identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
42D5DEA71FFF5F3BA507B1B90AAC5D61.text	42D5DEA71FFF5F3BA507B1B90AAC5D61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana ayotzintepecensis Moreno-Juárez & Villalobos & Álvarez 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana ayotzintepecensis sp. nov.</p>
            <p>Figs 4, 5</p>
            <p>Type material.</p>
            <p>Holotype: Mexico - Oaxaca • 1 ♂, CL 42.6 mm, CW 67.3 mm; Municipality of Ayotzintepec, Cajone River, south of Ayotzintepec town, stream tributary of Cajone River; 17°39'46"N, 96°07'51"W; alt. 128 m; 5 Mar. 2018; J.L. Villalobos, I.A. Toledano, E.G. Moreno leg; CNCR 34628. Paratype: 1 ♂, CL 43.4 mm, CW 67.4 mm; same as for holotype; CNCR 36323.</p>
            <p>Additional material examined.</p>
            <p> Mexico - Oaxaca • 2 ♂, CL 26.5-28.2 mm, CW 40.6-42.4 mm; same data as for holotype; CNCR 36324 . </p>
            <p>Description.</p>
            <p>Carapace with dorsal surface flat, finely punctate, with small black granulations on frontal and anterolateral areas; gastric and branchial regions little inflated; postfrontal portion lightly depressed, almost horizontal, reaching anteriorly superior frontal border. Postfrontal lobes low, but evident, delimited anteriorly by shallow depressions, separated by narrow and deep median groove. Gastric pits deep, wide. Cardiac region discernible. Cervical groove shallow, curved, deep posteriorly, straight anteriorly, becoming obsolete near anterolateral margin, forming shallow notch. Anterolateral margin prominent, armed with 22-24 rounded granules of similar size; granulated between orbit and cervical groove, shallow notch next to orbit. Posterior margin straight (Fig. 4A). In frontal view, superior frontal border straight, formed by low tubercles, divided by deep, V-shaped median notch, external angle internally projected almost touching internal orbital tooth; inferior frontal border thin, granulated, sinuous, more projected than superior one (Fig. 4B). Orbits with external angle slightly granulated, with deep basal notch; internal orbital tooth triangular, well developed, extending to interior of orbit floor as high keel. Basal article of antennal peduncle separated from front by orbital hiatus. Antennules and antennular fossae partially visible, slightly widening in middle portion by an undulation of inferior frontal border; interantennular septum concealed by inferior frontal border. Operculum of antennal gland as ovoidal, flat plate, with middle constriction and tuft of short bristles on lateral third. Epistome devoid of setae; pterygostomian region with low granules; epistomal tooth triangular, directed downwards. Opening of efferent branchial channel subrectangular, longer than wide, width/length ratio 0.68. Third maxilliped with trapezoidal ischium, slightly longer than wide; merus anterior margin rounded with shallow rounded notch in palp articulation; ratio exopod/ischium length 0.70.</p>
            <p>Chelipeds distinctly heterochelous in males, subequal size in females. Major chela right, merus subtriangular in cross section, superior margin rounded with short transversal rows of low granules; lower inner margin with longitudinal row of conical tubercles increasing in size distally. Carpus proximal half with row of small conical tubercles, distal with scattered tubercles, ending in short acute spine, median spine obtuse. Fingers moderately gaping, punctate, cutting margins with rounded teeth; fixed finger with row of variable sized subtriangular teeth, median ones larger; tips slightly crossing when closed. Palm slightly swollen (length/width ratio 1.34), inner surface smooth, rest of palm with scattered black granules (Fig. 4C). Dactylus moderately arched, slightly longer than palm (dactylus/palm ratio 1.05).</p>
            <p>G1 slender, proximal half cylindrical, becoming compressed distally, meso-distal lobe conical, well developed; principal axis with medial constriction on lateral surface, twisted mesially. In mesial view (Fig. 5B), distal half inclined towards cephalic surface. Caudo-marginal projection with distal lobe well developed, subrectangular with cephalic margin rounded, separated from proximal lobe by an incision; proximal lobe well developed, ax-shaped, 1.24 as higher than long. Meso-distal lobe arise from caudal corner of mesial surface, well developed, conical, with rounded apex. In caudal view (Fig. 5C), distal third straight, apical cavity distally directed, caudal surface ending distally in wide, shallow concavity. Caudo-marginal projection distal crest of mesial surface higher than lateral one. Meso-distal lobe well developed, conical, apex rounded. Mesial process reduced, spoon shaped, only partially visible. In cephalic view (Fig. 5A), mesial process reduced, with the distal surface excavated, transversal to principal axis of G1, without spine on lateral margin, laying over carina of inner surface of proximal lobe of caudo-marginal projection; cephalic border bilobed, internal lobe pyramidal, rounded not touching internal face of caudo-marginal projection distal lobe, lateral lobe rounded, subcylindrical, projected anteriorly more than internal one. Field of apical setae visible, cephalo-caudally elongated, setae brownish, shorter than distal crest of lateral surface. Caudo-marginal projection lobes cephalically directed, separated, distal one slightly curved mesially; proximal one oval shaped, lateromesially inclined; carina of inner surface not visible. Distal crest of lateral surface sharp, with some tufts of short setae; subdistal circular scar on lateral face of principal axis partially visible. In lateral view (Fig. 5D), distal half inclined cephalically. Caudo-marginal projection with the distal lobe partially visible, separated from the proximal lobe by incision, as long as lobe; inner surface of proximal lobe with semicircular strong carina, which extends over basal third of proximal surface. Mesial process lateral lobe subcylindrical, superior surface excavated, lateral border smooth, developing proximally rounded margin ending in subcircular scar. In distal view (Fig. 5E), apical cavity U-shaped, opening cephalically. Field of apical setae delimited by central crest and internal surface of lateral surface; 20-60 apical setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute, triangular internal lobe of mesial process, close to internal surface of distal lobe of caudo-marginal projection. Mesial process with distal surface excavated, as an anterior continuation of the field of setae, raised border delimit the lateral and internal lobes. Meso-distal lobe well developed, conical, apex rounded.</p>
            <p>Distribution.</p>
            <p>Only known from the type locality in northern Oaxaca, Mexico.</p>
            <p>Etymology.</p>
            <p> The specific epithet is taken from the town where the specimens were collected: near town of Ayotzintepec. The word is  Náhuatl “Ayotlzin-tepec” , and means "hill of the little turtles" ("en el cerro de las tortuguitas" in Spanish). </p>
            <p>Remarks.</p>
            <p> Tehuana ayotzintepec sp. nov. has the typical morphology of the species distributed throughout the Isthmus of Tehuantepec, with a reduced mesial process, cylindrical or spoon shaped, in a transversal position relative to the G1 principal axis and laying over the proximal lobe of the caudo-marginal projection. These characters, make the new species similar to  T. complanata ,  T. jacatepecensis , and  T. lamellifrons ; however, in  T. ayotzintepecensis sp. nov. the two lobes of the caudo-marginal projection are of the same length in mesial view, the proximal one has a subacute inner angle and the distal one is completely rounded. Geographically,  T. ayotzintepecensis sp. nov. and  T. jacatepecensis occur in the same general area along the Gulf of Mexico versant of the Sierra de  Juárez in northern Oaxaca. In contrast,  T. lamellifrons is distributed along the Pacific versant of the Sierra Madre Occidental in southern Oaxaca, and  T. complanata occurs in central Veracruz (Fig. 1). The phylogenetic tree is consistent with the morphological similarity as it shows a close relationship between the four species (Fig. 3). </p>
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	https://treatment.plazi.org/id/42D5DEA71FFF5F3BA507B1B90AAC5D61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
17AEDB39BF005BF19AB6F6F54493D1F6.text	17AEDB39BF005BF19AB6F6F54493D1F6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana chontalpaensis Villalobos & Alvarez 2003	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana chontalpaensis Villalobos &amp; Alvarez, 2003</p>
            <p>Figs 8B, 9B</p>
            <p> Tehuana chontalpaensis Villalobos &amp; Alvarez, 2003: 224, 228 (in key), figs 2, 4A.-  Álvarez et al. 2005: 191.-  Rodríguez and  Magalhães 2005: 356, tab. 1 (list).- Villalobos Hiriart and  Álvarez 2008: 280, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).- Alvarez and Villalobos 2016: 254, tab.8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list).-  Álvarez et al. 2020: 979, tab. 1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Tabasco • 1 ♂, holotype; LC 35.1 mm, AC 57.3 mm; Municipality of Huimanguillo, Carlos A.  
