identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
67E690268C765B8684F8C5E877CDE39C.text	67E690268C765B8684F8C5E877CDE39C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolephorus bengalensis (Dutt & Babu Rao 1959)	<div><p>Stolephorus bengalensis (Dutt &amp; Babu Rao, 1959)</p><p>[English name:  Hardenberg’s Anchovy] Figs 2, 3</p><p>Anchoviella baganensis bengalensis Dutt &amp; Babu Rao, 1959: 160 [original locality: Waltair and Kakinada, Andhra Pradesh, east coast of India; type locality: Kilakarai, Gulf of Mannar, India, based on the neotype designated by Hata et al. (2019)].</p><p>Stolephorus baganensis macrops (lapsus memoriae for  Stolephorus megalops) (not of Delsman): Whitehead 1967 (in part): 18 (Bay of Bengal).</p><p>Stolephorus insularis (not of Delsman): Whitehead et al. 1988 (in part): 413 (northern part of Indian Ocean); Young et al. 1994: 222, fig. 7 (Wangkun and Fangliao, Taiwan); Wongratana et al. 1999 (in part): 1736 (northern part of Indian Ocean); Gangan et al. 2020: 562, fig. 5 (Kochi, Kerala State, India).</p><p>Stolephorus bengalensis: Hata et al. 2019 (in part): 24, fig. 12 (Pakistan and India; neotype designation).</p><p>Neotype.</p><p>USNM 276476, 45.8 mm SL, Kilakarai, Gulf of Mannar, India, 20 Feb 1964, J. W. Reintjes and P. S. B. R. James.</p><p>Non-type specimens.</p><p>46 specimens, 30.8-58.7 mm SL. India: BMNH 1969.5.30.34-45, 12 specimens, Chennai, Tamil Nadu State; OCF-P 10435, 4 specimens, 30.8-38.1 mm SL, estuary of Hooghly River, West Bengal State (purchased in fish market in Kolkata, West Bengal State); USNM 204227, 21 specimens, 42.7-51.8 mm SL, Sassan Docks, Mumbai, Maharashtra State. Pakistan: KAUM-I. 69286, 50.0 mm SL, KAUM-I. 69287, 58.7 mm SL, KAUM-I. 69288, 50.5 mm SL, KAUM-I. 69289, 54.4 mm SL, KAUM-I. 69290, 49.0 mm SL, KAUM-I. 69291, 53.1 mm SL, KAUM-I. 69292, 47.3 mm SL, KAUM-I. 69294, 58.6 mm, KAUM-I. 69295, 58.6 mm, West Wharf, Karachi.</p><p>Diagnosis.</p><p>A species of  Stolephorus with the following combination of characters: 1UGR 16-19 (modally 18), 1LGR 23-27 (25), 1TGR 40-45 (44); 2UGR 11-15 (13), 2LGR 21-25 (23), 2TGR 33-39 (36); 3UGR 10-12 (11), 3LGR 13-15 (13), 3TGR 23-27 (24); 4UGR 7-9 (8), 4LGR 10-12 (11), 4TGR 17-21 (19); prepelvic scutes 5-7 (6); total vertebrae 40 or 41 (40); long maxilla, posterior tip just reaching or slightly short of posterior margin of opercle; predorsal scutes present; pelvic scute without spine; body scales deciduous; posterior border of pre-opercle concave, indented; paired dark patch on parietal area with little following pigmentation; distinct double pigment lines along dorsum posterior to dorsal fin; black spots below eye and on lower-jaw tip absent; anal-fin base long, 19.0-21.3% (20.2%) of SL; maximum orbit diameter 7.3-8.6% (8.1%) of SL; third dorsal-fin ray long, 18.5-19.9% (19.0%) of SL; pelvic fin rather long, 9.4-11.0% (10.2%) of SL, its posterior tip not reaching to vertical through dorsal-fin origin when depressed in specimens&gt; 50 mm SL; distance between posterior ends of supramaxilla and maxilla 5.3-6.6% (5.8%) of SL.</p><p>Description.</p><p>Data for neotype presented first, followed by data for non-type specimens in parentheses (if different). Counts and measurements, expressed as percentages of SL or HL, given in Tables 2 and 3. Body laterally compressed, elongate, deepest at dorsal-fin origin. Dorsal profile of head and body slightly convex from snout tip to dorsal-fin origin, gently lowering to uppermost point of caudal-fin base. Ventral profile of head and body slightly convex from lower jaw tip to pelvic-fin insertion, thereafter, slowly rising to lowermost point of caudal-fin base. Single spine-like scute just anterior to dorsal-fin origin. Abdomen somewhat rounded, covered with seven (four to seven) spine-like prepelvic scutes. Pelvic scute without spine. Postpelvic scutes absent. Anus just anterior to anal-fin origin. Snout tip rounded; snout length less than eye diameter. Mouth large, inferior, ventral to body axis, extending backwards beyond posterior margin of eye. Maxilla long, its posterior tip pointed, just reaching (or slightly short of) opercle posterior margin. Lower jaw slender. Single row of conical teeth on both jaws and palatine. Patch of fine conical teeth on pterygoid. Several distinct conical teeth on vomer. Several rows of conical teeth on upper edges of basihyal and basibranchial. Eye large, round, covered with adipose eyelid, positioned laterally on head dorsal to horizontal through pectoral-fin insertion, visible in dorsal view. Pupil round. Orbit elliptical. Nostrils close to each other, anterior to orbit. Posterior margin of pre-opercle concave, indented. Subopercle and opercle with smoothly rounded posterior margins. Gill membrane without serrations. Interorbital space flat, width less than eye diameter. Pseudobranchial filaments present, length of longest filament