taxonID	type	description	language	source
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	etymology	Etymology. Cystidiata, named after the inflated ectal excipulum cells.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	materials_examined	Holotype: HKAS 78941	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	diagnosis	Pileus loosely saddle-shaped to irregularly lobed, 1 – 3 cm high, 1 – 4 cm broad, margin reflexed and fused with stipe, hymenium slightly wrinkled to rugose, greyish to greyish brown when fresh, becoming brown to blackish when dry, receptacle surface even, grey to greyish brown when fresh, becoming brownish when dry. Stipe 3 – 5 cm long, 0.5 – 1 cm broad, lacunose, with rounded-ribs, glabrous, greyish to greyish brown when fresh, becoming brownish when dry, with white mycelium at the base.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	description	Medullary excipulum 90 – 210 μm broad, of textura intricata, hyaline, composed of 3 – 4 μm broad hyphae, walls thickened, J- in Melzer’s reagent. Ectal excipulum 60 – 110 μm broad, of textura angularis, composed of 20 – 50 × 20 – 30 μm, subglobose to ellipsoid, inflated cells, hyaline, terminal cells 10 – 35 × 8 – 20 μm, blue in cotton blue, J-. Stipitipellis 45 – 70 μm, of textura angularis, hyaline, terminal cells 8 – 44 × 5 – 20 μm, blue in cotton blue, J-. Asci arising from croziers, 8 - spored, subcylindrical to clavate, with apex rounded, 200 – 330 × 13 – 19 μm, J- in Melzer’s reagent, blue in cotton blue. Paraphyses filiform, 3 – 5 μm broad, slightly exceeding the asci, apex enlarged, 4 – 8 μm broad, hyaline, blue in cotton blue, J-. Ascospores [80 / 2 / 2, in H 2 O] 15 – 18.5 × 9 – 12 (– 12.5) μm [Q = (1.33 – 1.83), Qm = (1.62 ± 0.86)], ellipsoid, smooth-walled under the light microscope, uniguttulate, with large central globose to broadly ellipsoidal oil droplet. Asexual morph: Undetermined.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	biology_ecology	Habitat: Scattered or gregarious on the ground, under Pinus yunnanensis forests.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	distribution	Distribution: Currently only known in southwestern China.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	materials_examined	Specimen examined: CHINA, Yunnan Province, Gucheng County, on the ground, under Pinus yunnanensis Franch, alt. 2500 m, 16 Sep. 2012, Qi Zhao 1683 (HKAS 78941, holotype). CHINA, Yunnan Province, Lanping County, on the ground, alt. 2300 m, 16 Aug. 2011, Qi Zhao 1363 (HKAS 74316).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCFFF9FFF6EDD82FC83F299.taxon	discussion	Notes: Helvella cystidiata is characterized by its loosely saddle-shaped pileus with a margin reflexed and fused with stipe, a slightly wrinkled to rugose, greyish to grey-brown hymenium, an even, grey to greyish receptacle surface, a lacunose stipe with rounded-ribs, ectal excipulum comprised of ellipsoid to subglobose inflated cells and all tissues J- in Melzer’s reagent. Morphologically, Helvella cystidiata is similar to H. lacunosa, both sharing a margin flexed and fused with stipe of pileus and lacunose stipe, but the latter differs from the former as pubescent, yellowish brown to brownish when fresh. Ectal excipulum cells catenuliform in long fasciculate tufts (Afzelius 1783). Phylogenetically, Helvella cystidiata is allied with H. pseudolacunosa with strong statistical support (99 % ML, 1.00 BYPP, FIGURE 1) and (97 % ML, 1.00 BYPP, FIGURE 2). However, the latter species has a pinched hymenium, a creamy receptacle surface, and a lacunose stipe with an enlarged base (Ariyawansa et al. 2015).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	materials_examined	Neotype (here designated): SWEDEN, Lapland, Vindeljallets naturreservat, Gieravard, on the ground, under Betula nana, Salix sp., Dryas sp. forests, 20 Aug. 2010, K. Hansen, K. Gillen & I. Olariaga (KH. 10.97).