                <a title="Search Plazi for locations around (long -93.6625/lat 17.395834)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.6625&amp;materialsCitation.latitude=17.395834">Madrazo</a>
                 ,  
                <a title="Search Plazi for locations around (long -93.6625/lat 17.395834)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.6625&amp;materialsCitation.latitude=17.395834">Pueblo Viejo Stream</a>
                 ; 17°23'45"N, 93°39'45"W; alt. 135 m; 8 May 1997; J.L. Villalobos leg.; CNCR 18952  .  2 ♂, paratypes; LC 17.3-24.3 mm, AC 27.7-37.8 mm; same data as for holotype; 12 Jun. 1997; J.L. Villalobos, R. Robles leg.; CNCR 17093 .  1 ♂, 1 ♀, paratypes; LC 11.0- 14.8 mm, AC 17.0-23.0 mm; same collection data as for holotype; 12 Jun. 1997; J.L. Villalobos, R. Robles leg.; CNCR 17171 .   1 ♂, 2 ♀; LC 10.0-22.0 mm, AC 14.2-34.0 mm; Municipality of Huimanguillo, 3 km E of Carlos A. Madrazo, small tributary of  
                <a title="Search Plazi for locations around (long -93.68083/lat 17.397778)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.68083&amp;materialsCitation.latitude=17.397778">Pedregal-Tonala River</a>
                 ; 17°23'52"N, 93°40'51"W; alt. 116 m; 22 Jan. 1998; J.L. Villalobos, R. Robles leg.; CNCR 17290  .   Chiapas 1 ♂; LC 25 mm, AC 42 mm; Municipality of Ocozocoautla, Reserva de la  Biósfera Selva el  Ocote ,  Cerro Cola de Sapo ,  Frio Stream ; 18 Dic 2008; A.  García and M. Anzueto leg.; CNCR 25445  . 
            </p>
            <p>Diagnosis.</p>
            <p>As in Villalobos and Alvarez (2003).</p>
            <p>Distribution.</p>
            <p>Only known from the type locality and surroundings (Fig. 1).</p>
            <p>Remarks.</p>
            <p> As noted by Villalobos and Alvarez (2003)  Tehuana chontalpaensis is morphologically similar to  T. lamothei , a similarity that is consistent with their being sister species as shown in the molecular phylogeny (Fig. 3). </p>
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	https://treatment.plazi.org/id/17AEDB39BF005BF19AB6F6F54493D1F6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
66814882A7235BFEBDFEAFC3D6AEF630.text	66814882A7235BFEBDFEAFC3D6AEF630.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana col Moreno-Juárez & Villalobos & Álvarez 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana col sp. nov.</p>
            <p>Figs 6, 7</p>
            <p>Type material.</p>
            <p> Holotype: Mexico - Veracruz • 1 ♂, CL 30.4 mm, CW 50.3 mm; Municipality of San  Andrés Tuxtla, Col River at Cascadas Park; 18°38'29"N, 95°09'00"W; alt. 416 m; 25 Apr. 2017; J.L. Villalobos, I.A. Toledano, E.G. Moreno leg; CNCR 33928. Paratype: 1 ♂, CL 13.6 mm, CW 20.8 mm; same data as for holotype; CNCR 36325. </p>
            <p>Additional material examined.</p>
            <p> Mexico - Veracruz • 1 ♀, CL 19.6 mm, CW 29.5 mm; same data as for holotype; CNCR 36325 . </p>
            <p>Description.</p>
            <p> Carapace dorsal surface slightly concave, finely punctate, frontal and anterolateral surfaces with minute granulations; gastric and branchial regions slightly inflated; postfrontal portion depressed, almost horizontal, continued anteriorly to reach superior frontal border. Postfrontal lobes low, delimited anteriorly by shallow depressions, separated by narrow, deep median groove. Cardiac region hardly discernible. Cervical groove shallow, curved posteriorly, anterior  ¼ straight, becoming obsolete before anterolateral margin, not reaching it. Anterolateral margin prominent, armed with 21-23 conical granules of similar size; portion between orbit and cervical groove granulated, with shallow notch next to orbit. Posterolateral area of carapace with short setae; posterior margin widely concave (Fig. 6A). In frontal view, superior frontal border inclined towards central portion, formed by low tubercles, divided by moderately deep, narrow V-shaped, median notch; inferior frontal border continuous, sinuous, thinner, more projected than superior one (Fig. 6B). Orbits with external angle slightly granulated, with shallow basal notch; internal orbital tooth triangular, well developed, extending to interior of orbit floor as high keel. Basal article of antennal peduncle separated from front by orbital hiatus. Antennules and antennular fossae partially visible, slightly wider in the middle; interantennular septum concealed by inferior frontal border. Operculum of antennal gland as ovoidal, flat plate, with middle constriction, tuft of short bristles on lateral third. Epistome, area around buccal cavity and pterygostomian region with short setae; epistomal tooth covered by patch of short setae, triangular, directed downwards. Opening of branchial efferent channel subcircular, longer than wide, width/length ratio 0.68. Third maxilliped with ischium subrectangular, slightly longer than wide, anterior margin of merus rounded; ratio exopod/ischium length 0.81. </p>
            <p>Chelipeds moderately heterochelous in both sexes, more evident in males. Major chela right, merus subtriangular in cross section, superior margin rounded with short transversal rows of low granules; lower inner margin with longitudinal row of conical tubercles increasing in size distally (Fig. 6C).</p>
            <p>G1 slender, proximal half cylindrical, becoming compressed distally, meso-distal lobe on mesial surface conical, well developed; principal axis with medial constriction on lateral surface, twisted mesially. In mesial view (Fig. 7B), distal half inclined towards cephalic surface. Caudo-marginal projection with distal lobe well developed, subrectangular, cephalic margin rounded, separated from proximal lobe by an incision; proximal lobe well developed, ax-shaped, higher than long. Distal crest of MP, partially visible, some rounded and higher than apical cavity. Meso-distal lobe arising from caudal angle of mesial surface, well developed, conical, with subacute apex. In caudal view (Fig. 7C), distal third straight, apical cavity distally directed, caudal surface ending distally in a wide and shallow concavity. Caudo-marginal projection with distal crest of mesial surface as high or higher than lateral one. Mesial process as a longitudinal plate, caudally undulated, partially visible; subdistal circular scar on the base of the plate partially visible. In cephalic view (Fig. 7A), mesial process in longitudinal position relative to principal axis of G1, with an incipient rounded tooth on lateral margin, laying over carina of inner surface of proximal lobe of caudo-marginal projection; cephalic border bilobed, internal lobe pyramidal, rounded not touching internal face of the distal lobe of caudo-marginal projection, lateral lobe rounded, semicylindrical, projected anteriorly beyond internal one. Field of apical setae visible, cephalo-caudally elongated, setae brownish, shorter than distal crest of lateral surface. Lobes of caudo-marginal projection cephalically directed, separated, slightly curved mesially; proximal one oval shaped, lateromesially inclined, inner surface carina not visible. Distal crest of lateral surface sharp, smooth, with few short setae; subdistal circular scar on principal axis partially visible. In lateral view (Fig. 7D), distal lobe of caudo-marginal projection partially visible, separated from proximal one by long incision; inner surface of proximal lobe with circular or semicircular strong carina, extending over basal third of proximal surface. Mesial process as a longitudinal plate that close the apical cavity; lateral margin of smooth, superior angle rounded, median portion with an incipient rounded tooth, continued proximally to end in a subcircular scar. In distal view (Fig. 7E), Apical cavity U-shaped, opening cephalically, field of apical setae delimited by central and lateral crests; 20-60 apical setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute, triangular internal lobe of mesial process. Meso-distal lobe (MDL) well developed, conical, distal tip curved mesocephalically.</p>
            <p>Distribution.</p>
            <p>Only known from type locality.</p>
            <p>Etymology.</p>
            <p>The name of this species is taken from the River Col, Los Tuxtlas region of Veracruz, where the specimens were collected. We declare the specific epithet as noun in apposition.</p>
            <p>Remarks.</p>
