identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E20D10A853FFD9FF71FE4DFCFFFC69.text	03E20D10A853FFD9FF71FE4DFCFFFC69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratophysella brevisensillata Yosii 1961	<div><p>Ceratophysella brevisensillata Yosii, 1961</p> <p>Figs 1-9</p> <p>MATERIAL EXAMINED: 10 paratypes (?) on slides, formerly in alcoholic vial labeled: „ Hypogastrura (Ceratophysella) pseudarmata (Folsom) det. Yoshii, USA (Massachusetts), Arlington 15.XI.1950, leg. Bonet, on rain pools” (MNHG).</p> <p>REDESCRIPTION: Body length 1-1.3 mm. Color in alcohol grey to dark grey. Eye patches black, anal spines light. Granulation fine and uniform, with 14-22 granules between setae p 1 on abd. V. Dorsal chaetotaxy of B type, macrosetae p 2 on th. II-III set nearly in line with setae p 1, setae m 3 and m 4 on th. II usually present, setae a 2 longer than a 3, setae m 6 absent, setae p 1 and p 2 on abd. IV macro- and microsetae respectively, setae p 3 present (Figs 1-2). Arrangement of setae on head typical for the genus. Differentiation of dorsal setae into micro- and macrosetae distinct. Setae long, pointed, slightly curved and serrated (Fig. 3). Body sensilla (s) p 4 on th. II-III and p 5 on abd. I short, about 1/3-1/2 of microsetae. Body sensilla on abd. II-V and lateral parts of th. II-III long, but shorter than macrosetae (Fig. 3). Microsensilla (ms) on th. II present. Subcoxae I-III with 1, 2, 3 setae respectively.</p> <p>Ant. IV with simple apical vesicle, subapical organite (or), microsensillum (ms), 7 cylindrical sensilla (2 lateral and 5 dorsal), about 15-20 short curved flattened at tips sensilla in ventral file (Fig. 4). Ant. III-organ with two long (lateral) and two short (internal) curved sensilla. Microsensillum on ant. III present. Eversible sac between ant. III-IV present. Ant. I with 7 setae.</p> <p>Ocelli 8 + 8. Postantennal organ about twice as large as single ocellus, with four lobes of which the anterior pair larger than the posterior. Accessory boss present (Fig. 5).</p> <p>Labrum with 5, 5, 4 setae and without apical papillae. 4 prelabrals present. Labium and head of maxilla (Fig. 6) of the C. armata type. Outer lobe with 2 sublobal hairs.</p> <p>Tibiotarsi I, II, III with 19, 19, 18 setae respectively, tibiotarsal tenent hairs slightly longer than inner edge of claws and pointed. Claws with inner tooth and pair of indistinct lateral teeth. Empodial appendage with broad basal lamella and apical filament reaching inner tooth or slightly beyond (Fig. 9).</p> <p>Ventral tube with 4 + 4 setae.</p> <p>Furca well developed (Fig. 8). Dens/mucro ratio = ca. 2. Dens with 7 setae (2-4 inner modified). Mucro boat-like. Retinaculum with 4 + 4 teeth.</p> <p>Anal spines short, half as long as inner edge of claws III, situated on basal papillae (Figs 2, 7).</p> <p>REMARKS: The species was originally described from the north-eastern part of the USA. Later it was recorded from several localities within the same region (Christiansen &amp; Bellinger, 1980), from Alaska and Chukotka (Fjellberg, 1985) and Siberia (Babenko et al., 1994). Alaskan and East Palaearctic specimens treated as C. brevisensillata clearly differ from the above redescription of Yosii’s types by having longer anal spines and coarser integument granulation. They could be considered as a separate species if their morphology was homogeneous within the area. Unfortunately it is not so and many important diagnostic characters vary in different parts of the distributional range without forming clear geographical pattern. Thus, inner margin of claw III/anal spine ratio varies in Palaearctic specimens from 1.8: 1 to 0.9: 1, being usually about 1: 1. Yosii’s “a” measure is usually 10-12, but the whole range is 9-16. Chaetom differentiation into macro