taxonID	type	description	language	source
03F15F55CE59FFB2FF03D979FC3AFC60.taxon	synonymic_list	The genus comprises 13 species (including a new Japanese species described in this paper), of them six occur in the Nearctic region (Horstmann 2013 b) and seven in the Palaearctic region (Yu et al. 2016; Vas 2019). Three species, A. acutatus Khalaim, A. platyurus (Strobl, 1904) and A. rufobasalis Horstmann & Kolarov, are known from the Russian Far East (Khalaim 2007), and one species is described here from Japan. This is the first record of the genus Allophroides from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB2FF03D979FC3AFC60.taxon	discussion	Nothing is known about host range of any species in the genus.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB0FF03D8C3FAC0FD6E.taxon	description	(Figs 1 – 9)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB0FF03D8C3FAC0FD6E.taxon	materials_examined	Material examined. Holotype female (EUM) Japan, Hokkaido I., Sapporo, Hitsujigaoka, 43 ° 00 ’ N, 141 ° 24 ’ E, Malaise trap, 3 – 10. V. 2007, coll. K. Konishi. Paratype. JAPAN. 1 ♀ (ZISP) same data as holotype, but 43 ° 00 ’ 11 ” N, 141 ° 24 ’ 55 ” E, 16 – 23. V. 2011.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB0FF03D8C3FAC0FD6E.taxon	description	Description. Female. Body length 4.7 mm. Fore wing length 3.7 mm. Head, in dorsal view, roundly constricted posterior to eyes (Fig. 3); gena 0.9 – 0.95 × as long as eye width. Eyes with short sparse setae (Fig. 2). Clypeus about 3.0 × as broad as long, in front view lenticular, with ventrolateral margins distinctly impressed (Fig. 2), separated from face by deep and sharp furrow; clypeus with sparse shallow punctures, convex and finely granulate in upper 1 / 3, flat and polished in lower 2 / 3. Palpi formula 4 + 3. Mandible robust, weakly tapered in basal 0.3 and with upper and lower margins subparallel in apical 0.7; upper tooth slightly longer than the lower. Malar space 0.8 – 0.9 × as long as basal mandibular width. Antennal flagellum (Fig. 6) slightly tapered towards apex, with 23 – 25 flagellomeres; basal and median flagellomeres slightly elongate, subapical flagellomeres as long as broad; flagellomeres 4 to 6 (or 7) with distinct subapical finger-shaped structures on outer surface. Face with weak elongate median prominence. Face and frons distinctly granulate, dull, impunctate or with very fine inconspicuous punctures. Vertex and gena granulate, with fine and sparse punctures. Occipital carina complete. Mesoscutum finely and shallowly granulate, dull to weakly shining, with fine and sparse punctures. Notaulus with strong irregular wrinkles on anterolateral side of mesoscutum (Figs 3, 4). Scutellum with lateral longitudinal carinae present at its front 0.3 (Fig. 8). Mesopleuron finely punctate on nearly smooth and shining background in upper part and ventrally, strongly granulate (partly with irregular wrinkles), impunctate and dull in lower part, with irregular wrinkles peripherally (Fig. 4). Epicnemial carina with upper end distinctly curved to reach front margin of mesopleuron. Foveate groove moderately deep, weakly curved, extending in posterior 0.8 of mesopleuron, with strong transverse wrinkles (Fig. 4). Propodeal spiracle separated from pleural carina by about 1.5 × diameter of spiracle (Fig. 4). Propodeum almost entirely with rather strong irregular wrinkles; on the apical area wrinkles are mostly transverse (Fig. 8); basal area broad, transverse (in holotype) or quadrate (in paratype), about 0.3 × as long as apical area (Fig. 8); transverse carina strong laterally and indistinct mediodorsally, thus the basal area is not clearly delineated posteriorly (Fig. 8); apical area somewhat impressed; apical longitudinal carinae strong posteriorly and indistinct (because of irregular wrinkles) anteriorly. Fore wing (Fig. 7) with second recurrent vein (2 m-cu) distinctly postfurcal, unpigmented in anterior 0.55 and distinct in posterior 0.45. Intercubitus (2 rs-m) short and thick, shorter than abscissa of cubitus between intercubitus and second recurrent vein (abscissa of M between 2 rs-m and 2 m-cu). First abscissa of radius (Rs + 2 r) weakly curved, as long as width of pterostigma. Pterostigma broad, subtriangular. First and second abscissae of radius (Rs + 2 r and Rs) meeting at slightly acute angle. Metacarpus (R 1) not reaching apex of fore wing. Hind wing with nervellus (cu 1 & cu-a) weakly reclivous. Legs moderately slender. Tarsal claws not pectinate. First tergite 2.1 × as long as posteriorly broad, strongly longitudinally striate dorsally and laterally (Fig. 5), with only posterior dorsal margin smooth (Fig. 9); first tergite depressed, in cross-section centrally trapeziform (lateral carinae strong in anterior half of tergite, petiole dorsally flat). Glymma large and deep, situated in anterior 0.35 – 0.4 of tergite, joining by strong furrow to ventral part of postpetiole (Fig. 4). Second tergite distinctly transverse, 0.85 × as long as anteriorly broad (Fig. 9). Thyridial depression small, oval, transverse. Ovipositor robust, weakly and evenly bent upwards, with two dorsal subapical teeth and several distinct teeth ventrally (Fig. 5); sheath almost twice as long as first tergite (Fig. 1). Head, mesosoma and first tergite of metasoma black; lower 2 / 3 of clypeus reddish black. Mouthparts yellow, palpi brown. Mandible predominantly brownish yellow, reddish at base and with teeth reddish black. Antenna black, apical margins of scape and pedicel pale. Tegula brownish black, yellowish apically. Wings slightly infumate with brown. Pterostigma dark brown (in holotype) or brown (in paratype), with yellowish markings proximally and distally. Legs with all coxae black, reminder yellowish brown to dark brown. Metasoma posterior to first tergite dark brown. Male. Unknown.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB0FF03D8C3FAC0FD6E.taxon	etymology	Etymology. The species is named in honor of the Japanese expert in Ichneumonidae (Hymenoptera), Kyohei Watanabe (KPM-NK).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE59FFB0FF03D8C3FAC0FD6E.taxon	distribution	Distribution. Japan (Hokkaido I.). Comparison. The new species may easily be recognized by its robust ovipositor with two distinct dorsal subapical teeth (Fig. 5), antennal flagellum with 23 – 25 flagellomeres (Fig. 6), and simple tarsal claws. Similar ovipositor is known in the European A. boops (Gravenhorst), but in this species the ovipositor is much shorter (sheath about as long as first tergite), flagellum with 14 – 15 flagellomeres, and tarsal claws distinctly pectinate.