            <p> Tehuana col sp. nov. is morphologically similar to  T. poglayenorum which occurs in the same area in Los Tuxtlas region, with both exhibiting partially overlapping lobes of the caudo-marginal projection; however, they can be easily separated by the mesial process, irregular shape with a sinuous lateral margin in the former, versus a widely rounded plate with a proximal triangular tooth in the latter. Consistent with the morphology,  T. col sp. nov. and  T. poglayenorum are also genetically closely related (Fig. 3), and in turn they are related to  T. diabolis and  T. veracruzana . It is interesting to highlight that four clearly defined species of  Tehuana together with  Smalleyus tricristatus Alvarez, 1989 and  Pseudothelphusa parabelliana Alvarez, 1989 occur in Los Tuxtlas region which is small mountain range occupying an 80  × 33 km area in the coastal plain of southern Veracruz. </p>
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	https://treatment.plazi.org/id/66814882A7235BFEBDFEAFC3D6AEF630	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
27C50A5DF13E5283A5412709123F12BE.text	27C50A5DF13E5283A5412709123F12BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana complanata (Rathbun 1905)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana complanata (Rathbun, 1905)</p>
            <p>Figs 8D, 9D</p>
            <p> ? Pseudothelphusa bocourti .-Rathbun, 1898: 512 (in key), 513, 533 (list) [not  Boscia bocourti A. Milne-Edwards, 1866]. </p>
            <p> Pseudothelphusa complanata Rathbun, 1905: 303, fig. 3.- Coifmann 1939: 107 (list). </p>
            <p> "Pseudothelphusa"
complanata
 .-Pretzmann, 1965: 10 (list). </p>
            <p> Pseudothelphusa Pseudothelphusa (  Pseudothelphusa )  Pseudothelphusa lamellifrons gruneri .-Pretzmann, 1968: 7.- Pretzmann 1971: 22 (list). </p>
            <p> Pseudothelphusa (Pseudothelphusa) americana lamellifrons .-Bott, 1970: 334, pl. 2, figs 11-13. </p>
            <p> Potamocarcinus (Raddaus) bocourti complanata .-Pretzmann, 1971: 20.(list). </p>
            <p> Potamocarcinus (Raddaus) bocourti complanatus .-Pretzmann, 1972: 78, text fig. 18, fig. 542. </p>
            <p> Pseudothelphusa Pseudothelphusa (Tehuana) lamellifrons gruneri .-Pretzmann, 1972: 108, figs 621-623, 674-677. </p>
            <p> Pseudothelphusa (Tehuana) cordobensis Rodríguez &amp; Smalley, 1972: 77, fig. 8, pl. 5. </p>
            <p> Pseudothelphusa (Tehuana) complanata Türkay , 1978: 145, figs 2a, b, 3. </p>
            <p> Tehuana complanata Rodriguez, 1982: 131, fig. 85.- Villalobos 1982: 221 (list).- Villalobos-Hiriart et al. 1993: 284 (list).- Alvarez and Villalobos 1994: 730.-  Álvarez et al. 1999: 20, fig. 3 (map), 23, box 3 (list).-Villalobos and Alvarez 2003: 2003 (in key).- Villalobos Hiriart and  Álvarez 2008: 280, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).-  Álvarez et al. 2011a: appx. VIII.20, p. 13 (list).-  Mejía-Ortíz et al. 2011: 97, 136 (map 2).-  Álvarez et al. 2012: 1078, box 1 (list).- Cumberlidge et al. 2014: 144, tab. 3 (list).- Alvarez and Villalobos 2016: 254, tab.8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Veracruz • 1 ♂, holotype of  Pseudothelphusa (Tehuana) cordobensis Rodríguez and Smalley (1972); LC 38.2 mm, AC 59.0 mm; Municipality of  Córdoba Ojo de Agua,  
                <a title="Search Plazi for locations around (long -96.86361/lat 18.876389)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.86361&amp;materialsCitation.latitude=18.876389">Paraje Nuevo</a>
                 , 18°52'35"N, 96°51'49"W; alt. 661 m; 2 May 1953; A. Villalobos leg.; CNCR 311  .   1 ♂; LC 30.0 mm, AC 45.4 mm; Municipality of  Amatlán de Los Reyes,  
                <a title="Search Plazi for locations around (long -96.9/lat 18.783333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.9&amp;materialsCitation.latitude=18.783333">Lourdes River Cave</a>
                 ; 18°47'00"N, 96°54'00"W; alt. 439 m; 13 Jun. 1996; J. Herrera, E.  Ramírez leg.; CNCR 11958  .   3 ♂, 3 ♀; LC 15.4-27.9 mm, AC 22.9-42.7 mm; Municipality of  Amatlán de Los Reyes,  Amatlán II  
                <a title="Search Plazi for locations around (long -96.90528/lat 18.85639)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.90528&amp;materialsCitation.latitude=18.85639">Power Station</a>
                 ; 18°51'23"N, 96°54'19"W; alt. 730 m; 6 Jun. 1992; J. Herrera, E.  Ramírez leg.; CNCR 11957  .   1 ♀, LC 26.2 mm, AC 18.9 mm; Motzorongo, Municipality of Tezonapa,  
                <a title="Search Plazi for locations around (long -96.73139/lat 18.641666)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.73139&amp;materialsCitation.latitude=18.641666">Motzorongo River</a>
                 ; 18°38'30"N, 96°43'53"W; alt. 271 m; C. Pedraza, L.  García leg.; CNCR 34618  . 
            </p>
            <p>Diagnosis.</p>
            <p> G1, in cephalic view, with three protuberances on lateral surface, proximal one being the most developed. In mesial view, meso-distal lobe conical, with round and slender apex. In caudal view, median constriction forming a large lobe oriented proximally. Caudo-marginal projection with distal lobes separated by linear notch without leaving space in between. Distal lobe with rounded cephalic edge, shorter than proximal lobe. Proximal lobe ax-shaped, higher than distal (1.5  × ); cephalic border circular, caudal border straight; distal crest slightly laterally oriented, lobe with sloping appearance. Internal carina well marked, circular, its length covering at least ⅓ of internal surface. Mesial process reduced, in transversal position, without lateral spine; distal edge oriented cephalad. Internal angle developed in form of lobe, two-thirds as high as mesial process, wide, touching the internal surface of distal lobe (DL) of CMP. In distal view, mesial process concave, internal angle hidden below the proximal lobe (PL) of caudo-marginal projection. </p>
            <p>Distribution.</p>
            <p> Tehuana complanata is distributed around the city of Cordoba, Veracruz, Mexico (Fig. 1). </p>
            <p>Remarks.</p>
            <p> The recognition of the type locality of  Tehuana complanata has been problematic since Rathbun (1905) cited "Coban, Alta Vera Paz, Guatemala" as the type locality of  Pseudothelphusa complanata. Later,  Rodríguez and Smalley (1972) described  Pseudothelphusa (Tehuana) cordobensis from "Paraje Nuevo,  Córdoba , Veracruz" which fits the description of  T. complanata .  Türkay (1978) discussed this situation concluding that it was a labelling error by Bocourt who placed crabs from Veracruz, Mexico in a jar with specimens from Coban, Guatemala. Rodriguez (1982) synonymized  P. (T.) cordobensis under  T. complanata and designated the male from "Paraje Nuevo,  Córdoba , Veracruz" (CNCR 311) as the holotype. </p>
            <p> Morphologically, the G1 of  T. complanata is similar to that of  T. jacatepecensis ; both species have a broadly rounded to semicircular proximal lobe of the CMP, although in the former the distal and proximal lobes are subequal in length, whereas in the latter the proximal lobe is clearly shorter (Fig. 8D).  Tehuana complanata is also similar to  T. lamellifrons even when they are geographically distant within the genus range (Fig. 1); however, in the obtained phylogeny they appear as sister species (Fig. 3). </p>
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	https://treatment.plazi.org/id/27C50A5DF13E5283A5412709123F12BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
567BB8AE34935B6A8515FDDC80EE467B.text	567BB8AE34935B6A8515FDDC80EE467B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana diabolis (Pretzmann 1978)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana diabolis (Pretzmann, 1978)</p>
            <p>Figs 8E, 9E</p>
            <p> Pseudothelphusa (Tehuana) lamellifrons diabolis Pretzmann, 1978: 3.- Pretzmann 1980: 660, pl. 13, figs 56-60, pl. 17, fig. 77.- Rodriguez 1982: 210. </p>
            <p> Tehuana diabolis. - Villalobos-Hiriart et al. 1993: 284 (list).- Alvarez and Villalobos 1994: 730, 735.-  Álvarez and Villalobos 1997a: 416, 417, box 4.17 (list).-  Álvarez and Villalobos 1997b: 438, appx. 4.23 (list).-  Álvarez et al. 1999: 20, fig. 3 (map), 23, box 3 (list).-Villalobos and Alvarez 2003: 228 (in key).-Villalobos and  Álvarez 2008: 281, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).-  Álvarez et al. 2011a: appx. VIII.20, p. 13 (list); 2012: 1078, box 1 (list).- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list).-  Álvarez et al. 2020: 979, tab. 1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Veracruz • 2 ♀, CL 8.2-13.5 mm, CW 12.4-20.2 mm;  Municipality of Catemaco ,   Veracruz, Catemaco Lake, Playa Hermosa; 18°26'00"N, 95°04'60"W; alt. 351 m; 31 Aug. 1966; L. Holthuis, J. Cabrera leg.; CNCR 333. 3 ♂, CL 12.1-16.4 mm, CW 18.1-25.7 mm;  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Municipality of Catemaco</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Catemaco Lake</a>