and microsetae is strong in most Palaearctic regions but specimens from Kemerovo Province and eastern Tuva are characterized by weak differences in seta length with a 2 on th. II almost as long as a 1 and a 3. Intermediary conditions have been also seen. The single available specimen from the Kyrgyz mountains has rather short but strong (almost spine-like) macrosetae. Alaskan and Chukotka specimens have 2 sublobals on maxillary outer lobe and p 3 setae on abd. IV are usually absent. Populations from more western parts of Palaearctic differ by having only 1 sublobal hair and both p 2 and p 3 microsetae present on abd. IV. Nevertheless, two sublobals have been also seen in populations from Buryatia, eastern Tuva, north-eastern Altai and Kyrgyzstan and specimens without p 3 on one or both sides on abd. IV can be found within many studied populations. Just now we prefer to treat all these forms as a single polymorphic species with a wide Siberian-American distributional range being sure that more work and material are needed to clear up their status. However the existing distributional gap (see Fig. 21) between east and west American populations raises some doubts that they are conspecific.</p> </div>	http://treatment.plazi.org/id/03E20D10A853FFD9FF71FE4DFCFFFC69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Babenko, Anatoly;Skarżyński, Dariusz	Babenko, Anatoly, Skarżyński, Dariusz (2011): Ceratophysella lobata sp. n. from Siberia with notes on C. brevisensillata Yosii, 1961 (Collembola: Hypogastruridae). Revue suisse de Zoologie 118 (2): 257-264, DOI: 10.5962/bhl.part.117808, URL: http://www.biodiversitylibrary.org/part/117808
03E20D10A851FFDCFF71FC12FCE9FC09.text	03E20D10A851FFDCFF71FC12FCE9FC09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratophysella lobata Babenko & Skarżyński 2011	<div><p>Ceratophysella lobata sp. n.</p> <p>Figs 10-20</p> <p>Hypogastrura (Ceratophysella) brevisensillata: Fjellberg (1985): 37</p> <p>Hypogastrura (Ceratophysella) cf. brevisensillata: Babenko et al. (1994): 127</p> <p>MATERIAL EXAMINED: Holotype, female, Russia, Jakutia (Sakha Republic), <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.53334&amp;materialsCitation.latitude=63.216667" title="Search Plazi for locations around (long 139.53334/lat 63.216667)">Suntar-Khayata Mt. Range</a>, upper current of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.53334&amp;materialsCitation.latitude=63.216667" title="Search Plazi for locations around (long 139.53334/lat 63.216667)">Kyumyume River</a> (63° 13’N 139° 32’E), 1,800 m a.s.l, willow bushes with lichen cover, 9.VII.2002, leg. O. Makarova (MPGU). – Paratypes, 12 females, 11 males on slides and many specimens in alcohol, same data as holotype (MPGU, DBET and MNHG).</p> <p>OTHER MATERIAL: Russia: 3 females, Ural Mts., Perm’ Province, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.666668&amp;materialsCitation.latitude=58.833332" title="Search Plazi for locations around (long 58.666668/lat 58.833332)">State nature reserve “Basegi”</a>, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=58.666668&amp;materialsCitation.latitude=58.833332" title="Search Plazi for locations around (long 58.666668/lat 58.833332)">Srednii Baseg Mt.