taxonID	type	description	language	source
65633A03FF9C7034D4A2FD21BE5B0115.taxon	materials_examined	Type species: Lithosia gilveola Ochsenheimer, 1810 (a junior synonym of Phalaena palliatella Scopoli, 1763), by subsequent designation by Moore (1878).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9C7034D4A2FD21BE5B0115.taxon	diagnosis	Diagnosis. The genus was characterised and compared with Eilema in detail by Dubatolov & Zolotuhin (2011). No autapomorphies can be recognised but in the male genitalia, the combination of the following features is characteristic of the genus. (1) The vinculum is strongly elongate with thin arms and a membranous intravincular area bearing one or two coremata. (2) The valva has a long and wide dorsal part with a thin costa lacking processes. (3) The sacculus is relatively narrow with an elongate distal process well separated from the dorsal part of the valva. (4) The phallus vesica bears robust cornuti (Volynkin & Saldaitis 2021; Volynkin 2021 a).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9C7037D4A2FB3EBE38011F.taxon	discussion	Note. The male genitalia of the species of the group have the genital capsule structure typical of Manulea sensu stricto (illustrated by Dubatolov & Zolotuhin (2011 )) but the phallus vesica of most species bears a peculiar robust hook-like basal cornutus well separating them from other known taxa in Manulea (illustrated by Dubatolov & Zolotuhin 2011; Volynkin & Saldaitis 2021; Volynkin 2021 a, 2021 b). Nevertheless, one of the species has the external appearance and the male genital capsule very similar to others but its phallus vesica lacks a basal cornutus but bears a robust terminal one. Thus, the presence of a subbasal cornutus cannot be an autapomorphic feature of this complex of species, and, based on the morphological features, the latter should still be considered as just a species group within the genus Manulea and not a separate genus. However, the subgeneric placement of the group is still debatable since its vesica structure, especially the shape and size of the cornuti, is remarkably different from Manulea sensu stricto. Nevertheless, the taxonomic value of the cornuti shape in the subtribe Lithosiina is still unclear therefore we tentatively place the species group in the nominate subgenus of Manulea. The taxonomic placement of the group can be clarified in the future by using molecular methods through a multi-gene phylogenetic analysis including numerous groups and genera of the subtribe Lithosiina.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9C7037D4A2FB3EBE38011F.taxon	diagnosis	Diagnosis. Members of the species-group are superficially reminiscent of the Manulea costipuncta (Leech, 1890) / Manulea hokopo Volynkin, 2021 species pair (illustrated by Volynkin 2021 b) due to the presence of a small black spot on forewing costal margin medially but differ in the markedly smaller size and the narrower forewing. The male genital capsule of the species group is distinguished from the M. costipuncta / M. hokopo species pair in the shorter vinculum with two short sack-like coremata laterally, the distally tapered valva with a slenderer distal saccular process, and the pyramidal juxta, which is flattened and shield-like in the similar congeners. The vesica of the species group bears a robust cornutus basally or distally whereas the vesicae of M. costipuncta and M. hokopo bear a robust flattened cornutus sitting on the tip of an elongate conical diverticulum. In the female genitalia of the species group, the ductus bursae is gelatinous and rugose whereas it is heavily sclerotised and dorso-ventrally flattened in M. costipuncta and M. hokopo, and the anterior section of the corpus bursae is membranous and bears one or two signa whereas it is gelatinous with a broad sclerotised plate but without signa in the congeners. Additionally, the appendix bursae is relatively small (in proportion to the corpus bursae) and conical in M. tienmushanica species group whereas it is broader and globular in the M. costipuncta / M. hokopo species pair.