taxonID	type	description	language	source
03A1D418FF9A5A05FCD0F9B9FAFBFE27.taxon	description	(TABLES 2, 3; FIGS 1 – 4)	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9A5A05FCD0F9B9FAFBFE27.taxon	materials_examined	Material examined: Seven animals and four eggs. Specimens mounted on microscope slides in Hoyer’s medium (four animals + four eggs), processed for DNA sequencing (three animals). Population locality: 78 ° 40 ’ 33 ’’ N, 16 ° 38 ’ 49 ’’ E; 208 m a. s. l.: Norway, Svalbard, Fortet; moss on soil. Specimen depositories: Four animals (slides: NO. 393.393.2 - 4) and four eggs (slide: NO. 393.01) are deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Kraków, Poland.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9A5A05FCD0F9B9FAFBFE27.taxon	description	Description of the Norwegian population from Spitsbergen Animals (measurements and statistics in Table 2): Body transparent in smaller individuals (juveniles) and whitish in adults, after fixation in Hoyer’s medium transparent (Fig. 1 A). Eyes present in all specimens, visible after mounting in Hoyer’s medium. Cuticular pores (0.7 – 1.2 µm in diameter) present, clearly visible under PCM (Fig. 1 B – E) and scattered randomly on the entire body cuticle, including the external and internal surface of all legs. Patches of fine granulation present on internal and external surfaces of all legs I – III, as well as on legs IV and visible clearly in PCM (Fig. 1 C – E). A pulvinus present on the internal surfaces of legs I – III (Fig. 1 D). Granulation on legs IV is visible as a single large granulation patch on dorsal and lateral leg surfaces (Fig. 1 E). Claws slender, with flat and wide common tract, beginning with a visible stalk that connects the claws to the wide lunulae and ending with elongated branches (especially the primary branch; Fig. 2 A, B). Primary branches with distinct accessory points, visible in PCM (Fig. 2 A, B). Lunulae I – III smooth (Fig. 2 A), whereas lunulae IV with clear dentation (Fig. 2 B, D). A single continuous cuticular bar and double muscle attachments visible on each leg I – III (Fig. 2 C). Mouth anteroventral with ten peribuccal lamellae. Bucco-pharyngeal apparatus of the Macrobiotus - type (Fig. 3 A). Under PCM, oral cavity armature extremely reduced to only one large tooth present in the dorsal portion of the third band of teeth, whereas other bands of teeth invisible or absent (Fig. 3 A, B). Pharyngeal bulb spherical, with triangular apophyses, cuticular spikes, two rod-shaped macroplacoids (macroplacoid sequence: 2 <1) and a triangular small microplacoid (Fig. 3 A, D – E). The first macroplacoid with a weak central constriction, whereas the second macroplacoid is weakly subterminally constricted (Fig. 3 D, E). Eggs (measurements and statistics in Table 3): Eggs laid freely, whitish, spherical or slightly oval (Fig. 4 A, B). The spaces between processes are small, the surface between processes is of the persimilis - type, i. e. with the continuous smooth chorion, with no pores visible (Fig. 4 A, B). Egg processes single-walled (without reticulation caused by the labyrinthine layer) with dome-shaped basal part and thinner and elongated distal portions (Fig. 4 C – H). Internal septa are sometimes visible between the basal and the distal portion of the process in PCM (Fig. 4 H). The basal portions of the processes are pierced by pores that are arranged alternately with dark thickenings and around the process base (Fig. 4 A, B). The apical parts of the processes are flat but devoid of terminal discs, and are covered with short, thin and flexible filaments (Fig. 4 C – H).	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9A5A05FCD0F9B9FAFBFE27.taxon	biology_ecology	Reproduction: The population is dioecious (the examination of specimens freshly mounted in Hoyer’s medium revealed testis filled with spermatozoa), but no secondary sexual dimorphism has been observed. DNA sequences: All obtained DNA sequences were represented by a single haplotype per each marker: 18 S rRNA: MZ 463668, MZ 463669, MZ 463670. 