identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B81687E9F6547163EDD3730CFA95407B.text	B81687E9F6547163EDD3730CFA95407B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella pax Babenko & Shveenkova & Potapov 2022	<div><p>Philotella pax sp. nov.</p> <p>Figs 2–9</p> <p>Type material. Holotype: juvenile, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.3515&amp;materialsCitation.latitude=43.648" title="Search Plazi for locations around (long 132.3515/lat 43.648)">Southern Primorye</a>, Ussuri State Nature Reserve (Fig. 1, point 4), forest with Pinus koraiensis, 43°38.88’N 132°21.09’E, ~ 200 m alt., rotten wood, 22 July 2016. N. Kuznetsova &amp; M. Potapov leg. Paratypes: 3 juveniles, same data, but ~ 380 m alt., forest with P. koraiensis, litter, 22 July 2016.; 3 juveniles, same region and Nature Reserve, mixed forest, 43°38.2’N 132°20.93’E, ~ 380 m alt., litter, 23 July 2016. N. Kuznetsova &amp; M. Potapov leg.</p> <p>Additional material. 2 juveniles, same region, «Kedrovaya Pad» <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.48717&amp;materialsCitation.latitude=43.114666" title="Search Plazi for locations around (long 131.48717/lat 43.114666)">State Nature Reserve</a> (Fig. 1, point 3), mixed coniferous-broadleaf forest, 43°6.88’N 131°29.23’E, ~ 200 m alt., litter, 27 July 2016. N. Kuznetsova &amp; M. Potapov leg.; 2 juveniles, same region, ~ 30 km SE of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.15033&amp;materialsCitation.latitude=44.022" title="Search Plazi for locations around (long 134.15033/lat 44.022)">Chuguevka</a> (Fig. 1, point 7), mixed forest, 44°1.32’N 134°9.02’E, 500 m alt., 11 August 2017. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg.; 6 juveniles, same region and collectors, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.4985&amp;materialsCitation.latitude=43.668365" title="Search Plazi for locations around (long 132.4985/lat 43.668365)">Ussuri State Nature Reserve</a> (Fig. 1, point 4), valley of Anikin River, broadleaf forest, litter, 43°40.102’N 132°29.91’E, 13 August 2017.</p> <p>Diagnosis. A dark species characterized by the presence of 5+5 subequal ocelli; the second row of labral setae with three longer and two shorter setae; a labium of the first type; dorsal chaetotaxy with setae p2 present on Th. II–III but absent on Abd. I–V; the dorsolateral group of setae on Th. II–III with sensillum p3 set below two ordinary setae (a3 and m4); a complete tibiotarsal chaetotaxy and toothless unguis.</p> <p>Description. Length (without antennae) of available juveniles 0.33–0.42 mm, holotype — 0.40 mm. Habitus typical of the genus. Colour blue, uniformly dark. Tegument granulation very strong, especially on abdominal tip.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV blunt and elongate (Fig. 4); few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv slightly longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 11–12 setae, respectively.</p> <p>Head with 5+5 ocelli, slightly larger than surrounding secondary granules on ocular field (Fig. 5). PAO usually rounded, consisting of 8–10 vesicles, its axis to ocellus B ratio as 2.1–2.3: 1. Buccal cone short and blunt. Maxilla styliform, with at least one lamella bearing two small apical teeth (Fig. 6). Mandible delicate, with two tiny denticles on inner edge between apical and two basal teeth (Fig. 7). Labrum usually with a common set of 4/2352 setae of different length, distal pair and three setae of the next row being the longest (Fig. 8). Main part of labium of the first type with three ordinary setae in line along distal edge and 2+2 axial setae between fairly large sensorial elements (Fig. 9); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae as usual.</p> <p>Dorsal chaetotaxy symmetrical and rather reduced (Fig. 2). Ordinary setae short, needle-like, sensilla clearly longer than ordinary setae, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: head usually with unpaired seta d0 and only 2 setae on each ocular field; Th. I with 3+3 setae; Th. II–III with 3+3 axial setae (p2 present), dorso-lateral group consisting of two setae (a3 and m4) additionally to sensilla p 3 in backward position. Abd. I–III without p2, i.e. only 2+2 axial setae present, and with only one seta (a3) between axial group and sensillum p 4; both Abd. IV–V with only 2+2 axial setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae (Fig. 3). Furcal remnant in a form of two swellings, each with a seta, at anterior border of Abd. IV. Anal valves with two [?] hr-setae each.</p> <p>Legs I–III with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, 6 on trochanters, and 13, 12, 10–11 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Remarks. Unfortunately, only juveniles without any traces of a genital opening are present in the available material. However, based on a comparison of juvenile and adult individuals of other congeners, morphological rearrangements related to age can be assumed as not being too significant in the genus, failing to affect the main diagnostic characters, as a rule. The only notable exception is the ventral chaetotaxy of the abdomen, which is clearly more abundant in adults.</p> <p>Etymology. The species is named after the ancient goddess of peace in honour of those who uphold it. The name is a noun in apposition.</p> <p>Affinities. In general, P. pax sp. nov. seems to be especially similar to North Korean P. miracli. Their dorsal chaetotaxy is almost identical, except for the presence of two (a3 and m4) rather than three (a3, a4 and m4) setae in the dorsolateral group on Th. II–III. That this may be due to the juvenile condition of the type material cannot be ruled out, although such an assumption seems doubtful to us. The most significant difference between these two species lies in the structure of the labium: P. pax sp. nov. is characterized by a labium of the first type (see above), while the true P. miracli possesses a labial palp of the second type, with a triangular arrangement of setae A, C and D. A toothless unguis and a complete set of tibiotarsal setae also allows for P. pax sp. nov. to be distinguished from all three North Korean species described by Najt &amp; Weiner (1985), which lack setae M on the tibiotarsi.</p> </div>	http://treatment.plazi.org/id/B81687E9F6547163EDD3730CFA95407B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F6567162EDD372F7FD8542FB.text	B81687E9F6567162EDD372F7FD8542FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella oligotricha Babenko & Shveenkova & Potapov 2022	<div><p>Philotella oligotricha sp. nov.