taxonID	type	description	language	source
03992419FF88FF98FDD7FB2FF641FB9C.taxon	description	urn: lsid: zoobank. org: act: 718 AE 550 - 9 DB 0 - 4 AB 6 - 83 ED- 95549932 F 36 B	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF88FF98FDD7FB2FF641FB9C.taxon	type_taxon	Type species Rhytidaphora thailandica Reshchikov & Quicke gen. et sp. nov. by present designation.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF88FF98FDD7FB2FF641FB9C.taxon	diagnosis	Diagnosis Rhytidaphora gen. nov. can be distinguished from other genera of Euryproctini by the following combination of character states: lower mandibular tooth much longer than upper (Fig. 1 B); occipital carina broadly incomplete dorsally, joining hypostomal carina shortly before base of mandible; occiput with distinct concavity; fore wing without areolet (Fig. 2 A); tarsal claws distinctly pectinate (Fig. 2 B); T 1 dorsal carinae well-developed, continuing as dorsal carinae of T 2 (Fig. 2 F); S 1 very short (0.3 times as long as T 1); T 2 and T 3 immovably fused (Fig. 2 D); T 1 ‒ 5 with deep basal and subapical transverse striate impressions (Fig. 2 D, F); ovipositor rather short, straight, with subapical, dorsal notch (Fig. 2 D).	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF88FF98FDD7FB2FF641FB9C.taxon	etymology	Etymology The generic name Rhytidaphora derives from Greek “ ρυτίδα ” – “ wrinkle ” and “ φέρω ” – “ bear ”, referring to the sculpture of the metasoma. The gender is feminine.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF88FF98FDD7FB2FF641FB9C.taxon	description	Description MEASUREMENTS. Fore wing length 6.0 mm. HEAD. Clypeus flat in lateral view (Fig. 2 E), its lower margin blunt in its middle and slightly sharp laterally, bulging apically (Fig. 1 B). Face rather sparsely setose (Figs 1 B, 2 E). Anterior tentorial pit rather large (Fig. 1 B). Eyes bare (Fig. 2 C). Lower mandibular tooth much longer than upper (Fig. 1 B). Occipital carina broadly incomplete dorsally, joining hypostomal carina shortly before base of mandible. Occiput with distinct concavity. MESOSOMA. Epomia absent. Notauli shallow and indistinct (Fig. 2 C). Mesopleuron sparsely setose, not punctate (Fig. 2 C). Epicnemial carina joining anterior margin of mesopleuron (Fig. 2 C). Propodeum in lateral profile convexly rounded (Fig. 2 E), with carinae complete except for area apicalis, which is absent (fused with area superomedia) (Fig. 1 C). Fore wing without areolet (Fig. 2 A). Pterostigma receiving vein Rs + 2 r at its basal 0.35. Vein 2 m-cu with a single bulla (Fig. 2 A). Hind wing with cu-a receiving Cu 1 well below middle. Tarsal claws distinctly pectinate (Fig. 2 A). Fore tibia with apical tooth (Fig. 2 G – H). METASOMA. Without obvious punctures but densely finely setose (Fig. 2 D, F). T 1 as long as broad with strong dorsal longitudinal carina continuing in dorsal carinae of T 2 (Fig. 2 F). Glymma absent (Fig. 2 D). S 1 rather short, 0.3 times as long as T 1. T 2 and T 3 immovably fused (Fig. 2 D). T 1 ‒ 5 with distinct basal and subapical transverse impressions distinctly striate. Ovipositor very short, straight, with a long, shallow, subapical dorsal notch (Fig. 2 D). Species included The genus is described as monotypic.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	description	urn: lsid: zoobank. org: act: EC 8 F 8 A 55 - E 204 - 45 C 5 - 8750 - 7 D 26 E 24078 E 1 Figs 1 – 2	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	etymology	Etymology The specific name refers to the species distribution.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	materials_examined	Type material Holotype THAILAND • ♀; Nan Province, Doi Phu Kha National Park; 19 ° 12.3 ′ N, 101 ° 04 ′ E; 6 Sep. 2019; Malaise trap 5; W. Atsawasiramanee leg.; DNA voucher code CCDB 06324 - E 02; GenBank accession code OK 623371; CUMZ. Paratype THAILAND • 1 ♀; Lampang Province, Chae Son National Park, nature trail; 18 ° 50.172 ′ N, 99 ° 28.38 ′ E; 507 m a. s. l.; 14 – 21 Oct. 2007; Malaise trap; B. Kwannui and A. Sukpeng leg., T 5315; QSBG.