taxonID	type	description	language	source
13904BA524F85CCBA8D4BE005104B34C.taxon	description	Figs 2 A, 3, 4, 5	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
13904BA524F85CCBA8D4BE005104B34C.taxon	materials_examined	Material examined. Holotype: Thailand - Yala Province • ♂; Betong District, hill in evergreen forest, on forest floor; 1440 m a. s. l.; 5 ° 55 ' N, 101 ° 26 ' E; 22 May 2021; Wisut Sittichaya leg.; CUMZ. Paratype: Thailand - Yala Province • ♀; same District, elfin montane forest (Malaya Phytochorion province); 1430 m a. s. l.; 25 May 2022; Wisut Sittichaya leg.; CUMZ.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
13904BA524F85CCBA8D4BE005104B34C.taxon	diagnosis	Diagnosis. This new species seems to be particularly similar to M. siveci Mrsic, 1996, with which it shares most of the gonopodal characters. It differs from M. siveci by the wider body, 2.7 - 3.2 mm (vs smaller, 1.2 mm), the colour pattern which is uniformly red brown with lighter red brown paraterga (Fig. 3 A-F) (vs a light brown body with the collum and caudal edges of metazonae margined darker brown; Fig. 1 A), as well as the pleurosternal carinae present until segment 11 (vs until segment 5), the sternal lobe between ♂ coxae 4 with a pair of small cones laterally near base (Fig. 3 E, H, I) (vs absent, Fig. 1 D), and the tip of the gonopod with a denticulate margin (Figs 4 A, B, 5 C, D) (vs smooth and rounded; Fig. 1 E-G).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
13904BA524F85CCBA8D4BE005104B34C.taxon	description	Description. Length 29.3 (♂) or 36.2 mm (♀), width of midbody pro- and metazonae 2.1 and 2.7 mm (♂) or 2.7 and 3.2 mm (♀), respectively. Colouration of live animal rusty red (Fig. 2 A), edges of paraterga light red brown; antennae dark brownish, legs and venter contrasting light yellow (Fig. 2 A); colouration in alcohol, after one week of preservation, red brown (Fig. 3 A-F); edges of paraterga light red brown, head and antennae brown, legs, venter and a few basal antennomeres contrasting light yellow (Fig. 3 A-G). Clypeolabral region sparsely setose; epicranial suture distinct. Antennae long, extending caudally past metaterga 5 (♂) or metaterga 3 (♀) when stretched dorsally. In width, segment 3 <4 = collum <segment 2 = head <segment 5 <6 - 17, body gently and gradually tapering thereafter. Collum with three transverse rows of setae: 4 + 4 in anterior, 2 + 2 in intermediate, and 3 + 3 in posterior row, all mostly abraded, but still traceable as insertion points; lateral incisions absent; caudal corner of paraterga very broadly rounded, declined ventrad, produced slightly past rear tergal margin (Fig. 3 A, B). Tegument generally smooth and shining, prozonae finely shagreened, metaterga finely leathery and faintly rugulose (Fig. 3 A, C, F), surface below paraterga leathery and rugose (Fig. 3 B, D, E). Postcollum metaterga with two transverse rows of setae traceable at least as insertion points when setae broken off: 2 + 2 in anterior (presulcus) and 3 + 3 in posterior (post-sulcus) row. Tergal setae simple, slender, ca. 1 / 3 as long as metaterga. Axial line barely traceable both on pro- and metazonae. Paraterga rather well developed (Fig. 3 A, C, F), lying rather high (at upper 1 / 3 of body), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; lateral edge without incisions; caudal corner very narrowly rounded, not produced past rear tergal margin except in rings 2 and 3 (Fig. 3 A, B); posterior edge nearly straight. Paraterga 2 broad, anterior edge angular and rounded, lateral edge without incisions (Fig. 3 A). Calluses on paraterga rather narrow, delimited by a sulcus fully on dorsal side and in about posterior 2 / 3 on ventral side; on poreless rings more narrow than on pore-bearing ones in dorsal view (Fig. 3 B, D, E). Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 in front of posterior edge of metaterga. Transverse sulcus usually distinct (Fig. 3 A, C, F), complete on metaterga 5 - 17, narrow, line-shaped, rather deep, not reaching the bases of paraterga, very faintly ribbed at bottom, incomplete and nearly wanting on segment 18. Stricture between pro- and metazona wide, deep, ribbed at bottom down to base of paraterga starting with segment 5 (Fig. 3 A-E, F). Pleurosternal carinae complete crests with a sharp caudal tooth on rings 2 - 4, increasingly reduced and retaining a sharp caudal tooth on rings 5 and 6 thereafter, further retained as a small caudal tooth and increasingly reduced until segment 11, absent from segment 12 on (♂, ♀) (Fig. 3 B, D, E). Epiproct (Fig. 3 E-G) conical, flattened dorsoventrally, with two evident, but small, rounded, apical papillae; tip subtruncate; pre-apical papillae small, but evident, lying close to tip. Paraprocts regularly convex, each with premarginal sulci medially and two pairs of setigerous knobs at medial margin (Fig. 3 G). Hypoproct roundly subtrapeziform, setigerous knobs at caudal edge very small and well-separated (Fig. 3 G). Sterna sparsely setose, shining, cross-impressions shallow, without modifications; a single, linguiform, medially rather deeply notched sternal lobe between ♂ coxae 4, with a pair of small cones laterally near base (Fig. 3 E, H, I). A conspicuous and high ridge present in front of gonopod aperture. Legs long and slender (Fig. 3 B), midbody ones ca. 1.4 - 1.6 (♂) or 1.2 - 1.3 (♀) times as long as body height, without modifications, ♂ tarsal brushes absent. Gonopods (Figs 4 A-D, 5) simple; coxa a little curved caudad, densely setose distoventrally. Prefemur as usual, densely setose, about 1 / 3 as long as femorite + postfemoral part. Femorite rather stout, wider than prefemur or postfemur, slightly expanded distad, suberect, showing a distinct mesal groove / hollow (g), with a sulcus demarcating a postfemoral part; seminal groove running entirely mesally along fermorite, solenomere (sl) flagelliform, almost fully sheathed by solenophore (sph). Lamina medialis (lm) well developed, short and unciform, terminal lobe sheathing the tip of solenomerite. Lamina lateralis (ll) elevated, prominent, stout, expanded apically, denticulate at caudal edge (Figs 3 A, B, 4 C, D).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
