taxonID	type	description	language	source
03AA87D64219FF9BFCFAFCDCFEA9FBC0.taxon	synonymic_list	Type genus: Zigrasolabis Engel and Grimaldi, 2014.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D64219FF9BFCFAFCDCFEA9FBC0.taxon	diagnosis	Diagnosis. Temples typically with prominent, erect, bristles; flagellomere I longer than combined lengths of flagellomere II and III; tegmina and hind wings present and well-developed, lateral margins without longitudinal ridge (as in Labidurinae, ridge present in Nalinae); mesosternum rectangular, not strongly tapering posteriorly (strongly trapezoidally narrowed posteriorly in Allostethinae); metafemur longer than pronotum; tarsomere II slanted apically (not scapiform), with tarsomere III arising from slanted dorsal surface of tarsomere II, either not lobed nor expanded apically (as in Labidurinae), or with slight, narrowed extension; cercal forceps of female comparatively simple, those of male broadly arched; apices of male parameres bluntly rounded, incurved, not abruptly acute apically (abruptly acute and pointed apically in Labidurinae); paired genital lobes projected caudad, apically blunt, not exceeding parameres. Included genera. The subfamily currently includes Zigrasolabis Engel and Grimaldi and Tricholabidura, gen. nov. The genus Acantholabis Mao et al. perhaps also belongs here (Mao et al., 2020 a).	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D64219FF9BFCFAFCDCFEA9FBC0.taxon	discussion	Remarks. The genus can be placed in Labiduridae, as currently circumscribed, based on the following combination of features: Epidermapteran cervical sclerites (i. e., ‘ forficuloid neck’), thoracic segments of roughly equivalent widths (i. e., prothorax not prominently narrowed relative to pterothorax), tegmina and hind wings typically well developed (rarely secondarily absent), pronotum long and broad with meso- and metanota truncate posteriorly, prosternum subparallel (sometimes slightly constricted posteriorly), generally stout and not strongly compressed, male genitalia paired (paired lobes), with virgae bearing basal vesicle (vesicle not visible in current fossil, as preserved). Labidurid monophyly remains to be determined conclusively and we follow the current circumscription of the family pending future cladistic analyses and revisions of dermapteran classification. Putative apomorphies of Zigrasolabidinae are the rectangular mesosternum, male parameres incurved and bluntly rounded, and the loss of the lateral ridge of the tegmina. In addition, the combination of these features with flagellomere I longer than flagellomeres II and III, the slanted second tarsomere, and the erect bristles of the temple is a further unique combination in the lineage.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D64219FF9BFCFAFCDCFEA9FBC0.taxon	description	ZooBank LSID: This nomenclatural act is registered in ZooBank (www. zoobank. org), the official registry of the International Commission on Zoological Nomenclature, with the following LSID: urn: lsid: zoobank. org: act: 53014 F 06 - F 255 - 4279 - 9 ECE- 7 A 369248 B 6 A 4.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421AFF9AFFACFBCEFF50FE0D.taxon	type_taxon	Type species: Tricholabidura elongata sp. nov.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421AFF9AFFACFBCEFF50FE0D.taxon	etymology	Etymology. The new genus-group name is a combination of the Ancient Greek trikh ́ os (T P ῐ X ό ς, genitive of thríx / Ɵ ρ ῐϵ, meaning, “ hair ”), and Labidura Leach, itself a combination of the genitive of the noun labís (λ β ῐÇ, meaning, “ forceps ” or “ tongs ”; genitive labídos / λ ᾰ β ῐδΟ ς) and the noun our ̅ a (Ο ὐ P ᾱ, meaning, “ tail ”). The gender of the name is feminine.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421AFF9AFFACFBCEFF50FE0D.taxon	diagnosis	Diagnosis. Stout, modestly large earwigs (approximately 13.4 mm in length), somewhat dorsoventrally compressed; integument not chaetulose, with abundant, scattered, fine, short setae on tegmina and abdominal terga, including fine apical margin fringes of posteriorly directed setae (forming vibrissae-like fringes), and more widely scattered on other sclerites; temples of head with numerous long, erect bristles, such bristles less numerous but also present along pronotal margins and sparsely on thoracic sterna; integument matte and seemingly imbricate and impunctate (except setigerous punctures). Head slightly wider than long, not tumid, posterior border relatively concave with fine bristles, frons darker in colour making it distinctive among head regions; compound eyes large, separated from posterior border of head by slightly more than compound eye length; ecdysial cleavage lines faint; antenna with at least 30 antennomeres, total length more than one-half body length; scape sturdy, long, shorter than interocular distance but longer than compound eye length; pedicel small, length and width subequal; flagellomere I distinctly longer than wide, longer than combined lengths of flagellomeres II and III, flagellomere II slightly longer than wide, succeeding flagellomeres gradually more filiform; anterior ventral cervical sclerite narrow and greatly transverse, posterior cervical sclerite large, with apical margin broadly and gently protruded medially, surface with at least a couple prominent bristles, ́ lateral margins converging posteriorly, posteriorly narrower than prosternum. Pronotum subquadrate, slightly narrower than head, longitudinal groove of pronotum with numerous prominent and short setae, anterior margin gently convex with numerous prominent and elongate bristles, anterolateral corners relatively angulate, lateral margins slightly converging posteriorly, posterolateral angle rounded with short setae, posterior margin weakly convex; prosternum twice as long as wide, with procoxae positioned in posterior half, anterior margin gently convex and abutting posterior cervical sclerite, lateral margins straight and converging posteriorly, posterior margin straight, surface with some prominent bristles and shorter finer setae. Tegmina well-developed and long (extending to at least apical margin of abdominal tergum II), approximately rectangular, with blunt apices, surface with numerous fine setae; hind wings present and well-developed, squama as on tegmen. Femora neither compressed nor carinulate, with numerous prominent elongate setae; tibiae with abundant short setae, and several thickened, elongate spine-like bristles; tarsi trimerous with spines or bristle-like setae on surface, many extending beyond apex of each tarsomere; tarsomeres elongate, tarsomere shortest tarsomere but still long, tarsomere II more than 0.5 × length of tarsomere I, slightly extended beneath base of tarsomere III; pretarsal claws long and stout; arolium swollen. Abdomen long, comparatively slender, roughly parallel-sided; terga broader than long, with apical margins straight; pygidium slightly exposed, subtriangular, with short, acute, medioapical projection, apex acutely rounded; dorsal valvulae and gonapophyses long, slightly extended as preserved; antepenultimate sternum broader than long, apical margin straight; penultimate sternum broader than long, apical margin strongly convex medially, broadly and subtriangularly rounded medially; terminal sternum simple, apical margin weakly convex; female forceps symmetrical, broadly separated at bases, smooth and stout, parallelsided in proximal third, then angled and narrowing gradually to apex, apical third subfalcate, apex acutely rounded, without flanges, serration, or dentition.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421AFF9AFFACFBCEFF50FE0D.taxon	discussion	Remarks. The genus differs from Zigrasolabis in the larger body and more slender shape, antenna longer than half the body, more elongate legs and tarsomere II more elongate, abundant tegminal abdominal pubescence, longer tegmina, abundantly and finely setose abdominal terga, marginal vibrissae, exposed pygidium, apical sternum with medioapical margin produced as broadly rounded lobe, cerci somewhat broad in proximal third before tapering gently to acutely rounded apices, apical third subfalcate, and somewhat exposed valvulae.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421AFF9AFFACFBCEFF50FE0D.taxon	description	ZooBank LSID: This nomenclatural act is registered in ZooBank (www. zoobank. org), the official registry of the International Commission on Zoological Nomenclature, with the following LSID: urn: lsid: zoobank. org: act: D 35 DD 0 E 0 - 705 E- 435 A-A 47 A- C 60911528364.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421BFF9CFFACFD8CFEA5FEC0.taxon	etymology	Etymology. The specific epithet is taken from the Latin ̅ elong ̅ atus, meaning, “ elongate ”, and refers to the elongate antennae and legs, particularly the secondtarsomere.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421BFF9CFFACFD8CFEA5FEC0.taxon	materials_examined	Holotype. ♀, YKLP-AMB- 003, deposited in the research collection of the Yunnan Key Laboratory for Palaeobiology, Institute of Palaeontology, Yunnan University, Kunming, China. Locality and horizon. Hukawng Valley, Kachin State, northern Myanmar; upper Albian ̅ lower Cenomanian (ca. 98.79 ± 0.62 Ma). Diagnosis. As for the genus (vide supra).	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
