identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
DA1C381AFFA41953B7B2ECE851BBCD9F.text	DA1C381AFFA41953B7B2ECE851BBCD9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Piribelba rossica (Bulanova-Zachvatkina 1957)	<div><p>Piribelba rossica (Bulanova-Zachvatkina, 1957)</p> <p>Belba rossica Bulanova-Zachvatkina, 1957: 1170 (“ Belba rossica, sp. n. ”, with a single diagnostic character reported (the absence of proterosomal tubercles); an available name under ICZN Articles 11 and 13.1; types not designated, distribution not reported); Bulanova-Zachvatkina, 1962: 215, Figs 4.4, 4.5 (holotype not designated; distribution: European part of the [former] USSR, Siberia, Russian Far East); Bulanova-Zachvatkina, 1967: 19 (indication that the original type series was from the Moscow region).</p> <p>Piribelba rossica: Miko, 2021: 1583 (designated as type of Piribelba Miko, 2021).</p> <p>Adult (Figs 1–9)</p> <p>Measurements. Body length 690–790; length of notogaster 560–620; maximum notogastral width 380–420.</p> <p>Integument (Figs 1A–F, 3A–F, 4A–D, 5A, B, D–G, 6A–H, 7A–E, 8B, C, F, 9B–E, G, I, J). Reddish-brown to deep reddish brown in colour. Body surface microtuberculate, microtubercles most conspicuous on all enantiophyses, lateral part of podosoma and around leg acetabula. Surface of dorsal and ventral sides of body and leg segments with very thick reticulated cerotegument (Figs 1A–C, 3A–F, 4A–D, 5A, B, D–G, 6A–H, 7A–E, 9B–E, G, I, J). Notogaster of all specimens with exuvial scalps – in young specimens, it is often only thin nymphal skins, older specimens have a thick layer of dirt and debris (Fig. 8H); leg segments with loosely attached adherent debris.</p> <p>Prodorsum (Figs 1A, C, 2G, 4A–C, 5A, B, 6A, C, D, E, G, 8G–J, 9G, H). Rostrum slightly tapers in front. Propodolateral apophyses absent. Dorsosejugal areas with rounded short ridges in place of tubercles Ba (sejugal costulae, scos), in dorsal view partly over-covered by anterior edge of notogaster and sometimes difficult to observe, prodorsal tubercles absent. Parastigmatic apophyses Sa well developed, long, elongate triangular in shape and conspicuously pointed distally, Sp relatively small, rounded at tip. Anterobothridial ridges (abr) distinct. Costulae (cos) present, distally curved. Rostral setae (ro) of medium length, 73–88, thin and smooth. Lamellar setae (le) longer and distinctly thicker than rostral ones, 96–106, from dorsal view with rare dark small spines or scales on majority of its length (often they very rare and hardly visible) (Figs 8G–J). Interlamellar setae (in, 51–63) thick, blunt, very bumpy at the top. Exobothridial setae (ex, 40–55) thick, strongly bumpy, curved. Bothridial setae (bs, 222–305) slightly tuberous at the apex, with flagellate tip in young specimens, in old specimens often with a broken off top. Distinct fields of sigillae (muscle insertions) present on prodorsum, two anteriad – anteriomediad to bothridia and in, one in interbothridial area.</p> <p>Notogaster (Figs 1A, C; 2H, 4A–D, 5B, D, E, 6A–C, H, 8A–F, 9D, C, I–K). Notogaster irregularly oval, distinctly narrowed anteriorly. Broadest part of notogaster positioned slightly behind half of notogaster, anterior margin of notogaster covering dorsosejugal area in dorsal view. Notogastral setae c 1 (91–101) and c 2 (103–120) thin, hairy, slightly curved, directed forward; la, lm, lp short (62–99), noticeably curved; h 1, h 2, h 3 (53–67) thicker, strongly curved; p 1, p 2, p 3 shorter (43–50) than others, thick, curved, poorly visible in dorsal aspect. All notogastral setae slightly tuberous, with cerotegument. Setae la, lm, lp, h 1, h 2, h 3 often have one or two distal spines (Figs 2H, 8B, C, E). Distance c 1 – c 1 shorter than c 2 – c 2. A few microtubercles found between bases of setae c 1 and c 2. All lyrifissures (ia, im, ip, ih, ips) and opisthonotal gland openings distinct. Circumgastric row of muscle sigillae distinct, well visible; numerous muscle sigillae found anterior and posterior margins of notogaster.</p> <p>Gnathosoma (Figs 2D–F, 5C). Subcapitulum longer than wide (151–168 × 100–108). Subcapitular setae (a, m, h) similar in length (38–53), setiform, barbed. Adoral setae (13–15) setiform, thin, smooth. Palps (145–148) with setation 0-2-1-3-9(+ω); solenidion of palptarsi bacilliform, pressed to palptarsal surface mediodistally. Postpalpal setae (10–11) spiniform. Chelicerae (135–157) of shape and relative size as usual in Damaeidae, seta cha (42–45) with extended proximal part, bent outwards proximally and inwards distally, with short barbs on external curvature and attenuated tip, seta chb (30–33) straight, distally bent and with fringe of diminishing barbs. Trägårdh’s organ (32–35) elongate triangular.