taxonID	type	description	language	source
BA26D0AF4AC55303B0FC76FC0B9B17E7.taxon	description	Figs 1 A, 2	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
BA26D0AF4AC55303B0FC76FC0B9B17E7.taxon	description	Description. Tree or rarely shrub, unarmed or sometimes with blunt spine-like projections on young twigs (W. Hawthorne, personal communication), 5 - 35 m tall, stem to 40 cm in diameter. Branches covered with many minute round lenticels 0.1 - 0.2 mm, twigs, stipules, bud scales, all leaf axes and peduncles brown pubescent, often densely so. Foliage consisting of bipinnate leaves with opposite to sub-opposite pinnae with closely-spaced, distinctly discolourous leaflets, slightly wider in the middle with the apical pair distinctly distally projected, in the basal pair often one or both leaflets reduced to paraphyllidia. The inflorescences consisting of sub-spherical capitula emerging in axillary fascicles, often aggregated into pseudoracemes emerging from the perulate resting buds, with the leaves not fully developing and caducous. Stipules deltoid to linear, 2 - 8 x 1.0 - 1.8 mm, caducous, perulae deltoid, 2 - 4 x 1.8 - 3 mm. Leaves with (3 -) 5 - 7 (- 8) pairs of pinnae, apical pairs usually slightly longer and the basal pair sometimes much shorter with fewer leaflets, petiole 1.3 - 2.5 cm, pulvinate, adaxially shallowly canaliculate and often laterally and abaxially grooved, with a circular to elliptic sessile cupular nectary ca. 0.8 - 2.1 mm in diameter located ca. mid-petiole, rachis (1.8 -) 4.5 - 11 (- 14.5) cm, adaxially shallowly canaliculate and often laterally and abaxially grooved, with 2 - 5 circular to elliptic sessile cupular nectaries between or just below the distal pairs of pinnae ca. 0.5 mm in diameter, pinna-rachises (2.4 -) 3.5 - 8 (- 10.2) cm, proximally pulvinate, laterally compressed, with elliptic to circular sessile cupular nectaries of 0.2 - 0.8 x 0.2 - 0.5 mm between the upper 1 - 4 (- 15) leaflet pairs, pinnae with (7 -) 13 - 20 (- 25) leaflet pairs, very often the abaxial leaflet of the lowermost pair reduced to a paraphyllidium, leaflets sessile, pulvinate, inequilaterally hastate or weakly sigmoid, with acute or sometimes obtuse apex or (in subsp. Osodendron altissimum busiraensis) weakly rhombic with oblique base and mostly obtuse apex or sometimes apiculate, (4 -) 7 - 14 (- 18) x (1.5 -) 3 - 4 mm, asymmetrically palmately-pinnate secondary venation with 7 - 10 veins starting from leaflet base next to the mid-rib and (7 -) 15 - 22 major and intercalary secondary veins laterally from the mid-rib, brochidodromous to craspedodromous, distinctly prominent on both surfaces when dry (or obscure on adaxial surface in subsp. Osodendron altissimum busiraensis), tertiary venation finely reticulate, hardly visible on adaxial surface. Capitula ca. 35 - 50 - flowered, on peduncles 1.8 - 3.2 cm, dimorphic with 3 - 9 central flowers with broader base and longer staminal tubes, bracts lanceolate to lanceolate-spatulate, ca. 0.9 - 1.2 x 0.1 - 0.2 mm, ciliate in upper half, caducous. Peripheral flowers slenderly campanulate with pale green or greenish-white calyx and corolla, subsessile to shortly pedicellate, pedicel to ca. 0.5 mm, sparsely pubescent. Calyx 5 - merous, 2 - 3.2 mm long, the deltoid lobes ca. 0.5 x 0.5 mm, ciliate at the apex and often also a few scattered short hairs on the outer surface of calyx tube and lobes, otherwise glabrous, corolla 5 - merous, 5 - 7.5 mm long, glabrous, androecium of ca. 20 - 25 stamens, filaments white, ca. 12 - 17 mm long, fused into a tube at the base for ca. 2 - 3 mm, anthers yellow, basifixed, pollen