</p> <p>Figs 10–16</p> <p>Type material. Holotype: female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.48717&amp;materialsCitation.latitude=43.114666" title="Search Plazi for locations around (long 131.48717/lat 43.114666)">Southern Primorye</a>, «Kedrovaya Pad» State Nature Reserve (Fig. 1, point 3), mixed coniferous-broadleaf forest, 43°6.88’N 131°29.23’E, ~ 120 m alt., litter, 27 July 2016. N. Kuznetsova &amp; M. Potapov leg. Paratypes: 1 immature male, same habitat; 2 immature males and 1 juvenile, same region and <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.56134&amp;materialsCitation.latitude=43.101" title="Search Plazi for locations around (long 131.56134/lat 43.101)">Nature Reserve</a>, oak-wood, 43°6.06’N 131°33.68’E, ~ 150 m alt., litter, 28 July 2016; 1 female, same region, near <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.797&amp;materialsCitation.latitude=43.126" title="Search Plazi for locations around (long 132.797/lat 43.126)">Anisimovka</a>, Mount Khualaza (Fig. 1, point 5), greenwood forest, 43°7.56’N 132°47.82’E, ~ 400 m alt., litter and rotten wood, 21 July 2016; 1 immature female, same region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=132.3515&amp;materialsCitation.latitude=43.648" title="Search Plazi for locations around (long 132.3515/lat 43.648)">Ussuri State Nature Reserve</a> (Fig. 1, point 4), forest with P. koraiensis, 43°38.88’N 132°21.09’E, ~ 200 m. alt., 22 July 2016. N. Kuznetsova &amp; M. Potapov leg.</p> <p>Additional material. 5 females, Khabarovsk Territory, Lazo District, upper reaches of Katen River, Ko Mountains (Fig. 1, point 10), ~ 500 m alt., mixed forest, litter, 01 July 2018. A. Brinev leg.; 2 females, 1 immature male and 11 juveniles, Southern Primorye, Partisan District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=133.66783&amp;materialsCitation.latitude=43.305832" title="Search Plazi for locations around (long 133.66783/lat 43.305832)">Mount Olkhovaya</a> (Fig. 1, point 6), mixed forest in valley, 43°18.35’N 133°40.07’E, ~ 500 m alt., litter, 20 August 2018. M. Potapov, Yu. Shveenkova &amp; A. Kuprin leg.; 1 immature female and 1 immature male, Southern Primorye, Chuguev District, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.193&amp;materialsCitation.latitude=43.600666" title="Search Plazi for locations around (long 134.193/lat 43.600666)">National Park</a> «Zov Tigra», <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.193&amp;materialsCitation.latitude=43.600666" title="Search Plazi for locations around (long 134.193/lat 43.600666)">Mount Oblachnaya</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.193&amp;materialsCitation.latitude=43.600666" title="Search Plazi for locations around (long 134.193/lat 43.600666)">Ussuri River valley</a>, mixed forest, 43°36.04’N 134°11.58’E, ~ 550 m alt., litter, 19–20 September 2018. A. Kuprin leg.; 7 juveniles, Khabarovsk Territory, Komsomolsk State Nature Reserve, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.57&amp;materialsCitation.latitude=49.303333" title="Search Plazi for locations around (long 137.57/lat 49.303333)">Anyuinski National Park</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.57&amp;materialsCitation.latitude=49.303333" title="Search Plazi for locations around (long 137.57/lat 49.303333)">Tormasu River</a> ((Fig. 1, point 13), mixed coniferous-broadleaf forest, litter, 49°18.2’N 137°34.2’E, ~ 200 m alt., 07 August 2018. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg.</p> <p>Diagnosis. A dark species characterized by the presence of 3+3 subequal ocelli; the second row of labral setae with three longer and two shorter setae; a labium of the second type; reduced dorsal chaetotaxy without p2 on all terga from Th. II to Abd. V; a dorsolateral group of setae on Th. II–III with sensillum p3 set below two ordinary setae (a3 and m4); a complete tibiotarsal chaetotaxy and toothless unguis.</p> <p>Description. Length (without antennae) of available specimens 0.38–0.59 mm, holotype — 0.48 mm. Habitus typical of the genus. Colour dark blue, uniform. Tegument granulation very strong, especially on abdominal tip.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV blunt and elongate (Fig. 12); few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 11 setae, respectively.</p> <p>Head with 3+3 ocelli, clearly larger than surrounding secondary granules on ocular field. PAO rounded, consisting of 7–10 vesicles, its axis to ocellus B ratio as 1.6–2.0: 1. Buccal cone short and blunt. Maxilla styliform, with at least two lamellae, one of which bearing two hooks, another one with few apical denticles (Fig. 14). Mandible delicate, with five usual teeth, among them two tiny ones between apical and two basal teeth (Fig. 15). Labrum usually with a common set of 4/2352 setae of different length, distal pair and three setae of the next row being the longest (Fig. 13). Main part of labium of the second type with three ordinary setae in triangular position and 1+1 axial setae between sensorial elements (Fig. 16); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae.</p> <p>Dorsal chaetotaxy symmetrical and rather reduced (Fig. 10). Ordinary setae short, needle-like, sensilla longer, clearly differentiated, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: head usually with unpaired seta d0 and only 2 setae on each ocular field; Th. I with 3+3 setae; all terga from Th. II to Abd. IV with 2+2 axial setae (p2 absent); dorso-lateral group on Th. II–III consisting of 2 ordinary setae (a3 and m4) additionally to sensillum p 3 in backward position; Abd. V with only 1+1 setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae (Fig. 11). Furcal remnant in a form of two small warts, each with a seta, on anterior border of Abd. IV. Each anal valve usually with three setae hr, but variations frequent.</p> <p>Legs I–III with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, (5)6 on trochanters, and 13, 12, 11 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Remarks. The material from the Zeya State Nature Reserve (Amur Region, Fig. 1, point 1) contains three juvenile specimens of a very similar form, which differs from P. oligotricha sp. nov. by the presence of 5+5, not 3+3 ocelli. Other detected differences are minimal: the presence of three setae on the ocular field (vs only two setae in P. oligotricha sp. nov.) and 2+2 setae between sensilla on Abd. V (vs 1+1 setae in this position in P. oligotricha sp. nov.). Most likely, this is a different species, which cannot yet be described due to the lack of sufficient material.</p> <p>Etymology. The name of the new species reflects one of its most characteristic features, i.e. the relatively small number of dorsal setae; from ancient Greek όλίγος (few, little) &amp; τρίχα (seta).</p> <p>Affinities. Apart from the reduced number of ocelli, a character not always easy to correctly identify, P. oligotricha sp. nov. resembles P. deharvengi and P. tertia, the latter two species known from the neighbouring region. All these three species have an almost identical dorsal chaetotaxy with 2+2 axial setae on all terga from Th. II to Abd. III, and a backward position of sensilla in the dorsolateral groups of Th. II–III. Philotella oligotricha sp. nov. differs from P. deharvengi in having a different type of the labium (the first type in P. deharvengi, vs the second one in P. oligotricha sp. nov.) and 1+1 axial setae between sensilla only on Abd. V (vs 1+1 such setae on both Abd. IV and Abd. V in P. deharvengi). The chaetotaxy of the abdominal tip in P. tertia is characterized by the presence of 2+2 axial setae on both Abd. IV and Abd. V. Apart from this, both Korean species lack M-setae on all tibiotarsi and have an inner tooth on the unguis.