</p> <p>Epimeral and lateral podosomal regions (Figs 1B, C, 2A, B, 5A, B, 6C, D, 9B). Ventral tubercles absent. Epimeral setal formula variable: 3-1-3-4 or 3-1-4-4, or 3-1-3-5; left and right sides can differ in their formulas. Setae 1a, 1b, 1c, 2a, 3a, 3b, 3c, 3d, 4b situated on distinctly developed microtubercles. Epimeral setae comparatively long, smooth, strong, with attenuate tips, setae 4a, 4c, 4d slightly thinner than other epimeral setae. Epimeres with muscular sigillae, well visible, particularly closer to axial part. Discidia (dis) poorly developed, rounded at tip and projecting slightly laterally, poorly distinguishable in the lateral aspect.</p> <p>Anogenital region (Figs 1B–F, 5A, 6B, C, 9B). Six pairs of genital (g 1 – g 6, 30–54), one pair of aggenital (ag, 66– 70), two pairs of anal (an 1, an 2, 34–38), and three pairs of adanal (ad 1 – ad 3, 48–58) setae. Aggenital and adanal setae smooth, strong, subequal in length, similar to epimeral setae. Distance ad 3 – ad 3 longer than ad 2 – ad 2. Posteromedian part of genital plates with a few microtubercles; insertions of genital setae may be connected to one another by sclerotized longitudinal ridges; anal plates with a sclerotized longitudinal ridge on each. Genital setae g 1 distinctly longer (48–54) than others and nearly as long as adanal setae; anal setae similar in length to g 2 – g 6. Adanal lyrifissures (iad) apoanal – oblique and divergent from body axis posteriad. Anal lyrifissures (ian) present (Fig. 1F).</p> <p>Legs (Figs 3A–F, 5G, 7A–E, 9E). All legs moderately long, segments with distinctly swollen distal parts (moniliform), except tarsi, where swollen part is proximal. Legs IV slightly longer than body, legs I and III slightly shorter than body, legs II distinctly shorter than body (Table 1). All setae of trochanters, femora, genua, and tibiae (except v), and ft of tarsi strong, darkly pigmented, and mostly with very short barbs. Ventral setae of tibiae and rest setae of tarsi comparatively long and thin, pointed, smooth or slightly barbed. Formulas of leg setation and solenidia: I (1-7-4-4-20) [1-2-2], II (1-7-4-5-17) [1-1-2], III (2-5-3-4-17) [1–1–0], IV (2-5-3-4-14) [0-1-0]; homologies of setae and solenidia indicated in Table 2. Famulus of tarsi I normal, setiform, emergent, relatively long. Solenidia of genua I, II and III each coupled with companion setae d, as well as solenidia of tibiae II–IV. Length of solenidia on genua I–III and on tibiae II, III subequal to length of companion setae, tibial solenidia IV longer than respective companion setae.</p> <p>Larva (Figs 10, 11, 18A–C)</p> <p>Measurements. Body length 275–300; body width 150–170.</p> <p>Integument (Figs 10). Body oval, rounded posteriorly, weakly sclerotized. Body cuticle white-coloured. Cuticle of body and legs smooth, covered with thin layer of granular cerotegument. Round or oval granules small, up to 1–1.4 μm. Prodorsal and gastronotic setae (excluding ro, le, bs) without cerotegument.</p> <p>Prodorsum (Figs 10A, C) triangular, rostrum rounded. Setae strongly or weakly dark-coloured, inserted on small apophyses. Rostral (ro, 25–28) and lamellar (le, 28–30) setae strong, tapering at the end, barbed. Interlamellar setae (in, 18–20) thick, sharply pointed at the top, slightly barbed. Exobothridial setae (ex, 4–5) minute, smooth. Bothridial setae (bs, 166–180) flagellate, smooth.</p> <p>Gastronotum (Figs 10). Twelve pairs of gastronotic setae. Setae dark-coloured (except c 1 and c 3), inserted on apophyses (except c 3 and h 3). Setae c 1 (19–24) thin, light-coloured, smooth, inserted on separate small apophyses. Setae la, lm (19–25), h 1 (23–27), h 2 (22–24), da (37–40), dm (60–67) barbed, flagellate. Setae dp (180–195) longest, barbed, with long, thin tips. Setae lp (6–7) short, smooth, blunt. Setae c 3 and h 3 minute (3–4). Apophyses of setae dp located close to each other, connected. Cupules ia after to setae c 3, im ventral to setae lm, ih close to anal valves, ip between setae h 2 and h 3. Small apertures of opisthonotal glands (gla) visible lateral to setae h 3.</p> <p>Epimeral region (Figs 10B, C). Epimeral formula: 2-1-2 (except protective scale over Claparède’s organ). Epimeral setae short, smooth. Apodemes ap2, apsj porose.</p> <p>Gnathosoma (Figs 18A–C). Subcapitulum slightly longer than wide (70–75 × 66–71). Subcapitular setae (a, m, h, 22–25) setiform, barbed. Adoral setae (5–6) setiform, thin, smooth. Palps (77–84) with setation 0-1-1-3-9(+ω); seta v ʺ absent. All setae barbed (except on palptarsi). Solenidion of palptarsi bacilliform, pressed to palptarsal surface. Postpalpal setae (5) spiniform, smooth. Chelicerae (85–89) with two long, barbed setae (cha, 20–23; chb, 18–20), chb with long barbs in middistal part.