taxonID	type	description	language	source
42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	description	Figs 4, 5	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	materials_examined	Additional material examined. Mexico - Oaxaca • 2 ♂, CL 26.5 - 28.2 mm, CW 40.6 - 42.4 mm; same data as for holotype; CNCR 36324.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	description	Description. Carapace with dorsal surface flat, finely punctate, with small black granulations on frontal and anterolateral areas; gastric and branchial regions little inflated; postfrontal portion lightly depressed, almost horizontal, reaching anteriorly superior frontal border. Postfrontal lobes low, but evident, delimited anteriorly by shallow depressions, separated by narrow and deep median groove. Gastric pits deep, wide. Cardiac region discernible. Cervical groove shallow, curved, deep posteriorly, straight anteriorly, becoming obsolete near anterolateral margin, forming shallow notch. Anterolateral margin prominent, armed with 22 - 24 rounded granules of similar size; granulated between orbit and cervical groove, shallow notch next to orbit. Posterior margin straight (Fig. 4 A). In frontal view, superior frontal border straight, formed by low tubercles, divided by deep, V-shaped median notch, external angle internally projected almost touching internal orbital tooth; inferior frontal border thin, granulated, sinuous, more projected than superior one (Fig. 4 B). Orbits with external angle slightly granulated, with deep basal notch; internal orbital tooth triangular, well developed, extending to interior of orbit floor as high keel. Basal article of antennal peduncle separated from front by orbital hiatus. Antennules and antennular fossae partially visible, slightly widening in middle portion by an undulation of inferior frontal border; interantennular septum concealed by inferior frontal border. Operculum of antennal gland as ovoidal, flat plate, with middle constriction and tuft of short bristles on lateral third. Epistome devoid of setae; pterygostomian region with low granules; epistomal tooth triangular, directed downwards. Opening of efferent branchial channel subrectangular, longer than wide, width / length ratio 0.68. Third maxilliped with trapezoidal ischium, slightly longer than wide; merus anterior margin rounded with shallow rounded notch in palp articulation; ratio exopod / ischium length 0.70. Chelipeds distinctly heterochelous in males, subequal size in females. Major chela right, merus subtriangular in cross section, superior margin rounded with short transversal rows of low granules; lower inner margin with longitudinal row of conical tubercles increasing in size distally. Carpus proximal half with row of small conical tubercles, distal with scattered tubercles, ending in short acute spine, median spine obtuse. Fingers moderately gaping, punctate, cutting margins with rounded teeth; fixed finger with row of variable sized subtriangular teeth, median ones larger; tips slightly crossing when closed. Palm slightly swollen (length / width ratio 1.34), inner surface smooth, rest of palm with scattered black granules (Fig. 4 C). Dactylus moderately arched, slightly longer than palm (dactylus / palm ratio 1.05). G 1 slender, proximal half cylindrical, becoming compressed distally, meso-distal lobe conical, well developed; principal axis with medial constriction on lateral surface, twisted mesially. In mesial view (Fig. 5 B), distal half inclined towards cephalic surface. Caudo-marginal projection with distal lobe well developed, subrectangular with cephalic margin rounded, separated from proximal lobe by an incision; proximal lobe well developed, ax-shaped, 1.24 as higher than long. Meso-distal lobe arise from caudal corner of mesial surface, well developed, conical, with rounded apex. In caudal view (Fig. 5 C), distal third straight, apical cavity distally directed, caudal surface ending distally in wide, shallow concavity. Caudo-marginal projection distal crest of mesial surface higher than lateral one. Meso-distal lobe well developed, conical, apex rounded. Mesial process reduced, spoon shaped, only partially visible. In cephalic view (Fig. 5 A), mesial process reduced, with the distal surface excavated, transversal to principal axis of G 1, without spine on lateral margin, laying over carina of inner surface of proximal lobe of caudo-marginal projection; cephalic border bilobed, internal lobe pyramidal, rounded not touching internal face of caudo-marginal projection distal lobe, lateral lobe rounded, subcylindrical, projected anteriorly more than internal one. Field of apical setae