taxonID	type	description	language	source
5C1DEB197D455BAB9CE8CF89033551FC.taxon	description	Figs 2 A-C, 3, 4	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
5C1DEB197D455BAB9CE8CF89033551FC.taxon	materials_examined	Other material (n = 60). China. Fujian: 1 ♂ (TARI), 建陽 (Jianyang), 黃坑 (Huangkeng), 場头 (Changtou), 700 - 950 m, 23. VI. 1960, leg. 姜勝巧 (S. - Q. Jiang); 1 ♀ (TARI), 建陽 (Jianyang), 黃坑 (Huangkeng), 桂林 (Guiling), 290 m, 21. VI. 1960, leg. 姜勝巧 (S. - Q. Jiang); Taiwan. Nantou: 1 ♀ (TARI), Baibara (= Meiyuan, 眉原), 24. III. 1943, leg. A. Aoki; 1 ♂ (TARI), Horisha (= Puli, 埔里), 10. V. 1913, leg. M. Maki; 1 ♂ (TARI), same locality, 10. IV. 1919, leg. H. Kawamura (collector was not present on the card); 1 ♂ (TARI), Wanfengtsun (萬豐村), 2. IV. 2008, leg. W. - T. Liu; 1 ♂ (TARI), same but with " 18. IV. 2011 "; Pingtung: 1 ♂ (TARI), Shinsuiyei (= Chinshuiying, 浸水營), 17. III. 1926, leg. S. Issiki; 1 ♀ (TARI), Tahanshan (大漢山), 5. IV. 2009, leg. C. - F. Lee; 2 ♂, 1 ♀ (TARI), same but with " 26. III. 2013 "; 12 ♂, 2 ♀ (TARI), same but with " 18. IV. 2018 "; 3 ♂, 1 ♀ (TARI), same locality, 3. IV. 2012, leg. Y. - T. Chung; 6 ♂, 2 ♀ (TARI), same but with " 16. IV. 2013 "; 1 ♂, 1 ♀ (TARI), same but with " 10. V. 2013 "; 2 ♂ (TARI), same but with " 28. III. 2016 "; 2 ♂ (TARI), same but with " 10. IV. 2017 "; 3 ♂ (TARI), same but with " 17. IV. 2017 "; 1 ♂ (TARI), same but with " 5. IV. 2018 "; 2 ♂ (TARI), same but with " 9. IV. 2018 "; 1 ♂ (TARI), same but with " 23. IV. 2018 "; 2 ♂ (TARI), same but with " 27. IV. 2020 "; 1 ♂ (TARI), same but with " 20. III. 2021 "; 1 ♂ (TARI), same but with " 10. IV. 2021 "; 2 ♂ (TARI), same but with " 15. IV. 2021 "; 2 ♂, 1 ♀ (TARI), same locality, 29. IV. 2014, leg. J. - C. Chen.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
5C1DEB197D455BAB9CE8CF89033551FC.taxon	diagnosis	Diagnosis. Adults of Gonioctena (Asiphytodecta) subgeminata are easily distinguished from those of other consubgeneric species, G. (A.) tredecimmaculata, by the following combination of the characters: lacking black spots on the pronotum (Fig. 2 A) (with three black spots on the pronotum in G. (A.) tredecimmaculata (Fig. 2 D )), yellowish brown scutellum (Fig. 2 A) (black scutellum in G. (A.) tredecimmaculata (Fig. 2 D )), three black spots on suture of elytra, at basal 1 / 3, median of apical 1 / 3, and at apices (Fig. 2 A) (two black spots on suture of elytra, apical 1 / 3 and near apices in G. (A.) tredecimmaculata (Fig. 2 D )), punctures on elytra arranged into longitudinal striae (Fig. 2 A) (punctures on elytra entirely confused in G. (A.) tredecimmaculata (Fig. 2 D )), apical process of aedeagus apically wider and apical margin widely rounded (Fig. 3 B) (apically process of aedeagus apically narrow and apical margin narrowly rounded in G. (A.) tredecimmaculata (Figs 5 B, 6 B), slender gonocoxae with angular apices (Fig. 3 D) (wide gonocoxae with irregular apical margin in G. (A.) tredecimmaculata (Figs 5 D, 6 D )), sparse, short setae along outer margins of ventrites VIII (Fig. 3 E) (dense, short setae along outer margins of ventrite VIII in G. (A.) tredecimmaculata (Figs 5 E, 6 E )), relatively wider spermatheca (Fig. 3 F) (more slender spermatheca in G. (A.) tredecimmaculata (Figs 5 F, 6 F )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
5C1DEB197D455BAB9CE8CF89033551FC.taxon	biology_ecology	Biology. A large population of Gonioctena (Asiphytodecta) subgeminata was found in Tahanshan (大漢山), where Mr. Yi-Ting Chung (鍾奕霆) collected leaf beetles often. He collected early instar larvae feeding gregariously on young leaves (Fig. 4 A-C) on 18 March 2022 from Callerya reticulata. In the lab, mature larvae (Fig. 4 D) burrowed into the soil and built underground chambers for pupation on 22 March. Duration of the pupal stage (Fig. 4 E) was seven days. Newly emerged adults (Fig. 4 F) appeared in early April, but females failed to lay eggs. Thus, this species is likely univoltine since adults appear during spring (March to May).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