</p> <p>Legs (Figs 11). All legs shorter than body (Table 1). Tarsi and femora elongated, the longest among segments. Claws smooth. Most setae short, thick, heavily barbed, setae of tarsi setiform (except ft). Formulas of leg setation and solenidia: I (0-2-3-4-16) [1-1-1], II (0-2-3-3-13) [1-1-1], III (0-2-2-3-13) [1-1-0]; homologies of setae and solenidia indicated in Table 2. Famulus of tarsi I normal, setiform, emergent, relatively long. Solenidia of genua I–III each coupled with companion setae d, as well as solenidia of tibiae I–III. Solenidia σ pressed to the surface of genua, setae d on genua minute. Setae d about one eighth length of φ 1 on tibiae I. Setae d on tibiae II, III strong, barbed, subequal to length of companion solenidia.</p> <p>Protonymph (Figs 12, 13, 18D–F)</p> <p>Measurements. Body length 330–364; body width 192–200.</p> <p>Integument (Figs 12). Body oval, rounded posteriorly, weakly sclerotized. Body cuticle white-coloured. Cuticle of body and legs smooth, covered with thin layer of granular cerotegument. Round or oval granules small, up to 2–2.5 μm. Prodorsal and gastronotic setae (excluding ro, le, bs) without cerotegument.</p> <p>Prodorsum (Figs 12A, C) triangular, rostrum rounded. Curved cuticle thickenings are located laterally on the prodorsum, distal to the bothridia. Setae strongly or weakly dark-coloured, inserted on small apophyses. Rostral (34–37) and lamellar (38–42) setae strong, tapering at the end, barbed. Interlamellar setae (17–22) thick, sharply pointed at the top, smooth. Exobothridial setae (10) smooth. Bothridial setae (180–210) flagellate, smooth.</p> <p>Gastronotum (Figs 10). Twelve pairs of gastronotic setae. Setae dark-coloured (except c 3), inserted on apophyses. Setae c 1 (50–54), c 2 (74–80), la (56–60), lm (67–71), lp (48–51), h 3 (68–70), h 2 (35–37) barbed, flagellate. Setae h 1 (194–203) longest, barbed, with long, thin tips. Setae p 1 (19–21) short, slightly barbed. Setae c 3 (7–8), p 2, p 3 (9–11) short, smooth. Apophyses of setae c 1 separated, setae h 1 located close to each other, connected. Cupules ia after to setae c 3, im ventral to setae lm, ips close to anal valves, ih located above setae p 3, ip near setae h 2 and h 3. Small apertures of opisthonotal glands visible lateral to setae p 2 and p 3. Genital valves appear, with one pair of setae. Gastronotic region covered with particles of dirt (not covered with larval exuvium). Cornicle (k) poorly developed.</p> <p>Epimeral region (Figs 12B, C). Epimeral formula: 3-1-2-1. Epimeral setae short, smooth. Apodemes ap2, apsj, ap3 porose.</p> <p>Gnathosoma (Figs 18D–F). Subcapitulum longer than wide (90–95 × 70–76). Subcapitular setae (a, m, 24–27; h, 15–17) setiform, barbed (setae h smooth). Adoral setae (8–9) setiform, thin, smooth. Palps (90–100) with setation 0-2-1-3-9(+ω). All setae barbed (except on palptarsi). Solenidion of palptarsi bacilliform, pressed to palptarsal surface. Postpalpal setae (6–7) spiniform, smooth. Chelicerae (104–108) with two long, barbed setae (cha, 29–31; chb, 23–25), chb with long barbs in middistal part.</p> <p>Legs (Figs 13). All legs shorter than body (Table 1). Tarsi and femora elongated, the longest among segments. Claws smooth. Most setae of legs short, thick, heavily barbed, setae of tarsi setiform (except ft). Formulas of leg setation and solenidia: I (0-2-3-4-16) [1-1-2], II (0-2-3-3-13) [1-1-1], III (1-2-2-3-13) [1-1-0], IV (0-0-0-0-7) [0-0- 0]; homologies of setae and solenidia indicated in Table 2. Famulus of tarsi I normal, setiform, emergent, relatively long. Solenidia of genua I–III each coupled with companion setae d, as well as solenidia of tibiae I–III. Setae d about more one tenth length of φ 1 on tibiae I, smooth. Setae d on tibiae II, III and on genua I–III strong, barbed, subequal to length of companion solenidia.</p> <p>Deutonymph (Figs 14, 15, 18G–I, 19F, G)</p> <p>Measurements. Body length 425–450; body width 250–270.</p> <p>Integument (Figs 14). Body oval, rounded posteriorly, weakly sclerotized. Body cuticle white-coloured. Cuticle of body and legs smooth, covered with thin layer of granular cerotegument. Round or oval granules small, up to 1.5–2.8 μm. Prodorsal and gastronotic setae (excluding ro, le, bs) without cerotegument.</p> <p>Prodorsum (Figs 14A, C) triangular, rostrum rounded. Curved cuticle thickenings are located laterally on the prodorsum, distal to the bothridia. Setae strongly or weakly dark-coloured, inserted on small apophyses. Rostral (38–41) and lamellar (39–44) setae strong, tapering at the end, barbed. Interlamellar setae (19–21) thick, sharply pointed at the top, smooth. Exobothridial setae (14–15) smooth. Bothridial setae (195–220) flagellate, smooth.