aggregated into 32 - celled flattened polyads. Pistil ca. 13 - 18 mm long, the ovary ca. 3 mm with a few scattered hairs on distal half and the funnel-shaped stigma extending ca. 1 mm beyond the anthers. Central flowers similar, but sessile, broadly campanulate and staminal tube ca. 5.5 - 8.5 mm long (i. e. the stamens are approximately the same length, but the fused tube is longer than in peripheral flowers), the tube exserted beyond the corolla. Fruit a spirally-curved, twisted or coiled lomentiform pod, usually articulate between the seeds, breaking up tardily after abscission in water or on the forest floor, often with arenchymous swollen mesocarp for flotation, (10 -) 18 - 29 (- 30) x (0.8 -) 1.1 - 1.7 x 0.2 - 0.6 cm, (3 -) 9 - 20 (- 23) seeded. Seeds plano-compressed, nearly round or slightly elliptic, ca. 5 - 7.5 x 4 - 6 x 1.5 - 2 mm, chestnut brown with a small 2.1 - 3.2 x 1 - 1.5 mm elliptic to obovate or oblong open pleurogram.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
9FF091AF2B4D5B43954E3ACE0CEA1992.taxon	distribution	Distribution. Widespread across Upper and Lower Guinea and Congolia, from Guinea-Bissau in the west to the Democratic Republic of Congo in the east and Angola in the south (Fig. 2 A), also reported from Uganda and Zambia (Brenan 1959).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
A7DA518FBF2B555DB63558259082E9D1.taxon	distribution	Distribution. Local to the Busira river catchment in the western Democratic Republic of Congo (Fig. 2 A).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
B1C0D13B3EDF5C329DED71CDBDE5CB09.taxon	description	Fig. 3 A-F	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
B1C0D13B3EDF5C329DED71CDBDE5CB09.taxon	description	Description. Tree, up to 30 m tall, twigs with rows of lenticels and becoming shallowly grooved with age. Twigs, stipules, perulate resting buds, all leaf-axes and peduncles covered in a densely villose ferruginous indumentum. Foliage consisting of very finely divided bipinnately compound leaves, the tiny leaflets ciliate and somewhat discolourous. The inflorescences consisting of sub-spherical capitula emerging from axillary fascicles of coeval leaves, sometimes in compound ramiflorous pseudoracemes with the leaves not fully developing and caducous. Stipules narrowly deltoid or lanceolate to linear 3.5 - 4.3 (- 9) x ca. 1 mm, caducous leaving conspicuous glabrous scars, perulae deltoid, ca. 2 x 1.5 mm. Leaves with 10 - 34 pairs of pinnae, often with one or a few much reduced pinnae at the base, petiole 5 - 11 mm, pulvinate and adaxially flattened at the base, usually with a sessile or shortly stipitate cupular nectary 0.5 - 0.7 mm in diameter just below the lowermost pinna pair, rachis grooved and adaxially canaliculate although obscured by dense indumentum, with a 1 - 1.5 mm long deciduous mucro, 6.5 - 16 (- 21) cm, with 3 - 8 (- 11) interpinnal subsessile cupular nectaries apically, these sometimes button-shaped and on a 0.7 mm long stipe, pinnae obscurely pulvinate, with an adaxial ridge and a tiny 0.3 - 0.4 mm long mucro at the apex, 1.0 - 2.5 (- 4.5) cm, usually without nectaries, except on larger leaves (of juveniles or coppice shoots) with cupular nectaries between the upper 1 - 3 leaflet pairs, pinnae with (9 -) 27 - 40 (- 48) leaflet pairs, the abaxial leaflet of the lowermost pair usually reduced to a paraphyllidium, leaflets inequilaterally linear, sessile and pulvinate, with strongly oblique base and acute apex, 0.8 - 3 x 0.2 - 0.8 mm, venation obscure on the adaxial surface, palmately-pinnate brochidodromous, with three secondary veins emerging from the base next to the