taxonID	type	description	language	source
A0384152FF9A02384DE7FB8DFF5CF98C.taxon	description	(Figs 1 – 17)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9A02384DE7FB8DFF5CF98C.taxon	diagnosis	Diagnosis. Body elongated, small to large; pronotum usually widest in anterior 1 / 3, rarely in 1 / 2 of its length; subantennal fossae deep; apical labial palpomere articulated on the side of the preapical one; trigonium of penis not divided, parameres of tegmen without modifications.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9A02384DE7FB8DFF5CF98C.taxon	description	Redescription. Male. Body elongated, small to large (TL 2.4 – 6.9 mm), 1.9 – 2.4 × longer than wide, sides of elytra subparallel, moderately convex. Body yellowish-brown to dark brown, legs and antenna often lighter. Punctation on head and pronotum usually granulate, strong and dense, punctures often touching each other, punctation on elytra simple, not granulate. Body covered with short to long, adherent to erect setae. Head rather wide, W / L ca. 1.2, base of head covered with pronotum. Eyes big, oval, slightly protuberant, gently faceted. Distance from posterior margin of eye to occipital ridge distinctly longer than diameter of eye. Distance from the ventral margin of the eye to subgenal ridge (Figs 3 A, C) varies from 0.4 – 1.0 diameter of eye. Supraantennal ridges (Fig. 3 E) well-marked, slightly to significantly elevated, extending into clypeal margin. Subantennal fossae (Fig. 3 A) deep, wide, reach margin of eye and toward bottom of head, separated from eye with subocular carinae. Frontoclypeal suture absent. Clypeus short, strongly transverse, lateral margins strongly converging anteriorly, anterior edge straight to subtly semicircularly emarginated. Anteclypeus well visible, narrower and slightly longer than clypeus. Ventral side of head between subgenal ridges concave. Gular area convex, square. Submentum wider and slightly longer than mentum, flat, hexagonal, square to transverse, separated from gula. Antennae filiform; antennomere 1 cylindrical, broad, slightly flattened; antennomere 2 globular to cylindrical, the shortest; antennomere 3 narrow, ca. 2.2 × longer than antennomere 2, antennomeres 4 – 11 distinctly wider than antennomere 3, antennomere 4 slightly longer than antennomere 3, antennomeres 4 – 10 as long as antennomere 3, apical antennomere longer, as long as antennomeres 2 and 3 combined. Labrum (Fig. 4 C) narrower than clypeus, transversely rectangular, anterior margin deeply semicircularly indented, with long setae, front angles rounded. Mandibles (Fig. 4 A) slightly asymmetrical, narrowly triangular, ca. 2 × longer than width at bases, apices unidentate, acute; with short carina on the external edge, abruptly curved in apical third, with setae covering 2 / 3 of its length; internal edge with triangular denticle and carina with 3 smaller denticles, not covered with setae, denticles developed somewhat better on left mandible; prostheca absent; molar region without denticles or setation. Maxillae (Fig. 4 C): basistipes with distinct setation; maxillary palpi short; palpomere 1 poorly subconical, slightly longer than wide, palpomeres 2 and 3 about 2 × longer than palpomere 1, apical maxillary palpomere about same length as palpomere 3, slightly narrower, bulbous at base and pointed at apex, covered with setae at basal 2 / 3. Galea slightly elongated, apex with two types of setae — some of setae short and straight, forming fan-shaped structure, other setae much longer, “ feathered ”, and irregularly arranged. Lacinia elongated, covered with straight, sparse setae in the lower half and with longer, denser, bulkier on apex. Labium (Fig. 4 D): mentum subtrapezoidal, as wide as long; praementum strongly sclerotized, narrow at base, then strongly widening, anterior edge semicircular, covered with numerous straight setae. Labial palpomere 1 angular, long, without setae; palpomere 2 broadly oval, larger than palpomere 1; apical palpomere inserted laterally, oval, slightly shorter than palpomere 2, without setae. Pronotum (Fig. 5) transverse, subrectangular, narrower at base than in anterior part, in most species widest in anterior 1 / 3. Lateral edges rounded, even or crenulate. Anterior margin slightly rounded, not projecting beyond anterolateral angles, covering base of head. Anterolateral angles subtriangular, often acute and projecting. Posterolateral angles obtuse to acute, well-marked. Basal portion of pronotum with well-marked groove, basal margin of pronotum bisinuate (Fig. 3 D). Prosternal process elongate (Figs 3 B, C), ca. 3 – 4 × longer than wide, lanceolate to fusiform, apex narrow, bent ventrally, covered with setae. Procoxae long, subconical, procoxal cavities widely open. Scutellar shield triangular, subequilateral (slightly longer than it is wide), flat, with slightly rounded edges and acute apex. Elytra elongated, sides subparallel in proximal 1 / 2, angle between elytra and pronotum well-marked in dorsal view. Elytra distinctly wider than pronotum. Humeri pronounced, apices regularly rounded. Elytra in most species without longitudinal carinae. Epipleuron flat, narrow, slightly narrower than metepisternum at base, then regularly tapering towards apex of elytra, complete to apex. Sides of elytra not explanate. Hind wings (Fig. 6) ca. 2.2 × longer than its width; radial cell subtriangular; apical portion of r 3 well visible, straight; RML arcuate, forming acute angle with MP 1 + 2; CuA 1 long, oblique, joining MP 3 + 4 and CAS; anal field with well-markedAP 3 + 4. Medial field with 3 free veins; pigmented areas in radial cell, medial, radial, central and apical fields. Mesoventrite short, anterior edge with pair of oblique procoxal rests, suture present throughout mesoventral process. Prosternal process rest (Fig. 3 B) triangular, narrow to wide, with slightly rounded edges, slightly concave. Mesoventral process (Fig. 3 B) triangular to subrectangular, ca. 1.5 × longer than wide, apex bilobed. Mesocoxal cavities widely separated. Mesepisterna moderately wide, rhomboid, middle part with depression for procoxae. Mesepimeron narrower than mesepisternum, triangular. Mesocoxal cavities closed laterally by narrow processes of mesepimeron and metasternum. Mesocoxae weakly elongated, ovally transverse (ca. 1.6 × longer than wide), slightly protruding and triangular. Metaventrite long, convex. Metaventral discrimen present in posterior 1 / 4 – 1 / 2. Metepisternum wide, rhomboid. Metacoxae big, ca. 3.5 × longer than wide. Metacoxal plates touching each other mesally over the entire length. Legs long, femora not enlarged, not saltatorial, extending beyond elytral margin; tibiae slightly longer than femora, slender and, especially metatibiae, subtly S-shaped, slightly extended at apex; with well-marked paired longitudinal carinae; tibial spurs well developed, dorsal spurs ca. 1.5 × longer than ventral. Abdomen longer than wide, moderately convex. Ventrites 1 – 5 of subequal length, regularly covered with setae; ventrite 5 rounded to subtriangular, apex rounded. Morphology of terminal sclerites of males uniform (Figs 7 A, C). Sternite VIII transversely rectangular, Ushaped with very few setae and punctures concentrated on apical margin, distinctly wider than longer; sternite IX rectangular, consisting of subtriangular hemisternites, apical part covered with medium length setae, apodemes short and barely visible. Tergite VIII rectangular with rather distinct apodemes, numerous short setae in apical part; tergite IX elongated, apical part membranous, almost devoid of setae, with long apodemes. Aedeagus symmetrical. Tegmen with small styli, parameres widely rectangular, with rounded apical portions to narrowly triangular with pointed apices, with scarce punctures and numerous rows of delicate setae in apical part. Trigonium of penis rather narrow, apex rounded or pointed, shorter than parameroids or as long (rarely slightly longer) as parameroids, rarely trigonium widely oval, bulbous. Pala triangular, slightly longer than parameroids, in some species basal portion emarginated. Parameroids elongated, subtriangular to subrectangular, apices rounded, tempered or pointed, apical part covered with punctures. Female. Many species with significant sexual dimorphism. Pronotum often more transverse than in males, punctation of pronotum often sparser than in males, with shining interspaces. Punctation of elytra often coarser than in males. Hind wings often reduced — brachypterous (Fig. 6 B), rarely apterous, no infraspecific variation in wing reduction was observed. Base of elytra in some species closely fitting pronotal base, as a result, humeral portion may project anteriorly. Elytra of species exhibiting wing reduction with curved lateral sides and faintly marked humeri. Abdominal ventrite 4 (in some species also ventrite 3, and occasionally also 5) with paired oval or transverse patches of long, dense setae (Fig. 8). Apex of ventrite 5 usually deeply emarginated at apex (e. g. Figs 8 C, J). Tergite VII modified in some species, its posterior portion forming rectangular flap or concavity. Ovipositor (Fig. 9) long, branchlets absent; coxites narrow, membranous, styli apical; bursal sclerites present, either small, rod-like (Fig. 9 B) or larger, plate-like (Fig. 9 A).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9A02384DE7FB8DFF5CF98C.taxon	discussion	Remarks. Sharp (1878) erroneously described labial palpi of Atopida as “ not forked ”, i. e. the apical palpomere arising from the top of the penultimate one. This mistake was corrected by Broun (1893) who examined several species of Atopida, all with furcate labial palpi, i. e. with apical palpomere arising from the side of the penultimate one.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	description	(Figs 1 A – D, 3 A – B, 5 A – B, 8 A, 10 A, 11, 12 A, 17 A)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	materials_examined	Type material. Holotype (NHM): pinned into card, “ Type ” [round label with red border], “ New Zealand ” [handwritten], “ Atopida castanea White \ Zool. Ereb. & Terror \ Kaudi ... [illegible] ” [handwritten]. Other material examined. 1 female (NZAC): “ Tauraroa \ 24 - ii- 29 ”; “ Tauraroa \ Mt Auckland ”, “ coll. \ E. Fairborn \ 24 - ii- 1929 ”, “ A. E. Brookes \ Collection ”; 1 female (AMNZ): “ Mangamuka \ R. \ N. Auk. \ 10 - I- 24 O. E. C. ”, “ C. E. Clarke \ Collection ”; 2 females (AMNZ): “ Mangamuka \ Mt \ N. Auck. \ 9. I. 24 ”, “ C. E. Clarke \ Collection ”; 2 exx. (NZAC): “ NEW ZEALAND AK \ Sharps Bush \ Henderson \ 23 Jan 1982 \ P. Maddison ”, “ Beating ”; 1 ex. (NZAC): “ In tank \ Riverhead \ 16 Dec 1942 \ D McKenzie ”; 1 ex. (NZAC): “ Riverhead \ X light trap \ T. G. Edwards ”, “ 26 XII 54 ”, “ J. C. Watt \ Collection \ Ent. Div. \ DSIR, 1966. ”; 1 male (AMNZ): “ Albany \ 28. I. 1983 \ Elaine Power ”, “ Waitemata Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”; 1 male (AMNZ): “ Forest SWEPT \ Ngaiotonga \ 10. X. 1968 \ K. A. J. Wise ”; 1 ex. (NHM): “ Tairua ”, “ New Zealand \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”; 1 ex. (NZAC): “ Whangarei \ 28.12.26 ”, “ A. E. Brookes \ Collection ”; 1 ex. (NZAC): “ NEW ZEALAND CL \ Great Barrier Island \ 27 Jan 1980; G. Lowe \ Between Moco Valley & Stn Rk ”, “ In bush ”, “ G. Lowe \ Collection \ July 1980 ”; 1 male (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220 m, \ 7. xi. - 11. xii. 2001, P. \ Sutton & J. Gilbert. ”, “ Forest edge, Malaise \ trap. L 11043. ”; 1 male (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, \ Little Windy Hill, 220 m, \ 25. x. - 21. xi. 2002, \ P. Sutton. ”, “ Forest edge, Malaise \ trap. L 11053. ”; 4 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220 m, \ 11. xii. 2001 - 18. i. 2002, \ P. Sutton. ”, “ Forest edge, Malaise \ trap. L 11044. ”; 1 ex. (NZAC): “ NEW ZEALAND CL \ Little Barrier I \ 183 m 29 Feb 1976 \ A. K. Walker \ sweeping ”; 1 female (NZAC): “ NEW ZEALAND CL \ Little Barrier I \ Main track to \ Hauturu Mt 198 m \ 20 - 23 Feb 1976 ”; 1 ex. (NZAC): “ NEW ZEALAND CL \ Little \ Barrier I \ 17 Dec 1983 \ J. Playfair ”; 2 exx. (NZAC): “ Little Barrier \ Island ”, “ Collector \ 29 / 12 / 31 - 10 / 1 / 32 \ A. E. Brookes ”; 1 male (AMNZ): “ Summit Track \ 549 - 579 m \ Little Barrier I \ 19. I. 1983 \ K. A. J. Wise ”; 1 male (AMNZ): “ Summit Track \ 335 - 549 m \ Little Barrier I \ 19. I. 1983 \ K. A. J. Wise ”; 1 male (AMNZ): “ L. Barrier Is. \ XII. 1963 \ M. B. Paterson ”, “ off Rodney Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ M. B. Paterson \ Collection ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	diagnosis	Diagnosis. Body large (TL ca. 5.4 – 6.9 mm), maroon brown. Trigonium narrow, same length as parameroids; pala triangular, slightly longer than parameroids; tegmen with wide parameres. Similar to A. suturalis, but larger, elytra never contrasting yellowish, penis more elongated, pronotum widest in anterior 1 / 3 of its length, pronotum of females covered with coarse, granulate punctures also in posterior portion, no shining interspaces present.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	description	Redescription. Male. Body elongated, large (TL 5.5 – 6.9 mm), moderately convex. Body maroon brown, head and pronotum slightly darker, elytra, legs and antenna slightly lighter. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on elytra simple, punctures elongated, small and dense, punctures separated by 0.7 – 1.0 × diameter of a puncture. Body covered with moderately long yellowish setae, slightly ruffled on head and rather adherent on pronotum and elytra. Supraantennal ridges well-marked, slightly elevated. Pronotum widest in anterior 1 / 3, sides curved, anterolateral angles not produced, posterolateral angles obtuse, rounded. Elytra without carinae, humeri well-marked, elytral interspaces dull. Ventrite 5 rounded, apex truncate. Penis (L 0.91 mm, W 0.38 mm) elongated, trigonium narrow, apex truncate, same length as parameroids; parameroids elongated, apices obliquely truncate, with slightly irregular edge, apical part covered with punctures; pala triangular, slightly longer than parameroids. Tegmen (L 0.74 mm, W 0.64 mm) with wide, subrectangular to oval parameres, apices rounded, apical part covered with scarce punctures and numerous rows of delicate setae. Female. Body large (TL 5.4 – 6.5 mm), elongate, moderately convex, macropterous. Supraantennal ridges wellmarked, slightly elevated semicircularly above bases of antennae. Pronotum covered with coarse punctation, widest in anterior portion, anterolateral angles explanate, well-marked, posterolateral angles obtuse. Elytra covered with very short, adherent setae, shorter than in males, not reaching next puncture, elytral interspaces shining, punctation coarse, punctures separated by 0.5 × diameter of a puncture. Ventrite 5 rounded, with arcuately emarginated apex, tergite VII with transverse rectangular sclerotization. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite (L 0.40 mm, W 0.19 mm) consisting of two rod-like sclerites in anterior portion and two additional, well pigmented sclerites hooked at posterior end. Measurements and ratios. Males (n = 4) TL 5.50 – 6.9 mm (6.0 mm), PL 1.0 – 1.33 mm (1.13 mm), PW 1.60 – 2.15 mm (1.84 mm), EL 4.40 – 5.60 mm (4.91 mm), EW 2.40 – 3.21 mm (2.76 mm), TL / EW 2.07 – 2.33 (2.18), PW / PL 1.60 – 1.70 (1.63), EL / EW 1.69 – 1.97 (1.79), EL / PL 4.19 – 4.72 (4.37), EW / PW 1.49 – 1.53 (1.50). Females (n = 6) TL 5.40 – 6.50 mm (5.92 mm), PL 1.10 – 1.20 mm (1.13 mm), PW 1.70 – 2.19 mm (1.94 mm), EL 4.33 – 5.40 mm (4.81 mm), EW 2.30 – 3.12 mm (2.69 mm), TL / EW 2.06 – 2.35 (2.21), PW / PL 1.55 – 1.83 (1.71), EL / EW 1.67 – 1.88 (1.79), EL / PL 3.94 – 4.58 (4.25), EW / PW 1.28 – 1.42 (1.39).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	distribution	Distribution. The species occurs in northern part of North Island, in Northland, Auckland and Coromandel areas (Fig. 18 A).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF94023C4DE7F9C4FAB3FE23.taxon	discussion	Remarks. The species is rather rare in the collections, what is a bit surprising for a species described already in 1840 s. Probably its relatively big size attracted attention of White. Females that we identified as A. castanea differ considerably in the details of the 1) pronotum shape, and 2) elytral setation. Some of the specimens have distinctly explanate anterolateral angles (especially specimens from Mangamuka, Fig. 1 B), while other do not exhibit this feature (Fig. 1 D). Setation of elytra is either short and adherent or longer and semierect. As the morphology of bursal sclerite seems to be rather stable in all the specimens we assume the above mentioned differences should be attributed to infraspecific variation, although the problem needs to be studied more thoroughly in future.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	description	(Figs 1 E – F, 5 C – D, 8 B, 10 B, 12 B, 17 D)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	materials_examined	Type material. Holotype of A. hirta (NHM), male: card-mounted, “ 564 ”, “ Mount \ Manaia ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Atopida \ hirta. ” [in Broun’s hand]. Holotype of A. grahami, female (NHM): card-mounted, “ 3070. ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Rangiahua \ Hokianga ” [in Broun’s hand], “ Atopida \ grahami ” [in Broun’s hand]. Other material examined. 1 male (AMNZ): “ NEW ZEALAND, ND \ Waima \ 35 ° 34 ’ 173 ° 34 ’ \ 1. x. 1985 \ R. F. Gilbert ”, “ Swept track to \ Oraora saddle. ”; 1 female (NZAC): “ NEW ZEALAND, ND \ Waima Forest \ Wekaweka Tk \ 13 Jun 2000 \ R. Leschen, G. Hall \ leaf litter RL 583 \ 35 ° 34 ’ S, 173 ° 36 ’ E ”; 2 males (NZAC): “ NEW ZEALAND ND \ Long Louis \ Waima Forest \ 13 Feb 2008 \ R Leschen B Michaux ”, “ beating \ RL 13330 \ 35 ° 32.222 ’ S 173 ° 27.238 ’ E ”; 1 ex. (NHM): “ Tairua ”, “ New Zealand \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”; 9 males, 6 females (NZAC): “ NEW ZEALAND AK \ Waitakere Ra \ Sharps Bush \ Feb 1981 \ J. C. Watt ”; 1 female (NZAC): “ NEW ZEALAND AK \ Sharps Bush \ Herderson \ 23 Jan 1982 \ P. Maddison ”, “ Beating ”; 1 male (NZAC): “ NEW ZEALAND AK \ Sharps Bush \ Henderson \ 23 Jan 1982 \ J. Ramsay ”, “ Beating ”; 3 males (AMNZ): “ Sharps Bush \ Waitakere Ra. \ 24. I. 1982 \ K. A. J. Wise ”; 1 male (NZAC): “ NEW ZEALAND, ND \ Waipoua SF \ Waipoua R \ 30 Oct 1985 \ R. C. Craw ”; 2 exx. (DBET): New Zealand ND 31: Trounson Kauri Park, - 35.7221, 173.6470, 240 m, beating, 8.12.2016, leg. A. Kiałka; 1 female (NZAC): “ FERN FLAT RD. \ MAUNGATANIWHA \ RANGE 12000 FT. \ 20 FEB. 1967 \ B. M. May ”; 1 female (NZAC): “ L. Rotorua \ 1 - 3. II. 39 \ E. S. Gourlay ”; 1 female (NHM): “ NEW ZEALAND Kaipara \ Mt. Auckland 22 Feb 1986 \ beating dead wood C. Lyal \ BMNH {E} 1986 - 274 ”; 1 male (NHM): “ var / \ 564. ”, “ Tairua ”, “ New Zealand. \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ hirta coll. \ Grenn ”; 1 male (AMNZ): “ Waimatenui \ 19 - 3 - 47 \ C. E. C. [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”; 1 male (AMNZ): “ swept entance to walkway \ Manginangina Scenic Res. \ 12. XI. 1985 \ R. F. Gilbert \ NZMSINII 308600 \ 35 ° 12 ’ 173 ° 48 ’ ”; 1 male (AMNZ): “ NEW ZEALAND AK \ Atuanui \ Mt Auckland \ pitfall B 13 \ Feb 2002, A. Warren ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	diagnosis	Diagnosis. Body small (TL 2.2 – 3.5 mm), narrow, covered with erect setae, elytra often yellowish-brown with blackish suture. Trigonium narrow, rectangular, with parallel sides and truncate apex, same length as parameroids; pala triangular, slightly shorter than parameroids; tegmen elongated, subrectangular in outline, with subparallel outer edges of parameres. Size, colouration, and setation make this species similar to A. montana. Both species are distinguishable on the basis of morphology of genitalia.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	description	Redescription. Male. Body elongated, small (TL 2.4 – 3.5 mm), moderately convex. Body brown, legs and lower half of antenna yellowish brown. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on elytra simple, dense, punctures separated by 0.5 × diameter of a puncture. Body covered with yellowish setae, partly shorter and semi-erect, partly longer and protruding. Supraantennal ridges wellmarked, slightly elevated. Pronotum widest in anterior 1 / 3, sides curved, anterolateral angles distinctly produced, sharp and triangular, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex slightly emarginated. Penis (L 0.74 mm, W 0.26 mm) elongate and narrow, trigonium very slender, rectangular, with parallel sides and truncate apex, same length as parameroids; parameroids elongated, sides of parameroids subparallel, with few punctures in apical part; pala triangular, slightly shorter than parameroids, basal edge concave. Tegmen (L 0.67 mm, W 0.35 mm) narrow, sides subparallel, parameres triangular at apices, with scarce punctures and delicate setae on the inner, membranous edges. Female. Body small (TL 2.2 – 2.5 mm), macropterous. Supraantennal ridges well-marked, slightly elevated. Punctation becoming sparser in central and basal portion of pronotal disc, with shining interspaces, punctation on elytra denser, punctures separated by 0.3 × diameter of a puncture. Ventrite 5 rounded, with semicircularly emarginated apex, tergite VII with elongated subtriangular sclerotization in apical portion. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite consisting of two small lateral rod-like sclerites (L 0.15 mm) widened in posterior portion and additional faintly visible central sclerite. Measurements and ratios. Males (n = 4) TL 2.43 – 3.50 mm (3.11 mm), PL 0.43 – 0.67 mm (0.57 mm), PW 0.63 – 1.03 mm (0.90 mm), EL 1.93 – 2.93 mm (2.54 mm), EW 1.10 – 1.57 mm (1.41 mm), TL / EW 2.17 – 2.23 (2.21), PW / PL 1.46 – 1.72 (1.58), EL / EW 1.76 – 1.87 (1.80), EL / PL 4.15 – 4.89 (4.50), EW / PW 1.52 – 1.74 (1.59). Females (n = 10) TL 2.17 – 2.50 mm (2.37 mm), PL 0.37 – 0.43 mm (0.41 mm), PW 0.67 – 0.77 mm (0.72 mm), EL 1.80 – 2.00 mm (1.94 mm), EW 0.97 – 1.13 mm (1.05 mm), TL / EW 2.12 – 2.40 (2.26), PW / PL 1.62 – 1.92 (1.76), EL / EW 1.71 – 2.00 (1.86), EL / PL 4.38 – 5.00 (4.75), EW / PW 1.43 – 1.52 (1.45).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	distribution	Distribution. The species was described from Mount Manaia on the Whangerei Heads peninsula. It is widely distributed in northern part of North Island, mostly in Northland and Auckland areas, sometimes also in Bay of Plenty and Coromandel (Fig. 18 B).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF9702234DE7FE69FE7BFC2B.taxon	discussion	Remarks. Broun (1910) correctly noted that Atopida grahami is similar to A. hirta but differs in the sculpture of the thorax. What he did not realised is the fact that this difference was due to the sexual dimorphism, and A. grahami is in fact a female of A. hirta.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	description	(Figs 1 G – I, 5 E – F, 6 B, 8 C – D, 10 C, 12 C, 17 H)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	materials_examined	Type material. Holotype of A. impressa (NHM): card-mounted, “ 3418. ” [in Broun’s hand], “ Seaward Moss \ 30 / 9 / 10. ” [handwritten], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Atopida \ impressa ” [in Broun’s hand]. Holotype of A. pallidula (NHM): card-mounted, “ 4062. ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Bluff \ 1.12.1914 ” [in Broun’s hand], “ Atopida \ pallidula ” [in Broun’s hand]. Other material examined. 3 exx. (NZAC): “ NEW ZEALAND SL \ Slopedown, 580 m \ Mokoreta No 2 \ 23 Nov \ 12 Dec 1985 ”, “ B. I. P. Baratt \ Pit trap in \ shrubland ”; 12 exx. (NZAC): “ Rakeahua \ Valley ”, “ 19. ii. 68 \ G. Kuschel ”, “ Stewart I. \ Exp. Feb. 1968 ”; 2 exx. (NZAC): “ Rakeahua V. \ to m. v. light ”, “ 10. ii. 68 \ J. S. Dugdale ”, “ Stewart I. \ Exp. Feb. 1968 ”; 1 ex. (NZAC): “ Rakeahua \ Valley ”, “ South bank \ bush ”, “ 12 Feb 68 \ J. S. Dugdale ”; 1 ex. (NZAC): “ Mt. Rakeahua \ 1800 - 2100 ’ ”, “ 11. ii. 68 \ J. S. Dugdale ”; 4 exx. (NZAC): “ Mt. Rakeahua \ 1000 ’ - 2000 ’ ”, “ 10. ii. 68 \ G. Kuschel ”, “ Stewart I. \ Exp. Feb. 1968 ”; 1 ex. (NZAC): “ Mt. Rakeahua \ Stewart I 2000 ’ ”, “ 14. ii. 68 \ G. Kuschel ”, “ Stewart I. \ Exp. Feb. 1968 ”; 1 ex. (LUNZ): “ NEW ZEALAND, SI \ STEWART ISLAND \ Mt. Rakeahua \ 7. II. 1991 \ R. R. Scott ”, “ swept from Blechnum \ discolor ”; 4 exx. (LUNZ): “ NEW ZEALAND, SI \ STEWART ISLAND \ Christmas Village \ 6 - 7. ii. 1991 \ J. W. Early ”, “ swept in podocarp / \ hardwood forest ”; 1 ex. (LUNZ): “ Swept near hut \ Freshwater Creek Area \ Stewart Island, N. Z. \ 20. II. 1970 \ Ento. Dept. Field Trip ”; 1 male (AMNZ): “ Haldane \ Southland \ 24 - I- 47 C. E. C. [on the underside of the cardboard with the beetle] ”, “ Stewart I. \ Exp. Feb. 1968 ”, “ C. E. Clarke \ Collection ”; 5 exx. (NZAC): “ Port William \ Stewart I \ 7 Feb 69 \ A. C. Eyles ”, “ Stewart I. \ Exp. Feb. 1968 ”; 2 females (NZAC): “ Port \ William \ Stewart I ”, “ 4 Feb 69 \ B. A. Holloway ”, “ General \ Beating ”; 1 male (NZAC): “ Port William \ Stewart I \ 7 Feb 69 \ A. C. Eyles ”; 1 female (NZAC): “ Port William \ Stewart I. \ 7 Feb 69. \ B. A. Holloway ”, “ General \ beating ”; 2 females (NZAC): “ Collector \ Jan. 1943. \ A. E. Brookes ”, “ Stewart \ Island. N. Z. ”, “ A. E. Brookes \ Collection ”; 1 ex. (LUNZ): “ NEW ZEALAND, SI \ STEWART ISLAND \ Table Hill track \ 8. II. 1991 S. P., Womer \ under logs ”; 1 ex. (NZAC): “ Table Hill \ 1000 ’ 16. ii. 68 ”, “ Stewart I. \ Exp. Feb. 1968 ”, “ Beating ”; 2 males (AMNZ): “ Table Hill \ Stewart Is. \ 6 - 2 - 47 \ C. E. Clarke [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”; 1 female (NZAC): “ Big S. Cape I \ (NE) Feb 69 \ SW Stewart I \ B. A. Holloway ”, “ General \ Beating ”; 1 ex. (NZAC): “ Paterson \ Inlet ”, “ Little Glory \ Bay ”, “ Stewart I. \ 6 Feb 69 \ B. A. Holloway ”; 1 ex. (NZAC): “ Tisbury \ 9 / 10 / 10 ”; 1 ex. (NZAC): “ Wallacetown \ 18. II. 09 ”; 10 exx. (NZAC): “ Sealers Bay \ Codfish Is. ”, “ 12. xii. 66 \ J. I. Townsend ”, “ Weinmannia \ flowers ”; 1 female (NZAC): “ Sealers Bay \ Codfish Is. ”, “ 15. xii. 66 \ J. I. Townsend ”; 1 male (NZAC): “ NEW ZEALAND SI \ Stewart I \ Codfish I \ Sealers Bay \ Loop Track ”, “ 7 Dec 1981 \ B. A. Holloway \ Litter 81 / 209 ”; 2 males (NZAC): “ NEW ZEALAND SI \ Stewart I \ Codfish I \ Summit Tk, 250 m \ 28 Nov 1981 ”, “ B. A. Holloway \ on Gahnia ”; 1 male (NZAC): “ NEW ZEALAND SI \ Stewart I \ Codfish I \ Valley Track \ 24 - 30 Nov 1981 ”, “ B. A. Holloway \ on Gahnia ”; 1 male (NZAC): “ NEW ZEALAND SI \ Stewart I \ Codfish I \ Summit Tk, c 250 m \ 30 Nov 1981 ”, “ B. A. Holloway \ Sifted litter \ 81 / 199 ”; 1 female, 1 male (NZAC): “ Mt. Summit \ Codfish Isl. ”, “ 11. xii. 66 \ J. I. Townsend ”, “ On ferns \ at night ”; 1 female (NZAC): “ NEW ZEALAND SI \ Stewart I \ Little River \ West side \ 4 - 9 Jan 1976 ”, “ A. C. Harris \ Malaise trap ”; 2 males (NZAC): “ NEW ZEALAND SI \ Stewart I, W side \ Little River \ 26 Dec 1975 - \ 2 Jan 1976 ”, “ A. C. Harris \ Malaise trap ”; 1 female (NZAC): “ NEW ZEALAND SI \ Harry West Bay \ 24 Jan 2007 R. Leschen \ T Buckley K. Marske \ ex lichens ”, “ RL 1208 \ 46 55.917 S \ 168 1.032 E ”; 1 female (NZAC): “ Paterson \ Inlet ”, “ Little Glory \ Bay ”, “ Stewart I. \ 6 Feb 69 \ B. A. Kuschel ”; 1 female (NZAC): “ Pegasus \ 23. ii. 68 \ G. Kuschel. \ Ex. soil. ”, “ Stewart I. \ Exp. Feb. 1968 ”; 1 female (NZAC): “ Pegasus Ck. \ 24. ii. 68 \ G. Kuschel ”, “ Stewart I. \ Exp. Feb. 