taxonID	type	description	language	source
D679951B82344C65FF765C5EE637CD0D.taxon	description	(Figs. 1 – 3, 5)	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82344C65FF765C5EE637CD0D.taxon	type_taxon	Type species. Archaeomegalomus burmiticus sp. nov., here designated.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82344C65FF765C5EE637CD0D.taxon	diagnosis	Diagnosis. Archaeomegalomus gen. nov. can be distinguished from the other hemerobiid genera by the combination of the following characters: Antennae, relatively long; in forewing, costal space broad, narrowed distally; humeral veinlets recurrent, pectinately branched, most subcostal veinlets once to twice forked; Sc fused with RA at distal fourth; RP stemmed single origin; MP deeply forked; CuP deeply forked. In hind wing, basal crossvein connecting RP and M (1 r-m) long, slightly sigmoidal; CuP present, branched. Species included. Archaeomegalomus burmiticus sp. nov.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82344C65FF765C5EE637CD0D.taxon	etymology	Etymology. Combination of the Greek ‘ archaios ’ [ἀρχαῖος], meaning ‘ ancient’, and Megalomus, a genus-group name of Hemerobiidae, in reference to the similarities in wing venation. Gender masculine.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82344C65FF765C5EE637CD0D.taxon	discussion	Remarks. We assign this new genus to Hemerobiidae because of the presence of a pair of tibial spurs (absent or very short in Berothidae), slightly swollen tibia at the middle (usually not swollen in Berothidae), well developed recurrent and branched humeral vein in the forewing (usually undeveloped in Berothidae), and the hind wing CuA have long pectinate branches (long with short to very short branches in Berothidae) (see further details in Makarkin & Gröhn 2020).	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	description	(Figs. 1 – 3, 5)	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	diagnosis	Diagnosis. As for the genus (vide supra).	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	description	Description. Body. Length ca. 3.7 mm as preserved (measured from vertex to apex of the abdomen). Head. Eyes large, protruding laterally. Antennae: scapus relatively short, ca. 2 times as long as wide; pedicellus one-half length of scape; both scattered with thin setae; flagellum moniliform, relatively long, composed of 38 flagellomeres, with scattered fine setae on each segment. Maxillary palpi 4 or 5 segmented; labial palpi 3 segmented. Thorax. Pronotum rather short, poorly preserved, covered with thin setae. Mesothorax and metathorax covered with thin setae. Legs. Foreleg slender; procoxa poorly preserved, relatively long; profemur ca. 0.8 mm long, scarce setae; protibia ca. 0.5 mm long, slightly swollen at the middle, covered with dense thin setae, paired tibial spurs; protarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small. Midleg slender; mesocoxa short; mesofemur ca. 0.9 mm long, scarce setae; mesotibia ca. 0.9 mm long, only slightly swollen at the middle to apex, covered with dense thin setae, paired tibial spurs; mesotarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small. Hindleg slender; metacoxa short; metafemur ca. 0.9 mm long, scarce setae; metatibia ca. 1.5 mm long, slightly curved, only slightly swollen at the middle to apex, covered with dense thin setae, paired tibial spurs (Fig. 3 E); metatarsus five segmented, all tarsomeres covered with thin setae, pretarsal claws thin, small. Wings. Forewing hyaline (Fig. 2 A, B), oval with subacute apex, ca. 5.5 mm long, ca. 2.3 mm wide (right forewing); ca. 5.3 mm long, ca. 2.0 mm wide (left forewing). Trichosors present along the entire wing margin, one trichosor between adjacent veins; marginal portions of veins and trichosors with tufts of thin setae; thin setae sparsely present on dorsal and ventral surfaces of longitudinal veins, but absent on crossveins. Costal space broad, narrowed distally; humeral veinlets recurrent, pectinately branched, most subcostal veinlets once to twice forked. Sc fused with RA at distal fourth (Fig. 3 A). Subcostal space, proximal crossvein (1 sc-r) located at origin of RP, intermediate crossvein (2 sc-r) present. RP stemed single origin, with nine branches, most proximal branch deeply forked at the near stem, all branches terminally forked rather shallowly once or twice; five (right forewing) or six (left forewing) third series intra-crossveins and ten (left forewing) or eleven (right forewing) forth series intra-crossveins present. Three ra-rp crossveins (2 ra-rp, 3 ra-rp, and 4 ra-rp) present. M divided into MA and MP, MP deeply forked; two third series intra-crossveins and three fourth series intra-crossveins present. Three r-m crossveins (2 r-m, 3 r-m, and 4 r-m) present. Long radiomedial flexion line clearly discernible. Cu divided into CuA and CuP, CuP close to wing base; CuA pectinately branched, with four branches, most proximal branch deeply forked; CuP deeply forked; all branches terminally forked shallowly once or twice; one second series intra-crossvein and six fourth series intra-crossveins present. Three m-cu crossveins (1 m-cu, 2 m-cu, and 4 m-cu) present. Short mediocubital flexion line clearly discernible. Three anal veins present, poorly preserved; A 1 deeply forked. Two cu-a crossveins (1 cu-a and 4 cu-a) present. Cubitoanal flexion line clearly discernible. Hind wing hyaline (Fig. 2 C, D), elongate-ovoid, ca. 