</p> <p>Gastronotum (Figs 14). Twelve pairs of gastronotic setae. Setae dark-coloured (except c 3), inserted on apophyses. Setae c 1 (102–110), c 2 (140–150), la (68–73), lm (105–108), lp (90–93), h 3 (85–90), h 2 (64–67), p 1 (45–47) barbed, flagellate. Setae h 1 (230–247) longest, barbed, with long, thin tips. Setae c 3 (19–20), p 2, p 3 (14–17) short, smooth. Apophyses of setae c 1 separated, setae h 1 located close to each other, connected. Three pairs of genital (g, 12–14), one pair of aggenital (ag, 11–12) and three pairs of adanal (ad 1 – ad 3, 7–11) setae setiform. Some specimens show tiny anal setae. Cupules ia after to setae c 3, im ventral to setae lm, ih and ips located above setae p 3, ip near setae h 2 and p 2, iad close to anal valves. Small apertures of opisthonotal glands (gla) visible lateral to setae p 2 and p 3. Gastronotic region covered with protonymphal exuvium and particles of dirt. Cornicle (k) developed, conical, between setae la.</p> <p>Epimeral region (Figs 14B, C). Epimeral formula: 3-1-2-2. Epimeral setae short, smooth. Apodemes ap2, apsj, ap3 porose.</p> <p>Gnathosoma (Figs 18G–I). Subcapitulum longer than wide (120–130 × 105–110). Subcapitular setae (a, 37–41; m, 48–50; h, 21–24) setiform, barbed (setae h smooth). Adoral setae (9–11) setiform, thin, smooth. Palps (109–113) with setation 0-2-1-3-9(+ω). All setae barbed (except on palptarsi). Solenidion of palptarsi bacilliform, pressed to palptarsal surface. Postpalpal setae (10) spiniform, smooth. Chelicerae (120–124) with two long, barbed setae (cha, 34–37; chb, 25–27), chb with long barbs in middistal part.</p> <p>Legs (Figs 13, 19F, G). All legs shorter than body (Table 1). Tarsi and femora elongated, the longest among segments. Claws smooth. Most setae of legs short, thick, heavily barbed, setae of tarsi setiform (except ft). Formulas of leg setation and solenidia: I (1-4-4-5-16) [1-2-2], II (1-4-4-4-13) [1-1-2], III (2-3-3-4-13) [1-1-0], IV (1-2-2-3- 12) [0-1-0]; homologies of setae and solenidia indicated in Table 2. Famulus of tarsi I normal, setiform, emergent, relatively long. Solenidia of genua I–III each coupled with companion setae d, as well as solenidia of tibiae I–IV. Setae d about more one tenth length of φ 1 on tibiae I, smooth. Setae d on tibiae II, III and on genua I–III strong, barbed, subequal to length of companion solenidia. Solenidia φ on tibiae IV longer than respective companion setae.</p> <p>Tritonymph (Figs 16, 17, 18J–L, 19A–E)</p> <p>Measurements. Body length 650–720; body width 380–410.</p> <p>Integument (Figs 14). Body oval, rounded posteriorly, weakly sclerotized. Body cuticle white-coloured. Cuticle of body and legs partly smooth (gastronotum) and partly covered with fine pits (prodorsum). Body covered with thin layer of granular cerotegument. Round or oval cerotegument granules small, up to 1–3 μm. Prodorsal and gastronotic setae (excluding ro, le, bs) without cerotegument.</p> <p>Prodorsum (Figs 16A, C, 19E) triangular, rostrum rounded. Curved cuticle thickenings are located laterally on the prodorsum, distal to the bothridia. Setae strongly or weakly dark-coloured, inserted on small apophyses. Rostral (52–55) and lamellar (54–58) setae strong, tapering at the end, barbed. Interlamellar setae (28–31) thick, sharply pointed at the top, smooth. Exobothridial setae (26–28) smooth. Bothridial setae (250–270) flagellate, smooth.</p> <p>Gastronotum (Figs 16). Twelve pairs of gastronotic setae. Setae dark-coloured, inserted on apophyses. Setae c 1 (120–137), c 2 (138–160), la (120–125), lm (160–170), lp (182–193), h 3 (170–180), h 2 (100–112), p 1 (85–90) barbed, flagellate. Setae h 1 (280–290) longest, barbed, with long, thin tips. Setae c 3 (34–37), p 2, p 3 (17–20) short, smooth. Apophyses of setae c 1 separated, setae h 1 located close to each other, connected. Five pairs of genital (g, 15–17), one pair of aggenital (ag, 14–16), two pairs of anal (an, 13–15), and three pairs of adanal (ad 1 – ad 3, 18–22) and setae setiform. Cupules ia after to setae c 3, im ventral to setae lm, ih and ips located above setae p 3, ip near setae h 2 and p 2, iad close to anal valves. Small apertures of opisthonotal glands (gla) visible lateral to setae p 2 and p 3. Gastronotic region covered with protonymph and deutonymph exuvia and particles of dirt. Cornicle (k) developed, conical, between setae la.</p> <p>Epimeral region (Figs 16B, C). Epimeral formula: 3-1-3-3. Epimeral setae short, smooth. Apodemes ap2, apsj, ap3 porose (Figs 19A–D).