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB7FF03DFFEFD6CFA2A.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Shizuoka Pref., Honkawane-cho, Mt. Yamainudan, 35 ° 07 ’ 48 ” N, 138 ° 02 ’ 19 ” E, 1200 – 1440 m, 14. VI. 2008, coll. K. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB7FF03DFFEFD6CFA2A.taxon	distribution	Distribution. Holarctic region, Taiwan, Vietnam and India (Uttar Pradesh); in Japan known from Honshu, Shikoku and Kyushu islands (Khalaim 2015 c: 426).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB7FF03D812FAAAFB4A.taxon	description	Very little is known about host range of Barycnemis. In Europe, the genus has been recorded from the coleopteran genera Byrrhus L. (Byrrhidae) and Bledius Leach (Staphylinidae) (Horstmann 1981; Wyatt & Foster 1989), and in the Nearctic region one species was reared from a Pissodes species (Curculionidae) (Horstmann 2010).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB7FF03DD1EFE94F90A.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♂ (KPM-NK) Nagano Pref., Outaki-mura [Ôtaki], Mt. Ontake-san, Tanohara, 1800 m, 9. VIII. 2007, coll. K. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB7FF03DD1EFE94F90A.taxon	distribution	Distribution. Widespread Holarctic species; in Japan known from Hokkaido and Honshu islands (Khalaim 2015 c: 429).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5CFFB6FF03DC3EFCA0FE62.taxon	description	Species of Diaparsis have been reared from the beetle families Buprestidae, Cerambycidae, Chrysomelidae and Curculionidae, and in Europe one species has been reared from sawfly genera Pontania Costa and Phyllocolpa Benson (Hymenoptera: Tenthredinidae) (Yu et al. 2016).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DAD6FAEEFD42.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 2 ♀ (ZISP) Hyôgo Pref., Kobe, Rokko Mts, Maya Mt., 28. VIII – 25. IX. 2005, coll. S. A. Belokobylskij. 1 ♀ (ZISP) Yoyama Pref., Toyama City, Arimine, Jûrôdani [Inonedani], 1200 m, Malaise trap, 1 – 8. IX. 2009, coll. K. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DAD6FAEEFD42.taxon	distribution	Distribution. Russian Far East, South Korea; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03D9F6FB75FC4E.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Niigata Pref., Sado I., Hakuundai — Mt. Mvouken-zan, 38 ° 05 ’ 20 ” N, 138 ° 20 ’ 15 ” E, 830 – 1000 m, 10. X. 2010, coll. K. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03D9F6FB75FC4E.taxon	distribution	Distribution. Russian Far East; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DEABFB24F9BA.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Niigata Pref., Nagaoka, Kagimachi, sweep on Pinus, 4. V. 2015, coll. So Shimizu.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DEABFB24F9BA.taxon	distribution	Distribution. South Korea; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DDAFFA3DF89A.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 5 ♀ (4 ♀ in KPM-NK, 1 ♀ in ZISP) Niigata Pref., Nagaoka, Kagimachi [Kajimachi], sweep on Pinus, 4. V. 2015, coll. So Shimizu.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DDAFFA3DF89A.taxon	distribution	Distribution. Europe, South Korea; in Japan known from Honshu and Kyushu islands (Khalaim 2017 b: 46).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03DDAFFA3DF89A.taxon	biology_ecology	Biology. Reared from Xyela graeca Stein and probably X. curva Benson in Europe (Khalaim & Blank 2011).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5DFFB6FF03D8FBFC7EFABE.taxon	description	Species of the genus are parasitoids of xyelid sawflies (Xyelidae: mainly Xyela spp.) on coniferous plants in Europe (Khalaim & Blank 2011) and North America (Horstmann 2013 b).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DB9AFE98FE62.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀, 3 ♂ (1 ♀, 3 ♂ in KPM-NK; 1 ♂ in ZISP) Niigata Pref., Nagaoka, Kagimachi [Kajimachi], sweep on Pinus, 4. V. 2015, coll. So Shimizu. 7 ♂ (KPM-NK) Tokyo-to, Shibuya-ku, Yoyogi, Yoyogikoen, 35 ° 40 ’ 17 ” N, 139 ° 41 ’ 30 ” E, 29. III. 2016, coll. Yu Kato.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DB9AFE98FE62.taxon	distribution	Distribution. Russian Far East, South Korea, China (Liaoning); in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DAD6FBEBFD26.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Niigata Pref., Nagaoka, Kagimachi [Kajimachi], sweep on Pinus, 4. V. 2015, coll. So Shimizu.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DAD6FBEBFD26.taxon	distribution	Distribution. Europe, Turkey, Israel, Russian Siberia and Far East; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DAD6FBEBFD26.taxon	biology_ecology	Biology. Reared from Xyela julii (Brebisson) in Europe (Khalaim & Blank 2011).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03D812FBEEFC22.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 3 ♀ (2 ♀ in KPM-NK, 1 ♀ in ZISP) Niigata Pref., Nagaoka, Kagimachi [Kajimachi], sweep on Pinus, 4. V. 2015, coll. So Shimizu.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03D812FBEEFC22.taxon	distribution	Distribution. South Korea; in Japan known from Honshu I. (Khalaim 2017 b: 48).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB5FF03DF16FDA9FA92.taxon	description	In Europe, species of the genus are known as parasitoids of sap beetle larvae (Nitidulidae: Meligethes spp.) (Horstmann 1981; Yu et al. 2016).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB4FF03DEC6FF56F8D9.taxon	description	(Figs 10 – 19)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB4FF03DEC6FF56F8D9.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (ZISP) Sapporo, Usubetsu, Malaise trap, 29. IV – 24. V. 2012, coll. N. Kuhara.