taxonID	type	description	language	source
03CEA025FFE4FFB1FDC6FBB52B4EFBD3.taxon	description	Discodon is a morphologically diverse and heterogeneous genus distributed all across the continental parts of the Neotropical region and into the south-western United States. Gorham (1881) characterised the genus based on males as having the sides of the pronotum with a small notch, the last ventrite deeply divided into two halves, and the anterior claw of the protarsus with a broad basal lobe. The males of other genera of Silinae occurring in the Atlantic forests of Brazil – Silis Charpentier, 1825 and Pachymesia Westwood, 1849 – differ from Discodon for having the pronotum much wider than long and the lateral margins deeply notched, with variously shaped spines and projections, the anterior tarsal claw of the protarsus with a small basal lobe, the other claws simple, and the antennae sometimes distinctly swollen (Pachymesia). Other species from the Atlantic Forests currently combined in Incisosilis Pic, 1908, Polemius LeConte, 1851, Parasilis Gorham, 1885 and Malthinocantharis Pic, 1914 might need to be transferred to the aforementioned genera. The widespread Neotropical genus Polemius LeConte, 1851 is primarily distinguished from Discodon by the meso- and metatarsal claws of the males, which have a basal lobe in Polemius, but are deeply cleft in Discodon (Constantin 2017), but the phylogenetic relevance of this character is yet to be proven for the Neotropical fauna. The Discodon tricolor mimetic complex includes relatively large species (9.7 – 16.2 mm) characterised by the black body and legs, a similar orangish pattern on the pronotum and a pair of white to yellowish rounded or semi-circular spots in the middle of the elytra. It also includes some species with entirely black elytra that are here described as new species or have been treated as subspecies of D. tricolor.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE4FFB4FD8CF88C2DE3F893.taxon	description	Figs 2, 6 A, L, 7 A, 8 A, 9 A, 10 A, 11 A – C, 15 A	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE4FFB4FD8CF88C2DE3F893.taxon	diagnosis	Diagnosis Discodon tricolor differs from the other species for having most of the antennomeres yellow. Discodon tricolor is similar to D. vanini sp. nov. and D. neoteutonum sp. nov., differing from these species by the ventral wall of the tegmen (Fig. 11 A – C) longer, about as long as wide, and the median lobe with a narrow central sclerite; the last ventrite of the females (Fig. 15 A) with a distal margin almost straight, with a slight median projection.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE4FFB4FD8CF88C2DE3F893.taxon	materials_examined	Type material Holotype of Discodon tricolor BRAZIL • ♂; Minas Gerais, “ du midi de la capitale des mines ”; “ Silis tricolor Guér. icon. R. A. ”; MNHN, Paris EC 14297 (Fig. 2 A – C). Lectotype of Discodon albonotatum (here designated) BRAZIL • ♂; Rio de Janeiro, Itatiaya [Itatiaia]; 2400 m a. s. l.; Feb. 1899; E. Gounelle leg.; “ Museum Paris, Coll. M. Pic ”; MNHN, Paris EC 13550 (Fig. 2 D – F). Paralectotypes of Discodon albonotatum (3 specimens) BRAZIL • 1 ♀; São Paulo, Vale do Rio Pardo; Dec. 1898; E. Gounelle leg.; “ Museum Paris, Coll. M. Pic ”; MNHN, Paris EC 13549 • 1 ♂; Rio de Janeiro, Itatiaya [Itatiaia]; 2400 m a. s. l.; Feb. 1899; E. Gounelle leg.; “ Museum Paris, Coll. M. Pic ”; MNHN, Paris EC 13551 • 1 ♀; Rio de Janeiro, Itatiaya [Itatiaia]; 850 m a. s. l.; Feb. 1899; E. Gounelle leg.; “ Museum Paris, Coll. M. Pic ”; MNHN, Paris EC 13552. Other material examined (29 specimens) BRAZIL • 2 ♂♂, 1 ♀; Rio de Janeiro, Itatiaia; Feb. 1955; Dirings leg.; MZSP 46319 to 46321 • 5 ♂♂, 2 ♀♀; same collection data as for preceding; Mar. 1959; MZSP 46322 to 46328 • 1 ♂; same collection data as for preceding; Apr. 1959; MZSP 46329 • 1 ♂; same collection data as for preceding; Jan. 1963; MZSP 46330 • 2 ♂♂, 3 ♀♀; same collection data as for preceding; Jan. 1967; MZSP 46331 to 46335 • 1 ♀; same collection data as for preceding; 5 Feb. 1963; J. Halik leg.; MZSP 46336 • 2 ♂♂, 1 ♀; Rio de Janeiro, Parque Nacional do Itatiaia; above 1200 m a. s. l.; 5 Apr. 1964; C. E. and E. S. Ross legs.; NHMB [labelled as “ D. tricolor (Guér.) mit Typus verglichen ”] • 1 ♂; same collection data as for preceding; 4 Feb. 1927; Dr Seitz leg.; NHMB [labelled as “ D. albonotatum Pic ” by Wittmer] • 1 ♂; same collection data as for preceding, Maromba; 7 Apr. 1989; B. Harky leg.; MZSP 46337 • 1 ♂; “ Mar ” [Rio de Janeiro, Parque Nacional do Itatiaia, Maromba]; 1100 m a. s. l.; 11 Feb. 1926; Zikán leg.; NHMB • 1 ♂, 2 ♀♀; São Paulo, São José dos Barreiros, Parque Nacional Serra da Bocaina; 1400 m a. s. l.; 21 – 24 Mar. 1997; Pinto-da-Rocha, Campaner and Vanin leg.; MZSP 46338 to 46340 • 1 ♀; Minas Gerais [Passa Quatro]; 18 Mar. 22; [Zikán leg.]; NHMB • 1 ♂; “ America, Brasilia, coll. Rich Hicker ”; NHMB.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE4FFB4FD8CF88C2DE3F893.taxon	description	Re-description Body length: 10.4 – 14.0 mm. Coloration (Fig. 2): head pitch black, lustrous, except in front of antennae sockets and anterior margin of clypeus, dark brown; mandibles dark brown, darker at base and the tip; maxillary and labial palpi black, except apex of last palpomeres, dark brown; antennomeres I and sometimes II black, the latter testaceous to light brown. Pronotum (Figs 7 A, 8 A) lustrous, translucent, with broad irregular black band from anterior to posterior margin, wider posteriorly and narrower near anterior margin; background bright yellow to light orange with barely defined orange and brown lateral patches. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, large whitish to pale yellow round spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen pitch black, tarsal claws brown. Male (Fig. 2) Head short, slightly wider than long, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short; vertex flat; occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Figs 2 A, 6 A) slightly flattened dorsoventrally; antennomeres wider distally; antennomeres IX – XI distinctly narrower than preceding ones. Pronotum (Fig. 3 A) about 1.5 times as wide as long; anterior margin broadly arched, anterior angles indistinct; lateral margins with a shallow notch at posterior third, pronotum wider behind notch; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 3.4 times as long as wide, wider at apical third; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with a transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 A) broadly lobed basally, lobe with sinuous margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI deeply notched at posterior margin, V-shaped, lateral apices arched; lobes of ventrite VII (Fig. 10 A) elongate, oblong, concave, internal margins directed ventrally, distal margins rounded. Aedeagus (Fig. 11 A – C): ventral wall of tegmen short and broad, lateral margins slightly convergent, distal margin forming a pair of broadly rounded lobes; between lobes, small acute tip directed ventrally; fringe of long setae along lateral margins of tegmen dorsally; parameres broadly separated, robust, long and broad, parallel, curved ventrally, apex acute; median lobe short, membranous, partially retracted behind parameres; very long and narrow sclerite curved ventrally, apex acute. Female Similar to male; antennae (Fig. 6 L) shorter, antennomeres less broad; pronotum (Fig. 8 A) trapezoidal, much broader, 1.7 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 A) broad, lateral margins arched, distal margin sinuous, with a short and broad medial lobe. Note on nomenclature Delkeskamp (1977) and Wittmer (1977) both indicated Guérin-Méneville’s 1844 work as the original reference for Silis tricolor, since it contains the first published description of that species. Delkeskamp (1977) also cites “ Blanch. in Griffith, Anim. Kingdom 1, 1832, l. 36, fig. 10 ”, a plate published 12 years prior to Guérin-Méneville’s description. According to ICZN Art. 8.7, an illustration of a species published before 1930 that is accompanied by a new name does however already count as a valid description “ by indication ” and makes this name available, even without being accompanied by a description in words or an explicit statement that the name is new. We therefore regard the illustration in Griffith’s “ Animal Kingdom ” as the first available evidence of this species. As the species name Silis tricolor is explicitly attributed to “ Guérin ” on Griffith’s plate, F. E. Guérin-Méneville remains the author of the species (not Blanchard), but the year of description must be corrected to 1832.