5C1DEB197D455BAB9CE8CF89033551FC.taxon	distribution	Distribution. China, Taiwan.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
4F48034F44665EE19EAD81AEF20F0BAA.taxon	description	Figs 2 D-F, 5, 6, 7	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
4F48034F44665EE19EAD81AEF20F0BAA.taxon	materials_examined	Other material (n = 128). China. Fujian: 1 ♀ (TARI), 崇安 (Chongan), 星村 (Xingcun), 掛墩 (Guadun), 950 - 1210 m, 12. VI. 1960, leg. 左丞 (C. Zuo), 1 ♀ (TARI), 建陽 (Jianyang), 黃坑 (Huangkeng), 大竹欄 (Dazhulan) - 先鋒嶺 (Xianfengling), 950 - 1170 m, 2. V. 1060, leg. 馬成林 (C. - L. Ma); Taiwan. Chiayi: 1 ♀ (TARI), Arisan (= Alishan, 阿里山), 17. V. 1917, leg. T. Shiraki; 1 ♂ (TARI), Hsiting (隙頂), 1. VIII. 2018, leg. Y. - T. Chung; Hsinchu: 2 ♀ (TARI), Shiigao (name of the tribe), Chikuto (= Maopu, 茅埔), 27 - 30. VI. 1934, leg. M. Chujo; 1 ♀ (TARI), Shinchiku (= Hsinchu, 新竹), 1 - 30. VII. 1918, leg. J. Sonan, K. Miyake; 1 ♂ (TARI), Talu trail (大鹿林道), 22. X. 2008, leg. H. - J. Chen; 1 ♂ (TARI), same locality, 22. IX. 2012, leg. Y. - L. Lin; Hualien: 2 ♂, 2 ♀ (TARI), Chohsi (卓溪), 4. III. 2016, leg. S. - P. Wu; 3 ♂, 2 ♀ (KMNH), Hungyeh Wenchuan (紅葉溫泉), 14. VI. 1976, leg. H. Makihara; 1 ♀ (TARI), Mizuho (= Juisui, 瑞穗), 23. III. 1935, leg. M. Chujo; Ilan: 2 ♂, 3 ♀ (TARI), Ebosiyama (= Umutzushan, 烏帽子山), 17 - 21. V. 1933, leg. M. Chujo; 1 ♀ (TARI), Yingtzuling (鶯仔嶺), 15. IV. 2012, leg. Y. - L. Lin; Kaohsiung: 1 ♀ (TARI), Chienshan (建山), 23. VI. 2018, leg. B. - X. Guo; 1 ♂ (TARI), Hsiaokuanshan (小關山), 15. V. 2016, leg. B. - X. Guo; 2 ♂ (KMNH), Liu Kui (六龜), 29. III. 1986, leg. K. Baba; 2 ♂ (TARI), Paiyunshan (白雲山), 25. IV. 2016, leg. U. Ong; 1 ♂ (TARI), Tengchih (藤枝), 4. VII. 2011, leg. M. - H. Tsou; 1 ♀ (TARI), same locality, 18. IV. 2013, leg. B. - X. Guo; 1 ♀ (TARI), same but with " 3. VIII. 2013 "; 1 ♀ (TARI), same locality, 18. IV. 2013, leg. Y. - T. Chung; Nantou: 1 ♂, 2 ♀ (TARI), Honbukei (= Penpuchi, 本部溪), 7. VII. 1940, leg. M. Chujo; 1 ♂ (TARI), Huisun Experimental Forest Station (惠蓀林場), 22. IV. 2015, leg. B. - X. Guo; 1 ♀ (TARI), Lugu (鹿谷), 10. IX. 2 - 14, leg. H. - T. Shih; 1 ♀ (KMNH), Hsitou (溪頭), 31. III. 1980, leg. K. Sugiyama; 1 ♀ (TARI), Takeya (= Chienchityu, 乾溪仔), 8. VII. 1940, leg. M. Chujo; 1 ♂, 4 ♀ (TARI), Wanfengtsun (萬豐村), 2. IV. 2008, leg. W. - T. Liu; 2 ♂ (TARI), same but with " 9. VII. 2008 "; 1 ♀ (TARI), same but with " 23. IV. 2009 "; 1 ♂ (TARI), same but with " 13. IV. 2010 "; 2 ♂ (TARI), Wushe (霧社), 17. VIII. 1984, leg. K. C. Chou; Pingtung: 1 ♂ (TARI), Koshun (= Hengchun, 恆春), 25. IV. - 25. V. 1918, leg. J. Sonan, K. Miyake, M. Yoshino; 1 ♂ (TARI), Nanjenshan (南仁山), 24. II. 2009, leg. C. - F. Lee; 1 ♂ (TARI), Peihulushan (北湖呂山), 4. XI. 2009, leg. M. - H. Tsou; 2 ♂, 3 ♀ (TARI), Peitawushan (北大武山), 22. IV. 2013, leg. Y. - T. Chung; 1 ♂, 2 ♀ (TARI), same but with " 8. V. 2014 "; 1 ♂ (TARI), same but with " 2. V. 2016 "; 1 ♂ (TARI), same but with " 10. X. 2017 "; 1 ♀ (TARI), Raisha (= Laiyi, 來義), 13. III. 1926, leg. J. Sonan; 1 ♂ (TARI), Shouka (壽卡), 23. II. 2013, leg. W. - C. Liao; 1 ♀ (TARI), Tahanshan (大漢山), 17. VII. 2007, leg. M. - H. Tsou; 2 ♂ (TARI), same locality, 25. V. 2008, leg. C. - F. Lee; 1 ♂ (TARI), same locality, 28. VIII. 2010, leg. Y. - L. Lin; 1 ♂ (TARI), same locality, 26. VI. 2012, leg. Y. - T. Chung; 1 ♂ (TARI), same but with " 30. V. 2014 "; 2 ♀ (TARI), same but with " 18. III. 2016 "; 1 ♂ (TARI), same but with " 6. IX. 2017 "; 1 ♂ (TARI), same but with " 28. VI. 2020 "; 1 ♂ (TARI), same but with " 10. V. 2021 "; 4 ♂, 4 ♀ (TARI), Taiwu (泰武), 14. IX. 2017, leg. Y. - T. Chung; 1 ♀ (TARI), Wanlite (萬里得), 1. III. 2010, leg. M. - L. Cheng; Taichung: 1 ♂ (TARI), Hassenzan (= Pahsienshan, 八仙山), 4. VI. 1942, leg. A. Mutuura; 1 ♂ (TARI), Kukuan (谷關), 14. IV. 2019, leg. C. - C. Guo; 3 ♂ (TARI), same but with " 28. IV. 2019 "; 1 ♂, 1 ♀ (TARI), Wushihkeng (烏石坑), 13. VII. 2008, leg. M. - H. Tsou; 1 ♂ (TARI), same locality, 21. III. 2013, leg. C. - F. Lee; Tainan: 4 ♀ (TARI), Kantoushan (崁頭山), 13. III. 2010, leg. M. - H. Tsou; 5 ♂ (TARI), same but with " 8. IV. 2010 "; 3 ♂, 3 ♀ (TARI), Meiling (梅嶺), 24. IV. 2013, leg. B. - X. Guo; 