taxonID	type	description	language	source
03E18789FFFF0D09F4C4084E7FC4F951.taxon	description	(Figs 1 – 13)	en	Khaustov, Vladimir A., Döker, Ismail, Joharchi, Omid, Khaustov, Alexander A. (2022): Morphological ontogeny and complementary description of Neoseiulus subsolidus (Beglyarov) (Acari: Mesostigmata: Phytoseiidae). Zootaxa 5187 (1): 249-269, DOI: https://doi.org/10.11646/zootaxa.5187.1.14
03E18789FFFF0D09F4C4084E7FC4F951.taxon	description	FEMALE (n = 10) Figs 1 – 4, 13 (A – E, G) Dorsal idiosoma (Figs 1 A, 4 A – B, 13 A) 430 (410 – 452) long, 282 (272 – 303) wide at level of s 4. Dorsal shield rotund and dark brownish, well sclerotized, heavily reticulated from anterior edge to level of Z 4 seta and smooth or slightly reticulated posterior to Z 4. With 17 pairs of smooth setae except Z 5 weakly barbed with a blunt tip, and two interscutal setae r 3 and R 1 on soft cuticle, seven pairs of solenostomes (gd 1, gd 2, gd 4, gd 5, gd 6, gd 8, gd 9) and 15 pairs of lyrifissures (id 1, id 2, id 4, id 5, id 6, idm 1, idm 2, idm 3, idm 4, idm 5, idm 6, idl 1, idl 4, idx and is). Solenostomes gd 9 enlarged. Muscle attachment sites (sigillae) present from j 3 to Z 4 and clearly seen. Measurements of dorsal setae as follows: j 1 24 (22 – 27), j 3 52 (47 – 55), j 4 7 (6 – 8), j 5 7 (6 – 8), j 6 11 (10 – 13), J 2 20 (18 – 23), J 5 10 (9 – 10), z 2 30 (28 – 33), z 4 55 (50 – 60), z 5 8 (7 – 8), Z 1 60 (54 – 65), Z 4 95 (91 – 100), Z 5 74 (67 – 79), s 4 66 (63 – 72), S 2 68 (63 – 72), S 4 49 (44 – 54), S 5 38 (34 – 42), r 3 35 (31 – 38), R 1 22 (19 – 24). Venter (Figs 1 B, 4 C – D, 13 B – E). Sternal shield 56 (50 – 61) long from middle of anterior margin to posterior margin, 81 (76 – 85) wide (at the level of ST 2 – ST 2), heavily sclerotized and reticulated on lateral margins, with three pairs of setae (ST 1 – ST 3) and two pairs of lyrifissures (iv 1, iv 2). Setae ST 4 and lyrifissures iv 3 situated on ovalshaped and well sclerotized metasternal shields. Peritremes wide, with numerous chetoides, extending forward nearly to j 1 level. Genital shield heavily sclerotized, slightly reticulated, 81 (75 – 86) wide at level of setae ST 5 – ST 5. Pair of paragenital lyrifissures iv 5 present on soft cuticle. Four thick transverse sclerites present between posterior margin of genital and anterior margin of ventrianal shield. One pair of small sigillae (sgpa) near setae ZV 1. Ventrianal shield large, reticulated mostly in anterior part, 140 (131 – 151) long and 124 (114 – 136) wide at level of ZV 2, anterior margin straight. With three pairs of pre-anal setae (JV 1, JV 2, and ZV 2), one pair of para-anal setae (PA), unpaired postanal seta (PST), and a pair of small crescentic solenostomes (gv 3) posteromesad JV 2. Caudoventral area with four pairs of setae (ZV 1, ZV 3, JV 4, and JV 5) (in illustrated specimen with three setae on right side because abnormality) and six pairs of lyrifissures (five pairs of ivo, and a pair of ivp). Setae JV 5 35 (30 – 41) long. Two pairs of metapodal shield present on soft cuticle, primary 48 (43 – 58) and secondary 11 (9 – 13) long, respectively. Spermatheca (Figure 2 E, 13 C – D). Calyx thick walled, narrow basally and gradually expanding distally, Vshaped 22 (19 – 