</p> <p>Gnathosoma (Figs 18J–L). Subcapitulum longer than wide (148–158 × 130–138). Subcapitular setae (a, 44–50; m, 50–58; h, 36–40) setiform, barbed (setae h smooth). Adoral setae (12–16) setiform, thin, smooth. Palps (120– 130) with setation 0-2-1-3-9(+ω). All setae barbed (except on palptarsi). Solenidion of palptarsi bacilliform, pressed to palptarsal surface. Postpalpal setae (8–10) spiniform, smooth. Chelicerae (130–136) with two long, barbed setae (cha, 36–40; chb, 30–33), chb with long barbs in middistal part.</p> <p>Legs (Figs 13). All legs shorter than body (Table 1). Tarsi and femora elongated, the longest among segments. Claws smooth. Most setae of legs short, thick, heavily barbed, setae of tarsi setiform (except ft). Formulas of leg setation and solenidia: I (1-5-4-5-18) [1-2-2], II (1-5-4-5-15) [1-1-2], III (2-3-3-4-15) [1-1-0], IV (2-3-3-4-12) [0-1- 0]; homologies of setae and solenidia indicated in Table 2. Famulus of tarsi I normal, setiform, emergent, relatively long. Solenidia of genua I–III each coupled with companion setae d, as well as solenidia of tibiae I–IV. Setae d about more one sixth length of φ 1 on tibiae I, smooth. Setae d on tibiae II, III and on genua I–III strong, barbed, subequal to length of companion solenidia. Solenidia φ on tibiae IV longer than respective companion setae.</p></div> 	https://treatment.plazi.org/id/DA1C381AFFA41953B7B2ECE851BBCD9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Miko, Ladislav;Kolesnikov, Vasiliy B.;Ermilov, Sergey G.;Klimov, Pavel B.	Miko, Ladislav, Kolesnikov, Vasiliy B., Ermilov, Sergey G., Klimov, Pavel B. (2022): Taxonomy of European Damaeidae (Acari, Oribatida) XI. European species of the genus Piribelba Miko 2021: redescriptions of P. rossica (Bulanova-Zachvatkina 1957) and P. piriformis (Mihelčič, 1964) using morphology and DNA sequence data. Zootaxa 5187 (1): 169-210, DOI: https://doi.org/10.11646/zootaxa.5187.1.11
DA1C381AFFBF1962B7B2EDB95171CF87.text	DA1C381AFFBF1962B7B2EDB95171CF87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Piribelba piriformis (Mihelcic 1964)	<div><p>Piribelba piriformis (Mihelčič 1964)</p> <p>Belba piriformis Mihelčič 1964: 376, Figs. 10–12 (holotype not designated, distribution provided by author: Austria: Carinthia, East and South Tyrol).</p> <p>The species is very similar to P. rossica, but can be separated using morphology (see key to species below) and DNA sequence data: 12.0% uncorrected p-distances and 13.3% Kimura two-parameter distances (K2P). The text below, therefore focuses on measurements and diagnostic character states, without necessarily repeating all character states that are identical with P. rossica.</p> <p>Adult (Figs 21–24, 26–29)</p> <p>Measurements. Average total body length 630 (n=8: 605–665; type 730); ventral body length 585 (n=8: 571– 605; material of Mihelčič: n=8, 537–604; type 678); length of notogaster 425 (n=8: 406–440); maximum notogaster width 355 (n=8: 345–385), maximum prodorsum width 245 (n=8: 220–270). Specimens from Slovakia are slightly more robust, with roughly same average length of body having slightly broader notogaster (363).</p> <p>Integument (Figs 23, 24E, H, 29A). Dark reddish-brown, whole body surface under cerotegument densely microtuberculate. Adults carrying nymphal exuvial scalps covered by thick layer of organic and inorganic debris of characteristic shape (on top of notogaster, shifted forward and not prolonged posteriad, compared to Belba species, Fig. 29C). Most of the body covered by characteristic reticular cerotegument, including proximal segments of legs, lateral, sejugal, partly ventral parts of the body and more distal segments of legs covered by globular, columnar and amorphous cerotegument.</p> <p>Prodorsum (Figs 26A, E, 28A, D, 29A). Rostrum broadly rounded, propodolateral apophyses absent. Dorsosejugal areas with narrow, triangular to almost linear short ridges in place of tubercles Ba (sejugal costulae, scos), in dorsal view partly over-covered by anterior edge of notogaster and sometimes difficult to observe, in some specimens doubled, with additional second, smaller, parallel ridge present laterally, prodorsal tubercles absent. Distinct prodorsal grooves present. Anterobothridial ridges (abr) almost straight, short, distinct. Prodorsal costulae (cos) present on lateral prodorsum above legs I insertions, slightly curved, usually distinct but in some populations (Slovakia, Sivec) not well observable. Parastigmatic apophyses differently shaped, only Sa usually visible in dorsal view, relatively long, narrow, with conspicuously pointed tips, often