clearly stronger mid-rib on the adaxial side, 2 - 3 further secondary veins on each side and sparse reticulate tertiary venation. Capitula 1 - 3 per leaf axil, ca. 40 - 60 - flowered, on peduncles 1.8 - 3.5 (- 4.1) cm, dimorphic with 1 (- 3) central flower (s) that are more robust and with a longer, exserted staminal tube. Peripheral flowers ca. 0.7 mm broad at the base, the basal ones on a pedicel of ca. 0.4 - 0.6 mm, the others sub-sessile to sessile closer to the apex. Calyx slenderly campanulate, green, 5 - merous, 1.6 - 3.1 mm long, densely puberulous on outer surface, the deltoid lobes ca. 0.4 x 0.4 mm, corolla green, 4 - or 5 - merous, 3 - 5.5 mm long, pubescent, the lobes 1.8 - 2 mm long, stamens 10 - 14, filaments white, 14 mm long, basally fused into a tube ca. 3 mm long, anthers light yellow, basifixed, pollen released in 32 - celled polyads, pistil ca. 18 mm long with a sessile ovary ca. 2.5 mm long, puberulent on upper half with white hairs, stigma shallowly funnel-shaped. Central flower (s) similar to peripheral flowers, but ca. 1.8 mm broad at the base, calyx 2.0 - 2.4 mm long, corolla ca. 4.5 mm long, staminal tube ca. 6 mm long (i. e. the stamens are approximately the same length, but fused for a greater part than in peripheral flowers). Fruit an indehiscent woody pod up to 13 - 18 x 2 - 2.2 (- 2.5) x 0.3 - 0.6 cm, straight or slightly curved and with uniformly thick valves, except for the distinctly thickened margins, not swollen over the seeds, 16 - 24 seeded when well-fertilised, seeds rounded elliptical, only slightly laterally compressed, greenish light brown, 6 - 7.5 x 4 - 5 x 3 - 3.5 mm when dry, with a hard testa and an elongate closed pleurogram 5 - 6 x 2 - 2.5 mm.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
B1C0D13B3EDF5C329DED71CDBDE5CB09.taxon	distribution	Distribution. Upper Guinea, from Guinea-Bissau to Ghana (Fig. 3 G).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
B1C0D13B3EDF5C329DED71CDBDE5CB09.taxon	materials_examined	Representative specimens examined. Guinee-Bissau: Espirito Santo 1747 (K) & 1864 (K) & 1871 (K) & 2697 (K). Sierra Leone: X. van der Burgt 1994 (K).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
5D4D1D3766355BCDA201EDF8BB6ABED6.taxon	materials_examined	Type. Osodendron altissimum (Hook. f.) E. J. M. Koenen.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
5D4D1D3766355BCDA201EDF8BB6ABED6.taxon	diagnosis	Diagnosis. Osodendron differs from Robrichia by pods being either straight to twisted or coiled, but not contorted and ear-shaped.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
5D4D1D3766355BCDA201EDF8BB6ABED6.taxon	description	Description. Trees, or rarely shrubs, unarmed or sometimes with spine-like projections on twigs or spine-like outgrowths on adventitious roots, resting buds perulate with scales distinctly different from stipules. Indumentum of usually dense ferruginous pubescence on twigs, petioles, rachis and pinna rachises, stipules, bud scales and peduncles. Leaves with (3 -) 5 - 30 (- 34) pairs of pinnae, opposite or sometimes (the lowermost pairs) sub-opposite, with a single petiolar nectary usually present, as well as nectaries between at least some of the pinna pairs on abaxial surface of the rachis and often also between the upper 1 - many leaflet pairs on the pinna-rachises, the lowermost pair of pinnae usually distinctly shorter than others, pinnae with (7 -) 13 - 40 (- 48) leaflet pairs, one of the two leaflets of the lowermost pair usually reduced to a small paraphyllidium or lacking. Inflorescences sub-globose capitula, dimorphic, borne on peduncles arising from