taxonID	type	description	language	source
4D4D16FD37875A55908ED8588CDE43B5.taxon	description	Figs 2, 3, 6, 7	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
4D4D16FD37875A55908ED8588CDE43B5.taxon	materials_examined	Material studied. Holotype. 1 male (CUMZ-D 00149 - 1), Thailand, Uthaithani Province, Lan-Sak District, Hub Pa Tard; 15 ° 22 ' 37.13 " N, 99 ° 37 ' 49.98 " E; 119 m a. s. l.; 26 July 2020; P. Pimvichai, T. Backeljau and P. Prasankok leg. Paratypes. 3 males (CUMZ-D 00149 - 3), 3 females (CUMZ-D 00149 - 2); same data as holotype.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
4D4D16FD37875A55908ED8588CDE43B5.taxon	etymology	Etymology. The species epithet means " friend of the dragon " and refers to the type locality, which is shared with the " shocking pink dragon millipede ", Desmoxytes purpurosea Enghoff, Sutcharit & Panha, 2007.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
4D4D16FD37875A55908ED8588CDE43B5.taxon	diagnosis	Diagnosis. Differing from other species in the genus by having the tip of anterior gonopod crossing over with tip of opposite side, the two together delimiting a drop-shaped " window ", whereas in the other two species the tips of anterior gonopod are separated from each other; externally differing by its bright orange color on the anterior and posterior ends. In contrast, S. constrictus gen. et sp. nov. has a row of triangular dark brown spots middorsally on its body rings, while S. prasankokae gen. et sp. nov. has a row of rectangular dark brown spots middorsally on the body rings.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
4D4D16FD37875A55908ED8588CDE43B5.taxon	description	Description. Adult males with 41 or 42 podous rings, 1 apodous ring. Length ~ 3 cm, diameter 2.7 - 2.9 mm. Adult females with 41 - 43 podous rings, 1 - 3 apodous rings. Length ~ 3 cm, diameter 2.8 - 3.1 mm. Colour. Living animal mainly dark brown. Head, dorsal part of the first four body rings and telson orange, antenna light brown, middorsal metazona orange (Fig. 6 A, B). Anterior gonopods (Fig. 3 A, B, D, E) with extremely small triangular process between coxae (st), with high coxae, apically narrow, curving mesad, tip crossing over with tip of opposite side, the two together delimiting a drop-shaped " window ". Telopodite flattened apically rounded, curving backward, directed distad, as a thumb-like process (Fig. 3 B, arrow). Posterior gonopods (Fig. 3 C, F-K) simple, rounded, with short, smooth coxal part (pcx); telopodital part (pt) twice the length of the pcx, lateral margin folding mesad, forming a deep concavity, the inner lateral margin expanded into rounded lamella, apically forming a canopy (Fig. 3 J, arrow). Female vulvae (Fig. 3 L): simple, valves prominent, the right valve slightly larger than the left valve.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
4D4D16FD37875A55908ED8588CDE43B5.taxon	distribution	Distribution. Known only from the type locality in Uthaithani Province, Thailand (Fig. 7).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	description	Figs 2, 4, 6, 7	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	materials_examined	Material studied. Holotype. 1 male (CUMZ-D 00150 - 1), Thailand, Prachuap Khiri Khan Province, Kui Buri District, Ban Yang Chum; 12 ° 04 ' 19.64 " N, 99 ° 42 ' 57.63 " E; 149 m a. s. l.; 7 August 2014; C. Sutcharit leg. Paratypes. 1 male (CUMZ-D 00150 - 2), 2 females (CUMZ-D 00150 - 3); same data as holotype.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	etymology	Etymology. The species epithet is a Latin adjective derived from the verb constringere = " to tighten " and refers to the constricted anterior gonopod coxa.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	diagnosis	Diagnosis. Differing from other species in the genus by having the anterior gonopod coxae constricted at middle of lateral margin, while the other two species have no constriction at middle of lateral margin of anterior gonopod coxae; externally differing by having a middorsal row of triangular dark brown spots, with dark brown bands on both lateral sides running parallel to the dorsal one.