taxonID	type	description	language	source
8D6CD3C552FE68B46CC0086D3ABCEB4F.taxon	description	Description. Trees, scrambling shrubs or lianas, up to 50 m tall, monoecious, dioecious or putatively androdioecious. Indumentum, when present, of simple, fasciculate, stellate, or scale-like hairs. Leaves alternate, simple, distichous, margins entire, stipules absent. Inflorescences terminal or axillary, ramiflorous in leaf axils, on young or old leafless branches or cauliflorous, single to many-flowered, pedunculate or subsessile, bracts often present. Flowers bisexual or unisexual, actinomorphic, generally trimerous. Sepals in a single whorl, (2) 3 (4), valvate or imbricate in bud, free or basally to fully connate. Petals 3, 4 or 6, in 1 or 2 (3) whorls, generally differentiated into an inner and outer whorl alternating with the sepals, valvate or imbricate in bud, free, basally or fully connate. Stamens 3 to numerous, inserted onto a flat or convex receptacle; anthers generally exceeded by the connective apex, which forms a a protective cover at the top of the stamen; connective apex flat, extended (tongue shaped) or absent; filaments short or absent, free or rarely fused; staminodes absent or present. Carpels 1 to numerous, free or more rarely fused (syncarpous) in flower; stigma capitate, oblong or variously folded; ovules 1 to numerous, uni- or biseriate, basal or lateral. Fruit generally apocarpous, each carpel producing a single monocarp, or more rarely pseudosyncarpous (carpels fusing during fructification) or syncarpous (unilocular fruits resulting from syncarpous flowers), indehiscent or sometimes dehiscent; monocarps 1 to numerous, sessile to long-stipitate, cylindrical, globose, ovoid, ellipsoid, club-shaped or moniliform, 3 to over 40 mm in length, usually large; seeds 1 to numerous per monocarp, uni- or biseriate, or unordered in syncarpous species, sometimes with arilor sarcotesta; endosperm ruminate, hard.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8D6CD3C552FE68B46CC0086D3ABCEB4F.taxon	distribution	Distribution. Pantropical, from the Pacific and northern Australia to South East Asia (including southern China), India, Madagascar, tropical Africa, temperate eastern North America south to Central America and South America. 113 genera, and around 2550 species. In Cameroon, 28 genera and 163 species reported to date.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
554BD65EDEF8901726708EB1CD6A964F.taxon	description	Description. Same as species. A genus of lianas with three species from Central Africa, in the Democratic Republic of the Congo, Cameroon and Angola (Cabinda); one species in Cameroon, endemic. This genus was phylogenetically validated (Guo et al. 2017 b).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A438DDF76CE7AAE97B270DAB38B8B412.taxon	description	Fig. 4; Map 1 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A438DDF76CE7AAE97B270DAB38B8B412.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2980, Apr 1904: holotype: B [B 10 0153055]; isotypes: BM [BM 001125042]; BR [BR 000008800398]; [BR 000008800398]; G [G 00308361]; GOET [GOET 005676]; HBG [HBG- 502538]; K [K 000198949]; L [L. 1754813]; M [M- 0107910]; P [P 00363341]; S [S 03 - 2239]; WAG [WAG 0053550].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A438DDF76CE7AAE97B270DAB38B8B412.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent with short sericeous hairs. Leaves: petiole 3 - 4 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous, grooved, blade inserted on the side of the petiole, 7.5 - 10 cm long, 4 - 5 cm wide, elliptic, apex acuminate, acumen ca. 1 cm long, base obtuse, subcoriaceous, below sparsely pubescent to glabrous when young, glabrous when old, above glabrous when young and old, discolorous, whitish below (both when fresh and dry); midrib flat or sunken, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 8 to 13 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on young foliate branches, leaf-opposed or extra-axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 18 - 22 mm long, ca. 1 mm in diameter, pubescent; in fruit 20 - 40 mm long, 2 - 3 mm in diameter, pubescent; bracts not seen; sepals 3, valvate, free, 1 - 2 mm long, 1 - 2 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free, subequal; outer petals 3, valvate, 5 - 6 mm long, 3 - 4 mm wide, ovate, apex acute, base narrowed, margins flat, pubescent outside, pubescent inside; inner petals 3, valvate, 3 - 3.5 mm long, 2 - 3 mm wide, elliptic, apex obtuse, base truncate, margins flat, pubescent outside, pubescent inside; number of stamens not counted, number of rows not seen, 2 - 3 mm long, oblong; anthers not septate; connective discoid, pubescence not seen; staminodes absent; carpels not seen but free. Monocarps stipitate, stipes 5 - 10 mm long, 1 - 2 mm in diameter; monocarps 3 to 7, 10 - 15 mm long, 5 - 8 mm in diameter, ellipsoid, apex apiculate, pubescent, smooth, not ribbed, color unknown; seed 1, 13 - 15 mm long, 5 - 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A438DDF76CE7AAE97B270DAB38B8B412.taxon	distribution	Distribution. endemic to Cameroon; known from the South Region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A438DDF76CE7AAE97B270DAB38B8B412.taxon	materials_examined	Specimens examined. South Region: Bipindi, 3.05 ° N, 10.25 ° E, 01 January 1905, Zenker G. A. 2102 a (BM, BR, E, G, K, P); Bipindi, 3.05 ° N, 10.25 ° E, 01 January 1904, Zenker G. A. 3023 (BM, G, K, L, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 March 1914, Zenker G. A. 576 (MA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
89FD6ADFE6AD7755608DF8AD05229862.taxon	description	Description. Trees, up to 30 m tall, d. b. h. up to 50 cm; stilt roots or buttresses absent, slash yellow. Indumentum of simple, stellate and / or fasciculate hairs. Leaves: petiole 2 - 9 mm long, 1 - 2 mm in diameter; blade 3.5 - 29.5 cm long, 1.5 - 10.5 cm wide, elliptic to obovate, apex acuminate to acute, base narrowly cuneate to shortly attenuate to rounded; midrib sunken or flat; secondary veins 8 to 13 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 6 perianth parts in 2 whorls, 1 per inflorescence; pedicel 4 - 19 mm long; in fruit 10 - 27 mm long; bracts 1 to 2, basal and one upper towards the middle or lower half of pedicel, 2 - 8 mm long; sepals 3, valvate, free, 5 - 22 mm long, triangular, apex acute, base truncate; petals free; outer petals absent; inner petals 3, valvate, 12 - 34 mm long, 5 - 19 mm wide, ovate, margins inversely Y-shaped ridged, apex acute, base broad and concave; stamens 60 to 200, in 5 to 6 rows, 2 - 4 mm long, linear; connective tongue shaped or flattened, glabrous; staminodes absent; carpels free, 20 to 70, ovary 2 - 4 mm long, stigma lobed or cylindrical, pubescent. Monocarps stipitate, stipes 6 - 59 mm long, 5 to 55 monocarps, 18 - 35 mm long, 4 - 14 mm in diameter, ellipsoid to obovoid, apex sometimes mucronate, smooth, glossy; seed 1, 20 - 30 mm long, ca. 10 mm in diameter, ellipsoid; aril absent. A genus of eight species mostly distributed across west and central Africa, with one endemic species in Tanzania; four species occur in Cameroon, none endemic. The genus is easily identifiable when sterile by its yellow slash, and when fertile, by its leaf opposed or extra-axillary (terminal) flowers with 3 sepals and 3 petals, and stipitate monocarps with a single seed.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5B2566F66BC74F49FD4FC64E395C949.taxon	description	Figs 5, 7; Map 1 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5B2566F66BC74F49FD4FC64E395C949.taxon	materials_examined	Type. Angola. Cabinda, Munze, ring at Buco Zau, Gossweiler J. 6675, 11 Sep 1916: holotype K [not seen]: isotypes: BM [BM 000547034]; COI [COI 00004913]; LISC [LISC 000073, LISC 000072, LISC 000075, LISC 000074].