03AA87D6421BFF9CFFACFD8CFEA5FEC0.taxon	description	Description. ♀: Total length as preserved (excluding forceps and antenna) approximately 13.4 mm, with forceps total length approximately 16 mm; integument (as preserved) brown to light brown; with scattered punctures, particularly setigerous punctures, otherwise largely imbricate; head prognathous with length (to apex of clypeus) 1.8 mm, width (across compound eyes) 1.9 mm; surface not tumid except slightly tumid area on frons, otherwise relatively flat; ecdysial cleavage scar faint; compound eyes prominent, length 0.49 ̅ 0.53 mm, width between inner margins of compound eyes 1 mm; temple corner rounded with series of stiff, erect bristles, such bristles extending along posterior margin of head; clypeus with scattered, relatively long setae; maxillary palpus (Fig. 2 C, D) length 1.5 mm, with short setae and several long setae; labial palpus length 0.64 ̅ 0.7 mm; ligula and prementum highly sclerotized, length 0.36 mm; antenna elongate, more than half of total body length, length (as preserved) 9 mm, with scattered, minute setae, with at least 30 antennomeres (Fig. 2 A, B); scape shorter than distance between compound eyes, length 0.62 mm; pedicel length and width about 0.12 mm; flagellomere I length about 2.4 × width, length 0.27 mm, flagellomere II length 0.10 mm, flagellomere III length 0.16 mm; remaining flagellomeres gradually and progressively longer, shape gradually more filiform; pronotum subquadrate, 1.48 mm wide, 1.42 ̅ 1.49 mm long, slightly narrower than head, longitudinal groove of pronotum with numerous prominent and short setae, anterior margin gently convex with numerous prominent and elongate setae, anterolateral corners relatively angulate, lateral margins slightly converging posteriorly with scattered, short, suberect setae, posterolateral angle rounded with short setae, posterior border weakly convex. Tegmen long, rectangular, with numerous fine setae, length, as preserved, 2.83 ̅ 3.12 mm (one tegmen damaged and twisted), width 0.75 - ̅ 1.06 mm (wider medially, lateral margins slightly converging posteriorly in apical quarter); humeral angle rounded, relatively straight; lateral margin slightly convex but generally parallel with inner margin; hind wing present and fully developed, with posteriorly directed setae, squama length 1.05 ̅ 1.15 mm, width 0.57 ̅ 0.70 mm. Legs long, slender; femora swollen, with numerous elongate prominent setae; tibiae with abundant short setae and several thickened, elongate bristles; tarsi trimerous with spines or bristle-like setae (Fig. 3 C ̅ E). Abdomen long, with abundant subappressed, fine setae (Fig. 2 E); surface relatively smooth with scattered, shallow, coarse punctures; length approximately 8.5 mm (excluding forceps); segments transverse, maximum width 2.3 mm; basal segments overlapped by tegmina; lateral margins relatively straight; slightly wider at midlength; pygidium prominent (Fig. 3 B), basal width 0.23 ̅ 0.28 mm, medial length (excluding apical process) 0.42 mm, with prominent, slender, medioapical process, process length 0.12 ̅ 0.14 mm; gonapophyses elongate, slender, length 0.48 ̅ 0.56 mm; forceps nearly oval in cross section, glabrous, length 2.1 ̅ 2.3 mm, basal maximum width 0.51 mm, separation between bases 0.52 mm. ZooBank LSID: This nomenclatural act is registered in ZooBank (www. zoobank. org), the official registry of the International Commission on Zoological Nomenclature, with the following LSID: urn: lsid: zoobank. org: act: A 247327 C-DC 3 C- 4 BED-AE 9 B- BAA 4 ED 16 C 4 AC.	en	Ancheng Peng, Michael S. Engel, Yuhui Zhuang, Ziying Wu, Chen Feng, Yu Liu (2022): A new genus of striped earwigs allied to Zigrasolabis in mid- Cretaceous Kachin amber (Dermaptera: Labidurida). Cretaceous Researc 139: 1-7, DOI: 10.1016/j.cretres.2022.105305