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	description	Description Female (holotype) MEASUREMENTS. Body length 6.0 mm, fore wing length 6.0 mm, antenna length 7.5 mm. BODY. Matt and sparsely setose. HEAD. Antenna slender, as long as fore wing and with 37 flagellomeres. Scape 1.4 times as long as broad. First flagellomere with prominent basal annellus, 1.6 × as long as 2 nd and 1.9 × as long as 3 rd. Head narrowed behind eyes. Clypeus 0.4 times as high as wide, separated from face by distinct groove. Anterior tentorial pit rather large (Fig. 1 B). Malar space 0.5 times basal mandible width. Inner margins of eyes (Fig. 1 B) almost parallel, scarcely concave near antennal sockets. Lateral ocellus separated from eye margin by 1.2 times its widest diameter. Face matt with dense setiferous punctures (Fig. 1 B), with short yellow setosity. Frons with weak curved groove behind antennal sockets; with deep, moderately dense setiferous punctures. Occipital carina broadly incomplete dorsally, joining hypostomal carina shortly before base of mandible. Gena matt, with sparse punctures (Fig. 2 E). MESOSOMA. Mesoscutum shining, finely and densely punctate (Fig. 2 C). Pronotum with crenulate groove only anteriorly, weakly shiny and with fine, moderately dense puncturation. Mesopleuron weakly shiny, setose and without punctures (Fig. 2 E). Scuto-scutellar groove deep with steep posterior margin, triangular and gradually merging with mesoscutum anteriorly. Scutellum strongly convex, matt, without punctures. Juxtacoxal carina absent. Area superomedia and area apicalis fused, hexagonal (Fig. 1 C). Spiracles almost round. Wings slightly hyaline (Fig. 2 A). Fore wing with 1 cu-a interstitial. Areolet absent, veins 2 m-cu and 2 rs-m interstitial. Pterostigma receiving vein Rs + 2 r at its basal 0.35. 2 m-cu with a single, wide bulla anteriorly. Hind wing with cu-a receiving Cu 1 well below middle (Fig. 2 A). Propodeum smooth with strong lamelliform carination. Area basalis short and wide. Posterior transverse carina absent (Fig. 1 C). Claws pectinate (Fig. 2 B), fore claw with four teeth, mid- and hind claw with three teeth. Hind femur 4.2 times as long (including trochantellus) as maximally broad. Hind tibia 6.2 times as long as apically broad. First tarsomere of hind leg 8 times as long as broad and 2.5 times as long as second tarsomere. METASOMA. Without obvious punctures but densely finely setose (Fig. 2 D, F). T 1 as long as broad, with strong dorsal longitudinal carina continuing in dorsal carinae of T 2 (Fig. 2 F). Ventro-lateral carinae complete (Fig. 2 D). Glymma absent (Fig. 2 D). S 1 rather short, 0.3 times as long as T 1. T 2 and T 3 immovably fused, second suture deep and crenulate (Fig. 2 D, F). T 1 ‒ 5 with distinct basal and subapical transverse impressions distinctly striate (Fig. 2 F). Ovipositor sheath very short, sheaths 0.13 times as long as hind tibia. Ovipositor straight, with a long, shallow, subapical dorsal notch (Fig. 2 D). COLORATION. Yellow, except for the following which are marked with black: antenna (dorsally), teeth of mandible, dorsal part of head, pterostigma, hind coxa dorsoapically, hind femur and tibia apically, hind tarsus (except first tarsomere basally), lateral parts of mesoscutum entirely and its anterior medial part, pair of spots on T 3 and T 4 (Figs 1 – 2). Male Unknown.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	distribution	Distribution Thailand.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
03992419FF89FF95FE42FB94F14DFA67.taxon	description	Molecular analysis As in the only previous study in which Ctenopelmatinae were relatively well represented (Quicke et al. 2009), the molecular analysis did not recover any large ctenopelmatine tribes as monophyletic (Fig. 3). The placement of Scolomus within Ctenopelmatinae, even though this genus has been transferred to Metopiinae (Gauld & Wahl 2006), is consistent with the idea that Metopiinae may be derived from within the Ctenopelmatinae (Quicke et al. 2009). The Euryproctini as currently constituted were recovered spread across the tree in five separate clades but with low support. Nevertheless, nine of the included euryproctine genera formed a monophyletic grouping (Fig. 3, top) including Rhytidaphora gen. nov., and is therefore consistent with the placement of the new genus based on our morphological assessment.	en	Reshchikov, Alexey, Quicke, Donald L. J., Butcher, Buntika A. (2022): A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand. European Journal of Taxonomy 834 (1): 102-116, DOI: https://doi.org/10.5852/ejt.2022.834.1903, URL: http://dx.doi.org/10.5852/ejt.2022.834.1903
