identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
220587AFFFB4FFAF14C8FC6CFC4D1F4B.text	220587AFFFB4FFAF14C8FC6CFC4D1F4B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Formicidae Latreille 1809	<div><p>Family Formicidae Latreille, 1809</p> <p>Revision to the male-based key to global subfamilies. Due to the unconstricted condition of abdominal segment IV in † Desyopone hereon gen. et sp. nov., in addition to presence of the pleural sulcus and absence of the jugal lobe, it is necessary to revise couplet 8 from the global male-based key to subfamilies from Boudinot [42]. Modifications to this couplet are indicated by italics:</p> <p>8. Abdominal segment IV with cinctus (=constriction) between the pre- and postsclerite or jugal lobe present or oblique mesopleural sulcus absent or indistinct or petiole without tergosternal fusion... 9 (Ponerinae, Apomyrminae, Amblyoponinae, Proceratiinae, Dorylinae part, †Prionomyrmecini [Myrmeciinae], and Ectatomminae).</p> <p>–. Abdominal segment IV without a cinctus and jugal lobe absent and oblique mesopleural sulcus always present and petiole with complete tergosternal fusion, i.e., petiolar posttergites and poststernites fused along their lengths... 19 (“Formicomorpha” sensu Bolton [11], i.e., Aneuretinae, Dolichoderinae, Formicinae).</p> <p>Note also: An additional emendation is necessary: in couplet 11, the specimen that the lead author had previously identified as Dolioponera (Ponerini, CASENT 090028) and used for construction of the key is correctly identified as an aberrant member of Dorylinae; it may represent the unknown male of Vicinopone.</p> </div>	https://treatment.plazi.org/id/220587AFFFB4FFAF14C8FC6CFC4D1F4B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB4FFAC14C8F989FD3D1DEF.text	220587AFFFB4FFAC14C8F989FD3D1DEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aneuretinae Emery 1913	<div><p>Subfamily Aneuretinae Emery, 1913.</p> <p>Type genus. Aneuretus Emery, 1893.</p> <p>Diagnosis (all adult castes). All adults of Aneuretinae possess the following diagnostic plesiomorphies (1–6): (1) mandibles shovel-shaped (=“triangular”) [plesiomorphy of Poneroformicia, or at least Doryloformicia]; (2) meso- and metatibiae with one spur each [synapomorphy of clade Dolichoderomorpha]; (3) petiole with complete tergosternal fusion [synapomorphy of clade Dolichoderomorpha]; (4) petiole with an elongate anterior peduncle [possible synapomorphy of clade Myrmechoderines]; (5) helcium infraaxial [possible synapomorphy of clade Myrmechoderines]; and (6) abdominal segment IV unconstricted [synapomorphy of clade Dolichoderomorpha]. Males and queens of Aneuretinae share the following diagnostic traits (7–12): (7) near-complete fore wing venation, with only the subdiscal cell open; (8) crossvein 2rs-m furcal to post-furcal, i.e., 2r-sm meeting Rs at or distad 2r-rs; (9) well-prefurcal fore wing crossvein cu-a, i.e., cu-a meeting M+Cu proximad the split of M+Cu by more than one of its own lengths; (10) absence of free M in the hind wing after the juncture of rs-m and Mf1, i.e., the abscissa between Sc+ R +Rs and M+Cu linear, without a kink; (11) hind wing anal cell short, its length less than half that of the basal cell; and (12) absence of the hind wing jugal lobe. Males of Aneuretinae have the following diagnostic plesiomorphy (13): (13) genital gonocoxa and gonostylus not strongly differentiated in size, with the dorsal gonocoxal margin continuing more-or-less evenly to that of the gonostylus. Workers and queens of Aneuretinae share the following diagnostic plesiomorphies (14, 15): (14) mandible with biseriate dentition, i.e., with small teeth interspersed between large teeth [synapomorphy of Dolichoderomorpha]; (15) basal and masticatory margins of mandible not marked, i.e., these margins curving into one another, without a distinct angle [synapomorphy of Dolichoderomorpha].