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB4FF03DEC6FF56F8D9.taxon	diagnosis	Diagnosis. Head and mesosoma densely granulate, dull, impunctate. Head roundly constricted behind eyes in dorsal view (Fig. 15); lower part gena distinctly widened in lateral view (Fig. 14). Eye with short setae (Fig. 15). Clypeus lenticular. Mandible robust, with upper tooth somewhat longer than the upper tooth. Flagellum lenticular, with 19 flagellomeres (Fig. 12). Malar space distinctly longer than basal mandibular width (Fig. 14). Notaulus absent (Figs 14, 15). Foveate groove of mesopleuron indistinct, slightly impressed, with weak transverse wrinkles (Fig. 14). Propodeum with moderately broad rectangular basal area which is sometimes indistinct (Fig. 16). Fore wing (Fig. 13) with pterostigma broad, metacarpus (R 1) short and not reaching tip of the wing, and vein 2 m-cu strongly antefurcal. First tergite distinctly striate laterally (Fig. 17). Second tergite about 1.5 × as long as anteriorly broad (Fig. 18). Thyridial depression 1.5 – 2.0 × as long as broad, with posterior end rounded (Fig. 18). Ovipositor slender, weakly upcurved, with apical part very thin, needle-shaped (Fig. 19); sheath 1.25 × as long as first tergite. Antenna, head (except lower half of clypeus and mandible), mesosoma, all coxae and first metasomal segment brownish black to black; metasoma posterior to first tergite and all legs (except coxae) brown to dark brown (Fig. 10).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE5EFFB4FF03DEC6FF56F8D9.taxon	distribution	Distribution. Europe, Russian Far East, China (Shanxi); in Japan known from Hokkaido I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE50FFBBFF03DE6BFCDCF892.taxon	description	(Figs 20 – 21)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE50FFBBFF03DE6BFCDCF892.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (NIAES) Gunma Pref., Katashina-mura, Tsuchiide, 11. VII. 1988, coll. H. Makihara. Shikoku I.: 1 ♂ (NIAES) Tokushima Pref., Mt. Tsurugisan, 3 – 4. VI. 1981, coll. T. Goto. 1 ♂ (ZISP) Tokushima Pref., Mt. Tsurugisan, 1300 – 1500 m, 3. VI. 1981, coll. H. Takemoto. 1 ♂ (NIAES) same data, but 4. VI. 1981. 1 ♂ (NIAES) Ehime, Koyayama, Odamiyama, 1. V. 1997, coll. E. Yamamoto. Kyushu I.: 1 ♂ (NIAES) Fukuoka, Fukuoka-shi, Mt. Tachibana, 21. IV. 1977, coll. K. Konishi.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE50FFBBFF03DE6BFCDCF892.taxon	diagnosis	Diagnosis. Male (First record). Head with eyes displaced downwards in lateral view. Mesosoma compressed laterally (though less than in female). Malar space very short. Flagellum with 15 – 17 flagellomeres. Notaulus with a strong longitudinal wrinkle on anterolateral side of mesoscutum (as in female). Mesopleuron with almost linear narrow foveate groove (as in female); mesopleuron centrally (above foveate groove) with small but distinct punctures on nearly smooth background to virtually impunctate and shallowly granulate. Propodeum long; basal area long and narrow (as in female). First metasomal tergite in lateral view evenly convex, laterally striate (as in female).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE50FFBBFF03DE6BFCDCF892.taxon	distribution	Distribution. Japan (Honshu, Shikoku and Kyushu islands).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE51FFBAFF03DB9AFB75FD42.taxon	description	(Figs 22 – 28)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE51FFBAFF03DB9AFB75FD42.taxon	discussion	Remarks. This species was described in the identification key from a single female (Fig. 22) collected in Primorsky Reg. of Russia (Khalaim 2007: 585). Two females are recorded in this work from Japan; brief description and colour photographs of the species are provided.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE51FFBAFF03DB9AFB75FD42.taxon	diagnosis	Diagnosis. Flagellum with 14 flagellomeres, entirely dark brown to black (Fig. 24). Mesopleuron with thin, sharp and almost straight foveate groove in its anterior 0.6 (Fig. 25); mesopleuron distinctly punctate on smooth and shining background centrally (above foveate groove). Propodeum with basal area distinctly widened anteriorly; apical area in anterior part narrow, widened posteriorly (Fig. 26). First metasomal tergite polished, round in crosssection (Fig. 25). Second tergite very long, 3.2 times as long as broad anteriorly (Fig. 27). Thyridial depression very long (Fig. 27). Ovipositor with very weak dorsal subapical notch or depression (Fig. 28).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE51FFBAFF03DB9AFB75FD42.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 2 ♀ (NIAES) Yamagata Pref., Oguni-machi, Chôjagahara, 7. VI. 1980, coll. K. Maetô.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE51FFBAFF03DB9AFB75FD42.taxon	distribution	Distribution. Russian Far East; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE52FFB8FF03DC4CFF51FE3A.taxon	description	In Europe, some species of Probles have been recorded as parasitoids of the beetle families Attelabidae, Ciidae, Curculionidae, Endomycidae and Melandryidae (Horstmann 1971, 1981; Blommers & Vaal 2002), and one species probably parasitises in gall-forming sawfly of the genus Pontania Costa (Hymenoptera: Tenthredinidae) (Kopelke 1994).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D92FFB75FCFE.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Toyama Pref., Toyama City, Arimine, Jûrôdani [Inonedani], 1120 m, Malaise trap, 1 – 8. IX. 2009, coll. M. Watanabe. Same city, trap and collector, Arimine, Inonedani, 1120 m: 1 ♂ (KPM-NK) 11 – 16. VIII. 2009, 1 ♀ (KPM-NK) 8 – 15. IX. 2009. 1 ♀ (KPM-NK) Nagano Pref., Chiisagata-gun, Nagawa-cho, Daimon, Utsukushimatsu, 36.133 ° N, 138.232 ° E, 1300 – 1350 m, 25. VIII. 2011, coll. Shunpei Fujie.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D92FFB75FCFE.taxon	distribution	Distribution. Russian Far East; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D86BFDAAF972.taxon	description	(Figs 29 – 35)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D86BFDAAF972.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 2 ♂ (NIAES) Tomakomai City, Hokkaido University Experimental Forest, 18 – 20. VI. 1980, coll. H. Takemoto. Honshu I.: 1 ♀ (ZISP) Niigata Pref., Kurokawa-mura, Tainai, 3 – 4. VI. 1980, coll. K. Maetô. Shikoku I.: 1 ♀ (NIAES) Ehime Pref., Mt. Ishizuchi-san, 1300 – 1982 m, 5. VIII. 1983, coll. K. Konishi. Kyushu I.: 1 ♀ (NIAES) Miyazaki Pref., Takaharu-machi, Miike, 8. V. 1983, coll. K. Konishi. 1 ♂ (NIAES) Kagoshima Pref., Yakushima Is., Shirataniunsuikyô, 680 m, 15. V. 1983, coll. I. Kanazawa et al.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D86BFDAAF972.taxon	description	Variation. Antenna with 19 flagellomeres in female and 20 – 22 flagellomeres in male. Dorsolateral area of propodeum sometimes very shallowly sculptured and dull (Fig. 35).