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE4FFB4FD8CF88C2DE3F893.taxon	distribution	Distribution Brazil (Minas Gerais, Rio de Janeiro and São Paulo states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	description	urn: lsid: zoobank. org: act: 8 B 002 EDC-E 1 E 4 - 4403 - A 388 - BA 6 E 9826 EE 10 Figs 3 A, 4 B, M, 7 B, 8 B, 9 B, 10 B, 11 D – F, 15 B	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	diagnosis	Diagnosis Similar to D. tricolor and D. vanini sp. nov. It differs from these species by the antennae entirely black, larger whitish elytral spots (Fig. 3 A), the aedeagus (Fig. 11 D – F) with distal margin of tegmen shorter, apical lobes less pronounced; the median lobe with a very broad central sclerite; females with ventrite VII (Fig. 15 B) broader, with a more pronounced median projection.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	etymology	Etymology The epithet neoteutonum refers to the type locality of the species, Nova Teutônia, a district in the Seara municipality in Santa Catarina state, Brazil.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Santa Catarina, Nova Teutônia [Seara]; 27 º 11 ′ S, 52 º 23 ′ W; 300 – 500 m a. s. l.; Apr. 1947; Fritz Plaumann leg.; MZSP 46341 (Fig. 3 A). Paratypes (43 specimens) BRAZIL • 1 ♀; same collection data as for holotype; Feb. 1951; MZSP 46342 • 1 ♂, 1 ♀; same collection data as for preceding; Mar. 1951; MZSP 46343, 46344 • 4 ♂♂, 5 ♀♀; same collection data as for preceding; 14 Apr. 1951; NHMB • 1 ♀; same collection data as for preceding; 15 Mar. 1954; NHMB • 4 ♂♂; same collection data as for preceding; 9 Mar. 1957; NHMB • 2 ♂♂; same collection data as for preceding; Feb. 1965; MZSP 46345, 46346 • 3 ♂♂; same collection data as for preceding; Feb. 1966; MZSP 46347 to 46349 • 1 ♀; same collection data as for preceding; DZUP 320980 • 1 ♀; same collection data as for preceding; CEMT CUIABA 00118354 • 1 ♀; same collection data as for preceding; Nov. 1966; MZSP 46352 • 1 ♂; same collection data as for preceding; Dec. 1966; MZSP 46353 • 1 ♀; Santa Catarina, Rio Vermelho [São Bento do Sul]; Feb. 1961; Dirings leg.; MZSP 46354 • 1 ♂; Santa Catarina, Mafra; Rttr. leg.; NHMB • 1 ♂; Santa Catharina; Hicker leg.; NHMB • 9 ♂♂, 2 ♀♀; Rio Grande do Sul, Vila Oliva [Caxias do Sul]; 18 Feb. 1949; Becker leg.; NHMB • 1 ♂; same collection data as for preceding; 21 Feb. 1948; NHMB • 2 ♂♂; same collection data as for preceding; 24 Feb. 1948; NHMB.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	description	Description Body length: 10.4 – 13.0 mm. Coloration (Fig. 3 A): head pitch black, lustrous, except in front of antennae; sockets dark brown; mandibles dark brown, darker at the base and the tip; maxillary and labial palpi black; antennae black, except last three antennomeres, sometimes lighter, and ventral face of antennomere I dark brown. Pronotum (Figs 7 B, 8 B) lustrous, translucent, with a broad irregular black band from anterior to posterior margin, wider anteriorly and narrower near the middle; background bright yellow to light orange with barely defined orange to brownish lateral patches. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, a large sulphur to pale yellow hemispheric to triangular spot nearly reaching the lateral borders but not meeting at the suture. Thorax, legs and abdomen pitch black, tarsal claws dark brown. Male (Fig. 3 A) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short; vertex flat; occipital region convex, broadly rounded behind the eyes. Clypeus flat, anterior margin emarginate, with pair of lobes slightly projected anteriorly. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 B) slightly flattened dorsoventrally; antennomeres wider distally; antennomeres IX – XI slightly narrower than the preceding. Pronotum (Fig. 7 B) about 1.5 times as wide as long; anterior margin broadly arched, anterior angles indistinct; lateral margins with a shallow notch at posterior third, pronotum wider behind notch; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron about 4 times as long as wide, almost parallel, slightly wider in the middle; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with a transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 B) with tip pointing externally, broadly lobed basally, lobe with sinuous margin; posterior claws on meso- and metathoracic tarsal claws apparently split at the apex, with fine protruding tooth shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI deeply notched at posterior margin, V-shaped, lateral apices arched; ventrite VII (Fig. 10 B) elongate, oblong, strongly concave, internal margins arcuate, directed ventrally, distal margins projecting posteriorly, apices rounded. Aedeagus (Fig. 11 D – F): ventral wall of tegmen short and broad, lateral margins slightly convergent, distal margin forming a pair of broadly rounded lobes; between lobes, small acute tip directed ventrally; fringe of long setae along lateral margins of tegmen dorsally; parameres broadly separated, robust, long and broad, parallel, curved ventrally, apex rounded; median lobe short, membranous, partially retracted behind parameres; very broad central sclerite curved ventrally with broadly rounded apex. Female Similar to male; antennae (Fig. 6 M) longer, antennomeres broader; pronotum (Fig. 8 B) trapezoidal, broader, 1.6 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 B) broad, lateral margins arched, distal margin sinuous, with broad medial lobe projecting posteriorly.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE1FFBAFDC5FA4D2D9CFFEC.taxon	distribution	Distribution Brazil (Santa Catarina and Rio Grande do Sul states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	description	urn: lsid: zoobank. org: act: BF 23 FEEE-E 09 E- 4813 - B 496 - 268277095 C 24 Figs 3 B, 6 C, N, 7 C, 8 C, 9 C, 10 C, 11 G – I, 15 C	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	diagnosis	Diagnosis Similar to D. tricolor and D. neoteutonum sp. nov. It differs from these species by the last two or three antennomeres being orange to light brown (Fig. 3 B); the ventral wall of the tegmen (Fig. 11 G – I) less distinctly narrowed laterally, the central sclerite of the median lobe broader than in D. tricolor and narrower than in D. neoteutonum; the last ventrite (VII) of females with distal margin sinuate (Fig. 15 C).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	etymology	Etymology The specific epithet is patronymic, named in honour of the late Professor Sergio A. Vanin (1948 – 2020) in appreciation of his enormous and enduring contribution in the study and teaching of zoology, especially Coleoptera systematics.