1 ♂ (TARI), same but with " 5. X. 2015 "; 2 ♀ (TARI), same locality, 2. XI. 2015, leg. Y. - T. Chung; Taipei: 1 ♂ (TARI), Chungho (中和), 22. V. 2010, leg. Y. - L. Lin; 1 ♂ (TARI), Fushan (福山), 11. IV. 2021, leg. I. - H. Ku; 1 ♂ (TARI), Kotou (格頭), 27. III. 2010, leg. H. - J. Chen; 1 ♂ (TARI), Urai (= Wulai, 烏來), 14. V. 1933, leg. M. Chujo; 1 ♀ (TARI), same locality, 16. IV. 2008, leg. J. - F. Tsai; 1 ♀ (KMNH), Yangmingshan (陽明山), 28. IV. 1970, leg. M. Yamamoto; Taitung: 1 ♀ (TARI), Anshuo (安朔), 27. VII. 2011, leg. W. - T. Liu; 2 ♀ (TARI), Chipon (知本), 25. III. 1935, leg. M. Chujo; 1 ♂, 1 ♀ (TARI), same locality (= Tipon), 8. V. 1943, leg. M. Chujo; 1 ♂ (KMNH), Kueitien (歸田), 17. VI. 1976, leg. H. Makihara; 1 ♂, 1 ♀ (TARI), Liyuan (栗園), 19. VI. 2013, leg. C. - F. Lee; 1 ♂ (TARI), Motien (摩天), 18. VI. 2013, leg. J. - C. Chen; 3 ♂, 1 ♀ (TARI), Taimali (太麻里), 20. III. 2008, leg. P. - F. Wang; 1 ♀ (TARI), Taito (= Taitung, 台東), 25. II. - 27. III. 1919, leg. S. Inamura; Taoyuan: 1 ♂ (TARI), Hsuehwunao (雪霧鬧), 10. VI. 2016, leg. Y. - L. Lin.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
4F48034F44665EE19EAD81AEF20F0BAA.taxon	diagnosis	Diagnosis. Gonioctena (Asiphytodecta) tredecimmaculata is easily distinguished from the other consubgeneric species, G. (A.) subgeminata by the following combination of the characters: three black spots on pronotum (Fig. 2 D) (without black spots on pronotum in G. (A.) subgeminata (Fig. 2 A )), black scutellum (Fig. 2 D) (yellowish-brown scutellum in G. (A.) subgeminata (Fig. 2 A )), two black spots on suture of elytra, one at apical 1 / 3 and the other near apices (Fig. 2 D) (three black spots on suture of elytra, one at basal-median 1 / 3 median and apical 1 / 3, the other at apices in G. (A.) subgeminata (Fig. 2 A )), punctures on elytra entirely confused (Fig. 2 D) (punctures on elytra arranged into longitudinal striae in G. (A.) subgeminata (Fig. 2 A )), apical process of aedeagus apically narrow and apical margin narrowly rounded (Figs 5 B, 6 B) (apical process of aedeagus apically wider and apical margin widely rounded in G. (A.) subgeminata (Fig. 3 B )), wide gonocoxae with irregular apical margin (Figs 5 D, 6 D) (slender goncoxae with angular apices in G. (A.) subgeminata (Fig. 3 D )), dense, short setae along outer margins of ventrite VIII (Figs 5 E, 6 E) (sparse short setae along outer margins of ventrite VIII in G. (A.) subgeminata (Fig. 3 E )), more slender spermatheca (Figs 5 F, 6 F) (relatively wider spermatheca in G. (A.) subgeminata (Fig. 3 F )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
4F48034F44665EE19EAD81AEF20F0BAA.taxon	biology_ecology	Biology. Gonioctena (Asiphytodecta) tredecimmaculata populations are presumed to be univoltine and females are ovoviviparous. In Taiwan, overwintered adults appeared during early March. The females deposited larvae that were enclosed within the chorion (Fig. 7 B), and they hatched after several hours (Fig. 7 C). The larvae fed on leaves and the larval duration was eight days. Mature larvae (Fig. 7 D) burrowed into soil and built underground chambers for pupation. Duration of the pupal stage (Fig. 7 E) was 12 days. Newly emerged adults (Fig. 7 F) appeared in mid-April. The biology of this species in In Jiangxi, China (Shen and Xue 1986) is similar to that of Taiwanese populations. However, overwintered adults appeared during late April. The larvae required ten to 12 days in May to mature and pupal duration was 13 to 16 days. The adults emerged during early June.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
4F48034F44665EE19EAD81AEF20F0BAA.taxon	distribution	Distribution. China, Taiwan.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
67941529ED425AFAA5A4E164365834BE.taxon	description	Figs 8 D-F, 22	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