</p> <p>The dorsal chaetotaxy of P. oligotricha sp. nov. is also quite similar to that of P. porcella comb. nov. (central Alaska), the only known Nearctic representative of the genus, although the latter species is characterized by the anterior position of the sensillum in the dorsolateral group on Th. II–III, and the presence of 2+2 axial setae between sensilla on both Abd. IV and Abd. V.</p> </div>	http://treatment.plazi.org/id/B81687E9F6567162EDD372F7FD8542FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F659716EEDD377A5FE9C441F.text	B81687E9F659716EEDD377A5FE9C441F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella cedrina Babenko & Shveenkova & Potapov 2022	<div><p>Philotella cedrina sp. nov.</p> <p>Figs 17–21</p> <p>Type material. Holotype: female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.48717&amp;materialsCitation.latitude=43.114666" title="Search Plazi for locations around (long 131.48717/lat 43.114666)">Southern Primorye</a>, «Kedrovaya Pad» State Nature Reserve (Fig. 1, point 3), mixed coniferous-broadleaf forest, 43°6.88’N 131°29.23’E, ~ 120 m alt., litter, 27 July 2016. N. Kuznetsova &amp; M. Potapov leg. Paratype: immature male, same data as holotype.</p> <p>Diagnosis. A species of the genus Philotella characterized by the presence of 3+3 subequal ocelli; the second row of labral setae with two longer and three shorter setae; a labium of the first type; a dorsal chaetotaxy with setae p2 present on Th. II –III but absent on Abd. I –IV; 1+1 setae between sensilla on Abd. V; a complete tibiotarsal chaetotaxy and toothless unguis.</p> <p>Description. Length (without antennae) of a single available female— 0.34 mm. Habitus typical of the genus. Alive colour unknown, probably rather dark. Tegument granulation strong, especially on abdominal tip.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV clearly differentiated, blunt and prolonged; few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 11 setae, respectively.</p> <p>Head with 3+3 [?] ocelli, clearly larger than surrounding secondary granules on ocular field. PAO rounded, consisting of 7–9 vesicles, its axis to ocellus B ratio as 1.5–1.7: 1. Buccal cone short and blunt. Maxilla styliform, lamellae not clearly seen. Mandible delicate, with five teeth, two of which tiny and hardly detected. Labrum with a common set of 4/2352 setae of different length, the second row with two longer marginal setae and three smaller median ones (Fig. 17). Main part of labium of the first type with three ordinary setae set in a line and 2+2 axial setae between sensorial elements (Fig. 18); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae.</p> <p>Dorsal chaetotaxy almost symmetrical (Fig. 19). Ordinary setae short, needle-like, sensilla much longer, their number as usual: 22/11111, lateral sensilla on Th. II and dorsal ones on Abd. IV slightly thickened (Fig. 21), Th. II with lateral ms present as usual. Main characteristics: head usually with unpaired seta d0, ocular field with 3 setae; Th. I with 3+3(2) setae; Th. II–III with 3+3 axial setae (p2 present); dorso-lateral group consisting of 2 ordinary setae (a3 and m4) additionally to sensillum p 3 in backward position; Abd. I–IV with 2+2 axial setae, Abd. V with only 1+1 setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae. Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV. Unpaired anal valve with three hr-setae, paired ones with two such setae, each.</p> <p>Legs I–III with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, (5)6 on trochanters, and 13, 11, 10 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Remarks. Despite the limited material available, we venture to describe this species, since the holotype appears to be sexually mature (Fig. 20) and has a fairly characteristic morphology and a symmetrical chaetotaxy.</p> <p>Etymology. The name of the new species is given after its type locality, «Kedrovaya Pad» (Cedar Valley) State Nature Reserve; from Ancient Greek κέδρινος (kédrinos), cedrinus (a, um) in Latin.</p> <p>Affinities. Philotella cedrina sp. nov. is similar to P. pax sp. nov., except for a reduced number of ocelli, which are not always easy to correctly count. Both these species have a labium of the same type and an almost identical dorsal chaetotaxy. The only noticeable differences in the latter character are the presence of three setae on the ocular field (vs two such setae in P. pax sp. nov.), 1+1 setae between sensilla on Abd. V (vs 2+2 setae in this position in P. pax sp. nov.), and markedly thickened sensilla on Th. II and Abd. IV. In addition, these species are more or less easily distinguishable due to differing lengths of labral setae: in P. cedrina sp. nov., two marginal setae of the second row are longer than the others, while in P. pax sp. nov., there are three such setae in this row (cf. Fig. 17 and Fig. 8).</p> <p>Among the known species of the genus, P. cedrina sp. nov. also resembles the Korean P. miracli, in which the dorsal chaetotaxy is very similar to that of P. cedrina sp. nov., but differs in having three ordinary setae in addition to sensillum p 3 in the dorsolateral group on Th. II–III. Besides the number of ocelli, these species also differ in the types of the labium and the number of tibiotarsal setae.</p> <p>It is also noteworthy that clearly thickened sensilla on Th. II and Abd. IV, both characteristic of P. cedrina sp. nov., have previously been mentioned also for all three North Korean species. In our material, the differentiation of these sensilla was also detected in P. amurica sp. nov., being especially noticeable in P. alba sp. nov. (see descriptions below).</p> </div>	http://treatment.plazi.org/id/B81687E9F659716EEDD377A5FE9C441F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F65B716EEDD37651FBC942F3.text	B81687E9F65B716EEDD37651FBC942F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella aliniensis Babenko & Shveenkova & Potapov 2022	<div><p>Philotella aliniensis sp. nov.</p> <p>Figs 22–24</p> <p>Type material. Holotype: female, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=135.887&amp;materialsCitation.latitude=45.138332" title="Search Plazi for locations around (long 135.887/lat 45.138332)">Southern Primorye</a>, Sikhote-Alin State Nature Reserve (Fig. 1, point 9), coniferous forest with Rhododendron brachycarpum, 45°8.30’N 135°53.22’E, ~ 930 m alt., litter, 08 August 2017. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg. Paratypes: 1 female and 7 juveniles, same biotope; 1 immature female, same region, ~ 30 km SE of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.15033&amp;materialsCitation.latitude=44.022" title="Search Plazi for locations around (long 134.15033/lat 44.022)">Chuguevka</a> (Fig. 1, point 7), mixed forest, 44°1.32’N 134°9.02’E, ~ 500 m alt., 11 August 2017. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg.</p> <p>Diagnosis. A species of the genus Philotella characterized by the presence of (2)3+3 ocelli, coarse granulation, the second row of labral setae with two longer and three shorter setae; a labium of the second type, dorsal chaetotaxy with p2 present on Th. II –III but absent on Abd. I – V; three ordinary setae (a3, a4 and m4) in dorsolateral group on Th. II –III and two setae (a3 and p3) between axial group and sensilla p4 on Abd. I – III; a complete tibiotarsal chaetotaxy and toothless unguis.</p> <p>Description. Length (without antennae) up to 0.53 mm. Habitus typical of the genus. Alive colour unknown, probably rather dark. Tegument granulation very strong, especially on head and abdominal tip, but fields with larger granules also present on most terga.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV clearly differentiated, blunt and elongate; few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and (11)12 setae, respectively.</p> <p>Head with (2)3+3 [?] hardly visible ocelli, about as large as surrounding secondary granules on ocular field (Fig. 23). PAO rounded, consisting of 10–12 vesicles, its axis to ocellus B ratio as 2.1–2.6: 1. Buccal cone short and blunt. Maxilla styliform, lamellae not clearly seen. Mandible delicate, with five teeth, two of which tiny and hardly detected. Labrum with a common set of 4/2352 setae of different length, the second row with two longer marginal setae and three smaller median ones (cf. Fig. 17). Main part of labium of the second type with three ordinary setae set in a triangular arrangement and 1+1 axial setae between sensorial elements (Fig. 22); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae.</p> <p>Dorsal chaetotaxy almost symmetrical (Fig. 24). Ordinary setae short, needle-like, sensilla longer, their number as usual: 22/11111, Th. II with lateral ms present as usual. Main characteristics: head usually with unpaired seta d0, ocular field with 3 setae; Th. I with 3+3 setae; Th. II–III with 3+3 axial setae (p2 present); dorsolateral group consisting of 3 ordinary setae (a3, a4 and m4) additionally to sensillum p 3 in backward position; Abd. I–III with 2+2 axial setae (p2 absent) and two setae (a3 and p3) between axial group and sensilla, both Abd. IV–V with 2+2 setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae. Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV. Unpaired anal valve with three hr-setae, their number on paired valve variable (2–3).</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, (5)6 on trochanters, and 13, 12, 11 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Etymology. The new species is named after the region of its type locality, the Sikhote-Alin Mountains, based on the Manchu word “Alin” meaning mountain range.</p> <p>Affinities. The dorsal chaetotaxy of P. aliniensis sp. nov. is almost identical to that of P. miracli: the only noticeable differences in this character lies in the presence of two setae between the axial group and sensilla on Abd. I–III in P. aliniensis sp. nov., whereas only a3 is present in this position in P. miracli. However, the very coarse granulation of the integument, the reduced number of ocelli, and the complete chaetotaxy of the tibiotarsi, all characteristic of the new species, make it fairly easily distinguished among these two species.</p> </div>	http://treatment.plazi.org/id/B81687E9F65B716EEDD37651FBC942F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F65C7169EDD377A5FEF94273.text	B81687E9F65C7169EDD377A5FEF94273.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella amurica Babenko & Shveenkova & Potapov 2022	<div><p>Philotella amurica sp. nov.</p> <p>Figs 25–30</p> <p>Type material. Holotype: immature male, Khabarovsk Territory, Komsomolsk State Nature Reserve (Fig. 1, point 12), <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.42383&amp;materialsCitation.latitude=50.7375" title="Search Plazi for locations around (long 137.42383/lat 50.7375)">Gorin River</a>, mixed coniferous-broadleaf forest, 50°44.25’N 137°25.43’E, ~ 250 m alt., rotting wood, 10 August 2018. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg. Paratypes: 2 females, same data as for holotype.</p> <p>Diagnosis. A species of the genus Philotella characterized by the presence of 2–3 ocelli on each side of the head; a labrum with subequal setae in the second row; a labium of the second type; dorsal chaetotaxy with p2 present on Th. II –III but absent on Abd. I – V; three ordinary setae (a3, a4 and m4) in dorsolateral group on Th. II –III and two setae (a3 and p3) between axial group and sensilla on Abd. II – III; a slightly reduced tibiotarsal chaetotaxy without M-seta and a toothless unguis.</p> <p>Description. Length (without antennae) up to 0.49 mm, holotype — 0.43 mm. Habitus typical of the genus. Alive colour unknown, probably rather dark. Tegument granulation not especially coarse.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV clearly differentiated, blunt and elongate; few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 11 setae, respectively.</p> <p>Each side of head with 2–3 ocelli of different size, individual variations frequent. PAO rounded, consisting of 7–9 vesicles, its axis to ocellus B ratio as 2.3–2.4: 1 (Fig. 28). Buccal cone short and blunt. Maxilla styliform, lamellae not clearly seen. Mandible delicate, with five teeth, two of which tiny and hardly detected. Labrum with a common set of 4/2352 setae, slightly different in length (Fig. 26). Main part of labium of the second type with three ordinary setae set in a triangular arrangement and 1+1 axial setae between sensorial elements (Fig. 27); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae.</p> <p>Dorsal chaetotaxy almost symmetrical (Fig. 25). Ordinary setae short, needle-like, sensilla clearly differentiated, their number as usual: 22/11111, lateral sensilla on Th. II and dorsal ones on Abd. IV clearly thickened. Th. II with lateral ms present. Main characteristics: head usually with unpaired seta d0, ocular field with 3 setae; Th. I with 3+3 setae; Th. II–III with 3+3 axial setae (p2 present); dorsolateral group consisting of 3 ordinary setae (a3, a4 and m4) additionally to sensillum p 3 in backward position; Abd. I–III with 2+2 axial setae (p2 absent). Abd. I with only one seta between axial group and sensilla, whereas Abd. II–III always have two setae (a3 and p3) on this position. Abd. IV usually with 3+3 setae between sensilla, Abd. V with 2+2 such setae.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae (Fig. 30). Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV. Number of hr-setae on anal valves unstable in studied specimens.</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 5, 5, 5(6) on trochanters, and 12, 11, 10 setae on femora. Tibiotarsi with 18, 18, 17 setae, i.e. seta M absent (Fig. 29). Unguis toothless.</p> <p>Etymology. The new species is named after the region of its type locality, the lower reaches of the Amur.