axillary fascicles, sometimes arranged in short compound pseudoracemes with the leaves suppressed (not fully developing) and caducous as apparent from the presence of leaf scars in the pseudoracemes, these pseudoracemes developing below the foliage. Flowers sessile or shortly stipitate, 4 - or 5 - merous, with fused calyx and corolla, androecium consisting of 10 - 25 stamens that are fused in the lower part to form a staminal tube, pollen compound in 32 - celled polyads, central flowers more robust with a broader nectariferous base and longer staminal tube exserted beyond corolla tube. Fruits septate, indehiscent or tardily breaking up into articles, either lomentiform, twisted and strongly curved to coiled or weakly to not articulate and slightly curved to straight, seeds with a hard testa and open or closed pleurogram.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
5D4D1D3766355BCDA201EDF8BB6ABED6.taxon	distribution	Distribution and habitat. Three species in tropical Africa, from Senegal in the west to the Democratic Republic of Congo in the east and Zambia and Angola in the south. Typically occurring in rainforest and extending into the savannah zone in gallery forest.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
5D4D1D3766355BCDA201EDF8BB6ABED6.taxon	etymology	Etymology. The genus is named after " Oso ", a food that is prepared in West Africa (Ghana and Nigeria) by fermenting the seeds of the type species O. altissimum and grinding them into a protein-rich paste that is subsequently cooked and eaten as either a main food, a delicacy or as a condiment to flavour soups and stews (Popoola et al. 2004; Jolaoso et al. 2014).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
8280CC2267BA547895583B890922FA3B.taxon	description	Fig. 3 H-K	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
8280CC2267BA547895583B890922FA3B.taxon	description	Description. Tree or more rarely a shrub, unarmed or with spine-like outgrowths near the base of the bole perhaps associated with adventitious roots (W. Hawthorne, personal communication), to 30 m tall and to 1.25 m in stem diameter, twigs dark with densely scattered small pustular lenticels or sometimes with lighter coloured corky bark. Twigs, stipules, perulate resting buds, all leaf-axes and peduncles covered in dense ferruginous villose indumentum. Foliage consisting of finely divided bipinnate leaves, the leaflets variably ciliate, sometimes with only very few hairs on the margins, the lamina usually glabrous, but sometimes also with a few scattered hairs along the mid-rib on lower surface or rarely pilose or with appressed long hairs on both surfaces especially in lower half, usually distinctly discolourous. The inflorescences consisting of sub-spherical capitula emerging from axillary fascicles of coeval leaves, usually a few below the leaves in the axils of caducous leaves on the same shoots or sometimes also with a few short ramiflorous pseudoracemes, with all leaves caducous, lower down on the branch. Stipules elliptically oblong to asymetrically oblanceolate or falcate, 5 - 8 (- 9) x 2 - 3 mm, more scarcely pubescent or nearly glabrous on adaxial surface, caducous leaving conspicuous scars, perulae deltoid to ovoid, ca. 1.5 - 5 x 1.5 - 3.5 mm. Leaves with (10 -) 12 - 30 (- 35) pairs of pinnae, the lower 1 - 2 pairs of pinnae often shorter with fewer leaflets, petiole pulvinate and flattened on adaxial side at the base, (0.5 -) 0.8 - 2.2 (- 3) cm long, petiolar nectary usually present, cupular and circular, sessile or shortly stipitate and 0.6 - 1 mm in diameter, in variable position ranging from just above the