13904BA524F85CCBA8D4BE005104B34C.taxon	etymology	Etymology. To emphasize Hala-Bala Wildlife Sanctuary, the type locality. Noun in apposition.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
4E5486C31E6D5230A5A310AB0C51AEF0.taxon	description	Figs 6, 7, 8	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
4E5486C31E6D5230A5A310AB0C51AEF0.taxon	materials_examined	Material examined. Holotype: Thailand - Yala Province • ♂; Betong District, elfin montane forest (Malaya Phytochorion province); 1430 m a. s. l.; 25 May 2022; Wisut Sittichaya leg.; CUMZ.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
4E5486C31E6D5230A5A310AB0C51AEF0.taxon	diagnosis	Diagnosis. This new species is distinguished from its two congeners in sternal process between male coxae 4 linguiform with a rounded tip, and lamina lateralis of gonopodal solenophore triangular, apically bifid and protruded laterally.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
4E5486C31E6D5230A5A310AB0C51AEF0.taxon	description	Description. Length of holotype 31.5 mm, width of midbody pro- and metazonae 2.7 and 3.0 mm, respectively. Colouration of alcohol material after one week of preservation dark red brown (Fig. 6 A-F); paraterga paler, head and antennae light brown to brown (Fig. 6 A, B), legs and venter contrasting light yellow to brown (Fig. 6), antennae and legs increasingly darker brown distally (Fig. 6 B, E, G). All characters as in M. halabala sp. nov., except as follows. Antennae rather long, extending caudally past metaterga 4 when stretched dorsally. Collum with three transverse rows of setae: 4 + 4 in anterior, 2 + 2 in intermediate, and 3 + 3 in posterior row; with a small lateral setigerous incision near midway (Fig. 6 A, B). Paraterga 2 broad, anterior edge angular and rounded, lateral edge with a small notch at about 1 / 4 in front of caudal corner (Fig. 6 A). Calluses on paraterga rather narrow, delimited by a sulcus fully on dorsal side and in posterior half on ventral side; on poreless rings narrower than on pore-bearing ones in dorsal view (Fig. 6 B, D, E). Transverse sulcus distinct (Fig. 6 A, C, F), complete on metaterga 5 - 17, narrow, line-shaped, rather deep, not reaching the bases of paraterga, smooth at bottom, incomplete and nearly wanting on ring 18. Stricture between pro- and metazona wide, deep, beaded at bottom down to base of paraterga starting with segment 5 (Fig. 6 A-F). Pleurosternal carinae complete crests with a sharp caudal tooth on rings 2 - 4, increasingly reduced and retaining a sharp caudal tooth on rings 5 and 6 thereafter, retaining a small caudal tooth on ring 7, missing further on (Fig. 6 B, D, E). Hypoproct roundly subtriangular, setigerous knobs at caudal edge very small and well-separated (Fig. 6 G). Sterna moderately setose, shining, cross-impressions shallow, without modifications; an entire, large, linguiform, sternal lobe between ♂ coxae 4, with a pair of small denticles laterally near base (Figs 6 H, I, 7 E). An inconspicuous and low ridge present in front of gonopod aperture. Legs long and slender, midbody ones ca. 1.6 - 1.9 times as long as body height, without modifications, ♂ tarsal brushes absent. Gonopods (Figs 7 A-D, 8) rather simple; coxa almost straight caudad, densely setose distoventrally. Prefemur as usual, densely setose, about 1 / 3 as long as femorite + postfemoral part. Femorite stout, suberect, showing a distinct mesal groove / hollow (g), with a sulcus demarcating a postfemoral part; seminal groove running entirely mesally along fermorite, solenomere (sl) flagelliform, almost fully sheathed by solenophore (sph). Lamina medialis (lm) well developed, thick and large, unciform, terminal lobe sheathing the tip of solenomere. Lamina lateralis (ll) triangular in shape, protruding laterally, tapered apically, bifid at tip (Figs 7 C, D, 8 A, 8 C-D).	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
4E5486C31E6D5230A5A310AB0C51AEF0.taxon	etymology	Etymology. To emphasize " Malayorthomorpha hulutbeeda " which means " flat-back millipede " in Malay dialect, a noun in apposition. A Malay dialect language is mainly used in three provinces of southern Thailand where the holotype was obtained.	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
7C98A1427D1B537DA34B9149772328D3.taxon	description	Fig. 1	en	Likhitrakarn, Natdanai, Golovatch, Sergei I., Sittichaya, Wisut (2022): Review of the millipede genus Malayorthomorpha Mrsic, 1996 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of two new species from Thailand and a key to its species. ZooKeys 1118: 1-19, DOI: http://dx.doi.org/10.3897/zookeys.1118.89593, URL: http://dx.doi.org/10.3897/zookeys.1118.89593
