 2 more than twice length of 3 (V), 3–5 (V–VII) subequal lin length, sternites 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, convex at middle, sternite 7 (IX) (Fig. 67G) composed of pair of triangular, membranous structures.</p> <p>Aedeagus (Fig. 67H–J) 0.34 mm long, asymmetric, elongate; median lobe with laterally extended transverse basal capsule and slightly oval foramen, basoventral projection strongly enlarged, ventral stalk broad at base and apex, laterally expanded before apex and then dilated apically; dorsal lobe fusing with median lobe at base, elongate, in dorsal view bent to left at apical 1/4, then greatly curved to right and narrowing apically, parameres fused, broad and flattened, membranous.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species is allied with a number of species centered on T. dilophus by the relatively long and rounded postocular margins, simple male antennae, and a generally elongate and slender aedeagus. It is most similar to T. prolixicornis in the lack of additional spines along the discal carinae of the pronotum, the mesally expanded metafemora of the male, and the presence of a small preapical tubercle on the male mesotibiae. These two species can be readily separated by the relatively much shorter antennae, the head being distinctly wider than the pronotum, the shorter elytra and less prominent eyes, and different forms and structures of the metafemoral expansion and the aedeagus of T. grandiceps. Tribasodites mirabilis also has modified male metafemora, but otherwise it is readily separable by the unique modification of the metatibiae and form of the aedeagus.</p> <p>Distribution. Nyalam County, Tibet, SW China (Figs 86A, 92A, B).</p> <p>Etymology. The specific epithet is a Latin adjective meaning ‘large-headed’, indicating this species has a relatively larger head among the species of Tribasodites.</p> </div>	http://treatment.plazi.org/id/C822F213FF5024EACB88A4D177812183	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF5524D7CB88A1DB71AA25B2.text	C822F213FF5524D7CB88A1DB71AA25B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites gyirong Yin 2022	<div><p>Tribasodites gyirong sp. nov.</p> <p>(Figs 68, 86B, 89)</p> <p>Chinese common name: 吉Pûḃà甲</p> <p>Type material (11 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.346115&amp;materialsCitation.latitude=28.359722" title="Search Plazi for locations around (long 85.346115/lat 28.359722)">Gyirong Valley</a>, pass nr. Jifu Vill., 28°21’35”N, 85°20’46”E, 2600–2700 m, 19.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW吉 ė村ṉữ山ě ’ (SNUC). PARATYPES: CHINA: 4 ♂♂, 6 ♀♀, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length over 2.6 mm. Head subtruncate at base, slightly narrower than pronotum, tempus moderately short, rounded at posterolateral angle, vertex with transverse impression at apex, with relatively large, asetose foveae, with distinct mediobasal carina extending from head base anteriorly beyond level of posterior margin of eyes, with distinct lateral carina from head base to posterior margin of antennal tubercle; antenna elongate, with modified antennomeres 10–11, antennomere 11 with distinct basal projection. Pronotum with laterally carinate median and lateral longitudinal sulci, with pair of discal carinae, with two pairs of antebasal and distinct marginal spines. Discal stria of elytron long, extending posteriorly to approximately apical 1/4 of elytral length; disc finely punctate. Mesotibia with large, blade-like apical projection. Aedeagus strongly asymmetric; median lobe with extended basal capsule and elongate foramen, ventral stalk broad at middle and then abruptly narrowing and protruding apically, with forked apex, dorsal lobe large, plate-like, with elongate sclerite narrowing towards apex. Female. Body length slightly over 2.5 mm, antenna shorter than male, antenna and legs lacking modifications, genitalia as in Fig. 68K.</p> <p>Description. Male. Body (Fig. 68A) length 2.68–2.78 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 68B) roundly at base, much wider than long, length 0.52 mm, width across eyes 0.60 mm; vertex smooth, with transverse impression at apex, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina distinct, extending from head base anteriorly to slightly beyond level of posterior margin of eyes, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral margin rounded; frons weakly impressed between large, moderately raised antennal tubercles, anteriorly demarcated from clypeus by oblique frontal-clypeal ridge; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 57 ommatidia. Antenna elongate, length 1.49–1.52 mm, distinct club (Fig. 68C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, slightly elongate, 9 much wider than 8, transverse, 10 (Fig. 68D) broader and longer than 9, ventral surface with large basal impression, 11 largest, longer than 9 and 10 combined, with distinct basoventral projection, subbase of projection with round tubercle, apex with tufted setae.</p> <p>Pronotum (Fig. 68B) slightly wider than long, length 0.62 mm, width 0.63–0.64 mm, widest at middle; lateral margins rounded and with small denticles at apical 1/2, convergent basally and sub-parallel at basal 1/4; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of discal carinae and lateral longitudinal sulci; with outer pair of antebasal spines and inner pair of angulate tubercles, marginal spines distinct; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thick hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity broadly carinate.</p> <p>Elytra much wider than long, length 0.77–0.78 mm, width 0.96–0.97 mm; each elytron with three moderately large, asetose basal foveae; long discal stria extending posteriorly from outer basal fovea to approximately apical 1/4 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae moving medially and in large, setose shared impression; posterior margin roundly emarginate at middle, with narrow split.</p> <p>Legs elongate; procoxa with exceptionally long seta at base, protibia with tiny preapical denticle at mesal margin; mesotrochanter (Fig. 68E) with distinct ventral spine, mesotibia (Fig. 68F) with large blade-like projection before apex; metatrochanter (Fig. 68G) with long ventral projection strongly curved at apex, metatibia with long pencil-like apical tuft of setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.86–0.92 mm, width 0.87–0.90 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with large mediobasal and one pair of basolateral foveae, with pair of short discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, with long, oblique lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, 3–5 each with two pairs of small basolateral foveae and short lateral carinae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 68H) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 68I, J) 0.63 mm long, strongly asymmetric, elongate; median lobe with large extended basal capsule and elongate foramen, ventral stalk in dorsal view broad at middle and abruptly narrowing apically, with deeply forked, weakly sclerotized apex; dorsal lobe large, plate-like, apical part protruding and curved, in lateral view narrowing towards apex; parameres membranous, apically split into two parts.</p> <p>Female. Similar to male in external morphology; antenna shorter than that for male; antenna and legs lacking modifications; each compound eye composed of approximately 35 ommatidia. Measurements (as for male): body length 2.52–2.53 mm; length/width of head 0.52/ 0.60 mm, pronotum 0.54–0.55/ 0.59–0.60 mm, elytra 0.69– 0.70/ 0.88 mm; abdomen 0.80–0.86/ 0.81 mm; length of antenna 1.15–1.23 mm; genitalia weakly sclerotized (Fig. 68K), with elongate, forked lateral sclerites, maximum width 0.41 mm.