                 , El Zapotal; 18°25'00"N, 95°05'60"W; alt. 335 m; 18 Sep. 1954; A. Villalobos leg.; CNCR 334. 4 ♂, 6 ♀, CL 12.7-27.1 mm, CW 20.1-42.3 mm;  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Municipality of Catemaco</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Catemaco Lake</a>
                 , Las Margaritas Stream; 18°22'04"N, 95°01'01"W; alt. 345 m; 4 Aug. 1994; M.E. Camacho leg.; CNCR 12956. 4 ♂, CL 12.1-23.4 mm, CW 17.7-38.1 mm; same locality as previous record; 6 Aug. 1994; F.  Álvarez leg.; CNCR 12965. 1 ♂, CL 26.4 mm, CW 43.6 mm; same locality as previous record; 20 Apr. 2016; E. Moreno leg.; CNCR 34488. 1 ♂, 1 ♀, CL 12.9-74.6 mm, CW 11.9-44.7 mm;  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Municipality of Catemaco</a>
                 , 1 km S from Coyame; 18°25'50.6"N, 95°01'16"W; alt. 364 m; 6 Aug. 1994; J.L. Villalobos leg.; CNCR 12952. 2 ♀, CL 11.7-18.2 mm, CW 17.7-23.2 mm; same locality as previous record; 1 Aug. 1994; M.E. Camacho leg.; CNCR 12966. 2 ♀, CL 12.8-15.7 mm, CW 18.8-22.5 mm;  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Municipality of Catemaco</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.00166/lat 18.400555)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.00166&amp;materialsCitation.latitude=18.400555">Catemaco Lake</a>
                 , La Agayota; 18°24'02"N, 95°00'06"W; alt. 545 m; 1 Jul. 1986; F.  Álvarez leg.; CNCR 12907. 1 ♂, 4 ♀, CL 11.3-24.6 mm, CW 16.2-38.2 mm; same locality as previous record; 18 Jul. 1986; J.L. Villalobos leg.; CNCR 12911. 1 ♂, CL 13.1 mm, CW 20.1 mm; same locality as previous record; 4 Aug. 1994; F.  Álvarez leg.; CNCR 12954. 1 ♀, CL 26.3 mm, CW 42.4 mm; same locality as previous record; 18 Jul. 1986; F.  Álvarez leg.; CNCR 13125  . 
            </p>
            <p>Diagnosis.</p>
            <p>G1 slender, proximal half cylindrical, distal half becoming compressed. In caudal view, apical crest widely concave, mesial crest higher than lateral one, Meso-distal lobe conical, tip rounded. In mesial view, distal third of gonopod slightly inclined cephalically, caudo-marginal projection bilobed, lobes separated by V-shaped notch, distal lobe shorter than proximal one, rounded; proximal lobe ax-shaped, cephalic margin broadly rounded, internal margin straight. In cephalic view, caudo-marginal projection slightly curved mesially, meso-distal lobe prominent; mesial process as a widely rounded plate, distal margin rounded, lateral margin with triangular tooth, cephalic margin with rounded projection closing the apical cavity. In lateral view, mesial process oblique relative to apical cavity longitudinal axis, mesial crest clearly higher than lateral one. In apical view, apical cavity U-shaped, opening of sperm channel in caudal position, field of apical pore setae on lateral portion of cavity, caudal crest thicker than the rest.</p>
            <p>Type material.</p>
            <p>The holotype (NHMW 4068) and paratypes (NHMW 4069) are deposited in the Natural History Museum in Vienna, Austria.</p>
            <p>Type locality.</p>
            <p> México , Veracruz, Municipality of Catemaco, Catemaco Lake; 18°25'00"N, 95°06'00"W; alt. 325 m (Pretzmann 1980). </p>
            <p>Distribution.</p>
            <p>Only known from the north and eastern shores of Catemaco Lake, Veracruz, Mexico (Fig. 1).</p>
            <p>Remarks.</p>
            <p> A diagnosis, based on the description of G1 is provided for  T. diabolis since Pretzmann (1980) description and subsequent mentions of the species by other authors omitted important morphological details of G1. In  Tehuana diabolis G1 the proximal lobe of the caudo-marginal projection has an intermediate shape between those of  T. veracruzana and  T. poglayenorum (Fig. 8), and in the phylogenetic tree the three species appear also closely related (Fig. 3). Geographically,  T. diabolis distribution around Lake Catemaco is also between that of  T. poglayenorum to the northeast and that of  T. veracruzana to the south of Catemaco Lake in the town of Zapoapan de  Cabañas . </p>
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	https://treatment.plazi.org/id/567BB8AE34935B6A8515FDDC80EE467B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
2B98807D2D6E57FD8A6735F26EED5491.text	2B98807D2D6E57FD8A6735F26EED5491.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana jacatepecensis Villalobos & Alvarez 2003	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana jacatepecensis Villalobos &amp; Alvarez, 2003</p>
            <p>Figs 8H, 9H</p>
            <p> Tehuana jacatepecensis Villalobos &amp; Alvarez, 2003: 226, 228 (in key), figs 3, 4B.-  Rodríguez and  Magalhães 2005: 356, tab. 1 (list).- Villalobos Hiriart and  Álvarez 2008: 280, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 475, 477, fig. 11 (map).-  Mejía-Ortíz et al. 2011: 99, 136 (map 2).- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Oaxaca • 1 ♂, holotype; CL 30.5 mm, CW 48.0 mm; Municipality of Santa  María Jacatepec,  
                <a title="Search Plazi for locations around (long -96.21/lat 17.86028)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.21&amp;materialsCitation.latitude=17.86028">
Santo Domingo River in Santa 
Maria
Jacatepec
</a>
                 ; 17°51'37"N, 96°12'36"W; alt. 54 m; 23 May 1992; L. Huidobro, C. Rosas, D. Becerril, R. Palma leg.; CNCR 11920  .   2 ♂, 1 ♀, CL 11.9-25.9 mm, CW 17.6-39.9 mm; Municipality of San Juan Bautista Tuxtepec, km 165 highway Tuxtepec-Palomares,  
                <a title="Search Plazi for locations around (long -95.15972/lat 17.164165)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.15972&amp;materialsCitation.latitude=17.164165">El Zapote</a>
                 stream; 17°09'51"N, 95°09'35"W; alt. 167 m; 27 Sep. 1981; R. Lamothe leg.; CNCR 8817  .   3 ♂, 2 ♀, CL 15.9-24.4 mm, CW 24.1-37.2 mm; Municipality of Santa  María Jacatepec,  
                <a title="Search Plazi for locations around (long -96.133606/lat 17.894722)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.133606&amp;materialsCitation.latitude=17.894722">San Isidro El Naranjal</a>
                 ,  
                <a title="Search Plazi for locations around (long -96.133606/lat 17.894722)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.133606&amp;materialsCitation.latitude=17.894722">El Mazate</a>
                 waterfall; 17°53'41"N, 96°08'01"W; alt. 103 m; 3 Mar. 2018; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 34620  .   5 ♂, 3 ♀, CL 11.4-28.1 mm, CW 16.6-43.7 mm; Municipality of Santa  María Jacatepec, stream in  
                <a title="Search Plazi for locations around (long -96.12945/lat 17.892221)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.12945&amp;materialsCitation.latitude=17.892221">San Isidro El Naranjal</a>
                 ; 17°53'32"N, 96°07'46"W; alt. 84 m; 3 Mar. 2018; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 34622  .   3 ♂, 7 ♀, CL 7.4-11.9 mm, CW 10-17 mm; Municipality of San  José Chiltepec,  
                <a title="Search Plazi for locations around (long -96.05334/lat 17.907223)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.05334&amp;materialsCitation.latitude=17.907223">Arroyo de Pueblo Viejo</a>
                 ; 17°54'26"N, 96°03'12"W; alt. 79 m; 3 Mar. 2018; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 34640  .   1 ♂, 1 ♀, CL 16.7-35.8 mm, CW 24.6-43.3 mm; Municipality of San Juan Bautista Tuxtepec,  
                <a title="Search Plazi for locations around (long -95.1/lat 17.15)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.1&amp;materialsCitation.latitude=17.15">Tuxtepec-Palomares</a>
                 highway; 17°09'00"N, 95°06'00"W; alt. 96 m; collection data unknown; CNCR 8806  .   Veracruz • 2 ♂, 3 ♀, CL 9-43 mm, CW 12.8-67.3 mm; Municipality of Playa Vicente, Nueva Era; 1 km from  
                <a title="Search Plazi for locations around (long -95.81528/lat 17.689444)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.81528&amp;materialsCitation.latitude=17.689444">Santa</a>
                 Rosa ; 17°41'22"N, 95°48'55"W; alt. 111 m; 7 Mar. 2018; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 34624  .   3 ♂, 3 ♀, CL 11.3-21.6 mm, CW 15.6-31.2 mm; Municipality of Playa Vicente, El Tomate,  
                <a title="Search Plazi for locations around (long -95.86417/lat 17.69786)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.86417&amp;materialsCitation.latitude=17.69786">El Manantial Ranch</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.86417/lat 17.69786)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.86417&amp;materialsCitation.latitude=17.69786">Manzo River</a>
                 ; 17°41'52.3"N, 95°51'51"W; alt. 43 m; 6 Mar. 2018; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 34626  . 