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5B2566F66BC74F49FD4FC64E395C949.taxon	description	Description. Tree, 3 - 30 m tall, d. b. h. 3 - 50 cm; stilt roots or buttresses absent, slash yellow. Indumentum of simple, bifid and fasciculate hairs; old leafless branches glabrous, young foliate branches sparsely pubescent. Leaves: petiole 2 - 8 mm long, 1 - 2 mm in diameter, sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 3.5 - 26 cm long, 1.5 - 9.5 cm wide, elliptic to obovate, apex acuminate to acute, acumen 1 cm long, base narrowly cuneate to shortly attenuate, coriaceous to subcoriaceous, below pubescent when young and old with simple or bifid hairs pointing towards the leaf apex, above sparsely pubescent when young and old, concolorous; midrib sunken or flat, above sparsely pubescent to glabrous when young and old, below pubescent when young and old; secondary veins 8 to 13 pairs, sparsely pubescent below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 6 perianth parts in 2 whorls, 1 per inflorescence; pedicel 7 - 14 mm long, 1 - 2 mm in diameter, pubescent; in fruit 27 mm long, 2 - 3 mm in diameter, pubescent; bracts 1 - 2, basal and one upper towards the middle of pedicel, ca. 4 mm long, ca. 2 mm wide; sepals 3, valvate, free, 7 mm long, ca. 4 mm wide, triangular, apex acute, base truncate, green, pubescent outside, glabrous inside, margins flat; petals free; outer petals absent; inner petals 3, valvate, 15 - 33 mm long, 5 - 15 mm wide, ovate to inversely Y-shaped ridged, apex acute, base broad and concave, greenish yellow, margins flat, pubescent outside, glabrous inside; stamens 110 to 175, in 5 to 6 rows, 2 - 4 mm long, linear; connective tongue shaped, glabrous, yellow; staminodes absent; carpels free, 35 to 70, ovary 3 - 4 mm long, stigma lobed, pubescent. Monocarps stipitate, stipes 10 - 40 mm long, 1 - 2 mm in diameter; monocarps 3 to 34, 20 - 35 mm long, 9 - 14 mm in diameter, ellipsoid to obovoid, apex sometimes mucronate, sparsely pubescent, smooth, glossy, black when ripe; seed 1, ca. 30 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5B2566F66BC74F49FD4FC64E395C949.taxon	distribution	Distribution. From Nigeria (one collection) to the Republic of Congo and the extreme west of the Democratic Republic of Congo; in Cameroon known from the East, South, Littoral, Center and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5B2566F66BC74F49FD4FC64E395C949.taxon	materials_examined	Selected specimens examined. Central Region: near Ebolbom village 3 km est of Ngoumou 2 km north west of Otele, 3.59 ° N, 11.28 ° E, 02 May 2013, Couvreur T. L. P. 426 (WAG, YA); Ottotomo Forest Reserve 3 km after reserve base near small loggers road, 3.66 ° N, 11.28 ° E, 02 May 2013, Couvreur T. L. P. 437 (WAG, YA); Mefou Proposed National Park, 3.62 ° N, 11.57 ° E, 15 March 2004, Etuge M. 5139 (K, YA); Mbam Minkom, 3.96 ° N, 11.36 ° E, 19 September 2013, Kamdem N. 143 (YA); Nguila 1, 4.77 ° N, 11.75 ° E, 30 April 2017, Kamdem N. 521 (YA); Colline entre Tcherikoy et Sokelle II (30 km NW Eseka), 3.78 ° N, 10.96 ° E, 14 December 1973, Letouzey R. 12361 (P, YA). East Region: 77 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.72 ° E, 05 March 2019, Couvreur T. L. P. 1203 (MPU, WAG, YA); Deng Deng, 5.21 ° N, 13.44 ° E, 19 April 2016, Kamdem N. 422 (YA); 16 km E de Dimako, 4.38 ° N, 13.57 ° E, 15 December 1965, Leeuwenberg A. J. M. 7355 (BR, K, MO, P, PHA, WAG, YA); 15 km E of Dimako, 4.38 ° N, 13.57 ° E, 08 February 1966, Leeuwenberg A. J. M. 7787 (BR, C, K, MO, P, WAG, YA); Route Mintom I (70 km E de Djoum) - Alati (100 km SE de Djoum) - PK 63, 2.83 ° N, 13.35 ° E, 01 January 1973, Letouzey R. 11751 (P, YA). Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On Djuma-Djuma trail, 4.33 ° N, 10.24 ° E, 14 February 2014, Couvreur T. L. P. 621 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 20 June 2014, Couvreur T. L. P. 657 (WAG, YA). South Region: Ebolowa, 2.96 ° N, 11.28 ° E, 01 January 1925, Bates G. L. 1959 (BM, BR, MO); on road Lolodorf-Bipindi ca half way near Mbiguiligui village (Mbikiliki), 3.16 ° N, 10.53 ° E, 26 February 2018, Couvreur T. L. P. 1153 (P, WAG, YA); 22 km east from Lele village, 3.26 ° N, 10.10 ° E, 07 September 2013, Couvreur T. L. P. 469 (WAG, YA); ca 15 km east from Lele village, 2.26 ° N, 13.29 ° E, 09 September 2013, Couvreur T. L. P. 492 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.33 ° E, 11 February 2015, Couvreur T. L. P. 671 (WAG, YA); A 6 km a l'ouest de Masea (village situe a 50 km au SSW de Yokadouma), 3.14 ° N, 14.86 ° E, 05 July 1963, Letouzey R. 5412 (P, YA); Campo-Ma'an area road Nko-elon-Mvini Akok Beryat rock, 2.36 ° N, 10.25 ° E, 30 June 2001, van Andel T. R. 3784 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1909, Zenker G. A. 3839 (BM, BR, K, MO, P). South-West Region: Ekundu Kundu, 5.15 ° N, 8.883 ° E, 30 April 1996, Cheek M. 8297 (K, WAG, YA); Mungo river forest reserve North of Kumba-Tombel road entered ca 05 km West of Mungo bridge, 4.73 ° N, 9.55 ° E, 24 October 1998, Cheek M. 9354 (YA); Foot of Nyale Rock, 4.98 ° N, 9.616 ° E, 17 November 1998, Cheek M. 9654 (K, YA); on trail through palm oil plantation 3 km before lava flow and Seme Beach hotel when coming from Limbe, 4.05 ° N, 9.076 ° E, 18 October 2013, Couvreur T. L. P. 519 (WAG, YA); Kupe village to Loum State Forest, 4.73 ° N, 9.716 ° E, 30 May 1996, Etuge M. 2049 (K, WAG, YA); Nyale forest and rock, 5 ° N, 9.633 ° E, 15 February 1998, Etuge M. 4235 (K, YA); Edensueh forest, 5.25 ° N, 9.576 ° E, 30 November 2000, Etuge M. 4850 (K); Kumba-Mbonge road 500 m W of Meme River bridge between Bole and Mabonji, 4.55 ° N, 9.25 ° E, 07 July 1986, Thomas D. W. 6327 (MO); Baro village, 5.27 ° N, 9.21 ° E, 03 March 1988, Thomas D. W. 7494 (K, MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
78E08FEF25616D8471757CDF9603DA46.taxon	description	Fig. 7; Map 1 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
78E08FEF25616D8471757CDF9603DA46.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Thomson W. C 130, Dec 1863: holotype: K [K 000380204].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
78E08FEF25616D8471757CDF9603DA46.taxon	description	Description. Tree, 9 - 25 m tall, d. b. h. to 5 cm; stilt roots or buttresses absent, slash yellow. Indumentum of simple, bifid trifid, and fasciculate hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to pubescent. Leaves: petiole 2 - 9 mm long, ca. 2 mm in diameter, pubescent, slightly grooved, blade inserted on top of the petiole; blade 7 - 28 cm long, 2 - 9.5 cm wide, elliptic to obovate, apex acuminate, acumen ca. 1 cm long, base narrowly cuneate to shortly attenuate, coriaceous to papyraceous, below densely pubescent when young, sparsely pubescent when old, hairs simple, bifid or trifid hairs pointing in all directions, above glabrous when young and old, concolorous; midrib sunken or flat, above densely pubescent (pilose) to pubescent at least towards base when young, densely pubescent to pubescent at least towards base when old (rarely glabrous), below pubescent when young and old; secondary veins 8 to 12 pairs, glabrous below; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers bisexual with 6 perianth parts in 2 whorls, 1 per inflorescence; pedicel 5 - 11 mm long, ca. 1 mm in diameter, densely pubescent; in fruit 10 - 15 mm long, ca. 2 mm in diameter, densely pubescent; bracts 1 - 2, one basal and one upper towards the middle of pedicel, basal bract 4 - 8 mm long; sepals 3, valvate, free, 8 - 12 mm long, 4 - 6 mm wide, triangular, apex acute, base truncate, green, pubescent outside, glabrous inside, margins flat; petals free; outer petals absent; inner petals 3, valvate, 15 - 29 mm long, 6 - 14 mm wide, elliptic to inversely Y-shaped ridged, apex acute, base broad and concave, greenish yellow, margins flat, pubescent outside, pubescent in a small triangle at the base inside; stamens 145 to 160, in 5 to 6 rows, 2 mm long, linear; connective flattened, glabrous, yellow; staminodes absent; carpels free, 20 to 35, ovary ca. 2 mm long, stigma cylindrical, pubescent. Monocarps stipitate, stipes 6 - 20 mm long, ca. 1 mm in diameter; monocarps 3 to 27, 10 - 16 mm long, 4 - 9 mm in diameter, ellipsoid to obovoid, apex mucronate, sparsely pubescent, smooth, glossy, green turning red to black when ripe; seed 1, ca. 20 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
78E08FEF25616D8471757CDF9603DA46.taxon	distribution	Distribution. Known from Nigeria (one collection); in Cameroon known from the East, South, Center and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
78E08FEF25616D8471757CDF9603DA46.taxon	materials_examined	Specimens examined. Central Region: Nkolbisson shrubby low forest on summit of Mt Akockdoue Yaounde, 3.88 ° N, 11.45 ° E, 23 May 1970, Bos J. J. 6894 (BR, MO, P, WAG); Mont Mbam Minkon on trail 3 km from Nkol Nyada village, 3.96 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 414 (WAG, YA); Colline Akok Ndoue pres Nkolbisson 5 km WSW Yaounde, 3.88 ° N, 11.45 ° E, 23 May 1970, Farron C. 7335 (P, YA); Yaounde, 3.86 ° N, 11.51 ° E, Feburary 1895, Zenker G. A. 726 (P). Littoral Region: Foret de Ye Youme, 3.48 ° N, 12.3 ° E, 01 June 1917, Chevalier A. J. B. 33132 (P). South-West Region: Bambuko FR, 4.43 ° N, 9.116 ° E, 16 September 1951, Olorunfemi J. 30760 (K); Korup National Park, 5.28 ° N, 9.083 ° E, 03 April 1988, Thomas D. W. 7555 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