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Minas Gerais, Monte Verde; 28 Feb. 1964; J. Halik leg.; MZSP 46355 (Fig. 3 B). Paratypes (59 specimens) BRAZIL • 1 ♂, 1 ♀; same collection data as for holotype; MZSP 46356, 46357 • 1 ♂; same collection data as for preceding; 5 Dec. 1964; MZSP 46358 • 1 ♂; same collection data as for preceding; 10 Feb. 1965; MZSP 46359 • 1 ♀; same collection data as for preceding; DZUP 320981 • 2 ♂♂; same collection data as for preceding; 13 – 18 Jan. 1969; MZSP 46361, 46362 • 1 ♂; same collection data as for preceding; 16 Jan. 1969; MZSP 46363 • 1 ♀; same collection data as for preceding; UFMG-ICO- 2200000 • 1 ♀; same collection data as for preceding; 22 Feb. 1969; MZSP 46365 • 1 ♂; same collection data as for preceding; Nov. 1969; MZSP 46366 • 1 ♂, 1 ♀; same collection data as for preceding; Jan. – Feb. 1970; MZSP 46367, 46368, • 1 ♀; same collection data as for preceding; CEMT CUIABA 00118355 • 1 ♂; same collection data as for preceding; 24 Feb. 1971; MZSP 46370 • 1 ♂; São Paulo, Campos do Jordão; Jan. 1954; J. Lane leg.; MZSP 46371 • 1 ♀; same collection data as for preceding; Mar. – Apr. 2018; S. P. Rosa leg.; MZSP 46397 • 1 ♂; same collection data as for preceding; 1610 m a. s. l.; 12 Feb. 1952; NHMB • 1 ♂, 1 ♀; same collection data as for preceding; 1600 m a. s. l.; Mar. 1945; Wygodzinsky leg.; NHMB • 3 ♀♀; same collection data as for preceding, Capivari; Jan. 1961; Dirings leg.; MZSP 46372 to 46374 • 1 ♀; same collection data as for preceding; 10 – 22 Feb. 1981; S. A. and A. N. Vanin leg.; MZSP 46375 • 1 ♂; São Paulo, Cantareira [São Paulo]; 23 Mar. 1907; E. Garbe leg. [7772]; MZSP 46376 • 3 ♂♂, 2 ♀♀; same collection data as for preceding; Feb. 1962; J. Halik leg.; MZSP 46377 to 46381 • 1 ♀; same collection data as for preceding; 1 Mar. 1963; MZSP 46382 • 1 ♀; same collection data as for preceding; 6 Mar. 1968; MZSP 46383 • 1 ♂, 1 ♀; São Paulo, Ipiranga; 4 Mar. 1962; L. Stowbunenko leg.; MZSP 46384, 46385 • 1 ♂; São Paulo, Mata do Governo [Parque Estadual das Fontes do Ipiranga]; 3 Apr. 1931; R. Spitz leg.; MZSP 46386 • 1 ♂; Jundiaí, Reserva Biológica Serra do Japi; 23 º 14 ′ 20 ″ S, 46 º 57 ′ 27 ″ W; 23 – 25 Jan. 2012; E. A. Nascimento, G. Biffi and F. R. Fernandes leg.; MZSP 46387 • 1 ♀; Salesópolis, Estação Biológica de Boracéia; Apr. 1948; E. X. Rabello leg.; MZSP 46388 • 1 ♂; same collection data as for preceding; 21 – 22 Mar. 1973; S. Vanin and M. Jorge leg.; MZSP 46389 • 4 ♂♂, 1 ♀; São Paulo, Alto da Serra [Paranapiacaba, Santo André]; 28 Feb. 1912; G. E. Bryant and G. Bryant leg. 1919 - 147; NHMUK • 4 ♂♂, 2 ♀♀; same collection data as for preceding; 5 Mar. 1912; NHMUK • 1 ♂, 1 ♀; same collection data as for preceding; 7 Mar. 1912; NHMUK • 1 ♂, 1 ♀; same collection data as for preceding; 9 Mar. 1912; NHMUK • 1 ♂, 1 ♀; Paranapiacaba; Feb. 1962; L. Stowbunenko leg.; MZSP 46390, 46391 • 2 ♂♂, 2 ♀♀; same collection data as for preceding; 5 Mar. 1962; MZSP 46392 to 46395 • 1 ♂; Paraná, Ponta Grossa, Distrito de Itaicoca; 23 º 07 ′ 03.7 ″ S, 49 º 56 ′ 27.8 ″ W; 24 Mar. – 12 Apr. 2017; M. D. Almeida L. Tozetto and M. H. Santos leg.; MZSP 46396.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	description	Description Body length: 9.7 – 13.0 mm. Head pitch black, lustrous, except in lateral corners of clypeus, dark brown; mandibles dark brown, darker at base and tip; maxillary and labial palpi black; antennae black, last two or three antennomeres orange to light brown. Pronotum lustrous, translucent, with broad irregular black band from anterior to posterior margin, wider posteriorly and narrower near middle; background bright yellow to light orange with well-defined diagonal orange and brown patches laterally. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, large whitish to pale yellow round spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen pitch black, tarsal claws brown. Male (Fig. 3 B) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short; vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with a median incision. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 C) slightly flattened dorsoventrally; antennomeres III – VIII wider distally; antennomeres IX – XI narrower and less expanded distally than preceding ones. Pronotum (Fig. 7 C) about 1.5 times as wide as long; anterior margin almost straight, anterior angles arched; lateral margins with shallow notch at posterior third, pronotum wider behind notch; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 4.2 times as long as wide, wider at middle; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with a transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 C) broadly lobed basally, lobe with sinuous margin; posterior claws on meso- and metathoracic tarsal claws apparently split at the apex, with fine protruding tooth shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI deeply notched at posterior margin, V-shaped, lateral apices arched; ventrite VII (Fig. 10 C) elongate, oblong, strongly concave, internal margins arcuate, directed ventrally, distal margins projecting posteriorly, apices rounded. Aedeagus (Fig. 11 G – I): ventral wall of tegmen short and broad, lateral margins slightly convergent, distal margin forming a pair of broadly rounded lobes; between lobes, small acute tip directed ventrally; fringe of long setae along lateral margins of tegmen dorsally; parameres broadly separated, robust, long and broad, parallel, curved ventrally, apex rounded; median lobe short, membranous, partially retracted behind parameres; broad central sclerite curved ventrally, with apex narrowly rounded. Female Similar to male; antennae (Fig. 6 N) shorter, antennomeres slightly broader; pronotum (Fig. 8 C) trapezoidal, narrower, 1.4 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 C) broad, lateral margins broadly arched, distal margin sinuous, with broad and shallow medial lobe.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEFFFB9FD1FFD642D89FC76.taxon	distribution	Distribution Brazil (Minas Gerais, São Paulo and Paraná states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFECFFBFFDA9FEEE2C7AFFC9.taxon	description	Figs 5 A – C, 6 D, O, 7 D, 8 D, 9 D, 10 D, 11 J – L, 15 D	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFECFFBFFDA9FEEE2C7AFFC9.taxon	diagnosis	Diagnosis Similar to D. nigrocephalum by general coloration (Fig. 5 A – B). It differs especially by the elytra being smoother (Fig. 5 A), the pronotum (Figs 7 D, 8 D) with anterior and lateral margins broadly arched, the shape of the lobe of the tarsal claws (Fig. 9 D), the ventrite VII of males (Fig. 10 D) and females (Fig. 15 D) and the aedeagus (Fig. 11 J – L). The colour of the antennae (Fig. 5 A – B) of D. obscurior is usually light brown to orangish.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFECFFBFFDA9FEEE2C7AFFC9.taxon	materials_examined	Type material Lectotype (here designated) BRAZIL • ♀; Rio de Janeiro, Nova Friburgo; Feb. – Mar. 1903; E. Gounelle leg.; “ Muséum Paris, Coll. M. Pic ”; MNHN, Paris EC 13554 (Fig. 5 A – C). Paralectotype BRAZIL • 1 ♀; same collection data as for lectotype; MNHN, Paris EC 13555. Other material examined (38 specimens) BRAZIL • 28 ♂♂, 10 ♀♀; Rio de Janeiro, Petrópolis; 5 – 7 Mar. 1962; J. Bechyné leg.; MZSP 46398 to 46435.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFECFFBFFDA9FEEE2C7AFFC9.taxon	description	Re-description Body length: 11.0 – 13.0 mm. Coloration (Fig. 5 A – B): head pitch black, lustrous, except in lateral corners of clypeus, dark brown; mandibles dark brown, darker at base and tip; maxillary and labial palpi dark brown, light brown in last palpomeres; antennomere I black, the latter light brown, sometimes orangish. Pronotum (Figs 7 D, 8 D) lustrous, partly translucent, with broad irregular black band from anterior to posterior margin, black band usually with lateral projections in posterior third; background pale yellow with diffuse orange regions. Scutellum and elytra entirely pitch black, slightly lustrous. Thorax, legs and abdomen black, tarsal claws brown. Male (Fig. 5 A – B) Head short, slightly wider than long, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes large, rounded, slightly prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 D) long, slightly flattened dorsoventrally; antennomeres III – X narrowing proximally, sub-serrate; antennomeres without longitudinal lines. Pronotum (Fig. 7 D) wide, 1.5 times as wide as long; anterior margin, anterior angles and lateral margins broadly arched; lateral