67941529ED425AFAA5A4E164365834BE.taxon	materials_examined	Other material (n = 17). China. Heilongjiang: 1 ♂ (NMHUK), Erlungshan, 29. V. 1966, leg. P. M. Hammond; 1 ♀ (NMHUK), Habrin City (哈爾濱市), 10. VI. 1950; 5 ♂, 1 ♀ (NMHUK), same but with " 22. VI. 1952 "; 1 ♂, 1 ♀ (NMHUK), same but with " 29. VI. 1952 "; North Korea. 2 ♂ (TARI), Husen-Valley, Kankyo-Nando, 14. VII. 1937, leg. Y. Yano; 5 ♂, 2 ♀ (TARI), Mt. Myoko (= Mt. Myohyang, 妙香山), 25. VII. 1937, leg. M. Yamada; South Korea. 1 ♀ (TARI), Suigen, Keiki-Do, 3. VI. 1933, leg. D. Okamoto; Russia Far East. 1 ♀ (MNHUK), Primorskii krai Lazovski Zapovehik, 3 - 14. V. 2001, leg. M. Quest.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
67941529ED425AFAA5A4E164365834BE.taxon	diagnosis	Diagnosis. Adults of Gonioctena (Brachyphytodecta) fulva are similar to yellowish brown adults of G. (B.) scutellaris but differ in possessing a black head, thoracic and abdominal ventrites (Fig. 8 E) (yellowish brown head, thoracic and abdominal ventrites in G. (B.) scutellaris (Fig. 8 B )) and short apical process of aedeagus (Fig. 22 B) (extremely elongate apical process of aedeagus in G. (B.) scutellaris (Fig. 10 B )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
67941529ED425AFAA5A4E164365834BE.taxon	distribution	Distribution. China, Korea, Russia.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
432D91A423A65330A51FE3236898EF42.taxon	description	Figs 9 D-F, 12, 13	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
432D91A423A65330A51FE3236898EF42.taxon	description	Description. Length 5.3 - 6.1 mm, width 3.2 - 3.9 mm. Body color (Fig. 9 D-F) blackish brown; antennomeres I-V, sides of pronotum and hypomeron yellowish brown; elytra with wide yellowish brown outer margins and suture. Antennae (Fig. 12 A) with antennomere III slender, IV and V slightly swollen, VI and VII moderately swollen, VIII-X strongly swollen, XI elongate oval, length ratios of antennomeres I-XI 1.0: 0.5: 0.6: 0.5: 0.4: 0.4: 0.5: 0.6: 0.6: 0.6: 1.0, length to width ratios of antennomeres I-XI 2.1: 1.5: 2.2: 1.6: 1.5: 1.0: 1.0: 1.0: 1.1: 1.0: 1.8. Pronotum 2.3 x wider than long, lateral margins widest at base, convergent anteriorly, anterior angles strongly produced; anterior and lateral margins bordered, lateral margins barely visible in dorsal view; trichobothria absent on both anterior and posterior angles; disc covered with sparse fine punctures and mixed with finer punctures; both sides covered with much larger, denser punctures. Scutellum distinctly wider than long, narrowed posteriorly. Elytra 1.3 x longer than wide; lateral margins slightly wider posteriorly, widest near middle; humeral calli well developed; disc covered with rather regular coarse punctures arranged into single stria; interspaces covered with fine, sparse punctures. Hind wing well developed. Aedeagus (Fig. 12 B. C) with apical process wide and short in dorsal view, 0.1 x as long as aedeagus; lateral margins slightly narrowed in basal 1 / 3; strongly curved in lateral view; endophallic sclerite extremely elongate. Gonocoxae (Fig. 12 D) narrow, but apically narrowed, with few long setae along apical margins. Ventrite VIII (Fig. 12 E) transverse, with several long setae along outer margin. Receptacle of spermatheca (Fig. 12 F) slightly swollen, not separated from pump; pump short and curved; sclerotized proximal spermathecal duct moderately short.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
432D91A423A65330A51FE3236898EF42.taxon	diagnosis	Diagnosis. Gonioctena (Brachyphytodecta) liui sp. nov. can be distinguished from the other consubgeneric species, G. (B.) scutellaris, by the following combination of the characters: black head, most of pronotum (Fig. 9 D), and thoracic and abdominal ventrites (Fig. 9 E) (yellowish brown head and pronotum (Figs 8 A, 9 A), thoracic and abdominal ventrites (Figs 8 B, 9 B) in G. (B.) scutellaris), short, wide apical process of aedeagus (Fig. 12 B, C) (extremely elongate apical process of aedeagus in G. (B.) scutellaris (Fig. 10 B, C), and narrow gonocoxae covered with fewer setae (wide gonocoxae covered with more setae in G. (B.) scutellaris). Although Ge et al. (2007) drew only the aedeagi in dorsal and lateral views when G. issikii was mentioned, it is easily identified as this new species. Cho (2016) redescibed G. issikii in detail but it actually fits perfectly this new species. The type specimen was not studied by these authors mentioned above.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