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9C7037D4A2FB3EBE38011F.taxon	description	Description. Adult. Forewing length 11.5 – 17.0 mm in males and 15.0 – 16.5 mm in females. Sexual dimorphism limited: female with somewhat less elongate forewing apex than in male. Antenna ciliate in both sexes with longer cilia in male. Body and forewing upperside colouration from orange to pale ochreous-grey. Forewing elongate with slightly subapically downcurved costal and subproximally concave anal margins. Forewing upperside pattern consisting of black edge of costal margin and small round or rectangular black spot at costal margin medially. Forewing underside with intense black suffusion subbasally and medially. Hindwing paler than forewing, uniform, with black suffusion in certain species. Male genitalia. Uncus elongate and slender, laterally flattened, distally downcurved and apically pointed. Tegumen with broad arms fused in posterior three quarters. Tuba analis with thin and weakly sclerotised scaphium. Vinculum longer than tegumen with thin but heavily sclerotised arms, rectangular with rounded corners and shallow medial depression, bearing two short sack-like coremata. Transtillae thin, fused with each other. Valva lobular, somewhat dilated medially, distally tapered and apically rounded, with medially convex dorsal margin. Medial ventral plate of costa narrowly triangular setose with short crest in certain species. Sacculus narrow (ca. 1 / 2 – 1 / 3 of base of valva width) with elongate and slender, apically pointed and upcurved distal process. Juxta weakly sclerotised, pyramidal. Phallus long and broad (in proportion to genital capsule size), tubular, distally dilated in certain species. Vesica with robust basal or terminal cornutus and elongate dorsal and ventral diverticula distally bearing one or two clusters of spinules apically. Female genitalia. Papilla analis broadly rectangular with rounded corners, setose. Apophyses elongate and thin. Ductus bursae anteriorly dilated, gelatinous and rugose. Corpus bursae globular or elliptical with anterior section bearing one or two round or elliptical serrulate signa. Posterior section of corpus bursae gelatinous and rugose or with ban-like serrulate sclerotised plates. In certain species, corpus bursae with large heavily sclerotised lateral protrusion. Appendix bursae broadly conical, positioned postero-laterally, heavily sclerotised or membranous.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9C7037D4A2FB3EBE38011F.taxon	distribution	Distribution. Southwest and East mainland China and the island of Taiwan.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9F7031D4A2FA2EB84F06E3.taxon	description	(Figs 1 – 3, 9 – 11)	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9F7031D4A2FA2EB84F06E3.taxon	materials_examined	Type material examined. Holotype (Fig. 1): male, “ West Tien-mu-shan | Prov. Chekiang | 15.4.1932. H. Höne ” / pink label “ Typus ” / red label “ Holotypus | Lithosia | tienmushanica | Daniel. ” (ZFMK). Paratypes: 1 male, West Tien-mu-shan, Prov. Chekiang, 15. IV. 1932. H. Höne [leg.] / Allotypus / Allotypus Lithosia tienmushanica Daniel, unique number: ZFMKLep 153380, gen. prep. by Huang (ZFMK); 2 males, the same data as holotype (ZFMK); 1 male, same locality and collector as previous but 7. IV. 1932 (ZFMK); 1 male, same locality and collector as previous but 29. IV. 1932 (ZFMK); 1 male, Hoengshan, Prov. Hunan, 19. IV. 1933, H. Höne / Paratypus Lithosia tienmushanica Daniel 1947 / Sammlung Daniel (ZFMK); 1 male, West Tien-mu-shan (1600 m), Pz. Chekiang, 4. IV. 1932, H. Höne / Paratypus Lithosia tienmushanica Daniel 1947 / Sammlung Daniel, gen. prep. No.: ZSM Arct. 2019 - 968 (prepared by Volynkin) (MWM / ZSM). Additional material examined. MAINLAND CHINA: 1 male, same locality and collector as holotype but 18. IV. 