</p> <p>Comparative notes. Tribasodites gyirong is placed in the group of species centered on T. antennalis, which share a similar location of the male antennal modifications and the similar aedeagal forms. The male of T. gyirong is morphologically most similar to the sympatric T. pugiunculus by sharing most external characters. It can be readily separated from T. pugiunculus by the larger body size (2.68–2.78 mm vs. 2.30–2.41 mm), presence of a transverse impression on the vertex (lacking in T. pugiunculus), the relatively longer basal projections of antennomeres 11, and the broadly and deeply split apex of the ventral stalk of the aedeagus.</p> <p>Distribution. Gyirong County, Tibet, SW China (Figs 86B, 89).</p> <p>Etymology. This species is named after Gyirong County, where the type locality is situated.</p></div> 	http://treatment.plazi.org/id/C822F213FF5524D7CB88A1DB71AA25B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6B24D4CB88A57F719227F2.text	C822F213FF6B24D4CB88A57F719227F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites kiypu Yin 2022	<div><p>Tribasodites kiypu sp. nov.</p> <p>(Figs 69, 86B, 88, 89)</p> <p>Chinese common name: 吉âûḃà甲</p> <p>Type material (9 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.346115&amp;materialsCitation.latitude=28.359722" title="Search Plazi for locations around (long 85.346115/lat 28.359722)">Gyirong Valley</a>, pass nr. Jifu Vill., 28°21’35”N, 85°20’46”E, 2600–2700 m, 19.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉PW吉 ė村ṉữ山ě ’ (SNUC). PARATYPE: CHINA: 4 ♂♂, 4 ♀♀, same data as that of holotype (SNUC).</p> <p>Material not included in type series (5 exx.). 3 ♂♂, 1 ♀, same label data as of type series (SNUC); 1 ♂, same collectors and also from <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.263885&amp;materialsCitation.latitude=28.429722" title="Search Plazi for locations around (long 85.263885/lat 28.429722)">Gyirong Valley</a>, except ‘pass nr. Ru Kupuqiong, 28°25’47”N, 85°15’50”E, 3000 m, 20.vi.2021, ƱẪU喀NJ吉PWả库âAEṉữ ’ (SNUC).</p> <p>Diagnosis (based on type series). Male. Body length over 2.8 mm. Head sub-rounded at base, slightly narrower than pronotum, tempus long, with rounded posterolateral margin, vertex lacking sulcus, with relatively large, asetose foveae, with distinct mediobasal carina extending from head base anteriorly to slightly posterior to level of anterior margin of eyes, with distinct lateral carina from base to posterior margin of antennal tubercle; antenna elongate, lacking modifications, antennomeres slightly elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of longitudinal discal carinae composed of spines, with pair of large antebasal and distinct marginal spines. Discal stria of elytron shallow, extending posteriorly to pass half of elytral length; disc finely punctate. Mesotrochanter with blunt ventral spine, mesotibia with elongate spine at apex; metatrochanter with curved projection on ventral margin. Aedeagus elongate, asymmetric; median lobe with large basal capsule and broad foramen, ventral stalk abruptly narrowed and expanded at apex, dorsal lobe broad and flat, paramere membranous. Female. Body length slightly over 2.7 mm, antenna shorter than male, legs lacking spines or projections, genitalia as in Fig. 69H.</p> <p>Description (based on type series). Male. Body (Fig. 69A) length 2.82 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 69B) sub-rounded at base, slightly longer than wide, length 0.60–0.62 mm, width across eyes 0.58– 0.59 mm; vertex with rough surface, lacking sulcus, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina distinct, extending from head base anteriorly to slightly posterior to level of anterior margin of eyes, with pair of sinuate admesal carinae forming inner margins of reversed U-shaped impression that connect vertexal foveae, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral margin rounded; frons anteriorly demarcated from clypeus by oblique frontal-clypeal ridge, which extended anteriorly for short distance on clypeus, area between moderately raised antennal tubercles weakly impressed; clypeus with rough surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with granulate surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, weak median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 58 ommatidia. Antenna elongate, length 1.50–1.60 mm, indistinct club formed by slightly enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each slightly elongate, 8 smallest, 9 wider and longer than 8, subconical, 10 shorter than 9, subconical, 11 largest, slightly longer than 9 and 10 combined, subfusiform, impressed dorsally near apex.</p> <p>Pronotum (Fig. 69B) slightly wider than long, length 0.61–0.62 mm, width 0.62–0.64 mm, widest at middle; lateral margins rounded at middle, convergent apically and sub-parallel at basal 1/4; disc slightly convex, finely punctate, median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and short mediobasal carina, pair of discal longitudinal carinae (Fig. 69C) each composed of four to five distinct spines, with thin lateral longitudinal sulci; with pairs of large antebasal and smaller marginal spines; lateral antebasal foveae small and setose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, lateral antebasal hypomeral pit absent; margin of coxal cavity slightly carinate.</p> <p>Elytra much wider than long, length 0.77–0.80 mm, width 0.93–0.94 mm; each elytron with three moderately large, asetose basal foveae; discal stria extending posteriorly from outer basal fovea to pass half of elytral length; humerus prominent, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae broadly separated, originating from shared transverse opening, large lateral mesoventral foveae forked internally, with short, thickened mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long seta at base; mesotrochanter (Fig. 69D) with blunt ventral spine, mesotibia with elongate triangular apical spine; metatrochanter (Fig. 69E) with long, curved ventral protuberance.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.84–0.93 mm, width 0.84–0.86 mm. Tergite 1 (IV) approximately 1.3 times as long as 2 (V), lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, with pair of thin, sub-triangular discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 2 and 3 each with two pairs of small basolateral foveae and thin marginal carinae, 4 (VII) shorter than tergites 2 and 3 combined, with one pair of small basolateral foveae and pair of thin marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with broad setose basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, lacking lateral carina; midlength of sternites 2–5 (IV–VII) gradually shorter, sternites 3–4 each with broad basal sulcus and three pairs of basolateral foveae, 5 lacking basal sulcus, with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 69F, G) composed of pair of triangular, membranous structures.</p> <p>Aedeagus (Fig. 69F, G) 0.39 mm long, asymmetric, elongate; median lobe with broad basal capsule and large foramen, ventral stalk abruptly narrowed and strongly dilated at apex; dorsal lobe broad and flattened, parameres membranous, apical part weakly sclerotized.</p> <p>Female. Similar to male in external morphology; antenna shorter than male; legs lacking spines and projections; each compound eye composed of approximately 30 ommatidia. Measurements (as for male): body length 2.77–2.80 mm; length/width of head 0.60–0.61/ 0.57–0.58 mm, pronotum 0.61–0.62/ 0.61–0.63 mm, elytra 0.72–0.77/ 0.91– 0.92 mm; abdomen 0.87–0.88/ 0.87–0.88 mm; length of antenna 1.43–1.47 mm; genitalia weakly sclerotized (Fig. 69H), with broad lateral arms, maximum width 0.31 mm.