</p> <p>Affinities. Philotella amurica sp. nov. is the only known species in the genus in the study area that lacks a Mseta on the tibiotarsi, a character which is typical of all North Korean congeners: P. deharvengi, P. miracli and P. tertia. Among the latter trio, P. amurica sp. nov. resembles P. miracli particularly strongly in having a similar dorsal chaetotaxy and the labium of the same type, but it clearly differs in the number of ocelli (2–3, vs 5 in P. miracli). In addition, there are also some other consistent differences between these species: the presence of 2+2 setae between the axial group and sensilla on Abd. II–III (vs only 1+1 setae in this position in P. miracli) and 3+3 setae between sensilla on Abd. IV (vs only 2+2 axial setae in P. miracli).</p> <p>The dorsal chaetotaxy of P. amurica sp. nov. is even more similar to that of P. aliniensis sp. nov. (cf. Fig 25 &amp; Fig. 24). These species are also characterized by the same type of the labium, and a reduced number of ocelli, but it can easily be distinguished due to the significant differences in granulation and the presence/absence of a M-seta on the tibiotarsi.</p> </div>	http://treatment.plazi.org/id/B81687E9F65C7169EDD377A5FEF94273	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F65E7175EDD377A5FCFF4467.text	B81687E9F65E7175EDD377A5FCFF4467.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella alba Babenko & Shveenkova & Potapov 2022	<div><p>Philotella alba sp. nov.</p> <p>Figs 31–37</p> <p>Type material: Holotype: immature male, Khabarovsk Territory, Komsomolsk State Nature Reserve, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.70233&amp;materialsCitation.latitude=49.3635" title="Search Plazi for locations around (long 137.70233/lat 49.3635)">Anyuinski National Park</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.70233&amp;materialsCitation.latitude=49.3635" title="Search Plazi for locations around (long 137.70233/lat 49.3635)">Anyui River</a> (Fig. 1, point 14), mixed coniferous-broadleaf forest, 49°21.81’N, 137°42.14’E, ~ 200 m alt., litter, 08. July 2018. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg. Paratypes: 2 females, 3 immature males and 6 juveniles, same data as holotype; 1 female, 1 male and 3 juvenile, same region and <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=137.57&amp;materialsCitation.latitude=49.303333" title="Search Plazi for locations around (long 137.57/lat 49.303333)">National Park</a>, Tormasu River (Fig. 1, point 13), mixed coniferous-broadleaf forest, 49°18.2’N 137°34.2’E, ~ 200 m alt., litter, 07 August 2018. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg.</p> <p>Additional material. 3 females and 1 male, Khabarovsk Territory, Lazo District, mountain range «Arseniev’s granites»(Fig. 1, point 11), valley of Malyi Katen River, 500-600 m alt., mixed forest, litter, 08 July 2019. A. Brinev leg.; 1 immature male, same region, upper reaches of Katen River, Ko Mountains (Fig. 1, point 10), ~ 500 m alt., mixed forest, litter, 01 July 2018. All A. Brinev leg.</p> <p>Diagnosis. A colourless species of the genus Philotella characterized by: the presence of 2–3 ocelli on each side of the head; the labral setae in the second row similar in length; a labium of the second type; dorsal chaetotaxy with p2 present on Th. II–III but absent on Abd. I–V; three ordinary setae (a3, a4 and m4) in dorsolateral group on Th. II–III and usually two setae (a3 and p3) between axial group and sensilla p4 on Abd. I–III; a strongly thickened sensilla p6 on Th. II and p4 on Abd.4; a complete tibiotarsal chaetotaxy, and toothless unguis.</p> <p>Description. Length (without antennae) of mature specimens 0.34–0.49 mm. Uncoloured, ocular fields sometimes with few granules of dark pigment. Tegument granulation moderate, not especially coarse.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally, ventral separation more or less clear. Ant.4 with simple or slightly divided apical vesicle, external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV blunt and elongate. Antennal organ of Ant.3 typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 11 setae, respectively.</p> <p>Head with 2+2 or 3+3 small ocelli, clearly larger than secondary granules on ocular field (Figs 32–33). PAO usually rounded, more rarely slightly elliptic, consisting of 8–10 vesicles, its axis to ocellus B ratio as 2.0–2.2: 1 (Figs 32–33). Buccal cone short and blunt. Maxilla styliform with two small apical denticles, lamellae not clearly seen. Mandible typical, with five teeth along cutting edge. Labrum with a common set of 4/2352 setae of more or less similar length (Fig. 35). Main part of labium of the second type with three ordinary setae in triangular position and 1+1 axial setae between sensorial elements (Fig. 36); submentum and mentum with usual set of four setae each, i.e. 4+4. Head with 2+2 postlabial setae along ventral line as usual.</p> <p>Dorsal chaetotaxy usually symmetrical and rather reduced (Fig. 31). Ordinary setae short, needle-like, sensilla longer, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: sensilla on Abd. IV and lateral sensilla on Th. II short and thick (Figs 31, 34); Th. II–III with 3+3 axial setae (p2 present) and three setae (a3, a4 and m4) additionally to sensilla p 3 in dorsolateral position; Abd. I–III with 2+2 axial setae (p2 absent) and two setae (a3 and p3) between axial group and sensilla; Abd. IV and Abd. V with only 2+2 axial setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae. Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV (Fig. 37). Number of hr-setae on anal valves variable, probably depended on age.</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 7 setae on coxae, 6, 6, 6 on trochanters, and 13, 12, 11 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Remarks. In specimens from the type series 3+3 uncolored eyes can be seen, as a rule; traces of dark pigment on the ocular fields are less often observed. All specimens from more southern region (Fig. 1, points 10 and 11) have only 2+2 ocelli, which are clearly distinguished by the dark pigment. Otherwise, these individuals are identical to those from the type series.</p> <p>Etymology. The name of the new species reflects one of its characteristic features—the almost complete absence of dark pigment.</p> <p>Affinities. Philotella alba sp. nov. is difficult to compare to the known species of the genus as the main diagnostic features (sensorial equipment of the antennae, labrum and labium, maxillae and mandibles, ventral chaetotaxy) are not unique compared to other species of the genus. Its dorsal chaetotaxy with the presence of setae p2 only on the thoracic terga is also rather usual, being almost identical to that in P. miracli, P. aliniensis sp. nov., and P. amurica sp. nov. Nevertheless, there is one characteristic feature of the new species, i.e. a significant modification of the sensilla on Th. II and Abd. IV, which makes P. alba sp. nov. undoubtedly distinguished from all known species of the genus. A similar modification of the same sensilla is known for a number of species of Micranurida and Anurida Laboulbène, 1865, which can probably be considered as another piece of evidence of the existing generic classification of this part of Pseudachorutinae being artificial.