</p> <p>Remarks. The operational paleontological definition of Aneuretinae has relied on character states 2, 4, 6, 7 (regardless of subdiscal cell state), and 12. With the explicit recognition of character state 3—which was previously indicated for the “formicomorph subfamilies” by Bolton [11] (p. 16)—it is possible to reject the placement of † Desyopone hereon gen. et sp. nov. from the Aneuretinae. The condition of helcial axiality is here reinterpreted from Bolton [11] (p. 18), who described the helcium of Aneuretinae as “high on [the] anterior face of abdominal segment III”, which is interpretable as supraaxial sensu Keller [46]. Although the helcial tergite of worker Aneuretus is dorsoventrally short, it can be seen that the helcium is at the ventralmost position of the sternum, which does not broaden. An axial helcium is confirmed for the Baltic amber taxa † Paraneuretus and † Protaneuretus as well. Wing venation was observable for Aneuretus and † Paraneuretus. Finally, we recognize the mandibular character states 14 and 15 as critical for the identification of Aneuretinae. The states of the mandibles have not been previously remarked upon, but along with the conformation of the clypeus (not defined here), they form the gestalt of the Aneuretinae and Dolichoderinae, which was likely used by Wheeler [7] to place † Paraneuretus and † Protaneuretus, although his justifications were not made explicit. Further refinement of the aneuretine diagnosis via comparative phenomics and traditional comparative morphology is highly desirable.</p> </div>	https://treatment.plazi.org/id/220587AFFFB4FFAC14C8F989FD3D1DEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB7FFAC14C8FB2EFC781FD6.text	220587AFFFB7FFAC14C8FB2EFC781FD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ponerinae Lepeletier de Saint-Fargeau 1835	<div><p>Subfamily Ponerinae Lepeletier de Saint-Fargeau, 1835.</p> <p>Type genus. Ponera Latreille, 1804.</p> <p>Male diagnosis. Males of Ponerinae are best identified at the tribal level as there are as yet no clear male-based synapomorphies for the subfamily. Male Ponerinae share the following key diagnostic plesiomorphies: (1) wing venation complete or nearly complete, with at least four closed cells present; (2) petiole without tergosternal fusion; (3) abdominal segment III not petiolated; and (4) abdominal sternum IX without prongs or teeth.</p> <p>Remarks. The diagnosis provided for Ponerinae above and Platythyreini and Ponerini below collectively represent a revision of the global diagnosis for the subfamily of Boudinot [42]. Male Ponerinae have previously been diagnosed for the Malagasy region [47] and Japan [48, 49]. No single character has been discovered yet that uniquely identifies all male Ponerinae. Presence of posterolateral processes on the petiolar sternum which contact the outer margins of the helcial tergite, recognized as a ponerine synapomorphy for the female castes [46], are either poorly developed in males or obscured by the petiolar tergite, thus necessitating focused study. Notably, whereas female Ponerinae display a high degree of specialization with respect to mandibular and leg characters, these are universally lacking in the conspecific male.</p> </div>	https://treatment.plazi.org/id/220587AFFFB7FFAC14C8FB2EFC781FD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB7FFAD14C8F965FB5E1B63.text	220587AFFFB7FFAD14C8F965FB5E1B63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platythyreini Emery 1901	<div><p>Tribe Platythyreini Emery, 1901.</p> <p>Type genus. Platythyrea Roger, 1863.