432D91A423A65330A51FE3236898EF42.taxon	biology_ecology	Biology. Gonioctena (Brachyphytodecta) liui populations are presumed to be multivoltine during spring, and females are ovoviviparous. Host plants are one of only a few woody Fabaceae in Taiwan. They grow 7 - 9 m high (Fig. 13 A). They started blooming during April (Fig. 13 B). Most adults (Fig. 13 F) and larvae were collected from flower buds (Fig. 13 C). Females deposited larvae and the larvae (Fig. 13 D) fed on internal tissues of the flower buds through holes chewed by adults. They preferred to feed on the flower buds rather than young sprouts. The larval duration was eight days. Mature larvae (Fig. 13 D) burrowed into the soil and built underground chambers for pupation. The duration of the pupal stage (Fig. 13 E) was nine days at room temperature. Flowering season of the host plant occurs from April to June (Huang and Ohashi 1993), but the blooming period for individual trees is less than a month.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
432D91A423A65330A51FE3236898EF42.taxon	distribution	Distribution. This new species is restricted to Central Taiwan.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
FB00E6380FD858F195AB27478DCFDD56.taxon	description	Figs 8 A-C, 9 A-C, 10, 11	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
FB00E6380FD858F195AB27478DCFDD56.taxon	materials_examined	Other material (n = 150). China. Fujian: 5 ♂, 8 ♀ (NMHUK), Foochow (福州), 1935 - 1938, leg. M. S. Yang; Taiwan. Nantou: 2 ♂, 1 ♀ (TARI), Lienhuachih (蓮華池), 28. IV. 2016, leg. A. Li; Kinmen: Kinmen Island (金門): 14 ♂, 7 ♀ (TARI), Tsaitso trail (蔡厝古道), 20. IV. 2021, leg. C. - F. Lee; 4 ♂, 1 ♀ (TARI), Taiwushan (太武山), 14. IV. 2011, leg. Y. - J. Chang; 1 ♀ (TARI), same but with " 6. V. 2011 "; 5 ♂, 2 ♀ (TARI), same but with " 19. IV. 2015 "; 2 ♂, 4 ♀ (TARI), same but with " 29 - 30. IV. 2015 "; 24 ♂, 23 ♀ (TARI), same locality, 8. IV. 2021, leg. C. - F. Lee; 23 ♂, 27 ♀ (TARI), same but with " 15. IV. 2021 "; Dadan Island (大膽島): 1 ♀ (TARI), 9. V. 2016, leg. Y. - J. Chang; Matsu Islands: 1 ♂, 2 ♀ (TARI), Beigan Island (北竿), 1. V. 2018, leg. H. - T. Fang; 1 ♂ (TARI), Nangan Island (南竿), 24. V. 2009, leg. U. Ong.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
FB00E6380FD858F195AB27478DCFDD56.taxon	diagnosis	Diagnosis. Gonioctena (Brachyphytodecta) scutellaris can be distinguished from the other consubgeneric species, G. (B.) liui sp. nov. by the following combination of the characters: yellowish brown head and pronotum (Figs 8 A, C, 9 A), thoracic and abdominal ventrites (Figs 8 B, 9 B) (black head and most parts of pronotum (Fig. 9 D), thoracic and abdominal ventrites (Fig. 9 E) in G. (B.) liui sp. nov.), extremely elongate apical process of aedeagus (Fig. 10 B, C) (short, wide apical process of aedeagus in G. (B.) liui sp. nov. (Fig. 12 B, C )), and wide gonocoxae covered with more setae (Fig. 10 D) (narrow gonoxae covered with fewer setae in G. (B.) liui sp. nov. (Fig. 12 D )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
FB00E6380FD858F195AB27478DCFDD56.taxon	biology_ecology	Biology. Gonioctena (Brachyphytodecta) scutellaris are presumed to be multivoltine during spring and females are oviparous. In Taiwan, this species seems rare due to unpredictable sprouting times of host plants. Mr. Alex Li (李志穎) collected some adults (Fig. 11 F) and mature larvae on April 28, 2016, at Lienhuachih (蓮華池). However, we could not find this species around the same season in the following years because host plants had not sprouted. For example, we could not find sprouts of the host plants in late May 2021 at that location. The populations in small islands are more stable, where host plants began to sprout, and overwintered female laid eggs during middle of March 2021 in Kinmen Islands. They laid 12 eggs into two rows on very young leaves (Fig. 11 A). Larvae hatched in three-four days. Early instar larvae (Fig. 11 B) fed gregariously on young leaves but became solitary as they matured. The larval duration was nine days. Mature larvae (Fig. 11 C) burrowed into soil and built underground chambers for pupation. Duration of the pupal stage (Fig. 11 D) was nine days. The newly emerged adults (Fig. 11 F) appeared during early April and females started to lay eggs after a week. In Jiangxi, China, females began to lay eggs first and then laid larvae in early April (Zhang and Shen 1984).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