less than eye diameter. Gill rakers long, slender, rough, visible from side of head when mouth opened. Single row of asperities on anterior surface of gill rakers. Isthmus muscle long, reaching anteriorly to posterior margin of gill membranes. Urohyal hidden by isthmus muscle, not visible without dissection. Gill membrane on each side joined distally, most of isthmus muscle exposed, not covered by gill membrane. Scales on lateral surface of body thin, cycloid, deciduous, those on lateral body surface with several centrally continuous vertical grooves and several longitudinal striae anteriorly (Fig. 3). Head scales absent. Fins scaleless, except for broad triangular sheath of scales on caudal fin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray, slightly posterior to middle of body. Dorsal and anal fins with three anteriormost rays unbranched. First dorsal- and anal-fin rays minute. Anteriormost three rays of both dorsal and anal fins closely spaced. Anal-fin origin just below base of tenth (ninth to eleventh) dorsal-fin ray. Posterior tip of depressed anal fin not reaching caudal-fin base. Uppermost pectoral-fin ray unbranched, inserted below body axis. Posterior tip of pectoral fin not reaching to pelvic fin insertion. Dorsal, ventral and posterior margins of pectoral fin nearly linear. Pelvic fin shorter than pectoral fin, insertion anterior to vertical through dorsal-fin origin. Posterior tip of depressed pelvic fin not reaching to vertical through dorsal-fin origin. Caudal fin forked, posterior tips pointed.</p><p>*one specimen with 4 unbranched anal-fin rays.</p><p>Abbreviations: D-P1 (distance from dorsal-fin origin to pectoral-fin insertion); D-P2 (distance from dorsal-fin origin to pelvic-fin insertion); D-A (distance between origins of dorsal and anal fins); P1-P2 (distance between insertions of pectoral and pelvic fins); P2-A (distance between pelvic-fin insertion and anal-fin origin).</p><p>Colour of preserved specimens.</p><p>Body uniformly pale ivory. A pair of distinct dark patches on parietal region, with little pigmentation on occipital area. Double pigmented lines dorsally posterior to dorsal fin. A few melanophores scattered anteriorly on snout. No black spots below eye and on lower-jaw tip. Melanophores scattered along bases of dorsal and anal fins. All fins transparent, melanophores scattered along fin rays of caudal fin and anterior parts of dorsal and anal fins.</p><p>Distribution.</p><p>Stolephorus bengalensis is distributed in the northern Indian Ocean from Pakistan to the Bay of Bengal (Fig. 4). It is abundantly landed and marketed along the coast of the Bay of Bengal.</p><p>Morphological comparisons.</p><p>Stolephorus bengalensis has been considered conspecific with the three new species described herein, the four species being easily separable from all other congeners, except for  Stolephorus acinaces,  Stolephorus andhraensis Babu Rao, 1966,  Stolephorus carpentariae (De Vis, 1882),  Stolephorus hindustanensis Hata &amp; Motomura, 2022,  Stolephorus holodon (Boulenger, 1900),  Stolephorus ronquilloi Wongratana, 1983 and  Stolephorus tamilensis Gangan, Pavan-Kumar, Jahageerdar &amp; Jaiswar, 2020, the former having a concavely indented pre-opercular margin and lacking a spine on the pelvic scute (Whitehead et al. 1988; Wongratana et al. 1999; Kimura et al. 2009; Hata and Motomura 2018a, b, c, d, e, 2021a, b, c, 2022; Hata et al. 2019, 2020a, b, 2021; Gangan et al. 2020). However, the former four species are distinguished from the other seven by having a predorsal scute (vs. absent in the latter) and double dark lines on the dorsum posterior to the dorsal fin (vs. no lines on the dorsum, except in  S. hindustanensis and  S. ronquilloi). Moreover,  S. carpentariae also differs from  S. bengalensis and the three new species in having 19 or 20 branched anal-fin rays [16-18 (rarely 19 or 20) in the remaining five species] and the anal-fin origin located below the origin of the second to sixth dorsal-fin ray (vs. eighth to eleventh) (Whitehead et al. 1988; Wongratana et al. 1999; Gangan et al. 2020; Hata et al. 2020b).  Stolephorus bengalensis,  S. diabolus sp. nov.,  S. eclipsis sp. nov. and  S. eldorado sp. nov. resemble  Stolephorus baganensis Delsman, 1931,  Stolephorus dubiosus Wongratana, 1983 and  Stolephorus tri (Bleeker, 1852) in having a predorsal scute and double pigment lines on the dorsum behind the dorsal fin, but differ in having deciduous body scales (vs. body scales not deciduous) and lacking a spine on the pelvic scute (pelvic scute with a hard posteriorly projecting spine) (Whitehead et al. 1988; Wongratana et al. 1999; Hata et al. 2019). Comparisons of  S. bengalensis with  S. diabolus sp. nov.,  S. eclipsis sp. nov. and  S. eldorado sp. nov. are given in  “Comparisons” under each new species.</p><p>Molecular comparisons.