</p> <p>Comparative notes. This species is allied with T. dilophus, T. mirabilis and T. yatung, all described in this paper, in sharing relatively elongate habitus and postocular margins, and having simple male antennae. The male of Tribasodites kiypu can be readily separated from the other three species by the much larger body size (2.82 mm (2.52–2.65 mm for none-type specimens) vs. 2.14–2.15 mm for T. dilophus, 2.06 mm for T. mirabilis and 1.97–2.13 mm for T. yatung), and by the unique form of the aedeagus; from T. mirabilis also by the simple hind legs, which are strongly modified in the latter species.</p> <p>Distribution. Gyirong County, Tibet, SW China (Figs 69, 86B, 88, 89).</p> <p>Etymology. The specific epithet refers to Kiypu (= Ji Fu) Village, type locality of this species.</p> <p>Comments on none-type material. Three males and one female from the type locality, and one male from a nearby collecting site have the aedeagi identical to that of the holotype. They differ from the type series by the smaller body size, a slightly lighter body color, a smoother surface of the head dorsum, the U-shaped sulcus of the vertex not being ridged at its inner margins, the lack of an oblique frontal-clypeal ridge, and a thinner and shallower median sulcus of the pronotum. Note is that the pronotal spines, and spines/projections of the legs are, however, of same shapes as the type series. Such differences among the population from a same collecting site is not commonly detected for batrisines, these differences are temporarily considered as intraspecific variation within T. kiypu.</p> <p>The measurements of these ‘smaller-form’ individuals are given here for comparative purposes: 3 males from the type locality: each compound eye composed of approximately 45 ommatidia; body length 2.52–2.53 mm, length/ width of head 0.53–0.55/ 0.54 mm, pronotum 0.57–0.59/ 0.57–0.58 mm, elytra 0.76–0.77/ 0.88 mm; abdomen 0.73– 0.75/ 0.77 mm; length of antenna 1.41–1.43 mm; 1 female from the type locality: each compound eye composed of approximately 25 ommatidia; body length 2.56 mm, length/width of head 0.54/ 0.53 mm, pronotum 0.56/ 0.55 mm, elytra 0.70/ 0.81 mm; abdomen 0.79/ 0.75 mm; length of antenna 1.26 mm; 1 male from Ru Kupuqiong: each compound eye composed of approximately 48 ommatidia; body length 2.65 mm, length/width of head 0.55/ 0.54 mm, pronotum 0.58/ 0.59 mm, elytra 0.76/ 0.89 mm; abdomen 0.74/ 0.80 mm; length of antenna 1.43 mm.</p></div> 	http://treatment.plazi.org/id/C822F213FF6B24D4CB88A57F719227F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6824D2CB88A0B577A225DE.text	C822F213FF6824D2CB88A0B577A225DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites mirabilis Yin 2022	<div><p>Tribasodites mirabilis sp. nov.</p> <p>(Figs 70, 86B, 93E)</p> <p>Chinese common name: OiAEûḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Dinggyê County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.378334&amp;materialsCitation.latitude=27.903055" title="Search Plazi for locations around (long 87.378334/lat 27.903055)">Zhêntang Town</a>, Xiuxiongma Vill., 27°54’11”N, 87°22’42”E, 2700–3000 m, 25.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJŤĠ县Ŀ ƋWšŏ玛村 ’ (SNUC).</p> <p>Diagnosis. Male. Body length 2.06 mm. Head sub-rounded at base, approximately as wide as pronotum, tempus moderately long, with rounded posterolateral margin, vertex with shallow reversed U-shaped impression, with moderately large, setose foveae, with distinct mediobasal carina extending from head base anteriorly to before level of anterior margin of eyes, with thin lateral carina from base to posterior margin of antennal tubercle; antenna elongate, lacking modifications, antennomeres slightly elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of longitudinal discal carinae composed of spines, with pair of distinct antebasal and small marginal spines. Discal stria of elytron shallow, extending posteriorly before elytral midlength; disc finely punctate. Mesotibia with tiny preapical tubercle; metatrochanter with small protuberance on ventral margin, metafemur expanded before apex, metatibia with tuft of curved setae at apical 1/3. Aedeagus elongate, asymmetric; median lobe with large basal capsule and foramen, ventral stalk and dorsal lobe elongate, parameres broad, membranous.</p> <p>Description. Male. Body (Fig. 70A) length 2.06 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 70B) sub-rounded at base, as long as broad, length 0.45 mm, width across eyes 0.46 mm; vertex smooth, with shallow reversed U-shaped impression, vertexal foveae (dorsal tentorial pits) asetose, moderately large, mediobasal carina distinct, extending from head base anteriorly to slightly beyond level of anterior margin of eyes, lateral carina thin, shortly present at base; posterolateral margin rounded; frons anteriorly demarcated from clypeus by oblique frontal-clypeal ridge, area between moderately raised antennal tubercles weakly impressed; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, shortly branched below eye. Venter with smooth surface; small gular foveae (posterior tentorial pits) in single oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes small, each composed of approximately 14 ommatidia. Antenna elongate, length 1.08 mm, indistinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 shortest, sub-moniliform, 9 much wider and longer than 8, subconical, 10 wider than 9, subconical, 11 largest, slightly longer than 9 and 10 combined, sub-conical, anteromesal margin impressed.</p> <p>Pronotum (Fig. 70B) approximately as long as wide, length 0.46 mm, width 0.45 mm, widest at middle; lateral margins rounded at middle, convergent apically and basally; disc slightly convex, finely punctate, thin median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and short mediobasal carina, pair of discal longitudinal carinae (Fig. 70C) each composed of four distinct spines, with thin lateral longitudinal sulci; with pairs of distinct antebasal and small marginal spines; lateral antebasal foveae distinct and setose; with tiny outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity slightly carinate.</p> <p>Elytra much wider than long, length 0.56 mm, width 0.69 mm; each elytron with three moderately large, asetose basal foveae; discal stria extending posteriorly from outer basal fovea to before elytral midlength; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae forked internally, with short mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long setae at apex; mesotibia with tiny preapical tubercle; metatrochanter (Fig. 70D) with short curved ventral protuberance, metafemur (Fig. 70E) angularly expanded at apical 1/3, metatibia (Fig. 70F) concave at apical 2/5 of lateral margin, filled with tuft of long, curved setae.</p> <p>Abdomen constricted at base, widest at lateral margins of tergite 1 (IV), length 0.57 mm, width 0.64 mm. Tergite 1 (IV) approximately as long as 2 and 3 (V and VI) combined, lacking basal sulcus, with one mediobasal and two pairs of basolateral foveae, with complete thin inner and thickened outer marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined, 2–4 each with one pair of small basolateral foveae and pair of thin marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin angularly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, lateral carina absent; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, sternites 3 and 4 each with three pairs and 5 with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, convex at middle, sternite 7 (IX) (Fig. 70G) composed of pair of triangular, membranous structures.</p> <p>Aedeagus (Fig. 70H–J) 0.37 mm long, asymmetric, elongate; median lobe with broad basal capsule and large foramen, ventral stalk markedly expanded basolaterally and narrowing apically; dorsal lobe also broad at base and narrowing towards membranous apex, parameres fused, broad and flattened, membranous.