            </p>
            <p>Diagnosis.</p>
            <p>As in Villalobos and Alvarez (2003).</p>
            <p>Distribution.</p>
            <p> This species is distributed in and around the town of Santa  María Jacatepec in northern Oaxaca, Mexico (Fig. 1). </p>
            <p>Remarks.</p>
            <p> As noted by Villalobos and Alvarez (2003), the G1 of  T. jacatepecensis is morphologically similar to those of  T. complanata and  T. lamellifrons (Figs 8, 9), and coincidentally the three species together with  T. ayotzintepecensis sp. nov. are also closely related genetically forming a separate clade in the phylogenetic tree presented herein (Fig. 3). </p>
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	https://treatment.plazi.org/id/2B98807D2D6E57FD8A6735F26EED5491	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA97CE394291581BADF36919C49918C4.text	AA97CE394291581BADF36919C49918C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana lamellifrons (Rathbun 1893)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana lamellifrons (Rathbun, 1893)</p>
            <p>Figs 8F, 9F</p>
            <p> Pseudothelphusa lamellifrons Rathbun, 1893: 654, pl. 75, figs 2-5; 1898: 534, 537.- Young 1900: 221.- Rathbun 1905: 304.-Coiffman, 1939: 108. </p>
            <p> Potamocarcinus lamellifrons .-Ortmann, 1897: 317 (key). </p>
            <p> Pseudothelphusa (Pseudothelphusa) lamellifrons .-Pretzmann, 1965: 4. </p>
            <p> Pseudothelphusa (Tehuana) lamellifrons lamellifrons .-Pretzmann, 1971: 22 (list).- Pretzmann 1972: 107, figs 612-614, 662-664. </p>
            <p> Pseudothelphusa (Tehuana) lamellifrons .-  Rodríguez &amp; Smalley, 1972: 79, fig. 10.-  Türkay 1978: 144. </p>
            <p> Tehuana lamellifrons. -Rodriguez, 1982: 129, fig. 84.- Villalobos 1982: 220 (list).- Villalobos-Hiriart et al. 1993: 284 (list).- Alvarez and Villalobos 1994: 730.-  Álvarez and Villalobos 1997a: 416, 417, fig. 4.17 (list).-  Álvarez and Villalobos 1997b: 438, appx. 4.23 (list).-  Álvarez et al. 1999: 20, fig. 3 (map).-Villalobos and Alvarez 2003: 228 (in key).- Villalobos Hiriart and  Álvarez 2008: 282, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 475, 477, fig. 11 (map).- Guinot and Hendrickx 2014: 477.- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Oaxaca • 1 ♂, CL 50.4 mm, CW 80.6 mm; Municipality of  Asunción Ixtaltepec, Nizanda,  
                <a title="Search Plazi for locations around (long -95.035835/lat 16.687443)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.035835&amp;materialsCitation.latitude=16.687443">Cerro del Naranjo</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.035835/lat 16.687443)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.035835&amp;materialsCitation.latitude=16.687443">Naranjo</a>
                 stream; 16°41'14.8"N, 95°02'09"W; alt. 293 m; 15 Apr. 1999; D. Barreto, V.H. Reynoso leg.; CNCR 16875  .   1 ♂, CL 31.2 mm, CW 43.4 mm; Municipality of  Asunción Ixtaltepec,  
                <a title="Search Plazi for locations around (long -95.38139/lat 16.689999)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.38139&amp;materialsCitation.latitude=16.689999">Naranjo</a>
                 stream; 16°41'24"N, 95°22'53"W; alt. 639 m; 15 Sep. 1997; V.H. Reynoso leg.; CNCR 18951  .   3 ♂, 2 ♀, CL 8.9-40.8 mm, CW 12.4-65 mm; Municipality of  Asunción Ixtaltepec,  
                <a title="Search Plazi for locations around (long -95.01028/lat 16.658333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.01028&amp;materialsCitation.latitude=16.658333">Nizanda</a>
                 , stream; 16°39'30"N, 95°00'37"W; alt. 186 m; 26 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33939  . 
            </p>
            <p>Diagnosis.</p>
            <p>G1 slender, almost straight. Meso-distal lobe conical, in mesial view somewhat compressed caudo-cephalically. Mesial process in transversal position relative to G1 main axis, laying over proximal lobe of caudo-marginal projection, reduced, rounded, superior margin projected distally, without lateral tooth; in cephalic view internal angle triangular, pointing towards distal lobe of caudo-marginal projection (Figs 8F, 9F). Lobes of CMP separated by long incision, lobes not gaping; in cephalic view proximal lobe oblique relative to main axis of gonopod, distal lobe slightly curved laterally. Distal lobe of caudo-marginal projection simple, rounded, projected cephalically; proximal lobe broadly rounded extending proximally, with internal semicircular carina. Apical cavity with elongated field of setae next to lateral margin; opening of spermatic channel in caudal position.</p>
            <p>Type material.</p>
            <p>The syntypes are deposited in the National Museum of Natural History, Smithsonian Institution, Washington, D.C. (USNM 3289).</p>
            <p>Type locality.</p>
            <p>Mexico, Oaxaca, Municipality of Santo Domingo Tehuantepec, Tehuantepec, Tehuantepec River; 16°18'60"N, 95°13'60"W; alt. 60 m (Rathbun, 1893).</p>
            <p>Distribution.</p>
            <p>Along the Pacific versant of the Isthmus of Tehuantepec, in the drainage systems of the Tehuantepec and Zanatepec rivers, in SW Oaxaca, Mexico (Fig. 1).</p>
            <p>Remarks.</p>
            <p> A new diagnosis for  T. lamellifrons is here presented since those of Rathbun (1893) and  Rodríguez and Smalley (1972) are too short omitting important characters of the G1. Other remarks see those for  T. jacatepecensis . </p>
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	https://treatment.plazi.org/id/AA97CE394291581BADF36919C49918C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
628E2EB747D150F4AF7AD5300C15BB6B.text	628E2EB747D150F4AF7AD5300C15BB6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana lamothei Alvarez & Villalobos 1994	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tehuana lamothei Alvarez &amp; Villalobos, 1994</p>
            <p>Figs 8G, 9G</p>
            <p> Tehuana lamothei Alvarez &amp; Villalobos, 1994: 732, figs 2, 4c.-  Álvarez and Villalobos 1995: 93.-Villalobos and Alvarez 2003: 228 (in key).-  Rodríguez and  Magalhães 2005: 356, tab. 1 (list).- Villalobos Hiriart and  Álvarez 2008: 282, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).-  Álvarez et al. 2011b: 289.- Cumberlidge et al. 2014: 147, tab. 7 (list).- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Chiapas • 1 ♂, holotype; CL 18.2 mm, CW 27.5 mm; Municipality of  Ixtacomitán , 1 km from  Ixtacomitán ,  
                <a title="Search Plazi for locations around (long -93.1/lat 17.4)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.1&amp;materialsCitation.latitude=17.4">La Piedra</a>
                 stream; 17°24'00"N, 93°06'00"W; alt. 232 m; 4 Apr. 1986; J.L. Villalobos, J.C. Nates, A.  Cantú , D. Valle; CNCR 5604  .   2 ♂, CL 21-24 mm, CW 32-37.2 mm; Municipality of Tapilula, stream near  
                <a title="Search Plazi for locations around (long -93.02583/lat 17.268055)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.02583&amp;materialsCitation.latitude=17.268055">Tapilula</a>
                 ; 17°16'05"N, 93°01'33"W; alt. 780 m; 20 Apr. 1981; R. Lamothe leg.; CNCR 8812  . 