</p><p>Stolephorus bengalensis,  S. diabolus sp. nov. and  S. eldorado sp. nov. were divergent from each other by at least 3.5% COI-based mean uncorrected genetic distance (min-max = 3.5-7.7%) (Fig. 5). In contrast, each species was genetically uniform, with intraspecific differentiation not exceeding 1% (note:  Stolephorus sp. represented by a single specimen - see below), forming clear intraspecific versus interspecific genetic gaps. The ML phylogenetic tree using COI and cytochrome b markers (Fig. 5) was fully resolved, with interspecific relationships supported by bootstrap values above 80%. Each species formed a well-supported monophyletic group, in agreement with their genetic distinction, thereby confirming their taxonomic status, which was further supported by the morphological observations. The COI sequence of an unidentified larva collected from the Segara Anakan Lagoon, Central Java (Nuryanto et al. 2017), indicated either a range extension of  S. eclipsis sp. nov. or the presence of an unidentified species of  Stolephorus, related to  S. bengalensis .</p></div>	https://treatment.plazi.org/id/67E690268C765B8684F8C5E877CDE39C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hata, Harutaka;Lavoue, Sebastien;Motomura, Hiroyuki	Hata, Harutaka, Lavoue, Sebastien, Motomura, Hiroyuki (2022): Description of three new species previously identified as Stolephorus bengalensis (Dutt & Babu Rao, 1959) or Stolephorus insularis Hardenberg, 1933 and a re-description of S. bengalensis (Chordata, Osteichthyes, Clupeiformes, Engraulidae). ZooKeys 1121: 145-173, DOI: http://dx.doi.org/10.3897/zookeys.1121.84171, URL: http://dx.doi.org/10.3897/zookeys.1121.84171
89C301AA78C9595EA68CB5229FDCD7A8.text	89C301AA78C9595EA68CB5229FDCD7A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolephorus diabolus Hata & Lavoué & Motomura 2022	<div><p>Stolephorus diabolus sp. nov.</p><p>[New English name: Demon Anchovy] Figs 1C, 6</p><p>Stolephorus bengalensis (not of Dutt and Babu Rao): Hata et al. 2022: (in part) 34 (Singapore).</p><p>Holotype.</p><p>IPMB-I 13.00001, 49.7 mm SL, Teluk Bahang, Penang, Malaysia.</p><p>Paratypes.</p><p>14 specimens, 28.5-43.7 mm SL. USMFC 82-0017, 43.7 mm SL, collected with the holotype; USMFC 82-0057, 4 specimens, 40.1-41.1 mm SL, estuary of Merbok River, Jeti Semeling, Malaysia; ZUMT 62056, 5 specimens, 28.5-38.4 mm SL, KAUM-I. 163702, 36.3 mm SL, KAUM-I. 163703, 36.4 mm SL, NSMT-P 143554, 36.4 mm SL, NSMT-P 143555, 36.6 mm SL, Singapore.</p><p>Diagnosis.</p><p>A species of  Stolephorus with the following combination of characters: 1UGR 14-16 (modally 16), 1LGR 20-23 (22), 1TGR 35-38 (38); 2UGR 10 or 11 (11), 2LGR 19 or 20 (20), 2TGR 30 or 31 (31); 3UGR 8 or 9 (9), 3LGR 11 or 12 (12), 3TGR 20 or 21 (21); 4UGR 6 or 7 (7), 4LGR 9 or 10 (9), 4TGR 15-17 (17); prepelvic scutes 5-7 (6); total vertebrae 39; long maxilla, posterior tip just reaching or slightly short of posterior margin of opercle; predorsal scute present; pelvic scute without spine; body scales deciduous; posterior border of pre-opercle concave, indented; paired dark patch on parietal area with little following pigmentation; distinct double pigment lines along dorsum posterior to dorsal fin; black spots below eye and on lower-jaw tip absent; anal-fin base long, 19.8-22.3% (mean 20.7%) of SL; maximum orbit diameter 8.1-8.7% (8.3%) of SL; third dorsal-fin ray short, 17.0-18.5% (18.0%) of SL; pelvic fin rather long, 9.6-11.3% (10.0%) of SL, its posterior tip not reaching to vertical through dorsal-fin origin when depressed in specimens&gt; 40 mm SL; distance between posterior ends of supramaxilla and maxilla 5.7-6.4% (6.1%) of SL.</p><p>Description.</p><p>Data for holotype presented first, followed by data for paratypes in parentheses (if different). Counts and measurements, expressed as percentages of SL or HL, given in Tables 5 and 6. Body laterally compressed, elongate, deepest at dorsal-fin origin. Dorsal profile of head and body slightly convex from snout tip to dorsal-fin origin, gently lowering to uppermost point of caudal-fin base. Ventral profile of head and body slightly convex from lower jaw tip to pelvic-fin insertion, thereafter, slowly rising to lowermost point of caudal-fin base. Single spine-like scute just anterior to dorsal-fin origin. Abdomen somewhat rounded. Scutes on ventrum broken in holotype (five to seven spine-like prepelvic scutes on ventrum in paratypes). Pelvic scute without spine. Postpelvic scutes absent. Anus just anterior to anal-fin origin. Snout tip rounded; snout length less than eye diameter. Mouth large, inferior, ventral to body axis, extending backwards beyond posterior margin of eye. Maxilla long, its posterior tip broken in holotype (posterior pointed, just reaching or slightly short of opercle posterior margin in paratypes). Lower jaw slender. Single row of conical teeth on both jaws and palatine. Patch of fine conical teeth on pterygoid. Several