taxonID	type	description	language	source
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	description	(Figs 1, 2)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	materials_examined	Material examined. EAST MALAYSIA • 2 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87511, E 117.94206, 41.5 ± 7.7 m. a. s. l.; 17 May 2022, 19 h 22; calling underneath the bark of a large tree; coll. M. K. Tan & T. Robillard; SBH. 22.121, 122 (FRC)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	discussion	Remarks. This species was previously known from Lambir Hill in Sarawak and Kuala Belalong in Brunei Darussalam, both to the west of Borneo. Our specimens (Fig. 1) appear to resemble the descriptions from Lambir Hill by Gorochov (2017) and the figures in Tan & Wahab (2017). It should be noted that the pseudepiphallic lophi [posterolateral epiphallic lobes] appear narrower in lateral view in the Bruneian specimen (Tan & Wahab, 2017: Fig. 2 C). Here, we add a new locality record for this species in the east of Borneo and describe its calling song for the first time.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	biology_ecology	Ecology. This species seems to be relatively common. Multiples males were heard singing at night underneath barks of large trees.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	distribution	Distribution. Borneo, Sarawak State (Lambir Hill) and Sabah State (Sepilok), Brunei Darussalam (Kuala Belalong).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8DEDABFB0DFDBEFED2.taxon	materials_examined	Type locality. EAST MALAYSIA, Sarawak State, Lambir Hill Calling song (2 ♂, in the field, 26.5 ° C) (Fig. 2). The calling song consists of an echeme made up of 40 – 47 syllables. The first few syllables are of lower amplitude but the amplitude of subsequent syllables increases to a maximum and remains consistent. Each echeme has an average duration of 0.66 ± 0.03 s (0.61 – 0.71 s). The interval between consecutive echemes is 0.87 ± 0.14 s (0.70 – 1.15 s). Each syllable has an average duration of 11.6 ± 0.7 ms (10.9 – 12.9 ms) and the interval between consecutive syllables is 3.5 ± 0.8 ms (2.1 – 4.3 ms). The dominant frequency is 5.89 ± 0.06 kHz (5.72 – 5.91 kHz).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB6FF8CEDABFC60FCDBFBBD.taxon	type_taxon	Type species. Gryllus brachypterus Haan, by original designation	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	description	(Figs 3 – 6)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	materials_examined	Material examined. Holotype: EAST MALAYSIA • 1 ♂; Sabah State, Sabah State, Mount Silam, near Lahad Datu; N 4.97667, E 118.19135, 428.3 ± 7.9 m. a. s. l.; 13 May 2022, 21 h 36; on the ground along the trail; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.62 (FRC) Paratypes: • 1 ♂; Sabah State, Tabin Wildlife Reserve; N 5.19438, E 118.50294, 93.7 ± 11.0 m. a. s. l.; 15 May 2022, 19 h 35; calling from a burrow on the clayey bank of a forest stream; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.84 (ZRC) • 1 ♂, 1 ♀; Sabah State, Mount Silam, near Lahad Datu, lowland forest; N 4.97589, E 118.19060, 341 m. a. s. l.; 13 May 2022, 21 h 10, video of male calling song (TR 00022); male at the base of a large tree under the bark; female on the ground, near the entrance of burrow; coll. T. Robillard, M. K. Tan & R. Japir; TR 22 - 12; (MNHNEO-ENSIF 11147, MNHN-EO-ENSIF 11148) Other material examined: EAST MALAYSIA • 1 ♀ juvenile; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87395, E 117.93871 (SAB 14), 54 m. a. s. l.; 19 May 2022; coll. T. Robillard & M. K. Tan; TR 22 - 39 (MNHN)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	diagnosis	Diagnosis. This new species differs from other species of the nominal subgenus Duolandrevus by the shape of the pseudepiphallus [epiphallus], metanotal gland and FW venation. Its male genitalia are most similar to Duolandrevus (Duolandrevus) kalimantan Gorochov, 2016 from Kalimantan; but the new species differs by FW with a larger harp area and more strongly curved oblique veins (instead of mostly straight), the metanotal gland more transverse; dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe], when viewed laterally, with the apex rounded (instead of subacute). It is also similar to Duolandrevus (Duolandrevus) brachypterus (Haan, 1844) from Java; but differs by the FW without a mirror area, more strongly curved oblique veins and the shapes of pseudepiphallic posteromedial lophi and pseudepiphallic lophi [posteromedial and posterolateral epiphallic lobes, respectively]. From other Bornean species of the subgenus: this new species differs from Duolandrevus (Duolandrevus) kubah Gorochov, 2016 and Duolandrevus (Duolandrevus) sabah Gorochov, 2016 by the absence of mirror on the FW and the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of slender or humped, respectively); from Duolandrevus (Duolandrevus) balikpapan Gorochov, 2016 and Duolandrevus (Duolandrevus) spinicauda Gorochov, 2016 by the pseudepiphallic lophi [posterolateral epiphallic lobe] not elongated and slender; from Duolandrevus (Duolandrevus) matang Gorochov, 2017 by the dorsal projection of the pseudepiphallic lophi [posterolateral epiphallic lobe] broadly rounded (instead of hooked).