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D86BFDAAF972.taxon	discussion	Remarks. This species was originally described in the genus Diaparsis from Primorsky Reg. of Russia (Khalaim 2005) and it possesses the propodeum with clearly delimited rectangular basal area (basal longitudinal carinae sometimes indistinct) (Fig. 35). Such areolation of the propodeum is typical for the genus Probles, while in the genus Diaparsis propodeum is almost always with a single median longitudinal carina (basal keel). Other important characters, e. g. the S-curved foveate groove of mesopleuron (Fig. 32), first tergite with postpetiole strongly widened at base and much broader than the petiole (Fig. 33), second tergite with elongated thyridial depression (Fig. 33), and slender ovipositor with weak dorsal subapical depression (Figs 29, 30) also well correspond with the genus Probles. This genus is characterized by the glymma joining by furrow with the lower side of postpetiole, but in D. ecarinata the glymma is isolated (Fig. 32). In spite of this, I consider that this species belongs to Probles (comb. nov.) because the furrow between glymma and lower side of postpetiole sometimes indistinct or vanishing in other species of Probles and in the genus Stethantyx Townes which has a similar structure of the first tergite.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03D86BFDAAF972.taxon	distribution	Distribution. Russian Far East; in Japan known from Hokkaido, Honshu, Shikoku, Kyushu and Yakushima islands. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03DDD2FB24F81D.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Toyama Pref., Toyama City, Arimine, Jûrôdani [Inonedani], 1120 m, Malaise trap, 14 – 21. VII. 2009, coll. M. Watanabe. 1 ♀ (NIAES) Nagano Pref., Shimashimadant, 1000 – 1300 m, 28. VII. 1980, coll. H. Takemoto.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03DDD2FB24F81D.taxon	description	Variation. Female from Nagano Pref. has flagellum with 22 – 23 flagellomeres (number of flagellomeres in left and right antennae is different) and basal area of propodeum indistinct.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE53FFB8FF03DDD2FB24F81D.taxon	distribution	Distribution. South Korea; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE54FFBEFF03DC1AFB75FE8E.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (KPM-NK) Toyama Pref., Toyama City, Arimine, Inonedani, 1120 m, Malaise trap, 15 – 22. IX. 2009, coll. M. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE54FFBEFF03DC1AFB75FE8E.taxon	discussion	Remarks. This species is very similar to P. carinator, but female of P. hankaensis from Japan possesses malar space about 0.8 × as long as basal mandibular width, dorsolateral area of propodeum very shallowly granulate (nearly smooth) and weakly shining, propodeal spiracle separated from pleural carina by one diameter of spiracle, and ovipositor somewhat longer than in specimens of P. carinator from Japan. In addition to the diagnostic characters provided in the key to species (Khalaim 2014: 104), P. hankaensis differs from P. carinator by less slender first metasomal tergite with lateral sides of petiole partly striate (entirely smooth in P. carinator). Notably, the female of P. hankaensis was collected in the same locality where most specimens of P. carinator were collected.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE54FFBEFF03DC1AFB75FE8E.taxon	distribution	Distribution. Russian Far East; in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DABBFECEFD42.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (NIAES) Mt. Tarumae-san, Malaise trap, 21 – 26. VII. 1998, coll. K. Konishi. Honshu I.: 1 ♀ (KPM-NK) Toyama Pref., Toyama City, Arimine, Jûrôdani, 1120 m, Malaise trap, 21 – 28. VII. 2009, coll. M. Watanabe.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DABBFECEFD42.taxon	distribution	Distribution. South Korea,? Russia (Far East); in Japan known from Hokkaido and Honshu islands. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03D9F7FDC3FC22.taxon	materials_examined	Material examined. JAPAN. Honshu I.: 1 ♀ (NIAES) Nagano Pref., Tateshina-kôgen, 10. X. 1982, coll. K. Konishi.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03D9F7FDC3FC22.taxon	distribution	Distribution. Europe, Caucasus, Russian Siberia and Far East, Mongolia, China (Ningxia); in Japan known from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DF17FA27FB2E.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 2 ♀ (NIAES) Horonobe, Kamitoikan, Iwananosawa, Malaise trap, 25. VII – 27. IX. 1993, coll. M. Inoue.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DF17FA27FB2E.taxon	distribution	Distribution. Europe, Turkey, Russian Far East; in Japan known from Hokkaido I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DE1BFB55F89A.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (NIAES) Hokkaido Pref., Kumaishi, Ken’ichi-gawa [River], Iwafuchi-zawa, Malaise trap, 1 – 10. VIII. 1995, coll. Y. Ito & T. Ito. 2 ♂ (ZISP, KPM-NK) Hokkaido Pref., Sapporoshi, Maruyama, 43 ° 02 ’ 54 ” N, 141 ° 18 ’ 51 ” E, 50 – 210 m, 29. VII. 2009, coll. K. Watanabe. Honshu I.: 1 ♀ (ZISP) Aomori Pref., Aomori City, Moyasawa, 14. VIII. 1993, coll. T. Ichita. 1 ♂ (KPM-NK) Kanagawa Pref., Fujinocho, Mt. Jinba-yama, 35 ° 39 ’ 09 ” N, 139 ° 09 ’ 55 ” E, 700 – 830 m, 7. VI. 2008, coll. K. Watanabe. 1 ♀ (AEIC) Nagano Pref., Kamikochi, 24. VII. 1954, coll. Townes Family. 1 ♀ (KPM-NK) Yamanashi Pref., Hokuto-shi, Masutomi, Biwakubo-sawa, 24. VI. 2007, coll. Teriaki Ban. 1 ♂ (KPM-NK) Yamanashi Pref., Mt. Daibosatu, Sagashio-kosen, Hikawa-rindo, 16. VI. 2007, coll. K. Watanabe. Kyushu I.: 1 ♀ (NIAES) Miyazaki Pref., Miike [Lake], Miyakonojô, 20. V. 1982, coll. K. Konishi.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DE1BFB55F89A.taxon	description	Variation. Female from Miike is smaller than other specimens and possesses body 4.2 mm long, fore wing 3.2 mm long, flagellum with 26 – 27 flagellomeres and clypeus with fewer and sparser punctures. Males have flagellum with 26 – 31 flagellomeres.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DE1BFB55F89A.taxon	discussion	Remarks. This species was recorded from Hokkaido and Honshu islands by Khalaim (2007: 592) but without any details. In this paper, complete label data of these specimens are provided.