visible, cephalo-caudally elongated, setae brownish, shorter than distal crest of lateral surface. Caudo-marginal projection lobes cephalically directed, separated, distal one slightly curved mesially; proximal one oval shaped, lateromesially inclined; carina of inner surface not visible. Distal crest of lateral surface sharp, with some tufts of short setae; subdistal circular scar on lateral face of principal axis partially visible. In lateral view (Fig. 5 D), distal half inclined cephalically. Caudo-marginal projection with the distal lobe partially visible, separated from the proximal lobe by incision, as long as lobe; inner surface of proximal lobe with semicircular strong carina, which extends over basal third of proximal surface. Mesial process lateral lobe subcylindrical, superior surface excavated, lateral border smooth, developing proximally rounded margin ending in subcircular scar. In distal view (Fig. 5 E), apical cavity U-shaped, opening cephalically. Field of apical setae delimited by central crest and internal surface of lateral surface; 20 - 60 apical setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute, triangular internal lobe of mesial process, close to internal surface of distal lobe of caudo-marginal projection. Mesial process with distal surface excavated, as an anterior continuation of the field of setae, raised border delimit the lateral and internal lobes. Meso-distal lobe well developed, conical, apex rounded.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	distribution	Distribution. Only known from the type locality in northern Oaxaca, Mexico.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	etymology	Etymology. The specific epithet is taken from the town where the specimens were collected: near town of Ayotzintepec. The word is Nahuatl " Ayotlzin-tepec ", and means " hill of the little turtles " (" en el cerro de las tortuguitas " in Spanish).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
17AEDB39BF005BF19AB6F6F54493D1F6.taxon	description	Figs 8 B, 9 B	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
17AEDB39BF005BF19AB6F6F54493D1F6.taxon	materials_examined	Material examined. Mexico - Tabasco • 1 ♂, holotype; LC 35.1 mm, AC 57.3 mm; Municipality of Huimanguillo, Carlos A. Madrazo, Pueblo Viejo Stream; 17 ° 23 ' 45 " N, 93 ° 39 ' 45 " W; alt. 135 m; 8 May 1997; J. L. Villalobos leg.; CNCR 18952. 2 ♂, paratypes; LC 17.3 - 24.3 mm, AC 27.7 - 37.8 mm; same data as for holotype; 12 Jun. 1997; J. L. Villalobos, R. Robles leg.; CNCR 17093. 1 ♂, 1 ♀, paratypes; LC 11.0 - 14.8 mm, AC 17.0 - 23.0 mm; same collection data as for holotype; 12 Jun. 1997; J. L. Villalobos, R. Robles leg.; CNCR 17171. 1 ♂, 2 ♀; LC 10.0 - 22.0 mm, AC 14.2 - 34.0 mm; Municipality of Huimanguillo, 3 km E of Carlos A. Madrazo, small tributary of Pedregal-Tonala River; 17 ° 23 ' 52 " N, 93 ° 40 ' 51 " W; alt. 116 m; 22 Jan. 1998; J. L. Villalobos, R. Robles leg.; CNCR 17290. Chiapas 1 ♂; LC 25 mm, AC 42 mm; Municipality of Ocozocoautla, Reserva de la Biosfera Selva el Ocote, Cerro Cola de Sapo, Frio Stream; 18 Dic 2008; A. Garcia and M. Anzueto leg.; CNCR 25445.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
17AEDB39BF005BF19AB6F6F54493D1F6.taxon	diagnosis	Diagnosis. As in Villalobos and Alvarez (2003).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
17AEDB39BF005BF19AB6F6F54493D1F6.taxon	distribution	Distribution. Only known from the type locality and surroundings (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.taxon	description	Figs 6, 7	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.taxon	materials_examined	Additional material examined. Mexico - Veracruz • 1 ♀, CL 19.6 mm, CW 29.5 mm; same data as for holotype; CNCR 36325.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.taxon	description	Description. Carapace dorsal surface slightly concave, finely punctate, frontal and anterolateral surfaces with minute granulations; gastric and branchial regions slightly inflated; postfrontal portion depressed, almost horizontal, continued anteriorly to reach superior frontal border. Postfrontal lobes low, delimited anteriorly by shallow depressions, separated by narrow, deep median groove. Cardiac region hardly discernible. Cervical groove shallow, curved posteriorly, anterior 1 / 4 straight, becoming obsolete before anterolateral margin, not reaching it. Anterolateral margin prominent, armed with 21 - 23 conical granules of similar size; portion between orbit and cervical groove granulated, with shallow notch next to orbit. Posterolateral area of carapace with short setae; posterior margin widely concave (Fig. 6 A). In frontal view, superior frontal border