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	diagnosis	Pileus irregularly lobed to slightly saddle-shaped, 2 – 4 cm high, 2 – 5 cm broad, margin reflexed and fused with stipe; hymenium wrinkled to folded towards the center, taupe or nearly black when fresh, blackish when dry, receptacle surface glabrous, grey to pale greyish when fresh, becoming light blackish when dry. Stipe 3 – 10 cm long, 1.5 – 4 cm broad, lacunose, with sharp to rounded ribs, internally chambered, pubescent, taupe, yellowish brown to brownish when fresh, becoming dark brown when dry.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	description	Medullary excipulum 270 – 330 μm broad, of textura intricata, hyaline, composed of 3 – 4 μm broad hyphae, blue in cotton blue, J- in Melzer’s reagent. Ectal excipulum 50 – 110 μm broad, of textura angularis, outermost cells 15 – 37 × 7 – 12 μm, catenuliform in long fasciculate tufts, hyaline, evenly blue in cotton blue, J- in Melzer’s reagent. Stipitipellis 60 – 90 μm, hyaline, composed of cells of textura angularis, terminal cells 13 – 30 × 7 – 14 μm, clavate, with a yellow refractive content in Melzer’s reagent, blue in cotton blue. Asci arising from croziers, 8 - spored, subcylindrical to clavate, with apex rounded, 230 – 275 × 14 – 17 μm, J- in Melzer’s reagent, blue in cotton blue. Paraphyses filiform, 3 – 5 μm broad, slightly exceeding the asci, with a slightly yellow refractive content in Melzer’s reagent, blue in cotton blue, apex slightly enlarged, 5 – 7 μm broad. Ascospores [40 / 2 / 2, in H 2 O] (13) 14 – 16 (17) × 8 – 11 μm [Q = (1.36 – 1.81), Qm = (1.52 ± 0.09)], ellipsoid, smooth-walled under the light microscope, uniguttulate, with large central globose to broadly ellipsoidal oil droplet. Asexual morph: Undetermined.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	biology_ecology	Habitat: Scattered or gregarious on the ground, under Betula sp., Salix sp., Dryas sp. forests.	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	distribution	Distribution: Known in Europe (Dissing, 1966; Häffner, 1987) and China (Liu et al, 1985; Zhuang, 2004).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	materials_examined	Material examined: CHINA, Tibet, Linzhi County, Lulang Town, alt. 3300 m, 28 Jul. 2014, Qi Zhao 2111 (HKAS 87877), same location, 28 Jul. 2014, Qi Zhao 2112 (HKAS 87878).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
A21F8799FFCAFF9BFF6EDFCBFDC3F113.taxon	discussion	Notes: Helvella lacunosa was described by Afzelius (1783), and the type was studied by Abbott & Currah (1997). The distinctive characteristics of our collections, such as irregularly-lobed pileus with a dark brown to blackish hymenium, wrinkled to folded towards the center, a reflexed margin and fusing with stipe in several places, a glabrous, yellowish brown to brownish receptacle surface, and a lacunose, pubescent, yellowish brown, slightly greyish brown to brownish stipe, matches the character variation reported for Helvella lacunosa (Dissing 1966, Häffner 1987). Helvella lacunosa has been described and illustrated repeatedly by mycologists worldwide (e. g., Dissing 1966, Weber 1972, Häffner 1987, Abbott & Currah 1997, Zhuang 2004). The relationship and identity of Helvella lacunosa with other species of Helvella, e. g., H. crispa, H. dryophila, H. fusca, H. juniperi, H. maculata, H. palustris, H. phlebophora, H. pseudolacunosa and H. robusta, H. rugosa, H. sulcata, H. vespertina N. H. Nguyen & Vellinga and H. zhongtiaoensis, have been addressed several times (Dissing 1966, Häffner 1987, Abbott & Currah 1997, Nguyen et al. 2013, Ariyawansa et al. 2015).	en	Xu, Rong-Ju, Li, Lu, Zhao, Qi (2022): Helvella cystidiata sp. nov. (Helvellaceae, Ascomycota) from Tibetan Plateau, China. Phytotaxa 560 (1): 82-92, DOI: 10.11646/phytotaxa.560.1.6, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.6