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE55FFBEFF03DE1BFB55F89A.taxon	distribution	Distribution. Russian Far East; in Japan known from Hokkaido, Honshu and Kyushu islands.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DB9AFF56FBE0.taxon	materials_examined	Material examined. JAPAN. Kyushu I.: 1 ♀ (NIAES) Oita [Ôita] Pref., Mts Kuju, Mt. Daisen, 19. VII. 1978, coll. K. Setoya.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DB9AFF56FBE0.taxon	description	Variation. Female from Japan has clypeus with punctures somewhat denser than in material from the continent. Flagellum with 26 – 27 flagellomeres.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DB9AFF56FBE0.taxon	discussion	Remarks. This species is similar to P. kunashirica but differs from the latter by the features given in Table 1.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DB9AFF56FBE0.taxon	distribution	Distribution. Europe, Russian Siberia and Far East; in Japan known from Kyushu island. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DE39FAE9F984.taxon	materials_examined	Material examined. Japan. 3 ♀, 1 ♂ (2 ♀, 1 ♂ in NIAES; 1 ♀ in ZISP) Hokkaido I., Mt. Tarumae-san, Malaise trap, 12 – 26. vii. 1998, coll. K. Konishi.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DE39FAE9F984.taxon	description	Variation. Male from Japan is considerably smaller than females with body length 3.5 mm and fore wing 3.0 mm, while in both females from Japan the body is approximately 6.0 mm long and fore wing is 4.0 mm long.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DE39FAE9F984.taxon	discussion	Remarks. This species was mentioned from Hokkaido I. of Japan by Khalaim (2007: 592) but without any details. In this paper, complete label data of the examined specimens are provided.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBDFF03DE39FAE9F984.taxon	distribution	Distribution. North America, Europe, Middle Asia, Russian Far East; in Japan known from Hokkaido I.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE56FFBCFF03DDCDFEAEFF3E.taxon	description	Most host records of the genus are from the beetle families Chrysomelidae, Curculionidae and Nitidulidae (Coleoptera), but two species were reared from leaf mining Eriocraniidae (Lepidoptera) (Jordan 1998) and some species of the subgenus Pectinolochus probably parasitise tenthredinid sawflies (Hymenoptera: Tenthredinidae) (Yu et al. 2016).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03DA2BFF51FDD6.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (EUM) Bibai, 43 ° 19 ’ N, 141 ° 48 ’ E, soybean field, Yellow pan trap, 3 – 4. VI. 2003, coll. O. Nagaia.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03DA2BFF51FDD6.taxon	distribution	Distribution. Europe, Russian Far East, China (Qinghai); in Japan known from Hokkaido I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03DA2BFF51FDD6.taxon	biology_ecology	Biology. Parasitoid of birch leaf mining Eriocrania species (Lepidoptera: Eriocraniidae) in Europe (Jordan 1998).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03D983FE05FADB.taxon	description	(Figs 36 – 42)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03D983FE05FADB.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (ZISP) Sapporo, Hitsujigaoka, 43 ° 00 ’ N, 141 ° 24 ’ E, Malaise trap, 10 – 17. V. 2007, coll. K. Konishi. Honshu I.: 2 ♀ (EUM) Aomori, Nishimeya-mura, Kawaratai, 40 ° 31 ’ N, 140 ° 10 ’ E, Malaise trap, 29. V – 6. VI. 2013, coll. T. Nakamura. 1 ♀ (EUM) same data, but 3. VI. 2014. RUSSIA. Khabarovsk Reg.: 1 ♀ (ZISP) Khekhtsir, upper stream of Levaya River, 11. VI. 1983, coll. D. R. Kasparyan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03D983FE05FADB.taxon	diagnosis	Diagnosis. Head and mesosoma densely granulate, impunctate. Clypeus large, granulate in upper 0.7 (Fig. 37). Malar space about as long as basal mandibular width (Fig. 38). Antenna short, with 17 flagellomeres (Fig. 39); flagellomeres 4 to 7 with distinct subapical finger-shaped structures on outer surface (Fig. 39). Notaulus absent (Fig. 38). Foveate groove of mesopleuron shallow, oblique, with weak transverse wrinkles (Fig. 38). Propodeal spiracle very small, separated from pleural carina by one diameter of spiracle (Fig. 38). Propodeum with basal keel (Fig. 40). First tergite smooth laterally and dorsally (Figs 38, 41); postpetiole in dorsal view much wider than petiole (Fig. 41); glymma large, joining by distinct furrow to ventral part of postpetiole (Fig. 38). Thyridial depression very short, strongly transverse (Fig. 41). Ovipositor short, weakly upcurved, with weak dorsal subapical depression and more or less distinct rounded tooth before this depression (Fig. 42). Predominantly dark brown and black species (Fig. 36).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFBCFF03D983FE05FADB.taxon	distribution	Distribution. Russian Far East, South Korea; in Japan known from Hokkaido and Honshu islands. First record from Japan and Russia.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFA3FF03DE8FFAE1FF12.taxon	description	(Figs 43 – 49)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFA3FF03DE8FFAE1FF12.taxon	materials_examined	Material examined. JAPAN. Shikoku I.: 1 ♀ (EUM) Ehime Pref., 11 – 20. VI. 2012, coll. Eiji Yamamoto. 1 ♀ (EUM) Ehime Pref., Kumakogen, 21 – 30. VI. 2012, coll. Eiji Yamamoto.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFA3FF03DE8FFAE1FF12.taxon	diagnosis	Diagnosis. Gena and mesopleuron more or less smooth, distinctly punctate (Fig. 44). Clypeus large, lenticular. Malar space 0.7 – 0.9 × as long as basal mandibular width (Fig. 44). Antennal flagellum tapered towards apex. Notaulus distinct, with irregular wrinkles (Fig. 44). Foveate groove of mesopleuron deep and long, distinctly upcurved anteriorly, with transverse wrinkles (Fig. 45). Propodeal spiracle moderately large, separated from pleural carina by 1.5 – 2.0 × diameter of spiracle (Figs 44, 45). First tergite smooth laterally and dorsally (Figs 44, 46); postpetiole in dorsal view much wider than petiole; glymma large, joining by distinct furrow to ventral part of postpetiole (Figs 44, 46). Thyridial depression slightly elongate (Fig. 48). Ovipositor short and robust, apically strongly upcurved and with lower valve expanded ventrally; ovipositor with two dorsal subapical depressions or notches and two more or less distinct rounded teeth before these depressions (Fig. 47). Predominantly black species with metasoma posterior to first tergite and all legs (except coxae) predominantly yellowish brown to reddish brown (Fig. 43).