24); atrium swollen, minor duct present, major duct broad. Gnathosoma (Figs 2 A – D). Subcapitulum well sclerotized, with three pairs of hypostomal setae h 1, h 2, h 3 and one pair of palpcoxal setae pc, deutosternal groove with seven transverse rows of denticles. Palp 135 (129 – 138) long (distance between base of trochanter to apex of tarsus): trochanter with two ventral setae v 1 and v 2; femur with one spatulate anterolateral seta al, three dorsal (d 1 – d 3) and one posterolateral pl; genu with two short and thick anterolateral setae al 1, al 2, three dorsal (d 1 – d 3) and one posterolateral pl; tibia with one short anterolateral seta, eight dorsal (d 1 – d 8), two distal di 1, di 2, two ventral v 1, v 2, and one posterolateral pl; tarsus with 15 setae and two-tined apotele. Epistome triangular and smooth. Chelicera: movable digit 26 (23 – 28) long, with three teeth, fixed digit 26 (24 – 28) long, with nine teeth and pilus dentilis. Legs (Figs 3, 13 G). Leg I 319 (300 – 330), leg II 278 (266 – 289), leg III 291 (255 – 309), leg IV 367 (344 – 389) in length. Leg IV with a short and blunt-tip macroseta on basitarsus St IV 27 (26 – 29) long. Leg chaetotaxy provided in Table 2. MALE (n = 7, Figs 5 – 6, 13 F). Dorsal idiosoma (Figs 5 A, 6 A – B) 341 (324 – 377) long, 246 (229 – 261) wide at level of s 4. Dorsal shield rotund and dark brownish, well sclerotized, heavily reticulated from anterior edge to level of Z 4 seta and smooth or slightly reticulated posterior to Z 4. With 19 pairs of smooth setae except Z 5 barbed, seven pairs of solenostomes (gd 1, gd 2, gd 4, gd 5, gd 6, gd 8, gd 9) and 13 visible lyrifissures (id 1, id 2, id 4, id 5, idm 1, idm 2, idm 3, idm 4, idm 5, idm 6, idl 1, idl 3, idl 4). Solenostomes gd 9 enlarged as in female. Muscle attachment sites (sigillae) present from j 1 to S 4 and clearly seen. Measurements of dorsal setae as follows: j 1 18 (16 – 20), j 3 49 (40 – 55), j 4 7 (6 – 7), j 5 7 (6 – 8), j 6 10 (10 – 11), J 2 20 (17 – 23), J 5 8 (7 – 9), z 2 29 (26 – 32), z 4 50 (45 – 54), z 5 8 (7 – 9), Z 1 56 (48 – 62), Z 4 80 (72 – 87), Z 5 63 (59 – 67), s 4 61 (54 – 70), S 2 58 (53 – 61), S 4 44 (41 – 48), S 5 33 (27 – 40), r 3 31 (28 – 35), R 1 22 (19 – 25). Peritremes extending forward nearly to j 1 level. Venter (Figs 5 B, 6 C, 13 F). Sternogenital shield 121 (116 – 125) long from middle of anterior margin to posterior margin, 70 (66 – 74) wide at level of ST 2 – ST 2, lightly reticulated, with five pairs of smooth setae (ST 1 – ST 5) and three pairs of lyrifissures (iv 1, iv 2, iv 3). Ventrianal shield 144 (136 – 154) long, 167 (161 – 170) wide at level of anterior corners, well triangular, reticulated from anterior margin to level of JV 2, three specimens with five pairs of pre-anal setae (JV 1, JV 2, ZV 1, ZV 2, ZV 3); one specimen with five setae on left side and four setae on right side (JV 1 absent); two with five setae on right side and four on left side (ZV 3 absent); one with five setae on left side and three on right side (ZV 1 and ZV 3 absent), one pair of para-anal setae (PA), unpaired post-anal seta (PST), a pair of crescentic solenostomes (gv 3) posteromesad JV 2 and four pairs of lyrifissures (ivo, ivo, ivo, iv 5). A pair of JV 5 29 (26 – 