margins with shallow notch at posterior third; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 4.2 times as long as wide, wider medially; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 D) broadly lobed basally, lobe with sinuous to truncate margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI notched at posterior margin; ventrite VII (Fig. 10 D) with concave lobes, inner margins divergent, directed ventrally, with sharp borders; outer margins straight, apical margins truncate, meeting medially. Aedeagus (Fig. 11 J – L): ventral wall of tegmen short, lateral margins convergent from apical half; apex forming a pair of rounded lobes and salient border projecting ventrally; fringe of long setae along lateral margins of tegmen dorsally; parameres very long, robust, falciform, projecting dorsally, apex rounded with small acute tip; median lobe membranous, with very long and broad flattened central sclerite dorsally. Female Similar to male; antenna (Fig. 6 O) longer; pronotum (Fig. 8 D) more transverse, wider, 1.6 times as wide as long, sub-rectangular, anterior margin nearly straight, lateral margins without notch; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 D) broad, lateral and distal margins sinuous, apex without strong projections, with shallow lobes not strongly projecting posteriorly.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFECFFBFFDA9FEEE2C7AFFC9.taxon	distribution	Distribution Brazil (Rio de Janeiro state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	description	urn: lsid: zoobank. org: act: 66 D 5 B 02 D-DAD 1 - 47 BB-A 93 E- 0981 BECD 4 BB 7 Figs 3 C, 6 E, P, 7 E, 8 E, 9 E, 10 E, 12 A – C, 15 E	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	diagnosis	Diagnosis Differs from other species in the region by the entirely black antennae (Fig. 3 C), the males with antennomeres IX – XI with short longitudinal lines dorsally (Fig. 6 E), ventrite VII with apices truncate (Fig. 10 E), aedeagus (Fig. 12 A – C) with three distal projections on distal margin of tegmen, the central projection very long and curved ventrally; females with last ventrite (VII) (Fig. 15 E) broadly rounded with a pair of acute projections medially.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	etymology	Etymology The epithet lineaticorne refers to the longitudinal antennal lines on antennomeres IX – XI of the males.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	materials_examined	Type material Holotype BRAZIL • ♂; São Paulo, Pindamonhangaba, Eugênio Lefèvre; 28 Sep. 1962; Exp. Dep. Zool leg.; MZSP 46436 (Fig. 3 C). Paratypes (10 specimens) BRAZIL • 1 ♀; Minas Gerais, Itamonte, Instituto Alto Montana; 22 º 21 ′ 13 ″ S, 44 º 47 ′ 57 ″ W, 5 – 6 Nov. 2019; UC. Entomologia UNIFESP leg.; Malaise; MZSP 46444 • 1 ♀; São Paulo, Campos do Jordão; 22 º 40 ′ 51.4 ″ S, 45 º 35 ′ 30.5 ″ W; 1461 m a. s. l.; Oct. – Nov. 2019; S. P. Rosa leg.; MZSP 46445 • 1 ♂; same collection data as for preceding; 1 – 13 Oct. 2018; MZSP 46446 • 1 ♀; same collection data as for holotype; DZUP 320982 • 1 ♂; same collection data as for preceding; 26 Oct. 1962; MZSP 46438 • 1 ♀; same collection data as for preceding; 1200 m a. s. l.; 21 Dec. 1962; MZSP 46439 • 1 ♂, 3 ♀♀; São Paulo, Ipiranga; Oct. 1908; Luederwaldt leg. ‘ 20371 ’ [Luederwaldt’s collection record]; MZSP 46440 to 46443.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	description	Description Body length: 13.0 – 14.3 mm. Coloration (Fig. 3 C): head pitch black, lustrous, except in lateral corners of clypeus, light brown; mandibles light brown, darker at base and tip; maxillary and labial palpi dark brown to black, light brown at apex of last palpomeres; antennae entirely black. Pronotum (Figs 7 E, 8 E) lustrous, partly translucent, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; from median band, a pair of diagonal black bands; background pale yellow to light orange with barely defined orange patches. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, large whitish to pale yellow round spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen dark brown to black, tarsal claws brown. Male (Fig. 3 C) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes large, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 E) long, slightly flattened dorsoventrally; antennomeres III – IX wider distally, sub-serrate; antennomeres IX – XI with short longitudinal lines dorsally. Pronotum (Fig. 7 E) wide, about 1.5 times as wide as long; usually, anterior margin slightly arched, almost straight in middle, anterior angles arched; lateral margins slightly sinuate, with shallow notch at posterior third; anterior, posterior and lateral margins elevated; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 5 times as long as wide, almost parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 E) broadly lobed basally, lobe with rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI deeply notched at posterior margin, V-shaped, lateral apices arched; ventrite VII (Fig. 10 E) with parallel-sided lobes, distal margins truncate; ventrite lobes concave, internal margins directed ventrally. Aedeagus (Fig. 12 A – C): ventral wall of tegmen long and broad, apical margin convergent, forming a pair of lateral projections with truncate apices and very long central projection, strongly curved ventrally and with hook-like apex, usually visible between lobes of ventrite VII; fringe of long setae along lateral margins of tegmen dorsally; parameres robust, long and broad, parallel, apex acute; median lobe short, membranous, partially retracted behind parameres. Female Similar to male; antennae (Fig. 6) shorter, without longitudinal lines; pronotum wider, 1.6 times as wide as long, lateral margins without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII broad, lateral and distal margins broadly arched, distal margin with two long apical lobes projecting posteriorly at middle and U-shaped medial notch.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFEAFFBDFDFDFC832D2BF990.taxon	distribution	Distribution Brazil (Minas Gerais and São Paulo states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	description	urn: lsid: zoobank. org: act: 0 D 454 F 22 - 6 E 27 - 402 C-ADDB- 36 D 5483 D 57 FB Figs 3 D, 6 F, Q, 7 F, 8 F, 9 F, 10 F, 12 D – F, 15 F	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	diagnosis	Diagnosis Differs from all the other species by the elytra with the weakly defined longitudinal roughness and dark yellow spots (Fig. 3 D), the shape of the aedeagus (Fig. 12 D – F) and the last ventrite (VII) of the females (Fig. 15 F).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	etymology	Etymology The specific epithet aurimaculatum refers to the dark yellowish to golden rounded elytral spots.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Pindamonhangaba, Eugênio Lefèvre; 1200 m a. s. l.; 21 Dec. 1962; Exp. Dep. Zool. leg.; MZSP 46447 (Fig. 3 D). Paratypes (12 specimens) BRAZIL • 2 ♀♀; Rio de Janeiro, Itatiaia; 1100 m a. s. l.; Feb. 1963; MZSP 46448, 46449 • 2 ♀♀; same collection data as for preceding; Jan. 1968; MZSP 46450, 46451 • 1 ♂; same collection data as for preceding; 1300 m a. s. l.; Nov. 1950; NHMB • 1 ♀; same collection data as for preceding, Véu da Noiva, córrego Maromba; 22 º 26 ′ 10 ″ S, 44 º 37 ′ 28 ″ W; 1100 m a. s. l.; 2 – 3 Feb. 1997; Kury, Rocha and Mestre leg.; MZSP 46452 • 1 ♂, 2 ♀♀; same collection data as for holotype; MZSP 46453, 46454, 46457 • 1 ♀; same collection data as for holotype; DZUP 320983 • 1 ♀; same collection data as for holotype; CEMT CUIABA 00118356 • 1 ♀; same collection data as for holotype; 24 Jan. 1963; MZSP 45593.