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	discussion	Remarks. This new species is placed under the nominal subgenus Duolandrevus according to these characters: the hindwings well surpassing metanotal gland; the male anal plate without a bundle of strong setae directed dorsad; the male genitalia with dorsal denticle at or near base of each pseudepiphallic lophi [posterolateral epiphallic lobe].	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	etymology	Etymology. The species name refers to the rarity of finding landrevine males forming burrows on the clayey bank of a forest stream. Often, landrevines call among cavity and crevices of tree trunk or branches.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	description	Description. Typical of the genus, body dorso-ventrally compressed and not pubescent, dark red-brown in colouration. Head rostrum not pubescent, about 1.7 times as wide as scape, with apex broadly rounded (in dorsal view) (Fig. 4 A). Scapes brown (Fig. 4 A). Eyes medium-sized, rather rounded (in profile view). Maxillary palpi yellow brown; with apical (fifth) segment yellow, longer than third and subapical (fourth) segments, slightly enlarged apically, with apex obtusely rounded; subapical segment, widens slightly apically, slightly longer than third segment (Fig. 4 B). Head in anterior view slightly wider than tall (Fig. 4 C). Lateral ocelli posterior of scapes oval; median ocellus between scapes small circular (Fig. 4 C). Pronotum transverse, 1.7 times as broad as long; anterior margin as wide as posterior margin; dorsal disc with anterior margin lined with a row of setae and concave, and posterior margin feebly substraight (Fig. 4 A). Pronotal lateral lobe about 1.3 times as long as wide; anterior half about 1.6 times as tall as posterior half, with ventral margin straight (Fig. 4 B). TI with both tympana open and having oval tympanal membrane, inner one larger than outer one. TIII inner and outer margins with 4 – 5 stout articulated spurs (also known as movable spines) on each dorsal side; and 3 inner and ca. 9 outer basal spines, much smaller. TaIII with about 4 inner and 5 outer denticles. Male. FW glossy red-brown, harp area transparent (Figs 4 D, 4 E). Dorsal field not distinctly longer than lateral field, apical field stout (Fig. 4 E). Venation (Figs. 4 D, 4 E): no distinct mirror present; harp area very large with 6 oblique veins (3 strongly curved long posterior veins and 3 substraight small anterior veins). Lateral field wide, R and M closely-spaced. Sc parallel to R and M, with four longitudinal and parallel branches; third vein (from dorsal) bifurcate in middle. Hind wings well surpassing metanotal gland (Fig. 4 F). Metanotal gland with anterior margin with a row of strong setae, anterior lateral ends slightly swollen; two shallowly sunken divided from each other by a narrow longitudinal ridge; posterior margin broadly and faintly concave (Fig. 4 F). Anal plate triangular with truncated apex, with setae along posterior margin but without bundle of strong setae directed dorsad (Fig. 5 A). Male pseudepiphallus [epiphallus] (Figs 5 B – D) notched in the middle, anterior half not raised. Pseudepiphallic posteromedial lophi [posteromedial epiphallic lobules] triangular, not elongated with obtuse apex; these lobules separated from each other by very shallow and narrow notch, appears fused together with a groove. Pseudepiphallic lophi [posterolateral epiphallic lobes] forming narrow plate-like projection, pointing dorsad at the apex (when viewed laterally); with a shorter and smaller projection pointing posteriorly (when viewed laterally); ventrally folded internally (when viewed dorsally or ventrally), also with a small projection pointing interno-posteriorly. Dorsal and posterior projections with apices bearing a few setae; apex of dorsal projection more broadly rounded, apices of posterior projections narrowly obtuse. Ectophallic fold [rachis] when viewed ventrally faintly surpassing pseudepiphallic posteromedial lophi, but not reaching the posterior end of pseudepiphallic lophi; very slender with acute apex. Each ectophallic apodeme [endparamere] forked posteriorly, and forming a Y-shape structure. Endophallic sclerite [formula] when viewed ventrally fairly stout, narrow anteriorly, widened and truncated at the posterior end. Rami not fused together by the anterior ends. Measurements (2 ♂, in mm). PronL = 2.7 – 2.9 (mean = 2.8); PronW = 4.6 – 5.3 (5.0); FWL = 7.1; FIIIL = 12.7; TIIIL = 8.9.