distinct conical teeth on vomer. Several rows of conical teeth on upper edges of basihyal and basibranchial. Eye large, round, covered with adipose eyelid, positioned laterally on head dorsal to horizontal through pectoral-fin insertion, visible in dorsal view. Pupil round. Orbit elliptical. Nostrils close to each other, anterior to orbit. Posterior margin of pre-opercle concave, indented. Subopercle and opercle with smoothly rounded posterior margins. Gill membrane without serrations. Interorbital space flat, width less than eye diameter. Pseudobranchial filaments present, length of longest filament less than eye diameter. Gill rakers long, slender, rough, visible from side of head when mouth opened. Single row of asperities on anterior surface of gill rakers. Isthmus muscle long, reaching anteriorly to posterior margin of gill membranes. Urohyal hidden by isthmus muscle, not visible without dissection. Gill membrane on each side joined distally, most of isthmus muscle exposed, not covered by gill membrane. Body scales deciduous, completely lacking on specimens, except for prepelvic scutes. Head scales absent. Fins scaleless, except for broad triangular sheath of scales on caudal fin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray, slightly posterior to middle of body. Dorsal and anal fins with three anteriormost rays unbranched. First dorsal- and anal-fin rays minute. Anteriormost three rays of both dorsal and anal fins closely spaced. Anal-fin origin just below base of eighth (eighth to eleventh) dorsal-fin ray. Posterior tip of depressed anal fin not reaching caudal-fin base. Uppermost pectoral-fin ray unbranched, inserted below body axis. Posterior tip of pectoral fin not reaching to pelvic fin insertion. Dorsal, ventral and posterior margins of pectoral fin nearly linear. Pelvic fin shorter than pectoral fin, insertion anterior to vertical through dorsal-fin origin. Posterior tip of depressed pelvic fin not reaching to vertical through dorsal-fin origin (reaching to vertical through first to sixth dorsal-fin ray origin in some paratypes smaller than 40 mm SL). Caudal fin forked, posterior tips pointed.</p><p>Abbreviations: D-P1 (distance from dorsal-fin origin to pectoral-fin insertion); D-P2 (distance from dorsal-fin origin to pelvic-fin insertion); D-A (distance between origins of dorsal and anal fins); P1-P2 (distance between insertions of pectoral and pelvic fins); P2-A (distance between pelvic-fin insertion and anal-fin origin).</p><p>Colour of preserved specimens.</p><p>Body uniformly pale white. A pair of distinct dark patches on parietal region, with little pigmentation on occipital area. No black spots below eye and on lower-jaw tip. Melanophores scattered on posterior margins of scale pockets on dorsum. Double pigmented lines dorsally posterior to dorsal fin. Melanophores scattered along bases of dorsal and anal fins. All fins transparent, melanophores scattered along fin rays of caudal fin and anterior parts of dorsal and anal fins.</p><p>Distribution.</p><p>Stolephorus diabolus sp. nov. is currently known only from the western coast of the Peninsular Malaysia (Merbok River Estuary and Penang) and Singapore (Fig. 4).</p><p>Etymology.</p><p>The specific name " diabolus " is derived from Latin meaning  “demon”, in reference to the hard spine on the dorsum of the species.</p><p>Comparisons.</p><p>The new species is distinguished from  S. bengalensis,  S. eclipsis and  S. eldorado by lower gill raker counts: 1TGR, 35-38 in  S. diabolus (vs. 40 or more in the other three species); 2TGR, 30 or 31 in  S. diabolus [vs. 33 or more (rarely 30 or 31 in  S. eldorado)]; 3TGR, 20 or 21 in  S. diabolus [vs. 22 or more in the other three species (rarely 21 in  S. eldorado)]; and 4TGR, 15-17 in  S. diabolus (vs. 17 or more) (Fig. 7). Moreover,  S. diabolus has a shorter orbit diameter than  S. eldorado [maximum orbit diameter 8.1-8.7% (mean 8.3%) of SL in  S. diabolus vs. 8.2-9.9% (8.9%) in  S. eldorado; Fig. 8A]. Furthermore,  S. diabolus is distinguished from  S. bengalensis by having a shorter third dorsal-fin ray [17.0-18.5% (mean 18.0%) of SL in  S. diabolus vs. 18.5-19.9% (19.0%) in  S. bengalensis (Fig. 8B)] and lower total vertebral numbers [39 vs. 40 or 41 (modally 40) (Table 4)]. Detailed comparisons of  S. diabolus with  S. eclipsis and  S. eldorado are given in  “Comparisons” under each species.</p></div>	https://treatment.plazi.org/id/89C301AA78C9595EA68CB5229FDCD7A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hata, Harutaka;Lavoue, Sebastien;Motomura, Hiroyuki	Hata, Harutaka, Lavoue, Sebastien, Motomura, Hiroyuki (2022): Description of three new species previously identified as Stolephorus bengalensis (Dutt & Babu Rao, 1959) or Stolephorus insularis Hardenberg, 1933 and a re-description of S. bengalensis (Chordata, Osteichthyes, Clupeiformes, Engraulidae). ZooKeys 1121: 145-173, DOI: http://dx.doi.org/10.3897/zookeys.1121.84171, URL: http://dx.doi.org/10.3897/zookeys.1121.84171