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species is allied with T. dilophus, T. grandiceps, T. kiypu, T. prolixicornis, and T. yatung by the long, rounded postocular margins, the simple and elongate male antennae, and an elongate and slender aedeagus (except for T. dilophus, which has a much broader aedeagus). The male of T. mirabilis can be readily separated from that of these congeners by the strongly modified tibiae alone, as well as by the unique form of the aedeagus.</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 86B, 93E).</p> <p>Etymology. The specific epithet ‘ mirabilis (wonderful, astonishing)’ is a Latin adjective referring to the remarkably modified male hind legs of this species.</p> </div>	http://treatment.plazi.org/id/C822F213FF6824D2CB88A0B577A225DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6E24D0CB88A29E7649239E.text	C822F213FF6E24D0CB88A29E7649239E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites prolixicornis Yin 2022	<div><p>Tribasodites prolixicornis sp. nov.</p> <p>(Figs 71, 86C, 93F, G)</p> <p>Chinese common name: ḲƟûḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, China: Xizang, Dinggyê County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Zhêntang Town</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.40667&amp;materialsCitation.latitude=27.86389" title="Search Plazi for locations around (long 87.40667/lat 27.86389)">Ganma Zangbo Valley</a>, 27°51’50”N, 87°24’24”E, 2400 m, 28.vi.2021, Z. Peng leg., ƱẪŤĠ县ĿƋWH玛Ẫ布河ě ’ (SNUC).</p> <p>Diagnosis. Male. Body length slightly over 2.1 mm. Head sub-rounded at base, approximately as wide as pronotum, tempus moderately long, with rounded posterolateral margin, vertex with shallow reversed U-shaped impression, with small, indistinct foveae, with distinct mediobasal carina extending from head base anteriorly to level of anterior margin of eyes, with thin, faint lateral carina from base to posterior margin of antennal tubercle; antenna conspicuously elongate, lacking modifications, antennomeres each distinctly elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of longitudinal discal carinae, with pair of small antebasal and marginal spines. Discal stria of elytron shallow, extending posteriorly to apical 1/3 of elytral length; disc finely punctate. Mesotibia with small preapical tubercle; metafemur expanded before base, metatibia broadly and strongly sinuate along mesal margin. Aedeagus elongate, slightly asymmetric; median lobe with broad basal capsule and foramen, ventral stalk fusing with dorsal lobe at latter’s base, parameres broad, membranous.</p> <p>Description. Male. Body (Fig. 71A) length 2.16 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 71B) sub-rounded at base, as long as broad, length 0.49 mm, width across eyes 0.48 mm; vertex smooth, with shallow reversed U-shaped impression, vertexal foveae (dorsal tentorial pits) asetose, small, mediobasal carina distinct, extending from head base anteriorly to level of anterior margin of eyes, lateral carina thin, faint; posterolateral margin rounded; frons anteriorly demarcated from clypeus by faint frontal-clypeal ridge, area between moderately raised antennal tubercles weakly impressed; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with granulate surface; small gular foveae (posterior tentorial pits) in single oval pit, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 45 ommatidia. Antenna elongate, length 1.58 mm, indistinct club formed by slightly enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, antennomeres each markedly elongate, 8 shortest, 9 much wider and longer than 8, cylindrical, 10 slightly wider than 9, broadened at apical 1/2, 11 largest, slightly shorter than 9 and 10 combined, sub-conical, anterolateral margin impressed.</p> <p>Pronotum (Fig. 71B) approximately as long as wide, length 0.48 mm, width 0.46 mm, widest at middle; lateral margins rounded at middle, convergent apically and basally; disc slightly convex, finely punctate, thin median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and short mediobasal carina, pair of discal longitudinal carinae (Fig. 71C) avoid additional spines, with thin lateral longitudinal sulci; with pairs of small antebasal and marginal spines; lateral antebasal foveae small and asetose; with tiny outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, with shallow antebasal impression; margin of coxal cavity slightly carinate.</p> <p>Elytra slightly wider than long, length 0.68 mm, width 0.70 mm; each elytron with three moderately large, asetose basal foveae; discal stria extending posteriorly from outer basal fovea to apical 1/3 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae forked internally, with short mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long seta at apex; mesotibia (Fig. 71D) with small preapical tubercle; metatrochanter (Fig. 71E) with long curved ventral protuberance with long setae near base and setose apex, metafemur (Fig. 71F) strongly and broadly sinuate at mesal margin, covered with setae that are longer than those of pro- and mesofemur.</p> <p>Abdomen constricted near base, widest at lateral margins of tergite 1 (IV), length 0.59 mm, width 0.63 mm. Tergite 1 (IV) slightly longer than 2 and 3 (V and VI) combined, lacking basal sulcus, with one pair of mediobasal and two pairs of basolateral foveae, pair of thin and short discal carinae and complete, thin inner and thickened outer marginal carinae; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined, 2–4 each with one pair of small basolateral foveae and pair of thin marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin narrowly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and three pairs of basolateral foveae, lateral two pairs in large sockets, lateral carina absent; midlength of sternite 2 more than twice length of 3 (V), 3 as long as 4 (VI), 5 (VII) slightly longer than 4, sternites 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, convex at middle, sternite 7 (IX) (Fig. 71G) composed of pair of triangular, membranous structures.</p> <p>Aedeagus (Fig. 71H–J) 0.38 mm long, slightly asymmetric, elongate; median lobe with broad basal capsule and large foramen, basoventral projection long and enlarged, ventral stalk broad at base and apex, narrowed at middle; dorsal lobe fusing with median lobe at base, elongate, moderately curved, moderately expanded apically and with rounded apex, parameres fused, broad and flattened, membranous.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species is allied with a group of species centered on T. dilophus and appears to be morphologically most similar to T. grandiceps in the lack of additional spines along the discal carinae of the pronotum, the presence of a preapical tubercle on the male mesotibiae, and the expanded male metafemora. Tribasodites prolixicornis can be readily separated from T. grandiceps by the relatively much longer antennae, the head being only slightly wider than pronotum and with more strongly prominent eyes, the much longer elytra, the different form of the male metafemoral expansion, and the dorsal lobe of the aedeagus only slightly curved at the apical portion (strongly twisted in T. grandiceps).</p> <p>Distribution. Dinggyê County, Tibet, SW China (Figs 86C, 93F, G).</p> <p>Etymology. The specific name is combined from ‘ prolixus (stretched out, extended)’ and ‘ cornis (horned)’, referring to the markedly elongate antennae of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FF6E24D0CB88A29E7649239E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6C24DDCB88A75E70372492.text	C822F213FF6C24DDCB88A75E70372492.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites pugiunculus Yin 2022	<div><p>Tribasodites pugiunculus sp. nov.