1932 (ZFMK); 1 male, same locality and collector as holotype but 21. IV. 1932 (ZFMK); 1 male, China, Wuy Shan, Jiangxi – Fujian border, 50 km SE of Yingtan, 27 ° 56 ' N, 117 ° 25 ' E, 1600 m, III. 2002, Sinyaev & local coll. leg., gen. prep. No.: ZSM Arct. 2019 - 969 (prepared by Volynkin) (MWM / ZSM); 1 male, W Sichuan, 10 km S of Tianquan, 880 m, N 30 ° 00.456 ' E 102 ° 45.277 ', 23. IV. 2015, Floriani & Saldaitis, gen. prep. No.: AV 6779 (prepared by Volynkin) (AFM). TAIWAN: 1 male, Taiwan, Prov. Ilan [Yilan County], Ming-Chyr F. R. A., 1250 m, 24 ° 38 ' 98 '' N, 121 ° 28 ' 52 '' E, 6. IV. 1997, L. Peregovits & A. Kun leg., gen. prep. No.: ZSM Arct. 2021 - 197 (prepared by Volynkin) (MWM / ZSM). Notes. (1) The specimen supposed by Daniel to be a female and designated by him as ‘ allotype’ is a male in fact. (2) The single Taiwanese specimen examined displays some differences in both the superficial appearance and the male genitalia structures from the continental specimens of M. tienmushanica and the Taiwanese population may represent a distinct taxon. However, currently it is impossible to make a taxonomic decision and describe it due to the lack of a representative material.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9F7031D4A2FA2EB84F06E3.taxon	diagnosis	Diagnosis. The forewing length is 11.5 – 13.0 mm in males, smaller than all the other congeners (see measures below). In the male genitalia, the length of the dorsal diverticulum of the phallus vesica and the size of its apical cluster of spinules are variable (Figs 9 – 11). Manulea tienmushanica is most superficially reminiscent of M. diaforetica sp. n., the detailed comparison is provided below in the diagnosis of the latter species. In the male genitalia, the narrow vesica with two tubular diverticula bearing apical clusters of spinules is diagnostic for the species. The female is unknown.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF9F7031D4A2FA2EB84F06E3.taxon	distribution	Distribution. East (Zhejiang, Hunan, and the Jiangxi / Fujian border) (Daniel 1954) and Southwest (Sichuan) mainland China and the island of Taiwan.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF997031D4A2FC25BE4E0204.taxon	description	(Figs 4, 12)	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF997031D4A2FC25BE4E0204.taxon	materials_examined	Type material examined. Photographs of the holotype: male, “ Ʀ 川ŔM山 [Sichuan, Qingchengshan] | 1000 m | 中 ḄḨṩỄ ” [Chinese Academy of Sciences] / “ 1979 - V- 18 | ẌÞŮ; Ẫ 平 ” [collector: Ping Gao] / “ Eilema nigripuncta | Fang | ṪŽŮ; ḦƤffi ” [determiner: Fang Cheng-lai] / red label “ Holotype ” / blue label with a unique number “ IOZ (E) 209216 ” / green label with gen. prep. No.: “ L. 146 ” / “ ♂ ” (IZCAS). Additional material examined. MAINLAND CHINA: 3 males, China, Sichuan, Quingchenghoushan Mts, 70 km NW Chengdu, 1400 m, 14 – 15. V. 2005, S., V. & M. Murzin leg., gen. prep. Nos.: ZSM Arct. 2021 - 192, ZSM Arct. 2021 - 193 (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF997031D4A2FC25BE4E0204.taxon	diagnosis	Diagnosis. The forewing length is 15.0 mm in males. The species (Fig. 4) differs clearly from M. tienmushanica (Figs 1 – 3) in the pale ochreous body and wings colouration with rusty-brown tegula, patagia, and, cluster of androconial scales in the male forewing cell medially (whereas the forewing is uniform orange and lacks the cluster of androconial scales in the congener). The male genital capsule of M. nigripuncta (Fig. 12) differs from M. tienmushanica (Figs 9 – 11) in the shorter valva with a markedly shorter distal saccular process (in proportion to the tegumen-vinculum complex length). The vesica of M. nigripuncta is characterised by the strongly dilated medial chamber, the ventral diverticulum lacking the cluster of spinules, and the dorsal diverticulum bearing a bunch-like cluster of spinules directed distally whereas the medial chamber of the vesica of M. tienmushanica is tubular, the dorsal diverticulum bears a cluster of numerous spinules, and the ventral diverticulum bears a broader cluster of spinules curved laterally. The detailed comparison with M. salweena sp. n. is provided below under the diagnosis of the latter species. The female is unknown.