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	description	Description Body length: 11.7 – 13.0 mm. Coloration (Fig. 3 D): head pitch black, lustrous, except in lateral corners of clypeus, light brown; mandibles light brown, darker at tip; maxillary and labial palpi black, light brown at apex of last palpomeres; antennae entirely black. Pronotum (Figs 7 F, 8 F) lustrous, partly translucent, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; no lateral dark bands or patches; background pale yellow to light orange with barely defined orange regions. Scutellum and elytra pitch black; at mid-length of each elytron, pale-yellow round spot meeting lateral borders but not meeting at suture. Thorax, legs and abdomen dark brown to black, tarsal claws brown. Male (Fig. 3 D) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with shallow median incision. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 F) long, slightly flattened dorsoventrally; antennomeres slightly narrowing anteriorly, sub-serrate; antennomeres without longitudinal lines dorsally. Pronotum (Fig. 7 F) wide, about 1.4 times as wide as long; anterior margin slightly arched, anterior angles arched; lateral margins slightly sinuate, with deep notch at posterior third; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 4.8 times as long as wide, almost parallel; weakly defined longitudinal roughness, like costae, more visible anteriorly; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 F) broadly lobed basally, lobe with rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI slightly notched at posterior margin; ventrite VII (Fig. 10 F) with broad lobes, internal margins arched externally, apical margin broadly rounded. Aedeagus (Fig. 12 D – F): ventral wall of tegmen elongate, apical margins convergent, forming a pair of parallel lateral projections with rounded apices and a short central projection, strongly curved ventrally and with acute hook-like apex; fringe of long setae along lateral margins of tegmen dorsally; parameres very short, hidden between tegmen and median lobe; median lobe long, membranous, with a pair of strong sclerites, each with double acute apices; sides of median lobe with tuft of thick spine-like setae. Female Similar to male; antennae (Fig. 6 Q) shorter; pronotum (Fig. 8 F) wider, 1.6 times as wide as long, lateral margins without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 F) broad, lateral and distal margins broadly arched, distal margin with two short acute apical lobes projecting posteriorly at middle and V-shaped medial notch.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFE8FFA3FDD6FB482D37F969.taxon	distribution	Distribution Brazil (Rio de Janeiro and São Paulo states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	description	urn: lsid: zoobank. org: act: CECFCC 8 E- 713 B- 4 A 5 F-B 6 B 0 - 9995 F 9 A 776 B 3 Figs 3 E, 6 G, R, 7 G, 8 G, 9 G, 10 G, 12 G – I, 15 G	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	diagnosis	Diagnosis Similar to D. neoteutonum sp. nov. by the shape and coloration of the elytral spots. It differs from this species by the antennae being entirely black (Fig. 3 E), with the antennal lines in males (Fig. 3 G), the pronotum almost entirely black (Figs 7 G, 8 G), and the aedeagus with distal margin of the tegmen with three projecting lobes (Fig. 12 G – I); females with ventrite VII broad, with a sinuous distal margin and a less pronounced median projection (Fig. 15 G). The aedeagus of D. marginicolle sp. nov. is similar to that of D. aurimaculatum sp. nov., differing in the shape of the distal projections of the ventral wall of the tegmen; besides, the antennae of D. aurimaculatum do not have antennal lines and the elytral spots are rounder.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	etymology	Etymology The specific epithet refers to the barely defined dark-orangish spot in the margin of the pronotum.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Rio de Janeiro, Nova Friburgo, Macaé de Cima; Jan. 2000; P. Grossi leg.; CEMT CUIABA 00118359 (Fig. 3 E). Paratypes (4 specimens) BRAZIL • 1 ♂; same collection data as for holotype; MZSP 46458 • 2 ♀♀; Nova Friburgo, Mury; 1050 m a. s. l.; Mar. 1941; NHMB • 1 ♀; Terezópolis; Jan. 1980; A. Bello leg.; CEMT CUIABA 00118360.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	description	Description Body length: 11.0 – 13.0 mm. Coloration (Fig. 3 E): head pitch black, lustrous, except in corners of clypeus, dark brown; mandibles dark brown, darker at tip; maxillary and labial palpi black and antennae entirely black. Pronotum (Figs 7 G, 8 G) lustrous, mostly black, except for orangish to dark brown irregular patches near anterior and posterior angles. Scutellum and elytra pitch black; at mid-length of each elytron, large sulphur to greyish hemispheric to triangular spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen pitch black, tarsal claws dark brown. Male (Fig. 3 E) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short; vertex flat; occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, with a pair of lobes slightly projected anteriorly. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 G) sub-serrate, slightly flattened dorsoventrally; antennomeres III – VI wider distally, the latter nearly parallel, antennomere XI narrowing apically; antennomeres IX – XI with short longitudinal lines dorsally. Pronotum (Fig. 7 G) about 1.1 times as wide as long; anterior margin slightly arched, anterior angles rounded; lateral margins with shallow notch at posterior third, pronotum wider near notch; integument smooth, densely covered with very fine clear setae. Elytra long, each elytron about 5.5 times as long as wide, almost parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 G) with pointing tip, broadly lobed basally, lobe with rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI broadly concave at posterior margin; ventrite VII (Fig. 10 G) elongate, halves triangular, internal margins straight, lateral margins arched, apices rounded. Aedeagus (Fig. 12 G – I): ventral wall of tegmen elongate, apical margins convergent, forming a pair of parallel lateral projections with acute apices and central projection, strongly curved ventrally and with acute hook-like apex; fringe of long setae along lateral margins of tegmen dorsally; parameres very short, hidden between tegmen and median lobe; median lobe long, conical, membranous, with a pair of strong sclerites, each with double acute apices; sides of median lobe with a tuft of thick spine-like setae. Female Similar to male; antennae (Fig. 6) shorter, antennomeres broader, without antennal lines; pronotum (Fig. 8) trapezoidal, broader, 1.6 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VI with distal margin slightly arched, concave; ventrite VII (Fig. 15 G) broad and short, distal margin sinuous, with a broad medial lobe projecting posteriorly.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF6FFA0FDF9FBE32C7AFBA0.taxon	distribution	Distribution Brazil (Rio de Janeiro state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	description	urn: lsid: zoobank. org: act: 7 DC 62 ADA-E 43 F- 4874 - AB 7 D- 97 EED 692 DB 78 Figs 5 D, 6 H, S, 7 H, 8 H, 9 H, 10 H, 12 J – M, 15 H	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	diagnosis	Diagnosis Similar to D. obscurior and D. nigrocephalum by the general coloration. It differs especially by the weakly defined elytral costulae (Fig. 5 D) visible under unidirectional lighting, the shape of the pronotum (Figs 7 H, 8 H), which is less arched anteriorly, the tarsal claws of the males (Fig. 9 H), ventrite VII of the males (Fig. 10 H) and females (Fig. 15 H) and the aedeagus (Fig. 12 J – M).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	etymology	Etymology The name tenuecostatum refers to the weakly defined longitudinal elytral costulae in both males and females.