B61FE9B1A48559B99DA13C8F757769B6.text	B61FE9B1A48559B99DA13C8F757769B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolephorus eclipsis Hata & Lavoué & Motomura 2022	<div><p>Stolephorus eclipsis sp. nov.</p><p>[New English name: Eclipse Anchovy] Fig. 9</p><p>Holotype.</p><p>MZB 26452, 40.3 mm SL, Bintan Island, Riau Archipelago, Indonesia.</p><p>Paratypes.</p><p>28 specimens, 32.0-43.7 mm SL. LBRC-F 5039, 35.4 mm SL, LBRC-F 5040, 35.3 mm SL, LBRC-F 5041, 36.1 mm SL, Tanjungpinang, Bintan Island, Riau Archipelago, Indonesia; MZB 26440, 32.0 mm SL, MZB 26441, 36.1 mm SL, MZB 26442, 35.1 mm SL, MZB 26443, 34.7 mm SL, MZB 26444, 34.5 mm SL, MZB 26445, 36.3 mm SL, MZB 26446, 36.2 mm SL, MZB 26447, 38.8 mm SL, MZB 26448, 39.2 mm SL, MZB 26449, 37.7 mm SL, MZB 26450, 40.0 mm SL, MZB 26451, 36.8 mm SL, 26453, 36.4 mm SL, MZB 26454, 39.0 mm SL, MZB 26455, 41.3 mm SL, MZB 26456, 43.7 mm SL, MZB 26457, 39.8 mm SL, MZB 26458, 40.7 mm SL, MZB 26459, 43.2 mm SL, MZB 26460, 43.7 mm SL, MZB 26461, 5 specimens, 38.3-42.4 mm SL, collected with the holotype.</p><p>Diagnosis.</p><p>A species of  Stolephorus with the following combination of characters: 1UGR 19-21 (modally 20), 1LGR 26-30 (28), 1TGR 47-51 (47); 2UGR 13-16 (14), 2LGR 24-27 (25), 2TGR 37-42 (39); 3UGR 10-13 (12), 3LGR 14-16 (15), 3TGR 25-28 (27); 4UGR 8-11 (9), 4LGR 11-13 (12), 4TGR 19-24 (21); prepelvic scutes 5-7 (6); total vertebrae 38-39 (39); long maxilla, posterior tip just reaching or slightly short of posterior margin of opercle; predorsal scute present; pelvic scute without spine; body scales deciduous; posterior border of pre-opercle concave, indented; paired dark patch on parietal area with little following pigmentation; distinct double pigment lines along dorsum posterior to dorsal fin; black spots below eye and on lower-jaw tip absent; anal-fin base short, 17.6-19.3% (mean 18.6%) of SL; third dorsal-fin ray 16.5-18.8% (17.6%) of SL; pelvic fin short, 8.7-9.9% (9.4%) of SL, its posterior tip usually not reaching to vertical through dorsal-fin origin when depressed; distance between posterior ends of supramaxilla and maxilla 4.7-5.4% (5.1%) of SL; pre-dorsal-fin length 51.3-54.9% (53.4%) of SL; dorsal-fin base short, 13.1-14.5% (13.8%) of SL.</p><p>Description.</p><p>Data for holotype presented first, followed by data for paratypes in parentheses (if different). Counts and measurements, expressed as percentages of SL or HL, given in Tables 5 and 6. Body laterally compressed, elongate, deepest at dorsal-fin origin. Dorsal profile of head and body slightly convex from snout tip to dorsal-fin origin, gently lowering to uppermost point of caudal-fin base. Ventral profile of head and body slightly convex from lower jaw tip to pelvic-fin insertion, thereafter, slowly rising to lowermost point of caudal-fin base. Single spine-like scute just anterior to dorsal-fin origin. Abdomen somewhat rounded, covered with six (five to seven) spine-like prepelvic scutes. Pelvic scute without spine. Postpelvic scutes absent. Anus just anterior to anal-fin origin. Snout tip rounded; snout length less than eye diameter. Mouth large, inferior, ventral to body axis, extending backwards beyond posterior margin of eye. Maxilla long, its posterior tip pointed, just reaching (or slightly short of) opercle posterior margin. Lower jaw slender. Single row of conical teeth on both jaws and palatine. Patch of fine conical teeth on pterygoid. Several distinct conical teeth on vomer. Several rows of conical teeth on upper edges of basihyal and basibranchial. Eye large, round, covered with adipose eyelid, positioned laterally on head dorsal to horizontal through pectoral-fin insertion, visible in dorsal view. Pupil round. Orbit elliptical. Nostrils close to each other, anterior to orbit. Posterior margin of pre-opercle concave, indented. Subopercle and opercle with smoothly rounded posterior margins. Gill membrane without serrations. Interorbital space flat, width less than eye diameter. Pseudobranchial filaments present, length of longest filament less than eye diameter. Gill rakers long, slender, rough, visible from side of head when mouth opened. Single row of asperities on anterior surface of gill rakers. Isthmus muscle long, reaching anteriorly to posterior margin of gill membranes. Urohyal hidden by isthmus muscle, not visible without dissection. Gill membrane on each side joined distally, most of isthmus muscle exposed, not covered by gill membrane. Body scales deciduous, completely lacking on specimens, except for prepelvic scutes. Head scales absent. Fins scaleless, except for broad triangular sheath of scales on caudal fin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray, slightly posterior to middle of body. Dorsal and anal fins with three anteriormost rays unbranched. First dorsal- and