28 S rRNA: MZ 463674, MZ 463675, MZ 463676. ITS 2: MZ 463656, MZ 463657, MZ 463658. COI: MZ 460999, MZ 461000, MZ 461001.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	description	(TABLES 4, 5; FIGS 5 – 11) Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org: act: 8 A 12 E 93 C- 8729 - 4 B 13 - BD 36 - FC 507 DD 117 D 3	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	materials_examined	Material examined: Altogether 110 animals and 78 eggs. Specimens mounted on microscope slides in Hoyer’s medium (83 animals + 68 eggs), fixed on SEM stubs (20 animals + ten eggs + four buccal apparatuses), processed for DNA sequencing (three animals).	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	etymology	Etymology: The new subspecies is named after Greenland (from Danish Grønland), the territory where it was discovered.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	materials_examined	Type locality: 69 ° 15 ’ 17 ’’ N, 53 ° 30 ’ 46 ’’ W; 30 m a. s. l.: western coast of Greenland, Disko Island, Østerlien; moss on rock. Type depositories: Altogether 83 animals [slides: GL. 018. 2 – 3, 9 – 17, SEM stubs: 9.06, 12.15 (buccal apparatus), 16.19] and 68 eggs (slides: GL. 018. 1, 4 – 8, SEM stub: 16.19) are deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Kraków, Poland.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	description	Description of the new subspecies Animals (measurements and statistics in Table 4): Body colourless in juveniles and whitish in adults, after fixation in Hoyer’s medium transparent (Fig. 5 A). Eyes present, visible also after mounting the specimens on permanent slides in Hoyer’s medium. The entire cuticle covered with granulation visible in both PCM and SEM, arranged densely on the dorsum, and less densely on the venter and legs (Figs 5 B – G, 6 A, C, D, F, G, I). Only in some specimens the cuticular granulation can be less evident under PCM (Fig. 5 D). Oval cuticular pores present (0.5 – 1.4 µm in diameter) (Fig. 5 E – G). Patches of dense granulation present on internal and external surface of all legs I – III, as well as on legs IV and clearly visible (Fig. 6 A, B, D, E). A pulvinus present on the internal surface of legs I – III (Fig. 6 D, E). Granulation on legs IV is visible as a single large patch on dorsal and lateral leg surfaces (Fig. 6 G – H). Claws slender, with flat and wide common tract, beginning with a visible stalk that connects the claws to the wide lunulae and ending with elongated branches (especially the primary branch; Fig. 7 A – E). Primary branches with distinct accessory points, visible in PCM and SEM (Fig. 7 A – E). Lunulae I – III smooth (Fig. 7 A, C, D), whereas lunulae IV with clear dentation (Fig. 7 B, E). A single continuous cuticular bar (Fig. 7 A) and double muscle attachments visible on each leg I – III (Fig. 7 A, C, D). Mouth anteroventral with ten peribuccal lamellae. Bucco-pharyngeal apparatus of the Macrobiotus - type (Figs 8 A, 9 A). Oral cavity armature extremely reduced to only one large tooth present in the dorsal portion of the third band of teeth, whereas other bands of teeth are absent (Figs 8 A – D, 9 F). Pharyngeal bulb spherical, with triangular apophyses, cuticular spikes, two rod-shaped macroplacoids (macroplacoid sequence: 2 <1) and a triangular small microplacoid (Figs 8 A, E, 9 A, C, D). The first macroplacoid exhibits strong central constriction, whereas the second macroplacoid is subterminally constricted (Figs 8 E, 9 C, D). Two globular protuberances are present on the ventral side of the buccal apparatus, one on the buccal crown anteriorly to the beginning of ventral lamina, second posteriorly to the ventral lamina at the level of stylet support insertion point (character visible only under SEM; Fig. 9 A, B, E). Eggs (measurements and statistics in Table 5): Eggs laid freely, whitish, spherical or slightly oval (Figs 10 A – D, 11 A). The spaces between the processes are small and the surface of the egg between the processes is continuous and smooth, without any pores or reticulum, i. e. persimilis - type (Figs 10 A – D, 11 A – D). Between the