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Rio de Janeiro, Teresópolis, Parque Nacional da Serra dos Órgãos; 4 – 9 Nov. 2013; V. S. Ferreira and F. F. Barbosa leg.; MZSP 46459 (Fig. 5 D). Paratypes (11 specimens) BRAZIL • 3 ♂♂, 1 ♀; same collection data as for holotype; MZSP 46460, 46461, 46463, 46466 • 1 ♀; same collection data as for holotype; UFMG-ICO- 2200001 • 1 ♀; same collection data as for holotype; DZUP 320984 • 1 ♀; same collection data as for holotype; CEMT CUIABA 00118357 • 1 ♂, 1 ♀; same collection data as for preceding, Casa do Pesquisador; 16 Dec. 2016; Simeão Moraes leg.; MZSP 46467, 46468 • 1 ♀; [Parque Nacional do] Itatiaia; Nov. 1966; Dirings col.; MZSP 46469 • 1 ♀; Rio de Janeiro, Serra do Macaé; Nov. 1909; E. Garbe leg. [15,235]; MZSP 45594.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	description	Description Body length: 11.7 – 16.2 mm. Coloration (Fig. 5 D): head pitch black, lustrous, except in lateral corners and anterior margin of clypeus, dark brown; mandibles dark brown, darker at base and tip; maxillary and labial palpi dark brown, light brown in last palpomeres; antennae black at base, clearing apicad, antennomeres I – II black, III – V dark brown, VI – XI light brown, sometimes III – V light brown and the latter orangish. Pronotum (Figs 7 H, 8 H) lustrous, partly translucent, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; background pale yellow with diffuse orange regions. Scutellum and elytra entirely pitch black, slightly lustrous. Thorax, legs and abdomen black, tarsal claws brown. Male Head short, as long as wide, excluding eyes; integument smooth, densely covered with short and fine clear setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with rounded median incision. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 H) long, reaching elytral apical third, slightly flattened dorsoventrally; antennomeres narrowing proximally, sub-serrate, without longitudinal lines. Pronotum (Fig. 7 H) wide, about 1.5 times as wide as long; anterior margin slightly arched, anterior angles arched; lateral margins sinuate, almost parallel anteriorly, and widened before lateral deep notch at posterior third; integument smooth, densely covered with very fine yellow setae. Elytra very long, each elytron 5.6 times as long as wide, almost parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 H) broadly lobed basally, lobe with very broad rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI with posterior margin concave; ventrite VII (Fig. 10 H) with triangular concave lobes, inner margins nearly parallel, directed ventrally, outer margins convergent posteriorly, apical margins rounded, with tip directed internally. Aedeagus (Fig. 12 J – M) very robust, ventral wall of tegmen elongate, lateral margins constricted apically; apex forming a pair lobes with rounded projections, with strongly sinuous aspect; broad central projection, strongly flattened laterally with flat plaque-like projection ventrally, usually visible between ventrite lobes; fringe of long setae along lateral margins of tegmen dorsally; parameres very short, hidden between tegmen and median lobe; median lobe membranous, with strong falciform sclerites directed dorsally and central tufts of thick pubescence. Female Similar to male; antennae (Fig. 6 S) shorter; pronotum (Fig. 8 H) wider, 1.7 times as wide as long, subrectangular, anterior margin nearly straight, lateral margins without notch; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 H) long and broad, lateral and distal margins broadly arched, apex with two long and broad apical lobes projecting posteriorly at middle and U-shaped medial notch.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF5FFA7FDCDF9382C7AFBB7.taxon	distribution	Distribution Brazil (Rio de Janeiro state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF2FFA4FDDFF9232C49FEB5.taxon	description	Fig. 5 E – G	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF2FFA4FDDFF9232C49FEB5.taxon	diagnosis	Diagnosis Similar to D. obscurior and D. tenuecostatum sp. nov. by the general coloration. It differs from these species by the wider head (Fig. 5 F), apparently nearly as wide as the pronotum, the antennae black, and the elytra coriaceous, rough, without costulae (Fig. 5 E).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF2FFA4FDDFF9232C49FEB5.taxon	materials_examined	Type material Lectotype (here designated) BRAZIL • ♂; São Paulo, Vale do Rio Pardo; Dec. 1898; E. Gounelle leg.; “ Museum Paris Coll. M. Pic ”; MNHN, Paris EC 13556 (Fig. 5 E – G).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF2FFA4FDDFF9232C49FEB5.taxon	discussion	Remarks We were not able to examine the type specimen of D. nigrocephalum nor found similar specimens that could be attributed to this species. However, the photographs of the lectotype, deposited at the MNHN, Paris, enable the distinction between D. nigrocephalum on the one hand and D. obscurior and D. tenuecostatum sp. nov. on the other hand.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF2FFA4FDDFF9232C49FEB5.taxon	distribution	Distribution Brazil (São Paulo state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	description	urn: lsid: zoobank. org: act: A 319 DC 64 - B 2 FD- 4853 - A 80 E-CE 88 D 46 B 5 BC 2 Figs 3 F, 6 I, T, 7 I, 8 I, 9 I, 10 I, 13 A – C, 15 I	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	diagnosis	Diagnosis Antennae entirely black (Fig. 3 F), last ventrite (VII) of males with a strongly prominent acute tip directed ventrally (Fig. 10 I); the last ventrite of the females (VII) with a distal margin with two apical rounded lobes projecting posteriorly at middle (Fig. 15 I).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	etymology	Etymology The specific epithet refers to the Tamoio indigenous people, that inhabited the same region as D. tamoio sp. nov., currently comprising part of the coast of São Paulo and Rio de Janeiro states, Brazil.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	materials_examined	Type material Holotype BRAZIL • ♂; São Paulo, Salesópolis, Estação Biológica de Boracéia; 23 º 39 ′ 14 ″ S, 45 º 53 ′ 25 ″ W, 24 – 28 Mar. 2011; Expedição MZUSP / FFCLRP leg.; MZSP 46470 (Fig. 3 I). Paratypes (14 specimens) BRAZIL • 1 ♂; São Paulo, Salesópolis, Estação Biológica de Boracéia; 23 º 39 ′ 15.63 ″ S, 45 º 53 ′ 22.38 ″ W, 8 – 12 Mar. 2013; F. F. Albertoni leg.; lâmpada mista [mist lamp]; MZSP 46472 • 1 ♂; same collection data as for preceding; DZUP 320985 • 1 ♂; same collection data as for preceding; 23 º 39 ′ 14.5 ″ S, 45 º 53 ′ 21.7 ″ W; 16 Mar. 2008; F. R. Fernandes leg.; MZSP 46474 • 1 ♂; same collection data as for preceding; UFMG-ICO- 2200002 • 1 ♀; same collection data as for preceding; 23 º 39 ′ 02.6 ″ S, 45 º 53 ′ 32.1 ″ W; 23 – 27 Mar. 2012; F. F. Albertoni leg.; MZSP 46476 • 1 ♂; same collection data as for preceding; 23 º 39 ′ 14 ″ S, 45 º 53 ′ 25 ″ W; 24 – 28 Mar. 2011; Expedição MZUSP / FFCLRP leg.; MZSP 46471 • 1 ♂; same collection data as for preceding; 16 – 19 Feb. 2018; Expedição MZUSP leg.; MZSP 46477 • 1 ♀; Ubatuba, Parque Estadual da Serra do Mar, Núcleo Picinguaba; 3 – 14 Mar. 2008; F. Esteves and R. Feitosa leg.; MZSP 46478 • 1 ♂; Caraguatatuba, Reserva Florestal; 40 m a. s. l.; 2 Apr. 1962; Exp. Dep. Zool. leg.; MZSP 46479 • 1 ♂; same collection data as for preceding; 2 Apr. 1962; U. R. Martins, H. Reichardt and Silva leg.; MZSP 46480 • 1 ♀; same collection data as for preceding; CEMT CUIABA 00118358 • 1 ♂; São Sebastião; Dec. 1955; A. P. Silva leg.; Ferraciolli leg.; MZSP 46482 • 2 ♂♂; Rio de Janeiro, Parque Nacional do Itatiaia; 700 m a. s. l.; 6 Apr. 1947; W. Zikán leg.; NHMB.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	description	Description Body length: 10.4 – 13.6 mm. Coloration (Fig. 3 F): head pitch black, lustrous, except in lateral corners of clypeus, dark brown; mandibles dark brown, darker at base and tip; maxillary and labial palpi black; antennae entirely black. Pronotum (Figs 7 I, 8 I) lustrous, translucent, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; background pale yellow to light orange with barely defined orange and brown lateral patches. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, large whitish to pale yellow round spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen pitch black, tarsal claws brown. Male (Fig. 3 F) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes small, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 I) slightly flattened dorsoventrally; antennomeres III – IX wider distally, subserrate; antennomeres IX – XI slightly narrower than preceding ones. Pronotum (Fig. 7 I) variable, about 1.5 times as wide as long; usually, anterior margin slightly arched, almost straight in middle, anterior angles arched; lateral margins slightly sinuate, with deep notch at posterior third; sometimes lateral margins less sinuate and lateral notches shallow; integument smooth, densely covered with very fine yellow setae. Elytra long, each elytron 4.5 times as long as wide, almost parallel, wider at middle; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 I) broadly lobed basally, lobe with truncate margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI deeply notched at posterior margin, V-shaped, lateral apices arched; ventrite VII (Fig. 10 I) with broad lobes, internal and apical margins forming acute tip strongly projecting ventrally. Aedeagus (Fig. 13 A – C) triangular, ventral wall of tegmen parallel at base and strongly narrowing apically, apex acute; fringe of long setae along lateral margins of tegmen dorsally; parameres robust, very long, sinuous, apices acute, convergent; median lobe short, membranous, partially retracted behind parameres; central sclerite very long and slender, curved dorsally. Female Similar to male; antennomeres (Fig. 6 T) broader; pronotum (Fig. 8 I) trapezoidal, wider, 1.6 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VI not notched, distal margin slightly arched, concave; ventrite VII (Fig. 15 I) broad, lateral and distal margins broadly arched, distal margin with two apical rounded lobes projecting posteriorly at middle.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF1FFAAFD1AFC2F2D37FEC3.taxon	distribution	Distribution Brazil (São Paulo and Rio de Janeiro states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	description	urn: lsid: zoobank. org: act: 07 CD 44 CC-D 82 E- 44 B 0 - A 547 - 97 C 1 F 392 D 997 Figs 4 A, 6 J, U, 7 J, 8 J, 9 J, 10 J, 13 D – F, 15 J	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	diagnosis	Diagnosis Similar to D. vanini sp. nov. by the last antennomeres orangish (Fig. 4 A), but differs by the antennae shorter with longitudinal lines on the antennomeres IX – XI (Fig. 6 J, U), the pronotum with lateral margins not elevated (Figs 7 J, 8 J), by the elongate elytra (Fig. 4 A), and by the shape of ventrite VII of males (Fig. 10 J) and the aedeagus (Fig. 13 D – F). The single known male specimen of D. viridimontanum sp. nov. was fixed with an everted internal sac, which exacerbates the morphological comparison with other species.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	etymology	Etymology The specific epithet viridimontanum refers to the type locality of the species, Monte Verde (Minas Gerais state, Brazil), which translates to ‘ green hill’ from Portuguese.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	materials_examined	Type material Holotype BRAZIL • ♂; Minas Gerais, Monte Verde; 22 Feb. 1960; J. Halik leg.; MZSP 46483 (Fig 4 A). Paratype BRAZIL • 1 ♀; same collection data as for holotype; 2 Feb. 1970; MZSP 46484.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	description	Description Body length: 10.4 mm. Coloration (Fig. 4 A): head pitch black, lustrous, except in lateral corners of clypeus, light brown; mandibles light brown, darker at base and tip; maxillary and labial palpi dark brown to black, light brown at apex of last palpomeres; antennae black, except antennomeres IX – XI and apex of VIII, orangish. Pronotum (Figs 7 J, 8 J) lustrous, partly translucent, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; background pale yellow with barely diffuse orange patches. Scutellum and elytra pitch black, slightly lustrous; at mid-length of each elytron, rounded whitish spot nearly reaching lateral borders but not meeting at suture. Thorax, legs and abdomen dark brown to black, tarsal claws brown. Male (Fig. 4 A) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes large, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 J) short, slightly flattened dorsoventrally; antennomeres III – IX narrowing proximally, sub-serrate; antennomeres IX – XI with longitudinal lines dorsally. Pronotum (Fig. 7 J) wide, about 1.4 times as wide as long; anterior margin and anterior angles arched; lateral margins slightly sinuate, with shallow notch at posterior third; anterior, posterior and lateral margins not elevated; integument smooth, densely covered with very fine yellow setae. Elytra very long, each elytron 5.7 times as long as wide, almost parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with a transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 J) broadly lobed basally, lobe with very broad rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, coriaceous, densely covered with long setae; ventrite VI notched at posterior margin; ventrite VII (Fig. 10 J) with triangular lobes, inner margins nearly parallel, outer margins convergent posteriorly, apical margins rounded, with tip directed internally. Aedeagus (Fig. 13 D – F): ventral wall of tegmen elongate, lateral margins convergent from apical third; apex forming a pair of short, rounded lobes with central acute tip curved ventrally; fringe of long setae along the lateral margins of tegmen dorsally; parameres very short, hidden between tegmen and median lobe; median lobe membranous, with numerous spine-like sclerites and tufts of thick pubescence. Female Similar to male; antennae (Fig. 6 U) shorter; pronotum (Fig. 8 J) wider, 1.5 times as wide as long, lateral margins sinuate, without notches; tarsal claws without basal lobe or apical slit; ventrite VII (Fig. 15 J) with distal margin broadly arched, without projections or notches.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFEFFAEFDD3FE9E2C6EFFE7.taxon	distribution	Distribution Brazil (Minas Gerais state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	description	Figs 4 B – C, 6 K, 7 K, 9 K – L, 10 K, 13 G – J	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	diagnosis	Diagnosis Species readily distinguished by the strong modifications on the metathoracic legs (Figs 4 B, 9 L), the shape of the pronotum (Fig. 7 K), ventrite VII of the male (Fig. 10 K) and the aedeagus (Fig. 13 G – J).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	materials_examined	Type material Holotype BRAZIL • ♂; [Rio de Janeiro, Parque Nacional da] Serra dos Órgãos; 1200 m a. s. l.; 5 Feb. 1952; NHMB CANTH 00019527 (Fig. 4 B – C). Paratype BRAZIL • ♂; same collection data as for holotype; NHMB. Other material examined (1 specimen) BRAZIL • 1 ♂; Rio de Janeiro, Teresópolis; 13 – 16 Dec. 1957; Seabra and Alvarenga leg.; DZUP 376213.