            </p>
            <p>Diagnosis.</p>
            <p>As in Alvarez and Villalobos (1994).</p>
            <p>Distribution.</p>
            <p>Restricted to a small area in NE Chiapas, Mexico (Fig. 1).</p>
            <p>Remarks.</p>
            <p> As in  T. chontalpaensis . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/628E2EB747D150F4AF7AD5300C15BB6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
81534E809A365D9C9215455BE19F736A.text	81534E809A365D9C9215455BE19F736A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana poglayenorum (Pretzmann 1978)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Tehuana poglayenorum (Pretzmann, 1978)</p>
            <p>Figs 8I, 9I</p>
            <p> Pseudothelphusa (Tehuana) lamellifrons poglayenorum Pretzmann, 1978: 3; 1980: 660, pl. 12, figs 51-55. </p>
            <p> Tehuana poglayenorum .- Villalobos-Hiriart et al. 1993: 284 (list).-  Álvarez and Villalobos 1997a: 416, 417, box 4.17 (list); 1997b: 338, appx. 4.23 (list).-  Álvarez et al. 1999: 20, fig. 3 (map), 23, box 3 (list).-Villalobos and Alvarez 2003: 228 (in key).- Villalobos Hiriart and  Álvarez 2008: 283, 298 (list).-Villalobos and Alvarez 2010: 475, 477, fig. 11 (map).-  Álvarez et al. 2011a: appx. VIII.20, p. 13 (list); 2012: 1078, box 1 (list).- Cumberlidge et al. 2014: 147, tab. 7 (list).- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p> Tehuana poglayenora .- Ng et al. 2008: 177 (list) [error]. </p>
            <p> Tehuana lamellifrons .- Poettinger et al. 2016: 1722, tab. I (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Veracruz • 4 ♂, CL 11.4-26.0 mm, CW 17.0- 41.2 mm; Municipality of San  Andrés Tuxtla, Basura River; 18°31'55"N, 95°03'30"W; alt. 33 m; 19 Jul. 1998; R. Robles, C. Graham leg.; CNCR 17422. 1 ♀, CL 15.8 mm, CW 22.9 mm; same locality as previous record; 18 Jul. 1986; J.L. Villalobos, F.  Álvarez leg.; CNCR 13140. 3 ♂, 5 ♀, CL 8.9-28.8 mm, CW 13.1-45.2 mm; same locality as previous record; 4 Oct. 1994; J.L. Villalobos, F.  Álvarez leg.; CNCR 13187. 7 ♂, 1 ♀, CL 9.6-21.8 mm, CW 14.7-36.4 mm; same locality as previous record; 24 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33931. 4 ♂, 11 ♀, CL 11.8-19.5 mm, CW 12.8-33.2 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">Municipality of Santiago Tuxtla</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">Tapalapan River</a>
                 ; 18°32'00"N, 95°18'00"W; alt. 393 m; 17 Apr. 1957; H. Hobbs, A. Villalobos leg.; CNCR 336. 5 ♀, CL 25.1-36.1 mm, CW 40.6-59.1 mm; same locality as previous record; 23 May 1955; A. Villalobos leg.; CNCR 338. 4 ♂, 4 ♀, CL 7.3-14 mm, CW 10.8-20.3 mm; same locality as previous record; 17 April 1957; H. Hobbs, A. Villalobos leg.; CNCR 386. 2 ♂, CL 10.9-11.9 mm, CW 12.4-18.8 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">Otapan River</a>
                 ; 18°26'00"N, 95°12'00"W; alt. 379 m; 22 Sep. 1955; G.  Pérez leg.; CNCR 337. 2 ♂, CL 11.9-20.7 mm, CW 21.9-30.3 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 , Laguna Escondida; 18°35'00"N, 95°05'00"W; alt. 76 m; 1 Aug. 1985; J.L. Villalobos, M.D. Valle, P. Schmidtsdorf leg.; CNCR 4473. 15 ♂, 10 ♀, CL 7.4-21.7 mm, CW 10.4-34.9 mm; same locality as previous record 13 Jun. 1985; C. Nates, J.L. Villalobos leg.; CNCR 4709. 1 ♂, 3 ♀, CL 8.2-25.6 mm, CW 11.6-41 mm; same locality as previous record; 11 Jul. 1994; J.L. Villalobos, F.  Álvarez leg.; CNCR 5303. 4 ♂, 3 ♀, CL 25.4-32.3 mm, CW 42.2-53.2 mm; same locality as previous record; 24 Feb. 1989; M. Santiago leg.; CNCR 10220. 2 ♂, 2 ♀, CL 11-19.4 mm, CW 15.5-30.5 mm; same locality as previous record; 17 Jul. 1985; J.L. Villalobos, F.  Álvarez leg.; CNCR 12908. 6 ♂, 4 ♀, CL 11.7-18.7 mm, CW 17.7-45.9 mm; same locality as previous record; 5 Aug. 1994; J.L. Villalobos leg.; CNCR 12964. 2 ♂, 2 ♀, CL 6.8-11.3 mm, CW 9.3-17.0 mm; same locality as previous record; 10 Jul. 1986; F.  Álvarez leg.; CNCR 13138. 1 ♂, CL 20.0 mm, CW 32.3 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 , Playa Escondida; 18°35'00"N, 95°03'00"W; alt. 6 m; 28 Feb. 1986; A.  Cantú leg.; CNCR 5782. 3 ♂, 5 ♀, CL 13.3-17.2 mm, CW 20.0-27.0 mm; same locality as previous record; 28 Feb. 1986; A. Cervantes, J.  García , A.  Cantú leg.; CNCR 5788. 1 ♂, 4 ♀, CL 9.2-20.9 mm, CW 13.1-32.1 mm; same locality as previous record; 28 Feb. 1986; R. Lamothe leg.; CNCR 8821. 6 ♂, 11 ♀, CL 6.3-20.8 mm, CW 9.1-32.8 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 , trail to Laguna Escondida; 18°35'00"N, 95°04'00"W; alt. 108 m; 24 Feb. 1989; M. Santiago leg.; CNCR 10222. 1 ♂, 2 ♀, CL 11.8-21.7 mm, CW 17.3-35.1 mm; same locality as previous record; 3 Aug. 1994; F.  Álvarez , J.L. Villalobos leg.; CNCR 12967. 9 ♂, 5 ♀, CL 9.7-28.6 mm, CW 14.3-49.7 mm; same locality as previous record; 23 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33927. 13 ♂, 21 ♀, CL 9.9-30.0 mm, CW 14.7-50.4 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 ,  Lázaro Cárdenas ; 18°34'00"N, 95°06'00"W; alt. 359 m; 9 Jul. 1991; A. Cruz leg.; CNCR 12302. 1 ♂, CL 20.3 mm, CW 32.3 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">Cuetzalapan River</a>
                 ; 18°24'00"N, 95°00'00"W; alt. 220 m; 13 Jul. 1986; F.  Álvarez leg.; CNCR 12555. 2 ♂, 1 ♀, CL 11.3-23.6 mm, CW 18.0- 37.7 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">La Palma River</a>
                 ; 18°33'00"N, 95°03'00"W; alt. 77 m; 12 Jul. 1986; F.  Álvarez leg.; CNCR 12896. 1 ♂, 1 ♀, CL 21.1-23.0 mm, CW 35.0- 38.5 mm;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">Balzapote River</a>
                 ; 18°36'00"N, 95°04'00"W; alt. 54 m;  
                <a title="Search Plazi for locations around (long -95.066666/lat 18.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.066666&amp;materialsCitation.latitude=18.6">
Municipality of San 
Andres
Tuxtla
</a>
                 ,   Veracruz; 14 Jul. 1986; F.  Álvarez leg.; CNCR 12906. 7 ♂, 3 ♀, CL 13.0- 20.3 mm, CW 20.0- 32.5 mm; Municipality of San  Andrés Tuxtla,  Máquinas River; 18°37'00"N, 95°05'00"W; alt. 143 m; 1 Aug. 1994; F.  Álvarez , J.L. Villalobos leg.; CNCR 12961. 1 ♂, 2 ♀, CL 10.1-18.3 mm, CW 16.1-21.6 mm;  
                <a title="Search Plazi for locations around (long -95.35/lat 18.45)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.35&amp;materialsCitation.latitude=18.45">
Municipality of San 
Andres
Tuxtla
</a>
                 , Zacatal Lagoon; 18°35'00"N, 95°06'00"W; alt. 263 m; 14 Oct. 1994; F.  Álvarez , J.L. Villalobos leg.; CNCR 13188. 4 ♂, CL 11.8-23.4 mm, CW 17.5-37.2 mm;  
                <a title="Search Plazi for locations around (long -95.35/lat 18.45)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.35&amp;materialsCitation.latitude=18.45">Municipality of Santiago Tuxtla</a>
                 ,  
                <a title="Search Plazi for locations around (long -95.35/lat 18.45)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.35&amp;materialsCitation.latitude=18.45">Simapan River</a>