14A4DD3E8D685ACB5DB458937537E346.taxon	description	Fig. 6; Map 1 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
14A4DD3E8D685ACB5DB458937537E346.taxon	materials_examined	Type. Gabon. Ogooue-Lolo, Ikembele, Le Testu G. M. P. C. 8432, Oct 1930: lectotype, here designated: P [P 00267987]; isolectotypes: BM [BM 000547036]; BR [BR 0000006418700]; P [P 00362651, P 02005895, P 02005896].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
14A4DD3E8D685ACB5DB458937537E346.taxon	description	Description. Tree, 2 - 8 m tall, d. b. h. unknown; stilt roots or buttresses absent, slash yellow. Indumentum of simple, bifid, fasciculate or stellate hairs; old leafless branches glabrous, young foliate branches tomentose. Leaves: petiole 3 - 8 mm long, 1 - 2 mm in diameter, tomentose to sparsely pubescent, cylindrical, blade inserted on top of the petiole; blade 10 - 29.5 cm long, 3.5 - 10.5 cm wide, elliptic to obovate, apex acuminate to mucronate, acumen 1 - 2 cm long, base cuneate to rounded to acuminate, subcoriaceous, above glabrous when young and old, below pubescent when young and old, hairs simple, bifid and stellate pointing in all directions, concolorous; midrib sunken or flat, above glabrous when young and old, below pubescent when young and old; secondary veins 9 to 13 pairs, glabrous below; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 6 perianth parts in 2 whorls, 1 per inflorescence; pedicel 4 - 14 mm long, ca. 2 mm in diameter, sparsely pubescent; in fruit ca. 10 mm long, ca. 2 mm in diameter, pubescent; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 2 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, free, 5 - 9 mm long, 3 - 4 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free; outer petals absent; inner petals 3, valvate, 12 - 26 mm long, 7 - 12 mm wide, ovate to inversely Y-shaped ridged, apex acute, base broad and concave, yellow-green, margins flat, pubescent outside, pubescent towards margins inside; stamens 60 to 125, in 5 to 6 rows, ca. 2 mm long, linear; connective flattened, glabrous; staminodes absent; carpels free, 20 to 35, ovary ca. 3 mm long, stigma lobed, pubescent. Monocarps stipitate, stipes 8 - 19 mm long, ca. 1 mm in diameter; monocarps 8 to 20, 19 - 25 mm long, 10 - 14 mm in diameter, ellipsoid, apex mucronate, glabrous, smooth, glossy, green turning red to black when ripe; seeds ca. 20 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
14A4DD3E8D685ACB5DB458937537E346.taxon	distribution	Distribution. From Cameroon to Gabon, and one collection in northern Republic of Congo; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
14A4DD3E8D685ACB5DB458937537E346.taxon	materials_examined	Specimens examined. South Region: 15 km from Kribi Lolodorf road, 3.00 ° N, 10.02 ° E, 01 July 1969, Bos J. J. 4962 (MO, WAG); Mendoum, 2.22 ° N, 11.23 ° E, 13 February 1965, Raynal J. 13392 (P); Campo-Ma'an area Bifa, 2.67 ° N, 10.28 ° E, 13 October 2001, Tchouto Mbatchou G. P. BIFAX _ 150 (WAG); Campo-Ma'an area 2.73 ° N, 9.873 ° E, 16 August 2001, van Andel T. R. 3882 (WAG); Campo-Ma'an area near Boussebeliga creek bridge, 2.37 ° N, 9.822 ° E, 26 October 2001, van Andel T. R. 4216 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
13F5DC48F1F4EACCDC98C83030BBB9A8.taxon	description	Fig. 5; Map 1 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
13F5DC48F1F4EACCDC98C83030BBB9A8.taxon	materials_examined	Type. Sierra Leone. no region; no locality, Afzelius A. s. n., no date: holotype: B [B 10 0068937]; isotype: BM [BM 000547035].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
13F5DC48F1F4EACCDC98C83030BBB9A8.taxon	description	Description. Tree, 2 - 20 m tall, d. b. h. unknown; stilt roots or buttresses absent, slash yellow. Indumentum of simple, bifid, fasciculate or star hairs; old leafless branches glabrous, young foliate branches densely pubescent to tomentose. Leaves: petiole 3 - 8 mm long, 2 mm in diameter, densely pubescent to tomentose, grooved, blade inserted on the side of the petiole; blade 5 - 27 cm long, 2 - 8 cm wide, elliptic to obovate, apex acuminate, acumen 1 - 2 cm long, base cuneate to acuminate, coriaceous, below pubescent when young and old, hairs mainly bifid or stellate but some simple too, pointing in all directions, above pubescent when young and old, concolorous; midrib sunken or flat, above pubescent towards base when young, pubescent towards base when old, below pubescent when young, sparsely pubescent when old; secondary veins 8 to 13 pairs, pubescent below; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 6 perianth parts in 2 whorls, 1 per inflorescence; pedicel 9 - 19 mm long, ca. 2 mm in diameter, pubescent; in fruit 20 mm long, 4 - 5 mm in diameter, pubescent; bracts 2, one basal and one upper towards the middle of pedicel, basal bract 8 mm long, 4 mm wide; sepals 3, valvate, free, 9 - 22 mm long, 4 - 5 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free; outer petals absent; inner petals 3, valvate, 23 - 34 mm long, 8 - 19 mm wide, elliptic to inversely Y-shaped ridged, apex acute, base broad and concave, claw mm long, yellow, margins wavy, densely pubescent outside, glabrous inside; stamens 90 to 200, in 5 to 6 rows, 3 - 4 mm long, linear; connective flattened, glabrous; staminodes absent; carpels free, ca. 70, ca. ovary 3 mm long, stigma lobed, sparsely pubescent. Monocarps stipitate, stipes 19 - 59 mm long, 1 - 2 mm in diameter, monocarps 5 to 55, 18 - 23 mm long, 8 - 12 mm in diameter, obovoid, apex mucronate, sparsely pubescent, smooth, glossy, green turning red to black when ripe; seed 1, ca. 20 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
13F5DC48F1F4EACCDC98C83030BBB9A8.taxon	distribution	Distribution. A mainly West African species, from Sierra Leone to Cameroon; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
13F5DC48F1F4EACCDC98C83030BBB9A8.taxon	materials_examined	Specimens examined. South Region: Bipindi, 3.26 ° N, 10.20 ° E, 09 June 1928, Annet E. 174 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50245D2F19329D5C7F678F3C24262504.taxon	description	Description. Trees, 1 - 10 m tall, d. b. h. 2 - 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 7 - 20 mm long, 1 - 3 mm in diameter, blade 6 - 25 cm long, 4 - 19 cm wide, broadly obovate or obovate to broadly elliptic to elliptic, apex rounded or obtuse or shortly emarginated, base subcordate to rounded, discolorous, whitish below or concolorous; midrib sunken or flat; secondary veins 7 to 16 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; flowering peduncle sometimes present, short; pedicel 10 - 25 mm long; in fruit 15 - 50 mm long; bracts 2, all basal, 1 - 5 mm long; sepals 3, valvate, free, 3 - 4 mm long, triangular to ovate, apex acute, base truncate; petals free; outer petals longer than inner; outer petals 3, valvate, 10 - 15 mm long, 8 - 10 mm wide, ovate, apex acute, base truncate; inner petals 3, valvate, 8 - 10 mm long, 3 - 4 mm wide, narrowly oblong or narrowly elliptic, apex acute to obtuse, base truncate; stamens numerous (not counted), in 2 to 3 rows, 2 - 3 mm long, linear; connective discoid, shortly pubescent; staminodes absent; carpels free, numerous (not counted), ovary 1 - 2 mm long, stigma capitate, glabrous or pubescent. Fruit pseudosyncarpous, 20 - 50 mm long, 20 - 50 mm in diameter, obovoid to globose; monocarps sessile, completely fused between them, numerous (not counted); seed 1, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid, irregular in shape; aril absent. A mainly South American genus, one of the largest in Annonaceae with about 170 accepted species (Rainer 2001). In Africa, there are between three or four native species, with numerous subspecies and varieties and of which the taxonomy remains complicated (Robyns and Ghesquiere 1934; Sillans 1952; Le Thomas 1969 c). Annona glabra L. is probably of South American origin (Le Thomas 1969 b, 1969 c) but is naturalized along the coast of West and Central Africa. In Cameroon it is also found in mangrove areas, but is little collected (e. g. van der Burgt 130 (WAG )) We thus include it in the key, but do not provide a description. In addition, this genus contains the non-native edible species Annona squamosa L., A. muricata L. and A. reticulata L. (from South America), all of which can be found in cultivation (not included in the descriptions) in Cameroon.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