32) and a pair of lyrifissures ivp on soft cuticle. Gnathosoma (Figs 5 C, 6 D). Chelicera: movable digit 21 (19 – 22) long, with one tooth, fixed digit 21 (19 – 22) long, with six teeth and pilus dentilis. Spermatodactyl L-shaped with toe well developed. Legs. Leg I 286 (277 – 302), leg II 249 (242 – 258), leg III 264 (254 – 275), leg IV 336 (328 – 345) in length. Leg IV with short macroseta on basitarsus St IV 27 (23 – 28) long. Leg chaetotaxy as in female. LARVA (n = 1, Figs 7, 8, 13 H) Dorsal idiosoma (Fig. 7 A) 224 long, 156 wide at level of s 4. Podonotal shield weakly sclerotized, smooth, with nine pairs of setae (j 1, j 3, j 4, j 5, j 6, z 2, z 4, z 5 and s 4) and four pairs of lyrifissures (id 2, id 4, id 5, id 6), solenostomes absent. Setae j 3, j 6, z 4 and s 4 knobbed apically, remaining setae are pointed apically and setiform. Opisthonotal region with six pairs of setae Z 1, Z 4, Z 5, S 2, S 4, S 5 and one pair of “ setae nubs ” in position of setae J 5 (S 5, Z 5 and “ setae nubs ” situated on ventral side), with six pairs of lyrifussures (idm 1, idm 2, idm 3, idm 4 (situated on ventral side), idx, idl 1). Setae Z 4 apically knobbed and much longer than other dorsal setae. Measurements of dorsal setae as follows: j 1 26, j 3 37, j 4 7, j 5 6, j 6 57, z 2 26, z 4 49, z 5 13, Z 1 34, Z 4 113, Z 5 22, s 4 61, S 2 38, S 4 15, S 5 19. Venter (Figs 7 B, 13 H) without discernible shields. Coxisternal area with three pairs of setae ST 1 – ST 3 and without lyrifissures. Ventrianal area with four pairs of opisthogastric setae JV 1, JV 2, JV 5, ZV 2, one pair of paraanal setae (PA), unpaired post-anal seta (PST), and a pair of small rounded pre-anal solenostomes gv 3 and a pair of lyrifissures ivp. Measurements of ventral setae as follows: ST 1 23, ST 2 24, ST 3 22, JV 1 15, JV 2 19, JV 5 10, ZV 2 11, PA 16 and PST 13. Gnathosoma (Figs 7 C – E). Subcapitulum weakly sclerotized, with two pairs of hypostomal setae h 1 and h 2, deutosternal groove obscure, without discernible denticles. Palp 103 long (distance between base of trochanter to apex of tarsus): trochanter without setae; femur with one spatulate anterolateral seta al, two dorsal d 1, d 2 and one posterolateral pl; genu with one short and thick anterolateral seta al, three dorsal d 1, d 2, d 3 and one posterolateral pl; tibia with one short anterolateral seta al, six dorsal (d 1 – d 6), two distal di 1, di 2, two ventral v 1, v 2 and one posterolateral pl; tarsus with two dorsal setae d 1, d 2, six distal (di 1, di 3, di 5, di 6, di 7, di 8), three ventral (v 1 – v 3) and two-tined apotele. Chelicera: movable digit 17 long, with one tooth, fixed digit 17 long, with five teeth and short pilus dentilis. Legs (Fig. 8). Leg I 250, leg II 204 and leg III 222 in length. All legs without macrosetae. Leg chaetotaxy (except tarsus I) provided in Table 2. PROTONYMPH (n = 2, Figs 9 – 10) Dorsal idiosoma (Fig. 9 A) 253 - 263 long, 154 – 167 wide at level of s 4. Podonotal shield slightly reticulated mostly in anterior and lateral parts, muscle attachment sites (sigillae) present and clearly seen, with nine pairs of setae, four pairs of solenostomes (gd 1, gd 2, gd 4 (in illustrated specimen visible only on left side), gd 6) and two pairs