inclined towards central portion, formed by low tubercles, divided by moderately deep, narrow V-shaped, median notch; inferior frontal border continuous, sinuous, thinner, more projected than superior one (Fig. 6 B). Orbits with external angle slightly granulated, with shallow basal notch; internal orbital tooth triangular, well developed, extending to interior of orbit floor as high keel. Basal article of antennal peduncle separated from front by orbital hiatus. Antennules and antennular fossae partially visible, slightly wider in the middle; interantennular septum concealed by inferior frontal border. Operculum of antennal gland as ovoidal, flat plate, with middle constriction, tuft of short bristles on lateral third. Epistome, area around buccal cavity and pterygostomian region with short setae; epistomal tooth covered by patch of short setae, triangular, directed downwards. Opening of branchial efferent channel subcircular, longer than wide, width / length ratio 0.68. Third maxilliped with ischium subrectangular, slightly longer than wide, anterior margin of merus rounded; ratio exopod / ischium length 0.81. Chelipeds moderately heterochelous in both sexes, more evident in males. Major chela right, merus subtriangular in cross section, superior margin rounded with short transversal rows of low granules; lower inner margin with longitudinal row of conical tubercles increasing in size distally (Fig. 6 C). G 1 slender, proximal half cylindrical, becoming compressed distally, meso-distal lobe on mesial surface conical, well developed; principal axis with medial constriction on lateral surface, twisted mesially. In mesial view (Fig. 7 B), distal half inclined towards cephalic surface. Caudo-marginal projection with distal lobe well developed, subrectangular, cephalic margin rounded, separated from proximal lobe by an incision; proximal lobe well developed, ax-shaped, higher than long. Distal crest of MP, partially visible, some rounded and higher than apical cavity. Meso-distal lobe arising from caudal angle of mesial surface, well developed, conical, with subacute apex. In caudal view (Fig. 7 C), distal third straight, apical cavity distally directed, caudal surface ending distally in a wide and shallow concavity. Caudo-marginal projection with distal crest of mesial surface as high or higher than lateral one. Mesial process as a longitudinal plate, caudally undulated, partially visible; subdistal circular scar on the base of the plate partially visible. In cephalic view (Fig. 7 A), mesial process in longitudinal position relative to principal axis of G 1, with an incipient rounded tooth on lateral margin, laying over carina of inner surface of proximal lobe of caudo-marginal projection; cephalic border bilobed, internal lobe pyramidal, rounded not touching internal face of the distal lobe of caudo-marginal projection, lateral lobe rounded, semicylindrical, projected anteriorly beyond internal one. Field of apical setae visible, cephalo-caudally elongated, setae brownish, shorter than distal crest of lateral surface. Lobes of caudo-marginal projection cephalically directed, separated, slightly curved mesially; proximal one oval shaped, lateromesially inclined, inner surface carina not visible. Distal crest of lateral surface sharp, smooth, with few short setae; subdistal circular scar on principal axis partially visible. In lateral view (Fig. 7 D), distal lobe of caudo-marginal projection partially visible, separated from proximal one by long incision; inner surface of proximal lobe with circular or semicircular strong carina, extending over basal third of proximal surface. Mesial process as a longitudinal plate that close the apical cavity; lateral margin of smooth, superior angle rounded, median portion with an incipient rounded tooth, continued proximally to end in a subcircular scar. In distal view (Fig. 7 E), Apical cavity U-shaped, opening cephalically, field of apical setae delimited by central and lateral crests; 20 - 60 apical setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute, triangular internal lobe of mesial process. Meso-distal lobe (MDL) well developed, conical, distal tip curved mesocephalically.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.taxon	distribution	Distribution. Only known from type locality.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.taxon	etymology	Etymology. The name of this species is taken from the River Col, Los Tuxtlas region of Veracruz, where the specimens were collected. We declare the specific epithet as noun in apposition.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