</a> (58° 50’N 58° 40’E), alpine tundra, 23.VII.1990, leg. S. Esyunin (MPGU). – 2 females, same region but mixed forest with fern cover, 4.IX.1990, leg. S. Esyunin (MPGU). – 13 females and 3 males, Siberia, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.833336&amp;materialsCitation.latitude=68.13333" title="Search Plazi for locations around (long 91.833336/lat 68.13333)">Putorana plateau</a>, vicinity of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.833336&amp;materialsCitation.latitude=68.13333" title="Search Plazi for locations around (long 91.833336/lat 68.13333)">Yt-Kyuel’ lake</a> (68° 08’N 91° 50’E), 700-900 m a.s.l., nival desert, 28.VII-13.VIII.1996, leg. A. Babenko (MPGU). – 7 females and 1 male, Siberia, Taimyr peninsula, upper current of <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=87.416664&amp;materialsCitation.latitude=70.1" title="Search Plazi for locations around (long 87.416664/lat 70.1)">Nizhnaya Agapa River</a> (70° 06N 87° 25’E), tundra, 6.VII-5.VIII.1999, leg. A. Babenko (MPGU). – 2 females and 1 male, Chukotka, vicinity of El’gygytgyn Lake (67° 26’ 172° 10’E), tundra, 20.VIII.1974, leg. E. Bondarenko (MPGU). – 10 specimens, Chukotka, Aborigen, in fungi on dry ridge, 29.VII.1979, leg. A. Fjellberg (AF). – 9 specimens, USA, Alaska, Brook Range, W of Atigun Camp, dry alpine meadow, c. 1600 m a.s.l., 19.VIII.1976, leg. A. Fjellberg (AF). – 7 specimens, USA, Alaska,vicinity of Fairbanks, litter in aspen forest, 21.VII.1980, leg. A. Fjellberg (AF).</p> <p>DESCRIPTION: Body length 1-1.2 mm. Color in alcohol light to dark grey-brown. Eye patches black, anal spines light. Granulation fine and uniform, usually 12-15 granules between setae p 1 on abd. V. Dorsal chaetotaxy of B type, macrosetae p 2 on th. II-III set nearly in line with setae p 1, setae m 3 and m 4 on th. II usually present, setae a 2 slightly longer than a 3, setae m 6 absent, setae p 1 and p 2 on abd. IV macro- and micro-setae respectively, setae p 3 absent (Figs 10-11). Arrangement of setae on head typical for the genus. Differentiation of dorsal setae into micro- and macrosetae not strong and more pronounced on last abdominal terga. Setae short, fine, pointed, slightly curved and serrated. Body sensilla p 4 on th. II-III and p 5 on abd. I usually equal to microsetae, thick and sometimes curved. Body sensilla (s) on abd. II-V and lateral parts of th. II-III about as long as macrosetae (Fig 12). Microsensilla (ms) on th. II present. Subcoxae I-III with 1, 2, 3 setae respectively.</p> <p>Ant. IV with simple apical vesicle, subapical organite (or), microsensillum (ms), 5-7 (usually 7) cylindrical sensilla (2 lateral and 3-5 dorsal) (Fig. 15) and about 15 short curved flattened at tips sensilla in ventral file (Fig. 14). Ant. III-organ with two long (lateral) and two short (internal) curved sensilla (Fig. 15). Microsensillum on ant. III present. Eversible sac between ant. III-IV present. Ant. I with 7 setae.</p> <p>Ocelli 8 + 8. Postantennal organ about twice as large as single ocellus, with four lobes of which the anterior pair larger than the posterior. Accessory boss present (Fig. 13).</p> <p>Labrum with 5, 5, 4 setae and without apical papillae. 4 prelabrals present. Labium of the C. armata type. Maxillary head with prolonged denticulate lobe on lamella 5 which only slightly shorter than lamella 4 (Figs 16-17). Outer lobe with 2 sublobal hairs.</p> <p>Tibiotarsi I, II, III with 19, 19, 18 setae respectively, tibiotarsal tenent hairs nearly as long as inner edge of claws and usually pointed, sometimes truncate. Claws with inner tooth and pair of indistinct lateral teeth. Empodial appendage with broad basal lamella and apical filament reaching inner tooth or slightly beyond (Fig. 18).</p> <p>Ventral tube with 4 + 4 setae.</p> <p>Furca well developed. Dens/mucro ratio = ca. 2. Dens with 7 setae (2 inner modified). Mucro boat-like (Fig. 20). Retinaculum with 4 + 4 teeth.</p> <p>Anal spines equal to or slightly longer than inner edge of claws III, situated on basal papillae (Figs 11, 19).</p> <p>VARIABILITY: Siberian and available Alaskan material appears to be morphologically homogeneous throughout the distributional range. However Fjellberg (1985) mentioned that in Alaska “exact chaetotaxy and differentiation in macro/microchaetae is rather variable” and alpine specimens from Brook Range “frequently have long, hair-like p 5 sensilla on Abd.I”. It may indicate the presence of the third separate form there apart from C. lobata sp. n. and C. brevisensillata s. l.