</p> <p>(Figs 72, 86C, 89, 92C)</p> <p>Chinese common name: 匕ḇûḃà甲</p> <p>Type material (18 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Gyirong County, Gyirong Valley, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.346115&amp;materialsCitation.latitude=28.359722" title="Search Plazi for locations around (long 85.346115/lat 28.359722)">Jifu Vill.</a>, 28°21’35”N, 85°20’46”E, 2600–2700 m, 19.vi.2021, Z. Peng, Z. Yin &amp; W. Zhang leg., ƱẪU喀NJ吉P W吉ė村ṉữ山ě ’ (SNUC). PARATYPES: CHINA: 10 ♂♂, 6 ♀♀, same data as that of holotype; 1 ♂, ‘ China: Xizang A. R., Nyalam County, Zhangmu Town, pass nr. <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=85.97611&amp;materialsCitation.latitude=27.956112" title="Search Plazi for locations around (long 85.97611/lat 27.956112)">Lixin Village</a> (ca. 27°57’22”N, 85°58’34”E), 2600 m, 24.vii.2010, W.-X. Bi leg. [ƱẪỄŃ木县樟木WĒẾ村]’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.3–2.4 mm. Head sub-rectangular at base, approximately as wide as pronotum, tempus moderately short, rounded at posterolateral angle, vertex lacking sulcus, with relatively large, asetose foveae, with distinct mediobasal carina extending from head base anteriorly slightly beyond level of posterior margin of eyes, with distinct lateral carina from base to posterior margin of antennal tubercle; antenna elongate, with modified antennomeres 10–11, antennomere 11 with short basal projection. Pronotum with laterally carinate median and lateral longitudinal sulci, with pair of discal carinae, with two pairs of antebasal and distinct marginal spines. Discal stria of elytron long, extending posteriorly to more than apical 1/4 of elytral length; disc finely punctate. Mesotibia with large, blade-like apical projection. Aedeagus strongly asymmetric; median lobe with extended basal capsule and elongate foramen, ventral stalk broad at middle and then abruptly narrowing and protruding apically, dorsal lobe large, plate-like, with elongate sclerite narrowing towards apex. Female. Body length slightly over 2.2 mm, antenna shorter than male, antenna and legs lacking modifications, genitalia as in Fig. 72K.</p> <p>Description. Male. Body (Fig. 72A) length 2.30–2.41 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 72B) sub-rectangular at base, much wider than long, length 0.45 mm, width across eyes 0.53–0.55 mm; vertex smooth, lacking sulcus, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina distinct, extending from head base anteriorly to slightly beyond level of posterior margin of eyes, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral margin rounded; frons weakly impressed between large, moderately raised antennal tubercles, anteriorly demarcated from clypeus by frontalclypeal ridge; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 40–50 ommatidia. Antenna elongate, length 1.25–1.29 mm, distinct club (Fig. 72C) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, 8 smallest, moniliform, 9 much wider than 8, transverse, 10 (Fig. 72D) much broader and longer than 9, ventral surface occupied by large mesal impression, 11 largest, much longer than 9 and 10 combined, with short, subtriangular and slightly curved projection at base of mesal margin, base of projection with tiny semicircular tubercle.</p> <p>Pronotum (Fig. 72B) slightly wider than long, length 0.53 mm, width 0.54–0.57 mm, widest at middle; lateral margins rounded and with small denticles at apical 1/2, convergent basally and parallel at basal 1/4; disc slightly convex, finely punctate, broad median longitudinal sulcus with carinate sides, posteriorly confluent with oval antebasal impression and short mediobasal carina, with pair of discal carinae and lateral longitudinal sulci; with two pairs of antebasal and distinct marginal spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; thick hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity broadly carinate.</p> <p>Elytra much wider than long, length 0.67–0.71 mm, width 0.83–0.85 mm; each elytron with three moderately large, asetose basal foveae; long discal stria extending posteriorly from outer basal fovea to more than apical 1/4 of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared opening, large lateral mesoventral foveae not forked internally, with short mesoventral process, with complete marginal stria. Metaventrite prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, two lateral metaventral foveae moving medially and in large, setose shared impression; posterior margin roundly emarginate at middle, lacking split.</p> <p>Legs elongate; procoxa with exceptionally long seta at base, protibia with small preapical denticle at mesal margin; mesotrochanter (Fig. 72E) with distinct ventral spine, mesotibia (Fig. 72F) with broad, blade-like projection before apex; metatrochanter (Fig. 72G) with long ventral projection slightly to strongly curved at apex, metatibia with long pencil-like apical tuft of setae.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.72–0.74 mm, width 0.74–0.78 mm. Tergite 1 (IV) more than twice as long as 2 (V), lacking basal sulcus, with large mediobasal and one pair of basolateral foveae, with pair of short discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, with long, oblique lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, 3–5 each with two pairs of small basolateral foveae and short lateral carinae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 72H) weakly sclerotized, slightly oval.</p> <p>Aedeagus (Fig. 72I, J) 0.42–0.50 mm long, strongly asymmetric, elongate; median lobe with large extended basal capsule and elongate foramen, ventral stalk in dorsal view broad at middle and abruptly narrowing apically, with expanded, weakly sclerotized apex; dorsal lobe large, plate-like, apical part protruding and curved, narrowing towards apex; parameres membranous, apically split into two lobes.</p> <p>Female. Similar to male in external morphology; antenna approximately as long as that of male; antenna and legs lacking modifications; each compound eye composed of approximately 50 ommatidia. Measurements (as for male): body length 2.23–2.30 mm; length/width of head 0.43–0.44/ 0.53–0.55 mm, pronotum 0.50–0.51/ 0.54 mm, elytra 0.63–0.66/ 0.81–0.83 mm; abdomen 0.74/ 0.74–0.75 mm; length of antenna 1.12–1.27 mm; genitalia weakly sclerotized (Fig. 72K), maximum width 0.32 mm.</p> <p>Comparative notes. Tribasodites pugiunculus belongs to the group of species centered on T. antennalis which share similar locations of the male antennal modifications and general form of the aedeagus. The male of T. pugiunculus can be separated from the other species by the large blade-like projections near the apices of the mesotibia, where both T. antennalis and T. bari possess much smaller spines at the same position. Tribasodites pugiunculus further differs from both species by the short, apically narrowed basal projections of antennomeres 11, while the projections are much larger and broader in T. antennalis and much more greatly prolonged for in T. bari. The distinctive form of the aedeagus also leads to a confident identification of T. pugiunculus.</p> <p>Distribution. Gyirong and Nyalam County, Tibet, SW China (Figs 86C, 89, 92C).</p> <p>Etymology. The new species epithet ‘ pugiunculus (a small dagger)’ is a Latin noun referring to the blade-like apical projections on the male mesotibiae.</p> <p>Remarks. Parasitic Laboulbeniales were found on tergites 2 and 3 (IV and V) of the Nyalam specimen.</p></div> 	http://treatment.plazi.org/id/C822F213FF6C24DDCB88A75E70372492	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6124DBCB88A25571D9226A.text	C822F213FF6124DBCB88A25571D9226A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites vertexalis Yin 2022	<div><p>Tribasodites vertexalis sp. nov.