27C50A5DF13E5283A5412709123F12BE.taxon	description	Figs 8 D, 9 D	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
27C50A5DF13E5283A5412709123F12BE.taxon	materials_examined	Material examined. Mexico - Veracruz • 1 ♂, holotype of Pseudothelphusa (Tehuana) cordobensis Rodriguez and Smalley (1972); LC 38.2 mm, AC 59.0 mm; Municipality of Cordoba Ojo de Agua, Paraje Nuevo, 18 ° 52 ' 35 " N, 96 ° 51 ' 49 " W; alt. 661 m; 2 May 1953; A. Villalobos leg.; CNCR 311. 1 ♂; LC 30.0 mm, AC 45.4 mm; Municipality of Amatlan de Los Reyes, Lourdes River Cave; 18 ° 47 ' 00 " N, 96 ° 54 ' 00 " W; alt. 439 m; 13 Jun. 1996; J. Herrera, E. Ramirez leg.; CNCR 11958. 3 ♂, 3 ♀; LC 15.4 - 27.9 mm, AC 22.9 - 42.7 mm; Municipality of Amatlan de Los Reyes, Amatlan II Power Station; 18 ° 51 ' 23 " N, 96 ° 54 ' 19 " W; alt. 730 m; 6 Jun. 1992; J. Herrera, E. Ramirez leg.; CNCR 11957. 1 ♀, LC 26.2 mm, AC 18.9 mm; Motzorongo, Municipality of Tezonapa, Motzorongo River; 18 ° 38 ' 30 " N, 96 ° 43 ' 53 " W; alt. 271 m; C. Pedraza, L. Garcia leg.; CNCR 34618.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
27C50A5DF13E5283A5412709123F12BE.taxon	diagnosis	Diagnosis. G 1, in cephalic view, with three protuberances on lateral surface, proximal one being the most developed. In mesial view, meso-distal lobe conical, with round and slender apex. In caudal view, median constriction forming a large lobe oriented proximally. Caudo-marginal projection with distal lobes separated by linear notch without leaving space in between. Distal lobe with rounded cephalic edge, shorter than proximal lobe. Proximal lobe ax-shaped, higher than distal (1.5 x); cephalic border circular, caudal border straight; distal crest slightly laterally oriented, lobe with sloping appearance. Internal carina well marked, circular, its length covering at least ⅓ of internal surface. Mesial process reduced, in transversal position, without lateral spine; distal edge oriented cephalad. Internal angle developed in form of lobe, two-thirds as high as mesial process, wide, touching the internal surface of distal lobe (DL) of CMP. In distal view, mesial process concave, internal angle hidden below the proximal lobe (PL) of caudo-marginal projection.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
27C50A5DF13E5283A5412709123F12BE.taxon	distribution	Distribution. Tehuana complanata is distributed around the city of Cordoba, Veracruz, Mexico (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
567BB8AE34935B6A8515FDDC80EE467B.taxon	description	Figs 8 E, 9 E	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
567BB8AE34935B6A8515FDDC80EE467B.taxon	materials_examined	Material examined. Mexico - Veracruz • 2 ♀, CL 8.2 - 13.5 mm, CW 12.4 - 20.2 mm; Municipality of Catemaco, Veracruz, Catemaco Lake, Playa Hermosa; 18 ° 26 ' 00 " N, 95 ° 04 ' 60 " W; alt. 351 m; 31 Aug. 1966; L. Holthuis, J. Cabrera leg.; CNCR 333. 3 ♂, CL 12.1 - 16.4 mm, CW 18.1 - 25.7 mm; Municipality of Catemaco, Catemaco Lake, El Zapotal; 18 ° 25 ' 00 " N, 95 ° 05 ' 60 " W; alt. 335 m; 18 Sep. 1954; A. Villalobos leg.; CNCR 334. 4 ♂, 6 ♀, CL 12.7 - 27.1 mm, CW 20.1 - 42.3 mm; Municipality of Catemaco, Catemaco Lake, Las Margaritas Stream; 18 ° 22 ' 04 " N, 95 ° 01 ' 01 " W; alt. 345 m; 4 Aug. 1994; M. E. Camacho leg.; CNCR 12956. 4 ♂, CL 12.1 - 23.4 mm, CW 17.7 - 38.1 mm; same locality as previous record; 6 Aug. 1994; F. Alvarez leg.; CNCR 12965. 1 ♂, CL 26.4 mm, CW 43.6 mm; same locality as previous record; 20 Apr. 2016; E. Moreno leg.; CNCR 34488. 1 ♂, 1 ♀, CL 12.9 - 74.6 mm, CW 11.9 - 44.7 mm; Municipality of Catemaco, 1 km S from Coyame; 18 ° 25 ' 50.6 " N, 95 ° 01 ' 16 " W; alt. 364 m; 6 Aug. 1994; J. L. Villalobos leg.; CNCR 12952. 2 ♀, CL 11.7 - 18.2 mm, CW 17.7 - 23.2 mm; same locality as previous record; 1 Aug. 1994; M. E. Camacho leg.; CNCR 12966. 2 ♀, CL 12.8 - 15.7 mm, CW 18.8 - 22.5 mm; Municipality of Catemaco, Catemaco Lake, La Agayota; 18 ° 24 ' 02 " N, 95 ° 00 ' 06 " W; alt. 545 m; 1 Jul. 1986; F. Alvarez leg.; CNCR 12907. 1 ♂, 4 ♀, CL 11.3 - 24.6 mm, CW 16.2 - 38.2 mm; same locality as previous record; 18 Jul. 1986; J. L. Villalobos leg.; CNCR 12911. 1 ♂, CL 13.1 mm, CW 20.1 mm; same locality as previous record; 4 Aug. 1994; F. Alvarez leg.; CNCR 12954. 1 ♀, CL 26.3 mm, CW 42.4 mm; same locality as previous record; 18 Jul. 1986; F. Alvarez leg.; CNCR 13125.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