curved slightly lateroanterad; Sp much smaller, tubercular. Rostral setae (ro) of medium length, 62–86, thin, smooth. Lamellar setae (le) longer and distinctly thicker than rostral ones, 92–106, inserted on small tubercles or cuticular thickenings, from dorsal view with more or less distinct, sometimes dark small spines or scales on majority of its length. In different aspect (mediodorsal view) they appear flattened, proximally broadened, with spines visible only partly at external curvature (Figs 21C, 23E, 24D, 26C, 28D, 29B). In Slovak population from Sivec, lamellar setae are flattened, broadened proximally, with more transparent lateral parts (Fig. 26E), with spines and scales more transparent, present on dorsal surface of setae and more difficult to observe, easily to be interchanged with covering layer of granular cerotegument. Interlamellar setae (in, 36–52) erect, straight, thick, bluntly pointed, in lateral view appearing slightly flattened (Figs 22C, 26D, 28C). Exobothridial setae (ex, 48–58), strong, regularly curved, attenuated at ends (Figs 21C, 26D). Bothridial setae (bs, 170–278) long, setiform, distally with flagellate part which may be present or broken off, proximal part may be covered with fine layer of microgranular cerotegument (Fig 26D, 29A, F).</p> <p>Notogaster (Figs 21A, B, 23A, 26A, E, 28A, 29A, F). Notogaster elongated, ovoid, anteriorly narrowed, in lateral view however almost semicircular. Anterior margin of notogaster covering dorsosejugal areas in dorsal view. Notogastral setae thin, setiform, curved, distally attenuated, covered by layer of cerotegument proximally. Posterior setae more strongly curved or even curly distally, sometimes with one or two small, but distinct spines on external side of curvature. Setae medium long, their length reaching or almost reaching distance to following setal insertions, setae c 1 inserted closer to each other than other setae, distance c 1 – c 1 about twice shorter than c 2 – c 2 and shorter than c 1 – c 2. Setae of rows l slightly shorter (54–84) than rows c (60–89), h (69–91) and posterior setae (row p shorter but not observable in dorsal view).</p> <p>Gnathosoma (Figs 26B, F, 28D, 29F). Similar to that of P. rossica, but with apparent U-shaped less-sclerotised area on mentum (Figs 26B, 28B). This character is less visible on specimens from Slovak population from Sivec (Fig 26F). Palps (Figs 27F, 28D) with standard setation, identical to P. rossica.</p> <p>Epimeral and lateral podosomal regions (Figs 21A, 23B–D, 26B, F, 28B, 29F). Tecta of podocephalic fossa with short but distinct posterolateral angles, slightly projecting as short triangular projections (Fig. 26B). Without ventral tubercles, discidia reduced (absent or almost absent). Epimeral setal formula variable: 3-1-3or4-3or4; most of setae inserted on small tubercles or cuticular thickenings, setae rather strong, setiform or slightly flattened, attenuated towards tip (Fig. 21A, D). In Slovak population from Sivec, epimeral setae more distinctly flattened, with transparent margins (Figs 26F, H, 29F).</p> <p>Anogenital region (Figs 1B–F, 5A, 6B, C, 9B). Anal aperture slightly narrower than genital aperture. Standard set of anogenital setae present, aggenital and adanal setae quite strongly developed, setiform, attenuated towards tip (Fig. 21D). Genital and anal setae thinner and slightly shorter, smooth, g 1 usually significantly longer than other genital setae (Figs 21A, 22C, 28J), in Slovak population from Sivec length of genital setae subequal (Figs 26F, G, 29F).</p> <p>Legs (Figs 22A, B, E, 23F, 24B–G, 27, 28F–I, D–G). All legs generally of the same form and size as in P. rossica (for lengths of leg segment see Table 3). Setae strong and distally attenuated, serrated and/or covered by spines, on proximal segments curved and dark, on distal segments (tibiae, tarsi) ventral and lateral setae finer, straight, and more transparent. Setae generally thinner and more pointed on legs I, II than on legs III, IV, where they are stronger, more flattened and more distinctly serrated (with distinct dark spinuli, scales or barbs stronger on external curvature of setae). Setae d of genua and tibiae of legs III, IV, and sometimes also on femora III, IV broadened, blunt. Specimens from Slovak population from Sivec have dorsal setae of femora III, IV, genua IV and tibiae IV less coloured, with transparent serrated margins and darker thin “core”, serrated on both sides (Figs 27D, 29F, G). Formulas of leg setation and solenidia same as in rossica: I (1-7-4-4-20) [1-2-2], II (1-7-4-5-17) [1-1-2], III (2-5-3-4-17) [1-1-0], IV (2-5-3-4-14) [0-1-0]; however, three setae on trochanters III were observed in one case instead of two (Fig. 27E). Slight difference can be observed on length of solenidia: solenidia of genua III, tibiae III (both slightly) and tibiae IV (distinctly) longer than companion setae d. However, only solenidia ϕ 1 of tibiae I, inserted on distinct apophyses, long and tactile, solenidia of tibiae IV fine, setiform, but much shorter than tibiae IV and only slightly exceeding its distal ends (Fig 27C).