265E6A5E9A92D5FA0D6D436EDF0B572D.taxon	description	Fig. 8; Map 1 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
265E6A5E9A92D5FA0D6D436EDF0B572D.taxon	materials_examined	Type. Republic of the Congo. Pool; Bord de la M'Bote, Bouquet, A. 513, 12 Sep 1964: holotype: P [P 00363247].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
265E6A5E9A92D5FA0D6D436EDF0B572D.taxon	description	Description. Tree to shrub, 1 - 6 (8) m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches brown tomentose. Leaves: petiole 7 - 20 mm long, 2 - 3 mm in diameter, brown tomentose, grooved, blade inserted on top of the petiole; blade 6 - 20 cm long, 5 - 12 cm wide, obovate to elliptic, apex rounded or obtuse or shortly emarginate, base rounded to subcordate, papyraceous to coriaceous, below densely pubescent, curly hairs covering the whole leaf blade when young and old, above sparsely pubescent to glabrous when young, glabrous when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below densely pubescent when young and old; secondary veins 8 to 15 pairs, glabrous above; tertiary venation percurrent but also appearing reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 10 - 25 mm long, 1 - 2 mm in diameter, brown tomentose; in fruit 15 - 30 mm long, 3 - 4 mm in diameter, pubescent; bracts 2, all basal, 2 - 5 mm long, 2 - 3 mm wide; sepals 3, valvate, free, 3 - 4 mm long, 3 - 4 mm wide, triangular to ovate, apex acute, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, inner smaller than outer; outer petals 3, 10 - 15 mm long, 8 - 10 mm wide, ovate, apex acute, base truncate, yellow to green, margins flat, tomentose outside, glabrous inside; inner petals 3, valvate, 8 - 10 mm long, 3 - 4 mm wide, narrowly oblong or narrowly elliptic, apex acute, base truncate, yellow-green, margins flat, glabrous outside, glabrous inside; stamens numerous, rows not counted, 2 - 3 mm long, linear; connective discoid, shortly pubescent; staminodes absent; carpels free, numerous, ovary 1 - 2 mm long, stigma capitate, pubescent. Fruit pseudosyncarpous, 20 - 50 mm long, 20 - 50 mm in diameter, obovoid to globose, yellow orange at maturity; individual monocarps 20 to 30, sessile, completely fused between them; apex shortly pyramidal, brown tomentose, smooth, yellow to orange when ripe; seed 1, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
265E6A5E9A92D5FA0D6D436EDF0B572D.taxon	distribution	Distribution. A west and central African subspecies distributed from Guinea to Ivory Coast and from Cameroon to the Democratic Republic of the Congo and the Central African Republic; in Cameroon known from Adamaoua, Central, East, North, North-West, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
265E6A5E9A92D5FA0D6D436EDF0B572D.taxon	materials_examined	Specimens examined. Adamaoua Region: Meiganga, 6.52 ° N, 14.3 ° E, 06 February 1946, Aubreville A. 729 (P); Tignere, 7.37 ° N, 12.65 ° E, 01 March 1939, Jacques-Felix H. 3423 (P); Ngaoundere, 7.32 ° N, 13.58 ° E, 01 June 1939, Jacques-Felix H. 4012 (P); 11 km de Ngaoundere vers Ngakha, 7.32 ° N, 13.58 ° E, 06 March 1958, Letouzey R. 606 (P). Central Region: Mont Ngolep massif Ngoro 38 km N de Bafia, 5.09 ° N, 11.26 ° E, 21 April 1975, Ngameni B. K. 51 (P); Mont Ngoro a 58 km SW de Linte, 5.09 ° N, 11.26 ° E, 17 April 1982, Nkongmeneck B. A. 256 (P). East Region: Betare Oya, 5.5 ° N, 14.1 ° E, 02 March 1961, Breteler F. J. 1185 (P); Piste Moyenam Riviere Konbo, 4.58 ° N, 13.68 ° E, 29 February 1960, Letouzey R. 3187 (P); Piste Moyenam-riviere Koubou, 4.58 ° N, 13.68 ° E, 29 February 1960, Letouzey R. 3191 (P). North Region: Garoua, 9.3 ° N, 13.4 ° E, 10 February 1946, Aubreville A. 787 (P); Garoua, 9.3 ° N, 13.4 ° E, 11 February 1946, Aubreville A. 804 (P); 17 km N of Banyo along road to Mba, 6.91 ° N, 11.8 ° E, 29 February 1972, Leeuwenberg A. J. M. 9440 (WAG). West Region: Dschang, 5.45 ° N, 9.95 ° E, 13 April 1966, CNAD 317 (P); Nkounden, 5.7 ° N, 10.67 ° E, 01 May 1967, CNAD 808 (P); Bangwa ca. 15 km NW of Bagante, 5.2 ° N, 10.48 ° E, 30 April 1964, de Wilde W. J. J. O 2359 (P); Between Bangwa and Bangangte ca 8 km NW of Bangangte, 5.16 ° N, 10.5 ° E, 12 May 1964, de Wilde W. J. J. O 2589 (P, WAG); Batchingou, 5.13 ° N, 10.4 ° E, 01 January 1939, Jacques-Felix H. 3026 (P); Kontchankap, 5.58 ° N, 10.80 ° E, 01 February 1939, Jacques-Felix H. 3033 (P); Foumban, 5.72 ° N, 10.92 ° E, 01 February 1939, Jacques-Felix H. 3136 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DA07D13F1B38416E37AFCD4B89E2F4DE.taxon	description	Fig. 8; Map 1 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DA07D13F1B38416E37AFCD4B89E2F4DE.taxon	materials_examined	Type. Senegal: Roussillon 69, 1798: holotype: P [P 00363244].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DA07D13F1B38416E37AFCD4B89E2F4DE.taxon	description	Description. Tree to shrub, 1 - 10 m tall, d. b. h. 2 - 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches brown tomentose. Leaves: petiole 10 - 20 mm long, 1 - 2 mm in diameter, brown tomentose, grooved, blade inserted on top of the petiole; blade 7 - 25 cm long, 4 - 19 cm wide, broadly obovate to broadly elliptic, apex rounded, base subcordate, subcoriaceous to coriaceous, below densely pubescent with straight hairs to glabrescent when young and old, above sparsely pubescent to glabrous when young, glabrous when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below densely pubescent when young and old; secondary veins 7 to 16 pairs, glabrous above; tertiary venation percurrent but also appearing reticulate, dense. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 10 - 20 mm long, 1 - 2 mm in diameter, brown tomentose; in fruit 25 - 50 mm long, 3 - 4 mm in diameter, pubescent; bracts 2, all basal, 1 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, free, 3 - 4 mm long, 3 - 4 mm wide, broadly triangular to circular, apex acute, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, outer longer than inner; outer petals 3, 10 - 15 mm long, 8 - 9 mm wide, ovate, apex acute, base truncate, yellow to green, margins flat, tomentose outside, glabrous inside; inner petals 3, valvate, 8 - 10 mm long, 3 - 4 mm wide, narrowly oblong or narrowly elliptic, apex acute, base truncate, yellow-green, margins flat, glabrous outside, glabrous inside; stamens numerous (not counted), 2 - 3 mm long, linear; connective discoid, shortly pubescent; staminodes absent; carpels free, numerous (not counted), ovary 1 - 2 mm long, stigma capitate, glabrous. Fruit pseudosyncarpous; 20 - 50 mm long, 20 - 30 mm in diameter, obovoid to globose, yellow orange at maturity, monocarps sessile, numerous, apex flat, glabrous, smooth, yellow to orange when ripe; seed 1, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DA07D13F1B38416E37AFCD4B89E2F4DE.taxon	distribution	Distribution. A west, central and east African and northern Malagasy subspecies from Senegal to Mozambique; in Cameroon known from the Adamaoua, Central, East, Far North, North, North-West, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DA07D13F1B38416E37AFCD4B89E2F4DE.taxon	materials_examined	Specimens examined. Adamaoua Region: Dodeo, 7.48 ° N, 12.07 ° E, 01 March 1939, Jacques-Felix H. 3388 (P); Bountoun Mboun mountains ca 40 km N of Ngaoundere, 7.9 ° N, 13.48 ° E, 12 April 1977, Nordal I. 929 (P). Central Region: Bibbanga, 3.72 ° N, 10.3 ° E, 09 March 1927, Hedin L. 409 (P). East Region: Bertoua-Batouri, 4.58 ° N, 13.68 ° E, 01 January 1962, Vroumsia T. 116 (P). Far-North Region: Douzeye (c Bongor), 10.1 ° N, 15.28 ° E, 08 January 1968, Achoundong G. 1385 (P); Plaine de Maroua a 5 km au NO de Maroua, 10.6 ° N, 14.28 ° E, 18 August 1964, Biholong M. 28 (P); ca 5 km W of Maroua, 10.6 ° N, 14.28 ° E, 02 September 1964, de Wilde W. J. J. O 2966 (MO); Bogo (Maroua), 10.7 ° N, 14.61 ° E, 01 May 1939, Jacques-Felix H. 3737 (P); Reserve forestiere du Mayo Louti (10 km W de Mokolo), 10.7 ° N, 13.8 ° E, 10 September 1964, Letouzey R. 6779 (P); Mora, 11.0 ° N, 14.14 ° E, 01 January 1945, Vaillant A. 15 (P). North Region: Pitoa, 9.38 ° N, 13.50 ° E, 25 March 1974, Achoundong G. 3419 (P); Garoua, 9.3 ° N, 13.4 ° E, 04 August 1955, de Wit H. C. D 7182 (WAG); Ecole de faune de Garoua, 9.3 ° N, 13.4 ° E, 09 August 2000, Dong E. 391 (P). North-West Region: Piste Munka (= Munkep) 45 km NNW Wum, 6.8 ° N, 9.97 ° E, 09 July 1975, Letouzey R. 13988 (MO). South-West Region: Ndop Plain Hillside above Courtar Ndop Amp ref No 28, 6.02 ° N, 10.49 ° E, 01 March 1962, Brunt M. A. 51 (K). West Region: Bangwa, 5.2 ° N, 10.48 ° E, 12 May 