</p> <p>ETYMOLOGY: The name reflects the most characteristic feature of the new species - the presence of an additional lobe on one of the maxillary lamellae.</p> <p>DISTRIBUTION: The new species which seems to be a usual inhabitant of Subarctic Mountains was found in few remote Palaearctic regions from Ural to Chukotka and in Alaska (see map on Fig. 21 and material above).</p> <p>AFFINITIES: A combination of four features, viz. a full number of ocelli, a chaetotaxy of B type, shortened dorsal sensilla on three first terga and maxillae with prolonged lobe on lamella 5, distinguishes C. lobata sp. n. from all other known species of the ceratophysellan lineage. Thus, only C. brevisensillata and Bonetogastrura nivalis (Martynova) are characterized by the same type of sensillar differentiation, but both have the usual armata-type of maxillae and longer dorsal setae which are more clearly differentiated into macro/microsetae. Apart from this, B. nivalis has only 4 + 4 ocelli and partly reduced chaetotaxy (m 3 always and m 4 usually absent on th. II-III, abd. I-III without m-setae as a rule and p 3 often absent on abd. I-IV).</p> <p>Shortened dorsal sensilla on thorax similar to those in C. lobata sp. n. are also characteristic of C. bengtssoni (Ågren), C. microchaeta (Babenko) and B. variabilis (Christiansen) but maxillae of all these species are modified differently with broadened lamella 1 and without prolonged lobe on lamella 5.</p> <p>The same type of maxillae as in C. lobata sp. n. is known only for C. sigillata (Uzel), C. sibirica Martynova and C. pseudarmata (Folsom) 1. They can be easily distinguished from C. lobata sp. n. due to long dorsal sensilla on all terga and clearly clavate tibiotarsal tenent hairs (pointed or truncate in C. lobata sp. n.).</p> </div>	http://treatment.plazi.org/id/03E20D10A851FFDCFF71FC12FCE9FC09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Babenko, Anatoly;Skarżyński, Dariusz	Babenko, Anatoly, Skarżyński, Dariusz (2011): Ceratophysella lobata sp. n. from Siberia with notes on C. brevisensillata Yosii, 1961 (Collembola: Hypogastruridae). Revue suisse de Zoologie 118 (2): 257-264, DOI: 10.5962/bhl.part.117808, URL: http://www.biodiversitylibrary.org/part/117808
03E20D10A854FFDCFF6EFBCFFBB6FAAE.text	03E20D10A854FFDCFF6EFBCFFBB6FAAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratophysella Borner 1932	<div><p>The new species runs in the most recent key to the Palaearctic Ceratophysella species (Thibaud et al., 2004) to couplet 11 which needs to be modified as follows:</p> <p>11 Sensillum p 5 on abd. I small, spine-like; micro- and macrosetae clearly differentiated................................................ 11*</p> <p>– Sensillum p 5 on abd. I long, hair like; micro- and macrosetae weakly.. differentiated........................... microchaeta (Babenko, 1994)</p> <p>11* Maxillary head of the armata type without prolonged lobe on lamella 5........................................ brevisensillata Yosii, 1961</p> <p>– Maxillary head with prolonged denticulate lobe on lamella 5 (Figs 16-17)................................................... lobata sp. n.</p></div> 	http://treatment.plazi.org/id/03E20D10A854FFDCFF6EFBCFFBB6FAAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Babenko, Anatoly;Skarżyński, Dariusz	Babenko, Anatoly, Skarżyński, Dariusz (2011): Ceratophysella lobata sp. n. from Siberia with notes on C. brevisensillata Yosii, 1961 (Collembola: Hypogastruridae). Revue suisse de Zoologie 118 (2): 257-264, DOI: 10.5962/bhl.part.117808, URL: http://www.biodiversitylibrary.org/part/117808