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF997031D4A2FC25BE4E0204.taxon	distribution	Distribution. Known only from Sichuan Province of mainland China (Fang 2000).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF987030D4A2FF60B88E00A3.taxon	description	(Figs 5, 6, 13, 15)	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF987030D4A2FF60B88E00A3.taxon	materials_examined	Type material. Holotype (Figs 5, 13): male, “ China, NW. Yunnan | Nu Jiang valley | 42 km S. from Fugong | H- 2100 m | N ° 32.29 '' | E 98 ° 55.25 '' | 18. v. 2018 | Butvila & Saldaitis leg. ” / “ Slide | AV 6778 ♂ | A. Volynkin ” (WIGJ). Paratypes: MAINLAND CHINA: 2 males, 1 female, same data as holotype, gen. prep. No.: AV 4777 (prepared by Volynkin) (AFM, ASV); 1 female, China, SW Yunnan, Nujiang valley, S from Gongshan, H- 2100 – 2400 m, N 27 ° 43.42 '', E 98 ° 45.15 '' E, 15 – 16. V. 2018, Butvila & Saldaitis leg., gen. prep. No.: AV 4778 (prepared by Volynkin) (ASV); 4 males, 7 females, China, NW Yunnan Prov., Nujiang Lisu and Dulong auton. pref., Fugong County, Lishadi (= Walo), 42 km N of Fugong, 1390 m, 12 – 16. V. 1999, 27.15 ° N 98.55 ° E, leg. / ex coll. Dr R. Brechlin, gen. prep. No.: ZSM Arct. 2021 - 191 (male) (prepared by Volynkin) (MWM / ZSM).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF987030D4A2FF60B88E00A3.taxon	diagnosis	Diagnosis. The forewing length is 15.5 – 17.0 mm in males and 16.0 – 16.5 mm in females. The new species (Figs 5, 6) is superficially reminiscent of M. nigripuncta (Fig. 4) but distinguished by the more elongate forewing, and the pale ochreous-grey body and wings colouration with contrast paler veins whereas the body and the forewing ground colour of the congener is pale ochreous with markedly less contrast paler veins. The male genital capsules of the two species are similar (Figs 12, 13), but the uncus of M. salweena sp. n. is thicker than in M. nigripuncta, the valva is slightly broader (the length to width ratio is 2.28: 1 vs. 2.43: 1 in the congener), and the distal saccular process is markedly longer (in proportion to the valva length). The phallus of M. salweena sp. n. is somewhat shorter than in M. nigripuncta. The vesica configuration of the new species is similar to that of M. nigripuncta but the medial chamber of the former species is markedly narrower and bearing a short diverticulum ventrally, the ventral diverticulum is narrower, and the dorsal diverticulum is markedly longer (in proportion to the phallus size). Since the females of M. nigripuncta and M. tienmushanica are unknown, the female copulatory organ of M. salweena sp. n. is compared only to M. diaforetica sp. n. (the comparison of male genitalia of the two species is provided below in the diagnosis of the latter species). The female genitalia of M. salweena sp. n. (Fig. 15) differ clearly from M. diaforetica sp. n. (Fig. 16) in the markedly broader, longer and more rugose ductus bursae in the former. The corpus bursae of the new species is globular with three band-like and serrulate sclerotised transverse plates posteriorly and a round serrulate signum bursae anteriorly. Unlike M. salweena sp. n., the corpus bursae of M. diaforetica sp. n. is elliptical with a slightly dilated, gelatinous and rugose posterior section, the medial section of the corpus bursae is somewhat constricted with a heavily sclerotised right side bearing a large, triangular and apically rounded lateral protrusion which is basally fused with the appendix bursae, and the anterior section of the corpus bursae bears two elliptical signa laterally. The appendix bursae of the new species is membranous, broadly conical and apically rounded whereas it is markedly shorter and narrower and heavily sclerotised in M. diaforetica sp. n. Additionally, the apophysis anterioris of M. salweena sp. n. is somewhat longer (in proportion to the ovipositor size) than in the congener.