FB00E6380FD858F195AB27478DCFDD56.taxon	distribution	Distribution. China, Taiwan, including Kinmen Islands (Kinmen Island 金門島, Dadan Island 大膽島) and Matsu Islands (Kaoteng Island 高登島, Beigan Island 北竿島, and Nangan Island 南竿島, new records).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
D314E487A90150F687301A39E19BFD37.taxon	description	Figs 17, 18, 19	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
D314E487A90150F687301A39E19BFD37.taxon	materials_examined	Other material (n = 95). Taiwan. Matsu islands: 17 ♂, 17 ♀ (TARI), Nangan (南竿), 7. VI. 2017, leg. T. - C. Chen; 23 ♂, 35 ♀ (TARI), same locality, 8. VI. 2017, leg. Y. - L. Liu; 1 ♀ (TARI), same locality, 28. VI. 2017, leg. H. - T. Fang; 1 ♂, 1 ♀ (TARI), Beigan (北竿), 18. IV. 2020, C. - H. Tsieh.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
D314E487A90150F687301A39E19BFD37.taxon	diagnosis	Diagnosis. Gonioctena (Sinomela) nigroplagiata is easily separated from the other consubgeneric species, G. (S.) osawai by the following combination of the characters: no black spots on the dorsum, none or three or four pairs of black spots on the elytra (Figs 17) (one pair of black spots on pronotum and five pairs of black spots on elytra in G. (S.) osawai (Fig. 14 G); wide antennomere V, width subequal to length (Fig. 18 A) (elongate antennomere V, more than 2.0 x longer than wide in G. (S.) osawai (Fig. 20 A )); narrowly rounded apex of endophallic sclerites (Fig. 18 B) (bifurcate and asymmetrical apices of endophallic sclerites in G. (S.) osawai (Fig. 20 B )); wide gonocoxae covered with more setae (Fig. 18 D) (narrow gonocoxae covered with fewer setae in G. (S.) osawai (Fig. 20 D )); membranous ventrite VII with dense setae along apical margin (Fig. 18 E) (slightly sclerotized ventrites VIII with dense setae only on apices in G. (S.) osawai (Fig. 20 E )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
D314E487A90150F687301A39E19BFD37.taxon	biology_ecology	Biology. A mass outbreak of adults of this species occurred on Matsu Islands during June 2017. The large numbers of beetles made local citizens nervous and made the news. A number of specimens were sent to the authors for identification. Mr. Hua-Te Fang (方華德) collected larvae (Fig. 19 B) on 15 April 2021 at the locality. Mature larvae burrowed into soil and built underground chambers for pupation (Fig. 19 C) on 22 April. The first adults emerged from soil on 2 May and were entirely yellowish brown (Fig. 19 D). Black spots appeared after three days in some individuals and become stable within five days (Fig. 19 E, F). No adults were observed in the field by Mr. Fang on 11 May, 29 September, and 21 October. This species is presumed to be univoltine, and larvae are active only during spring.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
D314E487A90150F687301A39E19BFD37.taxon	distribution	Distribution. China, Japan, Taiwan (only on Matsu Islands, including Beigan Island 北竿島and Nangan Island 南竿島).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	description	Figs 14 G-I, 20, 21	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	materials_examined	Type. Holotype (sex undetermined, KMNH): " Nr Liukuei [p] (六龜) 溪南山 [h] (Chinanshan) / Kaohsiung Hs. Taiwan [p] / 20 I [h] V 19 [p] 91 [h] / W. Chen leg (Osawa) [p, w] // PHOTO [p, r] // HOLOTYPE [p, r] // Gonioctena / Gonioctena osawai / Kimoto, n. sp [h] / Det. S. Kimoto, 19 [p, w] // 2001822 IR 02 [p, w] // KMNHIR 200,093 [p, w] ". This holotype was deposited originally at the Biohistory Research Hall, Takatsuki, Osaka (Kimoto 1996). Now it is transferred to the KMNH.