anal-fin rays minute. Anteriormost three rays of both dorsal and anal fins closely-spaced. Anal-fin origin just below base of eighth (eighth to eleventh) dorsal-fin ray. Posterior tip of depressed anal fin not reaching caudal-fin base. Uppermost pectoral-fin ray unbranched, inserted below body axis. Posterior tip of pectoral fin not reaching to pelvic fin insertion. Dorsal, ventral and posterior margins of pectoral fin nearly linear. Pelvic fin shorter than pectoral fin, insertion anterior to vertical through dorsal-fin origin. Posterior tip of depressed pelvic fin not reaching to vertical through dorsal-fin origin (reaching to vertical through first to third dorsal-fin ray origin in some paratypes). Caudal fin forked, posterior tips pointed.</p><p>Colour of preserved specimens.</p><p>Body uniformly pale ivory. A pair of distinct dark patches on parietal region, with little pigmentation on occipital area. Double pigmented lines dorsally posterior to dorsal fin. A few melanophores scattered anteriorly on snout. No black spots below eye and on lower-jaw tip. Melanophores scattered along bases of dorsal and anal fins. All fins transparent, melanophores scattered along fin rays of caudal fin and anterior parts of dorsal and anal fins.</p><p>Distribution.</p><p>Stolephorus eclipsis sp. nov. is currently known only from Bintan Island, Riau Archipelago, Indonesia (Fig. 4).</p><p>Etymology.</p><p>The specific name " eclipsis " refers to eclipse, reminiscent of the concave pre-opercle of the new species.</p><p>Comparisons.</p><p>The new species differs from  S. bengalensis,  S. diabolus and  S. eldorado in having higher gill raker counts [1TGR, 47-51 or more in  S. eclipsis (vs. 47 or fewer in the other three species); 2TGR, 37-42 in  S. eclipsis (vs. 39 or fewer); 3TGR, 25-28 in  S. eclipsis (vs. 27 or fewer); and 4TGR, 19-24 in  S. eclipsis (vs. 22 or fewer) (Fig. 7)], a shorter anal-fin base (17.6-19.3% of SL in  S. eclipsis vs. 19.0-21.3% in  S. bengalensis, 19.8-22.3% in  S. diabolus and 19.0-22.3% in  S. eldorado; Fig. 8C) and pelvic fin [8.7-9.9% (mean 9.4%) of SL in  S. eclipsis vs. 9.4-11.0% (10.2%) in  S. bengalensis, 9.6-11.3% (10.0%) in  S. diabolus and 9.1-11.0% (10.0%) in  S. eldorado; Fig. 8D] and shorter distance between the posterior ends of the supramaxilla and maxilla [4.7-5.4% (5.1%) of SL in  S. eclipsis vs. 5.3-6.6% (5.8%) in  S. bengalensis, 5.7-6.4% (6.1%) in  S. diabolus and 5.0-6.3% (5.6%) in  S. eldorado; Fig. 8E].  Stolephorus eclipsis also differs from  S. bengalensis in having a shorter third dorsal-fin ray (16.5-18.8% of SL in  S. eclipsis vs. 18.5-19.9% in  S. bengalensis; Fig. 8B) and lower total vertebral number [38-39 (modally 39) vs. 40 or 41 (40) (Table 4)]. Moreover,  S. eclipsis is distinguished from  S. diabolus by a greater pre-dorsal-fin distance [51.3-54.9% (mean 53.4%) of SL in  S. eclipsis vs. 51.3-52.9% (52.1%) in  S. diabolus; Fig. 10A] and shorter dorsal-fin base (13.1-14.5% of SL vs. 13.9-16.6%; Fig. 10B) and postorbital head length (11.5-12.9% of SL vs. 12.8-14.2%; Fig. 10C).</p></div>	https://treatment.plazi.org/id/B61FE9B1A48559B99DA13C8F757769B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hata, Harutaka;Lavoue, Sebastien;Motomura, Hiroyuki	Hata, Harutaka, Lavoue, Sebastien, Motomura, Hiroyuki (2022): Description of three new species previously identified as Stolephorus bengalensis (Dutt & Babu Rao, 1959) or Stolephorus insularis Hardenberg, 1933 and a re-description of S. bengalensis (Chordata, Osteichthyes, Clupeiformes, Engraulidae). ZooKeys 1121: 145-173, DOI: http://dx.doi.org/10.3897/zookeys.1121.84171, URL: http://dx.doi.org/10.3897/zookeys.1121.84171
8FC6B0AE0FE25383B73FDA924E194586.text	8FC6B0AE0FE25383B73FDA924E194586.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stolephorus eldorado Hata & Lavoué & Motomura 2022	<div><p>Stolephorus eldorado sp. nov.</p><p>[New English name: El Dorado Anchovy] Figs 1A, B, D, E, F, 11</p><p>Stolephorus insularis (not of Delsman): Whitehead et al. 1988 (in part): 413 unnumbered fig. (Taiwan to Java Sea); Young et al. 1999: 222, fig. 7 (western coast of Taiwan); Wongratana et al. 1999 (in part): 1736, unnumbered fig. (Taiwan to Java Sea); Hata 2018: 41, unnumbered figs (Ha Long Bay, northern Vietnam).</p><p>Stolephorus tri (not of Bleeker): Zhang 2001: 129, fig. II-59 (Beihai City, Guangxi Province, China).</p><p>Stolephorus bengalensis (not of Dutt and Babu Rao): Hata et al. 2019 (in part): 24, fig. 12a, b (Taiwan; Hainan Island, China; Ha Long Bay, Vietnam; Gulf of Thailand; Songkhla, Thailand; Kuala Terengganu, Terengganu, Malaysia); Hata 2019: 206, unnumbered figs (Ke-tzu-liao, Ziguan District, Kaohsiung, Taiwan); Hata et al. 2022: (in part) 34 (Wenzhou City, Zheijiang Province, China).</p><p>Holotype.</p><p>KAUM-I. 94517, 44.4 mm SL, Ha Long Bay, Ha Long City, Quang Ninh District, Vietnam (purchased at fish market in Ha Long City), 24 Oct 2016; coll. by H. Hata and M. Matsunuma.