</p> <p>Male diagnosis. In addition to the ponerine plesiomorphies, male Platythyreini are distinguished by the following: (1) mandibles worker-like, shovel-shaped (=“triangular”) [plesiomorphy]; (2) face between antennal toruli distinctly raised, such that the antennal toruli are directed relatively laterad, and are situated dorsad a depression which may receive the antenna [synapomorphy, homoplastic in Ponerini]; (3) antennal toruli usually close to or indenting the posterior margin of clypeus, toruli never distant from the posterior margin by more than half of one of their diameters [apomorphy]; (4) meso- and metatibiae with two spurs each [plesiomorphy]; (5) jugal lobe present [plesiomorphy]; (6) helcium not distinctly infraaxial, being axial to more-or-less axial, i.e., at about or just slightly below the midheight of abdominal segment III [plesiomorphy]; (7) abdominal segment IV with cinctus [plesiomorphy]; (8) cuticle pruinose [synapomorphy, homoplastic in Ponerini and some Proceratiinae].</p> <p>Remarks. Because male Platythyreini have never been explicitly diagnosed, we found it necessary to provide a diagnosis in order to confirm the identification of the fossils in question. We observe that the mandibular form, tibial spur count, and cuticular sculpture of male Platythyreini are sufficient for identification at the global scale.</p> </div>	https://treatment.plazi.org/id/220587AFFFB7FFAD14C8F965FB5E1B63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB6FFA214C8FA97FBA7188D.text	220587AFFFB6FFA214C8FA97FBA7188D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desyopone Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot 2022	<div><p>Genus † Desyopone gen. nov. Boudinot and Perrichot</p> <p>Type species. † Desyopone hereon sp. nov., by present designation monotypy.</p> <p>ZooBank LSID: urn:lsid:zoobank.org:act: 7228E671-DF5E-44D1-ADE6-62FB6DB34220.</p> <p>Etymology. The genus name is a portmanteau of the traditional ponerine generic suffix, “-pone”, and the acronym for the Deutsches Elektronen-Synchrotron (DESY), whose storage ring and radiation beamline facilities were used to generate the phenomic data that were crucial for the correct identification of the new taxon.</p> <p>Diagnosis. † Desyopone has plesiomorphies 1–4 of Ponerinae and is identifiable as Ponerini at minimum due to the vestigial mandibles and infraaxial helcium. † Desyopone and Cryptopone are uniquely identified among all Ponerinae by: (1) subpetiolar process completely absent, with the poststernite low and nearly flat in profile. The new genus differs from the males of all known Cryptopone by the following: (2) head broader than long, excluding the compound eyes (vs. head narrower than long); (3) mandibles elongate (vs. short); (4) mandibles lobate (vs. spiniform); (5) mesospiracular sclerite evidently absent (vs. this sclerite present); (6) meso- and metatibiae with no spur and one spur, respectively (vs. two spurs each); and (7) petiolar peduncle long, about as long as node is tall (vs. peduncle short, considerable shorter than height of node).</p> <p>Remarks. The identity of Cryptopone is significantly clarified by the phenomic data from the new species and the phylogenomic revision of Branstetter and Longino [50]. Prior to this work, the diagnostic importance of the absent subpetiolar process was obscured by the inclusion of Wadeura guianensis in Cryptopone. Now it is clear that the absence of the subpetiolar process is a unique condition among extant Ponerinae that is shared between † Desyopone gen. nov. and Cryptopone, and thus constitutes a reasonable autapomorphy within the subfamily for the two genera. No known Cryptopone, however, matches the diagnostic character combination of † Desyopone gen. nov., with conditions 3–7 being apomorphic. Critically, the elongate peduncle of † D. hereon gen. et sp. nov. is nearly unique among Ponerinae; this condition is similarly derived in Harpegnathos and is approached by Dinoponera, some Odontomachus (e.g., O. chelifer, O. coquereli), and Platythyrea (although node at middle of segment rather than posterior).