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	description	Description. Adult males with 41 or 42 podous rings, 1 apodous ring. Length ~ 3 cm, diameter 3.6 - 3.7 mm. Adult females with 42 or 43 podous rings, 1 apodous ring. Length ~ 3 cm, diameter 4.1 - 4.5 mm. Colour. After 8 years in alcohol overall beige with 3 stripes from head to telson: a mid-dorsal row of triangular dark brown spots, with dark brown bands on both lateral side running parallel to the dorsal one (Fig. 2 J). Anterior gonopods (Fig. 4 A, B, D, E) with a small triangular process between coxae (st), with high coxae, broad at base, gradually narrowing towards tip, distinctly constricted at middle of lateral margin, apical part flattened, curving mesad, mesal margin basally straight, curving towards tip, posterior surface folding cover more than half of telopodite. Telopodite small, directed distad, thumb-like process, with longitudinal ridge from base to tip (Fig. 4 B, arrow). Posterior gonopods (Fig. 4 C, F-J) simple, rounded, with short, smooth coxal part (pcx); telopodital part (pt) twice the length of the pcx, lateral margin expanded, folding mesad, forming a concavity (not so deep as in the other two species). Female vulvae (Fig. 4 K): simple, valves prominent, the left valve slightly larger than the right valve.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
AA76D192049B577981E4BBE21E1D3389.taxon	distribution	Distribution. Known only from the type locality in Prachuap Khiri Khan Province, Thailand (Fig. 7).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
69F198FE3B5153688DC89F09B98B2D2B.taxon	description	Fig. 2	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
69F198FE3B5153688DC89F09B98B2D2B.taxon	etymology	Etymology. From Latin siliqua = pod, and referring to the podlike shape of the gonopod telopodite.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
69F198FE3B5153688DC89F09B98B2D2B.taxon	diagnosis	Diagnosis. Siliquobolellus species are characterised by (1) the unique shape of the telopodital part of their posterior gonopod, which forms a deep concavity. In contrast, the telopodital part of the posterior gonopod is extremely slender and sickle-shaped in Pseudospirobolellus, does not form a deep concavity in Coxobolellus, or is slender with a broad apical canopy in Benoitolus; (2) the telopodite of anterior gonopod (at) simple, directed distad, not reaching tip of coxal part of the anterior gonopod (cx). In contrast, the telopodite of anterior gonopod (at) is overreaching coxa in the other three genera; and (3) tip of anterior gonopod coxa narrowed, curving mesad. In contrast the tip of the anterior gonopod coxa directed distad in Coxobolellus, or is rounded in Pseudospirobolellus and Benoitolus.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
69F198FE3B5153688DC89F09B98B2D2B.taxon	description	General description. Head capsule smooth. Occipital furrow extending down between, but not beyond eyes; clypeal furrow reaching level of antennal sockets. Area below antennal sockets and eyes impressed, forming part of antennal furrow. Incisura lateralis (IL) open (Fig. 2 A). 2 + 2 labral teeth, a row of labral setae, 3 + 3 supralabral setae. Diameter of eyes ca half of interocular space; 7 vertical rows of ommatidia, 4 horizontal rows, 24 - 26 ommatidia per eye. Antennae short, not reaching beyond collum when stretched back, accommodated in a shallow furrow composed of a horizontal segment in the head capsule and a vertical segment in the mandibular cardo and stipes. Antennomere lengths 2> 6> 1> 5> 3 = 4> 7 in Siliquobolellus amicusdraconis gen. et sp. nov. and S. constrictus gen. et sp. nov. and 2> 1> 6> 5> 3 = 4> 7 in S. prasankokae gen. et sp. nov.; antennomere 1 glabrous, 2 and 3 with some ventral setae, 4, 5 and 6 densely setose; 4 apical sensilla. Mandibles: stipes (Mst) broad at base, apically gradually narrowed, triangular. Gnathochilarium (Fig. 2 B): each stipe (Gst) with 3 apical setae; each lamella lingualis with 2 setae, one behind the other. Basal part of mentum (Me) transversely wrinkled; basal part of stipites longitudinally wrinkled. Collum smooth, with a marginal furrow along lateral part of anterior margin; lateral lobes narrowly rounded, extending as far ventrad as the ventral margin of body ring 2. Body rings 2 - 3 ventrally concave, hence with distinct ventrolateral " corners ". Body rings very smooth, parallel-sided in dorsal view. Prozona smooth. ' Tergo-pleural' suture visible on pro- and mesozona; mesozona ventrally with fine oblique striae, dorsally punctate; metazona ventrally with fine longitudinal striae, otherwise smooth. " Pleural " parts of rings with fine oblique striae. Sterna transversely striate. Ozopores from ring 6, situated in mesozona, ~ 1 / 2 pore diameter in front of metazona. Telson smooth; preanal ring (Pre) with slightly recurved dorsal