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	description	Re-description Body length: 13.0 mm. Coloration (Fig. 4 B): head black, lustrous, except in lateral corners of clypeus, orangish; mandibles testaceous at base, gradually darker apicad; maxillary and labial palpi black, light brown at apex of last palpomeres; antennae entirely black. Pronotum (Fig. 7 K) lustrous, partly translucent, with broad irregular dark brown to black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; background pale yellow with diffuse orange patches. Scutellum and elytra dark brown to black, slightly lustrous; at mid-length of each elytron, long, irregular yellow to orangish spot extending from lateral borders to suture. Thorax, legs and abdomen dark brown to black, tarsal claws brown. Male (Fig. 4 B) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine yellow setae; frons short, vertex flat, occipital region convex, broadly rounded behind eyes. Clypeus flat, anterior margin emarginate, slightly projected anteriorly with median incision. Eyes large, rounded, prominent. Mandibles falciform, acute, without accessory teeth. Last maxillary and labial palpomeres securiform. Antennae (Fig. 6 K) long, distinctly flattened dorsoventrally; antennomeres III – IX slightly narrowing proximally, sub-serrate; antennomeres VIII – XI with longitudinal lines dorsally. Pronotum (Fig. 7 K) wide, about 1.3 times as wide as long; anterior margin and anterior angles arched; lateral margins nearly parallel anteriorly, with rounded lobe, followed by deep notch at posterior third and broadly rounded lobe posteriorly; anterior, posterior and lateral margins elevated; integument smooth, densely covered with very fine yellow and black setae. Elytra very long, each elytron 5.5 times as long as wide, almost parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs densely pubescent, covered with long and thick setae; fore leg slender; mid femur slightly arcuate at base and swollen at apex, mid tibia curved at apex; hind leg (Fig. 9 L) strongly modified, coxa with strong unciform projection, trochanter oblong, femur strongly curved and swollen, narrower proximally, tibia widened and flattened distally; all tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base; anterior prothoracic tarsal claws (Fig. 9 K) broadly lobed basally, lobe with very broad rounded margin; posterior claws on meso- and metathoracic tarsal claws apparently split at apex, with fine protruding tooth slightly shorter than claws. Abdomen weakly sclerotised, especially at middle of the ventrites, coriaceous, densely covered with long setae; ventrite VI notched at posterior margin, U-shaped; ventrite VII (Fig. 10 K) strongly notched at inner margins, exposing part of aedeagus, inside densely covered with thick and long yellow setae, brush-like, outer margins arched posteriorly, apical margins acute, with tip directed internally. Aedeagus (Fig. 13 G – J) partly exposed between halves of ventrite VII; ventral wall of tegmen short and broad, with elongate central projection curved dorsally; lateral margins of ventral wall rounded and divergent, with long and thick setae on distal margin and inside; central projection parallel and with a pair of longitudinal ridges proximally, and widened, flattened, with small triangular spine distally, apex with longitudinal slit; parameres short and stout, apex truncate; median lobe short, membranous, with a pair of strong triangular dorsal sclerites and brush-like projections laterally and ventrally. Female Unknown.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	distribution	Distribution Brazil (Rio de Janeiro state) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFFBFFADFDD7FD702FB8FCFD.taxon	discussion	Remarks Wittmer (1952) erected the subgenus Acanthodiscodon to include the single species D. (A.) crassipes, defined by the strong modifications in the hind legs. Given the vast morphological variation seen in Neotropical species of Discodon, especially in the aedeagus structures, it is not justifiable to separate a single species D. crassipes in its own subgenus Acanthodiscodon, which is herein synonymised with Discodon (syn. nov.).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	description	Figs 4 D – G, 7 K, 14 A – C	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	diagnosis	Diagnosis Distinguished from the other species by the elytra black with unclearly defined lighter spots and the legs reddish brown with black tarsi (Fig. 4 D, F).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	materials_examined	Type material Syntypes (2 specimens) BRAZIL • 1 ♂; [Minas Gerais, Passa Quatro]; “ 16 / 12. / 1915 // ex coll. Zikan / Bras. ”; “ Naturhistorisches Museum Basel, Coll. W. Wittmer ”; 16 Dec. 1915; NHMB • 1 ♀; Rio Grande do Sul; “ coll. Richard Hicker, Wien ”; “ Naturhistorisches Museum Basel, Coll. W. Wittmer ”; NHMB (Fig 4 D – G).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	description	Re-description Body length: 12.0 mm. Coloration: head black, lustrous, except in lateral corners of clypeus, orangish; maxillary palpi dark brown; antennomeres I – VIII dark brown, IX – XI orangish. Pronotum (Fig. 8 K) lustrous, with broad irregular black band from anterior to posterior margin, wider anteriorly and near posterior margin, and narrower near anterior half; background testaceous with diffuse orange patches. Scutellum and elytra black, slightly lustrous; at mid-length of each elytron, small and weakly defined round yellowish spot near lateral margins. Legs reddish brown, tarsomeres II – V black. Male (Fig. 4 D – E) Head short, nearly as long as wide, excluding eyes; integument smooth, densely covered with short and fine setae; frons short. Eyes large, rounded, prominent. Last maxillary palpomeres securiform. Antennae long, probably reaching the middle of elytra, slightly flattened dorsoventrally; antennomeres III – IX slightly narrowing proximally, sub-serrate. Pronotum wide, about 1.5 times as wide as long; anterior margin and anterior angles broadly arched; lateral margins sinuate broader posteriorly; anterior, posterior and lateral margins elevated; integument smooth, densely covered with very fine setae. Elytra long, each elytron 5 times as long as wide, nearly parallel; integument coriaceous, densely covered with short and fine decumbent setae, and much longer thick erect setae. Legs slender, densely pubescent, covered with long and thick setae; tarsi flattened dorsoventrally, fourth tarsomere with transversal slit at base. Aedeagus (Fig. 14 A – C): ventral wall of tegmen elongate, lateral margins convergent from apical third; apex forming a pair of short, rounded lobes with central acute tip curved ventrally; fringe of long setae along lateral margins of tegmen dorsally; parameres very short, hidden between tegmen and median lobe; median lobe membranous, with numerous spine-like sclerites and tufts of thick pubescence. Female Similar to male; pronotum (Fig. 8 K) wider, lateral margins without notches; tarsal claws without basal lobe or apical slit; ventrite VII (Fig. 15 K) broad, lateral and distal margins broadly arched, distal margin with two apical rounded lobes projecting posteriorly at middle.	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	distribution	Distribution Brazil (Minas Gerais and Rio Grande do Sul states) (Fig. 16).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
03CEA025FFF8FF93FDB0FD962B27FF3E.taxon	discussion	Remarks This taxon was described as a variety (“ v. ”) of D. albonotatum by Pic (1930). Pic did not specify the number of specimens examined, but he stated that they were “ ex Hicker ”, i. e., from the collection of his contemporary Richard Hicker (1872 – 1965). Hicker’s Cantharidae collection was acquired by Walter Wittmer and is now at NHMB. Two specimens matching Pic’s description were found there. A female from “ Rio Grande do Sul ” labelled as coming from R. Hicker’s collection and a male with a handwritten label by Zikan from W. Wittmer’s collection, which was likely acquired by W. Wittmer via R. Hicker and is therefore here included as a syntype. During a visit at MNHN, and on photos of M. Pic’s Discodon boxes kindly provided by R. Constantin, no specimen attributable to D. testaceipes was found. The name D. albonotatum testaceipes is here treated as subspecific and available in accordance with ICZN Art. 45.6.4, as it is currently widely practised for names described as “ var. ” by this author. But based on the differences outlined above, we conclude that it should be treated as a full species (stat. nov.).	en	Biffi, Gabriel, Geiser, Michael (2022): A revision of Discodon tricolor (Guérin-Méneville) and its mimics from the Atlantic forests of Brazil (Coleoptera: Cantharidae). European Journal of Taxonomy 834 (1): 148-189, DOI: https://doi.org/10.5852/ejt.2022.834.1907, URL: http://zoobank.org/c2df7ac2-3d99-43ff-bb36-cef8e8747160