567BB8AE34935B6A8515FDDC80EE467B.taxon	diagnosis	Diagnosis. G 1 slender, proximal half cylindrical, distal half becoming compressed. In caudal view, apical crest widely concave, mesial crest higher than lateral one, Meso-distal lobe conical, tip rounded. In mesial view, distal third of gonopod slightly inclined cephalically, caudo-marginal projection bilobed, lobes separated by V-shaped notch, distal lobe shorter than proximal one, rounded; proximal lobe ax-shaped, cephalic margin broadly rounded, internal margin straight. In cephalic view, caudo-marginal projection slightly curved mesially, meso-distal lobe prominent; mesial process as a widely rounded plate, distal margin rounded, lateral margin with triangular tooth, cephalic margin with rounded projection closing the apical cavity. In lateral view, mesial process oblique relative to apical cavity longitudinal axis, mesial crest clearly higher than lateral one. In apical view, apical cavity U-shaped, opening of sperm channel in caudal position, field of apical pore setae on lateral portion of cavity, caudal crest thicker than the rest.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
567BB8AE34935B6A8515FDDC80EE467B.taxon	distribution	Distribution. Only known from the north and eastern shores of Catemaco Lake, Veracruz, Mexico (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
2B98807D2D6E57FD8A6735F26EED5491.taxon	description	Figs 8 H, 9 H	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
2B98807D2D6E57FD8A6735F26EED5491.taxon	materials_examined	Material examined. Mexico - Oaxaca • 1 ♂, holotype; CL 30.5 mm, CW 48.0 mm; Municipality of Santa Maria Jacatepec, Santo Domingo River in Santa Maria Jacatepec; 17 ° 51 ' 37 " N, 96 ° 12 ' 36 " W; alt. 54 m; 23 May 1992; L. Huidobro, C. Rosas, D. Becerril, R. Palma leg.; CNCR 11920. 2 ♂, 1 ♀, CL 11.9 - 25.9 mm, CW 17.6 - 39.9 mm; Municipality of San Juan Bautista Tuxtepec, km 165 highway Tuxtepec-Palomares, El Zapote stream; 17 ° 09 ' 51 " N, 95 ° 09 ' 35 " W; alt. 167 m; 27 Sep. 1981; R. Lamothe leg.; CNCR 8817. 3 ♂, 2 ♀, CL 15.9 - 24.4 mm, CW 24.1 - 37.2 mm; Municipality of Santa Maria Jacatepec, San Isidro El Naranjal, El Mazate waterfall; 17 ° 53 ' 41 " N, 96 ° 08 ' 01 " W; alt. 103 m; 3 Mar. 2018; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 34620. 5 ♂, 3 ♀, CL 11.4 - 28.1 mm, CW 16.6 - 43.7 mm; Municipality of Santa Maria Jacatepec, stream in San Isidro El Naranjal; 17 ° 53 ' 32 " N, 96 ° 07 ' 46 " W; alt. 84 m; 3 Mar. 2018; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 34622. 3 ♂, 7 ♀, CL 7.4 - 11.9 mm, CW 10 - 17 mm; Municipality of San Jose Chiltepec, Arroyo de Pueblo Viejo; 17 ° 54 ' 26 " N, 96 ° 03 ' 12 " W; alt. 79 m; 3 Mar. 2018; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 34640. 1 ♂, 1 ♀, CL 16.7 - 35.8 mm, CW 24.6 - 43.3 mm; Municipality of San Juan Bautista Tuxtepec, Tuxtepec-Palomares highway; 17 ° 09 ' 00 " N, 95 ° 06 ' 00 " W; alt. 96 m; collection data unknown; CNCR 8806. Veracruz • 2 ♂, 3 ♀, CL 9 - 43 mm, CW 12.8 - 67.3 mm; Municipality of Playa Vicente, Nueva Era; 1 km from Santa Rosa; 17 ° 41 ' 22 " N, 95 ° 48 ' 55 " W; alt. 111 m; 7 Mar. 2018; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 34624. 3 ♂, 3 ♀, CL 11.3 - 21.6 mm, CW 15.6 - 31.2 mm; Municipality of Playa Vicente, El Tomate, El Manantial Ranch, Manzo River; 17 ° 41 ' 52.3 " N, 95 ° 51 ' 51 " W; alt. 43 m; 6 Mar. 2018; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 34626.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
2B98807D2D6E57FD8A6735F26EED5491.taxon	diagnosis	Diagnosis. As in Villalobos and Alvarez (2003).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
2B98807D2D6E57FD8A6735F26EED5491.taxon	distribution	Distribution. This species is distributed in and around the town of Santa Maria Jacatepec in northern Oaxaca, Mexico (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA97CE394291581BADF36919C49918C4.taxon	description	Figs 8 F, 9 F	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA97CE394291581BADF36919C49918C4.taxon	materials_examined	Material examined. Mexico - Oaxaca • 1 ♂, CL 50.4 mm, CW 80.6 mm; Municipality of Asuncion Ixtaltepec, Nizanda, Cerro del Naranjo, Naranjo stream; 16 ° 41 ' 14.8 " N, 95 ° 02 ' 09 " W; alt. 293 m; 15 Apr. 1999; D. Barreto, V. H. Reynoso leg.; CNCR 16875. 1 ♂, CL 31.2 mm, CW 43.4 mm; Municipality of Asuncion Ixtaltepec, Naranjo stream; 16 ° 41 ' 24 " N, 95 ° 22 ' 53 " W; alt. 639 m; 15 Sep. 1997; V. H. Reynoso leg.; CNCR 18951. 3 ♂, 2 ♀, CL 8.9 - 40.8 mm, CW 12.4 - 65 mm; Municipality of Asuncion Ixtaltepec, Nizanda, stream; 16 ° 39 ' 30 " N, 95 ° 00 ' 37 " W; alt. 186 m; 26 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33939.