processes on the egg surface, lightrefracting dots are usually visible in PCM, resembling micropores (Fig. 10 A – D). Egg processes single-walled (without reticulation caused by labyrinthine layer) with dome-shaped basal part with distal part being thinner and elongated (Figs 10 E – P, 11 A – D). Internal septa are sometimes visible between basal and distal portion of the process in PCM (Fig. 10 E – P). The basal portions of the processes are pierced by pores of uniform size (1.1 – 1.8 µm in diameter) that are arranged alternately with dark thickenings around the process base (Figs 10 A – D, 11 B – D). In SEM, a reticulate internal structure is visible inside the pores and it seems to be a remnant of the reduced labyrinthine layer (Fig. 11 B – D). The apical parts of the processes are flat but devoid of terminal discs and are covered with short, thin and flexible filaments (Figs 10 E – P, 11 E, F).	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	biology_ecology	Reproduction: The population is dioecious (the examination of specimens freshly mounted in Hoyer’s medium revealed testes filled with spermatozoa), but no secondary sexual dimorphism has been observed. DNA sequences: All obtained DNA sequences were represented by a single haplotype per each marker: 18 S rRNA: MZ 463662, MZ 463663, MZ 463664. 28 S rRNA: MZ 463677, MZ 463678, MZ 463679. ITS 2: MZ 463653, MZ 463654, MZ 463655. COI: MZ 461005, MZ 461006, MZ 461007.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF9C5A09FCC6FDF6FAE9FB9C.taxon	diagnosis	Differential diagnosis: Macrobiotus a. groenlandicus, known only from its locus typicus in Disko Island, Greenland, shares with M. a. ariekammensis the elongated primary branches of all claws, only one tooth in the third band of teeth in the oral cavity, and single-layer egg processes surrounded by a crown of pores and thickenings around their bases. However, it differs from M. a. ariekammensis, which is known only from a few localities in Svalbard (Norway) and Poland, by: the presence of a strong pronounced constriction in the first macroplacoid (first macroplacoid weakly constricted in M. a. ariekammensis), the presence of light-refracting dots resembling micropores on the egg surface (egg surface smooth in M. a. ariekammensis) and by the presence of fine granulation on the body cuticle visible in PCM and SEM (the body granulation absent or not visible in PCM in M. a. ariekammensis).	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	description	(TABLES 6, 7; FIGS 12 – 18)	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	materials_examined	Material examined: Altogether 66 animals, and 15 eggs. Specimens mounted on microscope slides in Hoyer’s medium (53 animals + ten eggs), fixed on SEM stubs (ten + five), processed for DNA sequencing (three animals). Population locality: 41 ° 32 ’ 37.98 ’’ N, 75 ° 10 ’ 2.28 ’’ E; 2288 m a. s. l.: Kyrgyzstan, Chui, Kegeti, moss on rock. Specimens depositories: Altogether 53 animals (slides: KG. 062.006. 1, 9 – 14, SEM stub: 18.07) and ten eggs (slides: KG. 062. 5 – 8, SEM stub: 18.07) are deposited at the Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Kraków, Poland.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	description	Description of the Kyrgyz Republic population Animals (measurements and statistics in Table 6): Body whitish in adults and colourless in smaller individuals, after fixation in Hoyer’s medium transparent (Fig. 12 A). After mounting in Hoyer’s medium eyes present in all specimens. Small, oval pores (0.5 – 0.8 µm in diameter), visible under PCM and SEM (Fig. 12 B, C), scattered randomly on the entire body cuticle, including the external and internal surface of all legs (Fig. 13 A – F). Extremely fine body granulation (c. 60 nm in diameter), visible only in SEM, present on the entire dorsocaudal cuticle (Fig. 12 C). Patches of dense granulation present on the internal and external surfaces of all legs I – III and clearly visible both in PCM and SEM (Fig. 13 A – D). A cuticular bulge, resembling a pulvinus, is present on the internal surfaces