of lyrifissures (id 2, id 5). Podonotal and pygidial shields separated by interscutal membrane, which bears three pairs of setae (J 2, Z 1 and S 2), six pairs of mesonotal sclerites (biggest with seta J 2), five pairs of lyrifissures (idm 2, idm 3, is 1, idl 1, and idl 3); setae r 3 and R 1 situated on soft cuticle. Pygidial shield well reticulated, with five pairs of setae (J 5, Z 4, Z 5, S 4, S 5), two pairs of solenostomes (gd 8, gd 9) and two pairs of lyrifissures (idx, idl 4), setae Z 4 and Z 5 barbed. Measurements of dorsal setae as follows: j 1 22 – 24, j 3 42 – 45, j 4 7, j 5 6 – 7, j 6 23 – 27, J 2 39 – 41, J 5 8 – 9, z 2 24 – 25, z 4 52, z 5 9 – 10, Z 1 40 – 48, Z 4 69, Z 5 47 – 49, s 4 59 – 63, S 2 50 – 52, S 4 27 – 31, S 5 22 – 26, r 3 31 – 32, R 1 19 – 20. Venter (Fig. 9 B). Coxisternal region without discernible shields, with three pairs of setae (ST 1 – ST 3) and two pairs of lyrifissures (iv 1, iv 2). Ventrianal region with four pairs of setae (JV 1, JV 2, JV 5, ZV 2), one pair of para-anal setae (PA) and unpaired post-anal seta (PST), with one pair of pre-anal circular solenostomes gv 3, posterior to JV 2 and five pairs of lyrifissures (iv 5, ivo, ivo, ivo, ivp) on soft cuticle surrounding anal shield. Measurements of ventral setae as follows: ST 1 20 – 21, ST 2 19 – 21, ST 3 19 – 22, JV 1 16, JV 2 15 – 17, JV 5 23 – 25, ZV 2 15, PA 14, PST 15 – 16. Peritremes short, slightly beyond the level of seta z 5. Gnathosoma (Figs. 9 C – E). Subcapitulum weakly sclerotized, with three pairs of hypostomal h 1, h 2, h 3 and one pair of palpcoxal setae pc, deutosternal groove with eight transverse rows of denticles, each row with two or three teeth. Palp 122 – 114 long (distance between base of trochanter to apex of tarsus): trochanter with one ventral seta v; femur, genu and tibia as in larva; tarsus with three dorsal (d 1 – d 3), nine distal (di 1 – di 9), three ventral (v 1 – v 3) and two-tined apotele. Chelicera: movable digit 19 long, with two teeth, fixed digit 19 long, with six teeth and pilus dentilis. Legs (Fig. 10). Leg I 251 – 257, Leg II 219 – 222, Leg III 223 – 227 and Leg IV 265 – 269. Latter with a macroseta, ST IV 43 – 44. Leg chaetotaxy (except tarsus I) provided in Table 2. DEUTONYMPH FEMALE (n = 4, Figs 11 – 12, 13 I) Dorsal idiosoma (Figs 11 A, 13 I) 310 (296 – 325) long, 178 (170 – 187) wide at level of s 4. Podonotal and opisthonotal shields fused at the level of R 1 creating schizodorsal shield. Latter reticulated, with 17 pairs of smooth setae except Z 4 and Z 5 barbed; seven pairs of solenostomes (gd 1, gd 2, gd 4, gd 5, gd 6, gd 8, gd 9) and 14 pairs of lyrifissures (id 2, id 4, id 5, id 6, idm 1, idm 2, idm 3, idm 5, idm 6, idl 1, idl 3, idl 4, idx and is), setae r 3 and R 1 on interscutal membrane. Muscle attachment sites (sigillae) present and clearly seen. Measurements of dorsal setae as follows: j 1 24 (24 – 25), j 3 46 (45 – 50), j 4 8 (7 – 9), j 5 7 (7 – 8), j 6 15 (13 – 17), J 2 30 (27 – 35), J 5 7 (6 – 9), z 2 27 (25 – 29), z 4 51 (47 – 55), z 5 8 (8 – 9), Z 1 53 (51 – 57), Z 4 71 (55 – 80), Z 5 53 (47 – 60), s 4 68 (65 – 71), S 2 58 (55 – 63), S 4 35 (32 – 41), S 5 29 (27 – 34), r 3 36 (34 – 39), R 1 22 (20 – 23). Venter (Figure 11 B). Coxisternal