</p><p>Paratypes.</p><p>57 specimens, 37.5-58.8 mm SL. Taiwan: ASIZP 73957, 51.8 mm SL, Fangyan, Changhua (23°57'42.8"N, 120°17'39.8"E); KAUM-I. 110282, 49.5 mm SL, KAUM-I. 113142, 54.0 mm SL, KAUM-I. 113143, 45.5 mm SL, KAUM-I. 113144, 44.3 mm SL, KAUM-I. 113145, 46.3 mm SL, KAUM-I. 113146, 37.5 mm SL, KAUM-I. 113147, 47.3 mm SL, KAUM-I. 113148, 55.3 mm SL, KAUM-I. 113149, 49.4 mm SL, KAUM-I. 113150, 45.9 mm SL, KAUM-I. 113151, 47.3 mm SL, off Ke-tzu-liao, Ziguan District, Kaohsiung. China: BMNH 1965.4.1.981-983, 3 specimens, 58.1-58.8 mm SL, Stanley, Hong Kong. Vietnam: FRLM 49725, 46.9 mm SL, KAUM-I. 67322, 46.7 mm SL, KAUM-I. 67405, 45.6 mm SL, KAUM-I. 94509, 41.4 mm SL, KAUM-I. 94518, 43.7 mm SL, KAUM-I. 94519, 38.8 mm SL, KAUM-I. 94520, 41.7 mm SL, KAUM-I. 94521, 43.4 mm SL, Ha Long Bay, Ha Long, Quang Ninh Province. Thailand: CAS 46931, 8 specimens, 44.4-46.7 mm SL, between Bangsaen and Chol Buri, Chol Buri, Gulf of Thailand; CAS 230414, 4 specimens, 39.9-45.8 mm SL, Lem Nam Point, south tip of Lem Nam Peninsula, Gulf of Thailand (12°02'55"N, 102°35'35"E), approx. 0.6 m depth; KAUM-I. 23190, 48.2 mm SL, Gulf of Thailand (obtained at fish market in Mahachai, Samut Prakan Province), trawl; NSMT-P 142790, 47.9 mm SL, Ko Maeo Island, off Songkhla; URM-P 12398, 3 specimens, 43.0-45.5 mm SL, Song Khula; URM-P 13635, 11 specimens, 46.2-49.5 mm SL, Ang Sila. Indonesia: BMNH 1965.10.20.42-47, 6 specimens, 40.6-43.5 mm SL, 20 miles (approx. 32 km) east of Tegal, Java.</p><p>Diagnosis.</p><p>A species of  Stolephorus with the following combination of characters: 1UGR 16-21 (modally 18), 1LGR 23-28 (25), 1TGR 40-47 (42); 2UGR 10-14 (13), 2LGR 20-24 (23), 2TGR 33-38 (rarely 30) (modally 36); 3UGR 8-12 (modally 10), 3LGR 12-14 (13), 3TGR 20-26 (23); 4UGR 7-10 (8), 4LGR 9-12 (11), 4TGR 16-22 (18); prepelvic scutes 5-7 (6); total vertebrae 38-40 (39); long maxilla, posterior tip just reaching or slightly short of posterior margin of opercle; predorsal scutes present; pelvic scute without spine; body scales deciduous; posterior border of pre-opercle concave, indented; paired dark patch on parietal area with little following pigmentation; distinct double pigment lines along dorsum posterior to dorsal fin; black spots below eye and on lower-jaw tip absent; anal-fin base long, 19.0-22.3% (20.4%) of SL; orbit rather long, 8.2-9.9% (8.9%) of SL; third dorsal-fin ray short, 15.9-18.6% (17.4%) of SL; pelvic fin rather long, 9.1-11.0% (10.0%) of SL, its posterior tip usually not reaching to vertical through dorsal-fin origin when depressed in individuals&gt; 50 mm SL; distance between posterior ends of supramaxilla and maxilla 5.0-6.3% (5.6%) of SL.</p><p>Description.</p><p>Data for holotype presented first, followed by data for paratypes in parentheses (if different). Counts and measurements, expressed as percentages of SL or HL, given in Tables 2 and 3. Body laterally compressed, elongate, deepest at dorsal-fin origin. Dorsal profile of head and body slightly convex from snout tip to dorsal-fin origin, gently lowering to uppermost point of caudal-fin base. Ventral profile of head and body slightly convex from lower jaw tip to pelvic-fin insertion, thereafter, slowly rising to lowermost point of caudal-fin base. Single spine-like scute just anterior to dorsal-fin origin. Abdomen somewhat rounded, covered with six (five to seven) spine-like prepelvic scutes. Pelvic scute without spine. Postpelvic scutes absent. Anus just anterior to anal-fin origin. Snout tip rounded; snout length less than eye diameter. Mouth large, inferior, ventral to body axis, extending backwards beyond posterior margin of eye. Maxilla long, its posterior tip pointed, just reaching (or slightly short of) opercle posterior margin. Lower jaw slender. Single row of conical teeth on both jaws and palatine. Patch of fine conical teeth on pterygoid. Several distinct conical teeth on vomer. Several rows of conical teeth on upper edges of basihyal and basibranchial. Eye large, round, covered with adipose eyelid, positioned laterally on head dorsal to horizontal through pectoral-fin insertion, visible in dorsal view. Pupil round. Orbit elliptical. Nostrils close to each other, anterior to orbit. Posterior margin of pre-opercle concave, indented. Subopercle and opercle with smoothly rounded posterior margins. Gill membrane without serrations. Interorbital space flat, width less than eye diameter. Pseudobranchial filaments present, length of longest filament less than eye diameter. Gill rakers long, slender, rough, visible from side of head when mouth opened. Single row of asperities on anterior surface of gill rakers. Isthmus muscle long, reaching anteriorly to posterior margin of gill membranes. Urohyal hidden by isthmus muscle, not visible without dissection. Gill membrane on each side joined distally, most