                 ; 18°27'00"N, 95°21'00"W; alt. 336 m; 3 May 1995; J.L. Villalobos leg.; CNCR 13346  . 
            </p>
            <p>Diagnosis.</p>
            <p>G1 slender, in mesial view distal third inclined cephalically. Meso-distal lobe conical, tip rounded, large relative to size of other apical structures (Fig. 9I). Mesial process well developed; lateral margin with triangular, acute tooth; distal margin broadly rounded; internal angle with triangular projection in cephalic position. Lobes of the caudo-marginal projection elongated, overlapping along most of their length; proximal lobe curved cephalically, distal lobe straight, oriented forward or with the tip moderated curved proximally, slightly longer than proximal one. In mesial view, distal lobe with superior margin inclined proximally; distally, falling over proximal lobe. Internal carina of proximal lobe rounded, prominent, separated from lobe. Apical cavity with elongated field of setae, opening of spermatic channel in caudal position.</p>
            <p>Type material.</p>
            <p>The holotype (NHMW 4066) and paratypes (NHMW 4067) are deposited in the Natural History Museum in Vienna, Austria.</p>
            <p>Type locality.</p>
            <p> Mexico, Veracruz, Municipality of San  Andrés Tuxtla, Basura River; 18°32'00"N, 95°03'00"W; alt. 33 m (Pretzmann, 1978). </p>
            <p>Distribution.</p>
            <p> Tehuana poglayenorum is distributed in the north-central section, of the Los Tuxtlas region. </p>
            <p>Remarks.</p>
            <p> Tehuana poglayenorum is the most widely distributed and abundant freshwater crab in the whole Los Tuxtlas Mountain Range. It belongs to clade c, where all the species from Los Tuxtlas are grouped (Fig. 1). We noted there small morphological variations through its distribution range: in the carapace with setae or without them, and in the G1 in the caudo-marginal projection and mesial process. </p>
            <p> Poettinger et al. (2016) presented the sequence of the16S mitochondrial gene (KU578859) as belonging to  T. lamellifrons ; however, the two females in the lot NHMW 4067 (from  Río Basura, state of Veracruz) are the paratypes of  T. poglayenorum designated by Pretzman (1978, 1980). | </p>
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	https://treatment.plazi.org/id/81534E809A365D9C9215455BE19F736A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
AA0A15F3CFCC5A6EA126DB2DBB6231EF.text	AA0A15F3CFCC5A6EA126DB2DBB6231EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana Rodriguez & Smalley in Smalley 1970	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> Genus  
Tehuana 
Rodriguez
&amp; Smalley in Smalley, 1970
</p>
            <p> Pseudothelphusa (Tehuana) Rodríguez &amp; Smalley in Smalley, 1970: 106 (in key).- Pretzmann 1971: 22.- Pretzmann 1972: 107; 1980: 660.-  Rodríguez and Smalley 1972: 77.-  Türkay 1978: 144. </p>
            <p> Tehuana .-Rodriguez, 1982: 129; 1986: 66.- Villalobos-Hiriart et al. 1993: 285 (list).- Alvarez and Villalobos 1994: 730.-Alvarez et al. 1996: 129.-Villalobos and Alvarez 2003: 228.-  Álvarez et al. 2005: 191.- Ng et al. 2008: 177 (list).- Villalobos Hiriart and  Álvarez 2008: 279.-Villalobos and Alvarez 2010: 474.-  Álvarez et al. 2011b: 289.-  Mejía-Ortíz et al. 2011: 19 (list).- Guinot and Hendrickx 2014: 477, 478, tab. 1.- Alvarez and Villalobos 2016: 244.- Villalobos-Hiriart et al. 2019: 156, tab.1 (list).-  Álvarez et al. 2020: 12, tab. 4, 20 (list). </p>
            <p>Type species.</p>
            <p> Pseudothelphusa lamellifrons Rathbun, 1893 [by original designation]. </p>
            <p>Diagnosis.</p>
            <p> Carapace with dorsal surface flat, smooth, punctate, with small granulations adjacent to antero-lateral margin. Front vertically deflexed, smooth, superior border formed by low rounded tubercles, medially divided by V-shaped notch reaching inferior margin; inferior margin not distinguishable dorsally, in ventral view projected, widely bilobed. Third maxillipeds with ischium trapezoidal, slightly longer than wide; merus smaller than ischium, anterolateral margin rounded, with anterior shallow concavity; ratio exopod/ischium length 0.6 to 0.8. G1 slender, proximal half cylindrical, compressed distally, with a meso-distal lobe conical, well developed on mesial surface; principal axis with medial constriction on lateral surface and twisted towards medial suture of ventral sternites (Fig. 2). In mesial view, proximal half with marginal plate closing spermatic channel, border facing marginal suture with proximal fringe of long and stout setae; distal half inclined towards cephalic surface; caudal surface fused with marginal plate and expanded cephalically to form the bilobed caudo-marginal projection, distal lobe well-developed (except in  T. lamothei where it is reduced), subrectangular with cephalic border rounded, distal portion can be curved proximally and separated from proximal lobe by an incision; proximal lobe well developed or reduced, trapezoidal, circular, ax-shaped or elongated proximally to reach and sometimes overlap gonopod principal axis (Fig. 2B). Meso-distal lobe arising from distocaudal angle of mesial surface, conical, subacute, or flattened caudo-cephalically, distally rounded. In cephalic view, mesial process in longitudinal position, oval shaped, closing apical cavity, distal margin widely rounded, lateral margin straight, armed with strong median spine (  T. chontalpaensis ,  T. lamothei ), or with moderate or reduced triangular tooth (  T. diabolis ,  T. col sp. nov.,  T. poglayenorum ,  T. veracruzana ); or in transversal position, reduced, spoon shaped, with lateral margin smooth without spine or tooth (  T. ayotzintepecensis sp. nov.,  T. complanata ,  T. jacatepecensis ,  T. lamellifrons ). Caudo-marginal projection with the two lobes directed cephalically, distal one slightly curved mesially; proximal one, ax-shaped or elongated proximally, and inclined in different gradations; carina of inner surface partially visible. Meso-distal lobe arise from caudal corner of G1 mesial surface, commonly is well developed, conical, with subacute or rounded apex. In lateral view, distal half inclined cephalically. Caudo-marginal projection with distal lobe (partially or totally visible; inner surface of proximal lobe with circular or semicircular strong carina, extending to different extents over proximal surface. In caudal view, distal third straight, apical cavity distally directed, caudal surface ending distally in a wide and shallow concavity. Caudo-marginal projection with the distal crest of mesial surface higher or at the same level as lateral. Meso-distal lobe well developed, conical, with the apex rounded or subacute rounded, its position with respect the distal crest of mesial surface, could variate through different species. Mesial process as a longitudinal apical plate or transversal reduced with the distal surface excavated, in the first case, oval shaped, the distal margin could be widely rounded, and the lateral margin straight and armed with a strong median spine, or with a moderate or reduced triangular tooth; in the second, only partially visible. In distal view, apical cavity U-shaped, opening cephalically. Field of apical setae with 20 to 60 setae; aperture of spermatic channel in caudal position; central crest ending cephalically in acute and triangular or rounded internal angle of mesial process (Fig. 2D, E). </p>