</p> </div>	http://treatment.plazi.org/id/B81687E9F65E7175EDD377A5FCFF4467	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F6407175EDD37699FE7342F3.text	B81687E9F6407175EDD37699FE7342F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella pallida Babenko & Shveenkova & Potapov 2022	<div><p>Philotella pallida sp. nov.</p> <p>Figs 38–43</p> <p>Type material. Holotype: female, Khabarovsk Territory, Lazo District, mountain range «Arseniev’s granites», valley of Malyi Katen River (Fig. 1, point 11), 500–600 m alt., mixed forest, litter, 08 July 2019. A. Brinev leg. Paratypes: 4 females and 1 male, same data as holotype.</p> <p>Diagnosis. A species of the genus Philotella characterized by a pale colouration; the presence of 3+3 slightly coloured ocelli; the labral setae almost equal in length; a labium of the second type, dorsal chaetotaxy with p2 setae present on all terga from Th. II to Abd. III; four ordinary setae (a3, a4, m4 and p4) in dorsolateral group on Th. II–III additionally to sensillum p3 set slightly above seta p4 and usually two setae (a3 and p3) between axial group and sensillum p4 on Abd. I–III; a complete tibiotarsal chaetotaxy, and toothless unguis.</p> <p>Description. Length (without antennae) of available specimens 0.44–0.63 mm, holotype — 0.52 mm. Habitus typical of the genus. Colour light blue, pigment granules sparsely distributed all over dorsal side of body. Tegument granulation moderate, not especially coarse.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV clearly differentiated, blunt and elongate (Fig. 39); few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 12 setae, respectively.</p> <p>Head with 3+3 ocelli, slightly larger than surrounding secondary granules on ocular field. PAO rounded, consisting of 7–9 vesicles, its axis to ocellus B ratio as 1.7–2.2: 1. Buccal cone not especially short. Maxilla styliform, with two small denticles on apex and at least two lamella bearing two apical teeth (Fig. 40). Mandible delicate, with usual five teeth (Fig. 41). Labrum with a common set of 4/2352 setae of similar length, distal pair and two marginal setae of the next row being slightly longer. Main part of labium of the second type with three ordinary setae in triangular position and 1+1 axial setae between sensorial elements (Fig. 42); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae as usual.</p> <p>Dorsal chaetotaxy rather complete, often with some abnormalities (Fig. 38). Ordinary setae short, needle-like, sensilla longer, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: Th. II–Abd. III with 3+3 axial setae (p2 present), dorso-lateral group on Th. II–III usually consisting of four setae (a3, a4, m4 and p4) additionally to sensilla p3, sometimes one of them absent. Abd. I–III with or without p3, i.e. 2+2 or 3+3 p-setae present between sensilla. Abd. V with 2+2 axial setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae (Fig. 43). Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV. Unpaired anal valve usually with three setae hr, number of these setae on paired vales variable.</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, 6 on trochanters, and 13, 12, 11 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Etymology. The name of the new species reflects its light coloration, from the Latin pallidus.</p> <p>Affinities. Philotella pallida sp. nov. is characterized by a quite complete dorsal chaetotaxy, which is comparable only with that of P. caeca sp. nov. (see below). The latter species also has setae p2 on all terga from Th. II to Abd. III, and a complete set of setae (a3, a4, m4, S = p3 and p4) in dorsolateral position on Th. II–III. These two species can easily be distinguished by colouration (light bluish-grey in P. pallida sp. nov., vs colourless in P. caeca sp. nov.), granulation (moderate in P. pallida sp. nov., vs coarse in P. caeca sp. nov.), presence/absence of ocelli, and the number of lobes and shape of the PAO (rounded with 7–9 lobes in P. pallida sp. nov., vs elliptic with 12–15 lobes in P. caeca sp. nov.).</p> </div>	http://treatment.plazi.org/id/B81687E9F6407175EDD37699FE7342F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F6427177EDD377A5FD73424F.text	B81687E9F6427177EDD377A5FD73424F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella caeca Babenko & Shveenkova & Potapov 2022	<div><p>Philotella caeca sp. nov.</p> <p>Figs 44–48</p> <p>Type material. Holotype: female, Khabarovsk Territory, Lazo District, mountain range «Arseniev’s granites», valley of Malyi Katen River (Fig. 1, point 11), ~ 900 m alt., coniferous forest belt, rotten wood, 08 July 2019. A. Brinev leg. Paratypes: 1 female, same habitat, litter; 2 females and 1 juvenile, same area but ~ 1500 m alt., Mount Arseniev, Pinus pumila belt, litter, 08 July 2019. A. Brinev leg.</p> <p>Diagnosis. A colourless species of the genus Philotella characterized by coarse cuticular granulation; the absence of ocelli; an elliptic PAO with 12–15 lobes; a labrum with subequal setae; a labium of the second type; a complete dorsal chaetotaxy with p2 setae present on all terga from Th. II to Abd. IV; tibiotarsi with 19–19–18 setae, and a toothless unguis.</p> <p>Description. Length (without antennae) of available specimens 0.54–0.79 mm. Habitus typical of the genus. White, without traces of dark pigment. Tegument granulation rather coarse.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally. Ant. IV with simple or slightly divided apical vesicle; external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV blunt and elongate; few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present. Ant. I–II with 7 and 12 setae, respectively.</p> <p>Ocelli absent, PAO as a wide ellipse, consisting of 12–15 vesicles (Fig. 45). Buccal cone not especially short. Maxilla styliform, with two small denticles on apex, lamella not clearly seen. Mandible with usual five teeth. Labrum usually with a common set of 4/2352 setae of similar length, but abnormalities frequent. Main part of labium of the second type with three ordinary setae in triangular position and 1+1 axial setae between sensorial elements (Fig. 46); submentum and mentum with usual set of four setae each. Head ventrally with 2+2 postlabial setae as usual.</p> <p>Dorsal chaetotaxy complete (Fig. 44). Ordinary setae short, needle-like, sensilla longer, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: Th. II–Abd. IV with 3+3 axial setae (p2 present); dorsolateral group on Th. II–III usually consisting of four setae (a3, a4, m4 and p4) additionally to sensilla p3 set in forward position, sometimes one of them absent; Abd. I–III with p3 present, i.e. three p-setae present between axial line and sensilla; Abd. V with 2+2 axial setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae (Fig. 47). Furcal remnant in a form of two swellings, each with a seta and 2–3 tiny setules, on anterior border of Abd. IV (Fig. 48). Number of setae hr on anal valves in available mature specimens unstable.</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 6 setae on coxae, 6, 6, 6 on trochanters, and 12, 11, 10 setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless.</p> <p>Etymology. This is the first described species of the genus without ocelli, the name reflecting this: from the Latin caecus —blind.