1964, de Wilde W. J. J. O 2389 (P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76F54C97EAC425FDC68E01A3AEFC94A0.taxon	description	Description. Trees, 4 - 30 m tall, d. b. h. up to 80 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 3 - 10 mm long, 2 - 5 mm in diameter, blade 20 - 50 cm long, 7 - 18 cm wide, oblong to obovate, apex rounded or abruptly acuminate, base subcordate forming two small lobes on top of the petiole, concolorous; midrib sunken or flat; secondary veins 10 to 20 pairs; tertiary venation reticulate. Individuals androdioecious or dioecious; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls. 5 to 20 or more per inflorescence; flowering peduncle long, up to 2 - 4 m, woody, hanging or semi erect; pedicel 10 - 70 mm long; in fruit 25 - 100 mm long; bracts 2 - 4, basal or inserted along the pedicel, 1 - 5 mm long; sepals 3, valvate, free, 3 - 4 mm long, triangular to ovate, apex acute, base truncate; petals free; outer petals longer than inner; outer petals 3, valvate, 10 - 15 mm long, 8 - 10 mm wide, ovate, apex acute, base truncate; inner petals 3, valvate, 8 - 10 mm long, 3 - 4 mm wide, narrowly oblong or narrowly elliptic, apex acute to obtuse, base truncate; stamens 65 to 700, in 2 to 3 rows, 2 - 3 mm long, linear; connective discoid, shortly pubescent; staminodes absent; carpels free, 180 to 260, ovary 1 - 2 mm long, stigma capitate, glabrous or pubescent. Fruit pseudosyncarpous, 20 - 50 mm long, 20 - 50 mm in diameter, obovoid to globose; monocarps sessile, completely fused, 250 to 500; seed 1, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent. A genus of trees with four species distributed in Central Africa, one widespread and common across its range (A. mannii) and three others mainly in Gabon (two endemic); in Cameroon two species, none endemic. Anonidium usambarense R. E. Fr, endemic to Tanzania, is in fact a Polyceratocarpus species (probably Polyceratocarpus scheffleri Engl. & Diels, Couvreur, pers. obs.). Finally, one study (Focho et al. 2010) reports the presence of Anonidium floribundum Pellegr. in the Mount Cameroon area. However, no herbarium collection is available to confirm this and we do not consider it present in Cameroon for now.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BEA57CEDAD8A032048B859E1D77D952C.taxon	description	Figs 9, 10; Map 1 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BEA57CEDAD8A032048B859E1D77D952C.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Mann G. 2231, 1863: lectotype, sheet here designated: K [K 000198881]; isotypes: B [B 100153003]; K [K 000198882]; P [P 00363249].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BEA57CEDAD8A032048B859E1D77D952C.taxon	description	Description. Tree, 8 - 30 m tall, d. b. h. 30 - 80 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 3 - 7 mm long, 2 - 3 mm in diameter, sparsely pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 20 - 45 cm long, 7 - 18 cm wide, obovate to oblong-elliptic, apex rounded to abruptly acuminate, acumen 2 - 3 cm long, base rounded to shortly attenuate generally covering part of the petiole, subcoriaceous to coriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 10 to 20 pairs, glabrous above; tertiary venation reticulate, indistinct. Individuals androdioecious; inflorescences cauliflorous or on old leafless branches, axillary; peduncle 0.3 - 4 m long, up to 2 - 3 cm in diameter towards the base, woody, hanging, sparsely pubescent to glabrous. Flowers with 9 perianth parts in 3 whorls, 10 to 20 per inflorescence, male and female inflorescences similar; pedicel 10 - 85 mm long, 2 - 4 mm in diameter, glabrous; in fruit 25 - 100 mm long, 10 - 20 mm in diameter, woody, glabrous; bracts 2 - 4, several basal and one upper towards the middle of pedicel, basal bracts 7 - 12 mm long, 7 - 12 mm wide; upper bract 10 - 12 mm long, 12 - 25 mm wide, clasping the pedicel; sepals 3, valvate, basally fused, 15 - 22 mm long, 17 - 20 mm wide, triangular to ovate, apex acute, base truncate, green, tomentose outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 25 - 50 mm long, 20 - 40 mm wide, elliptic to obovate, apex acute or rounded, base truncate, yellow to green, margins flat, pubescent outside, pubescent inside; inner petals 3, valvate, 12 - 35 mm long, 15 - 25 mm wide, elliptic, apex rounded, base attenuate to truncate, yellow-green, margins flat, pubescent outside, pubescent inside; stamens in male flowers: numerous on a conical receptacle, 4 - 4.5 mm long, oblong, connective truncate, sparsely pubescent, green to cream-yellow; stamens in hermaphrodite flowers: 65 to 100, in 3 - 4 rows, 2 - 2.5 mm long, oblong; connective truncate, sparsely pubescent, green to cream - yellow; staminodes absent; carpels free, 180 - 210, ovary 3 - 4 mm long, stigma capitate, glabrous. Fruit pseudosyncarpous, 250 - 500 mm long, 100 - 300 mm in diameter; individual monocarps 150 to 180, sessile, completely fused between them; apex shortly pyramidal, glabrous, smooth, glossy, yellow at maturity with white pulp; seed 1, 30 - 50 mm long, 17 - 30 mm in diameter, flattened to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BEA57CEDAD8A032048B859E1D77D952C.taxon	distribution	Distribution. A central African species, from southeastern Nigeria to the Republic of Congo, and in the Democratic Republic of the Congo; in Cameroon known from the East, South, Central, Littoral, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BEA57CEDAD8A032048B859E1D77D952C.taxon	materials_examined	Selected specimens examined. Central Region: Eastern sector of M'fou Nat Park Footpath running E of bridge SE of Ndanan 1, 3.61 ° N, 11.58 ° E, 22 October 2002, Cheek M. 11248 (K, YA); near Otele, 3.43 ° N, 11.14 ° E, 25 February 2007, Couvreur T. L. P. 106 (WAG, YA); Mont Mbam Minkon on trail 3 km from Nkol Nyada village, 3.96 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 415 (WAG, YA); Ottotomo Forest Reserve on top of small hill in front of reserve house, 3.65 ° N, 11.29 ° E, 25 June 2013, Couvreur T. L. P. 452 (WAG, YA); on trail to Oveng Lodge hotel near parking just behind the village of Oveng 30 km on road from Mbalmayo to Sangmeli 3.41 ° N, 11.70 ° E, 09 February 2014, Couvreur T. L. P. 609 (WAG, YA); Ottotomo Forest reserve 7 km north-west from Ngoumou 30 km south west from Yaounde, 3.65 ° N, 11.28 ° E, 24 February 2016, Couvreur T. L. P. 987 (WAG, YA); SSW of M'Balmayo, 3.52 ° N, 11.5 ° E, 27 February 1964, de Wilde W. J. J. O 1968 (B, B, BR, K, MO, P, WAG, YA); Mbam-Minkom Village de Nkolniada, 3.96 ° N, 11.40 ° E, 26 July 2012, Droissart V. 1416 (MO); Nanga Eboko, 4.68 ° N, 12.36 ° E, 17 February 1927, Hedin L. 36 (P); Mbam Minkom, 3.96 ° N, 11.36 ° E, 19 September 2013, Kamdem N. 142 (YA); Nkila, 4.68 ° N, 12.37 ° E, 12 March 1959, Letouzey R. 1547 (P, YA); Ekom, 3.85 ° N, 11.7 ° E, 16 February 1947, Letouzey R. 194 (P). East Region: Toungrelo, 4.33 ° N, 13.53 ° E, 09 January 1962, Breteler F. J. 2454 (K, P, WAG); 81 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.17 ° N, 14.69 ° E, 05 March 2019, Couvreur T. L. P. 1207 (MPU, WAG, YA); Somalomo, 3.32 ° N, 12.71 ° E, 18 March 2016, Kamdem N. 415 (YA); Lomie, 3.11 ° N, 13.58 ° E, 01 December 2016, Kamdem N. 454 (YA). Littoral Region: Nkam, 4.35 ° N, 10.67 ° E, 13 June 1927, Hedin L. 1337 (P). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 696 (WAG, YA); 17 km On the newly reconstructed road from Ebolowa to Minkok, 2.75 ° N, 11.25 ° E, 29 January 1975, de Wilde J. J. F. E 7930 (BR, K, MO, P, U, WAG, YA); Massif de Ngovayang village de Atog Boga, 3.25 ° N, 10.49 ° E, 05 September 2015, Droissart V. 2158 (BRLU); Essam, 4.68 ° N, 12.37 ° E, 13 February 1959, Letouzey R. 1276 (P); Essam (Nanga Eboko), 4.68 ° N, 12.37 ° E, 14 February 1959, Letouzey R. 1385 (P); Campo-Ma'an area Nsengou, 2.18 ° N, 10.58 ° E, 05 February 2001, Tchouto Mbatchou G. P. 3129 (KRIBI, WAG). South-West Region: Colline de Bokwa 42 km SE Mamfe, 5.71 ° N, 9.643 ° E, 07 December 1986, Achoundong G. 1325 (YA); Nyasoso, 4.86 ° N, 9.7 ° E, 03 June 1996, Cable S. 2801 (K, YA); Nature trail, 4.81 ° N, 9.683 ° E, 15 January 1995, Cheek M. 6009 (K, WAG); Kupe Mount, 4.82 ° N, 9.683 ° E, 20 November 1995, Cheek M. 7896 (K, WAG, YA); Nyasoso village at base of My Kupe forest reserve along nature trail, 4.82 ° N, 9.686 ° E, 04 April 2016, Couvreur T. L. P. 1053 b (WAG, YA); Cameroon Mountain, 4.08 ° N, 9.1 ° E, 29 December 1983, Thomas D. W. 2850 (MO, WAG). West Region: Pres Bandounga a 40 km au NW de Ndikinimeki, 4.98 ° N, 10.55 ° E, 12 February 1972, Letouzey R. 11202 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D05AEB35FCBAC11DC2A8AEDBACD7A31A.taxon	description	Figs 9, 10; Map 1 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D05AEB35FCBAC11DC2A8AEDBACD7A31A.taxon	materials_examined	Type. Democratic Republic of the Congo. Bas-Congo; Ganda-Sundi, de Briey J. 86, 1911: lectotype, here designated: BR [BR 8822635]; isolectotype: BR [BR 8822642].