</p> <p>(Figs 73, 86C, 100B)</p> <p>Chinese common name: ḔĦûḃà甲</p> <p>Type material (1 ex.). HOLOTYPE: CHINA: ♂, ‘ <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.46618&amp;materialsCitation.latitude=29.583559" title="Search Plazi for locations around (long 95.46618/lat 29.583559)">China</a>: Xizang, Mêdog County, 96K, 1460 m, 29°35’0.81”N, 95°27’58.26”E, 17.iii.2017, X.-B. Song leg. [ƱẪẸṘ 96K]’ (SNUC).</p> <p>Diagnosis. Male. Body length 2.40 mm. Head rounded triangular, as broad as pronotum, tempus moderately long, vertex and frons strongly modified, each with large projection at middle; antenna lacking modifications. Pronotum with laterally carinate median and lateral longitudinal sulci, discal carinae composed row of two small discal and one large antebasal spine; lateral margin denticulate at anterior half, with distinct marginal spine. Discal stria of elytron shallow and long, extending posteriorly to apical 1/4 of elytral length; disc finely punctate. Fore legs simple; mesotrochanter with apically curved ventral spine, mesotibia with distinct apical spine; metatrochanter with broad, curved ventral projection. Aedeagus strongly asymmetric, median lobe constricted at base, ventral stalk strongly broad at middle, dorsal lobe weakly sclerotized, parameres flattened, membranous.</p> <p>Description. Male. Body (Fig. 73A) length 2.40 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 73B) rounded triangular, much wider than long, length 0.48 mm, width across eyes 0.59 mm; vertex with large projection at middle, area anterior and lateral projection impressed and with pair of nude foveae, with tufts of setae at anterolateral margins of impression, postocular area with oval patch covered by dense, short setae; frons anteriorly demarcated from clypeus by complete frontal-clypeal ridge, which merged at middle and extending anteriorly to apex of clypeus, with strongly raised tubercle covered by long posteriorly pointed setae at middle; clypeus with smooth surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with rough surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 55 ommatidia. Antenna lacking modification, length 1.26 mm, club (Fig. 73C) loosely formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, sub-moniliform, 8 smallest, 9 much larger than 8, elongate, 10 slightly broader and as long as 9, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform, with indistinct line separating apical 2/5 from basal 3/5.</p> <p>Pronotum (Fig. 73B) slightly wider than long, length 0.53 mm, width 0.58 mm, widest at middle; lateral margins rounded at middle, convergent apically and parallel at basal 1/4; disc moderately convex, finely punctate, median longitudinal sulcus with carinate sides, followed posteriorly by oval impression and short mediobasal carina, with pair of mesally carinate lateral longitudinal sulci, discal carinae composed of row of two distinct discal and one large antebasal spine; lateral margin with large marginal and five to six smaller spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge complete, extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity broadly carinate.</p> <p>Elytra wider than long, length 0.73 mm, width 0.97 mm, widest at middle, narrowing form middle towards apex; each elytron with three large, asetose basal foveae; discal stria shallow and long, extending posteriorly from outer basal fovea to apical 1/4 of elytral length; humerus broadly prominent, with small denticle, subhumeral fovea small, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae relatively broadly separated, large lateral mesoventral foveae not forked internally, with short mesoventral process. Metaventrite slightly prominent admesally, impressed at middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina of meso- and metaventrite complete.</p> <p>Legs moderately elongate; protrochanter with exceptionally long seta on ventral margin; mesotrochanter with apically curved ventral spine (Fig. 73D), mesotibia (Fig. 73E) with triangular apical spine; metatrochanter (Fig. 73F) with broad, curved projection on ventral margin.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.63 mm, width 0.88 mm. Tergite 1 (IV) less than twice as long as 2 (V), basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with pair of long discal carinae, inner marginal carina hardly oblique, outer one thicker, both complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, 4 with small marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–4 each with two pairs of small basolateral foveae and small tubercle at middle, 5 with one pair of basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) composed of several membranous structures.</p> <p>Aedeagus (Fig. 73G–I) 0.35 mm long, strongly asymmetric; median lobe constricted at base, with rounded triangular foramen, ventral stalk strongly broad at middle, flattened apically and narrowing towards apex, dorsal lobe weakly sclerotized, extended to right side (in ventral view), parameres membranous, broad and plate-like.</p> <p>Female. Unknown.</p> <p>Comparative notes. Several different forms of aedeagi have been found within the possibly heterogenous Tribasodites. This species possesses an aedeagus of a form typical for the genera Hingstoniella, Besuchetaceus Yin &amp; Li, and Sinotrisus Yin &amp; Li, creating concerns about the relationships between these groups. At this moment no discrete character can be found to support a placement of this species elsewhere other than in Tribasodites. Among the Tibetan species, the habitus of T. vertexalis is relatively much stouter than the others, the male vertex is strongly modified with a large central excavation, and has lateral rounded areas covered by dense short setae. These characters in combination with the simple male antennae, the discal carina of the pronotum composed of three large spines, and the triangular apical spines of the mesotibiae readily separates this species from the others.</p> <p>Distribution. Mêdog County, Tibet, SW China (Figs 86C, 100B).</p> <p>Etymology. The species epithet refers to the strongly modified male vertex of this species.</p></div> 	http://treatment.plazi.org/id/C822F213FF6124DBCB88A25571D9226A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6724D9CB88A5CA71D120BE.text	C822F213FF6724D9CB88A5CA71D120BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tribasodites yatung Yin 2022	<div><p>Tribasodites yatung sp. nov.</p> <p>(Figs 74, 86C, 94A)</p> <p>Chinese common name: 亚东ûḃà甲</p> <p>Type material (2 exx.). HOLOTYPE: CHINA: ♂, ‘China: Xizang, Yatung County, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.94556&amp;materialsCitation.latitude=27.432499" title="Search Plazi for locations around (long 88.94556/lat 27.432499)">Xiayadong Township</a>, 27°25’57”N, 88°56’44”E, 3400 m, 18.viii.2010, Jian-Qing Zhu leg. [ƱẪ亚东县T亚东乡]’ (SNUC). PARATYPE: CHINA: 1 ♂, same data as that of holotype (SNUC).</p> <p>Diagnosis. Male. Body length approximately 2.0 mm. Head sub-rounded at base, approximately as wide as pronotum, tempus moderately long, with rounded posterolateral margin, vertex lacking sulcus, with relatively large, asetose foveae, with distinct mediobasal carina extending from head base anteriorly to level of anterior margin of eyes, with distinct lateral carina from base to posterior margin of antennal tubercle; antenna elongate, lacking modifications, antennomeres slightly elongate. Pronotum with laterally carinate median and shallow lateral longitudinal sulci, with pair of longitudinal discal carinae composed of spines, with pair of large antebasal and small marginal spines. Discal stria of elytron shallow, extending posteriorly to pass half of elytral length; disc finely punctate. Mesotrochanter with distinct ventral spine, mesotibia with triangular projection at apex; metatrochanter with long protuberance on ventral margin. Aedeagus elongate, asymmetric; median lobe with large basal capsule and broad foramen, ventral stalk and dorsal lobe extremely elongate, parameres broad, membranous.