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	biology_ecology	Ecology. One of the males was found calling from a burrow on the clayey bank of a forest stream (Fig. 3). Another male was found on the ground along the trail. These suggest that this species is a more ground-dwelling landrevine.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	distribution	Distribution. Borneo, Sabah State: Mount Silam near Lahad Datu, Tabin Wildlife Reserve.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB1FF87EDABFF59FD37F9E2.taxon	materials_examined	Type locality. EAST MALAYSIA, Sabah State, Mount Silam near Lahad Datu. Calling song (1 ♂, in the field, 23.3 ° C) (Fig. 6). The calling song consists of an echeme made up of 32 – 48 syllables. The first few syllables are of lower amplitude but the amplitude of subsequent syllables increases to a maximum and remains relatively consistent. Each echeme has an average duration of 0.60 ± 0.07 s (0.46 – 0.70 s). The interval between consecutive echemes is 5.01 ± 1.84 s (3.19 – 8.47 s). Each syllable has an average duration of 9.1 ± 0.5 ms (8.5 – 10.0 ms) and the interval between consecutive syllables is 5.6 ± 0.6 ms (4.4 – 6.1 ms). The dominant frequency is 3.90 ± 0.03 kHz (3.86 – 3.95 kHz).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBDFF87EDABF9B0FBE1F81D.taxon	type_taxon	Type species. Odontogryllodes brevicauda Chopard, 1969, by original designation.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBDFF87EDABF9B0FBE1F81D.taxon	discussion	Remarks. This genus previously consisted of six species from Malay Peninsula, Sumatra and Java prior to the new species described from Borneo. These crickets are characterised by a cylindrical body; males with short FWs and completely lack stridulatory apparatus; metanotal gland present; tympanum absent; females with short ovipositor (Gorochov, 2016). Tan & Kamaruddin (2016) presented a key to species.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	description	(Figs 7 – 11)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	materials_examined	Material examined. Holotype: EAST MALAYSIA • ♂; Sabah State, Mount Silam, near Lahad Datu; N 4.96786, E 118.17203, 722.3 ± 6.7 m. a. s. l.; 12 May 2022, 20 h 22; on a branch of a small tree; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.33 (FRC) Paratypes: EAST MALAYSIA • 1 ♂; Sabah State, Mount Silam, near Lahad Datu; N 4.96609, E 118.17285, 717.8 ± 8.6 m. a. s. l.; 12 May 2022, 20 h 05; on a foliage of a small tree; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.29 (ZRC) • 1 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87342, E 117.94214, 64.4 ± 7.1 m. a. s. l.; 17 May 2022, 21 h 36; on a branch of a small tree; coll. M. K. Tan & T. Robillard; SBH. 22.135 (ZRC) • 1 ♀; Sabah State, Sandakan, Sepilok, Kabili Sepilok Forest Reserve; N 5.86984, E 117.93806, 68.3 ± 6.5 m. a. s. l.; 3 October 2019, 20 h 19; on a branch; coll. M. K. Tan & J. Lee Yukang; SDK. 19.100 (MNHN) Other material examined: EAST MALAYSIA • 1 ♀; Sabah State, Tabin Wildlife Reserve; N 5.19541, E 118.50370, 117.7 ± 6.7 m. a. s. l.; 15 May 2022, 21 h 32; on a branch near the ground; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.92 (FRC) [need a male from the same locality to confirm the species]	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	diagnosis	Diagnosis. The new species differs from all of the congeners by the anterior half of the male pseudepiphallus [epiphallus] curved upwards and with lateral squarish lobes (these two lateral lobes have spine-like setae); and by the pseudepiphallic lophi [posterolateral epiphallic lobe] forming narrow-triangular dorsal lobules pointing dorsad with subacute apices (in lateral view), and ventral lobules stout and obtuse (all lobules with setae); the male FW with the apical area of the dorsal field narrowly-obtuse.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	discussion	Remarks. This represents the first record of Odontogryllodes in Borneo (previously known from Malay Peninsula, Sumatra and Java).