identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
13320672CD6C5740B1F64CEDE0A1337D.text	13320672CD6C5740B1F64CEDE0A1337D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bocchus rex Olmi & Eggs & Capradossi & van de Kamp & Perkovsky & Guglielmino & Vasilenko 2022	<div><p>Bocchus rex sp. nov.</p> <p>Type material.</p> <p>Holotype (= specimen 1; Fig. 1): f#, in SIZK: Ukraine: Les-1, specimen in upper Eocene Rovno amber, collected in Lisove amber mine in Volyn Region of W Ukraine, 9 km east of Manevichi (the former Manevichi district, now Kamen-Kashirsky district). Horizon: Priabonian (35-38 Ma). Paratype (= specimen 2; Figs 2, 3): 1f#, in SMNS: UKR-1, specimen in Rovno amber (unknown locality).</p> <p>Diagnosis.</p> <p>Macropterous female of Bocchus (Figs 1a-d, 2a-c) with OOL more than three times as long as OPL (Fig. 1b); epicnemium concealed (Fig. 1d); notauli incomplete, reaching about 0.75 × length of mesoscutum (Figs 1b, 2c); fore wing with one dark transverse band (Figs 1a, 2c); petiole distinctly visible (Fig. 1a); enlarged claw (Figs 1e, f, 2d, 4a) with one long row of small teeth, in addition to one lamella; protarsomere 5 (Figs 1e, f, 2d, 4a) with distal apex broad and dark pigmented, with one preapical lamella and inner band, without bristles on inner margin.</p> <p>Description of the female</p> <p>(Figs 1 - 4a). Fully winged (Figs 1a-d, 2a-c); length 2.8-3.2 mm (holotype 3.2 mm). Holotype ferruginous-black; paratype black. Antenna clavate (Fig. 1a, b), without ADOs; antennomeres in following proportions: 9:6:6:5:5:4.5:4.5:4.5:4.5:6; antennomere 9 slightly longer than broad (4.5:3). Head dull, completely granulate, not reticulate rugose; frontal line complete; occipital carina complete; POL = 3; OL = 2; OOL = 5; OPL = 6.5; TL = 6; greatest breadth of lateral ocellus shorter than POL (2:3). Mandible quadridentate, with one smaller intermediate tooth (Fig. 1d). Mesosoma longer than head (18:7), shorter than metasoma (18:23). Pronotum crossed by strong transverse impression, with sculpture not distinct, laterally with some longitudinal keels; pronotal tubercle reaching tegula. Mesoscutum dull, granulate. Notauli incomplete, reaching about 0.80 × length of mesoscutum (Figs 1b, 2c). Sculpture of mesoscutellum and metanotum not distinctly visible. Epicnemium concealed (Fig. 1d). Metapectal-propodeal complex not distinctly visible. Fore wing with one dark broad transverse band (Figs 1a, 2c); distal part of 2r-rs&amp;Rs vein much longer than proximal part (17:5). Petiole very long, much shorter than rest of metasoma (4:19). Proleg ratio: 12 (procoxa): 10 (protrochanter): 24 (profemur): 17 (protibia): 13 (protarsomere 1): 2 (protarsomere 2): 3 (protarsomere 3): 7 (protarsomere 4): 11 (protarsomere 5). Enlarged claw (Figs 1e, f, 2d, 4a) with one long row of small teeth (number not distinct) + one lamella. Protarsomere 5 (Figs 1e, f, 2d, 4a) with distal apex broad and dark pigmented, with one preapical lamella and inner band, without bristles on inner margin. Rudimentary claw present, although hardly visible. Mesoleg ratio: 9 (mesocoxa): 4 (mesotrochanter): 19 (mesofemur): 21 (mesotibia): 30 (tarsus). Metaleg ratio: 10 (metacoxa): 6 (metatrochanter): 27 (metafemur): 30 (metatibia): 25 (tarsus). Tibial spurs 1,?, 1.</p> <p>Male. Unknown.</p> <p>Hosts. Unknown.</p> <p>Etymology.</p> <p>Bocchus was the name of two kings of Mauretania (the first being father-in-law to the Numidian King Jugurtha) and Bocchus rex is an appropriate epithet of this nice species with its regal look.</p> <p>State of preservation of paratype</p> <p>(= specimen 2). The head and thorax are well preserved but each have a fissure dorsally (Fig. 3c, e). The metasoma has been completely crushed (Figs 2c, 3a, c, e, g, Suppl. material 1: Video S1). Its size and form can be roughly estimated from the cavity formed in the amber. However, cavities are also present laterally between the head and the thorax (Figs 2a-c, 3a, c, e, g, i, Suppl. material 1: Video S1). Unfortunately, the resolution of the SR-µCT scan was too low to 3D-reconstruct the chelae in great detail. In the head, the partially preserved optical ganglion complex, presumably the optic lobes (= medullae), is clearly visible (Fig. 3b, d, f, h, j, Suppl. material 1: Video S1). It is the second report of optic lobe preservation for Rovno amber arthropods (cf. fig. 1 of Sukhomlyn et al. 2022). Other internal structures are partially preserved; however, it is unclear whether they belong to the optical system or different brain regions, or are partially preserved muscles of the mouthparts (Fig. 3b, d, f, h, j, Suppl. material 1: Video S1). In addition, many muscles in the thorax, the legs and, occasionally, in the metasoma are still preserved (Fig. 3b, d, f, h, j, Suppl. material 1: Video S1). However, as the depicted set of muscles is by no means complete and plays no role in species identification, it will not be discussed further here.</p> <p>Remarks.</p> <p>After the description of Bocchus rex sp. nov., the key published by Perkovsky et al. (2020) can be modified as follows.</p> </div>	http://treatment.plazi.org/id/13320672CD6C5740B1F64CEDE0A1337D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Olmi, Massimo;Eggs, Benjamin;Capradossi, Leonardo;van de Kamp, Thomas;Perkovsky, Evgeny E.;Guglielmino, Adalgisa;Vasilenko, Dmitry V.	Olmi, Massimo, Eggs, Benjamin, Capradossi, Leonardo, van de Kamp, Thomas, Perkovsky, Evgeny E., Guglielmino, Adalgisa, Vasilenko, Dmitry V. (2022): A new species of Bocchus from upper Eocene Rovno amber (Hymenoptera, Dryinidae). Journal of Hymenoptera Research 92: 257-272, DOI: http://dx.doi.org/10.3897/jhr.92.87084, URL: http://dx.doi.org/10.3897/jhr.92.87084