</p> <p>Description. Male. Body (Fig. 74A) length 1.97–2.13 mm; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.</p> <p>Head (Fig. 74B) sub-rounded at base, slightly broader than long, length 0.40–0.44 mm, width across eyes 0.44– 0.46 mm; vertex smooth, lacking sulcus, vertexal foveae (dorsal tentorial pits) asetose, relatively large, mediobasal carina distinct, extending from head base anteriorly to slightly beyond level of anterior margin of eyes, lateral carina distinct, extending from base to posterior margin of antennal tubercle; posterolateral margin rounded; frons anteriorly demarcated from clypeus by oblique frontal-clypeal ridge, which merged at middle and extended anteriorly for short distance on clypeus, area between moderately raised antennal tubercles weakly impressed; clypeus with smooth surface, its entire anterior margin strongly carinate and moderately raised; ocular-mandibular carina complete. Venter with smooth surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 20 ommatidia. Antenna elongate, length 1.00– 1.1 mm, indistinct club formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–8 each slightly elongate, 8 shortest, 9 much wider and longer than 8, subconical, 10 shorter than 9, subconical, 11 largest, slightly longer than 9 and 10 combined, subfusiform, impressed at anteromesal margin.</p> <p>Pronotum (Fig. 74B) approximately as long as wide, length 0.43–0.47 mm, width 0.42–0.44 mm, widest at middle; lateral margins rounded at middle, convergent apically and sub-parallel at basal 1/4; disc slightly convex, finely punctate, thin median longitudinal sulcus with slightly carinate margins, posteriorly confluent with oval antebasal impression and short mediobasal carina, pair of discal longitudinal carinae (Fig. 74C) each composed of five distinct spines, with thin lateral longitudinal sulci; with pairs of large antebasal and small marginal spines; lateral antebasal foveae small and setose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; distinct hypomeral ridge extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity slightly carinate.</p> <p>Elytra much wider than long, length 0.59–0.61 mm, width 0.71–0.72 mm; each elytron with three moderately large, asetose basal foveae; discal stria extending posteriorly from outer basal fovea to pass half of elytral length; humerus denticulate, subhumeral fovea present, carinate marginal stria extending from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae narrowly separated, originating from shared transverse opening, large lateral mesoventral foveae forked internally, with short mesoventral process, with complete marginal stria. Metaventrite moderately prominent admesally, inclined towards middle, with well-developed lateral mesocoxal foveae, with two lateral metaventral foveae, posterior margin roundly emarginate and with narrow split at middle.</p> <p>Legs elongate; procoxa with exceptionally long seta at base; mesotrochanter (Fig. 74D) with blunt ventral spine, mesotibia (Fig. 74E) with large, rounded triangular apical projection; metatrochanter (Fig. 74F) with long, strongly curved ventral protuberance accompanied by thickened long seta, metatibia with short pencil-like tuft of setae at apex.</p> <p>Abdomen widest at lateral margins of tergite 1 (IV), length 0.54–0.62 mm, width 0.62–0.71 mm. Tergite 1 (IV) approximately twice as long as 2 (V), lacking basal sulcus, with one mediobasal and two pairs of basolateral foveae, with pair of broad, triangular discal carinae, marginal carinae complete, oblique inner carina thinner than outer one; tergite 2 (V) slightly longer than 3 (VI), with two pairs of small basolateral foveae and thin marginal carinae, 3 (VI) with one pair of small basolateral foveae and thin marginal carinae, 4 (VII) shorter than tergites 2 and 3 combined, with one pair of small basolateral foveae and thin, incomplete marginal carinae; tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of mediobasal and two pairs of basolateral foveae, with short lateral carina; midlength of sternites 2–4 (IV–VI) gradually shorter, 5 (VII) slightly longer than 4, sternites 3–4 each with three pairs, two pairs and 5 with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) (Fig. 74G) composed of pair of triangular, membranous structures.</p> <p>Aedeagus (Fig. 74H–J) 0.36 mm long, asymmetric, elongate; median lobe with broad basal capsule and large foramen, ventral stalk markedly narrowed and extremely elongate; dorsal lobe also extremely elongate at base approximately 7.5 times as broad as median lobe, narrowing apically and with rounded apex, parameres broad and flattened, membranous.</p> <p>Female. Unknown.</p> <p>Comparative notes. This species is allied with T. dilophus by the similar external morphology, and the pronotum with the discal carinae bearing five distinct spines and large antebasal spines. Tribasodites yatung can be separated by the relatively thicker antennae, the mesotibiae with large, rounded triangular projections at the apices (short apical spurs in T. dilophus), the long and straight ventral protuberances of the metatrochanters (short and strongly curved projections in T. dilophus), as well as an extremely elongate ventral stalk and dorsal lobe of the aedeagus (notably broad aedeagus in T. dilophus).</p> <p>Distribution. Yatung County, Tibet, SW China (Figs 86C, 94A).</p> <p>Etymology. This species is named after Yatung County, where its type locality is situated.</p></div> 	http://treatment.plazi.org/id/C822F213FF6724D9CB88A5CA71D120BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6524D8CB88A67F742627D6.text	C822F213FF6524D8CB88A67F742627D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisinus Raffray 1894	<div><p>Trisinus Raffray, 1894</p> <p>Chinese common name: 幻Ɵà甲¤</p> <p>Trisinus Raffray, 1894b: 265. Type species: Trisinus cratocerus Raffray, 1894b: 265 (designated by Lucas 1920: 657).</p> <p>= Batrisoplisus Raffray, 1908: 180. Type species: Batrisus antennatus Weise, 1877: 97 (monotypy). Synonymized by Yin et al. 2012c: 62.</p> <p>Trisinus was synonymized with Batrisoplisus by Yin et al. (2012c), and is currently defined by the presence of an oblique inner pair of carinae on tergite 1 (IV), and the unique form of the aedeagus. The males of most described species have characteristically modified antennae, but the new species described here possesses a modified tergite 2 (V).</p> <p>The genus is represented by 26 species distributed in Japan (21 spp., including the Ryukyu Islands), China (3 spp. in Yunnan), Singapore (1 sp.), and Indonesia (1 sp. in Sumatra) (Newton 2018). The first species from Tibet is described here.</p> </div>	http://treatment.plazi.org/id/C822F213FF6524D8CB88A67F742627D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