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	etymology	Etymology. The species name refers to the numerous prickle-like setae on the pseudepiphallus; in Latin from spîna (“ thorn, spine, prickle ”) + - fer (“ carrying ”).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	description	Description. Typical of the genus, body slightly cylindrical and densely pubescent, red-brown in colouration (Fig. 7). Head rostrum very densely pubescent, about 1.2 times as wide as scapes, with apex truncated (in dorsal view) (Fig. 8 A). Scapes yellow brown. Eyes fairly small, vertically elongated (in profile view) (Fig. 8 B). Maxillary palpi with apical (fifth) segment yellow, swollen and pyriform, longer than third and subapical (fourth) segments, with apex obtusely rounded; subapical segment pale-brown, widens apically, slightly longer than third segment; third segment pale-brown, cylindrical (Fig. 8 B). Head in anterior view 1.1 times as tall than wide (Fig. 8 C). Face dark brown, clypeus whitish. Median ocellus small and circular, between scapes; lateral ocelli small and circular, posterior of scapes. Pronotum 1.1 times as long as broad; very densely setose but anterior and posterior margins with strong hairs (Fig. 8 A). Dorsal disc slightly widening with posterior margin 1.2 times as wide as anterior margin; with anterior margin feebly concave in the middle and posterior margin feebly convex in the middle (Fig. 8 A). Pronotal lateral lobe about 1.3 – 1.4 times as long as wide; anterior half about 2.2 times as tall as posterior half; with ventral margin straight (Fig. 8 B). TI without tympanum. TIII inner and outer margins with 4 stout articulated apical spurs (also known as movable spines) on each dorsal side; with 7 (3 small and 4 longer) inner and 6 (2 small and 4 longer) outer subapical spines. TaIII with two rows of denticles, 4 – 5 on each side. Male. FW red-brown, densely pubescent (not as dense as pronotum) between veins; barely reaching posterior end of metanotum; spaced well apart and not touching each other (Figs 8 D, 8 E). Dorsal field distinctly longer than lateral field; venations obsolete, except R and M straight, widely spaced apart and parallel; apical area narrowlyobtuse (Fig. 8 D). Lateral field wide, with four longitudinal veins; dorsal two longitudinal veins most spaced-apart, dorsal most longitudinal vein straight to faintly sinuous, second and third veins (from dorsal) parallel and gently curved in the middle (Fig. 8 E). Hind wings absent. Metanotal gland seemingly under-developed. Anal plate triangular with truncated apex, with sparse setae along posterior margin (Fig. 9 A). Male pseudepiphallus [epiphallus] (Figs 9 B – D) in dorsal view notched in the middle; anterior half curved upwards and with two lateral squarish lobes; lateral lobe with spine-like setae. Pseudepiphallic posteromedial lophi [posteromedial epiphallic lobe] stout with obtuse apex; these lobules separated from each other by shallow and narrow notch. Pseudepiphallic lophi [posterolateral epiphallic lobe] forming dorsal lobules in lateral view narrow-triangular and pointing dorsad with subacute apices, with spine-like setae on the dorsal and ventral (fewer) margins; ventral lobules stout and obtuse, also with a few setae. Ectophallic fold [rachis] barely reaching apex of pseudepiphallic posteromedial lophi [posteromedial epiphallic lobe] (when viewed ventrally), slender and faintly raised and tapering to subacute apex (when viewed laterally); vertically lamellar plates of the ectophallic fold [rachis] having microscopic ridges on their lateral sides. Each ectophallic apodeme [endparamere] forked posteriorly, and together with its long anterior apodeme forming a Y-shape structure; inner branch shorter and with obtuse apex, outer branch longer (reaching bottom of pseudepiphallic notch) and with truncated apex. Endophallic sclerite [formula] elongated; faintly tapering posteriorly into obtuse to truncated apex (when viewed ventrally). Rami not fused together by the anterior ends. Female. Habitus not different from males (Figs 10 A, 10 B). FW characteristic of genus, small and rounded, with 7 – 8 longitudinal veins in both dorsal and lateral fields; FWs not touching each other. Subgenital plate typical of genus, broader than long, with posterior margin concave (Fig. 10 C). Ovipositor slightly curved, short, barely reaching middle of cerci; having characteristic denticulate lateral edges of the distal part of upper valves (Fig. 10 D). Measurements (3 ♂, 1 ♀, in mm). PronL = ♂ 2.8 – 2.9 (2.9), ♀ 2.9; PronW = ♂ 3.2 – 3.3 (3.3), ♀ 3.3; FWL = ♂ 2.2 – 2.4 (2.3), ♀ 1.4; FIIIL = ♂ 7.6 – 8.3 (8.0), ♀ 8.4; TIIIL = ♂ 4.6, ♀ 4.9; OL = 4.6.