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA97CE394291581BADF36919C49918C4.taxon	diagnosis	Diagnosis. G 1 slender, almost straight. Meso-distal lobe conical, in mesial view somewhat compressed caudo-cephalically. Mesial process in transversal position relative to G 1 main axis, laying over proximal lobe of caudo-marginal projection, reduced, rounded, superior margin projected distally, without lateral tooth; in cephalic view internal angle triangular, pointing towards distal lobe of caudo-marginal projection (Figs 8 F, 9 F). Lobes of CMP separated by long incision, lobes not gaping; in cephalic view proximal lobe oblique relative to main axis of gonopod, distal lobe slightly curved laterally. Distal lobe of caudo-marginal projection simple, rounded, projected cephalically; proximal lobe broadly rounded extending proximally, with internal semicircular carina. Apical cavity with elongated field of setae next to lateral margin; opening of spermatic channel in caudal position.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA97CE394291581BADF36919C49918C4.taxon	distribution	Distribution. Along the Pacific versant of the Isthmus of Tehuantepec, in the drainage systems of the Tehuantepec and Zanatepec rivers, in SW Oaxaca, Mexico (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
628E2EB747D150F4AF7AD5300C15BB6B.taxon	description	Figs 8 G, 9 G	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
628E2EB747D150F4AF7AD5300C15BB6B.taxon	materials_examined	Material examined. Mexico - Chiapas • 1 ♂, holotype; CL 18.2 mm, CW 27.5 mm; Municipality of Ixtacomitan, 1 km from Ixtacomitan, La Piedra stream; 17 ° 24 ' 00 " N, 93 ° 06 ' 00 " W; alt. 232 m; 4 Apr. 1986; J. L. Villalobos, J. C. Nates, A. Cantu, D. Valle; CNCR 5604. 2 ♂, CL 21 - 24 mm, CW 32 - 37.2 mm; Municipality of Tapilula, stream near Tapilula; 17 ° 16 ' 05 " N, 93 ° 01 ' 33 " W; alt. 780 m; 20 Apr. 1981; R. Lamothe leg.; CNCR 8812.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
628E2EB747D150F4AF7AD5300C15BB6B.taxon	diagnosis	Diagnosis. As in Alvarez and Villalobos (1994).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
628E2EB747D150F4AF7AD5300C15BB6B.taxon	distribution	Distribution. Restricted to a small area in NE Chiapas, Mexico (Fig. 1).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
81534E809A365D9C9215455BE19F736A.taxon	description	Figs 8 I, 9 I	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
81534E809A365D9C9215455BE19F736A.taxon	materials_examined	Material examined. Mexico - Veracruz • 4 ♂, CL 11.4 - 26.0 mm, CW 17.0 - 41.2 mm; Municipality of San Andres Tuxtla, Basura River; 18 ° 31 ' 55 " N, 95 ° 03 ' 30 " W; alt. 33 m; 19 Jul. 1998; R. Robles, C. Graham leg.; CNCR 17422. 1 ♀, CL 15.8 mm, CW 22.9 mm; same locality as previous record; 18 Jul. 1986; J. L. Villalobos, F. Alvarez leg.; CNCR 13140. 3 ♂, 5 ♀, CL 8.9 - 28.8 mm, CW 13.1 - 45.2 mm; same locality as previous record; 4 Oct. 1994; J. L. Villalobos, F. Alvarez leg.; CNCR 13187. 7 ♂, 1 ♀, CL 9.6 - 21.8 mm, CW 14.7 - 36.4 mm; same locality as previous record; 24 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33931. 4 ♂, 11 ♀, CL 11.8 - 19.5 mm, CW 12.8 - 33.2 mm; Municipality of Santiago Tuxtla, Tapalapan River; 18 ° 32 ' 00 " N, 95 ° 18 ' 00 " W; alt. 393 m; 17 Apr. 1957; H. Hobbs, A. Villalobos leg.; CNCR 336. 5 ♀, CL 25.1 - 36.1 mm, CW 40.6 - 59.1 mm; same locality as previous record; 23 May 1955; A. Villalobos leg.; CNCR 338. 4 ♂, 4 ♀, CL 7.3 - 14 mm, CW 10.8 - 20.3 mm; same locality as previous record; 17 April 1957; H. Hobbs, A. Villalobos leg.; CNCR 386. 2 ♂, CL 10.9 - 11.9 mm, CW 12.4 - 18.8 mm; Municipality of San Andres Tuxtla, Otapan River; 18 ° 26 ' 00 " N, 95 ° 12 ' 00 " W; alt. 379 m; 22 Sep. 1955; G. Perez leg.; CNCR 337. 2 ♂, CL 11.9 - 20.7 mm, CW 21.9 - 30.3 mm; Municipality of San Andres Tuxtla, Laguna Escondida; 18 ° 35 ' 00 " N, 95 ° 05 ' 00 " W; alt. 76 m; 1 Aug. 1985; J. L. Villalobos, M. D. Valle, P. Schmidtsdorf leg.; CNCR 4473. 15 ♂, 10 ♀, CL 7.4 - 21.7 mm, CW 10.4 - 34.9 mm; same locality as previous record 13 Jun. 1985; C. Nates, J. L. Villalobos leg.; CNCR 4709. 1 ♂, 3 ♀, CL 8.2 - 25.6 mm, CW 11.6 - 41 mm; same locality as previous record; 11 Jul. 1994; J. L. Villalobos, F. Alvarez leg.; CNCR 5303. 4 ♂, 3 ♀, CL 25.4 - 32.3 mm, CW 42.2 - 53.2 mm; same locality as previous record; 24 Feb. 1989; M. Santiago leg.; CNCR 10220. 2 ♂, 2 ♀, CL 11 - 19.4 mm, CW 15.5 - 30.5 mm; same locality as previous record; 17 Jul. 1985; J. L. Villalobos, F. Alvarez leg.; CNCR 12908. 6 ♂, 4 ♀, CL 11.7 - 18.7 mm, CW 17.7 - 45.9 mm; same locality as previous record; 5 Aug. 1994; J. L. Villalobos leg.; CNCR 12964. 2 ♂, 2 ♀, CL 6.8 - 11.3 mm, CW 9.3 - 17.0 mm; same locality as previous record; 10 Jul. 1986; F. Alvarez leg.; CNCR 13138. 1 ♂, CL 20.0 mm, CW 32.3 mm; Municipality of San Andres Tuxtla, Playa Escondida; 18 ° 35 ' 00 " N, 95 ° 03 ' 00 " W; alt. 6 m; 28 Feb. 1986; A. Cantu leg.; CNCR 5782. 