C822F213FF6424C6CB88A0977195254A.text	C822F213FF6424C6CB88A0977195254A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trisinus quadrilobus Yin 2022	<div><p>Trisinus quadrilobus sp. nov.</p> <p>(Figs 75, 87, 67C, 100A)</p> <p>Chinese common name: Żú幻Ɵà甲</p> <p>Type material (14 exx.). HOLOTYPE: CHINA: ♂, ‘ China: Xizang, Nyingchi City, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.000275&amp;materialsCitation.latitude=30.019722" title="Search Plazi for locations around (long 95.000275/lat 30.019722)">Pailong Village</a>, 30°01’11”N, 95°00’01”E, 2069 m, 15.iii.2017, X.-B. Song leg. [中ǞƱ Ẫffż 市ḦË乡]’ (SNUC). PARATYPES: CHINA: 7 ♂♂, 3 ♀♀, same data as that of holotype; 1 ♂, 2 ♀♀, ‘ China: <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.50278&amp;materialsCitation.latitude=29.685833" title="Search Plazi for locations around (long 95.50278/lat 29.685833)">Xiang</a>, Mêdog County, nr. 80K, 29°41’09”N, 95°30’10”E, alt. 2330 m, mixed leaf litter, sifted, 09.vii.2018, Cheng, Peng &amp; Shen leg. (中ǞƱẪẸṘ县 80K ṉ ữ)’ (all paratypes in SNUC).</p> <p>Diagnosis. Male. Body length approximately 1.85–1.90 mm. Head subtruncate at base; vertex with complete reversed U-shaped sulcus connecting small, asetose foveae, sulcus extending anteriorly at middle towards frons, with moderately long mediobasal carina; antenna elongate, lacking modifications, with scattered exceptionally long setae; all antennomeres slightly elongate, club moderately distinct. Pronotum with five longitudinal striae. Discal stria of elytron thin, extending posteriorly to approximately apical 2/5 of elytral length. Fore and hind legs simple, mesotrochanter with thin protuberance on ventral margin, mesotibia with distinct triangular spine at apex. Abdomen with long tergite 1 (IV) long, as long as tergites 2–4 (V–VII) combined; tergite 2 with two admesal and two lateral projections; sternite 2 (IV) with lamellar process at middle. Aedeagus strongly asymmetric, horn-shaped; ventral stalk broad at base, narrowing apically, dorsal lobe membranous at apex and strongly twisted and extended at basal portion. Female. Body length slightly over 1.85 mm, antenna shorter than male, legs and abdomen lacking modifications, genitalia as in Fig. 75I.</p> <p>Description. Male. Body (Fig. 75A) length 1.86–1.91 mm; color dark reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively long pubescence, antenna with scattered suberect, exceptionally long setae (up to 0.17 mm).</p> <p>Head (Fig. 75B) subtruncate at base, approximately as long as wide, length 0.40–0.41 mm, width across eyes 0.44–0.46 mm; vertex roughly punctate, with relatively small, asetose vertexal foveae (dorsal tentorial pits), with deep reversed U-shaped sulcus connecting foveae, sulcus at middle of anterior margin extending anteriorly towards apical part of frons, mediobasal carina extending from near head base anteriorly to level of anterior margin of eyes, antennal tubercles slightly raised, with setose postantennal foveae; frons broadly and shallowly impressed medially, confluent with clypeus; clypeus with smooth surface, its anterior margin carinate and strongly raised; ocular-mandibular carina complete. Venter with two tiny gular foveae (posterior tentorial pits) in single opening, with thin median carina extending from fovea anteriorly to mouthparts. Eyes moderately prominent, composed of approximately 25 ommatidia. Antenna 0.94–0.97 mm long, lacking modifications, club formed by apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each elongate, 8 smallest, much shorter and narrower than 7, 9–11 successively larger, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform.</p> <p>Pronotum (Fig. 75B) approximately as long as wide, length 0.44 mm, width 0.43–0.48 mm, widest at middle; lateral margins rounded; disc weakly convex, sparsely with sparse large punctures, with median longitudinal sulcus shorter than semicircular lateral sulci in dorsal view, with pair of discal longitudinal sulci; lacking median antebasal fovea, with complete, deep transverse antebasal sulcus connecting lateral antebasal foveae, small antebasal tubercles present; outer and inner pair of basolateral foveae distinct. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; hypomeral ridge present at basal 1/2, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity weakly carinate.</p> <p>Elytra much broader than long, length 0.61–0.62 mm, width 0.70–0.72 mm, truncate at bases; each elytron with two moderately-sized, widely separated basal foveae, subbasal fovea absent; humerus slightly prominent; discal stria extending from outer basal fovea to apical 2/5 of elytral length; subhumeral fovea present, marginal stria extending posteriorly from fovea to posterior margin of elytron.</p> <p>Mesoventrite short, demarcated from metaventrite by ridged anterior edges of impressed areas where large, setose lateral mesocoxal foveae situated at mesal margins, with pair of admesal carinae; setose median mesoventral foveae narrowly separated and originating from shared central opening, lateral mesoventral foveae large and setose, forked internally. Metaventrite broadly impressed at middle, with pair of large lateral mesocoxal foveae and setose lateral metaventral foveae; area mesal to metacoxa longitudinally carinate, posterior margin broadly emarginate, lacking split at middle.</p> <p>Legs elongate, fore and hind legs simple. Mesotrochanter with thin protuberance on ventral margin, mesotibia (Fig. 75C) with large triangular spine at apex.</p> <p>Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length 0.48–0.51 mm, width 0.63–0.66 mm, with modified tergite 2 (V) and sternite 2 (IV). Tergite 1 (IV) approximately as long as tergites 2–4 (V–VII) combined, densely punctate; lacking basal sulcus, with two mediobasal foveae and one pair of large basolateral foveae, discal carinae distinct, with complete and oblique inner marginal carinae, posterior margin with row of dense setae; tergite 2 (V) (Fig. 75D) heavily modified, central part strongly concave, with pairs of admesal and lateral projections, areas between projections impressed, with basolateral foveae lateral to admesal projections, surface of lateral projections with short, dense sensory setae; tergite 3 (VI) short, 4 (VII) three times as long as 3 along middle, tergite 5 (VIII) semicircular, posterior margin roundly emarginate at middle, tergite 3–5 (V–VIII) each with one pair of small basolateral foveae. Sternite 2 (IV) (Fig. 75E) with large, transverse mediobasal and distinct basolateral foveae, lacking basal impression or sulcus, with pair of moderately long lateral carinae, with lamellar process at middle; midlength of sternite 2 (IV) longer than 3–5 (V–VII) combined, 3–5 each with one pair of small basolateral foveae, sternite 6 (VIII) transverse, posterior margin sinuate, sternite 7 (IX) (Fig. 75F) suboval, moderately sclerotized, apex rounded and with few long setae along apical margin.</p> <p>Aedeagus (Fig. 75G, H) 0.20 mm long, strongly asymmetric and horn-like, with distinct basoventral projection; ventral stalk of median lobe in lateral view broad, rounded at apex; dorsal lobe membranous and lamellar at apex, basal half strongly extended and twisted; parameres reduced to single membranous structure.</p> <p>Female. Similar to male in external morphology; antenna shorter; legs and abdomen lacking modifications; each compound eye composed of 11 ommatidia. Measurements (as for male): body length 1.86–1.87 mm; length/width of head 0.41/ 0.46–0.47 mm, pronotum 0.44/ 0.43 mm, elytra 0.53–0.55/ 0.69–0.70 mm; abdomen 0.53–0.55/ 0.63–0.64 mm; length of antenna 0.87–0.89 mm; genitalia membranous, lacking obvious sclerotized structures.</p> <p>Comparative notes. The male of the new species can be readily separated from those of all congeners by the roughly punctate head, the unique modifications of tergite 2 (V) and sternite 2 (IV), and the form of the aedeagus.</p> <p>Distribution. Chagyib District (Nyingchi) and Mêdog County, Xizang, SW China (Figs 87, 67C, 100A).</p> <p>Etymology. The specific epithet refers to the presence of four projections of male tergite 2 (V).</p></div> 	http://treatment.plazi.org/id/C822F213FF6424C6CB88A0977195254A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yin, Zi-Wei	Yin, Zi-Wei (2022): The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae). Zootaxa 5111 (1): 1-211, DOI: https://doi.org/10.11646/zootaxa.5111.1.1, URL: http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3