of isthmus muscle exposed, not covered by gill membrane. Body scales deciduous, completely lacking on all specimens, except for prepelvic scutes. Head scales absent. Fins scaleless, except for broad triangular sheath of scales on caudal fin. Dorsal-fin origin posterior to vertical through base of last pelvic-fin ray, slightly posterior to middle of body. Dorsal and anal fins with three anteriormost rays unbranched. First dorsal- and anal-fin rays minute. Anteriormost three rays of both dorsal and anal fins closely spaced. Anal-fin origin just below base of ninth (eighth to tenth) dorsal-fin ray. Posterior tip of depressed anal fin not reaching caudal-fin base. Uppermost pectoral-fin ray unbranched, inserted below body axis. Posterior tip of pectoral fin not reaching to pelvic fin insertion. Dorsal, ventral and posterior margins of pectoral fin nearly linear. Pelvic fin shorter than pectoral fin, insertion anterior to vertical through dorsal-fin origin. Posterior tip of depressed pelvic fin not reaching to vertical through dorsal-fin origin (reaching to vertical through first to fourth dorsal-fin ray origin in some paratypes smaller than 50 mm SL). Caudal fin forked, posterior tips pointed.</p><p>Colour of fresh specimens.</p><p>(based on colour photographs of KAUM-I. 67322, 46.7 mm SL, KAUM-I. 67405, 45.6 mm SL, KAUM-I. 94517, 44.4 mm SL, KAUM-I. 94521, 43.4 mm SL and KAUM-I. 110282, 49.5 mm SL). Body yellowish milky-white, a silver longitudinal band, of width slightly less than pupil diameter, extending from just above posterior tip of pectoral fin to caudal-fin base. Caudal fin yellow with black posterior margin. Melanophores scattered along caudal-fin rays, ventral surface of caudal peduncle and bases of dorsal and anal fins. Fin rays of dorsal and anal fins yellow. A few melanophores scattered on snout and fin rays of anterior part of dorsal fin. Fin rays and fin membrane of pectoral and pelvic fins transparent whitish, lacking melanophores. A pair of dark patches on parietal region, with little pigmentation on occipital area. Distinct double pigment lines on dorsum from end of dorsal-fin base to caudal-fin base. Body wholly yellowish when freshly caught (Fig. 11E), quickly becoming white after death (Figs 11F-H).</p><p>Colour of preserved specimens.</p><p>Body uniformly pale white. A pair of distinct dark patches on parietal region, with little pigmentation on occipital area. Melanophores scattered on posterior margins of scale pockets on dorsum. Double pigmented lines dorsally posterior to dorsal fin. A few melanophores scattered anteriorly on snout. No black spots below eye and on lower-jaw tip. Melanophores scattered along bases of dorsal and anal fins. All fins transparent, with melanophores scattered along fin rays of caudal fin and anterior parts of dorsal and anal fins.</p><p>Distribution.</p><p>Stolephorus eldorado sp. nov. is distributed in the western Pacific from Taiwan to Java, Indonesia (Fig. 4). The species is abundantly caught by trawl and marketed fresh in northern Vietnam. It is a set net bycatch in south-western Taiwan.</p><p>Etymology.</p><p>The specific name " eldorado ", referring to the mythical city of gold, reflects the bright yellow colouration of the new species.</p><p>Morphological comparisons.</p><p>Stolephorus eldorado sp. nov. has been previously identified as  S. insularis or  S. bengalensis (together with  S. bengalensis,  S. diabolus and  S. eclipsis as recognised here) (e.g. Whitehead et al. 1988; Wongratana et al. 1999; Hata et al. 2019). However,  S. eldorado is distinguished from  S. diabolus and  S. eclipsis by having an intermediate number of gill rakers on each gill arch (Table 2; Fig. 7). More detailed comparisons of  S. eldorado with the latter two species are given in  “Comparisons” under each species.</p><p>Although  S. eldorado sp. nov. closely resembles  S. bengalensis in having very similar numbers of gill rakers on each gill arch, the former differs from the latter in having a greater orbit diameter [maximum orbit diameter 8.2-9.9% (mean 8.9%) of SL vs. 7.3-8.6% (8.1%) in  S. bengalensis (Fig. 8A)], shorter third dorsal-fin ray [15.9-18.6% (mean 17.4%) of SL vs. 18.5-19.9% (19.0%)] (Fig. 8B) and fewer total vertebrae [38-40 (modally 39) vs. 40 or 41 (40)] (Table 4).</p></div>	https://treatment.plazi.org/id/8FC6B0AE0FE25383B73FDA924E194586	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hata, Harutaka;Lavoue, Sebastien;Motomura, Hiroyuki	Hata, Harutaka, Lavoue, Sebastien, Motomura, Hiroyuki (2022): Description of three new species previously identified as Stolephorus bengalensis (Dutt & Babu Rao, 1959) or Stolephorus insularis Hardenberg, 1933 and a re-description of S. bengalensis (Chordata, Osteichthyes, Clupeiformes, Engraulidae). ZooKeys 1121: 145-173, DOI: http://dx.doi.org/10.3897/zookeys.1121.84171, URL: http://dx.doi.org/10.3897/zookeys.1121.84171