</p> <p>Remarks. In our material, there are two more undescribed forms which are characterized by the absence of a dark pigment on the body. One of them, from the Amur Region (Zeya State Nature Reserve, Fig. 1, point 1), like P. caeca sp. nov., lacks ocelli, but distinguished by rounded PAO with a smaller number of lobes, a posterior position of sensilla in dorsolateral groups on Th. II–III (i.e., p4 setae absent), and shortened and slightly widened sensilla on Abd. IV. The second one from Southern Primorye (Fig. 1, point 5) has 3+3 small and uncoloured ocelli, and its dorsal and ventral chaetotaxy is almost identical to that of P. caeca sp. nov. Most likely, these specimens represent distinct species, but the shortage of material does not allow us to describe them.</p> <p>Affinities. The species possesses all diagnostic features of the genus Philotella, but lacks ocelli. This combination of characters is not known among the described species of the genus, although Najt &amp; Weiner (1997) mentioned the existence of a similar form in New Caledonia.</p> </div>	http://treatment.plazi.org/id/B81687E9F6427177EDD377A5FD73424F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F6447171EDD377A5FECC42F4.text	B81687E9F6447171EDD377A5FECC42F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella porcella (Fjellberg 1985) Babenko & Shveenkova & Potapov 2022	<div><p>Philotella porcella (Fjellberg, 1985) comb. nov.</p> <p>Figs 49–55</p> <p>Studied material: 1 female, Far East, Amur Region, ~ 3 km west of Arkhara (Fig. 1, point 2), broadleaf forest (Quercus, Tilia, Betula, Ulmus), litter, 17 August 2014, M. Potapov &amp; N. Kuznetsova leg.; 1 male and 1 juvenile, Khabarovsk Territory, Komsomolsk State Nature Reserve, Gorin River (Fig. 1, point 12), mixed coniferous-broadleaf forest, 50°44.25’N 137°25.43’E, ~ 250 m alt., rotting wood, 10 August 2018. N. Kuznetsova, A. Kuprin &amp; A. Geraskina leg.; 6 juveniles, same region, State Nature Reserve and collectors, Tormasu River (Fig. 1, point 13), mixed coniferous-broadleaf forest, litter, 49°18.2’N 137°34.2’E, ~ 200 m alt., 07 August 2018; 7 females, 2 males and 3 juveniles, Khabarovsk Territory, Lazo District, mountain range «Arseniev’s granites» (Fig. 1, point 11), valley of Malyi Katen River, ~ 500–600 m alt., mixed forest, litter and soil, 08 July 2019; 3 females, 2 males and 4 juveniles, same area, but ~ 900 m alt., forest belt, rotten wood, 08 July 2019; 3 females, 1 male and 3 juveniles, Southern Primorye, Partisan District, Mount Olkhovaya (Fig. 1, point 6), mixed forest in valley, 43°18.35’N 133°40.07’E, ~ 500 m alt., litter, 20 August 2018; 1 juvenile, same area, but 43°20.83’N 133°39.38’E, ~ 1600 m alt., P. pumila belt, soil, 20 August 2018. M. Potapov, Yu. Shveenkova &amp; A. Kuprin leg.</p> <p>Description. Length (without antennae) 0.5–0.8 mm. Colour dark blue, uniform dark. Tegument granulation strong, especially on abdominal tip.</p> <p>Antennae clearly shorter than head, Ant. III–IV fused dorsally, ventral separation more or less clear. Ant. IV with slightly divided apical vesicle, external ms, subapical or and seta i present; six sensilla (S1–S4, S7–S8) on dorsal side of Ant. IV blunt and elongate (Fig. 54), few sensilliform setae of various shape also present on its ventral side. Antennal organ of Ant. III typical, inner sensilla small, sgv longer than sgd and curved, ventral ms present.Ant. I–II with 7 and (11)12 setae, respectively.</p> <p>Head with 5+5 small ocelli, about as large as secondary granules on ocular field. PAO usually rounded, more rarely slightly elliptic, consisting of 8–10 vesicles, its axis to ocellus B ratio as 1.8–2.0: 1 (Fig 50). Buccal cone short and blunt. Maxilla styliform, lamellae not clearly seen. Mandible typical, with five teeth along cutting edge. Labrum with a common set of 4/2352 setae of different length, proximal pair of labral setae being the longest (Fig. 51). Main part of labium of the first type with three ordinary setae in line and 2+2 axial setae between sensorial elements (Fig.52); submentum and mentum with usual set of four setae each, i.e. 4+4. Perilabial area with 5+5 setae, 2+2 postlabial setae along ventral line present as usual.</p> <p>Dorsal chaetotaxy usually symmetrical and rather reduced (Fig. 49). Ordinary setae short, needle-like, sensilla longer, their number as usual: 22/11111, lateral ms present on Th. II. Main characteristics: head with unpaired seta d0 and only 2 setae on ocular field; Th. I usually with 2+2 setae, but variant with 3+3 setae also seen; only 2+2 axial (dorso-internal) setae present on all terga from Th. II to Abd. V; Th. II–III with only two setae additionally to p3 (=S) in dorso-external position, p3 set clearly in front of p4 on both thoracic terga; Abd. I–III with only one seta between axial group and sensillum p4; both Abd. IV–V with only 2+2 axial setae between sensilla.</p> <p>Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, ventral side of Abd. II–III without unpaired axial setae. Furcal remnant in a form of two swellings, each with a seta, on anterior border of Abd. IV (Fig. 55). Number of setae hr on anal valves variable, probably depended on age.</p> <p>Legs 1–3 with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 6, 7 setae on coxae, 6, 6, 6 on trochanters, and 13, 12, 11(12) setae on femora. Tibiotarsi with most complete set of setae: 19, 19, 18, seta M present. Unguis toothless (Fig. 53).</p> <p>Remarks. The conspecificity of this Far Eastern form with the true P. porcella comb. nov., the latter species described from central Alaska, raises certain doubts. However, it is impossible to reliably distinguish it from the Alaskan species on the basis of the available description. The existing differences, for example, the number of setae on Th. I, are insignificant and variable. The most characteristic feature of P. porcella comb. nov. (and the above Far Eastern specimens as well) is the anterior position of the sensilla in the dorsolateral groups on Th. II–III. This position of sensilla makes it distinguished from all known species of the genus with a reduced chaetom, but it has been noted for P. obesa, P. pallida sp. nov. and P. caeca sp. nov., which have a more complete set of setae on the thoracic segments.</p> <p>In addition to the peculiar thoracic chaetotaxy, fairly good diagnostic features of P. cf. porcella are a strongly elongate pair of proximal setae on the labrum (a character unique for the set of species under consideration, but not known for Alaskan individuals), and a labium of the first type (shared only with P. deharvengi, P. pax sp. nov. and P. cedrina sp. nov.).</p> </div>	http://treatment.plazi.org/id/B81687E9F6447171EDD377A5FECC42F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
