taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037395/files/figure.png	http://doi.org/10.5281/zenodo.7037395	FIGURE 2. Holotype of Tlalocohyla celeste sp. nov. (UCR 23700). (A) Dorsal view. (B) Ventral view.	FIGURE 2. Holotype of Tlalocohyla celeste sp. nov. (UCR 23700). (A) Dorsal view. (B) Ventral view.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037397/files/figure.png	http://doi.org/10.5281/zenodo.7037397	FIGURE 3. Holotype of Tlalocohyla celeste sp. nov. (UCR 23700) (A) Dorsal view of head in living specimen. (B) Lateral view of head in preserved specimen. (C) Lateral view in preserved specimen, note the tympanum. Scale bar = 2 mm. (D) Right hand in ventral view. (E) Right foot in ventral view.	FIGURE 3. Holotype of Tlalocohyla celeste sp. nov. (UCR 23700) (A) Dorsal view of head in living specimen. (B) Lateral view of head in preserved specimen. (C) Lateral view in preserved specimen, note the tympanum. Scale bar = 2 mm. (D) Right hand in ventral view. (E) Right foot in ventral view.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037399/files/figure.png	http://doi.org/10.5281/zenodo.7037399	FIGURE 4. Variation in dorsal pattern of Tlalocohyla celeste sp. nov. (A–B) male holotype of Tlalocohyla celeste sp. nov. (UCR 23700) in life. (C) Size differences between females and males. (D) Typical nocturnal pattern of males. (E) Female with the oocytes visible by transparency in the body. (F) Diurnal pattern of adults, males and females, note the white iris chance. (G) Juvenile. (H) Diurnal pattern with red-brown iris.	FIGURE 4. Variation in dorsal pattern of Tlalocohyla celeste sp. nov. (A–B) male holotype of Tlalocohyla celeste sp. nov. (UCR 23700) in life. (C) Size differences between females and males. (D) Typical nocturnal pattern of males. (E) Female with the oocytes visible by transparency in the body. (F) Diurnal pattern of adults, males and females, note the white iris chance. (G) Juvenile. (H) Diurnal pattern with red-brown iris.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037406/files/figure.png	http://doi.org/10.5281/zenodo.7037406	FIGURE 5. Variation and diagnostic characteristics in Tlalocohyla celeste sp. nov. (A-B) pattern change and green tones change of two uncollected males. (C) Ventral view in diurnal pattern. (D) ventral view of female (UCR 23701), note green bones (black arrow) and white peritoneum (blue arrow). (E) Spotted pattern in juvenile. (F) Axillary membrane (red arrow).	FIGURE 5. Variation and diagnostic characteristics in Tlalocohyla celeste sp. nov. (A-B) pattern change and green tones change of two uncollected males. (C) Ventral view in diurnal pattern. (D) ventral view of female (UCR 23701), note green bones (black arrow) and white peritoneum (blue arrow). (E) Spotted pattern in juvenile. (F) Axillary membrane (red arrow).	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037410/files/figure.png	http://doi.org/10.5281/zenodo.7037410	FIGURE 6. Ventral view Tlalocohyla celeste sp. nov. (A) Ventrals skin texture, and cloacal view. (B) Liver of recently preserved specimen (ECB-Anf-f50-08-01,01.). (C) Nuptial pads of the left hand in Tlalocohyla celeste sp. nov. male holotype (UCR 23700), in life, and preserved.	FIGURE 6. Ventral view Tlalocohyla celeste sp. nov. (A) Ventrals skin texture, and cloacal view. (B) Liver of recently preserved specimen (ECB-Anf-f50-08-01,01.). (C) Nuptial pads of the left hand in Tlalocohyla celeste sp. nov. male holotype (UCR 23700), in life, and preserved.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037412/files/figure.png	http://doi.org/10.5281/zenodo.7037412	FIGURE 7. Vocal sac of uncollected males of Tlalocohyla celeste sp. nov. (A) Ventral view and vocal sac deflated. (B–D). Calling males perched on vegetation in different positions, note the vocal sac extension.	FIGURE 7. Vocal sac of uncollected males of Tlalocohyla celeste sp. nov. (A) Ventral view and vocal sac deflated. (B–D). Calling males perched on vegetation in different positions, note the vocal sac extension.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037393/files/figure.png	http://doi.org/10.5281/zenodo.7037393	FIGURE 1. Bayesian phylogenetic tree within the Hylidae family based on 12S and 16S mitochondrial DNA genes. Posterior probabilities in nodes, asterisks represent support of>95. Branches are collapsed to illustrate genus-level relationships, see expanded branches on Suppl. material. 2. The scale bar refers to the estimated substitutions per site.	FIGURE 1. Bayesian phylogenetic tree within the Hylidae family based on 12S and 16S mitochondrial DNA genes. Posterior probabilities in nodes, asterisks represent support of>95. Branches are collapsed to illustrate genus-level relationships, see expanded branches on Suppl. material. 2. The scale bar refers to the estimated substitutions per site.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037420/files/figure.png	http://doi.org/10.5281/zenodo.7037420	FIGURE 11. Metamorphosis of Tlalocohyla celeste sp. nov. (A) Tadpole at stage 35. (B) Metamorph of 11 mm of SVL. (C) Recently metamorphosed juvenile of 7 mm. (D) Juvenile.	