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	materials_examined	Other material (n = 36). Taiwan. Kaohsiung: 1 ♂ (TARI), Hsiaokuanshan (小關山), 15. V. 2016, leg. B. - X. Guo; Pingtung: 2 ♀ (TARI), Chichia (七佳), 27. VII. 2018, leg. Y. - T. Chung; 1 ♂, 1 ♀ (TARI), same but with " 17. VII. 2019 "; 2 ♂, 2 ♀ (TARI), same locality, 18. VII. 2019, leg. B. - X. Guo; 1 ♀ (TARI), Lilungshan (里龍山), 4. VIII. 2009, leg. J. - C. Chen; 2 ♂ (TARI), same but with " 15. VIII. 2009 "; 1 ♂ (TARI), same locality and date, le. g S. - F. Yu; 2 ♂ (TARI), same but with " leg. M. - H. Tsou; 6 ♂, 2 ♀ (TARI), Laochichia (老七佳), 15. VII. 2021, leg. Y. - T. Chung; 5 ♂, 6 ♀ (TARI), same but with " 30. VIII. 2021 "; 1 ♂ (TARI), Tahanshan (大漢山), 30. V. 2012, leg. J. - C. Chen; 1 ♀ (TARI), same locality, 4. VI. 2016, leg. Y. - F. Hsu; 1 ♂ (TARI), same locality, 14. IV. 2020, leg. Y. - T. Chung;	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	diagnosis	Diagnosis. Gonioctena (Sinomela) osawai is easily separated from the other consubgeneric species, G. (S.) nigroplagiata by the following combination of the characters: one pair of black spots on pronotum and five pairs of black spots on the elytra (Fig. 14 G) (no black spots on the dorsum, or three or four pairs of black spots on the elytra in G. (S.) nigroplagiata (Fig. 17 )); elongate antennomere V, more than 2.0 x longer than wide (Fig. 20 A) (wide antennomere V, width subequal to length in G. (S.) nigroplagiata (Fig. 18 A )); bifurcate and asymmetric apices of endophallic sclerites (Fig. 20 B) (narrowly rounded apex of endophallic sclerites in G. (S.) nigroplagiata (Fig. 18 B )); narrow gonocoxae covered with fewer setae (Fig. 20 D) (wide gonocoxae covered with more setae n in G. (S.) nigroplagiata (Fig. 18 D )); slightly sclertozied ventrites VIII with dense setae only on apices (Fig. 20 E) (membranous ventrite VII with dense setae along apical margin in G. (S.) nigroplagiata (Fig. 18 E )).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	biology_ecology	Biology. Gonioctena (Sinomela) osawai populations are presumed to be univoltine during summer and females are ovoviviparous. Sprouting season of the host plant is during summer. Adults (Fig. 21 F) were found and brought into the laboratory on 15 July 2021. Females deposited larvae (Fig. 21 B) the following day. The larval duration (Fig. 21 B-D) was 12 days. Mature larvae (Fig. 21 D) burrowed into the soil and built underground chambers for pupation. The duration of the pupal stage (Fig. 21 E) was 13 days.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
48474EC347BA5A27A614B4252EC7244B.taxon	distribution	Distribution. South Taiwan (Kaohsiung and Pingtung counties).	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	description	Figs 14 A-F, 15, 16	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	materials_examined	Type. Holotype ♂ (TARI, original designation): " Hoorin [h] (= Fenglin, 鳳林) / FORMOSA [p] / 10. VIII. 1934 [h] / COL. [p] M. Kamikawa [h, w] // Phytodecta / Phytodecta kamikawai / Chujo [h] / DET. M. CHUJO [p, w] // Holo / Type [h, w; circle card with red letters and border but fade out] // 697 [p, w] ".	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	materials_examined	Other material (n = 77). Taiwan. Hsinchu: 1 ♀ (TARI), Lupi (魯壁), 12. III. 2009, leg. H. Lee; 1 ♂ (TARI), Mamei (馬美), 4. V. 2008, leg. S. - F. Yu; 1 ♀ (TARI), Wufeng (五峰), 17. III. 2009, leg. S. - F. Yu; Hualien: 1 ♂ (TARI), Jinma tunnel (金馬隧道), 5. V. 2014, leg. J. - F. Tsai; 1 ♂, 1 ♂ (TARI), Pilu (碧綠), 17. V. 2009, leg. M. - H. Tsou; 7 ♂, 2 ♀ (TARI), same locality, 1. VI. 2009, leg. C. - F. Lee; 2 ♀ (TARI), same locality, 22. VI. 2009, leg. U. Ong; Ilan: 1 ♀ (TARI), Fushan Botanical Park (福山植物園), 8. V. 2008, leg. M. - H. Tsou; 1 ♂ (TARI), Suchi trail (四季林道), 19. V. 2010, leg. H. - J. Chen; Kaohsiung: 2 ♂ (TARI), Chungchihkuan (中之關), 3. VII. 2009, leg. S. - F. Yu; 1 ♂ (TARI), Erhchituan (二集團), 1. IV. 2015, leg. B. - X. Guo; 2 ♂ (TARI), Tengchih (藤枝), 2 - 5. VI. 2008, leg. C. - F. Lee; 1 ♂ (TARI), same locality, 