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	biology_ecology	Ecology. This species is found among branches and tree trunks, usually around and above breast height (rather than near the ground).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	distribution	Distribution. Borneo, Sabah State: Mount Silam near Lahad Datu, Kabili Sepilok Forest Reserve (including Rainforest Discovery Centre).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFBEFF83EDABFF59FD19FA5B.taxon	materials_examined	Type locality. EAST MALAYSIA, Sabah State, Mount Silam near Lahad Datu. Calling song. The species is probably mute.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB9FF83EDABF9BBFB6AF83D.taxon	type_taxon	Type species. Brevimunda variegata Gorochov, 2007, by original designation	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFB9FF83EDABF9BBFB6AF83D.taxon	discussion	Remarks. This small genus consists of two species, both from Mount Kinabalu in Borneo: B. kinabalu Gorochov, 2007 and B. variegata Gorochov, 2007. The genus is characterised by the TI with only a small inner tympanum; the apical segment of the maxillary palpi particularly widened; the lateral field of the FW with parallel longitudinal veins; the hindwings shorter than FWs; and the male anal plate short and with a pair of characteristic small transverse lobes at the upper (proximal) half. Below, a species from the eastern part of Sabah is described, first for the genus, but it also exhibits some deviations from the above-mentioned generic characters.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	description	(Figs 12 – 14)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	materials_examined	Material examined. Holotype: EAST MALAYSIA • ♂; Sabah State, Sepagaya Waterfall, near Lahad Datu; N 4.98364, E 118.14537, 172.1 ± 6.4 m. a. s. l.; 13 May 2022, 12 h 10; on a foliage of an understory plant; coll. M. K. Tan & T. Robillard; SBH. 22.57 (FRC) Paratype: 1 ♂; Sabah State, Sandakan, Sepilok, Kabili Sepilok Forest Reserve; N 5.86948, E 117.93913, 36.4 ± 8.8 m. a. s. l.; 1 October 2019, 19 h 49; on a foliage; coll. M. K. Tan, R. Japir & J. Lee Yukang; SDK. 19.72 (MNHN)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	diagnosis	Diagnosis. The new species differs from currently known Brevimunda by the apical segment of the maxillary palpi less strongly widened; the hind wings surpassing the FWs; by the pseudepiphallic lophi [apical epiphallic lobe] more elongated, slenderer and pointing dorsad (vs. broad in B. variegata) and nearly perpendicular to the pseudepiphallus [epiphallus] (vs. pointing posterio-dorsad in B. kinabalu); the pseudepiphallic paramere [ectoparamere] in ventral view more elongated and slenderer. The new species also differs from B. variegata by the cream-coloured / yellow spot on the posterior margin of pronotal lateral lobe (instead of anterior-ventral angle) and from B. kinabalu by a darker overall colouration. The pseudepiphallic lophi [apical epiphallic lobe] of the new species is also slender and pointing dorsad, somewhat resemble that of Mistshenkoana pileata Gorochov, 2008 from the Solomon Islands, but the new species differs from the latter by its distinct darker colouration. Generic remarks. This new species certainly bears similarities with currently known Brevimunda such as the apical segment of the maxillary palpi strongly widened (but not so strongly widened as described by Gorochov [2007]); the lateral field of the FW with longitudinal veins parallel; and the male anal plate short and with a pair of characteristic small transverse lobes at upper (proximal) half. However, there also some key differences: the apical segment of the maxillary palpi (described above); the TI with both tympana open (instead of a small inner tympanum and no outer tympanum); the hind wings surpassing the FWs.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	etymology	Etymology. The species name refers to the characteristic the three elongated white / cream-coloured spots along M on the FW: at the basal third, in the middle (largest) and near the apical third.