</p> </div>	https://treatment.plazi.org/id/220587AFFFB6FFA214C8FA97FBA7188D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB6FFAD14C8FDB2FB6D1C40.text	220587AFFFB6FFAD14C8FDB2FB6D1C40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ponerini Lepeletier de Saint-Fargeau 1835	<div><p>Tribe Ponerini Lepeletier de Saint-Fargeau, 1835.</p> <p>Type genus. Ponera Latreille, 1804.</p> <p>Male diagnosis. In addition to the ponerine plesiomorphies, male Ponerini are distinguished by the following: (1) mandibles vestigial, with an enlarged mandalus, and being variably lobate, spatulate, spiniform, or nub-like (no exceptions known) [synapomorphy, homoplastic among Formicidae]; (2) face between antennal toruli not distinctly raised, thus toruli directed more-or-less dorsally; if the intertorular region is raised, then this region is grooved in appearance due to impression of the supraclypeal area (=frontal triangle) (Plectroctena and Psalidomyrmex with medial torular arches raised, but not face; Hagensia, Megaponera, Ophthalmopone, Simopelta, some Euponera, and some Odontomachus with intertorular region raised) [plesiomorphy]; (3) antennal toruli usually distant from posterior clypeal margin (some Brachyponera, many Leptogenys, Megaponera, and Ophthalmopone have toruli that are close to the clypeal margin) [plesiomorphy]; (4) meso- and metatibiae with two, one, or no spurs; (5) jugal lobe present or absent; (6) helcium distinctly infraaxial, i.e., situated well below the midheight of abdominal segment III (Simopelta is an exception due to softening and reduction in size of the metasoma) [synapomorphy, homoplastic among Formicidae]; (7) abdominal segment IV with or without cinctus; (8) cuticle usually not pruinose, being shiny and variably sculptured (Belonopelta, Hagensia, Megaponera, and Ophthalmopone are exceptions) [plesiomorphy].</p> <p>Remarks. Definitive infraaxiality in males, i.e., with abdominal tergum III rising high above the petiole, is a strong diagnostic condition for Ponerinae, as this is an infrequent apomorphic condition at the subfamily level. It also occurs in Dolichoderinae and Formicinae, and to some extent in Myrmeciinae and various Myrmicinae. Discothyrea (Proceratiinae) may approach infraaxiality, but the third abdominal tergum is low.</p> </div>	https://treatment.plazi.org/id/220587AFFFB6FFAD14C8FDB2FB6D1C40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
220587AFFFB9FFA614C9FE4CFB541E29.text	220587AFFFB9FFA614C9FE4CFB541E29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Desyopone hereon Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot 2022	<div><p>† Desyopone hereon sp. nov. Boudinot and Perrichot</p> <p>(Figures 1–5)</p> <p>ZooBank LSID: urn: lsid:zoobank.org:act: 9E345965-6AA1-42D8-A468-187BABBB38D2.</p> <p>Etymology. The specific epithet gratefully recognizes the Helmholtz-Zentrum Hereon, the research center which funds and operates the Imaging Beamline (P05) at DESY, thus making the present work possible.</p> <p>Holotype. Male (m), MAIG 6016 (Figures 1B, 2A,B, 3 and 4), deposited in the Museum of Amber Inclusions at University of Gda´nsk, Poland.</p> <p>Paratypes. 12 males (m). Synincluded with holotype in amber piece MAIG 6016 (Figures 1A,C,D, 2C,D and 5).</p> <p>Type locality. Exact locality unknown in the Bashilo river gorge near Weldiya, Semien Wollo Zone, Amhara Region, Ethiopia.</p> <p>Type horizon. A fine siltstone/mudstone of Early Miocene age (16–23 Ma).</p> <p>Material. Holotype and paratypes 1–4 are nearly complete, preserved without apparent distortion but with integument entirely covered by a white, opaque (bacterial?) coat for paratypes 1, 2, 4 (Figures 1A,C and 2A). Paratypes 5–12 mostly complete but variously preserved, more or less distorted by apparent dehydration (e.g., Figure 1D).</p> <p>Diagnosis. † Desyopone hereon is uniquely identifiable among all Ponerinae by the distinctly elongate petiolar peduncle and the enlarged, lobate mandibles.