of legs I – III (Fig. 13 C, D). Cuticular granulation on legs IV present and always clearly visible both in PCM and SEM (Fig. 13 E, F). Claws slender, with flat and wide common tract, beginning with an evident stalk that connects the claws to the wide lunulae and ending with extremely elongated branches (especially the primary branch; Fig. 14 A, B, D, E). Primary branches with indistinct accessory points, barely visible in PCM, but clearly visible in SEM (Fig. 14 A, B, D, E). Lunulae I – III smooth (Fig. 14 A, D), whereas lunulae IV with clear dentation (Fig. 14 B, E). Mouth anteroventral with ten peribuccal lamellae (Fig. 16 A, B). Bucco-pharyngeal apparatus of the Macrobiotus - type (Fig. 15 A). Under PCM, only the second and third band of teeth visible, with the second band being faintly marked (Fig. 15 B, C). However, under SEM, all of the three bands of teeth are visible, with the first band being situated at the base of peribuccal lamellae and composed of several irregular rows of small granular teeth surrounding the oral cavity (Fig. 16 A, C). The second band of teeth is situated between the ring fold and the third band of teeth, and is comprises of small cones, barely visible in PCM (Figs 15 B, 16 B; note: in Fig 16 B, only distal portion of these teeth are visible from behind the ring fold; due to unsuitable positioned specimen it was impossible to get better image in SEM). The teeth of the third band are located within the posterior portion of the oral cavity, between the second band of teeth and the buccal tube opening (Figs 15 B – D, 16 A, B). The third band of teeth is discontinuous and divided into a dorsal and a ventral portion. Under PCM, the dorsal teeth form a transversal ridge weakly divided into two granular teeth, whereas the ventral teeth are smaller and faintly visible as two separate lateral transverse ridges with granular / roundish thickening at their medial extremities (Fig 15 B – D). In SEM, both the dorsal and the ventral portion of the third band of teeth are visible as one fused ridge with two evident teeth extending from the medial portion of the ridge (Fig 16 A, B). Pharyngeal bulb spherical, with triangular apophyses, cuticular spikes, two rodshaped macroplacoids (macroplacoid sequence: 2 <1) and a triangular small microplacoid (Fig. 16 A, E). The first macroplacoid exhibits weak central constriction, whereas the second macroplacoid is sub-terminally and weakly constricted (Figs 15 E). Eggs (measurements and statistics in Table 7): Eggs laid freely, whitish, spherical or slightly oval (Figs 17 A, B, 18 A). Although the spaces between processes are small, the surface between processes is of the persimilis - type, i. e. with the continuous smooth chorion, with few, randomly distributed pores (Figs 17 A, B, 18 B – D). Egg processes single-walled (without reticulation caused by labyrinthine layer) with domeshaped basal part and rigid spine-like distal part (Figs 17 A – F, 18 A – F). In PCM, the basal and distal portions are clearly separated from with single internal septum (Fig 17 C – F). The bases of egg processes are pierced with pores of uniform size (0.3 – 0.7 µm in diameter), distributed evenly around the base and most often arranged in two rows (Figs 17 A, B, 18 B – F). In PCM, short, dark thickenings are sometimes visible around the process bases below or at the same level as the lower ring of pores (Fig 17 A, B). The apical part of the processes is devoid of terminal discs and is covered with short, thin and flexible filaments (Figs 17 C – F, 18 A – F).	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	biology_ecology	Reproduction: The population is dioecious (the examination of specimens freshly mounted in Hoyer’s medium revealed testes filled with spermatozoa), but no secondary sexual dimorphism has been observed.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	description	DNA sequences: All obtained DNA sequences were represented by a single haplotype per each marker: 18 S rRNA: MZ 463665, MZ 463666, MZ 463667. 28 S rRNA: MZ 463671, MZ 463672, MZ 463673. ITS 2: MZ 463659, MZ 463660, MZ 463661. COI: MZ 461002, MZ 461003, MZ 461004.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