FIGURE 11. Metamorphosis of Tlalocohyla celeste sp. nov. (A) Tadpole at stage 35. (B) Metamorph of 11 mm of SVL. (C) Recently metamorphosed juvenile of 7 mm. (D) Juvenile.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037423/files/figure.png	http://doi.org/10.5281/zenodo.7037423	FIGURE 12. Tlalocohyla celeste sp. nov. tadpole at stage 34 (35) according to Gosner (1960). (A) Lateral view, (B) dorsal view, (C) ventral view, (D) oral disc.	FIGURE 12. Tlalocohyla celeste sp. nov. tadpole at stage 34 (35) according to Gosner (1960). (A) Lateral view, (B) dorsal view, (C) ventral view, (D) oral disc.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037414/files/figure.png	http://doi.org/10.5281/zenodo.7037414	FIGURE 8. Calls repertory of Tlalocohyla celeste sp. nov. Call type I: (A) oscillogram of a call series type I, first growing in amplitude and the second and third with similar amplitude notes; (B) oscillogram and (C) spectrogram of a call type I with 13 notes; (D) power spectrum and (E) oscillogram of a single note. Call type II: (F) oscillogram and (G) spectrogram of a call group: call type I following call type II; (H) oscillogram and (I) power spectrum of the single squeak-like note in call Type II. Dotted red line indicates the dominant frequency.	FIGURE 8. Calls repertory of Tlalocohyla celeste sp. nov. Call type I: (A) oscillogram of a call series type I, first growing in amplitude and the second and third with similar amplitude notes; (B) oscillogram and (C) spectrogram of a call type I with 13 notes; (D) power spectrum and (E) oscillogram of a single note. Call type II: (F) oscillogram and (G) spectrogram of a call group: call type I following call type II; (H) oscillogram and (I) power spectrum of the single squeak-like note in call Type II. Dotted red line indicates the dominant frequency.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037416/files/figure.png	http://doi.org/10.5281/zenodo.7037416	FIGURE 9. Call repertoire of Tlalocohyla celeste sp. nov. Call type III: (A) oscillogram and (B) spectrogram of a call series: call type I following call type III; (C) oscillogram and (D) power spectrum of notes in call type III. Dotted red line indicates the dominant frequency. (E) Oscillogram of call groups in Tlalocohyla celeste sp. nov. Showing a combination of the three call types (type I in orange, type II in green and type III in red) emitted by different males during the chorus.	FIGURE 9. Call repertoire of Tlalocohyla celeste sp. nov. Call type III: (A) oscillogram and (B) spectrogram of a call series: call type I following call type III; (C) oscillogram and (D) power spectrum of notes in call type III. Dotted red line indicates the dominant frequency. (E) Oscillogram of call groups in Tlalocohyla celeste sp. nov. Showing a combination of the three call types (type I in orange, type II in green and type III in red) emitted by different males during the chorus.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037418/files/figure.png	http://doi.org/10.5281/zenodo.7037418	FIGURE 10. Oviposition process of Tlalocohyla celeste sp. nov. (A) A pair in axillary amplexus with egg deposition. (B-D) Oviposition at the end of different leaves. (E) 5 days of development. (F) 9 days of development.	FIGURE 10. Oviposition process of Tlalocohyla celeste sp. nov. (A) A pair in axillary amplexus with egg deposition. (B-D) Oviposition at the end of different leaves. (E) 5 days of development. (F) 9 days of development.	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037425/files/figure.png	http://doi.org/10.5281/zenodo.7037425	FIGURE 13. Geographic distribution of the genus Tlalocohyla in Mesoamerica. Inset, map of Costa Rica, showing the distribution of Tlalocohyla loquax (green circles) and Tlalocohyla celeste sp. nov. (red circle). Sources: (GBIF.org 2022).	FIGURE 13. Geographic distribution of the genus Tlalocohyla in Mesoamerica. Inset, map of Costa Rica, showing the distribution of Tlalocohyla loquax (green circles) and Tlalocohyla celeste sp. nov. (red circle). Sources: (GBIF.org 2022).	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
E5316619D55E4C46FF28FCEBA642FE08.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7037427/files/figure.png	http://doi.org/10.5281/zenodo.7037427	FIGURE 14. Habitat where Tlalocohyla celeste sp. nov. is found. General view of the north (A) and southwest (B) sector of the wetland. The habitat is dominated by the grass Rhynchospora corymbosa (C), but other species of plants tolerant to flooded environments can be used by the frog for its reproductive activity (D).	FIGURE 14. Habitat where Tlalocohyla celeste sp. nov. is found. General view of the north (A) and southwest (B) sector of the wetland. The habitat is dominated by the grass Rhynchospora corymbosa (C), but other species of plants tolerant to flooded environments can be used by the frog for its reproductive activity (D).	2022-08-31	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex		Zenodo	biologists	Varela-Soto, Donald;Abarca, Juan G.;Brenes-Mora, Esteban;Aspinall, Valeria;Leenders, Twan;Shepack, Alex			