</p> <p>Description. Measurements: Holotype (paratypes) (taken from paratypes 1,2,4)—BL 3.11 (3.00–3.50), HW 0.57 (0.50–0.60), HWE 0.70 (0.66–0.70), SL 0.12 (0.10–0.12), EL 0.23, OLL 0.08 (0.08), OIL 0.16 (0.16), WL 1.25 (1.25), ML 0.52 (0.50–0.56), MW 0.49, FWL (2.75), PH 0.26 (0.30), PL ~0.40 (0.40), GL 1.20 (1.00–1.50), GW 0.62 (0.65–0.80), HWI 81 (76–86), SI 21 (20), OCI 50 (50), PI 154 (133).</p> <p>Head (Figures 2–4): Head capsule ovoid-elliptical in lateral view; in full-face view, posterior head margin broadly and evenly convex to compound eyes; oral region of head, i.e., the malar areas, clypeus, and mouthparts, narrower than distance between compound eyes; postgenal bridge short, about 2/5 the length of the head in full-face view as measured from the postocciput to the hypostoma; malar areas distinctly developed; clypeus medially bulging, laterally depressed. Compound eyes situated almost entirely in anterior half of head; eyes subspherical and relatively small, with their length being about 1/3 head length. Ocelli distant from compound eyes, with the lateral ocelli separated from the compound eyes by slightly more than one compound eye length. Antennal toruli located at about head midlength in full-face view, distinctly posterad the posterior clypeal margin. Antennal scapes short, just barely longer than wide, with their length distinctly &lt;2 lateral ocellus diameters; scape length slightly more than one pedicel length, but not more than two. Pedicel about 1/3 the length of the first flagellomere. Flagellum narrow and long, with their length greatly exceeding mesosoma length. Mandibles flat and lobate in appearance, without distinct masticatory and basal margins; medial mandibular margin convex, curving more-or-less evenly around apex, which does not have incurvature; mandible length slightly greater than compound eye length. Labrum and paraglossae dangling at rest, both distinctly narrower than the distance between the mandibular bases. Maxillary stipes without transverse ridging.</p> <p>Mesosoma (Figures 2 and 3): Pronotum short and simple but with distinct muscular convexity as seen in lateral and dorsal views; posterad the anteromedian pronotal lobe (“nuchal lobe”), pronotum in the form of a simple arch, without distinct dorsal and ventral surfaces. Propleurae widely emarginate posteromedially, together forming a broad arch for the prosternum. Prosternum with basisternum apparently arcuate anteriorly; prosternal process developed. Mesoscutum somewhat narrow, with the anteroposterior length slightly greater than the lateromedial width. Notauli developed, and Y-shaped; meeting in the posterior half of the mesoscutum. Parapsides developed, although indistinct. Scutoscutellar sulcus broad, with at least five cross-costae. Mesoscutellum simple, convex, longer than broad in full-face view. Oblique mesopleural sulcus developed. Spiracular lobe (ventrad wing insertions) absent. Mesopleural area divided into dorsal and ventral regions; both regions dorsoventrally taller than anteroposteriorly long. Mesosternal and metasternal regions without processes. Propodeum convex, without armature or distinct sculpturation. Propodeal lobes developed, weak.</p> <p>Legs: Mesotibiae with no spur, metatibiae with a single, pectinate spur. Pretarsal claws narrow and simple, without additional teeth. Arolia well-developed but not very large.