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFA3FF03DE8FFAE1FF12.taxon	discussion	Remarks. Female from Kumakogen, Ehime Pref. possesses abnormal first metasomal tergite which is abruptly curved posteriorly in lateral view (Fig. 46) and has a pair of strong spiracular prominences (Fig. 48). Another female from Ehime Pref. bears a parasitic larva of a mite belonging to Parasitengona (Acariformes: Trombidiformes) on dorsolateral side of the fifth tergite (Figs 43, 49).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE57FFA3FF03DE8FFAE1FF12.taxon	distribution	Distribution. Russian Siberia and Far East; in Japan known from Shikoku I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE49FFA7FF03DEEAFE69FF12.taxon	description	(Figs 50 – 54)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE49FFA7FF03DEEAFE69FF12.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: Sapporo, Hitsujigaoka, 43 ° 00 ’ N, 141 ° 24 ’ E, Malaise trap, coll. K. Konishi: 2 ♀ (EUM) 27. IX – 4. X. 2007, 1 ♀ (EUM) 4 – 11. X. 2007, 4 ♀ (EUM) 11 – 18. X. 2007, 7 ♀ (EUM) 25. X – 1. XI. 2007, 4 ♀ (2 ♀ in EUM, 2 ♀ in ZISP) 1 – 8. XI. 2007, 1 ♀ (EUM) 8 – 22. XI. 2007, 4 ♀ (EUM) 1 – 8. IX. 2008, 3 ♀ (EUM) 25. IX – 2. X. 2009, 2 ♀ (EUM) 23. X – 6. XI. 2009, 3 ♀ (EUM) 5 – 12. X. 2010, 1 ♀ (EUM) 12 – 19. X. 2010, 1 ♀ (EUM) 19 – 26. X. 2010, 1 ♀ (EUM) 26. X – 2. XI. 2010, 1 ♀ (EUM) 24 – 31. X. 2011, 1 ♀ (EUM) 7 – 21. XI. 2011, 2 ♀ (EUM) 22 – 29. X. 2013. 1 ♀ (EUM) same locality, 16. X. 2015, coll. Yu Hisasue. 1 ♀ (EUM) Sapporo, Hitsujigaoka, 43 ° 00 ’ 01 ” N, 141 ° 24 ’ 57 ” E, around nest of Lasius (D.) nipponensis, 17. X. 2015, coll. K. Konishi. 1 ♀ (EUM) Chitose, Bifue [Campground], Kusabue-rindô, 42.7154 ° N, 141.2143 ° E, Malaise trap, 27. X – 10. XI. 2012, coll. N. Kuhara. 2 ♀ (EUM) Kyowa, Shinsen-numa, 42 ° 54 ’ N, 140 ° 35 ’ E, 11. IX. 2010, coll. K. Konishi. 1 ♀ (EUM) Naganuma, Malaise trap, 26 – 30. IX. 2003, coll. A. Iwasaki. Honshu I.: 11 ♀, 6 ♂ (10 ♀, 5 ♂ in EUM; 1 ♀, 1 ♂ in ZISP) Tochigi Pref., Nikkô-shi, Yumoto, 22. IX. 2015, coll. Juriya Okayasu. Shikoku I.: 2 ♀, 1 ♂ (EUM) Ehime Pref., Matsuyama-shi, Mt. Ishizuchi, 1400 – 1600 m, 10 – 11. IX. 2017, coll. Yu Hisasue. 2 ♀ (EUM) Ehime Pref., Mt. Saragamine, 9. X. 1979, coll. Y. Seiyama. 2 ♀ (EUM) Ehime Pref., Odamiyama, Koyayama, 28. X and 19. XI. 2000, coll. E. Yamamoto. 1 ♀ (EUM) Ehime Pref., Odamiyama, Hontani, 21 – 31. X. 2008, coll. E. Yamamoto. 1 ♀ (EUM) Tokushima, Sanagouchi, Ôkawara, 33 ° 57 ’ 28 ” N, 134 ° 25 ’ 10 ” E, Malaise trap, 12 – 17. VI. 2016, coll. K. Ôhara. 1 ♀ (EUM) Kôchi Pref., Agawagun, Ino-cho, Teragawa, 33 ° 47 ’ N, 133 ° 12 ’ E, 5. XI. 2016, coll. Keita Kuroda.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE49FFA7FF03DEEAFE69FF12.taxon	diagnosis	Diagnosis. Antennal flagellum filiform, slender, brownish black or black (Fig. 51). Malar space short, about half as long as basal mandibular width (Fig. 52). Head and mesosoma evenly granulate, dull, impunctate (Fig. 52). Mesopleuron with very short and thin foveate groove (Fig. 52). Propodeum usually with narrow basal area which is often indistinct because of longitudinal wrinkles (Fig. 53). Metacarpus almost reaching tip of the fore wing (Fig. 50). First metasomal tergite very slender, laterally smooth (Fig. 52). Second tergite about as long as anteriorly broad. Metasoma dark brown (Fig. 50). Ovipositor short, robust, with weak dorsal subapical depression and more or less distinct rounded teeth before and behind this depression, with fine teeth ventrally (Fig. 54).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE49FFA7FF03DEEAFE69FF12.taxon	distribution	Distribution. Russian Far East; in Japan known from Hokkaido, Honshu and Shikoku islands.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE49FFA7FF03DEEAFE69FF12.taxon	biology_ecology	Biology. One female from Sapporo was collected around nest of Lasius (Dendrolasius) nipponensis Forel (Hymenoptera: Formicidae).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DA47FE98FCB6.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (EUM) Sapporo, Usubetsu, 42.9348 ° N, 141.1206 ° E, small stream, Malaise trap, 29. IV – 24. V. 2012, coll. N. Kuhara. Additional material. Honshu I.: 1 ♀ (EUM) Aomori, Nishimeya-mura, Kawaratai, 43 ° 31 ’ N, 141 ° 10 ’ E, Malaise trap, 21 – 30. VI. 2013, coll. T. Nakamura.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DA47FE98FCB6.taxon	diagnosis	Diagnosis. Flagellum short, with 17 – 18 short flagellomeres. Mesopleuron entirely granulate, with weak punctures centrally; foveate groove absent or indistinct. Metacarpus short, not reaching tip of the wing. First metasomal tergite slender, polished. Second tergite slightly elongate. Ovipositor sheath distinctly shorter than first tergite.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DA47FE98FCB6.taxon	description	Variation. Malar space in Japanese specimens 0.7 – 0.8 × as long as basal mandibular width. Female from Honshu I. has metacarpus long and reaching tip of the wing, vein 2 rs-m short and thick, and mesopleuron centrally nearly smooth and finely punctate. Thus, this specimen may represent either an undescribed species or variation of T. impunctator.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DA47FE98FCB6.taxon	distribution	Distribution. Russian Far East; in Japan known from Hokkaido I. and probably from Honshu I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03D8A3FB6AFA92.taxon	description	(Figs 55 – 57)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03D8A3FB6AFA92.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: Sapporo, Hitsujigaoka, Malaise trap, 43 ° 00 ’ N, 141 ° 24 ’ E, coll. K. Konishi: 1 ♀ (EUM) 4 – 11. VI. 2003, 1 ♀ (ZISP) 11 – 18. VI. 2003, 1 ♀ (EUM) 18 – 25. VI. 2003, 8 ♀ (EUM) 28. V – 4. VI. 2003, 3 ♀ and 1 ♂ (EUM) 15 – 22. V. 2009, 1 ♀ (EUM) 4 – 11. VII. 2011.