</p> <p>Juvenile instars One deutonymph is available in slide D of Mihelčič (Fig. 20D 4), observable only from lateral view.</p> <p>Deutonymph (Figs 22F, 25)</p> <p>Measurements. Total (maximum) body length 389, ventral body length 372; body width impossible to measure. All measurements for deutonymph should be taken as approximative, as the specimen is damaged, and some parts (such as distal parts of some setae, or whole setae bs) were difficult to observe and measure.</p> <p>Integument (Fig. 25A, B). Body cuticle variably coloured, notogaster including anogenital region weakly sclerotized, white-coloured. Prodorsum, lateral parts of podosoma and legs with thicker cuticle, light reddish-brown. Body and legs smooth, covered with thin layer of granular cerotegument. Round or oval granules small (1.5–2.8). Prodorsal and gastronotic setae (excluding ro, le, bs) as in rossica, without cerotegument.</p> <p>Prodorsum (Figs 22F, 25A, B). In lateral view with rather distinct prodorsal protuberances, visible straight fold or cuticular thickening above acetabula I in place of prodorsal costulae of adult. Bothridia ear-like, with elongated projections. Setae le (44) strong, barbed, setae ro finer, shorter, smooth. Interlamellar setae (18) strong, thorn-like, pointed, smooth. Exobothridial setae (19) subequal in length to in, rather strong, smooth, larger and thicker than in rossica. Bothridial setae (approx. 102) broken, very difficult to observe, long, setiform with attenuated distal part.</p> <p>Gastronotum (Figs 22F, 25B) Gastronotic setae generally similar to rossica, appearing however generally slightly shorter and more barbed. Longer setae proximally darker, lm, lp, h 1, h 3 inserted on distinct apophyses. Setae of rows l and h (and probably also c 1, c 2 which were very difficult to observe) proximally distinctly barbed, distally flagellate, with very fine, sometimes highly curly ends. Lengths could be measured only approximately, as distal parts of some may be incomplete or broken and/or very curly, approximate measures as follows: c 2 (98), la (59), lm (56), lp (107), h 1 (180), h 2 (52), h 3 (70), p 1 (35). Setae c 3 much shorter (18), fine, smooth. Apertures of opisthonotal glands (gla) well visible, below setae lm. Cornicle (k) short, distally curly (Fig. 22F), positioned between setae la.</p> <p>Legs (Figs 22F, 25B, C). Leg segments and claws similarly formed as in rossica, all legs shorter than body (legs: I—337, II—238, III—297, IV—351). Surface of femora I, trochanters and femora IV with distinct and dense punctuation (pores?) (Fig. 22F). Leg setae also with similar patterns to rossica, dorsal and lateral setae mostly very strong, darkly coloured, serrate or strongly barbed, ventral setae finer, on tarsi setiform, lightly coloured or transparent. Formulas of leg setation and solenidia identical to rossica: I (1-4-4-5-16) [1-2-2], II (1-4-4-4-13) [1-1- 2], III (2-3-3-4-13) [1-1-0], IV (1-2-2-3-12) [0-1-0]. Except of tibiae I and IV, solenidia of genua and tarsi difficult to observe, shorter than companion setae d, which is usually very strong, dark, distinctly serrate or barbed. Tibial solenidia φ of legs I long (over 120), tactile, solenidia φ of tibiae IV (Fig. 25D) much shorter (42), but clearly exceeding length of companion setae d (30). Dorsal setae of femora I (38) and genua I, and lateral setae of tibiae I (42) very strong, remarkably barbed (several rows of relatively long barbs or scales present), companion setae d on tibiae short, thin, minute but well observable. Setae ft ʺ (66) much stronger and longer than ft ʹ, proximally covered up to 2/3 of its length by layer of granular cerotegument (Fig 25C). Famulus of tarsi I emergent, quite fine, setiform. On legs II, lateral setae of tibiae strongly developed, long, setae ft ʺ again strong, long. Legs III with setae ev ʹ and setae d of femora, genua and tibiae very strong, curved, distinctly barbed, on genua and femora clearly overlooking the distal ends of their segments, setae ft ʺ again longer, on both sides with long hairs. Dorsal setae on femora, genua (21) and tibiae of legs IV developed similarly as on legs III, setae d on tibiae (30) particularly strong, with several rows of long spines/barbs. Setae ft ʺ (52) long, with several rows of long barbs on both sides (Fig 25E).