3 ♂, 5 ♀, CL 13.3 - 17.2 mm, CW 20.0 - 27.0 mm; same locality as previous record; 28 Feb. 1986; A. Cervantes, J. Garcia, A. Cantu leg.; CNCR 5788. 1 ♂, 4 ♀, CL 9.2 - 20.9 mm, CW 13.1 - 32.1 mm; same locality as previous record; 28 Feb. 1986; R. Lamothe leg.; CNCR 8821. 6 ♂, 11 ♀, CL 6.3 - 20.8 mm, CW 9.1 - 32.8 mm; Municipality of San Andres Tuxtla, trail to Laguna Escondida; 18 ° 35 ' 00 " N, 95 ° 04 ' 00 " W; alt. 108 m; 24 Feb. 1989; M. Santiago leg.; CNCR 10222. 1 ♂, 2 ♀, CL 11.8 - 21.7 mm, CW 17.3 - 35.1 mm; same locality as previous record; 3 Aug. 1994; F. Alvarez, J. L. Villalobos leg.; CNCR 12967. 9 ♂, 5 ♀, CL 9.7 - 28.6 mm, CW 14.3 - 49.7 mm; same locality as previous record; 23 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33927. 13 ♂, 21 ♀, CL 9.9 - 30.0 mm, CW 14.7 - 50.4 mm; Municipality of San Andres Tuxtla, Lazaro Cardenas; 18 ° 34 ' 00 " N, 95 ° 06 ' 00 " W; alt. 359 m; 9 Jul. 1991; A. Cruz leg.; CNCR 12302. 1 ♂, CL 20.3 mm, CW 32.3 mm; Municipality of San Andres Tuxtla, Cuetzalapan River; 18 ° 24 ' 00 " N, 95 ° 00 ' 00 " W; alt. 220 m; 13 Jul. 1986; F. Alvarez leg.; CNCR 12555. 2 ♂, 1 ♀, CL 11.3 - 23.6 mm, CW 18.0 - 37.7 mm; Municipality of San Andres Tuxtla, La Palma River; 18 ° 33 ' 00 " N, 95 ° 03 ' 00 " W; alt. 77 m; 12 Jul. 1986; F. Alvarez leg.; CNCR 12896. 1 ♂, 1 ♀, CL 21.1 - 23.0 mm, CW 35.0 - 38.5 mm; Balzapote River; 18 ° 36 ' 00 " N, 95 ° 04 ' 00 " W; alt. 54 m; Municipality of San Andres Tuxtla, Veracruz; 14 Jul. 1986; F. Alvarez leg.; CNCR 12906. 7 ♂, 3 ♀, CL 13.0 - 20.3 mm, CW 20.0 - 32.5 mm; Municipality of San Andres Tuxtla, Maquinas River; 18 ° 37 ' 00 " N, 95 ° 05 ' 00 " W; alt. 143 m; 1 Aug. 1994; F. Alvarez, J. L. Villalobos leg.; CNCR 12961. 1 ♂, 2 ♀, CL 10.1 - 18.3 mm, CW 16.1 - 21.6 mm; Municipality of San Andres Tuxtla, Zacatal Lagoon; 18 ° 35 ' 00 " N, 95 ° 06 ' 00 " W; alt. 263 m; 14 Oct. 1994; F. Alvarez, J. L. Villalobos leg.; CNCR 13188. 4 ♂, CL 11.8 - 23.4 mm, CW 17.5 - 37.2 mm; Municipality of Santiago Tuxtla, Simapan River; 18 ° 27 ' 00 " N, 95 ° 21 ' 00 " W; alt. 336 m; 3 May 1995; J. L. Villalobos leg.; CNCR 13346.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
81534E809A365D9C9215455BE19F736A.taxon	diagnosis	Diagnosis. G 1 slender, in mesial view distal third inclined cephalically. Meso-distal lobe conical, tip rounded, large relative to size of other apical structures (Fig. 9 I). Mesial process well developed; lateral margin with triangular, acute tooth; distal margin broadly rounded; internal angle with triangular projection in cephalic position. Lobes of the caudo-marginal projection elongated, overlapping along most of their length; proximal lobe curved cephalically, distal lobe straight, oriented forward or with the tip moderated curved proximally, slightly longer than proximal one. In mesial view, distal lobe with superior margin inclined proximally; distally, falling over proximal lobe. Internal carina of proximal lobe rounded, prominent, separated from lobe. Apical cavity with elongated field of setae, opening of spermatic channel in caudal position.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
81534E809A365D9C9215455BE19F736A.taxon	distribution	Distribution. Tehuana poglayenorum is distributed in the north-central section, of the Los Tuxtlas region.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA0A15F3CFCC5A6EA126DB2DBB6231EF.taxon	diagnosis	Diagnosis. Carapace with dorsal surface flat, smooth, punctate, with small granulations adjacent to antero-lateral margin. Front vertically deflexed, smooth, superior border formed by low rounded tubercles, medially divided by V-shaped notch reaching inferior margin; inferior margin not distinguishable dorsally, in ventral view projected, widely bilobed. Third maxillipeds with ischium trapezoidal, slightly longer than wide; merus smaller than ischium, anterolateral margin rounded, with anterior shallow concavity; ratio exopod / ischium length 0.6 to 0.8. G 1 slender, proximal half cylindrical, compressed distally, with a meso-distal lobe conical, well developed on mesial surface; principal axis with medial constriction on lateral surface and twisted towards medial suture of ventral sternites (Fig. 2). In mesial view, proximal half with marginal plate closing spermatic channel, border facing marginal suture with proximal fringe of long and stout setae; distal half inclined towards cephalic surface; caudal surface fused with marginal plate and expanded cephalically to form the bilobed caudo-marginal projection, distal lobe well-developed (except in T. lamothei where it is reduced), subrectangular with cephalic border rounded, distal portion can be curved proximally and separated from