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03D8A3FB6AFA92.taxon	diagnosis	Diagnosis. Antennal flagellum short, brownish black or black. Head and mesosoma granulate, dull, impunctate (Fig. 56). Mesopleuron with foveate groove usually represented by weak oblique wrinkles or stronger granulation (Fig. 56). Propodeum with fine basal keel which is sometimes indistinct (Fig. 57). Metacarpus not reaching tip of fore wing (Fig. 55). First metasomal tergite slender, usually finely striate laterally before glymma (Fig. 56). Second tergite about as long as anteriorly broad. Metasoma dark brown (Fig. 55). Ovipositor slender, long, with small dorsal subapical notch (Fig. 55).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03D8A3FB6AFA92.taxon	distribution	Distribution. Russian Far East; in Japan known from Hokkaido I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DEC7FCB9F852.taxon	description	(Figs 58 – 60)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DEC7FCB9F852.taxon	materials_examined	Material examined. JAPAN. Shikoku I.: 1 ♀ (EUM) Ehime Pref., Omogo, 2. VI. 1978, coll. Y. Seiyama. 1 ♀ (ZISP) Ehime Pref., Odamiyama, Keikoku, 16. VI. 2000, coll. E. Yamamoto. 1 ♀ (EUM) Ehime Pref., Imabari City, Mt. Narabara, 11. VI. 2006, coll. Yûgo Satô. Additional material. Shikoku I.: 2 ♀ (EUM) Ehime Pref., Matsuyama, flight intercept, 11 – 17. VI. 2016, coll. H. Yoshitomi. 1 ♀ (ZISP) Ehime Pref., Takanawa Peninsula, 14 – 15. V. 2016, coll. K. Kuroda. 1 ♀ (EUM) Ehime Pref., Komenono, Ehime University Field Station, 33 ° 54 ’ N, 132 ° 54 ’ E, 6. VI. 2015, coll. K. Konishi.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DEC7FCB9F852.taxon	description	Variation. Typical females have flagellum with 23 flagellomeres, malar space about 0.7 × as long as basal mandibular width, second tergite 0.9 – 1.0 × as long as broad and ovipositor sheath 1.2 – 1.3 × as long as first tergite (Fig. 58), while four females included as additional material are smaller, have flagellum with 18 – 19 flagellomeres, malar space 0.5 – 0.7 × as long as basal mandibular width and mesopleuron centrally polished and almost impunctate.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4CFFA7FF03DEC7FCB9F852.taxon	distribution	Distribution. Japan (Kyushu and Shikoku islands).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFA5FF03DFCDFEA9FA10.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (EUM) Kamikawa Distr., Kurodake-Sounkyo, 24. VII. 1956, coll. M. Miyatake.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFA5FF03DFCDFEA9FA10.taxon	description	Variation. Female from Japan virtually lacks foveate groove; mesopleuron just with a slightly stronger granulation indicating path of the foveate groove.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFA5FF03DFCDFEA9FA10.taxon	distribution	Distribution. Europe, Russian Far East; in Japan known from Hokkaido I. First record from Japan.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFA5FF03DFCDFEA9FA10.taxon	biology_ecology	Biology. Reared from unknown larva in mine on Betula pendula Roth (= verrucosa Ehrh.) in France (Aubert 1997: 108).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFAAFF03DD41FC65FE62.taxon	description	(Figs 61 – 68)	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFAAFF03DD41FC65FE62.taxon	materials_examined	Material examined. Holotype female (EUM) Japan, Shikoku I., Ehime Pref., Mt. Kasatori, 21 – 31. V. 2013, coll. Eiji Yamamoto.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFAAFF03DD41FC65FE62.taxon	description	Description. Female. Body length 4.0 mm. Fore wing length 3.7 mm. Head, in dorsal view, weakly rounded and strongly constricted posterior to eyes (Fig. 64); gena 0.75 × as long as eye width. Eyes with short sparse setae (Fig. 63). Clypeus about 2.6 × as broad as long, in front view almost lenticular, with lower margin slightly truncate (Fig. 63), separated from face by deep impression, bent backwards in lower 0.3 (i. e. with transverse bend); clypeus smooth in lower 0.3 and distinctly punctate on finely granulate background in upper 0.7. Mandible moderately robust, slightly tapered in basal 0.4 and with upper and lower margins subparallel in apical 0.6; upper tooth somewhat longer than the lower. Malar space about 0.9 × as long as basal mandibular width. Antennal flagellum (Fig. 62) filiform, with 18 flagellomeres; subbasal flagellomeres 1.6 – 1.8 × and subapical flagellomeres about 1.2 × as long as broad; flagellomeres 3 to 5 with distinct subapical finger-shaped structures on outer surface (Fig. 62). Face with elongate median prominence. Face, frons, vertex and gena distinctly granulate, dull, virtually impunctate (gena with very fine inconspicuous punctures). Occipital carina complete. Mesosoma entirely granulate, dull to weakly shining, mostly impunctate; with more or less distinct punctures on mesopleuron centrally. Notaulus with short and weak longitudinal wrinkle on anterolateral side of mesoscutum (Fig. 64). Scutellum with lateral longitudinal carinae present at its front 0.2. Epicnemial carina with upper end distinctly curved to reach front margin of mesopleuron. Foveate groove completely absent (Fig. 67). Propodeal spiracle separated from pleural carina by distance shorter than one diameter of spiracle (Fig. 67). Propodeum with basal keel which is about 0.3 × as long as apical area (Fig. 68); apical area slightly impressed along midline, rounded anteriorly; apical longitudinal carinae reaching transverse carina anteriorly (Fig. 68). Fore wing with second recurrent vein (2 m-cu) distinctly postfurcal, unpigmented in anterior half and distinct in posterior half. Intercubitus (2 rs-m) moderately long and thick, about as long as abscissa of cubitus between intercubitus and second recurrent vein (abscissa of M between 2 rs-m and 2 m-cu). First abscissa of radius (Rs + 2 r) straight, longer than width of pterostigma. First and second abscissae of radius (Rs + 2 r and Rs) meeting at slightly acute angle. Metacarpus (R 1) not reaching apex of fore wing. Hind wing with nervellus (cu 1 & cu-a) slightly reclivous, almost vertical. Legs slender. Tarsal claws not pectinate. First tergite 2.9 × as long as posteriorly broad, predominantly smooth, petiole with shallow longitudinal striae in posterior part dorsally and laterally, and with deep longitudinal impression dorsally in posterior part; first tergite, in cross-section centrally, trapeziform (lateral carinae strong in anterior half of tergite, petiole dorsally flat). Glymma large and deep, situated somewhat behind centre of tergite, joining by strong furrow to ventral part of postpetiole. Second tergite as long as anteriorly broad (Fig. 65). Thyridial depression about as long as broad (Fig. 65). Ovipositor long and slender, weakly bent upwards in basal 0.8 and strongly upcurved in apical 0.2, with two distinct dorsal subapical teeth and several fine teeth ventrally (Fig. 66); sheath about 2.4 × as long as first tergite (Fig. 61). Head, mesosoma and first tergite of metasoma black; lower 0.3 of clypeus, mouthparts and mandible (teeth reddish black) yellow-black. Antenna black, apical margins of scape and pedicel brownish. Tegula and pterostigma brown. Legs predominantly yellow-brown, all coxae and hind femur centrally darkened with brown. Metasoma posterior to first tergite dark brown. Male. Unknown.