</p> <p>Fore wings (Figures 2C,D and 5): Costal vein (C) present, complete. Rsf1 diverging from Sc+ R +Rs well proximad pterostigma, with Sc+ R abscissa about 1/3 pterostigma length. Pterostigma well-developed, long, and narrow, with its length&gt; 5 × its width. Rf distad pterostigma tubular. M+Cu tubular. Mf1 diverging from M+Cu at or slightly distad crossvein cu-a; this abscissa very weakly curved and meeting Rf1 at a distinct oblique angle. Rs+M tubular. Rsf2–3 diverging from Rs+M proximad 1m-cu. Crossvein 1r-rs absent. Crossvein 2r-rs anterior juncture at 2/3 length of pterostigma; this crossvein meeting Rsf proximad crossvein 2rs-m, which is tubular (paratype 4 has 2rs-m duplicated on the right wing, see Figure 2D). Rsf4+ tubular, meeting Rf distally). Mf2 (=abscissa between Rs+M and 1m-cu) short, shorter than 2r-rs and 2rs-m. Mf3 similar in length to but distinctly shorter than Rsf2–3. Mf4+ tubular proximally, becoming nebulous distally, with this occurring at a distance that is about 3 × the length of 2rs-m. Cuf3 (=abscissa of Cu after 1m-cu) joined to 1A posteriorly; 1A is tubular for its entire length. Submarginal cells 1 and 2 similar in size and shape, but with 2 distinctly smaller than 1. Marginal cell 1 long and narrow, with a length that is about 1.5 × pterostigma length. Discal cell 1 subrhomboidal, its length slightly less than 2 × its anteroposterior width. Subdiscal cell closed.</p> <p>Hind wings (Figures 2C and 5): Wing with eight distal hamuli. Jugal lobe absent. C not distinctly developed. R splitting from Sc+ R +Rs distad crossvein 1rs-m; Rf incomplete, not reaching anterior wing margin. Rsf tubular for a distance that is about 2 × the length of Sc+ R +Rs distad 1rs-m. M+Cu splitting well distad crossvein cu-a. Mf1 meeting rs-m at a broad, oblique angle; Mf developed as a stud distad this juncture. Cuf developed distad the split of M+Cu, but exact condition uncertain. Crossvein cu-a long, slightly longer than the length of Mf1; this crossvein situated proximad split of M+Cu by about twice its length. Anal vein (A) tubular past its juncture with cu-a. Anal cell relatively long; M+Cu proximad cu-a distinctly longer than M+Cu distad cu-a.</p> <p>Metasoma (Figures 2B and 3A,D,E): Petiole nodiform and distinctly pedunculate, albeit without a marked inflection between its anterior portion and the anterior surface of the petiolar node; peduncle about 2/5 petiole length; petiolar node height about 3/5 entire petiole length; node broad and convex; posterior collar well-developed; tergosternal fusion absent, laterotergites present; sternum low and very weakly sinuate in lateral view, without an anteroventral (=subpetiolar) process, nor with a posterior process; posterior sternal margin distinctly notched. Helcium infraaxial (=below abdominal segment III midheight); helcial tergite broad and overlapping sternite laterally. Prora not distinctly developed. Gastral segments homonomous in appearance and gradually decreasing in length from abdominal segment III to VIII. Abdominal segment IV without cinctus (=constriction). Abdominal spiracles IV–VIII obscured by preceding tergites. Abdominal tergum VIII apparently simple. Abdominal sternum IX lobate and somewhat narrow, with a length that is about twice its width. Cerci (=pygostyles) developed.</p> <p>Genitalia (Figures 2B and 3A): Cupula present (only visible in µ- CT cross-sections). Gonopods longer than tall. Gonostyli broad proximally, indistinctly set off from gonocoxa, and lobate in appearance, being narrowly rounded apically. Lateropenites (=digiti) thickened apically, downcurved. Penites (=penisvalvae) apparently longer than tall, curving strongly to their narrowly lobate apices.</p> <p>Setation: Body with two primary hair classes: (1) short pubescence, which covers the head and all segments of the antennae, mesosoma, legs, and metasoma; (2) long hairs, which are sparse on all body regions, but are denser on the metasoma.</p></div> 	https://treatment.plazi.org/id/220587AFFFB9FFA614C9FE4CFB541E29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Boudinot, Brendon E.;Richter, Adrian K.;Hammel, Jörg U.;Szwedo, Jacek;Bojarski, Błażej;Perrichot, Vincent	Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej, Perrichot, Vincent (2022): Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae). Insects 73 (796): 1-19, DOI: 10.3390/insects13090796, URL: http://dx.doi.org/10.3390/insects13090796