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF987030D4A2FF60B88E00A3.taxon	distribution	Distribution. Known only from Nujiang Valley in north-western Yunnan Province of mainland China.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF987030D4A2FF60B88E00A3.taxon	etymology	Etymology. The specific epithet originates from the Salween (= Nujiang) River, upper stream of which the type locality is in. The name is a noun in apposition.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF98703DD4A2F9E4BB3107E7.taxon	description	(Figs 7, 8, 14, 16)	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF98703DD4A2F9E4BB3107E7.taxon	materials_examined	Type material. Holotype (Figs 7, 14); male, “ China, Yunnan | 5 km N Hutiaoxia | 220 km N Dali | 2200 m, 23 – 27. IV. 1998 | ex coll. Dr. R. Brechlin ” / “ Slide | ZSM Arct. | 2021 - 195 ♂ | A. Volynkin ” (MWM / ZSM). Paratypes. MAINLAND CHINA: 16 males, 3 females with same data as in holotype, ZSM Arct. 2021 - 194 (male), ZSM Arct. 2021 - 196 (female) (prepared by Volynkin) (MWM / ZSM); 1 female, N Yunnan, North from Yongseng, 2050 m, N 26 ° 31.14 ' E 100 ° 44.52 ', 11. V. 2018, Butvila & Saldaitis leg., gen. prep. No.: AV 6780 (prepared by Volynkin) (AFM).	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF98703DD4A2F9E4BB3107E7.taxon	diagnosis	Diagnosis. The forewing length is 15.0 – 15.5 mm in males and 15.0 mm in females. Manulea diaforetica sp. n. (Figs 7, 8) is superficially reminiscent of M. tienmushanica (Figs 1 – 3) but differs in the larger size, the forewing with a more elongate apex and the more oblique tornus, and the smaller black costal spot. Additionally, the male of the new species is distinguished from M. tienmushanica by the presence of a cluster of androconial scales in the cell proximally, and the intense ochreous-yellow suffusion on the hindwing medially-anteriorly. The male genital capsule of M. diaforetica sp. n. (Fig. 14) is most similar to M. salweena sp. n. (Fig. 13) but differs in the narrower uncus, the longer valva (in proportion to the tegumen-uncus complex length), and the more elongate and upcurved distal saccular process. The phallus of the new species is longer than in M. salweena sp. n. and evenly tubular whereas it is distally dilated in the congener. The vesica of M. diaforetica sp. n. (Fig. 13) differs clearly from other known relatives (Figs 9 – 13) in the lack of the subbasal cornutus, and the presence of a robust, thorn-like terminal cornutus. The vesica configuration of the new species is fundamentally different from other species in the group and is characterised by the broad and downcurved subbasal diverticulum, and the main chamber having three short conical diverticula lacking clusters of spinules. The detailed comparison of the female genitalia of M. diaforetica sp. n. (Fig. 16) with M. salweena sp. n. (Fig. 15) is provided above in the diagnosis of the latter species.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF98703DD4A2F9E4BB3107E7.taxon	distribution	Distribution. Known only from the type locality in northern Yunnan Province of mainland China.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
65633A03FF98703DD4A2F9E4BB3107E7.taxon	etymology	Etymology. The specific epithet originates from the Greek ‘ διαφορετικός’ meaning ‘ different’ and refers to the phallus and vesica structures significantly different from other species in the species group. The name is a noun in apposition.	en	Volynkin, Anton V., Saldaitis, Aidas, Huang, Si-Yao (2022): Taxonomic review of Manulea (M.) tienmushanica (Daniel), comb. nov., and its allies with descriptions of two new species from mainland China (Lepidoptera Erebidae: Arctiinae). Zootaxa 5182 (4): 389-398, DOI: https://doi.org/10.11646/zootaxa.5182.4.5