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFAAFF03DD41FC65FE62.taxon	etymology	Etymology. The species is named after its similarity to T. serratus Khalaim & Lee.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE4EFFAAFF03DD41FC65FE62.taxon	distribution	Distribution. Japan (Shikoku I.). Comparison. In the key to the species occurring in Korea (Khalaim et al. 2014), the new species runs to T. uncinatus Khalaim & Lee in couplet 9, but differs from this species by the ovipositor with two distinct dorsal subapical teeth and several fine teeth ventrally (Fig. 66) (apically thin and with a weak dorsal notch in T. uncinatus). In the shape of the ovipositor, T. serratoides sp. nov. resembles another Korean species — T. serratus Khalaim & Lee (Khalaim et al. 2014). In the key to the species of the Russian Far East and Japan (Khalaim 2012), T. serratoides sp. nov. runs to T. juxtus in couplet 8, but in the new species the gena is shorter and less rounded posterior to eyes (Fig. 64), and ovipositor is with distinct dorsal subapical teeth (Fig. 66). Tersilochus serratoides sp. nov. also differs from the occurring in China T. ningxiator Khalaim & Sheng (Khalaim & Sheng 2009) by completely lacking foveate groove of the mesopleuron (Fig. 67) and longer ovipositor (Fig. 61), and from T. runatus Khalaim & Sheng (Khalaim & Sheng 2009) by shorter gena, flagellum with 18 flagellomeres (15 – 16 in T. runatus) and more slender ovipositor (Fig. 61).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03D8A3FBD0FBB3.taxon	materials_examined	Material examined. Russia. 1 ♀ (ZISP) Primorsky Reg., 10 km SE of Partizansk, Novitskoe, 11. VII. 1996, coll. S. A. Belokobylskij.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03D8A3FBD0FBB3.taxon	distribution	Distribution. South Korea, Russia (south of Far East). First record from Russia.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03DFA7FA24FABE.taxon	materials_examined	Material examined. RUSSIA. Vologda Prov.: 1 ♀ (ZISP) Kaduy Distr., Zaozerye, pine forest, Hieracium, 26. VIII. 2014, coll. N. S. Kolesova.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03DFA7FA24FABE.taxon	distribution	Distribution. Holarctic species; in Palaearctic region known from Europe,? Iran, Russian Siberia and Far East.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03DAD7FEFAFCFE.taxon	materials_examined	Material examined. JAPAN. Hokkaido I.: 1 ♀ (EUM) Sapporo, Hitsujigaoka, 43 ° 00 ’ 27 ” N, 141 ° 24 ’ 53 ” E, Malaise trap, 15 – 22. VI. 2010, coll. K. Konishi. 1 ♀ (EUM) Chitose, Bifue, Kusabue-rindô, 42.7154 ° N, 141.2143 ° E, Malaise trap, 21. VI – 8. VII. 2012, coll. N. Kuhara. Honshu I.: 4 ♀ (EUM) Aomori, Nishimeya-mura, Kawaratai, 43 ° 31 ’ N, 141 ° 10 ’ E, Malaise trap, 12 – 21. VI. 2013, coll. T. Nakamura. 1 ♀ (EUM) same data, but 30. VI – 15. VII. 2013. Shikoku I.: 1 ♀ (EUM) Ehime Pref., Kumakogen, Yurano, 31. V. 2007, coll. E. Yamamoto.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFAAFF03DAD7FEFAFCFE.taxon	distribution	Distribution. Russian Far East, China (Liaoning); in Japan known from Hokkaido, Honshu, Shikoku and Kyushu islands.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFA9FF03DE96FBD0FE8F.taxon	materials_examined	Material examined. RUSSIA. Sakhalin Prov.: 3 ♀ (ZISP) SW Kunashir I., Ivanovsky Cape, Grozovoe, broadleaved forest, 23 – 25. V. 2013, coll. Yu. & L. Sundukov. 1 ♂ (ZISP) SW Kunashir I., Ivanovsky Cape, 26 – 27. V. 2013, coll. Yu. & L. Sundukov.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFA9FF03DE96FBD0FE8F.taxon	diagnosis	Diagnosis. Head, in dorsal view, distinctly rounded posterior to eyes (Fig. 64); gena about 0.8 × as long as eye width. Antennal flagellum more or less filiform, brownish black or black, with 17 – 19 flagellomeres in female (Fig. 73) and 20 flagellomeres in male (Fig. 72). Malar space short, about half as long as basal mandibular width. Head and mesosoma evenly granulate, dull, impunctate (Figs 71, 75). Notaulus absent or with weak wrinkle or wrinkles on anterolateral side of mesoscutum (Figs 71, 75). Foveate groove of mesopleuron absent or shallow, with stronger granulation and weak wrinkles (Fig. 75). Propodeum mediodorsally with longitudinal wrinkles, thus basal area is indistinct (Fig. 76). Metacarpus not reaching tip of the fore wing (Figs 69, 70). First metasomal tergite smooth laterally and dorsally (sometimes with short striae laterally before glymma), trapeziform in cross-section centrally; glymma deep (Fig. 75). Second tergite about as long as anteriorly broad. Metasoma dark brown to brownish black, ventrally yellowish (Figs 69, 70). Ovipositor with strong dorsal subapical notch (Fig. 74), sheath about 1.8 × as long as first tergite (Fig. 69).	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFA9FF03DE96FBD0FE8F.taxon	description	Description of male (Fig. 70) (First record). Similar to female. Antenna with 20 flagellomeres (Fig. 72); flagellomeres 3 to 6 with distinct subapical finger-shaped structures on outer surface (Fig. 72). Malar space about 0.4 × as long as basal mandibular width. Hypostomal carina absent, this area finely striate. Notaulus and foveate groove absent. First tergite of metasoma laterally mostly smooth, before glymma partly striate. Thyridial depression transverse.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
03F15F55CE41FFA9FF03DE96FBD0FE8F.taxon	distribution	Distribution. South Korea, Russia (south of Far East). First record from Russia.	en	Khalaim, Andrey I. (2022): Tersilochinae (Hymenoptera: Ichneumonidae) of Japan: two new species, a new combination and 23 new country records. Zootaxa 5174 (4): 301-330, DOI: https://doi.org/10.11646/zootaxa.5174.4.1