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D05AEB35FCBAC11DC2A8AEDBACD7A31A.taxon	description	Description. Tree, 15 - 25 m tall, d. b. h. up to 35 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 3 - 10 mm long, 2 - 5 mm in diameter, sparsely pubescent, soon becoming glabrous, slightly grooved, blade inserted on top of the petiole; blade 37 - 50 cm long, 10 - 16 cm wide, oblong, apex rounded to abruptly acuminate, acumen 2 - 3.5 cm long, base rounded, subcordate, subcoriaceous to coriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 14 to 16 pairs, glabrous above; tertiary venation reticulate. Individuals dioecious; inflorescences cauliflorous or on leafless branches, axillary; peduncle 50 - 115 mm long, 3 - 10 mm in diameter, woody, hanging or semi erect from the trunk, glabrous. Flowers with 9 perianth parts in 3 whorls, 1 to 10 per inflorescence, male and female inflorescences similar; pedicel 10 - 35 mm long, 2 - 4 mm in diameter, glabrous; bracts, 1 to 2 basal and one upper towards the upper of pedicel, basal bracts 5 - 10 mm long, 5 - 10 mm wide; upper bract directly under the calyx, amplexicaul, 12 - 21 mm long, 10 - 20 mm wide; sepals 3, valvate, free, 20 - 40 mm long, 14 - 18 mm wide, triangular, apex long acuminate, gradually tapering into an acute apex, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 35 - 60 mm long, 10 - 18 mm wide, narrowly elliptic to obovate, apex acute to attenuate, base attenuate, green turning yellow, margins flat, pubescent outside, pubescent inside; inner petals 3, valvate, 30 - 50 mm long, 9 - 16 mm wide, narrowly elliptic, apex acute, base truncate, green turning yellow, margins flat, pubescent outside, pubescent inside; stamens in male flowers: 500 to 700 inserted on a conical receptacle, in 14 to 17 rows, 3.5 - 5 mm long, oblong, connective truncate, sparsely pubescent, green to cream-yellow; carpels in female flower (see notes) 220 to 260, ovary 2 - 3 mm long, stigma capitate, glabrous; staminodes absent. Whole fruits not seen, label information: ca. 20 cm long, ca. 10 cm in diameter [taken from descriptions on specimens Halle & Villiers 4505, P 02032580], white [taken from Cheek 10240 P 00956188]; seed 1, 40 - 48 mm long, 17 - 23 mm wide, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D05AEB35FCBAC11DC2A8AEDBACD7A31A.taxon	distribution	Distribution. A central African species, known from Cameroon, Gabon, Republic of the Congo and Angola (Cabinda); in Cameroon known from the Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D05AEB35FCBAC11DC2A8AEDBACD7A31A.taxon	materials_examined	Specimens examined. Littoral Region: Loum Forest Reserve, 4.73 ° N, 9.731 ° E, 16 April 2005, Onana J. M. 3101 (K). South-West Region: Banga, 4.55 ° N, 9.416 ° E, 01 March 1956, Binuyo A. 35606 (FHO); Southern Bakundu FR, 4.48 ° N, 9.350 ° E, 13 March 1948, Brenan J. P. M. 9410 (K, P); Just west of Loum, 4.73 ° N, 9.729 ° E, 03 December 1999, Cheek M. 10240 (K, MO, P, WAG, YA); S Bakundu FR, 4.49 ° N, 9.374 ° E, 09 April 1951, Olorunfemi J. 30508 (K); 5 km S of Kumba on Buea / Douala Road, 4.65 ° N, 9.39 ° E, 21 June 1983, Thomas D. W. 2188 (MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
91F3F7576770ECF800A012263827E596.taxon	description	Description. Lianas, up to 30 m tall, d. b. h. up to 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs or absent. Leaves: petiole 1 - 15 mm long, 1 - 2 mm in diameter; blade 7 - 26 cm long, 2.5 - 14 cm wide, elliptic to ovate to obovate to oblong, apex acuminate to acute, base decurrent to subcordate, concolorous; midrib sunken or flat; secondary veins 7 to 16 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 90 per inflorescence; pedicel 2 - 25 mm long; in fruit 2 - 25 mm long; bracts 2, all basal, minute, soon falling; sepals 3, valvate, free, 1 - 15 mm long, triangular, apex acute, base truncate; petals free, sub equal; outer petals 3, valvate, 5 - 35 mm long, 1 - 14 mm wide, ovate to elliptic to linear to tubular, apex acute to rounded, base broad and concave; inner petals 3, valvate, 5 - 30 mm long, 1 - 9 mm wide, ovate to elliptic to linear to tubular, apex acute to rounded, base broad and concave, forming a pollination chamber over the receptacle; stamens 15 to 70, in 2 to 5 rows, 2 - 3 mm long, linear or cuneiform; connective discoid, glabrous or pubescent; staminodes absent; carpels free, 3 to 32, ovary 1 - 4 mm long, stigma bilobed or cylindrical, pubescent or glabrous. Monocarps sessile or substipitate, stipe, when present 1 - 25 mm long, 1 to 20 monocarps, 6 - 60 mm long, 5 - 25 mm in diameter, ellipsoid to obovoid, apex rounded to apiculate, smooth or verrucose; seed 1 to 2, 5 - 25 mm long, 5 - 15 mm in diameter, ellipsoid or flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2B299694A87256B14DAEC650D481C23E.taxon	description	Fig. 11; Map 2 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2B299694A87256B14DAEC650D481C23E.taxon	materials_examined	Type. Cameroon. Central Region; Yaounde, Zenker G. A. 690, 1896: holotype: B [B 10 0153007]; isotypes: BM [BM 000546848]; COI [COI 00004927]; P [P 00363375, P 00363376]; K [K 000198859, K 000198860].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2B299694A87256B14DAEC650D481C23E.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent with short appressed hairs. Leaves: petiole 3 - 5 mm long, 1 - 2 mm in diameter, pubescent with short appressed hairs to glabrous, grooved, blade inserted on the side of the petiole; blade 7.5 - 10 cm long, 2.5 - 6 cm wide, oblong to elliptic, apex acuminate, acumen 0.5 - 1 cm long, base cuneate to rounded, coriaceous to subcoriaceous, below sparsely pubescent with short appressed hairs to glabrous when young, glabrous when old, above glabrous when young and old, concolorous; midrib sunken or flat, above glabrous when young and old, below pubescent with short appressed hairs when young and old; secondary veins 10 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; hook-shaped peduncle 15 - 20 mm long, sparsely pubescent; pedicel 7 - 10 mm long, 1 - 2 mm in diameter, sparsely pubescent; in fruit 7 - 13 mm long, 2 - 3 mm in diameter, glabrous; bracts all basal, minute; sepals 3, valvate, free, 3 - 5 mm long, 2 - 3 mm wide, triangular, apex acute, base truncate, green, pubescent outside, glabrous inside, margins flat; petals free, sub equal, green turning red-orange; outer petals 3, 15 - 30 mm long, 4 - 9 mm wide, narrowly elliptic to narrowly ovate, apex attenuate, base broad and concave, white to light green, margins flat, tomentose outside, tomentose with a glabrous base inside; inner petals 3, valvate, 15 - 30 mm long, 3 - 6 mm wide, linear to narrowly elliptic, apex acute, base broad and concave, white to light green, margins flat, tomentose outside, tomentose with a glabrous base inside; stamens 15 to 20, in 2 to 3 rows, 2 - 3 mm long, cuneiform; connective discoid, glabrous; staminodes absent; carpels free, 8 to 10, ovary ca. 2 mm long, stigma cylindrical, glabrous. Monocarps sessile, 4 to 6, 15 - 20 mm long, 7 - 13 mm in diameter, ellipsoid to oblong, apex rounded, glabrous, smooth, red when ripe, not ribbed; seeds 1 to 2 per monocarp, 10 - 13 mm long, 5 - 8 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2B299694A87256B14DAEC650D481C23E.