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	description	Description. Habitus typical of the genus, medium sized, head, pronotum and FW mostly dark-coloured (Fig. 12). Head with dorsum flattened, pubescent. Rostrum dark-coloured with two lateral white stripes, about 0.7 times as wide as scapes, with apex roundly truncated (in dorsal view) (Fig. 13 A). Scapes dark-coloured. Eyes distinctly projected anteriorly in dorsal view (Fig. 13 A). Median ocellus round and small; lateral ocelli oval, located near eyes. Maxillary palpi cream-coloured; with apical segment longest, dorso-posteriorly dark-coloured, flattened and widened, with obliquely-truncated apex; with subapical segment very short, cylindrical and expanding apically, dark-coloured at both ends; third segment longer than subapical segment, cylindrical (Fig. 13 B). Face in anterior view 1.1 times taller than wide; dorsum half black, ventral half (including clypeus) cream-coloured (Fig. 13 C). Gena cream-coloured with tint of dark (Fig. 13 B). Pronotal disc dark brown, pubescent; about as wide as long, barely widening posteriorly (posterior margin 1.1 times as wide as anterior margin); finely pubescent, with a row of strong setae along posterior and anterior margins; anterior margin of disc broadly concave; posterior margin of disc angularly convex (Fig. 13 A). Pronotal lateral lobe 1.4 times as long as high, dark-coloured, with a cream-coloured spot in the middle along the posterior margin (Fig. 13 B). Metanotal gland absent (Fig. 13 D). TI slightly swollen; with inner tympanum open and having elongate, oblong tympanal membrane; with outer tympanum also open and having smaller oval tympanal membrane; TIII with 5 inner and 5 outer long spines in distal half, with small denticles before most proximal spines. Legs generally cream-coloured with irregular and sparse dark spots; Fs with more dark spots in the apical half; Ts generally with more dark spots. FIII with a black ring at apical third, knees dark-coloured. TIII with dark rings. TaIII with basal half cream-coloured and posterior half dark-coloured. Thoracic and abdominal segments dark coloured. Male. FW (Figs 13 E, 13 F) covering abdomen and surpassing apex of FIII. Colouration: yellow brown, with darker infumated spots; at the base with a diagonal white stripe across FW; along M with three elongated white / cream-coloured spots at the basal third, in the middle (largest) and near apical third. Lateral field with intercalary triangle transparent, otherwise dark-coloured with dorsal half of cells between branches of Sc cream-coloured. Venation: without stridulatory organs, with longitudinal veins parallel; lateral field with R and M narrowly spaced apart, parallel and with transverse veins (especially in the basal half); branches of Sc parallel. Hind wings clearly surpassing FWs. Anal plate at anterior half with a pair of characteristic small transverse lobes at upper (proximal) half yellow in colouration, posterior half with two rounded indentation, cream-coloured with dark margins; cerci cream-coloured; posterior margin emarginated in the middle (Fig. 14 A). Subgenital plate typical of subgenus, cream-coloured with small dark spots. Male genitalia as shown in (Figs 14 B – D): pseudepiphallus [epiphallus] elongated, tapering slightly posteriorly, divided into two parts at posterior end and roundly and narrowly excised; posterior end produced into pseudepiphallic lophi [apical epiphallic lobe]. Pseudepiphallic lophi [apical epiphallic lobe] in dorsal view long and slender and pointing posterior-externally with apex obtuse; in lateral view with also slender, pointing dorsad (nearly perpendicular to pseudepiphallus [epiphallus]), apex rounded and with one or a few strong setae. Pseudepiphallic paramere [ectoparamere] in ventral view elongated and slender (more so than congener), slightly curved, anterior end widened lamellate fan-like. Ectophallic fold [rachis] small, slender and straight, taper posteriorly with apex obtuse. Female. Unknown. Measurements (2 ♂, in mm). PronL = 2.1 – 2.2 (2.2); PronW = 2.2; FWL = 10.3 – 11.2 (10.8); HWT = 3.3; FIIIL = 9.3; TIIIL = 9.5.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	biology_ecology	Ecology. Unknown.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	distribution	Distribution. Borneo, Sabah State: Sepagaya Waterfall, near Lahad Datu, Kabili Sepilok Forest Reserve	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA4FF9DEDABFF59FDF5F93A.taxon	materials_examined	Type locality. EAST MALAYSIA, Sabah State, Sepagaya Waterfall, near Lahad Datu Calling song. This species is mute.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF9AEDABFF59FDE4FDDB.taxon	type_taxon	Type species. Varitrella palawanensis Gorochov, by original designation	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF9AEDABFF59FDE4FDDB.taxon	discussion	Remarks. This subgenus consists of 19 species from Indochina, Malay Peninsula, Java, Borneo, Philippines and Sulawesi. It can be distinguished from the nominotypical subgenus by the absence of metanotal gland in males, the apical part of the ectophallic fold [rachis] in the male genitalia widened and with comparatively smaller lobes (including a virga-like or finger-like median process), and the ectoparameres (if they are developed) probably originated from the lateral lobes of the pseudepiphallus [epiphallus] but not from the lateral lobes of the ectophallic fold [rachis] (Gorochov & Tan, 2014).