area with four pairs of setae, ST 1 – ST 4 and three pairs of lyrifissures (iv 1, iv 2, iv 3) situated on weakly sclerotized sternogenital shield. Length of sternogenital shield (distance between setae ST 1 – ST 5) 138 (136 – 141), width (distance between setae ST 2 – ST 2) 69 (67 – 70). Genital area with pair of setae ST 5 and a pair of para-genital poroid iv 5 on soft cuticle. Ventrianal region with seven pairs of setae (JV 1, JV 2, JV 4, JV 5, ZV 1, ZV 2, ZV 3), one pair of para-anal setae (PA) and unpaired post-anal seta (PST), with one pair of pre-anal circular solenostomes gv 3 (situated on small weakly sclerotized platelets), posterior to JV 2 and three visible pairs of lyrifissures (ivo, ivo, ivp) on soft cuticle surrounding anal shield. Opisthogastric area with one pair of metapodal plate. Measurements of ventral setae as follows: ST 1 26 (25 – 28), ST 2 25 (24 – 26), ST 3 22 (22 – 23), ST 4 21 (20 – 22), ST 5 22, JV 1 15 (15 – 16), JV 2 18 (18 – 19), JV 4 12, JV 5 30 (28 – 33), ZV 1 14 (13 – 14), ZV 2 16 (16 – 17), ZV 3 11 (10 – 12), PA 14 (13 – 14), PST 14 (13 – 15). Peritremes extending beyond level of j 3 setae. Gnathosoma (Fig. 11 C – E). Subcapitulum slightly reticulated, with three pairs of hypostomal h 1, h 2, h 3 and one pair of palpcoxal setae pc, deutosternal groove with seven transverse rows of denticles. Palp 122 (121 – 123) long (distance between base of trochanter to apex of tarsus): chaetotaxy as in female. Chelicera: movable digit 20 (20 - 21) long, with three teeth, fixed digit 20 (19 – 21) long, with eight teeth and pilus dentilis. Legs (Figure 12). Leg I 280 (266 – 287), Leg II 254 (251 – 261), Leg III 268 (258 – 273), Leg IV 330 (325 – 335) long. Latter with a blunt tip macroseta STIV 39 (37 – 41) long. Leg chaetotaxy (except tarsus I) provided in Table 2.	en	Khaustov, Vladimir A., Döker, Ismail, Joharchi, Omid, Khaustov, Alexander A. (2022): Morphological ontogeny and complementary description of Neoseiulus subsolidus (Beglyarov) (Acari: Mesostigmata: Phytoseiidae). Zootaxa 5187 (1): 249-269, DOI: https://doi.org/10.11646/zootaxa.5187.1.14
03E18789FFFF0D09F4C4084E7FC4F951.taxon	materials_examined	Specimens examined: 4 females and 2 males from Salix sp. in northern part of Tyumen region, Russia, 11 August 2019; 4 females from Salix sp. in north part of Tyumen region, Russia, 8 August 2019; 3 females, 1 male, 1 larva, 2 protonymph and 4 deutonymph from Salix sp. in Altai republic, Russia, 50 ° 13 ’ N, 87 ° 48 ’ E, 1472 m, 30 July 2020; 2 males from Salix sp. in Chelyabinsk region, Tyulyuk village, Russia, 54 ° 36 ′ N, 58 ° 46 ′ E, 18 August 2020; 5 females and 3 males from Salix sp. in Altai republic, Russia, 50 ° 32 ’ N, 87 ° 45 ’, 1546 m, 19 July 2021. All material collected by V. A. Khaustov. Geographical distribution: Belarus, Estonia, Finland, Latvia, Lithuania, Russia, Ukraine (Beglyarov 1981; Kuznetsov & Petrov 1984; Kolodochka 2006; Demite et al. 2022).	en	Khaustov, Vladimir A., Döker, Ismail, Joharchi, Omid, Khaustov, Alexander A. (2022): Morphological ontogeny and complementary description of Neoseiulus subsolidus (Beglyarov) (Acari: Mesostigmata: Phytoseiidae). Zootaxa 5187 (1): 249-269, DOI: https://doi.org/10.11646/zootaxa.5187.1.14