1968 ”, “ litter ”; 1 female (NZAC): “ Twilight Bay \ Port Pegasus ”, “ 22. ii. 68 \ G. Kuschel ”, “ Stewart I. \ Exp. Feb. 1968 ”, “ moss ”; 2 males (NZAC): “ New Zealand SI \ Golden Bay \ Stewart Island \ 10 - Jan- 96 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 2 males (NZAC): “ New Zealand SI \ Halfmoon Bay \ Stewart Island \ 11 - Jan- 96 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 3 males (NZAC): “ NEW ZEALAND SL \ Slopedown, 580 m \ Mokoreta No 2 \ 23 Nov- \ 12 Dec 1985 ”, “ B. I. P. Barratt \ Pit trap in \ shrubland ”, “ Duplicate \ specimens \ in alcohol ”; 4 exx. (NMPC): “ NEW ZEALAND: Stewart Island 0.65 \ km NNW of Fred’s Camp Hut \ 46 ° 55.51 ’ E 167 ° 58.32 ’ E; 70 m \ 18 - 22. i. 2016; Seidel, Sýkora & \ Fikáček lgt. 2016 - NZ 010 ”, “ flight intercept trap in sparse \ hardwood-podocarp forest \ numerous ferns in understory ”; 2 exx. (NMPC): “ NEW ZEALAND: Stewart Island 0.5 \ km N of Fred’s Camp Hut \ 46 ° 55.54 ’ E 167 ° 58.49 ’ E; 30 m \ 18 - 22. i. 2016; Seidel, Sýkora & \ Fikáček lgt. 2016 - NZ 009 ”, “ flight intercept trap in mossy \ hardwood forest with numerous \ fallen / decaying trunks ”; 26 exx. (NMPC): “ NEW ZEALAND: Stewart Island \ end of Fern Gully Tk. W of Oban, \ 17. - 21. i. 2016; 46 ° 53.34 ’ S \ 168 ° 5.51 ’ E; 65 m; Seidel, Sýkora \ & Fikáček lgt. 2016 - NZ 005 ”, “ baited pitfall traps (rotten squid) in \ dense young forest with numerous \ ferns in understory ”; 1 ex. (NMPC): “ NEW ZEALAND: Steward Island \ Northwest Circuit Tk. 3.2 km W of \ Oban, 17. - 21. i. 2016; \ 46 ° 53.80 ’ S; 168 ° 5.18 ’ E; 95 m \ Seidel & Fikáček lgt. 2016 - NZ 006 ”, “ sifting of leaf litter in hardwood- \ podocarp forest with sparse ferns \ and many mosses ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	diagnosis	Diagnosis. Body of moderate size (TL 3.8 – 4.6 mm), setation short, adherent, punctation of pronotum not granulate, females apterous. Male genitalia with wide, rounded apical portion of trigonium. Tergite VII in females with oval depression; this structure is unknown in any other Atopida species. Bursal sclerite with paired rod-like sclerites and additional small sclerite between the rods. A. suffusa is the only species with similar aedeagus and bursal sclerite but its dorsum is covered with long, erect setae.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	description	Redescription. Male. Body elongated, moderately large (TL 3.9 – 4.6 mm), convex. Body yellowish-brown to dark brown. Punctation on head subtly granulate, punctures touching each other, on pronotum not granulate in central portion of the disc, sometimes granulate on sides, but granules flat and sparse, punctures separated by 0.3 – 1.0 × diameter of a puncture, punctation on elytra simple, punctures separated by 0.5 – 0.7 × diameter of a puncture. Body covered with very short, yellowish, strongly adherent setae. Supraantennal ridges well-marked, slightly elevated. Pronotum widest slightly before middle of its length, sides curved, anterolateral angles slightly produced, posterolateral angles obtuse, rounded. Elytra in some specimens with two faint longitudinal carinae, humeri wellmarked. Ventrite 5 rounded, apex truncate. Penis elongate (L 0.59 mm, W 0.25 mm), with very wide trigonium, rounded in apical portion, a little shorter than parameroids, covered with very short setae; parameroids widely triangular, axe-shaped, apices rounded, apical part covered with punctures; pala triangular, slightly longer than parameroids. Tegmen (L 0.57 mm, W 0.39 mm) relatively wide, with triangular parameres, covered with numerous minute setae in apical portion. Female. Body medium sized (TL 3.8 – 4.4 mm). Supraantennal ridges well-marked, only slightly elevated. Punctation on disc of pronotum sparser than in males, punctation on elytra coarser, punctures separated by 0.5 × diameter of a puncture. Ventrite 5 rounded, with deep, semicircular emargination at apex, tergite VII with oval depression (Fig. 8 D) in apical portion. Ventrite 4 with two large, oblique setal patches. Bursal sclerite consisting of two lateral rods (L 0.40 mm) widened in posterior portion and armed with denticles on outer parts of anterior portion, additional small, v-shaped sclerite present between rods. Measurements and ratios. Males (n = 10) TL 3.85 – 4.60 mm (4.10 mm), PL 0.70 – 0.85 mm (0.74 mm), PW 1.10 – 1.35 mm (1.23 mm), EL 3.10 – 3.70 mm (3.35 mm), EW 1.75 – 2.15 mm (1.93 mm), TL / EW 2.02 – 2.23 (2.13), PW / PL 1.53 – 1.79 (1.66), EL / EW 1.64 – 1.82 (1.73), EL / PL 4.31 – 4.93 (4.53), EW / PW 1.52 – 1.68 (1.57). Females (n = 10) TL 3.80 – 4.35 mm (4.06 mm), PL 0.70 – 0.85 mm (0.76 mm), PW 1.35 – 1.50 mm (1.42 mm), EL 3.15 – 3.55 mm (3.30 mm), EW 1.85 – 2.15 mm (2.02 mm), TL / EW 1.88 – 2.08 (2.01), PW / PL 1.71 – 2.00 (1.87), EL / EW 1.56 – 1.71 (1.64), EL / PL 3.94 – 4.73 (4.34), EW / PW 1.37 – 1.52 (1.42).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	distribution	Distribution. Very common in southern part of South Island, especially on Stewart Island, present also in Southland area (Fig. 18 C).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8802214DE7FC61FD91FDCB.taxon	discussion	Remarks. The species is very distinct in terms of morphology, and we agree with Broun’s (1914) conclusion that “ there is no similar species ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	description	(Figs 1 J – K, 3 C – D, 5 G – H, 8 E, 9 A, 10 D, 12 D, 17 O)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	materials_examined	Type material. Holotype, male (NZAC): “ New Zealand FD \ South Arm \ L Manapouri \ 14 - Dec- 98 ”, “ General \ beating ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; paratypes, 5 exx. (NZAC): same data as holotype; paratype, 1 ex. (LUNZ): “ NEW ZEALAND FD \ Fiordland Nat Pk \ Murchison Mts \ Plateau Ck \ 990 m ”, “ 3. XII. 1980 \ R. M. Emberson \ C. A. Muir ”, “ Malaise trap \ N. menziesii \ forest edge ”; paratypes, 7 exx. (NZAC): “ Upper Holly- \ ford Val. \ Homer ”, “ 12. xii. 66 \ A. K. Walker ”, “ Olearia \ ilicifolia ”; paratype, 1 ex. (NZAC): “ Hidden falls \ Tr. 14. xii. 66 ”, “ Lower \ Hollyford Val. \ A. K. Walker ”, “ General \ beating ”; paratype, 1 ex. (NZAC): “ Hollyford Val. \ 13. I. 67 \ A. K. Walker ”, “ at night \ off ferns ”; paratypes, 2 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ Gut hut \ 22 - 28 Nov 1981 ”, “ C. F. Butcher \ Malaise trap ”; paratypes, 2 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 600 m \ 30 Nov 1981 ”, “ C. F. Butcher \ sweeping \ at night ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 29 Nov 1981 \ C. F. Butcher ”, “ ex pan trap \ in tussock ”; paratypes, 13 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850 m \ 30 Nov 1981 \ C. F. Butcher ”, “ sweeping \ at night ”; paratypes, 12 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850 m \ 30 Nov 1981 \ C. F. Butcher ”, “ sweeping \ tussock ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850 m \ 30 Nov 1981 \ C. F. Butcher ”, “ Insect spray \ rock face ”; paratypes, 4 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono \ 27 Nov 1981 ”, “ C. F. Butcher ”; paratypes, 3 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono \ 27 - 30 Nov 1981 ”, “ C. F. Butcher \ Pan trap ”; paratypes, 4 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono, 853 m \ 27 Nov 1981 ”, “ C. F. Butcher \ Mats and mosses \ 81 / 189 ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono, 853 m \ 30 Nov 1981 ”, “ C. F. Butcher \ mats, moss and \ tussock 81 / 186 ”; paratype, 1 ex. (NZAC): “ NEW ZEALAN FD \ Camp 1020 m \ Tutoko Bench \ Darran Mts ”, “ 13 Jan 1977 \ T. K. Crosby and \ J. S. Dugdale \ Malaise trap ”; paratype, 1 ex. (NZAC): “ Alpine \ Garden ”, “ Homer Tunnel \ to Milford Rd. ”, “ 12. xii. 66 \ A. K. Walker ”, “ General beating ”; paratype, 1 ex. (NZAC): “ Bold Peak, \ 27 - 12 to 1 - 1 - 45 \ E. S. Gourlay ”, “ Atopida \ lawsoni Shp. \ Id. by J. Armstrong ” paratype, 1 ex. (NZAC): “ Fox Glacier \ 14.12.60 \ S. Westland, ”, “ J. I. Townsend \ & P. R. Kettle ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND WD \ Mt Tuhua, 1067 m \ E. side of \ L. Kaniere \ 20 Nov 1984 ”, “ B. A. Holloway \ Moss and \ lichen 84 / 75 ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Bauza I \ Mar 1984 \ C. F. Butcher ”; paratype, 1 ex. (NZAC): “ Flat Top Mt. Hunter \ Mts. L. Manapouri \ 4000 ’ Clark. ”, “ A. E. Brookes \ Collection ”; paratype, 1 ex. (NZAC): “ S of Cascade \ Eglinton R \ 30. X. 66 J. I. T ”, “ moss under \ beech trees ”; paratype, 1 ex. (NZAC): “ New Zealand OL \ S. Mavora Lake \ East side \ 16 - xii- 2013 ”, “ Sifted \ ground moss \ Nothofagus \ forest ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; paratype, 1 ex. (NZAC): “ New Zealand SL \ Princhester Base \ Takitimu Forest \ 10 - Dec- 04 ”, “ On low \ vegetation by \ stream ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; paratype, 1 ex. (NZAC): “ 13 - I- 66 \ Takitimus 2500 ’ \ A. C. Eyles ”, “ Celmisia \ (sticky) ”; paratype, 1 ex. (AMNZ): “ Routeburn \ R. \ 4 - I- 45 \ C. E. Clarke ”, “ C. E. Clarke \ Collection ”; paratype, 1 ex. (AMNZ): “ Upper \ Routeburn R. \ 18 - I- 26 \ C. E. Clarke ”; paratypes, 2 exx. (NMPC): “ NEW ZEALAND: Fiordland (FD) \ Borland Road 24 km NWW of \ Monowal, 45 ° 41.13 ’ S \ 167 ° 20.29 ’ E; 320 m; 24 - 27. i. 2016; \ Seidel, Sykora & Fikáček lgt. \ 2016 - NZ 024 ”, “ sifting of leaf litter, moss and \ rotten logs in degraded remnants \ of Nothofagus forest with \ numerous rotten logs ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	diagnosis	Diagnosis. Body small to moderately large (TL 3.3 – 4.5 mm), colouration variable. Similar to some other Atopida from South Island, certain identification is possible only on the basis of genital structures: trigonium much shorter than parameroids, curved laterally, parameroids triangularly pointed at apices, bursal sclerite subtriangular in outline, with paired tubercles in central portion.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	description	Description. Male. Body elongated, moderately large (TL 3.7 – 4.5 mm), moderately convex. Body colouration variable, from yellowish to brown, head and pronotum slightly darker, elytra yellowish with dark suture or uniformly yellowish to dark brown. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on pronotum becoming more sparse towards the middle and base, surface of pronotum uneven, in some specimens forming two tubercles in basal portion of pronotum, punctation on elytra simple, coarse, punctures separated by 0.3 – 0.7 × diameter of a puncture. Body covered with moderately long, protruding, yellowish setae, more numerous and ruffled on head and pronotum. Supraantennal ridges well-marked, elevated, forming triangular flap. Pronotum widest slightly before middle of its length, sides curved, lateral carinae crenulate, anterolateral angles produced, triangular, pointed, posterolateral angles right-angled. Base of pronotum with longitudinal depression along posterior margin. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex subtly truncate. Penis (L 0.70 mm, W 0.36 mm) elongate, with explanate lateral lobes, trigonium very short, narrow, curved laterally at ~ right angle; parameroids elongated, sinuate, outer edges subtly denticulate, apices pointed, covered with punctures; pala triangular, very short, significantly shorter than parameroids. Tegmen (L 0.67 mm, W 0.46 mm) with narrow, triangular parameres, covered with scarce punctures and numerous denticles. Female. Body small to moderately big (TL 3.3 – 4.1 mm), brachypterous. Supraantennal ridges well-marked, elevated, forming triangular flaps in some specimens. Colouration and punctation like in males, but central portion of pronotal disc sparsely punctate, with shining interspaces. Ventrite 5 rounded, with very shallow emargination at apex. Lateral portions of ventrites 3 and 4 rather big, oblique setal patches, small oblique patches visible also on ventrite 5. Bursal sclerite big (L 0.38 mm, W 0.32 mm) consisting of two interconnected plates — one with pair of tubercles visible in central portion of sclerite, the second forming hook-like processes resembling long spines in dorsal view. Measurements and ratios. Males (n = 10) TL 3.65 – 4.45 mm (4.05 mm), PL 0.65 – 0.75 mm (0.69 mm), PW 1.10 – 1.30 mm (1.19 mm), EL 3.00 – 3.70 mm (3.36 mm), EW 1.70 – 2.00 mm (1.89 mm), TL / EW 1.92 – 2.25 (2.14), PW / PL 1.57 – 1.86 (1.73), EL / EW 1.59 – 1.89 (1.78), EL / PL 4.33 – 5.46 (4.88), EW / PW 1.50 – 1.68 (1.59). Females (n = 10) TL 3.30 – 4.05 mm (3.82 mm), PL 0.60 – 0.75 mm (0.67 mm), PW 1.10 – 1.40 mm (1.24 mm), EL 2.75 – 3.30 mm (3.14 mm), EW 1.70 – 2.05 mm (1.83 mm), TL / EW 1.94 – 2.31 (2.09), PW / PL 1.60 – 2.08 (1.86), EL / EW 1.61 – 1.89 (1.71), EL / PL 4.40 – 5.25 (4.72), EW / PW 1.40 – 1.58 (1.48).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	distribution	Distribution. The species is distributed mostly in south-west areas of South Island — Fiordland and Otago Lakes, rarely occurs also in Westland area (Fig. 18 D).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8A02204DE7FD01FAE7FAFF.taxon	etymology	Etymology. Latin insularis = insular, an allusion to the fact that the species is common on Secretary Island.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	description	(Figs 1 L – M, 3 E, 5 G – H, 13 A, 17 L)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	materials_examined	Type material. Holotype, 1 ex. (NZAC): “ Rapahoe \ 3.11.40 \ E. S. Gourlay ”, “ var. ”, “ E. S. Gourlay \ Dec. 1970 \ Ent. Div. ”; paratype, 1 ex. (NZAC): same data as holotype; paratype, 1 ex. (NZAC): “ SWEPT BEECH FOREST \ AND STREAM \ BY COBB RESERVOIR \ 18 DEC. 1967 \ B. M. MAY ”; paratype, 1 ex. (NZAC): “ Mt Peel. 1200 m \ Cobb V. Nelson \ Dec 67 \ S Bridge ”, “ moss ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	diagnosis	Diagnosis. Externally resembling other species with crenulate lateral pronotal carinae that inhabit South Island, like A. insularis sp. nov., A. tuhua sp. nov. etc. Certain identification possible only after examination of male genitalia: trigonium as long as parameroids, triangular, pointed at apex, parameres with sharp denticles on inner margins.