proximal lobe by an incision; proximal lobe well developed or reduced, trapezoidal, circular, ax-shaped or elongated proximally to reach and sometimes overlap gonopod principal axis (Fig. 2 B). Meso-distal lobe arising from distocaudal angle of mesial surface, conical, subacute, or flattened caudo-cephalically, distally rounded. In cephalic view, mesial process in longitudinal position, oval shaped, closing apical cavity, distal margin widely rounded, lateral margin straight, armed with strong median spine (T. chontalpaensis, T. lamothei), or with moderate or reduced triangular tooth (T. diabolis, T. col sp. nov., T. poglayenorum, T. veracruzana); or in transversal position, reduced, spoon shaped, with lateral margin smooth without spine or tooth (T. ayotzintepecensis sp. nov., T. complanata, T. jacatepecensis, T. lamellifrons). Caudo-marginal projection with the two lobes directed cephalically, distal one slightly curved mesially; proximal one, ax-shaped or elongated proximally, and inclined in different gradations; carina of inner surface partially visible. Meso-distal lobe arise from caudal corner of G 1 mesial surface, commonly is well developed, conical, with subacute or rounded apex. In lateral view, distal half inclined cephalically. Caudo-marginal projection with distal lobe (partially or totally visible; inner surface of proximal lobe with circular or semicircular strong carina, extending to different extents over proximal surface. In caudal view, distal third straight, apical cavity distally directed, caudal surface ending distally in a wide and shallow concavity. Caudo-marginal projection with the distal crest of mesial surface higher or at the same level as lateral. Meso-distal lobe well developed, conical, with the apex rounded or subacute rounded, its position with respect the distal crest of mesial surface, could variate through different species. Mesial process as a longitudinal apical plate or transversal reduced with the distal surface excavated, in the first case, oval shaped, the distal margin could be widely rounded, and the lateral margin straight and armed with a strong median spine, or with a moderate or reduced triangular tooth; in the second, only partially visible. In distal view, apical cavity U-shaped, opening cephalically. Field of apical setae with 20 to 60 setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute and triangular or rounded internal angle of mesial process (Fig. 2 D, E).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA0A15F3CFCC5A6EA126DB2DBB6231EF.taxon	distribution	Distribution. The species of Tehuana are distributed in southeastern Mexico covering the Isthmus of Tehuantepec region (Fig. 1) in Chiapas, Oaxaca, Tabasco, and Veracruz.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
E6C45F73407E58EB982CA120DF10A90C.taxon	description	Figs 8 J, 9 J	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
E6C45F73407E58EB982CA120DF10A90C.taxon	materials_examined	Material examined. Mexico - Veracruz • 1 ♂, holotype; CL 48.0 mm, CW 28.2 mm; Municipality of Catemaco, Zapoapan de Cabanas Stream; 18 ° 20 ' 00 " N, 95 ° 05 ' 48 " W; alt. 518 m; 15 Apr. 1957; A. Villalobos, H. H. Hobbs leg.; CNCR 335. 1 ♂, 1 ♀, paratypes; CL 23.8 - 28.2 mm, CW 37.2 - 47.5 mm; same data as for holotype; CNCR 335. 1 ♂, 1 ♀, CL 15.7 - 27.8 mm, CW 24.8 - 45.2 mm; Municipality of Catemaco, stream near Zapoapan de Cabanas; 18 ° 20 ' 09 " N, 95 ° 02 ' 22 " W; alt. 629 m; 25 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33934. 1 ♂, 3 ♀, CL 12.1 - 20.5 mm, CW 18.4 - 32.5 mm; Municipality of Catemaco, stream near Zapoapan de Cabanas; 18 ° 20 ' 32 " N, 95 ° 04 ' 13 " W; alt. 364 m; 25 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33932. 1 ♂, 2 ♀, CL 9.2 - 12.5 mm, CW 14.5 - 21.6 mm; Municipality of Catemaco, stream near Zapoapan de Cabanas; 18 ° 19 ' 18 " N, 95 ° 03 ' 02 " W; alt. 364 m; 25 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33937. 1 ♂, 1 ♀, CL 9.8 - 20.5 mm, CW 14.4 - 32.2 mm; Municipality of Catemaco, road Zapoapan de Cabanas-Zoteapan; 18 ° 17 ' 52 " N, 94 ° 58 ' 06 " W; alt. 848 m; 25 Apr. 2017; J. L. Villalobos, I. A. Toledano, E. Moreno leg.; CNCR 33955.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
E6C45F73407E58EB982CA120DF10A90C.taxon	diagnosis	Diagnosis. As in Rodriguez and Smalley (1972).	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
E6C45F73407E58EB982CA120DF10A90C.taxon	distribution	Distribution. This species occurs in the southeastern portion of the Los Tuxtlas Mountain Range, in an area that starts sloping towards the coastal plain of southern Veracruz.	en	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