</p> <p>Remarks</p> <p>Out of all studied P. piriformis specimens, mites collected from Sivec, Slovakia, in 1989 (two damaged specimens), show several remarkable differences, mentioned in the redescriptions above. The most conspicuous is the flattened lamellar setae with transparent lateral parts and irregular margins, and the broad, flat, partially transparent, curved dorsal setae on femora, genua and tibiae of leg IV (Figs 26E–H, 27B, D, 29F–G). The two specimens have also shorter and flattened epimeral setae, genital setae of subequal size (g 1 is not significantly longer than other ones), and these specimens seem to have also less developed prodorsal costulae. Given that the differences between rossica and piriformis are of similar rank or even more subtle, these specimens may represent a new species. On the other hand, the flattened lamellar setae were observed in some specimens from Austria; certain similarity also exists in the development of leg setae (Fig. 28C), although this similarity is intermediary between the typical piriformis and the Sivec specimens. At the same time, material collected in other locations of Slovakia (Fig 29A, B), not far from Sivec have a perfect match with the typical piriformis. While substantial genetic differences between rossica and piriformis were detected (see below), we could did not do molecular work for the Sivec specimens, so their status cannot be confirmed. We therefore propose to treat the Sivec material as a distinct morphotype “lanceata”, until its specific or subspecific status can be confirmed.</p> </div>	https://treatment.plazi.org/id/DA1C381AFFBF1962B7B2EDB95171CF87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Miko, Ladislav;Kolesnikov, Vasiliy B.;Ermilov, Sergey G.;Klimov, Pavel B.	Miko, Ladislav, Kolesnikov, Vasiliy B., Ermilov, Sergey G., Klimov, Pavel B. (2022): Taxonomy of European Damaeidae (Acari, Oribatida) XI. European species of the genus Piribelba Miko 2021: redescriptions of P. rossica (Bulanova-Zachvatkina 1957) and P. piriformis (Mihelčič, 1964) using morphology and DNA sequence data. Zootaxa 5187 (1): 169-210, DOI: https://doi.org/10.11646/zootaxa.5187.1.11
DA1C381AFF80196CB7B2EBB050CDCF53.text	DA1C381AFF80196CB7B2EBB050CDCF53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Piribelba Miko 2021	<div><p>Key to species of the genus Piribelba</p> <p>1. Notogastral setae setiform, straight or slightly bent, without attenuated and flagelliform tips. Setae of anogenital shield strong and quite long, adanal setae reaching level of insertions of subsequent pairs. Body length 550–670; Mongolia, China............................................................................. P. crassisetosa (Bayartogtokh, 2000).</p> <p>– Notogastral setae slightly to strongly bent, fine, attenuated up to curly tips. Setae of anogenital shield fine and shorter, adanal setae not reaching level of insertions of subsequent pair...................................................... 2</p> <p>2. Lamellar setae with rare, dark, small scales or bristles (often difficult to see) on dorsal side. Lamellar setae setiform, not flattened or extended. Prodorsal costulae anteriorly strongly bent inwards. Dorsal setae of proximal segments of legs III and IV strong, with spinuli, scales or bristles, but not flattened. Body length 690–790; Russia................................................................................................. P. rossica (Bulanova-Zachvatkina, 1962).</p> <p>– Lamellar setae covered by well visible multiple, dark, scales or bristles on dorsal side. Lamellar setae sometimes proximally distinctly flattened and broadened. Prodorsal costulae slightly bent. Dorsal setae of proximal segments of leg III and IV often flattened, broadened, lanceolate, with dentate margins, usually covered by dense spinuli or bristles. Body length 605–730; Europe (Austria, Italy, Slovakia)................................................. P. piriformis (Mihelčič, 1964).</p> </div>	https://treatment.plazi.org/id/DA1C381AFF80196CB7B2EBB050CDCF53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Miko, Ladislav;Kolesnikov, Vasiliy B.;Ermilov, Sergey G.;Klimov, Pavel B.	Miko, Ladislav, Kolesnikov, Vasiliy B., Ermilov, Sergey G., Klimov, Pavel B. (2022): Taxonomy of European Damaeidae (Acari, Oribatida) XI. European species of the genus Piribelba Miko 2021: redescriptions of P. rossica (Bulanova-Zachvatkina 1957) and P. piriformis (Mihelčič, 1964) using morphology and DNA sequence data. Zootaxa 5187 (1): 169-210, DOI: https://doi.org/10.11646/zootaxa.5187.1.11