B81687E9F646717DEDD377A5FE9343EB.text	B81687E9F646717DEDD377A5FE9343EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philotella Najt & Weiner 1985	<div><p>A key to the East Palaearctic species of the genus Philotella *</p> <p>1 Th. II–III with 1+1 axial setae (Fig. 58)........................................... japonica (Tamura) comb. nov.</p> <p>- Th. II–III with more axial setae.......................................................................... 2</p> <p>2 Th. II–III with 3+3 axial setae (p2 present) (Fig. 56)......................................................... 3</p> <p>- Th. II–III with 2+2 axial setae (p2 absent) (Fig. 57)......................................................... 12</p> <p>3 Abd. I–III with 3+3 axial setae (p2 present) (Fig. 59)......................................................... 4</p> <p>- Abd. I–III with 2+2 axial setae (p2 absent) (Fig. 60)......................................................... 7</p> <p>4 Tibiotarsi with 18–18–17 setae, i.e. seta M absent (Fig. 61).................................. olgae Kniss &amp; Thibaud</p> <p>- Tibiotarsi with 19–19–18 setae, i.e. seta M present.......................................................... 5 5 Dark coloured species with 5+5 ocelli; dorsolateral group on Th. II–III with two ordinary setae additionally to sensillum (Fig. 62); Abd. I–III with one seta between axial group and sensillum (Fig. 64)............................ obesa Babenko</p> <p>- White species with or without ocelli; dorsolateral group on Th. II–III with 3–4 ordinary setae additionally to sensillum (Fig. 63); Abd. I–III with two seta between axial group and sensillum (Fig. 65)........................................ 6 6 3+3 uncoloured ocelli present; PAO rounded with 7–9 vesicles (Fig. 66)............................. pallida sp. nov.</p> <p>- Ocelli absent; PAO elliptical with 12–15 vesicles (Fig. 67)......................................... caeca sp. nov.</p> <p>7 Tibiotarsi with 18–18–17 setae, i.e. seta M absent (Fig. 61).................................................... 8</p> <p>- Tibiotarsi with 19–19–18 setae, i.e. seta M present.......................................................... 9</p> <p>8 5+5 ocelli; Abd. I–III with one setae between axial group and sensillum (Fig. 68); Abd. IV with 2+2 setae between sensilla (Fig. 70)............................................................................... miracli Najt &amp; Weiner</p> <p>- (2)3+3 ocelli; Abd. II–III with two setae (a3 and p3) between axial group and sensillum (Fig. 69); Abd. IV with 3+3 setae between sensilla (Fig. 71)................................................................... amurica sp. nov.</p> <p>9 Main part of labium with 2+2 setae between sensorial elements, i.e. labium of the first type (Fig. 72)................. 10</p> <p>- Main part of labium with 1+1 setae between sensorial elements, i.e. labium of the second type (Fig. 73)............... 11</p> <p>10 3+3 ocelli present; two marginal setae of the second row on labrum clearly longer (Fig. 74); lateral sensilla on Th. II and sensilla of Abd. IV shorter and clearly thickened (Fig. 76)............................................... cedrina sp. nov.</p> <p>- 5+5 ocelli present; three setae in the second labral row are longer (Fig. 75); all dorsal sensilla similar (Fig. 77)...................................................................................................... pax sp. nov.</p> <p>11 Lateral sensilla on Th. II and dorsal ones on Abd. IV short and strongly thickened (Fig. 78); granulation moderate.................................................................................................. alba sp. nov.</p> <p>- All sensilla on terga similar in shape; tegument granulation very coarse............................ aliniensis sp. nov.</p> <p>12 Dorsolateral group on Th. II–III with sensilla p 3 in forward position (Fig. 79)............ porcella (Fjellberg) comb. nov.</p> <p>- Dorsolateral group on Th. II–III with sensilla p 3 in backward position below ordinary setae (Fig. 80)................. 13</p> <p>13 Dark blue coloured species with 3+3 or 5+5 ocelli.......................................................... 14</p> <p>- Body «bluish gray», paler ventrally; 2+2 ocelli present.............................. grana (Lee &amp; Kim) comb. nov.</p> <p>14 Tibiotarsi with 18–18–17 setae, i.e. seta M absent (Fig. 61); 5+5 ocelli......................................... 15</p> <p>- Tibiotarsi with 19–19–18 setae, i.e. seta M present; 3+3 ocelli.................................. oligotricha sp. nov.</p> <p>15 Labium of the first type with 2+2 setae between sensorial elements (Fig. 72); Abd. IV– V with only 1+1 setae between sensilla (Fig. 81)....................................................................... deharvengi Najt &amp; Weiner</p> <p>- Labium of the second type with 1+1 setae between sensorial elements (Fig. 73); Abd. IV– V with 2+2 setae between sensilla (Fig. 82)............................................................................ tertia Najt &amp; Weiner</p> <p>* Species of the Micranurida / Philotella complex of the region with elongated antennal sensilla not included in the key:</p> <p>Two more species of this complex, which position in the existing generic classification is not clear, are known in East Asia. These are M. montana Lee &amp; Kim, 1984 and M. tuberculata Lee &amp; Kim, 1984, both described from South Korea and both showing elongated antennal sensilla. Therefore, neither fits the diagnosis of the genus Micranurida sensu Deharveng (1982). This was well known to the authors, who wrote that the situation “… prevents us from assigning the present taxon neither to Rusekella nor to Micranurida sensu Deharveng ”, but “… to leave it, for the time, with Micranurida sensu Massoud which certainly is polyphyletic ” (p. 18). Najt &amp; Weiner (1985) also considered that these species did not fit the diagnosis of the genus Philotella, even when it was expanded to include species with reduced ocelli.</p> <p>M. montana is a large species (up to 2.3 mm) with 5+5 ocelli and very peculiar chaetotaxy. For instance, it has a2 setae on Th. II–III, which is not at all characteristic of this generic complex (Fjellberg 1985a). The dorsal chaetotaxy of M. tuberculata, which has 2+2 ocelli, is more similar to that of species of the genus Philotella, but its serrate maxilla with 8–9 minute teeth (p. 18) is not typical of the genus.</p> <p>Species that do not correspond to the diagnosis of the genus Micranurida sensu Deharveng (1982), but are listed at www.collembola.org within this genus, are also present in other areas of the Holarctic. In Europe, these are M. anophthalmica and M. forsslundi; in North America, M. furcifera and M. valiana Fjellberg, 1985. All of them have elongated antennal sensilla, which is more consistent with the existing diagnosis of Philotella. Obviously, while maintaining the validity of the latter genus, in the future it will be necessary to clarify the generic status of all forms listed above.</p> </div>	http://treatment.plazi.org/id/B81687E9F646717DEDD377A5FE9343EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Babenko, Anatoly;Shveenkova, Yulia;Potapov, Mikhail	Babenko, Anatoly, Shveenkova, Yulia, Potapov, Mikhail (2022): New species of the genus Philotella Najt & Weiner, 1985 (Collembola, Neanuridae, Pseudachorutinae) from the Russian Far East. Zootaxa 5174 (5): 525-550, DOI: https://doi.org/10.11646/zootaxa.5174.5.3