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	description	Description. Male. Body elongated, moderate size (TL 4.1 – 4.4 mm), moderately convex, surface of head and pronotum uneven. Body yellowish-brown to almost black. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other, with the exception of central portion of disc, where punctures are more sparse, with shining interspaces; punctation on elytra simple, punctures separated by 0.3 – 0.5 × diameter of a puncture. Body covered with moderately long, erect setae. Supraantennal ridges projecting to form long, triangular outgrowths (Fig. 3 E). Sides of pronotum subparallel to slightly rounded, widest in half of its length, sides crenulate, anterolateral angles distinctly produced, posterolateral angles pointed. Elytra without carinae, humeri well-marked. Ventrite 5 subtriangular, with rounded apex. Penis (L 0.51 mm, W 0.26 mm) moderately elongated, with explanate lateral lobes, trigonium rather narrow, triangular, pointed at apex, as long as parameroids; parameroids elongated, apices pointing outwards, covered with numerous punctures, rounded at apices; pala triangular, a bit longer than parameroids. Tegmen (L 0.45 mm, W 0.34 mm) wide, rectangular, parameres triangular, with scarce punctures and sharp denticles on inner edges and on apices. Female unknown. Measurements and ratios. Males (n = 4) TL 4.10 – 4.35 mm (4.18 mm), PL 0.70 – 0.80 mm (0.73 mm), PW 1.15 – 1.30 mm (1.25 mm), EL 3.35 – 3.55 mm (3.45 mm), EW 2.00 – 2.05 mm (2.03 mm), TL / EW 2.02 – 2.12 (2.06), PW / PL 1.63 – 1.86 (1.73), EL / EW 1.68 – 1.73 (1.70), EL / PL 4.44 – 4.93 (4.77), EW / PW 1.54 – 1.74 (1.62).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	distribution	Distribution. The species is known from localities in Nelson and Buller areas of South Island (Fig. 18 E).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	etymology	Etymology. Dedicated to Richard A. B. Leschen, one of the leading contemporary coleopterists, who encouraged us to study New Zealand marsh beetles and helped during our New Zealand trip in 2016.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8B02274DE7FA15FCD1FD5B.taxon	discussion	Remarks. Due to low number of known specimens, little is known on the variability of the species, which nevertheless seems to be significant, at least in colouration. Specimens from the area of Cobb Reservoir are yellowish, while those collected in Rapahoe are almost black.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	description	(Figs 1 N – O, 5 J – K, 8 F, 10 E, 13 B, 14, 17 E)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	materials_examined	Type material. Two syntypes, males (NHM): card-mounted, “ Mt Owen \ 27 - 12 - 1914 ” [in Broun’s hand], “ Atopida \ montana ” [in Broun’s hand], “ 4063. ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ” Other material examined. 1 female (NHM): “ var. \ 4063 ”, “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Glen Hope \ 14.3.1915 ” [in Broun’s hand]; 1 ex. (NZAC): “ Dun Track \ 14.2.42 \ E. S. Gourlay ”; 2 exx. (LUNZ): “ NEW ZEALAND BR \ Mt Robert Carpark \ St Arnaud \ 11 Jan 2014 \ SDJ Brown ”, “ Extracted from beech \ litter \ 880 m \ 41.8219 ° S 172 8071 ° E ”; 2 exx. (LUNZ): “ NEW ZEALAND, BR \ Lake Rotoroa \ nr camp ground \ 10. I. 1993 \ J. W. M. Marris ”, “ beaten from foliage, \ Nothofagus / Podocarpus \ forest ”; 1 male (NZAC): “ L. Rotoroa \ 1 - 3 - II- 39 \ E. S. Gourlay ”; 1 ex. (LUNZ): “ NEW ZEALAND, NN \ nr Takaka, The \ Grove Scen Res, \ Nov 1985 \ J. W. Early ”, “ yellow pan \ trap in \ bush ”; 1 ex. (NZAC): “ Canan \ W Nelson \ South Is. ”, “ Collector \ 10 - 13 I. 1949 \ A. E. Brookes ”; 1 ex. (NZAC): “ Pigeon Saddle \ Takaka I. […] \ Nelson ”, “ A. E. Brookes \ Collection ”; 1 ex. (NZAC): “ Owen River \ NN \ E. S. Gourlay ”; 1 ex. (NZAC): “ Dun Mt. Summit \ Nel. 19. xii. 68 \ J. C. Watt ”; 1 male (NZAC): “ Dun Mt. 3000 ’ \ 7. I. 34 \ E. S. Gourlay ”; 1 male (NZAC): “ Dun Mt. 2000 ’ \ 7.2.33 \ E. S. Gourlay ”; 1 male (NZAC): “ Dun Mt. 2000 ft. \ 15.1.28. \ E. S. Gourlay ”; 1 male (NZAC): “ Third. House \ Dunn Track. \ 10 / 1 / 42 ”, “ Atopida \ montana Broun \ Det. E. S. Gourlay ”; 1 male (NZAC): “ Third House \ Dunn Track \ A. Hamilton \ 10 / 1 / 42 ”; 1 male (NZAC): “ Third House \ Dun Mt. Track \ 2000 ’. 30.12.53 \ E. S. Gourlay ”; 1 male (NZAC): “ Third House \ Dun Mt. Track \ 3000 ’. 30.12.53 \ E. S. Gourlay ”; 1 ex. (NZAC): “ Bealy Vall. \ 9 - 11 - I- 57 \ E. S. Gourlay ”; 1 ex. (NZAC): “ Fox Glacier \ 14.12.60 \ S. Westland ”, “ J. I. Townsend \ & P. R. Kettle ”; 1 ex. (NMPC): “ NEW ZEALAND: Marlborough \ Pelorus Bridge, 30. xi. 2010, \ 41 ° 30.490 ’ S 173 ° 56.747 ’ E \ night collecting on trunks, rotten \ wood and fungi \ Fikacek & Leschen RL 1515 ”; 2 males (NZAC): “ Canaan Nelson \ 17 - 26.1.49 ”, “ A. E. Brookes \ Collection ”; 1 male (NZAC): “ NEW ZEALAND, BR \ Mt Robert \ 5 Feb 1982 \ C. F. Butcher ”, “ Sweeping ”; 1 male (NZAC): “ mt. Owen \ 27.12.1914. ”, “ Atopida \ montana ”, “ T. Broun \ Collection ”, “ A. E. Brookes \ Collection ”; 1 male (NZAC): “ Opouri \ Nelson \ 10 Jan 69 \ J. S. Dugdale ”; 1 male (NZAC): “ Pittosporum \ tenuifolium ”, “ Whangamoa \ 700 ’ 29 - 11 - 88 \ G. Kuschel ”; 1 male (NZAC): “ Takaka Valley \ Nelson \ 26.11.63 ”; 1 male (NZAC): “ Canaan Nel. \ 17 - 26.1.49 ”, “ A. E. Brookes \ Collection ”; 1 male (NZAC): “ Canaan Nel. \ Dec. 1949 \ O’Connor ”, “ A. E. Brookes \ Collection ”; 1 male (NZAC): “ Puramahoi \ Collingwood \ 26.10.65 \ J. I. Townsend ”; 1 male (NZAC): “ Puramahoi \ Takaka Dist. Nel. \ 26.10.65 \ J. I. Townsend ”, “ 65 / 524 ”, “ litter ”; 1 male (NHM): “ Nothofagus \ forest litter ”, “ BR: L. Rotoiti \ St. Arnaud Ra \ 1550 m. \ 25. xii. 1983 ”, “ NEW ZEALAND \ B. M. 1984 - 80 \ P. M. Hammond ”; 1 male (AMNZ): “ NEW ZEALAND BR \ L. Rotoiti \ beech forest \ emergence trap \ 12 - 20. xii. 2003 \ D. Hartnett ”; 1 male (AMNZ): “ Whangapeka Vall \ 2 - 12 - 34 \ E. S. Gourlay ”, “ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”, “ Atopida \ montana ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	diagnosis	Diagnosis. Body small to moderately big (TL 3.3 – 4.3 mm), narrow, covered with erect setae; pronotum densely covered with granulate punctures. Trigonium narrow, rounded at apex, slightly shorter than parameroids. Bursal sclerite consisting of two long rods, setal patches present on ventrites 3 – 5 of females small, rounded; apical portion of ventrite 5 with 4 tubercles. Externally resembling A. hirta from North Island, distinguishable on the basis of morphology of genitalia. Probably a species complex, see remarks below.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	description	Redescription. Male. Body elongated, small to moderately large (TL 3.3 – 4.3 mm), moderately convex. Body yellowish brown, head and pronotum slightly darker, often with black-brown stripe along elytral suture. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on elytra simple, coarse, punctures separated by 0.3 – 0.5 × diameter of a puncture. Body covered with rather long, yellowish, protruding, ruffled setae. Supraantennal ridges well-marked, slightly elevated. Pronotum widest in the anterior portion, slightly before middle of its length, sides curved, anterolateral angles slightly produced, posterolateral angles right-angled. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex triangular. Penis (L 0.45 mm, W 0.19 mm) elongated, trigonium rather narrow, sides subparallel, apex rounded, slightly shorter than parameroids; parameroids elongated, apices obliquely truncate, apical part covered with punctures; pala rectangular, slightly longer than parameroids. Tegmen (L 0.43 mm, W 0.26 mm) with wide rectangular parameres, rounded apices, scarce punctures and numerous rows of delicate setae in apical part. Female. Body moderately big (TL 3.6 mm), brachypterous, coloration yellowish-brown, elytra with darkened suture. Supraantennal ridges marked, not elevated. Basal portion of pronotal disc with impunctate, triangular area. Ventrite 5 rounded, with 4 small tubercles in apical portion. Lateral portions of ventrites 3 and 4 with small, oval setal patches, small patch-like area without distinct setation present also on ventrite 5. Bursal sclerite (L 0.2 mm) consisting of two elongated rods, triangularly widened in middle of their length, truncate posteriorly and pointed in anterior portion. Measurements and ratios. Males (n = 9) TL 3.30 – 4.30 mm (3.74 mm), PL 0.55 – 0.80 mm (0.67 mm), PW 1.00 – 1.35 mm (1.13 mm), EL 2.70 – 3.50 mm (3.05 mm), EW 1.55 – 2.05 mm (1.71 mm), TL / EW 2.10 – 2.29 (2.19), PW / PL 1.50 – 1.93 (1.71), EL / EW 1.71 – 1.83 (1.79), EL / PL 4.13 – 5.00 (4.60), EW / PW 1.42 – 1.62 (1.51). Female (n = 1) TL 3.55 mm, PL 0.70 mm, PW 1.15 mm, EL 2.80 mm, EW 1.75 mm, TL / EW 2.0, PW / PL 1.6, EL / EW 1.6, EL / PL 4.0, EW / PW 1.5.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	distribution	Distribution. Distributed in several areas of South Island: Marlborough Sounds, Nelson, Buller, and Westland (Fig. 18 F).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8D02244DE7FF24FEA1FD67.taxon	discussion	Remarks. While all males examined are very uniform in terms of morphology, it is not the case of females. The description of female is based on a specimen from Glenhope, identified by T. Broun. Its punctation on pronotum is very similar to that of the holotype. Among numerous female specimens that we initially identified as A. montana, several striking differences were noticed after a closer examination, including level of wing reduction, differences in punctation of pronotal disc (almost smooth and impunctate in some specimens), and even more striking differences in arrangement of abdominal setal patches. A. montana is apparently a species complex that needs further studies in order to recognize particular species it includes. Currently we distinguished three additional female morphotypes, tentatively named after the localities of specimens: 1) ‘ Mt Robinson’ (Figs 14 A, D, G) — brachypterous; setal patches present on ventrite 4, elongated; bursal sclerite consisting of triangular sclerites (L 0.17 mm); 2) ‘ Port Underwood’ (Figs 14 B, E, H) — brachypterous; setal patches present on ventrite 4, very small, rounded; ventrite 5 with two pointed, triangular denticles on apex; bursal sclerite consisting of triangular sclerites (L 0.23 mm); 3) ‘ Pelorus Bridge’ (Figs 14 C, F, I) — apterous; abdomen without setal patches (!); bursal sclerite with two rods (L 0.21 mm).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	description	(Figs 1 P – Q, 2 A, 5 L – M, 7 C, 8 G, 10 F, 13 C, 17 M)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	materials_examined	Type material. Holotype, male (NZAC): “ Mt Domett \ NN. 1250 m \ Bush edge \ 3 Dec. 71 ”, “ J. S. Dugdale ”; paratypes, 5 exx. (NZAC): same data as holotype; paratypes, 4 exx. (NZAC): “ Lochnagar \ Ridge Lake \ 3050 ’ ”, “ Paparoa \ Range ”, “ 3 Dec 69 \ J. S. Dugdale ”, “ sweeping \ tussocks ”; paratype, 1 ex. (NZAC): “ Lochnagar \ Ridge ”, “ Paparoa \ Range ”, “ 3500 ’ - 3800 ’ \ 2 - 10 Dec 61 \ J. S. Dugdale ”; paratypes, 6 exx. (NZAC): “ Mt Domett \ NN. 1250 m \ G. Kuschel \ Nov-Dec 71 ”, “ Litter ”; paratype, 1 ex. (NZAC): “ Mt Domett \ 1066 - 1219 m \ NN Nov 71 \ G. W. Ramsay ”; paratypes, 2 exx. (NZAC): “ Mt Domett \ NN. 1463 m \ 1 Dec 71 \ J. S. Dugdale ”, “ moss ”; paratype, 1 ex. (NZAC): “ Mt Dewar 1060 m \ J. G. McBurney ”, “ Paparoa Range \ Dec. 1969 ”, “ Dracophyllum \ Beating \ longifolium ”; paratype, 1 ex. (NZAC): “ Mt Dewar 1100 m \ J. I. Townsend ”, “ Paparoa Range \ Dec. 1969 ”, “ 69 / 254 \ moss ”; paratype, 1 ex. (NZAC): “ Mt Dewar 1130 m \ J. C. Watt \ 8 Dec 69 ”, “ Paparoa Range \ Dec. 1969 ”, “ 69 / 254 \ moss ”; paratype, 1 ex. (NZAC): “ Mt. Dewar 1060 m \ J. I. Townsend ”, “ Paparoa Range \ Dec. 1969 ”, “ Swept at \ night ”; paratypes, 2 exx. (NZAC): “ Mt Dewar 1280 m \ J. C. Watt ”, “ 11 Dec 69 ”; paratypes, 2 exx. (NZAC): “ Canaan 2800 ’ \ Takaka Hill \ 4.12.64 litter \ G. Kuschel ”; paratype, 1 ex. (NZAC): “ Lake Paringa \ 6 - 10.12.60 \ S. Westland ”, “ J. I. Townsend \ & P. R. Kettle ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	diagnosis	Diagnosis. Body of moderate size (TL 3.8 – 4.7 mm), colouration variable, elytral interspaces shining. Certain identification possible only on the basis of genital characters: trigonium of penis triangularly pointed at apex, slightly shorter than parameroids, parameroids with triangular apices, basal portion of pala emarginated; central portion of bursal sclerite with single tubercle.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	description	Description. Male. Body elongated, moderately big (TL 4.2 – 4.7 mm), moderately convex. Coloration variable: from entirely yellowish brown to black, sometimes elytra yellowish with black suture. Punctation on head weakly granulate, punctures separated by ca. 0.3 × diameter of a puncture. Punctation on pronotum granulate on sides, punctures close to each other, punctures separated by ca. 0.5 × diameter of a puncture and simple in the middle, getting more sparse towards the end of pronotum, separated by 1.0 – 3.0 × diameters of a puncture. Punctation on elytra simple, coarse, punctures separated by 0.3 – 1.0 (rarely 2.0) × diameter of a puncture. Body very shiny, covered with two types of whitish, protruding setae: medium long, and visibly longer and even more protruding. Supraantennal ridges well-marked, elevated. Pronotum widest slightly before middle of its length, sides bisinuate, anterolateral angles produced, pointed triangularly, posterolateral angles right-angled; base of pronotum with longitudinal depression along posterior margin. Lateral carinae slightly crenulate. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex delicately truncate. Penis (L 0.76 mm, W 0.59 mm) relatively wide, with explanate lateral lobes, trigonium pointed at apex, triangular, slightly shorter than parameroids; parameroids elongated, apices with serrated edge, covered with punctures, apices triangular, pointed; pala very wide, triangular, about same length parameroids, with shallow emargination in basal portion. Tegmen (L 0.69 mm, W 0.52 mm) with wide triangular parameres, apices acute, with scarce punctures and setae. Female. Body of moderate size (TL 3.8 – 4.5 mm), brachypterous. Coloration and punctation like in males. Supraantennal ridges well-marked, elevated. Pronotum more transverse than in males. Ventrite 5 rounded, with small semicircular emargination at apex. Lateral portions of ventrites 3 and 4 with large, oblique setal patches. Bursal sclerite (L 0.20 mm, W 0.23 mm), consisting of interconnected plates — two lateral ones, wrapped around and single central plate with median tubercle. Measurements and ratios. Males (n = 7) TL 4.20 – 4.65 mm (4.44 mm), PL 0.65 – 0.80 mm (0.74 mm), PW 1.25 – 1.40 mm (1.31 mm), EL 3.50 – 3.90 mm (3.68 mm), EW 1.90 – 2.15 mm (1.99 mm), TL / EW 2.09 – 2.33 (2.24), PW / PL 1.67 – 1.92 (1.79), EL / EW 1.74 – 1.95 (1.85), EL / PL 4.67 – 5.62 (5.02), EW / PW 1.41 – 1.60 (1.51). Females (n = 5) TL 3.80 – 4.45 mm (4.15 mm), PL 0.60 – 0.70 mm (0.66 mm), PW 1.15 – 1.45 mm (1.35 mm), EL 3.25 – 3.70 mm (3.49 mm), EW 1.90 – 2.15 mm (2.04 mm), TL / EW 1.95 – 2.10 (2.03), PW / PL 1.92 – 2.15 (2.04), EL / EW 1.66 – 1.76 (1.71), EL / PL 5.14 – 5.42 (5.29), EW / PW 1.46 – 1.65 (1.52).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	distribution	Distribution. Recorded from scattered localities on west coast of South Island — Nelson, Buler, and Westland areas (Fig. 18 G).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8F022B4DE7FCADFB36FD2F.taxon	etymology	Etymology. After Paparoa Range, one of the areas where type specimens were collected.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	description	(Figs 2 B, 5 N, 13 D, 17 N)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	materials_examined	Type material. Holotype, male (NZAC): “ Lake Paringa \ 6 - 10.12.60 \ S. Westland ”, “ J. I. Townsend \ & P. R. Kettle ”; paratypes, 16 exx. (NZAC): same data as holotype.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	diagnosis	Diagnosis. Body small to moderately big (TL 3.4 – 4.2 mm), colouration variable, from yellowish brown to dark brown, elytral suture often darkened. Certain identification possible on the basis of male genital characters: lateral plates of penis with minute denticles on outer edge in apical portion, trigonium narrow, much narrower than parameroids, curved laterally; parameroids elongated, apices truncate, with serrated edges, tegmen subrectangular with narrow parameres.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	description	Description. Male. Body elongated, small to moderately big (TL 3.4 – 4.2 mm), moderately convex. Two types of body colouration — lighter and darker: yellowish brown with darker stripe along elytral suture, head and pronotum darker brown and uniformly dark brown body with lighter brown legs. Punctation on head and pronotum dense, granulate, punctures touching each other, sparser on tubercles in lower part of pronotum. Punctation on elytra simple, coarse, punctures separated by 0.5 × diameter of a puncture. Body covered with medium long, protruding, yellowish setae, pointed towards the middle of pronotum. Supraantennal ridges well-marked, elevated triangularly. Pronotum relatively narrow, widest slightly before middle of its length, sides slightly curved, lateral carinae crenulate, anterolateral angles produced, pointed triangularly, posterolateral angles right-angled; base of pronotum with longitudinal depression along posterior margin. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex truncate. Penis (L 0.65 mm, W 0.39 mm) elongate, with explanate lateral lobes, outer margin of apical portions of apical plates with minute denticles; trigonium narrow, curved laterally, usually at right angle, shorter than parameroids; parameroids elongated, apices truncate, with serrated edges, apical part covered with punctures; pala wide, triangular, much shorter than parameroids. Tegmen (L 0.67 mm, W 0.39 mm) subrectangular, with narrow rectangular parameres, apices truncate, with scarce punctures and few setae. Female unknown. Measurements and ratios. Males (n = 6) TL 3.40 – 4.20 mm (3.90 mm), PL 0.60 – 0.75 mm (0.68 mm), PW 0.95 – 1.15 mm (1.04 mm), EL 2.90 – 3.45 mm (3.28 mm), EW 1.65 – 1.95 mm (1.84 mm), TL / EW 2.06 – 2.15 (2.12), PW / PL 1.36 – 1.64 (1.53) EL / EW 1.74 – 1.84 (1.78), EL / PL 4.53 – 5.00 (4.81), EW / PW 1.68 – 2.05 (1.77).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	distribution	Distribution. Known only from the type locality in Westland area of South Island (Fig. 18 L).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF80022B4DE7FD65FC73F8AF.taxon	etymology	Etymology. After Lake Paringa area, where the type locality is located.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	description	(Figs 2 C – D, 5 O – P, 8 H – I, 9 A, 10 G, 15 A, 17 C)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	materials_examined	Type material. Syntype (NHM): card-mounted, “ Atopida proba \ Type D. S. \ New Zealand Broun ” [written at base of card in Sharp’s hand], “ Type \ H. T. ” [round label with red border], “ New Zealand. [red underline] ”, “ Sharp Coll. \ 1905 - 313. ” Syntype (NHM): card-mounted, “ Atopida proba \ Ind. typ. D. S. \ New Zealand Broun ” [written at base of card in Sharp’s hand], “ New Zealand. [red underline] ”, “ Sharp Coll. \ 1905 - 313. ” Syntype (NHM): cardmounted, “ Atopida proba \ Ind. Type D. S. \ New Zealand Broun ” [written at base of card in Sharp’s hand], “ New Zealand. [red underline] ”, “ Sharp Coll. \ 1905 - 313. ” Holotype of Atopida sinuata (NHM): card-mounted, “ 2524. ” [in Broun’s hand], “ Howick ”, “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Atopida \ sinuata ” [in Broun’s hand]. Syntypes of Atopida basalis (NHM): 2 exx., card-mounted, “ 3244 ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Kaitoke. \ Wellington ” [in Broun’s hand], “ Atopida \ basalis ” [in Broun’s hand]. Other material examined. 1 ex. (NZAC): “ NEW ZEALAND CL \ Thames, Kirikiri \ Saddle, 400 m \ 12 Feb 1979 \ J. S. Dugdale ”, “ to light ”; 2 exx. (NZAC): “ NEW ZEALAND CL \ Thames, 400 m \ Kirikiri Sdle \ 12 Feb 1979 \ J. S. Dugdale ”, “ ex sprayed logs \ with polypores ”; 3 males (NZAC): “ NEW ZEALAND CL \ Thames, Kirikiri \ Saddle, 450 m \ 12 Feb 1979 \ J. S. Dugdale ”, “ to light ”; 1 ex. (NZAC): “ Mt. Hobson \ 304 m. \ Gt. Barrier \ I. 27 Nov 57. ”, “ J. C. Watt ”, “ Lepto- \ spermum \ scoparium \ ericoides ”; 1 ex. (NZAC): “ Ohakune Harris ”; 1 ex. (NZAC): “ NEW ZEALAND WO \ Mt Pirongia 500 m \ 16 Jan 1977 \ A. K. Walker and \ W. H. Benn ”, “ Litter \ 77 / 3 ”; 1 ex. (NMPC): “ NEW ZEALAND: Waikato (WO) \ Pirongia Forest Park \ Mangakara Nature Walk; 177 m \ 37.9667 ° S, 175 / 149 ° E \ 18. xi. - 21. xi. 2019 \ M. Fikáček & M. Seidel lgt. MMO 3 ”, “ Lowland broadleaf forest with \ podocarps, nikau palms and \ Ripogonum lianas; sparse \ understory: baited pitfall traps \ (rotten squid) ”; 1 ex. (NZAC): “ coll. \ A. C. O’Connor \ Feb. 1919 ”, “ Karori \ Wellington ”, “ 3244 ”, “ Atopida \ basalis \ Broun ”, “ A. E. Brookes \ Collection ”; 1 ex. (LUNZ): “ NEW ZEALAND, BP \ Te Koau 243 m \ NZMS 260 Z 14 778862 \ 24. x. 1992 - 31. i. 1993 \ R. M. Emberson ”, “ pitfall traps, mixed \ broadleaf / podocarp forest ”; 4 exx. (NZAC): “ NEW ZEALAND, BP \ Te Koau \ 1 Dec 1992 – \ 31 Jan 1993 \ J. S. Dugdale ”, “ Malaise trap ”; 4 males (NZAC): “ NEW ZEALAND, BP \ Orete Forest \ Te Puia Hut \ 19 Oct – \ 23 Nov 1992 ”, “ G. Hall \ Malaise trap ”; 1 ex. (LUNZ): “ NEW ZEALAND, TK \ Mount Egmont NP \ Stoney River \ 350 m, 25. xii. 1985 \ under stones ”, “ R. M. Emberson \ P. Syrett ”; 6 exx. (CMNC): “ N. Zealand, TK \ 21 km N. Ohura \ Dec. 14, 1983 \ L. Masner ”, “ Undergrowth in \ Nothofagus forest ”; 2 exx. (NZAC): “ New Zealand WN \ Otaki Forks \ 30 - Dec- 80 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 2 exx. (NZAC): “ NEW ZEALAND, WN \ Orongorongo V, W. N. \ 15 Dec 1994 ”, “ Hard beech \ forest litter \ Tullgren # 57 ”; 2 exx. (NZAC): “ New Zealand WN \ Pakuratahi Forks \ Kaitoke \ 06 - Jan- 93 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 ex. (NHM): “ N. Auck. \ 1 27 ”, “ NEW ZEALAND \ C. E. Clarke \ Collection \ B. M. 1957 - 24. ”; 1 male (AMNZ): “ Whangarei \ Falls \ 5. I. 24 [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”; 1 ex. (NHM): “ Whangarei \ 6 1 1927 \ C. E. Clarke ”; 1 ex. (NZAC): “ Kaimai \ 1500 ’ \ 3 - I- 48 ”, “ A. E. Brookes \ Collection ”; 1 ex. (NZAC): “ Parna Bay \ Whangarei \ Fairhum. Dec. 27 ”, “ A. E. Brookes \ Collection ”; 1 ex. (NZAC): “ Balance Reserve \ Manawatu Gorge \ 11 / 1 / 67 ”, “ Atopida \ suffusa ”, “ R. M. Bull \ Collection ”; 1 female (NZAC): “ 745 ”, “ NEW ZEALAND. \ Wainuiomata \ 745. \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1981 - 315 ”; 3 males (AMNZ): “ Silverstream \ 22. XII. 1923 ”, “ Hutt Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”; 1 male (AMNZ): “ Silverstream \ Wellington \ 15.12.23 \ Coll. T. [illegible] [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”, “ Atopida \ proba \ Sharp \ Id. by J. Armstrong ”; 2 males (NZAC): “ Okiwi \ Great Barrier \ Island ”, “ 24 Nov 57 \ J. C. Watt ”, “ beating \ flowering \ Leptospermum ”; 1 male (NZAC): “ Unuwhao 800 ’ \ Spirits Bay ”, “ Forest \ Litter ”; 1 male (NZAC): “ NEW ZEALAND RI \ Ruehine nr \ Rangiwahia \ 4 Jan 1975 \ J. C. Watt ”; 1 male (NZAC): “ NEW ZEALAND BP \ Kaimai Ra \ Lindeman lookout \ and of Lindeman Rd \ 4 Nov 2005 ”, “ R Leschen S Nomura \ beating RL 992 \ 37 ° 54 ’ S \ 175 ° 87 ’ E ”; 1 male (NZAC): “ Wharite \ 2 - 2 - 58 \ J. I. Townsend ”; 1 male (AMNZ): “ NEW ZEALAND AK \ Mataitai Forest S. A. \ pitfall B 1 \ Jan 2002 \ A. Warren ”; 2 males (NZAC): “ NEW ZEALAND CL \ Little Barrier I \ Awaroa Stm \ 15 Feb 1977 \ J. A. Macdonald ”, “ MV light trap \ at base of \ waterfall ”; 3 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220 m, \ 11. xii. 2001 - 18. i. 2002, \ P. Sutton. ”, “ Forest edge, Malaise \ trap. L 21044 ”; 6 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I, Little \ Windy Hill, 220 m, \ 7. xi. - 11. xii. 2001, \ P. Sutton & J. Gilbert. ”, “ Forest edge, Malaise \ trap. L 11043. ”; 3 males (AMNZ): “ NEW ZEALAND, CL \ Great Barrier I \ Mt Hobson, 500 m \ 17 Dec 2003 \ J. W. Early ”, “ Agathis / podocarp \ forest \ Swept. L 12446 ”; 1 male (AMNZ): “ Waikaraka \ Valley \ Parva Bay \ 14. i. 24 [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”, “ Atopida \ proba \ Sharp \ Id. by J. Armstrong ”; 1 male (AMNZ): “ swept understorey \ Waikaraka Strm. Vy. \ S. of Puketi Forest \ 14. XI. 1985 \ R. F. Gilbert ”, “ Bay of Island Co \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”; 1 male (AMNZ): “ Wayby Gorge \ N. Auck. \ 28.12.26 [on the underside of the cardboard with the beetle] ”, “ Rodney Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”; 2 males (AMNZ): “ Clevedon \ 27.2.34 [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	diagnosis	Diagnosis. Body of moderate size to large (TL 3.9 – 5.0 mm), uniformly brown. Setation of head and pronotum adherent, covered with somewhat flattened setae, with no suberect setae, what makes this species easy to distinguish from other Atopida.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	description	Redescription. Male. Body elongated, moderate size to large (TL 3.9 – 5.0 mm), moderately convex. Body uniformly brown, antennae tone lighter at base. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on elytra simple, punctures separated by 0.5 – 1.0 × diameter of a puncture, interspaces microreticulate, dull. Body covered with moderately long, adherent, yellowish setae, arranged into kind of pattern along the middle of pronotum and sometimes head. Supraantennal ridges well-marked, elevated. Pronotum widest in anterior 1 / 3, sides curved, anterolateral angles not or only subtly produced, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded. Penis long (L 0.63 mm, W 0.24 mm), trigonium narrow, as long as or slightly shorter than parameroids, slightly bent upwards, apex truncate; parameroids elongated, apices broadening and obliquely truncate, slightly serrated, apical part covered with few punctures; pala triangular, much longer than parameroids. Tegmen (L 0.52 mm, W 0.34 mm) with moderately wide subrectangular parameres, apices triangularly narrowing, with scarce punctures and delicate setae in apical part. Female. Body moderately big to large (TL 4.3 – 5.0 mm), macropterous. Supraantennal ridges well-marked, elevated. Pronotum strongly transverse, much stronger than in males, punctures along basal margin larger and sparser than on remaining portions of elytra. Setation on elytra very short, setae nor reaching the next puncture, elytral interspaces shining, punctures separated by ca. 0.5 × diameter of a puncture. Ventrite 5 rounded, apex rounded, tergite VII with rectangular sclerotization in apical portion. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite consisting of pair of small (L 0.15 mm) rod-like sclerites hooked in posterior portion. Measurements and ratios. Males (n = 10) TL 3.90 – 4.95 mm (4.50 mm), PL 0.70 – 0.85 mm (0.77 mm), PW 1.00 – 1.40 mm (1.22 mm), EL 3.15 – 4.10 mm (3.73 mm), EW 1.70 – 2.20 mm (2.04 mm), TL / EW 2.08 – 2.35 (2.21), PW / PL 1.33 – 2.00 (1.58), EL / EW 1.76 – 1.95 (1.83), EL / PL 4.50 – 5.79 (4.84), EW / PW 1.48 – 2.15 (1.67). Females (n = 10) TL 4.25 – 5.00 mm (4.63 mm), PL 0.75 – 0.90 mm (0.80 mm), PW 1.30 – 1.55 mm (1.45 mm), EL 3.45 – 4.15 mm (3.79 mm), EW 1.85 – 2.15 mm (2.01 mm), TL / EW 2.21 – 2.38 (2.30), PW / PL 1.67 – 2.07 (1.82), EL / EW 1.76 – 1.97 (1.89), EL / PL 4.11 – 5.47 (4.77), EW / PW 1.34 – 1.46 (1.39).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	distribution	Distribution. Common in North Island, known from scattered localities in Northland, Auckland, Coromandel, Waikato, Bay of Plenty, Gisborne, Taranaki, Rangitikei and Wellington areas (Fig. 18 H).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8102284DE7FF24FD73FBF3.taxon	discussion	Remarks. Broun (1912) correctly noted the similarities of A. sinuata and A. basalis, although he did not notice that these are males and females of the same species.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	description	(Figs 2 E – F, 5 Q – R, 8 J, 10 H – I, 15 B, 17 I)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	materials_examined	Type material. Holotype (NHM): “ 3071 ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Greymouth \ J. H. Larris ” [in Broun’s hand], “ Atopida \ suffusa ” [in Broun’s hand]. Other material examined. 2 exx. (NZAC): “ L. Wahapo \ 15 - I- 57, \ E. S. Gourlay ”; 7 females (NHM): “ WD: 6 km SW. \ Mahitahi \ swamp forest \ 5. ii. 1984 ”, “ NEW ZEALAND \ B. M. 1984 - 80 \ P. M. Hammond ”; 2 females (NMPC): “ NEW ZEALAND: Westland \ Three Mile Lagoon 3.7 km SW of \ Okaroto, 43 ° 14.5 ’ S 170 ° 7.5 ’ E, \ 0 m, 8. xii. 2012, Fikáček, Hájek \ & Leschen lgt. NZ 44 ”, “ sifting of ordinary leaf litter in low \ swampy coastal bushes on the \ dune behind the beach ”; 2 females (NMPC): “ NEW ZEALAND: Westland \ Westland NP; Lake Matheson \ 5 km NWW of Fox Glacier vill. \ 43 ° 26.3 ’ S 169 ° 58.0 ’ E, 120 m. \ 9. xii. 2012 \ Fikáček, Hájek & Leschen lgt. NZ 45 ”, “ sifting of leaf litter and mosses in \ the podocarp-broadleaf forest \ and its margins with tree ferns \ along E coast of the lake ”; 1 ex. (NMPC): “ NEW ZEALAND: Westland \ Ship Creek (Tauperikaka) \ 16 km NE Haast, 10 - 12. xii. 2012 \ 43 ° 45.7 ’ S 169 ° 8.7 ’ E, 0 - 10 m \ M. Fikáček, J. Hájek & R. Leschen lgt. ”; 1 female (NMPC): “ NEW ZEALAND: Westland \ Kellys Creek @ Otira Hwy., 3 km \ N of Otira, 350 m, 3 - 14. xii. 2012 \ 42 ° 48.2 ’ S 171 ° 34.4 ’ E \ Fikáček, Hájek & Leschen NZ 50 ”, “ sifting of mosses + accumulated \ leaf litter in the mossy forest on \ the flood plain of small mountain \ river + slightly above in \ Nothofagus-podocarp forest ”; 4 males (NMPC): “ NEW ZEALAND: Westland \ Lake Mapourika 9.5 km SE of \ Okaroto, 43 ° 17.8 ’ S 170 ° 13.5 ’ E, \ 160 m, 10. xii. 2012, Fikáček, \ Hájek & Leschen lgt. NZ 47 ”, “ on flowering Cordyline and \ Weinmannia ”; 1 female (CMNC): “ N. Zealand, WN \ Akatarawa saddle \ Dec. 17, 1983 \ L. Masner, s. s. ”; 1 male (AMNZ): “ Waiho Gorge \ S. Westland \ 16. I. 25 [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	diagnosis	Diagnosis. Body moderately sized to large (TL 4.0 – 5.1 mm), rather stout, covered with long, erect setae. Externally somewhat resembling A. villosa, but easy to distinguish on the basis of genitalia. Penis with wide, bulbous trigonium and bursal sclerite with long lateral rods and small, v-shaped central sclerite are known only in A. impressa. Easy to distinguish from A. impressa on the basis of external morphology, especially erect setation of dorsum.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	description	Redescription. Male. Body moderately sized to large (TL 4.00 – 5.10 mm) elongated, moderate size, moderately convex. Body brown, elytra a bit lighter than head and pronotum, antennae and legs lighter. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, granules a bit flattened in central portion of pronotal disc, separated by ca. 0.5 diameter; punctation on elytra simple, punctures separated by 0.5 – 1.0 × diameter of a puncture, interspaces shining. Body covered with long, erect, yellowish setae. Supraantennal ridges well-marked, elevated. Pronotum widest slightly before middle of its length, sides curved, anterolateral angles not produced, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded. Penis elongated (L 0.62 mm, W 0.24 mm), trigonium strongly widened in apical portion, oval, shorter than parameroids; parameroids elongated, apices obliquely truncate, axe-shaped, apical part covered with few punctures; pala subrectangular, narrow, ca. as long as parameroids. Tegmen (L 0.63 mm, W 0.42 mm) with narrow, subtriangular parameres, narrowing at apices, covered with numerous punctures and setation near inner margin of apical portion. Female. Body moderately big (TL 4.0 – 4.6 mm), brachypterous. Supraantennal ridges marked, not elevated. Pronotum more transverse than in males, punctation of elytra coarser. Ventrite 5 rounded, with deeply, semicircularly emarginated apex. Central portion of ventrite 4 with two oblique setal patches. Bursal sclerite consisting of two lateral rods (L 0.30 mm) widened in posterior portion and armed with small denticles on outer parts of anterior portion, additional small, v-shaped sclerite present between rods. Measurements and ratios. Males (n = 6) TL 4.00 – 5.10 mm (4.56 mm), PL 0.70 – 0.85 mm (0.78 mm), PW 1.35 – 1.65 mm (1.47 mm), EL 3.60 – 4.25 mm (3.83 mm), EW 2.05 – 2.35 mm (2.18 mm), TL / EW 1.95 – 2.23 (2.09), PW / PL 1.75 – 2.06 (1.87), EL / EW 1.67 – 1.84 (1.76), EL / PL 4.50 – 5.36 (4.90), EW / PW 1.42 – 1.56 (1.49). Females (n = 4) TL 4.00 – 4.56 mm (4.26 mm), PL 0.75 – 0.85 mm (0.81 mm), PW 1.40 – 1.60 mm (1.53 mm), EL 3.35 – 3.65 mm (3.50 mm), EW 2.10 – 2.25 mm (2.16 mm), TL / EW 1.86 – 2.05 (1.97), PW / PL 1.87 – 1.88 (1.88), EL / EW 1.56 – 1.67 (1.62), EL / PL 4.24 – 4.47 (4.31), EW / PW 1.34 – 1.54 (1.42).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	distribution	Distribution. Described from Greymouth in Buller area, but most of the specimens examined were collected in various localities in Westland area of South Island (Fig. 18 I).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF83022F4DE7FB19FE44FA9B.taxon	discussion	Remarks. The species is an unusual example of a marsh beetle with genital structures (both aedeagus and bursal sclerite) hardly distinguishable from another species from which it can be easily separated on the basis of external morphology. Usually species of marsh beetles that are externally almost identical differ considerably at least in male genital features.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF84022C4DE7FA31FC9BFC2B.taxon	description	(Figs 2 G – H, 3 F, 4, 5 S – T, 6 A, 7 A – B, 8 K – L, 10 J – K, 15 C, 17 B)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF84022C4DE7FA31FC9BFC2B.taxon	materials_examined	Type material. Holotype of Ptinus suturalis (NHM): pinned into card, “ Type ” [round label with red border], “ Ptinus suturalis White \ Zool. Ereb & Terror. \ Leptosp. flowers \ Hooker and Smith ” [handwritten]. Holotype of A. lawsoni, female (NHM): card-mounted, “ Atopida Lawsoni \ Type D. S. \ Auckland Lawson ” [written at base of card in Sharp’s hand], “ Type \ H. T. ” [round label with red border], “ Auckland, [red underline] \ New Zealand. ”, “ Sharp Coll. \ 1905 - 313. ” Syntypes of A. brouni, 2 exx. (NHM): card-mounted, “ Atopida brouni \ Type D. S. \ Tairua Broun ” [written at base of card in Sharp’s hand], “ Type \ H. T. ” [round label with red border], “ Tairua, [red underline] \ New Zealand. \ Broun. ”, “ Sharp Coll. \ 1905 - 313. ” Syntype, 1 ex. (NHM): card-mounted, “ Atopida brouni \ Ind. typ. \ D. S. \ Tairua Broun ” [written at base of card in Sharp’s hand], “ Tairua, [red underline] \ New Zealand. \ Broun. ”, “ Sharp Coll. \ 1905 - 313. ”, “ = suturalis White ” [handwritten]. Other material examined. 3 males (AMNZ): “ Wayby Gorge \ N. Auck. \ 28.12.26 [on the underside of the cardboard with the beetle] ”, “ C. E. Clarke \ Collection ”, “ Atopida \ suturalis \ White \ = brouni Shp. \ Id. by J. Armstrong \ comp. Cot. brouni ”; 1 ex. (CMNC): “ NEW ZEALAND \ North Is. Huia \ 17 - 27. XI. 1981 \ H & A Howden ”; 7 exx. (DBET): “ New Zealand AK 1: \ Waikatere Range, \ Waiatarua W, - 36.9372, \ 174.5575, 260 m, beating \ in wet forest, 26.11.2016, \ leg. R. Ruta ”; 14 exx. (DBET): “ New Zealand ND 24: \ Waipoua Forest, Yakas \ Track, 7.12. 2016, 300 m \ beating, leg. R. Ruta ”; 1 ex. (DBET): “ New Zealand ND 25: \ Waipoua Forest, Yakas \ Track, 300 m, 7.12.2016, \ litter sample, leg. R. Ruta ”; 2 exx. (DBET): “ New Zealand ND 29: \ Toatoa Track, - 35.67, \ 173.56, 270 m, beating, \ 7.12.2016, leg. R. Ruta ”; 17 exx. (CMNC): “ N. Zealand, ND \ Waipoua Kauri Forest \ Dec. 11 & 12, 1983 \ L. Masner ”; 4 exx. (CMNC): “ NEW ZEALAND: N. Is. \ Waipoua S. Forest \ 20 - 23. XI. 1981 \ H & A Howden ”; 2 exx. (NZAC): “ Waipoua \ S. F. Oct. 67 \ Auckland ”, “ at light \ camp ”, “ J. C. Watt ”; 1 ex. (NZAC): “ Waipoua S. F. \ Auckland \ 12 Oct. 67 ”, “ at light \ J. C. Watt ”; 1 ex. (LUNZ): “ Waipoua For. \ 7.12.61 \ G. Kuschel ”; 1 ex. (LUNZ): “ Waipoua S. F. \ 16. X. 67 ”, “ to m / v light \ J. S. Dugdale ”; 1 ex. (LUNZ): “ NEW ZEALAND ND \ Waipoua SF \ 4 Feb 1975 \ sweeping ”; 2 exx. (NZAC): “ Waipoua Beach \ 18 Oct. 67 \ Auckland ”, “ at light ”, “ J. C. Watt ”; 1 ex. (NZAC): “ NEW ZEALAND ND \ Waipoua Forest \ 1 - 2 Dec 1989 \ D. J. Bickel \ Pan traps ”; 1 ex. (NZAC): “ NEW ZEALAND ND \ Waipoua SF \ Waipoua River \ 31 Oct 1985 \ R. C. Craw ”; 1 female (NZAC): “ NEW ZEALAND ND \ Waipoua SF \ 26 Nov 1980 \ G. Kuschel ”; 1 male (NZAC): “ Waipoua \ S. F. Oct. 67 \ Auckland ”, “ m / v. \ light. ”, “ J. S. Dugdale ”; 1 male (NZAC): “ 988 ”, “ New Zealand ND \ Waipoua Forest \ 25.11.94 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 2 males (NZAC): “ NEW ZEALAND ND \ Waipoua SF \ 26 Nov 1980 \ G. Kuschel ”; 1 male (NZAC): “ NEW ZEALAND, ND \ Waipoua SF \ Lookout \ 29 Oct 1980 \ G. Kuschel ”; 1 male (NZAC): “ NEW ZEALAND, ND \ Waipoua SF \ Lookout \ 29 Oct 1980 \ G. Kuschel ”; 1 male (NZAC): “ Kawerua \ Beach ”, “ 19. x. 67 \ J. S. Dugdale ”, “ Waipoua S. F. ”; 1 ex. (NHM): “ Hunua G. \ nr Papakura \ 13 - 12 - 36 \ A. Richardson ”, “ NEW ZEALAND \ C. E. Clarke \ Collection \ B. M. 1957 - 24. ”; 2 exx. (LUNZ), 1 female (NZAC): “ Hunua Falls \ J. C. Watt \ 2 Dec 61 AK ”, “ Dacrydium \ cupressinum \ Podocarpus \ totara ”; 3 exx. (NZAC): “ Hunua Ra. \ 2.12.61 \ G. Kuschel ”; 2 males (AMNZ): “ Hunua \ Gorge \ 13 - 12 - 36 \ A. Richardson [on the underside of the cardboard with the beetle] ”, “ HUNUA G. \ nr. Papakura \ 13 - 12.36 \ A. Richardson ”, “ Manukau Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”, “ C. ocularis ”; 1 male (NHM): “ Waitakere Ra \ Destruction \ Gully \ 2. xi. 1985 \ C & J Lyal ”, “ General \ beating ”, “ NEW ZEALAND \ AK (CL 94) \ BM 1986 - 274 ”; 2 females (NZAC): “ NEW ZEALAND \ AK (CL 94) \ BM 1986 - 274 ”, “ Waitakere Ra \ Destruction \ Gully \ 2. xi. 1985 \ C & J Lyal ”, “ General \ beating ”; 1 male (AMNZ): “ 31.12.47 \ Waitakere \ Ras. C. T. ”, “ Waitemata Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”, “ Atopida \ sp. ”; 1 female (NZAC): “ Waitakeri \ 30 - 12 - 15 ”, “ 560 ”, “ T. Broun \ Collection ”, “ A. E. Brookes \ Collection ”: 1 male (NZAC): “ Waitakeri \ 31 - 12 - 14 ”, “ 561 ”, “ T. Broun \ Collection ”; 1 female (NZAC): “ NEW ZEALAND AK \ Waitakere Ra \ Fairy Falls Trk \ 4 Dec 1979 ”, “ C. F. Butcher and \ M. F. Tocker \ On flowers of \ Leptospermum \ scoparium ”; 1 female (NZAC): “ NEW ZEALAND AK \ Waitakere Range \ Cutty Grass Trk \ 2 Jan 1984 \ R. C. & J. P. N. Craw ”; 1 male (NZAC): “ WAITAKERES \ AUCKLAND \ 9. DEC. 1953 \ A. E. BROOKES. ”; 1 male (NZAC): “ Anawhata Rd. \ Waitakere \ 1 Jan 1945 \ M. W. Carter ”; 2 exx. (NHM): “ NEW ZEALAND \ ND (CL 55) \ BM 1986 - 274 ”, “ Omapere \ 30. X. 1985 \ C & J Lyal ”; 3 exx. (NZAC): “ Pandora ”, “ Spirits Bay \ Northland \ Nov 67 \ J. I. Townsend ”; 1 ex. (NZAC): “ NEW ZEALAND ND \ Unuwhao, 270 m \ 25 Nov 198 X \ G. Kuschel ”, “ sifted litter \ and decayed \ wood 82 / 125 ”; 2 exx. (NZAC): “ Uauwhao 1000 ’ \ Spirits Bay ”, “ 18.1.66 \ A. K. Walker ”, “ Rhopalostylis \ sapida ”; 1 ex. (NZAC): “ Unuwhao 243 ’ \ Spirits Bay ”, “ 18. I. 66 \ A. K. Walker ”; 10 males, 2 females (NZAC): “ NEW ZEALAND ND \ Unuwhao, 270 m \ 25 Nov 1982 \ G. Kuschel ”, “ Sifted litter \ and decayed \ wood 82 / 125 ”, “ Duplicate \ specimens \ in alcohol ”; 2 males (NZAC): “ NEW ZEALAND ND \ Unuwhao, 270 m \ 26 Nov 1982 \ G. Kuschel ”; 1 ex. (NZAC): “ NEW ZEALAND CL \ Little Barrier I \ 183 m 29 Feb 1976 \ A. K. Walker \ sweeping ”; 1 ex. (NZAC): “ NEW ZEALAND AK \ Riverhead State \ forest Reserve \ 11 Nov 1980 \ M. F. Tocker ”, “ on \ Myrsine \ australis ”; 1 ex. (NZAC): “ NEW ZEALAND \ TV \ 28 Dec 1997 \ P A Maddison ”; 5 exx. (NZAC): “ NEW ZEALAND ND \ Paihia Opua SF \ 22 Jan 1981 \ G Kuschel ”; 1 ex. (NZAC): “ NEW ZEALAND, TK \ Piopio, Mangaotaki \ Bush Walk \ 8 Dec 2000, G. Hall \ R. Hoare, T. Sirey ”, “ under logs \ Podocarp forest \ 38 ° 28 ’ S, 174 ° 51 ’ E ”; 1 ex. (NZAC): “ NEW ZEALAND ND \ Warawara camp \ 15 December 2007 \ T R Buckley D Seldon \ R Hoare ”, “ to MVL \ TB 107 \ 35 22.167 S \ 173 17.012 E ”; 1 ex. (NZAC): “ NEW ZEALAND CL \ Rd E of Tapu \ 450 m Summit \ 3 Dec 1989 \ D. J. Bickel ”; 2 exx. (NZAC): “ Swanson \ X Totara \ 18 - 12 - 55 J. C. Watt \ 439 B ”; 3 exx. (NZAC): “ Swanson \ X Karamu \ 18 - 12 - 55 J. C. Watt \ 439 B ”; 1 ex. (NZAC): “ Swanson \ X Olearia \ 18 - 12 - 55 J. C. Watt \ 439 B ”; 1 