03A1D418FF905A17FCFBFB4FFC02FE1A.taxon	discussion	PHYLOGENY The phylogenetic reconstruction (Fig. 19) shows three well-supported distinct lineages constituting three separate genera within superclade I (sensu Stec et al., 2021 a) of the family Macrobiotidae: the clade comprising Macrobiotus species, and further two monophyletic groups: one corresponding to the genus Mesobiotus Vecchi et al., 2016, and the other representing Sisubiotus Stec et al., 2021 a (Fig. 19). Macrobiotus is divided into three well-supported subclades: A, B and C, sensu Stec et al. (2021 a). All of the three newly found populations investigated in this study, M. a. ariekammensis, M. a. groenlandicus and M. kirghizicus, are nested in subclade A, which contains species of the Macrobiotus hufelandi morphogroup sensu Stec et al. (2021 a) and Macrobiotus basiatus Nelson et al., 2020, which exhibits unique egg morphology. Subclade B comprises three species complexes delineated by Stec et al. (2021 a). As in Stec et al. (2021 a) and Vecchi & Stec (2021), the Macrobiotus pallari complex and the Macrobiotus pseudohufelandi complex are monophyletic also in the present study (Fig. 19). However, the Macrobiotus persimilis complex, which was monophyletic in the two earlier studies, appears to be paraphyletic in the current analysis (Fig. 19). Thus, further studies are needed to clarify the phyletic character of the latter species complex. Subclade C comprises species of the Macrobiotus hufelandi morphogroup. SPECIES DELIMITATION AND GENETIC DISTANCES The PTP analysis identified 49 and 55 putative species in ML and BI approach, respectively. The ASAP analysis, on the other hand, identified 48 putative species. These results are in line with the general inspection of the tree terminals and the morphological information that would suggest also 48 species among the ingroup taxa. However, for two out of the three newly found populations analysed in this study, both PTP approaches were not congruent with ASAP results. The PTP approaches indicated that M. a. ariekammensis and M. a. groenlandicus constitute a single species, whereas the ASAP analysis identified them as separate entities. Uncorrected pairwise distances between the three newly found populations analysed in this study are as follows: • 18 S rRNA: 0.2 % for M. a. ariekammensis and M. a. groenlandicus; 0.1 % for M. a. ariekammensis and M. kirghizicus; 0.3 % for M. a. groenlandicus and M. kirghizicus. • 28 S rRNA: 0.1 % for M. a. ariekammensis and M. a. groenlandicus; 0.3 % for M. a. ariekammensis in PCM (E) and SEM (F). Filled flat arrowheads indicate granulation patch on the external leg surface, empty indented arrowheads indicate cuticular bulge (pulvini), filled indented arrowhead indicates cuticular bar, empty flat arrowheads indicate granulation patch on the internal leg surface. Scale bars in µm. and M. kirghizicus; 0.1 % for M. a. groenlandicus and M. kirghizicus. • ITS 2: 0.3 % to 0.8 % for M. a. ariekammensis and M. a. groenlandicus; 6.1 % for M. a. ariekammensis and M. kirghizicus; 6.3 % for M. a. groenlandicus and M. kirghizicus. • COI: 3.3 % for M. a. ariekammensis and M. a. groenlandicus; 16.3 % for M. a. ariekammensis and M. kirghizicus; 16.4 % for M. a. groenlandicus and M. kirghizicus. Given the discrepancies between the PTP and ASAP species delineation results, shallow genetic divergence and low p - distances in COI and ITS 2 between M. a. ariekammensis and M. a. groenlandicus, we interpreted the morphological differences between the two taxa as intraspecific variability, hence the later taxon is described here as a subspecies rather than a separate species.	en	Stec, Daniel, Vončina, Katarzyna, Kristensen, Reinhardt Møbjerg, Michalczyk, Łukasz (2022): The Macrobiotus ariekammensis species complex provides evidence for parallel evolution of claw elongation in macrobiotid tardigrades. Zoological Journal of the Linnean Society 195: 1067-1099, DOI: 10.1093/zoolinnean/zlab101
