taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994458/files/figure.png	http://doi.org/10.5281/zenodo.6994458	Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23).	Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23).	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994460/files/figure.png	http://doi.org/10.5281/zenodo.6994460	Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape.	Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994466/files/figure.png	http://doi.org/10.5281/zenodo.6994466	Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina.	Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994470/files/figure.png	http://doi.org/10.5281/zenodo.6994470	Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse	Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994472/files/figure.png	http://doi.org/10.5281/zenodo.6994472	Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole.	Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994474/files/figure.png	http://doi.org/10.5281/zenodo.6994474	Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere.	Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994482/files/figure.png	http://doi.org/10.5281/zenodo.6994482	Figure 11. Couplet 1: mesonotum with (1-1) or without (1’-1) transverse ridge; abdominal segment IV with (1-2) or without (1’-2) cinctus; prora large (1-3) or small (1’-3). (1-1) †G. ufv-05, ANTWEB103848 (J. Chaul, AntWeb), mesosoma in profile view. (1’-1, 1’-3) †G. indet., JWJ-Bu19 (V. Perrichot, AntWeb), mesosoma in profile view. (1-2) †G. sternorhabda sp. nov. paratype, UFV-LABECOL-009656, metasoma in ventrolateral view. (1’-2) †G. gracilis, CASENT0741232, metasoma in lateral view. (1-3) †G. pilosa holotype, JZC-Bu225 (P. Barden, AntWeb), anterior metasoma in profile view.	Figure 11. Couplet 1: mesonotum with (1-1) or without (1’-1) transverse ridge; abdominal segment IV with (1-2) or without (1’-2) cinctus; prora large (1-3) or small (1’-3). (1-1) †G. ufv-05, ANTWEB103848 (J. Chaul, AntWeb), mesosoma in profile view. (1’-1, 1’-3) †G. indet., JWJ-Bu19 (V. Perrichot, AntWeb), mesosoma in profile view. (1-2) †G. sternorhabda sp. nov. paratype, UFV-LABECOL-009656, metasoma in ventrolateral view. (1’-2) †G. gracilis, CASENT0741232, metasoma in lateral view. (1-3) †G. pilosa holotype, JZC-Bu225 (P. Barden, AntWeb), anterior metasoma in profile view.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFE8AA26D2FEFE34FC09947B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994480/files/figure.png	http://doi.org/10.5281/zenodo.6994480	Figure 10. Features hidden by appendages, including artefacts caused by close proximity of limbs. †Gerontoformica gracilis: A, B, D, adult, CASENT0741232; C, pupa (CASENT0741231). A, C, mesosoma in lateral view with the legs distal to the coxae digitally removed, with the exception of the fore leg of (C), which is not cropped from the render. B, D, head in facial view with the scapes digitally rendered (B) and removed (D). Abbreviations: mspa, mesopectal artefact; sca, scapal artefact.	Figure 10. Features hidden by appendages, including artefacts caused by close proximity of limbs. †Gerontoformica gracilis: A, B, D, adult, CASENT0741232; C, pupa (CASENT0741231). A, C, mesosoma in lateral view with the legs distal to the coxae digitally removed, with the exception of the fore leg of (C), which is not cropped from the render. B, D, head in facial view with the scapes digitally rendered (B) and removed (D). Abbreviations: mspa, mesopectal artefact; sca, scapal artefact.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994458/files/figure.png	http://doi.org/10.5281/zenodo.6994458	Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23).	Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23).	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994460/files/figure.png	http://doi.org/10.5281/zenodo.6994460	Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape.	Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994476/files/figure.png	http://doi.org/10.5281/zenodo.6994476	Figure 8. Photomicrographs of †G. gracilis showing setation and certain key features. Specimen CASENT0741232. A, body in ventrolateral anterior oblique view. B, body in dorsolateral posterior oblique view. C, detail of head and prothorax in ventrolateral anterior oblique view. D, detail of mid and hind legs in ventrolateral anterior oblique view. Abbreviations: AII–VII, abdominal segments II–VII; AVsp, spiracle of abdominal segment V; ar, arolium; ca, calcar; cav, velum of calcar; ce, compound eye; fc, frontal carina; mc, cranial condyle of mandible; md, mandible; mspfm, prefemur of mesothoracic leg; msptc, tooth of mesothoracic leg pretarsal claw; mstbspa, anterior tibial spur of mesothoracic leg; mstbspp, posterior tibial spur of mesothoracic leg; mtpfm, prefemur of metathoracic leg; mtplglvf, metapleural gland ventral flange; pd, pedicel; pdcs, prodisticoxal suture; pl, labial palp; plo, plantar lobe; pm, maxillary palp; pnt, pronotum; ppd, propodeum; ppdsp, propodeal spiracle; ppl, propleuron; pr, prora; ptsp, spiracle of the petiole; sc, scape.	