ex. (NZAC): “ Swanson \ X Tanakaha \ 18 - 12 - 55 J. C. Watt \ 439 B ”; 5 exx. (NAZC): “ Swanson \ X Kanuka \ 18 - 12 - 55 J. C. Watt \ 439 B ”; 2 exx. (NZAC): “ Swanson \ Auckland \ 13 Dec 58 ”, “ J. C. Watt ”; 2 males (NZAC): “ Swanson \ Auckland \ 13 Dec 58 ”, “ J. C. Watt ”; 1 female (NZAC): “ Ex light trap \ Titirangi \ NOV / 51 \ C. R. Thomas. ”; 5 exx. (NZAC): “ Ex light trap \ Titirangi \ NOV / 52 \ C. R. Thomas. ”; 1 female (NZAC): “ Ex light trap \ Titirangi \ DEC / 52 \ C. R. Thomas. ”; 2 exx. (NZAC): “ Ex light trap \ Titirangi \ DEC / 53 \ C. R. Thomas. ”; 1 male (NZAC): “ Titirangi \ 28 Nov 1942 \ M. W. Carter ”; 1 ex. (NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 15 Oct 1978 \ P. A. Maddison ”, “ At \ light ”; 2 females (NZAC): “ Titirangi ”, “ P. E. B. 27 - 12 - 49 ”, “ A. E. Brookes \ Collection ”; 2 females (NZAC): “ NEW ZEALAND AK \ Titirangi \ Minnehaha Res \ 9 Dec 1980 \ P. A. Maddison ”; 1 female (NZAC): “ NEW ZEALAND AK \ Titirangi \ Minnehaha Res \ 9 Dec 1980 \ P. A. Maddison ”, “ Beating \ streamside \ shrubs ”; 1 female (NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 10 Dec 1978 \ P. A. Maddison ”, “ In house ”; 2 males (NZAC): “ NEW ZEALAND AK \ Auckland \ Titirangi \ 15 Dec 1978 \ P. A. Maddison ”, “ At light ”; 2 exx. (NZAC): “ Waimatenui \ Northland \ North Is. ”, “ Collector \ 10.12.1951 \ A. E. Brookes ”; 1 female (NZAC): “ Waimatenui \ V. 1715.10.12.69 \ Brooks ”, “ A. E. Brookes \ Collection ”; 1 female (NHM): “ Paparoa ”, “ Atopida \ lawsoni ”, “ New Zealand. \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ lawsoni \ Broun coll. ”; 1 male, 1 female (NZAC): “ 980 ”, “ New Zealand ND \ Radar Bush \ Te Paki \ 4.12.95 ”, “ In forest \ litter ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 female (NZAC): “ NEW ZEALAND ND \ Te Paki Trig \ 23 Nov 1982 \ G. Kuschel ”, “ Sifted litter \ and decayed \ wood 82 / 126 ”; 2 males (NZAC): “ NEW ZEALAND ND \ Te Paki Trig \ 25 Nov 1982 \ G. Kuschel ”, “ Sifted litter \ and decayed \ wood 82 / 117 ”; 1 female (NZAC): “ New Zealand WN \ Pakuratahi Forks \ Katioke \ 7 - 10 Feb 92 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 female (AMNZ): “ NEW ZEALAND, ND \ Waima R catchment \ Mangatete Stm \ 15. x. 1985, R. F. Gilbert \ Swept foliage by stream ”, “ AMNZ 126025 \ AUCKLAND \ MUZEUM \ NEW ZEALAND ”; 1 male (NZAC): “ NEW ZEALAND ND \ Long Louis \ Waima Forest \ 13 Feb 2008 \ R Leschen B Michaux ”, “ beating \ RL 13330 \ 35 ° 32.222 ’ S 173 ° 27.238 ’ E ”; 1 male (AMNZ): “ ex. Folliage \ Walkers Bsh \ Nov. 1966. MBP. ”, “ Waitemata Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ M. B. Paterson \ Collection ”, “ AMNZ 126073 \ AUCKLAND \ MUZEUM \ NEW ZEALAND ”; 1 male (NZAC): “ NORTHCOVE ”, “ 561 ”, “ T. Broun \ Collection ”; 1 male (NZAC): “ New Zealand ND \ Omahuta SF \ Nov- 94 ”, “ In leaf \ litter ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 male (NZAC): “ NEW ZEALAND, ND \ Pukenui F \ 14 Sep 1998 \ G. Hall \ under logs ”; 1 male (NZAC): “ NEW ZEALAND, AK \ Hillsborough \ 2001 \ D. S. Seldon \ Pittosporum tree ”; 1 male (NZAC): “ NEW ZEALAND WN \ Stakas Valley \ 23 Feb 1975 \ B. A. Hallaway ”; 1 male (AMNZ): “ Albany \ Hobson Block \ 11. XI. 1982 \ R. F. Gilbert ”; 1 male (AMNZ): “ NEW ZEALAND AK \ Atuanui \ Mt Auckland \ pitfall F 25 \ Jan 2002, A. Warren ”; 2 males (AMNZ): “ NEW ZEALAND, ND \ E of Puketi SF \ Mangirangina Scientific \ Reserve, 12. xi. 1985 \ R. F. Gilbert ”, “ Swept entrance to \ walkway ”; 1 male (AMNZ): “ swept understorey \ of high forest \ Waikaraka Valley \ S. of Puketi Forest \ 13. XI. 1985 \ R. F. Gilbert \ NZMSI NII 273527 \ 35 ° 16 ’ 173 ° 48 ’ ”; 1 male (AMNZ): “ swept understorey \ Waikaraka Strm. Vy. \ S. of Puketi Forest \ 14. XI. 1985 ”; 1 male (AMNZ): “ NEW ZEALAND AK \ Mataitai Forest S. A. \ pitfall D 25 \ Jan 2002 \ A. Warren ”; 2 males (AMNZ): “ MANGAMUKA G. \ G. Z. 64 \ R. A. CUMBER ”; 1 male (AMNZ): “ Mangamuka ”, “ Mangonui Co. \ North Island \ NEW ZEALAND \ AUCKLAND MUSEUM ”, “ C. E. Clarke \ Collection ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF84022C4DE7FA31FC9BFC2B.taxon	diagnosis	Diagnosis. Body relatively large (TL 4.5 – 5.3 mm), light brown to brown, head and pronotum usually a tone darker than elytra, setation of elytra adherent. Externally resembling A. castanea but pronotum of females with sparse punctation in basal half, elytra of females often yellowish and contrasting with dark brown pronotum and head. Penis moderately elongated, trigonium relatively wide, with rounded apex and parallel sides, as long as parameroids; pala slightly shorter than parameroids. Bursal sclerite long, with hooked processes in central portion.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF84022C4DE7FA31FC9BFC2B.taxon	description	Redescription. Male. Body elongated, moderate to large (TL 4.5 – 5.3 mm), moderately convex. Body brown, elytra often lighter than head and pronotum. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other, punctation on elytra simple, punctures separated by 0.5 – 1.0 × diameter of a puncture. Body covered with relatively short setae, semi-erect on head and pronotum, and adherent on elytra. Supraantennal ridges well-marked, elevated. Pronotum widest in anterior 1 / 3, sides curved, anterolateral angles slightly produced, posterolateral angles slightly obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, with triangularly rounded apex. Penis (L 0.68 mm, W 0.29 mm) moderately elongated, trigonium narrow, rectangular, with rounded apex and parallel sides, as long as or slightly longer than parameroids; parameroids elongated, apices pointing outwards, covered with numerous punctures, inner edge slightly irregular; pala triangular, about the same length as parameroids. Tegmen (L 0.59 mm, W 0.33 mm) moderately wide, parameres slightly tapered, with scarce punctures and very few delicate setae on the inner edge. Female. Body moderate to large (TL 4.5 – 5.2 mm), elongate, moderately convex, macropterous. Body brown, head and pronotum darker, elytra contrasting yellowish-brown, legs and antenna lighter. Supraantennal ridges wellmarked, slightly elevated triangularly above bases of antennae. Pronotum with peculiar punctation — punctures in basal portion sparse, punctures separated by 2.0 × diameter of a puncture, interspaces glabrous, remaining areas of pronotum covered with coarse, granulate punctures. Anterolateral angles of pronotum more produced than in males, triangular. Elytra covered with very short, adherent setae, distinctly shorter than in males, not reaching next puncture, punctation of elytra coarser than in males. Ventrite 5 rounded, with triangularly emarginated apex. Lateral portions of ventrites 3 and 4 with small, oval setal patches. Bursal sclerite long (L 0.40 mm), anterior and posterior portion with two subparallel rod-like structures, central portion subtriangular, with well visible hooked projections on sides. Measurements and ratios. Males (n = 10) TL 4.50 – 5.30 mm (4.98 mm), PL 0.75 – 0.90 mm (0.86 mm), PW 1.25 – 1.45 mm (1.39 mm), EL 3.75 – 4.50 mm (4.16 mm), EW 1.95 – 2.40 mm (2.15 mm), TL / EW 2.21 – 2.38 (2.32), PW / PL 1.53 – 1.73 (1.62), EL / EW 1.86 – 2.00 (1.93), EL / PL 4.41 – 5.29 (4.87), EW / PW 1.48 – 1.68 (1.55). Females (n = 10) TL 4.50 – 5.20 mm (4.76 mm), PL 0.75 – 0.95 mm (0.84 mm), PW 1.35 – 1.60 mm (1.48 mm), EL 3.80 – 4.20 mm (3.98 mm), EW 1.95 – 2.25 mm (2.11 mm), TL / EW 2.14 – 2.36 (2.11), PW / PL 1.67 – 1.93 (1.76), EL / EW 1.81 – 1.98 (1.89), EL / PL 4.42 – 5.07 (4.74), EW / PW 1.34 C 1.50 (1.43).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF84022C4DE7FA31FC9BFC2B.taxon	distribution	Distribution. Widely distributed in North Island, common in Northland and Auckland areas, less common in Coromandel, Waikato, and Wellington areas (Fig. 18 J).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	description	(Figs 2 I – J, 5 U – V, 8 M, 10 L, 15 D, 17 P)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	materials_examined	Type material. Holotype, male (NZAC): “ NEW ZEALAND WD \ Mt Tuhua, 1067 m \ E side of \ L. Kaniere \ 20 Nov 1984 ”, “ B. A. Holloway \ Moss and \ lichen 84 / 75 ”; paratypes, 7 exx. (NZAC): same data as holotype.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	diagnosis	Diagnosis. Body small to moderately big (TL 3.1 – 3.8 mm), dark brown, head, pronotum, and scutellar shield slightly darker than elytra. Sides of elytra crenulate. Certain identification possible on the basis of genital characters: parameroids narrow, pointed at apices, parameres wide, apices obliquely truncate; bursal sclerite large, in dorsal view with well-marked sharp lateral processes and central tubercle.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	description	Description. Male. Body elongated, small to moderately big (TL 3.4 – 3.8 mm), moderately convex, surface of head and pronotum uneven. Body dark brown, head, pronotum, and scutellar shield slightly darker than elytra, elytra sometimes darkened along suture. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other, with the exception of basal portion of pronotum where the punctures are more sparse, with shining interspaces; punctation on elytra simple, punctures separated by 0.5 × diameter of a puncture. Body covered with moderately long, erect setae. Supraantennal ridges well-marked, elevated, but not forming subtriangular flaps. Pronotum widest in anterior 1 / 3 of its length, lateral carinae crenulate, anterolateral angles distinctly produced, posterolateral angles pointed. Elytra without carinae, humeri well-marked. Ventrite 5 with regularly rounded apex. Penis (L 0.70 mm, W 0.35 mm) moderately elongated, with explanate lateral lobes, trigonium very narrow, curved laterally, apex rounded, distinctly shorter than parameroids; parameroids elongated, apices pointed, narrowly triangular, covered with numerous punctures, inner edge denticulate; pala triangular, shorter than parameroids. Tegmen (L 0.58 mm, W 0.45 mm) moderately wide, parameres relatively wide, subrectangular, obliquely truncate at apices, covered with punctures and very few delicate setae near outer edge. Female. Body small to moderately big (TL 3.1 – 3.6 mm), brachypterous. Supraantennal ridges well-marked, elevated. Pronotum more transverse than in males, disc with sparser punctation. Sides of elytra more rounded. Ventrite 5 rounded, apex truncate. Lateral portions of ventrites 3 and 4 with small, slightly oblique setal patches, ventrite 5 with two very small circular patches. Bursal sclerite big (L 0.46 mm, W 0.24 mm), elongated, consisting of two interconnected plate-like sclerites that are fused, anterior portion triangular, central part with a single tubercle, posterior portion rounded, in dorsal view with two subtriangular denticles on lateral margins, in lateral view these structures resemble hooks. Measurements and ratios. Males (n = 4) TL 3.40 – 3.75 mm (3.59 mm), PL 0.60 – 0.65 mm (0.63 mm), PW 0.95 – 1.10 mm (1.03 mm), EL 2.80 – 3.00 mm (2.92 mm), EW 1.65 – 1.70 mm (1.69 mm), TL / EW 2.06 – 2.21 (2.13), PW / PL 1.58 – 1.69 (1.64), EL / EW 1.70 – 1.76 (1.73), EL / PL 4.54 – 4.89 (4.68), EW / PW 1.55 – 1.74 (1.65). Females (n = 3) TL 3.14 – 3.60 mm (3.30 mm), PL 0.50 – 0.70 mm (0.58 mm), PW 1.05 – 1.15 mm (1.07 mm), EL 2.60 – 2.95 mm (2.72 mm), EW 1.55 – 1.85 mm (1.72 mm), TL / EW 1.80 – 2.03 (1.93), PW / PL 1.64 – 2.00 (1.85), EL / EW 1.49 – 1.68 (1.59), EL / PL 4.21 – 5.20 (4.71), EW / PW 1.55 – 1.67 (1.61).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	distribution	Distribution. Known only from the type locality in Westland area of South Island (Fig. 18 L).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FF8702134DE7FC61FD37FE07.taxon	etymology	Etymology. From type locality, Mt. Tuhua.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	description	(Figs 2 K – L, 5 W – X, 8 N, 9 B, 10 M – N, 16 A, 17 F)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	materials_examined	Type material. Holotype (NHM): card-mounted, “ 4061. ” [in Broun’s hand], “ New Zealand. [red underline] \ Broun Coll. \ Brit. Mus. \ 1922 - 482. ”, “ Pakuratahi \ 2.1.1915 ” [in Broun’s hand], “ Atopida \ villosa ” [in Broun’s hand]. Other material examined. 1 ex. (NZAC): “ Akatarawa \ 17. Nov. 51 \ E. S. Gourlay ”; 1 ex. (NZAC): “ Silverstream \ WN. 19 Nov 51 \ E. S. Gourlay ”; 3 exx. (NZAC): “ New Zealand WN \ Keith George Res \ Silverstream \ 08 - Nov- 92 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 male (NZAC): “ New Zealand WN \ Keith George Res \ Silverstream \ 18 - Nov- 96 ”, “ General \ beating ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 ex. (NHM): “ NEW ZEALAND \ Days Bay \ Wellington \ 19. XI. 1931 \ G. V. Hudson ”, “ Pres. by \ Com. Inst. Ent. \ B. M. 1948 - 525. ”; 2 exx. (NZAC): “ New Zealand WN \ Pakuratahi Forks \ Kaitoke \ 19 - Dec- 92 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 female (NZAC): “ 981 ”, “ New Zealand WN \ Pakurathi Forks \ 18 - Dec- 94 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 female (NZAC): “ New Zealand WN \ Pakuratahi Forks \ Kaitoke \ 19 - Nov- 96 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 6 exx. (NZAC): “ New Zealand WN \ Pakuratahi Forks \ Wellington \ 17 - Nov- 96 ”, “ General \ beating ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 male (NZAC): “ 1116 ”, “ New Zealand WN \ Karori Reservoir \ Wellington \ 20 - Nov- 96 ”, “ Amongst submerged \ leaves in \ small stream ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 ex. (NHM): “ NEW ZEALAND \ Korokoro \ XI. 1943 \ G. V. Hudson ”, “ Pres. by \ Com. Inst. Ent. \ B. M. 1948 - 525. ”; 3 males (NZAC): “ NEW ZEALAND \ Korokoro \ 1039. \ 12. xi. 1938 \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”; 1 ex. (NZAC): “ Mt. Holdsworth \ 2000 ’ \ Tararuas \ 19. xi. 68 \ S. Edrige ”, “ moss \ 68 / 203 ”; 1 male (NHM): “ 7 - ii- 31 ”, “ NEW ZEALAND \ Days Bay \ Wellington \ 1039. \ 17. xi. 1931 r. \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”; 1 male (NZAC): “ New Zealand WN \ Khandallah \ Domain \ 17 - Aug- 92 ”, “ In Tawa \ litter ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 1 male (NZAC): “ 1116 ”, “ New Zealand WN \ Eastbourne \ 15 - Nov- 95 ”, “ Ex John Nunn \ Collection \ Donated 2015 ”; 2 males (NZAC): “ 1039 ”, “ NEW ZEALAND \ Wainuiomata \ XII. 1919. \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”; 1 male (NHM): “ 103 ”, “ NEW ZEALAND \ Gollans Valley \ 11. XI. 1939 \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”; 1 male (NHM): “ NEW ZEALAND \ Gollans Valley \ 1039. \ Wellington \ xii. 1933 \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”; 1 male (NHM): “ NEW ZEALAND \ Karori (?) \ G. V. Hudson ”, “ Pres. by \ Comm Inst Ent \ B. M. 1948 - 525. ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	diagnosis	Diagnosis. Body moderately big to large (TL 4.3 – 5.4 mm), stout, covered with very long, erect setae. Male genitalia similar to those of A. montana, bursal sclerite consisting of long, paired rods, hooked in posterior portion. Easy to distinguish from other Atopida on the basis of external features: long, erect setation, and females with peculiar concavity of elytral base fitting basal portion of pronotum.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	description	Redescription. Male. Body moderately elongated, moderately big to large (TL 4.6 – 5.4 mm), moderately convex. Body dark brown, head, pronotum, and scutellar shield slightly darker than elytra. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other; punctation on elytra simple, punctures separated by 0.5 – 1.0 × diameter of a puncture. Body covered with long, erect setae. Supraantennal ridges well-marked, elevated. Pronotum widest slightly before middle, sides rounded, anterolateral angles only subtly produced, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 with regularly rounded apex. Penis (L 0.76 mm, W 0.29 mm) moderately elongated, trigonium narrow, with rounded apex and parallel sides, slightly shorter than parameroids. Pala triangular, as long as parameroids; parameroids elongated, lateral edges subparallel, apices pointing outwards, covered with numerous punctures, inner edge slightly crenulate. Tegmen (L 0.65 mm, W 0.38 mm) moderately wide, parameres subtriangular, narrowing towards apices, with scarce punctures and very few delicate setae on the inner edge. Female. Body moderately big (TL 4.3 – 4.8 mm), stouter than males, brachypterous. Supraantennal ridges wellmarked, elevated. Punctation of elytra coarser, punctures separated by 0.3 × diameter of a puncture, basal portion of elytra with concavity fitting base of pronotum. Ventrite 5 triangular, with deep, subtriangular emargination at apex. Ventrites 3 and 4 with paired setal patches forming long transverse stripes close to posterior margins of segments. Bursal sclerite consisting of two long (L 0.40 mm) rods pointed in anterior portion and hooked posteriorly. Measurements and ratios. Males (n = 7) TL 4.55 – 5.40 mm (4.91 mm), PL 0.80 – 0.90 mm (0.85 mm), PW 1.45 – 1.70 mm (1.57 mm), EL 3.75 – 4.35 mm (4.01 mm), EW 2.25 – 2.45 mm (2.35 mm), TL / EW 2.02 – 2.20 (2.09), PW / PL 1.78 – 1.94 (1.85), EL / EW 1.64 – 1.78 (1.70), EL / PL 4.44 – 5.00 (4.72), EW / PW 1.44 – 1.59 (1.50). Females (n = 4) TL 4.25 – 4.80 mm (4.50 mm), PL 0.80 – 0.95 mm (0.86 mm), PW 1.55 – 1.75 mm (1.64 mm), EL 3.40 – 3.85 mm (3.60 mm), EW 2.15 – 2.35 mm (2.26 mm), TL / EW 1.93 – 2.04 (1.99), PW / PL 1.84 – 1.94 (1.90), EL / EW 1.55 – 1.64 (1.59), EL / PL 4.05 – 4.29 (4.18), EW / PW 1.34 – 1.42 (1.38).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	distribution	Distribution. Known from numerous localities in Wellington area of North Island (Fig. 18 K).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFB802124DE7FE4DFCB2FCBF.taxon	discussion	Remarks. Broun (1921) noted his specimen was apterous and had peculiar structure of elytral base; the holotype was found in decaying leaves on the ground (Broun 1921).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	description	(Figs 2 M, 5 Y, 16 B, 17 J)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	materials_examined	Type material. Holotype, male (NZAC): “ Waipoua S. F. \ Auckland \ Prov. 19 Oct 67 \ J. C. Watt ”, “ Beating \ flowers ”, “ Leptospermum \ scoparium ”; paratypes, 14 exx. (NZAC): same data as holotype; paratype, 1 ex. (NZAC): “ NEW ZEALAND ND \ Waipoua \ Waipoua Stm \ 26 Nov 1980 \ T. K. Crosby ”; paratypes, 2 exx. (CMNC): “ NEW ZEALAND: N. Is. \ Waipoua S. Forest \ 20 - 23. XI. 1981 \ H & A Howden ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	diagnosis	Diagnosis. Body small (TL 3.0 – 3.3 mm), narrow. Setation of dorsum long, erect, pronotum widest in the middle of its length. Penis same width over the entire length, lateral sides subparallel, trigonium subtriangular, rounded at apex, slightly shorter than parameroids; pala rectangular, shorter than parameroids; tegmen rectangular, with triangularly pointed parameres.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	description	Description. Male. Body elongated, small (TL 3.03 – 3.30 mm) and narrow. Body light brown, head, pronotum, and scutellar shield slightly darker than elytra, outer edges and adsutural portion of elytra often darkened, antennae darkening towards apices. Head and pronotum opaque, evenly covered with strong and dense, granulate, doughnutshaped punctures, punctures touching each other. Elytra shining, punctation on elytra simple, punctures separated by 0.5 × diameter of a puncture. Body densely covered with moderately long, protruding, yellowish setae. Supraantennal ridges well-marked, not elevated. Pronotum widest in the middle of its length, sides curved, anterolateral angles slightly produced, posterolateral angles slightly obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex delicately truncate. Penis (L 0.51 mm, W 0.17 mm) same width over the entire length, trigonium medium width, subtriangular, shorter than parameroids. Pala rectangular, slightly shorter than parameroids. Parameroids elongated, rectangular, apices wide, apical part covered with punctures. Tegmen (L 0.48 mm, W 0.21 mm) narrow, same width over the entire length, with triangular parameres, acute apices, scarce punctures and setae. Female unknown. Measurements and ratios. Males (n = 10) TL 3.03 – 3.30 mm (3.19 mm), PL 0.50 – 0.57 mm (0.55 mm), PW 0.87 – 0.97 mm (0.92 mm), EL 2.40 – 2.77 mm (2.59 mm), EW 1.40 – 1.53 mm (1.49 mm), TL / EW 2.07 – 2.29 (2.14), PW / PL 1.53 – 1.80 (1.68), EL / EW 1.60 – 1.84 (1.73), EL / PL 4.24 – 5.00 (4.74), EW / PW 1.56 – 1.73 (1.63).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	distribution	Distribution. Known only from Waipoua State Forest in Northland area of North Island (Fig. 18 L).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02114DE7FF24FF71FB6F.taxon	etymology	Etymology. After type locality, Waipoua State Forest, a well-known area of occurrence of well-preserved kauri forests.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	description	(Figs 2 N, 5 Z, 16 C, 17 K)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	materials_examined	Type material. Holotype, male (NZAC): “ Lee Valley \ Nel. 15. X. 66 \ A. K. Walker ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	diagnosis	Diagnosis. Body moderately big (TL 4.0 mm), externally resembling A. insularis sp. nov. and other species with crenulate lateral carinae of pronotum. Certain identification possible on the basis of male genitalia: trigonium with triangularly pointed apex, parameroids widened at apices, pala much longer than parameroids.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	description	Description. Male. Body elongated, moderately big (TL 4.0 mm), moderately convex. Body light brown, slightly darker stripe along elytral suture. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, elongated indentation in anterior part of pronotum and round indentation in posterior part, punctation on elytra simple, coarse, punctures separated by 0.5 × diameter of a puncture. Body covered with medium long, erect, yellowish setae, more numerous and ruffled on head and pronotum. Supraantennal ridges well-marked, triangularly elevated. Pronotum widest in the middle of its length, sides curved, lateral carinae crenulate, anterolateral angles produced, triangular, posterolateral angles obtuse. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex delicately truncate. Penis (L 0.72 mm, W 0.27 mm) with rather wide trigonium, tapering towards the end, pointed, almost same length as parameroids; parameroids elongated, rectangular, apices wide, apical part covered with punctures; pala oblong oval, ca. 3 × longer than parameroids. Tegmen (L 0.74 mm, W 0.40 mm) with wide triangular parameres, sparsely punctured, apices rounded, with numerous minute denticles at apices. Female unknown. Measurements and ratios. Male (n = 1) TL 4.00 mm, PL 0.70 mm, PW 1.13 mm, EL 3.25 mm, EW 2.00 mm, TL / EW 2.0, PW / PL 1.6, EL / EW 1.6, EL / PL 4.6, EW / PW 1.8.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	distribution	Distribution. Known only from the type locality in Nelson area of South Island (Fig. 18 L).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBA02104DE7FAA4FB82FF52.taxon	etymology	Etymology. The species is dedicated to Anette K. Walker, a New Zealand entomologist, collector of the holotype of A. walkerae sp. nov. and many other specimens examined during our studies.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	description	(Figs 2 O – P, 5 AA – BB, 8 O, 10 O, 16 D, 17 G)	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	materials_examined	Type material. Holotype, male (CMNC): “ N. Zealand, WD \ Westland N. P. \ Franz Josef Glacier \ Jan. 2, 1984 \ L. Masner ”; paratype, male (CMNC): same data as holotype; 1 male (CMNC): “ N. Zealand, WD \ Lake Mahinapua \ 10 km S. Hokitiwa \ Dec. 30, 1983 \ L. Masner ”; 3 females (LUNZ): “ NEW ZEALAND, WD \ Westland NP \ L. Wombat track \ 180 - 240 m \ 11. i. 1982 ”, “ J. W. Early \ swept ferns in \ podocarp / broad \ leaf forest ”.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	diagnosis	Diagnosis. Body moderately big (TL 4.2 – 4.9 mm), central portion of pronotal disc with normal, not granulate punctation in males, and almost impunctate in females, dorsum covered with erect setae. Penis similar like in A. montana, bursal sclerite with paired elongated triangular sclerites.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	description	Description. Male. Body elongated, medium sized (TL 4.2 – 4.9 mm), convex. Body yellowish brown, elytra often with dark suture. Punctation on head and pronotum granulate, strong and dense, punctures doughnut-shaped, touching each other, punctation flattened and sparse in central portion of disc, separated by ca. 0.5 diameter; punctation on elytra simple, coarse, punctures separated by ca. 0.5 × diameter of a puncture. Body covered with long, erect setae. Supraantennal ridges well-marked, elevated. Pronotum widest slightly before middle, sides bisinuate, anterolateral angles produced outwards, pointed, triangular, posterolateral angles right angled. Elytra without carinae, humeri well-marked. Ventrite 5 with regularly rounded apex. Penis (L 0.76 mm, W 0.29 mm) moderately elongated, trigonium narrow, rectangular, with rounded apex and parallel sides, slightly shorter than parameroids; parameroids elongated, apices pointing outwards, covered with numerous punctures, inner edge slightly crenulate; pala triangular, about the same length as parameroids. Tegmen (L 0.63 mm, W 0.41 mm) moderately wide, rectangular, parameres slightly tapered, subtriangular, with scarce punctures and very few delicate setae on the inner edge. Female. Body relatively large (TL 4.80 – 4.85 mm), brachypterous. Supraantennal ridges well-marked, elevated. Central portion of pronotal disc smooth, with only sparse punctures, punctures separated by ca. 4.0 × diameter of a puncture, interspaces with subtle secondary punctation, consisting of minute punctures, punctation of elytra coarser than in males, punctures separated by 0.3 × diameter of a puncture. Base of elytra with concavity fitting pronotum but not as well developed as in A. villosa. Ventrite 5 subtriangular, with semicircularly emarginated apex. Lateral portions of ventrite 4 with rather big, oval setal patches. Bursal sclerite with two triangular sclerites (L 0.17 mm) consisting of two small sclerites hooked in posterior portion. Measurements and ratios. Males (n = 3) TL 4.15 – 4.90 mm (4.62 mm), PL 0.75 – 1.0 mm (0.87 mm), PW 1.15 – 1.55 mm (1.40 mm), EL 3.35 – 4.0 mm (3.77 mm), EW 1.95 – 2.35 mm (2.17 mm), TL / EW 2.09 – 2.18 (2.16), PW / PL 1.53 – 1.76 (1.65), EL / EW 1.68 – 1.82 (1.77), EL / PL 3.95 – 4.71 (4.59), EW / PW 1.47 – 1.70 (1.58). Females (n = 2) TL 4.80 – 4.85 mm (4.83 mm), PL 0.90 – 0.95 mm (0.93 mm), PW 1.60 – 1.65 mm (1.63 mm), EL 3.85 – 3.90 mm (3.88 mm), EW 2.30 mm, TL / EW 2.09 – 2.11 (2.10), PW / PL 1.74 – 1.78 (1.76), EL / EW 1.67 – 1.70 (1.68), EL / PL 4.05 – 4.33 (4.19), EW / PW 1.39 – 1.44 (1.42).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	distribution	Distribution. Restricted to Westland area of South Island (Fig. 18 L).	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
A0384152FFBB02104DE7FEF9FACEF9FA.taxon	etymology	Etymology. Specific epithet refers to the area where the species occurs — Westland area of South Island.	en	Kiałka, Agata, Ruta, Rafał (2022): Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae). Zootaxa 5174 (4): 401-443, DOI: 10.11646/zootaxa.5174.4.6