2. X. 2008, leg. M. - H. Tsou; 1 ♂ (TARI), same but with " 26. V. 2009 "; 1 ♂ (TARI), same locality, 19. III. 2013, leg. Y. - T. Chung; 1 ♂ (TARI), same locality, 8. II. 2014, leg. W. - C. Liao; 2 ♂ (TARI), same but with " 28. III. 2015 "; 1 ♂ (TARI), Tona trail (多納林道), 13. II. 2013, leg. B. - X. Guo; 3 ♂ (TARI), same locality, 3. II. 2013, leg. W. - C. Liao; Nantou: 1 ♂ (TARI), Chingching (清境), 6. IV. 2010, leg. Y. - T. Wang; 1 ♂ (TARI), Kuantaoshan (關刀山), 19. IV. 2014, leg. Y. - L. Lin; 1 ♂ (TARI), Tatachia (塔塔加), 9. VI. 2009, leg. C. - F. Lee; 2 ♂ (TARI), same but with " 27. IV. 2010 "; Pingtung: 2 ♀ (TARI), Tahanshan (大漢山), 8. V. 2009, leg. U. Ong; 1 ♂ (TARI), same but with " 1. VIII. 2009 "; 1 ♂, 1 ♀ (TARI), same locality, 24. IV. 2013, leg. Y. - T. Chung; 1 ♂ (TARI), same but with " 17. III. 2014 "; 1 ♂ (TARI), same but with " 10. VI. 2014 "; 1 ♂ (TARI), same but with " 24. II. 2015 "; 1 ♂ (TARI), same but with " 14. II. 2016 "; 1 ♀ (TARI), same but with " 9. IV. 2018 "; 2 ♂, 1 ♀ (TARI), same but with " 29. VI. 2018 "; Taichung: 1 ♂ (TARI), Piluhsi (畢祿溪), 1. VII. 2008, leg. M. - H. Tsou; 1 ♂ (TARI), Sungmao trail (松茂林道), 20. V. 2012, leg. T. - H. Lee; 1 ♂ (TARI), Wuling (武陵), 30. VI. 2008, leg. S. - F. Yu; Taipei: 1 ♀ (TARI), Manyuehyuan (滿月圓), 7. VI. 2010, leg. C. - L. Chiang; Taitung: 1 ♂ (TARI), Liyuan (栗園), 23. VI. 2010, leg. M. - H. Tsou; 1 ♂ (TARI), Hsiangyang (向陽), 28. III. 2014, leg. W. - C. Huang; Taoyuan: 2 ♂ (TARI), Hsuanyuan (萱源), 21. IV. 2010, leg. S. - F. Yu; 4 ♂ (TARI), Hsuehwunao (雪霧鬧), 2. IV. 2011, leg. M. - H. Tsou; 2 ♀ (TARI), Paling (巴陵), 29 - 30. IV. 2009, leg. M. - H. Tsou; 2 ♂, 1 ♀ (TARI), same but with " 23. V. 2009 "; 4 ♂ (TARI), same but with " 21. III. 2010 "; 1 ♀ (TARI), same but with " 28. III. 2010 "; 2 ♂ (TARI), same but with " 20. III. 2011 "; 1 ♂ (TARI), Tungyanshan (東眼山), 12. IV. 2007, leg. H. Lee; 1 ♀ (TARI), same but with " 2. V. 2009 ".	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	diagnosis	Diagnosis. Gonioctena (s. str.) kamikawai is the only member of the subgenus Gonioctena Gonioctena in the Taiwan fauna. Thus it is easily recognized by its subgeneric character -- trichobothria present on posterior angles of pronotum (trichobothria absent on anterior and posterior angles of pronotum in the subgenera Asiphytodecta and Brachyphytodecta; trichobothira present on anterior and posterior angles of the pronotum in the subgenus Asiphytodecta Sinomela). In addition, this species is characterized by its color patterns, shape of aedeagus, and apical processes of endophallic sclerites.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	biology_ecology	Biology. Females laid more than 280 eggs at a time (Fig. 16 A). Larvae hatched in 5 - 7 days. The early instar larvae (Fig. 16 B-D) gregariously fed on young leaves but became solitary as mature larvae. Mature larvae (Fig. 16 E) burrowed into the soil and built underground chambers for pupation. The duration of the pupal stage was ten days. Newly emerged adults (Fig. 16 F) appeared after May and went into a resting stage after feeding. Three hibernating adults were found during May 2011 at Wulai (烏來), North Taiwan. They became active and fed on leaves on 15 January 2012. The following observations suggest that populations of Gonioctena (s. str.) kamikawai are multivoltine. Many larvae were found on 21 March 2010 at Paling (巴陵), North Taiwan; females were observed laying eggs on 10 May 2010, at Peitawushan (北大武山), South Taiwan. These adults were likely the first full generation of the season.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
08A7B1F5C91756E2B10637414A88ED3E.taxon	distribution	Distribution. Endemic to Taiwan. This species is widespread in lowlands and mid-elevations.	en	Lee, Chi-Feng, Hsieh, Chia-Hung (2022): Integrative taxonomy of the leaf-beetle genus Gonioctena Chevrolat, 1836 in Taiwan (Coleoptera, Chrysomelidae, Chrysomelinae, Gonioctenini) reveals new synonymies and one new species. ZooKeys 1120: 1-46, DOI: http://dx.doi.org/10.3897/zookeys.1120.87526, URL: http://dx.doi.org/10.3897/zookeys.1120.87526