pulvinule to just below the basal pair of pinnae, rachis canaliculate although often obscured due to dense pubescence, 8 - 15 (- 22) cm long, with 1 - 4 (- 6) cupular nectaries between the upper pairs of pinnae, these 0.3 - 0.6 mm in diameter, pinnae pulvinate and with an adaxial ridge, (0.5 -) 2.5 - 5 (- 7.5) cm long, nearly always the abaxial leaflet of the lowermost pair reduced to a paraphyllidium and the adaxial one somewhat smaller than the other leaflets, with elliptical nectaries ca. 0.1 - 0.25 mm in diameter between the upper 1 - 3 leaflet pairs, pinnae with (17 -) 24 - 35 (- 42) leaflet pairs, leaflets sessile, pulvinate, asymmetrically oblong to oblanceolate, base oblique to sometimes hastate, apex rounded to acute, (2 -) 3 - 6 (- 9) x (0.3 -) 0.5 - 1.2 (- 2) mm, venation palmately-pinnate brochidodromous, with 2 - 4 basal veins adaxially next to the mid-rib and 5 - 8 further secondary veins on each side and reticulate tertiary venation, apart from the mid-rib, all venation often obscure in smaller leaves, otherwise the secondary venation prominent, sometimes only on the lower surface. Capitula 2 - 4 arranged in fascicles, on peduncles ca. 2.7 - 4.5 cm, dimorphic with ca. 50 - 70 peripheral flowers and a single elongated central flower. Bracteoles spatulate, ca. 0.5 mm long, pubescent on outer surface. Flowers white, the peripheral ones on pedicels 0.75 - 2.5 mm. Calyx slenderly campanulate, green, 5 - merous, 2 - 2.5 mm long, densely puberulous on outer surface, the deltoid lobes ca. 0.25 mm long, corolla 4 - 4.5 mm long, pubescent on outer surface, the lobes 1.8 - 2 mm long, stamens 10 - 14, the filaments white, 11 - 14 mm long, basally fused into a tube 1.5 - 2.5 mm long, anthers basifixed, pollen released in 32 - celled polyads, pistil on a ca. 0.25 - 0.75 (- 1.25) mm long stipe, ovary ca. 1 - 1.5 mm long, pubescent, style 12 - 16 mm long, stigma shallowly funnel-shaped. Central flower (s) similar to peripheral flowers, but sessile and longer, calyx ca. 2.5 mm long, corolla ca. 6 - 6.5 mm long, staminal tube 4 - 6 mm long, exserted beyond corolla tube. Fruit a dark brown to black indehiscent pod that may tardily disintegrate into 1 - multiple-seeded parts, the slightly thickened margins usually straight or also often articulate especially around aborted seeds, the valves papery and thin or with somewhat thickened mesocarp, but not really ligneous, when ripe, swollen over the seeds, when well-fertilised, 12 - 28 (- 32) - seeded, (4.5 -) 10 - 16 (- 20) x 0.9 - 1.3 x 0.4 - 0.7 cm. Seeds yellowish-brown, 5.5 - 7.5 x 3 - 4.5 x 2 - 3 mm with hard testa and a darker closed pleurogram of 4.5 - 5.5 x 1.2 - 1.5 mm.	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
8280CC2267BA547895583B890922FA3B.taxon	distribution	Distribution. Lower Guinea and Congolia in Cameroon, Central African Republic, Gabon, Republic of the Congo, Democratic Republic of Congo, Angola (Fig. 3 G).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
8280CC2267BA547895583B890922FA3B.taxon	materials_examined	Representative specimens examined. Cameroon: J. Mildbread 8584 (K); Democratic Republic of Congo: Luidi, Thysville, Lusolo, 16 October 1959, P. Compere 610 (BR), Isangi, 21 February 1950, Callens 2363 (BR), Petite vallees d'affluente de la Belanzovi, pres son confluent avec la Lubimbe, 26 January 1949, A. Michelson 877 (BR, K).	en	Koenen, Erik J. M. (2022): Osodendron gen. nov. (Leguminosae, Caesalpinioideae), a new genus of mimosoid legumes of tropical Africa. PhytoKeys 205: 453-470, DOI: http://dx.doi.org/10.3897/phytokeys.205.82821, URL: http://dx.doi.org/10.3897/phytokeys.205.82821