profile, with short process protruding to vertical tangent to anal valves or slightly beyond (Fig. 2 C-E). Anal valves (Av) smooth, rounded (Fig. 2 C-E). Subanal scale (Sub) broadly triangular (Fig. 2 F). Legs (Fig. 2 G, H): length of midbody legs 55 - 60 % of body diameter in males, 40 - 48 % of body diameter in females. Prefemur basally constricted and longer than other podomeres. First and second legs with 2 or 3 prefemoral, 2 or 3 femoral, 2 or 3 postfemoral, and 2 - 4 tibial setae, and 4 or 5 ventral and 1 dorsal apical setae on tarsi, numbers of setae reaching constancy from pair 3: in males each podomere from postfemur to tibia with 1 seta; tarsi with 1 ventral apical and 1 dorsal apical seta; in females each leg podomere from coxae to tibia with 1 seta, tarsi with 1 - 3 ventral and 1 dorsal apical setae. Claw slender and sharp. Male sexual characters. Apical part of coxa and entire ventral surface of prefemur from third to the last body rings with large ventral soft pad. Body ring 7 entirely fused ventrally, no trace of a suture. Tip of anterior gonopods visible when the animal is stretched out (not when it is rolled up). Anterior gonopods (Figs 3 - 5 A, B, D, E) with a fairly small triangular mesal sternal process (st). Coxa broad at base, apically gradually narrowed, curving mesad, posterior surface folding over telopodite, for accommodation of telopodite. Telopodite gradually narrowed towards tip, directed distad, not reaching tip of anterior gonopod coxa (cx). Posterior gonopods (Figs 3 - 5 C, F) simple, rounded, with short, smooth coxal part (pcx); with prominent opening of efferent groove (oeg) distomesally of pcx; telopodital part (pt) twice the length of the pcx, lateral margin folding mesad, forming a deep concavity. Female vulvae (Figs 3 L, 4 K, 5 K) simple, valves prominent. Species descriptions	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	description	Figs 2, 5, 6, 7	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	materials_examined	Material studied. Holotype. 1 male (CUMZ-D 00148 - 1), Thailand, Lampang Province, Ngao District, Pha Thai; 18 ° 36 ' 01.05 " N, 99 ° 54 ' 01.00 " E; 350 m a. s. l.; 28 July 2020; P. Pimvichai, P. Prasankok and S. Saratan leg. Paratypes. 6 females (CUMZ-D 00148 - 2); same data as holotype.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	etymology	Etymology. The species honors Associate Professor Dr. Pongpun Prasankok, biologist and devoted millipede collector.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	diagnosis	Diagnosis. Differing from other species in the genus by having the posterior gonopod apically with a short, sharp protrusion; externally differing by having a row of rectangular dark brown spots middorsally on the body rings.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	description	Description. Adult male with 41 podous rings, 1 apodous ring. Length ~ 3 cm, diameter ~ 3 mm. Adult females with 40 or 41 podous rings, 1 apodous ring. Length ~ 3 cm, diameter 3.2 - 3.6 mm. Color of living animal: collum and telson beige; antenna and leg light brown; dorsal part of body rings yellowish brown, with a row of rectangular dark brown spots middorsally on the body rings; head and lateral part of body rings dark brown (Fig. 2 I, 6 C, D). Anterior gonopods (Fig. 5 A, B, D, E) with high coxae, apically narrow, curving mesad, mesal margin sigmoid, posterior surface folding to cover more than half of telopodite, with strong ridge laterally. Telopodite with thick process, with strong ridge distomesally (Fig. 5 B, arrow). Posterior gonopods (Fig. 5 C, F-J) simple, rounded, with short, smooth coxal part (pcx); telopodital part (pt) twice the length of the pcx, lateral margin folding mesad, forming a deep concavity, the inner lateral margin expanded into a rounded lamella, apically forming a canopy (Fig. 5 I, arrow), apically with a short, sharp protrusion (Fig. 5 C, G, arrows). Female vulvae (Fig. 5 K): simple, valves prominent, the right valve slightly larger than the left valve.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
2E9151E1675E59D8840599C81BBF9B62.taxon	distribution	Distribution. Known only from the type locality in Lampang Province, Thailand (Fig. 7).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak, Backeljau, Thierry (2022): A new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species. Zoosystematics and Evolution 98 (2): 313-326, DOI: http://dx.doi.org/10.3897/zse.98.90032, URL: http://dx.doi.org/10.3897/zse.98.90032