            <p>Remarks.</p>
            <p> The phylogeny presented shows  Tehuana to be closely related to  Disparithelphusa (Fig. 2); however, there are strong differences in the G1 morphology: in  Tehuana it is stouter, the lateral notch on the main shaft is in the middle, the internal surface of the proximal lobe of the caudo-marginal projection is marked with a strong carina, and the meso-distal lobe arises from the distocaudal corner of the mesial surface.  Álvarez et al. (2020) in their phylogenetic analysis of the family also recovered  Tehuana and  Disparithelphusa as sister taxa. </p>
            <p>Distribution.</p>
            <p> The species of  Tehuana are distributed in southeastern Mexico covering the Isthmus of Tehuantepec region (Fig. 1) in Chiapas, Oaxaca, Tabasco, and Veracruz. </p>
            <p>Species included.</p>
            <p> Tehuana ayotzintepecensis sp. nov.;  T. col sp. nov.;  T. complanata (Rathbun, 1905);  T. chontalpaensis Villalobos &amp; Alvarez, 2003;  T. diabolis (Pretzmann, 1978);  T. jacatepecensis Villalobos &amp; Alvarez, 2003;  T. lamellifrons (Rathbun, 1893);  T. lamothei Alvarez &amp; Villalobos, 1994;  T. poglayenorum (Pretzmann, 1978);  Tehuana veracruzana (  Rodríguez &amp; Smalley in Smalley, 1970). </p>
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</html>
	https://treatment.plazi.org/id/AA0A15F3CFCC5A6EA126DB2DBB6231EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
E6C45F73407E58EB982CA120DF10A90C.text	E6C45F73407E58EB982CA120DF10A90C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tehuana veracruzana (Rodriguez & Smalley in Smalley 1970)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div>
            <p> 
Tehuana veracruzana (
Rodriguez
&amp; Smalley in Smalley 1970)
</p>
            <p>Figs 8J, 9J</p>
            <p> Pseudothelphusa (Tehuana) veracruzana Rodríguez &amp; Smalley in Smalley, 1970: 100, fig. 11. Pretzmann 1972: 108.-  Rodríguez and Smalley 1972: 77, fig. 9.-  Türkay 1978: 148, fig. 4.- Guinot and Hendrickx 2014: 478, tab. 1 (list). </p>
            <p> Tehuana veracruzana .-Rodriguez, 1982: 131, fig. 86.- Villalobos 1982: 221 (list).-  Rodríguez 1986: 59, fig. 10l, 61, fig.12- Villalobos-Hiriart et al. 1993: 284.-  Álvarez and Villalobos 1995: 93; 1997a: 416, 417, box 4.17 (list); 1997b: 338, appx. 4.23 (list).-  Álvarez et al. 1999: 20, fig. 3 (map), 23, box 3 (list).- Villalobos and  Álvarez 2003: 228 (in key).- Villalobos Hiriart and  Álvarez 2008: 284, 298 (list).- Ng et al. 2008: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).-  Álvarez et al. 2011a: appx. VIII.20, p. 13 (list); 2012: 1078, box 1 (list).- Guinot and Hendrickx 2014: 477.- Cumberlidge et al. 2014: 147, tab. 7 (list).- Alvarez and Villalobos 2016: 254, tab. 8.1 (list).- Villalobos-Hiriart et al. 2019: 156, tab.1 (list). </p>
            <p>Material examined.</p>
            <p>
                  Mexico - Veracruz • 1 ♂, holotype; CL 48.0 mm, CW 28.2 mm; Municipality of Catemaco,  
                <a title="Search Plazi for locations around (long -95.09667/lat 18.333334)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.09667&amp;materialsCitation.latitude=18.333334">
Zapoapan de 
Cabanas
Stream
</a>
                 ; 18°20'00"N, 95°05'48"W; alt. 518 m; 15 Apr. 1957; A. Villalobos, H.H. Hobbs leg.; CNCR 335  .  1 ♂, 1 ♀, paratypes; CL 23.8-28.2 mm, CW 37.2-47.5 mm; same data as for holotype; CNCR 335 .   1 ♂, 1 ♀, CL 15.7-27.8 mm, CW 24.8-45.2 mm; Municipality of Catemaco, stream near  
                <a title="Search Plazi for locations around (long -95.039444/lat 18.335835)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.039444&amp;materialsCitation.latitude=18.335835">
Zapoapan de 
Cabanas</a>
                 ; 18°20'09"N, 95°02'22"W; alt. 629 m; 25 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33934  .   1 ♂, 3 ♀, CL 12.1-20.5 mm, CW 18.4-32.5 mm; Municipality of Catemaco, stream near  
                <a title="Search Plazi for locations around (long -95.070274/lat 18.342222)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.070274&amp;materialsCitation.latitude=18.342222">
Zapoapan de 
Cabanas</a>
                 ; 18°20'32"N, 95°04'13"W; alt. 364 m; 25 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33932  .   1 ♂, 2 ♀, CL 9.2-12.5 mm, CW 14.5-21.6 mm; Municipality of Catemaco, stream near  
                <a title="Search Plazi for locations around (long -95.05056/lat 18.321667)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.05056&amp;materialsCitation.latitude=18.321667">
Zapoapan de 
Cabanas</a>
                 ; 18°19'18"N, 95°03'02"W; alt. 364 m; 25 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33937  .   1 ♂, 1 ♀, CL 9.8-20.5 mm, CW 14.4-32.2 mm; Municipality of Catemaco, road  
                <a title="Search Plazi for locations around (long -94.96833/lat 18.297777)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.96833&amp;materialsCitation.latitude=18.297777">
Zapoapan de 
Cabanas-Zoteapan</a>
                 ; 18°17'52"N, 94°58'06"W; alt. 848 m; 25 Apr. 2017; J.L. Villalobos, I.A. Toledano, E. Moreno leg.; CNCR 33955  . 
            </p>
            <p>Diagnosis.</p>
            <p> As in  Rodríguez and Smalley (1972). </p>
            <p>Distribution.</p>
            <p>This species occurs in the southeastern portion of the Los Tuxtlas Mountain Range, in an area that starts sloping towards the coastal plain of southern Veracruz.</p>
            <p>Remarks.</p>
            <p> Tehuana veracruzana is easily distinguishable from the rest of its congeners due to the very large proximal lobe of the CMP of the G1 (Fig. 8). Although it exhibits an extreme form of the G1 it is genetically closely related to the other species from Los Tuxtlas (Fig. 1). </p>
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	https://treatment.plazi.org/id/E6C45F73407E58EB982CA120DF10A90C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Moreno-Juarez, Eric G.;Villalobos, Jose Luis;Alvarez, Fernando	Moreno-Juarez, Eric G., Villalobos, Jose Luis, Alvarez, Fernando (2022): Revision of the freshwater crabs of the genus Tehuana Rodriguez & Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species. ZooKeys 1117: 1-35, DOI: http://dx.doi.org/10.3897/zookeys.1117.85362, URL: http://dx.doi.org/10.3897/zookeys.1117.85362