4.6 mm long, ca. 1.8 mm wide (right hind wing); dark brown on pterostigmal area. Trichosors present from the pterostigmal area of costal margin to along the entire wing margin, one trichosor between adjacent veins; marginal portions of veins and trichosors with tufts of thin setae; thin setae sparsely present on dorsal and ventral surfaces of longitudinal veins, but absent on crossveins. Costal space narrowed; humeral lobe triangular, humeral plate bears dense bristles; almost subcostal veinlets simple, five veinlets forked at distal (left hind wing). Subcostal space one crossvein (3 sc-r) present (Fig. 3 B). RP divided into seven branches; most proximal branche (RP 1) loop like vein at the near stem (right hind wing), deeply forked; one second series intra-crossvein and eight third series intra-crossveins present. Two ra-rp crossveins (1 ra-rp and 3 ra-rp) present, 1 ra-rp long and slightly sinuate. M divided into MA and MP, one intra-crossvein (3 ma-mp) present. Three r-m crossveins (1 r-m, 2 r-m, and 3 r-m) present, basal 1 r-m long and sinuate. Cu divided into CuA and CuP; CuP close to wing base: CuA pectinately branched, with six (right hind wing) or five (left hind wing) branches; one third series intra-crossvein present. Two m-cu crossveins (1 m-cu and 3 m-cu) present. Anal vein poorly preserved; A 1 deeply forked. Two cu-a crossveins (1 cu-a and 3 cu-a) present. Abdomen. Visible in right lateral view (Fig. 3 F), poorly preserved, but posterior part rather well preserved. Tergite 6 and 7 developed each posterior margin. Terminal segment unclear. Gonocoxite 9 visible, scattered setae on surface; gonostylus of gonocoxite 9 well developed. Ectoproct poorly preserved.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	materials_examined	Material. HOLOTYPE: a nearly complete female adult (Fig. 1), partly unobservable due to cracks, preserved in approximately 14 mm × 7.5 mm × 4 mm flat drop-shaped, yellow, transparent amber piece; specimen accession number AMNH Bu-SY 31. Type locality and horizon. Hukawng Valley (26 ° 20 ′ N, 96 ° 36 ′ E), Kachin State, northern Myanmar; midCretaceous, upper Albian to lower Cenomanian.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	etymology	Etymology. The specific epithet ‘ burmiticus ’, noun in apposition, refers to the occurrence of the fossil in Burmese amber (burmite) from northern Myanmar.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82364C60FF765F29E5E0CAEF.taxon	discussion	Remarks. The distal parts of Sc and RA in the forewing are separated in nearly all extant Hemerobiidae, except for Adelphohemerobius Oswald, 1994 (see Oswald 1994: fig. 1; Monserrat 1997: fig. 116). In contrast, in the Mesozoic Hemerobiidae, Hemeroberotha sinefurca Makarkin, 2020 is the only species in which the distal part of Sc and RA in the forewing are definitely separated, while the others are fused (Table 1). This trait is probably a plesiomorphic condition in the Hemerobiidae.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82334C6EFF765D1DE3FCCA06.taxon	description	(Fig. 4) Description. The body and wings are not clearly visible due to many cracks. Length ca. 3.6 mm as preserved (measured from vertex to apex of the abdomen). Head poorly preserved; eyes large; antennae composed at least 27 flagellomeres with scattered fine setae on each segment. Thorax covered with dense thin setae; pronotum rather short. All legs slender, covered with dense thin setae; all tibia slightly swollen at the middle with paired tibial spurs. Wings poorly preserved; forewing hyaline, oval, ca. 3.5 mm long; trichosors present along the entire wing margin, one trichosor between adjacent veins, marginal portions of veins and trichosors with tufts of thin setae, humeral veinlets recurrent, costal space rather broad; hind wing hyaline, oval, ca. 3.0 mm long. Abdomen poorly preserved; terminal segment scattered with thin setae; gonostylus of gonocoxite 9 developed.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82334C6EFF765D1DE3FCCA06.taxon	materials_examined	Material. Incomplete female adult (Fig. 4), indiscernible many morphological details due to numerous cracks and overlapping wings and body, preserved in approximately 15 mm × 11 mm × 2.5 mm flat arcwise, dark yellow, somewhat transparent amber piece with mumerous debris and pigmentation; specimen accession number AMNH Bu-SY 32.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
D679951B82334C6EFF765D1DE3FCCA06.taxon	discussion	Remarks. We assign this undetermined specimen to the Hemerobiidae because of the presence of a pair of tibial spurs, slightly swollen tibia at the middle, and recurred humeral veinlets. This undetermined specimen can be distinguished from the three burmite hemerobiids by the number of flagellomeres (this specimen has at least 27 or 28 flagellomeres, whereas there are 38 in Archaeomegalomus, 19 in Hemeroberotha) and the costal space in the forewing (this specimen is rather broad, whereas it is very broad in Cretoneuronema). Thus, we withheld identification of the specimen as we could not confirm the important traits of the wing venation, although we were able to differentiate it from the three previously recorded fossil species found in Burmese amber.	en	Nakamine, Hiroshi, Yamamoto, Shûhei, Takahashi, Yui (2022): Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae). Zootaxa 5178 (4): 380-390, DOI: https://doi.org/10.11646/zootaxa.5178.4.4