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF98EDABFDABFDE0FDAE.taxon	description	(Figs 15, 16)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF98EDABFDABFDE0FDAE.taxon	materials_examined	Material examined. Holotype: EAST MALAYSIA • ♂; Sabah State, Sandakan, Sepilok, Kabili Sepilok Forest Reserve; N 5.86985, E 117.93805, 69.8 ± 7.7 m. a. s. l.; 1 October 2019; on a foliage; coll. M. K. Tan, R. Japir & J. Lee Yukang; SDK. 19.78 (FRC) Other material examined: EAST MALAYSIA • 1 ♂; Sabah State, Sandakan, Sepilok, Rainforest Discovery Centre; N 5.87435, E 117.94205, 49.9 ± 5.7 m. a. s. l.; 17 May 2022, 20 h 17; among the branch and foliage, feeding on the fruits of Leea; coll. M. K. Tan & T. Robillard; SBH. 22.132 (MNHN)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF98EDABFDABFDE0FDAE.taxon	discussion	Remarks. This represents the second specimen for this species, having been previously described by a single holotype. The male resembles the holotype but we also illustrated the spermatophore (Fig. 15 C).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF98EDABFDABFDE0FDAE.taxon	biology_ecology	Ecology. The recently collected specimen was observed to feed on the fruits of Leea, but it probably also fed on the flowers and leaves of this common plant at the Rainforest Discovery Centre (Fig. 16).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA0FF98EDABFDABFDE0FDAE.taxon	distribution	Distribution. Known only from the type locality, Sepilok (Borneo, Sabah State). Calling song. The calling song of the holotype was described in Tan et al. (2020). The call structure and call parameters of Varitrella (Cantotrella) suikei are distinct from those of Varitrella (Cantotrella) tabin sp. nov.: the former consists of irregular trill instead of well-defined echeme-sequences (even though each echeme is made up of two syllables). The echeme and syllable durations of Varitrella (Cantotrella) suikei are shorter in duration; and the dominant frequency is slightly higher.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	description	(Figs 17 – 19)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	materials_examined	Material examined. Holotype: EAST MALAYSIA • ♂; Sabah State, Tabin Wildlife Reserve; N 5.19447, E 118.50259, 80.1 ± 7.8 m. a. s. l.; 15 May 2022, 22 h 57; on a foliage of a tree along the forest edge; coll. M. K. Tan, T. Robillard & R. Japir; SBH. 22.101 (FRC)	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	diagnosis	Diagnosis. The new species is most similar to Varitrella (Cantotrella) sukau Gorochov, 2014 from Sukau in Kinabatangan by the shape of male genitalia, but differs by the shape of ectoparamere. It also differs from V. sukau by the shape and pattern of anal plate. From species found nearby, this new species also differs from Varitrella (Cantotrella) tawau Gorochov, 2014 from Tawau (south of Tabin Wildlife Reserve) by the oval patch on anal plate nearly touching other and fusing (instead of widely spaced apart), the dorso-anterior pseudepiphallic spine (ds) larger and the pseudepiphallic paramere [ectoparamere] not short and membranous; and from Varitrella (Cantotrella) suikei from Sepilok (north of Tabin Wildlife Reserve) by the dorso-anterior pseudepiphallic spine (ds) and the latero-apical lobular parts larger and more robust as well as the calling song (see above). The pronotum and FW colouration are somewhat similar to Varitrella (Cantotrella) orion Tan & Gorochov, 2014.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	etymology	Etymology. The species is named after its type locality, Tabin Wildlife Reserve; noun in apposition.