Figure 8. Photomicrographs of †G. gracilis showing setation and certain key features. Specimen CASENT0741232. A, body in ventrolateral anterior oblique view. B, body in dorsolateral posterior oblique view. C, detail of head and prothorax in ventrolateral anterior oblique view. D, detail of mid and hind legs in ventrolateral anterior oblique view. Abbreviations: AII–VII, abdominal segments II–VII; AVsp, spiracle of abdominal segment V; ar, arolium; ca, calcar; cav, velum of calcar; ce, compound eye; fc, frontal carina; mc, cranial condyle of mandible; md, mandible; mspfm, prefemur of mesothoracic leg; msptc, tooth of mesothoracic leg pretarsal claw; mstbspa, anterior tibial spur of mesothoracic leg; mstbspp, posterior tibial spur of mesothoracic leg; mtpfm, prefemur of metathoracic leg; mtplglvf, metapleural gland ventral flange; pd, pedicel; pdcs, prodisticoxal suture; pl, labial palp; plo, plantar lobe; pm, maxillary palp; pnt, pronotum; ppd, propodeum; ppdsp, propodeal spiracle; ppl, propleuron; pr, prora; ptsp, spiracle of the petiole; sc, scape.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994478/files/figure.png	http://doi.org/10.5281/zenodo.6994478	Figure 9. Exceptional soft tissue preservation. The adult specimen of †G. gracilis displays exceptional preservation of soft tissues, including identifiable neuropils (A), countable glomeruli of the antennal lobe (A’), the digestive tract, glands, and muscle (B), down to the scale of individual muscular cross-striations (B’).	Figure 9. Exceptional soft tissue preservation. The adult specimen of †G. gracilis displays exceptional preservation of soft tissues, including identifiable neuropils (A), countable glomeruli of the antennal lobe (A’), the digestive tract, glands, and muscle (B), down to the scale of individual muscular cross-striations (B’).	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994480/files/figure.png	http://doi.org/10.5281/zenodo.6994480	Figure 10. Features hidden by appendages, including artefacts caused by close proximity of limbs. †Gerontoformica gracilis: A, B, D, adult, CASENT0741232; C, pupa (CASENT0741231). A, C, mesosoma in lateral view with the legs distal to the coxae digitally removed, with the exception of the fore leg of (C), which is not cropped from the render. B, D, head in facial view with the scapes digitally rendered (B) and removed (D). Abbreviations: mspa, mesopectal artefact; sca, scapal artefact.	Figure 10. Features hidden by appendages, including artefacts caused by close proximity of limbs. †Gerontoformica gracilis: A, B, D, adult, CASENT0741232; C, pupa (CASENT0741231). A, C, mesosoma in lateral view with the legs distal to the coxae digitally removed, with the exception of the fore leg of (C), which is not cropped from the render. B, D, head in facial view with the scapes digitally rendered (B) and removed (D). Abbreviations: mspa, mesopectal artefact; sca, scapal artefact.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994482/files/figure.png	http://doi.org/10.5281/zenodo.6994482	Figure 11. Couplet 1: mesonotum with (1-1) or without (1’-1) transverse ridge; abdominal segment IV with (1-2) or without (1’-2) cinctus; prora large (1-3) or small (1’-3). (1-1) †G. ufv-05, ANTWEB103848 (J. Chaul, AntWeb), mesosoma in profile view. (1’-1, 1’-3) †G. indet., JWJ-Bu19 (V. Perrichot, AntWeb), mesosoma in profile view. (1-2) †G. sternorhabda sp. nov. paratype, UFV-LABECOL-009656, metasoma in ventrolateral view. (1’-2) †G. gracilis, CASENT0741232, metasoma in lateral view. (1-3) †G. pilosa holotype, JZC-Bu225 (P. Barden, AntWeb), anterior metasoma in profile view.	Figure 11. Couplet 1: mesonotum with (1-1) or without (1’-1) transverse ridge; abdominal segment IV with (1-2) or without (1’-2) cinctus; prora large (1-3) or small (1’-3). (1-1) †G. ufv-05, ANTWEB103848 (J. Chaul, AntWeb), mesosoma in profile view. (1’-1, 1’-3) †G. indet., JWJ-Bu19 (V. Perrichot, AntWeb), mesosoma in profile view. (1-2) †G. sternorhabda sp. nov. paratype, UFV-LABECOL-009656, metasoma in ventrolateral view. (1’-2) †G. gracilis, CASENT0741232, metasoma in lateral view. (1-3) †G. pilosa holotype, JZC-Bu225 (P. Barden, AntWeb), anterior metasoma in profile view.