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	description	Description. Habitus very typical of the genus, medium sized, generally grey-brown. Head with dorsum slightly flattened, very finely pubescent (Fig. 17 A). Rostrum dark-coloured, about 0.8 times as wide as scapes, with apex broadly rounded (in dorsal view) (Fig. 17 B). Eyes projected anteriorly in dorsal view (Fig. 17 B). Median ocellus round and small; lateral ocelli oval, located near eyes. Maxillary palpi with apical segment longest, distinctly flattened and oblong with subobtuse apex; with subapical segment cylindrical and slightly expanding apically; third segment very short and stout (Fig. 17 A). Gena greyish with tint of black patches (Fig. 17 A). Face in anterior view 1.1 times as tall as wide; with a black patch between antennal grooves (Fig. 17 C). Pronotal disc brown with numerous dark spots (especially distinct along anterior margin); about 1.3 times as wide as long, faintly widening posteriorly (posterior margin 1.2 times as wide as anterior margin); finely and densely pubescent; anterior margin of disc broadly concave; posterior margin of disc nearly straight (Fig. 17 B). Pronotal lateral lobe 1.4 times as long as high; with a longitudinal dark band in dorsal half, numerous black spots of different sizes in ventral half; with ventral margin faintly convex (Fig. 17 A). Metanotal gland absent (Fig. 17 D). TI slightly swollen; with inner tympanum slit-like; with outer tympanum open and having oval tympanal membrane. TIII with 6 inner and 5 outer long spines; with 2 – 4 denticles between spines; 4 denticles before most proximal spines; and with inner spurs (apical spines) longer than corresponding outer ones. Legs generally pale brown with irregular and sparse dark spots. Thoracic and abdominal segments yellow brown to brown. Male. FW (Fig. 17 E) covering abdomen and barely surpassing apex of FIII. Colouration: yellow brown, with darker infumation. Dorsal field of FW yellow brown with basal area, proximal part of region of chords, and apical area almost dark brown, and with large white spot between disto-lateral edge of mirror and stock of MP + CuA 1. Lateral field of FW paler, light brown with blackish marks near Sc between branches of Sc. Venation: diagonal straight, with 5 oblique veins in harp area; mirror 1.7 times as long as wide, dividing vein typical of subgenus; lateral field around 16 branches on Sc (Fig. 17 E). Hind wings clearly surpassing FWs. Anal plate with anterior half broad and black; at base with two whitish oval parts fusing in the anterior half and appearing as cranium-shaped; posterior half truncated, faintly emarginated in the middle (Fig. 17 F). Subgenital plate typical of subgenus. Male genitalia as shown in (Fig. 18): Pseudepiphallus [epiphallus] stout, in profile view tall. Pseudepiphallus [epiphallus] with dorso-anterior pseudepiphallic spine (ds) strongly sclerotized with acute apex; elongated and slender in dorsal view; elongated pointing dorsad in profile view, with anterior margin straight and posterior margin slightly curved. Posterior end of pseudepiphallus [epiphallus] produced into two latero-apical lobular parts (al), in dorsal view broad, with apex obtuse; deeply V-emarginated between latero-apical lobules in dorsal view; in profile with posterior part rounded with a dorso-apical spinule and a row of widely-spaced small spinules on dorsal margin, dorsal margin convex in profile view. Pseudepiphallic paramere [ectoparamere] elongated, tapered abruptly into a slender and curved hooked-like apical third with apex acute, and pointing ventrad posteriorly. Ectophallic fold [rachis] rather long, slightly surpassing pseudepiphallus [epiphallus], produced into two broadly-obtuse lateral lobes (lo) at posterior end, with apical emargination wider between lateral lobes in dorsal view. Endophallic sclerite [formula] elongated, typical of genus. Spermatophore oval. Female. Unknown. Measurements (♂, in mm). PronL = 3.0; PronW = 3.3; FWL = 18.1; FIIIL = 13.2; TIIIL = 12.9.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	biology_ecology	Ecology. Unknown.	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
7A5A878EFFA2FF94EDABFD69FE2CFBDE.taxon	distribution	Distribution. Known only from type locality, Tabin Wildlife Reserve (Borneo, Sabah State). Calling song (1 male, in captivity, 32.0 ° C) (Fig. 19). The calling song consists of a well-defined echemesequence made up of 6 or 7 echemes. The echeme-sequence has an average duration of 1.23 ± 0.10 s (1.10 – 1.39 s). The interval between consecutive echeme-sequence is 7.4 ± 1.2 s (6.1 – 10.5 s). Each echeme consists of two closelyspaced syllables and has an average duration of 114.7 ± 11.1 ms (100.1 – 127.9 ms). The interval between consecutive echemes is 92.3 ± 11.6 ms (72.6 – 107.5 ms). Each syllable has an average duration of 45.4 ± 5.0 ms (36.3 – 53.1 ms) and the interval between consecutive syllables is 17.5 ± 4.5 ms (9.5 – 22.2 ms). The dominant frequency is 6.28 ± 0.03 kHz (6.20 – 6.32 kHz).	en	Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C., Robillard, Tony (2022): New taxa and notes on bark and bush crickets (Orthoptera, Grylloidea, Gryllidae Landrevinae and Podoscirtinae) from Sabah. Zootaxa 5178 (3): 201-228, DOI: https://doi.org/10.11646/zootaxa.5178.3.1