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994464/files/figure.png	http://doi.org/10.5281/zenodo.6994464	Figure 3. Summary of the evidence for eusocial evolution in stem groups. A, overview of the phylogenetic relationships of the total clade Formicidae, with key biological traits mapped. F+, winged females; f-, wingless females; D, developmental simplification of the mesosoma; and B*, brood care. Solid squares, trait confirmed as present; empty squares, trait confirmed as absent; half-filled squares, trait confirmed as variable; question marks, state uncertain; black square with asterisk, new evidence provided in the present study. B, exemplars of females with the full and complex complement of flight sclerites, namely the mesoscutum (green), mesoscutellum (blue), and metanotum (magenta). C, exemplars of wingless †Zigrasimeciinae demonstrating independent convergence on the extremely simplified mesosomal form of various crown Formicidae, illustrated in this case by Discothyrea (Proceratiinae). D, exemplars of theAntennoclypeata, showing a similar degree of sclerite reduction between †Brownimecia and crown Formicidae. Notes: The topology in 1A is summarized from Johnson et al. (2013), Barden & Grimaldi (2016)and Boudinot et al.(2020a). Images for rows 1B, C were downloaded from AntWeb(2021); taxon names, imagers and unique specimen identifiers are listed from left to right, top to bottom are as follows: †Zigrasimecia tonsora Barden & Grimaldi, 2013 (P. Barden, ANTWEB1008098); †Dhagnathos autokrator Perrichot et al., 2020 (V.Perrichot, FANTWEB00022); †Linguamyrmex rhinocerus Miao & Wang, 2019 (V. Perrichot, FANTWEB00016); †Protozigrasimecia chauli Cao et al., 2020c (H. Cao, FANTWEB00051); †Zigrasimecia ufv-01 (J. Chaul, ANTWEB10141055); Discothyrea bobi Chaul, 2020 (J. Chaul, UFV-LABECOL-000032); †Brachyponera croceicornis (Emery, 1900) (F. Esteves, CASENT0916594); Brachyponera croceicornis (Emery, 1900) (A. Nobile, CASENT0172432); †Brownimecia clavata Grimaldi et al., 1997 (V. Perrichot, AMNH-NJ667).	Figure 3. Summary of the evidence for eusocial evolution in stem groups. A, overview of the phylogenetic relationships of the total clade Formicidae, with key biological traits mapped. F+, winged females; f-, wingless females; D, developmental simplification of the mesosoma; and B*, brood care. Solid squares, trait confirmed as present; empty squares, trait confirmed as absent; half-filled squares, trait confirmed as variable; question marks, state uncertain; black square with asterisk, new evidence provided in the present study. B, exemplars of females with the full and complex complement of flight sclerites, namely the mesoscutum (green), mesoscutellum (blue), and metanotum (magenta). C, exemplars of wingless †Zigrasimeciinae demonstrating independent convergence on the extremely simplified mesosomal form of various crown Formicidae, illustrated in this case by Discothyrea (Proceratiinae). D, exemplars of theAntennoclypeata, showing a similar degree of sclerite reduction between †Brownimecia and crown Formicidae. Notes: The topology in 1A is summarized from Johnson et al. (2013), Barden & Grimaldi (2016)and Boudinot et al.(2020a). Images for rows 1B, C were downloaded from AntWeb(2021); taxon names, imagers and unique specimen identifiers are listed from left to right, top to bottom are as follows: †Zigrasimecia tonsora Barden & Grimaldi, 2013 (P. Barden, ANTWEB1008098); †Dhagnathos autokrator Perrichot et al., 2020 (V.Perrichot, FANTWEB00022); †Linguamyrmex rhinocerus Miao & Wang, 2019 (V. Perrichot, FANTWEB00016); †Protozigrasimecia chauli Cao et al., 2020c (H. Cao, FANTWEB00051); †Zigrasimecia ufv-01 (J. Chaul, ANTWEB10141055); Discothyrea bobi Chaul, 2020 (J. Chaul, UFV-LABECOL-000032); †Brachyponera croceicornis (Emery, 1900) (F. Esteves, CASENT0916594); Brachyponera croceicornis (Emery, 1900) (A. Nobile, CASENT0172432); †Brownimecia clavata Grimaldi et al., 1997 (V. Perrichot, AMNH-NJ667).	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994474/files/figure.png	http://doi.org/10.5281/zenodo.6994474	Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere.	Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
038287F4FFEFAA3ED106FB00FCEA9156.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6994472/files/figure.png	http://doi.org/10.5281/zenodo.6994472	Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole.	Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole.	2022-02-05	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei		Zenodo	biologists	Boudinot, Brendon E.;Richter, Adrian;Katzke, Julian;Chaul, Júlio C. M.;Keller, Roberto A.;Economo, Evan P.;Beutel, Rolf Georg;Yamamoto, Shûhei			