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of Congo and the Democratic Republic of the Congo; in Cameroon known from the Central, East, Littoral, South, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2B299694A87256B14DAEC650D481C23E.taxon	materials_examined	Specimens examined. Central Region: Badjob, 3.68 ° N, 10.68 ° E, 21 December 1963, de Wilde W. J. J. O 1602 (BR, MO, P, WAG, YA); Bank Nyong River near the new bridge ca 65 km SSW of Eseka, 3.46 ° N, 10.5 ° E, 17 June 1964, de Wilde W. J. J. O 2720 (WAG); Yaounde, 3.87 ° N, 11.52 ° E, 1896, Zenker G. A. 690 (B, K, P). East Region: Ebaka (Bertoua), 4.93 ° N, 13.32 ° E, 24 May 1961, Breteler F. J. 1429 (BR, K, M, P, WAG, YA); Ndo Riv (Bertoua), 4.58 ° N, 13.68 ° E, 12 December 1961, Breteler F. J. 2210 (BR, K, P, WAG, YA); Doume Riv (Batouri), 4.23 ° N, 13.45 ° E, 15 April 1962, Breteler F. J. 2799 (K, P, WAG); Goyoum, 5.22 ° N, 13.38 ° E, 29 January 1961, Breteler F. J. 968 (A, BR, K, M, P, WAG); Nguelemendouka, 4.38 ° N, 12.92 ° E, 04 April 1962, de Bruijn J. s. n. (WAG [WAG 0175010]); Rives du Dja pres Ndongo a 40 km WNW de Moloundou, 2.15 ° N, 14.86 ° E, 18 March 1973, Letouzey R. 12141 (P, WAG, YA); Berge arbustive et broussailleuse du fleuve Sanaga au Nord de Goyoum, 5.24 ° N, 13.36 ° E, 29 January 1961, Letouzey R. 3309 (P, YA); Rives du Dja entre les rivieres Meu et Edjune, 3.41 ° N, 13.33 ° E, 12 April 1961, Letouzey R. 3772 (P, YA); Rives de la Kadei entre Mindourou et Dongongo (40 km SSE de Batouri), 4.13 ° N, 14.60 ° E, 25 April 1962, Letouzey R. 4859 (P, YA); Betare Oya, 5.59 ° N, 14.08 ° E, Tisserant C. 3651 (P). Littoral Region: Manengouba mount base 4 km WNW Of Nkongsamba, 4.96 ° N, 9.883 ° E, 09 September 1971, Leeuwenberg A. J. M. 8319 (B, BR, MO, P, U, WAG, YA). South Region: Ebom, 3.1 ° N, 10.73 ° E, 13 August 1996, Elad M. 510 (WAG); Nyabesan, 2.4 ° N, 10.4 ° E, 05 March 1963, Raynal J. 10240 (P, YA). South-West Region: Likombe, 4.11 ° N, 9.183 ° E, 19 February 1995, Cable S. 1309 (K, MO, WAG, YA); Likombe, 4.11 ° N, 9.183 ° E, 19 February 1995, Cable S. 1310 (K, WAG, YA); Road to NLO Mt from Kodmin, 5 ° N, 9.683 ° E, 23 January 1998, Cheek M. 9063 (K, YA); Nyasoso, 4.81 ° N, 9.683 ° E, 08 February 1995, Elad M. 132 (K, YA); Mt Cameroon south slope Transect 8, 4.07 ° N, 9.015 ° E, 16 November 1985, Gentry A. H. 52942 (MO, P); Nzee Mbeng trail from Ngomin to Nzee Mbeng, 5.83 ° N, 9.716 ° E, 10 February 1998, Gosline W. G. 99 (K, YA); Ndum, 4.83 ° N, 9.7 ° E, 31 January 1995, Groves M. 21 (K, MO, WAG, YA); Nyasoso, 4.81 ° N, 9.683 ° E, 08 February 1995, Groves M. 77 (K, YA); South slope of mount north of Mt Etinde Forest, 4.08 ° N, 9.133 ° E, 20 March 1988, Nemba J. 953 (MO, P). West Region: Bali Ngemba grassland and forest patches northeast of Mantum, 5.82 ° N, 10.08 ° E, 12 April 2004, Etuge M. 5373 (K, MO, P, WAG, YA); Dschang, 5.45 ° N, 9.95 ° E, 01 May 1960, Jacques-Felix H. 5211 (K, P, WAG); Ngwenfon 35 km NW de Foumban, 5.72 ° N, 10.92 ° E, 11 December 1974, Letouzey R. 13495 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F317A1C8F3C49298C44AC9F49F2F8489.taxon	description	Fig. 11; Map 2 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F317A1C8F3C49298C44AC9F49F2F8489.taxon	materials_examined	Type. Gabon. Ogooue-Lolo; Lastoursville, Le Testu G. M. P. C. 7116, Mar 1929: lectotype, sheet here designated: P [P 02034091]; isotypes: BR [BR 0000008820792, BR 0000008809940, BR 0000008809933]; IFAN [IFAN 01625]; LISC [LISC 000367]; P [P 02034088]; K [K 000198858].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F317A1C8F3C49298C44AC9F49F2F8489.taxon	description	Description. Differs from the type variety by the presence of numerous densely packed flowers (6 - 15 versus 1 - 4).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F317A1C8F3C49298C44AC9F49F2F8489.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of Congo and the Democratic Republic of the Congo, the Artabotrys multiflorus variety is known from Cameroon and Gabon; in Cameroon known from the Central, East, Littoral, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F317A1C8F3C49298C44AC9F49F2F8489.taxon	materials_examined	Specimens examined. Central Region: Pres Yaounde, 3.86 ° N, 11.45 ° E, 11 March 1981, Meijer D. 15033 (MO, WAG). West Region: Bali- Ngemba FR, 5.81 ° N, 10.08 ° E, 13 April 2002, Onana J. M. 2027 (K, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
616B5D493D76665F94FE2F9890150781.taxon	description	Fig. 12; Map 2 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
616B5D493D76665F94FE2F9890150781.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Lukolela, Dewevre A. P. 819, 31 Mar 1896: holotype: BR [BR 0000008820808]; isotype: B [B 10 0153012].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
616B5D493D76665F94FE2F9890150781.taxon	description	Description. Liana, height unknown, d. b. h. ca. 8 cm. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 10 - 15 mm long, ca. 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 8 - 26 cm long, 4 - 11 cm wide, elliptic to obovate, apex acute to acuminate, acumen 0.5 - 1 cm long, base acute to decurrent, coriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 13 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence, hook-shaped peduncle 10 - 15 mm long; pedicel 10 - 25 mm long, 2 - 3 mm in diameter, glabrous; in fruit 12 - 20 mm long, 2 - 3 mm in diameter, glabrous; bracts 1 to 2, all basal, basal bracts 2 - 3 mm long, 1 - 2 mm wide; sepals 3, valvate, free, 5 - 8 mm long, 3 - 7 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free, inner shorter than outer, green turning yellow; outer petals 3, 15 - 25 mm long, 8 - 14 mm wide, ovate, apex acute, base broad and concave, margins flat, tomentose outside, tomentose with a glabrous base inside; inner petals 3, valvate, 8 - 17 mm long, 5 - 8 mm wide, elliptic to rhombic, apex acute, base broad and concave forming a chamber over the receptacle, margins flat, tomentose outside, tomentose with a glabrous base inside; stamens numerous, number of rows unknown, 2 - 3 mm long, oblong; connective discoid, glabrous; staminodes absent; carpels free, 15 to 20, ovary 3 - 4 mm long, stigma tubular, sparsely pubescent. Monocarps stipitate, stipes 1 - 2 mm long, 1 - 3 mm in diameter, but gradually widening into seed-bearing part; monocarps 15 to 20, to 60 mm long, ca. 25 mm in diameter, obovoid, apex acute to rounded, glabrous, smooth, glossy, green when ripe; seeds 1 to 2 per monocarp, 19 - 22 mm long, 10 - 15 mm in diameter, ellipsoid, laterally flattened; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
616B5D493D76665F94FE2F9890150781.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of Congo and in the Democratic Republic of the Congo; in Cameroon known from the Central, East, Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
616B5D493D76665F94FE2F9890150781.taxon	materials_examined	Specimens examined. Central Region: near Ebolbom village 4 km est of Ngoumou 3 km north west of Otele, 3.60 ° N, 11.28 ° E, 02 May 2013, Couvreur T. L. P. 431 (MPU, WAG, YA). East Region: Sur axe Lomie-Ngoila-Souanke a 15 km au SSW de Ngola, 2.51 ° N, 13.86 ° E, 22 February 1973, Letouzey R. 12016 (P, YA). Littoral Region: Tissongo, 3.57 ° N, 9.869 ° E, 09 July 1976, McKey D. B. 111 (K). South Region: Ebolowa-Yaounde, 3.00 ° N, 10.92 ° E, 12 January 1914, Mildbraed G. W. J. 7727 (B). South-West Region: Nyasoso, 4.86 ° N, 9.7 ° E, 04 June 1996, Cable S. 2851 (K, YA); Along path from village Mt Etinde summit, 4.05 ° N, 9.15 ° E, 02 December 1993, Cable S. 332 (K); Kupe Rock saddle, 4.78 ° N, 9.716 ° E, 11 July 1996, Cable S. 3804 (K, YA); Kupe village, 4.78 ° N, 9.716 ° E, 17 July 1996, Cable S. 3894 (K, YA); Upper Boando, 4.05 ° N, 9.153 ° E, 08 December 1993, Cable S. 475 (K, YA); Mabeta Moliwe reserve 3 - 5 km east of Limbe, 4.00 ° N, 9.256 ° E, 03 July 1992, Cheek M. 3470 (P); Kupe village, 4.77 ° N, 9.688 ° E, 29 November 1999, Gosline W. G. 241 (K); Pente E Mont 6 km E Bomana 35 km NW Limbe Alt 950 m, 4.27 ° N, 9.112 ° E, 11 December 1984, Villiers J. - F. 2441 (P, YA); Saddle between Mt Etinde and Cameroon, 4.08 ° N, 9.116 ° E, 28 October 1992, Wheatley J. I. 644 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0B7F7A75C29993375610D89ED300726.taxon	description	Figs 13, 14; Map 2 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0B7F7A75C29993375610D89ED300726.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 510, 1 Jan 1914: lectotype, sheet here designated: P [P 00363381]; isotypes: B [B 10 0153019]; BR [BR 0000008820693]; P [P 00363377]; U [U 0000237]; WAG [WAG 0000084, WAG 0000085].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0B7F7A75C29993375610D89ED300726.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches pubescent, young foliate branches tomentose to densely pubescent with long, up to 2 mm, ferruginous hairs. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, densely pubescent, grooved, blade inserted on top of the petiole; blade 15 - 24 cm long, 6 - 7 cm wide, elliptic, apex acuminate to acute, acumen 0.5 - 1 cm long, base subcordate, coriaceous, below densely pubescent when young and old, above sparsely pubescent when young, glabrous when old, concolorous; midrib impressed, above glabrous or sparsely pubescent when young, glabrous when old, below densely pubescent when young and old; secondary veins 13 to 16 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences, ramiflorous on old leafless branches, generally without leaves, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 10 to 15 per inflorescence, hook-shaped peduncle 12 - 17 mm long; pedicel 3 - 4 mm long, ca. 1 mm in diameter, densely pubescent; bracts ca. 2, all basal, basal bracts 5 mm long, 4 mm wide; sepals 3, valvate, free, 5 - 12 mm long, 3 - 7 mm wide, triangular, apex acute, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 20 - 35 mm long, 6 - 10 mm wide, elliptic, apex acute, base broad and concave, margins flat, densely pubescent outside, glabrous inside; inner petals 3, valvate, 15 - 25 mm long, 6 - 9 mm wide, elliptic, apex acute, base broad and concave, margins flat, densely pubescent outside, pubescent inside; stamens 50 to 70, in 3 to 4 rows, 1 - 2 mm long, cuneiform; connective discoid, pubescent; staminodes absent; carpels free, 25 to 32, ovary 1 - 2 mm long, stigma cylindrical, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0B7F7A75C29993375610D89ED300726.taxon	distribution	Distribution. endemic to Cameroon; known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0B7F7A75C29993375610D89ED300726.taxon	materials_examined	Specimens examined. South Region: Lele, 2.29 ° N, 13.34 ° E, 06 September 2013, Couvreur T. L. P. 453 (WAG, YA); Campo-Ma'an area 2.28 ° N, 9.866 ° E, 03 October 2001, van Andel T. R. 4128 (KRIBI, U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1914, Zenker G. A. 2087 (WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1914, Zenker G. A. 510 (B, P, U, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
24F4E1ED3C67E272984C245AF2BC1C4B.taxon	description	Fig. 11; Map 2 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
24F4E1ED3C67E272984C245AF2BC1C4B.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2801, 1904: lectotype, sheet here designated: B [B 10 0153021]; isotypes: B [B 10 0153022]; COI [COI 00004928]; GOET [GOET 005674]; HBG [HBG 502547]; K [K 000198855]; MO [MO- 216862]; P [P 00363364]; S [S-G- 7465]; WAG [WAG 0053175]; WU [WU 0025886].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
24F4E1ED3C67E272984C245AF2BC1C4B.taxon	description	Description. Liana, up to 10 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent with short appressed hairs. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous, grooved, blade inserted on the side of the petiole; blade 10 - 20 cm long, 3.5 - 6 cm wide, ovate to elliptic, apex acuminate to acute, acumen 0.5 - 1 cm long, base acute, subcoriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above sparsely pubescent to glabrous when young, glabrous when old, below sparsely pubescent to pubescent when young, sparsely pubescent when old; secondary veins 9 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches. Flowers with 9 perianth parts in 3 whorls, leaf opposed, 1 to 3 per inflorescence, hook-shaped peduncle 16 - 25 mm long; pedicel 2 - 5 mm long, 1 - 2 mm in diameter, sparsely pubescent with short appressed hairs; in fruit 2 - 3 mm long, ca. 2 mm in diameter, glabrous; bracts 2 (?), all basal, basal bracts not seen; sepals 3, valvate, free, 10 - 15 mm long, 5 - 8 mm wide, triangular, apex acute, base truncate, green turning light reddish, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 30 - 35 mm long, 7 - 12 mm wide, elliptic, apex acute, base attenuate (rounded), green, margins flat but recurved outwards in vivo, pubescent to densely pubescent outside, pubescent to sparsely pubescent inside; inner petals 3, valvate, 15 - 25 mm long, 2 - 6 mm wide, elliptic to oblong, apex acute, base broad and concave, margins flat, but recurved outwards in vivo, densely pubescent to pubescent outside, pubescent to sparsely pubescent inside; stamens numerous, number of rows not seen, 2 mm long, cuneiform; connective discoid, glabrous; staminodes absent; carpels free, 12 to 17, ovary 3 - 4 mm long, stigma cylindrical, pubescent. Monocarps sessile, 7 to 9, 20 - 25 mm long, 10 - 12 mm in diameter, ellipsoid, apex long apiculate and slightly curved, glabrous, smooth, not ribbed; red when ripe, seeds 2 per monocarp, 8 - 11 mm long, 4 - 6 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
24F4E1ED3C67E272984C245AF2BC1C4B.taxon	distribution	Distribution. A west and central African species, from Sierra Leone to Benin and from Cameroon to the Democratic Republic of the Congo; in Cameroon known from Central, East, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
24F4E1ED3C67E272984C245AF2BC1C4B.taxon	materials_examined	Specimens examined. Central Region: Route Ndanan I-Ndangan I, 3.62 ° N, 11.58 ° E, 10 March 2004, Cheek M. 11641 (K, YA); Yangafok II 25 km ENE de Bafia, 4.93 ° N, 11.37 ° E, 26 November 1969, Letouzey R. 9607 (P, YA). East Region: 82 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.73 ° E, 06 March 2019, Couvreur T. L. P. 1214 (MPU, WAG, YA). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.46 ° N, 10.35 ° E, 14 February 2015, Couvreur T. L. P. 710 (WAG, YA); Ebom, 3.1 ° N, 10.73 ° E, 25 February 1997, Elad M. 580 (KRIBI, WAG); Ebom, 3.1 ° N, 10.71 ° E, 26 February 1997, Parren M. P. E. 4 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1904, Zenker G. A. 2801 (B, BR, K, L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1907, Zenker G. A. 3320 (P). South-West Region: Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.096 ° E, 03 April 2016, Couvreur T. L. P. 1044 (MPU, WAG, YA); Korup National Park, 5.28 ° N, 9.083 ° E, 03 April 1988, Thomas D. W. 7578 (MO, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0FAAE4B79F3DE80B1DCEDC1E666159B8.taxon	description	Fig. 14; Map 2 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0FAAE4B79F3DE80B1DCEDC1E666159B8.taxon	materials_examined	Type. Cameroon. East Region; Bitya, near Dja river, Bates G. L. 1792, Sep 1922: holotype: P [00363370].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0FAAE4B79F3DE80B1DCEDC1E666159B8.taxon	description	Description. Differs from the type variety by its densely brown tomentose and shortly hirsute branches and petioles, pubescent lower side of leaf blades and tomentose petals.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0FAAE4B79F3DE80B1DCEDC1E666159B8.taxon	distribution	Distribution. A west and central African species, from Sierra Leone to Benin and from Cameroon to the Democratic Republic of the Congo; in Cameroon known from Central, East, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0FAAE4B79F3DE80B1DCEDC1E666159B8.taxon	materials_examined	Specimens examined. East Region: Village Djang 40 km west of Bertoua, 4.58 ° N, 13.35 ° E, 15 May 1962, Breteler F. J. 2956 (P, WAG). South Region: Bitye near R Ja, 3.02 ° N, 12.37 ° E, 01 September 1922, Bates G. L. 1792 (P); Rive du Ntem a Ebianemeyong 60 km east de Campo, 2.42 ° N, 10.33 ° E, 12 April 1970, Letouzey R. 10370 (P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5D213C87FEC1E9AF33EEF01A833E995.taxon	description	Fig. 15; Map 2 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5D213C87FEC1E9AF33EEF01A833E995.taxon	materials_examined	Type. Cameroon. Central Region; Nkidi forest, Jacques-Felix H. 2490, Nov 1938: lectotype, sheet here designated: P [P 00363361]; isotypes: K; P [P 00363359, P 00363362].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5D213C87FEC1E9AF33EEF01A833E995.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 4 - 7 mm long, ca. 1 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 8 - 13 cm long, 3 - 14 cm wide, oblong to elliptic, apex acute, acumen ca. 0.5 cm long, base decurrent to acute, coriaceous, below glabrous when young and old, above glabrous when young and old, shiny when dried, concolorous; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 12 pairs, glabrous above; tertiary venation reticulate. Flowers bisexual with 9 perianth parts in 3 whorls. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 3 to 6 per inflorescence, hook-shaped peduncle 15 - 20 mm long; pedicel 10 - 25 mm long, ca. 1 mm in diameter, glabrous; in fruit 20 - 30 mm long, 1 - 2 mm in diameter, glabrous; bracts 1 to 2, all basal, basal bracts ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 1 - 2 mm long, ca. 1 mm wide, triangular, apex acute, base truncate, glabrous outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 20 - 35 mm long, 1 - 2 mm wide, linear to narrowly ovate, apex rounded, base broad and concave, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 15 - 25 mm long, 1 - 3 mm wide, linear, apex rounded, base broad and concave, margins flat, densely pubescent outside, glabrous inside; stamens 50 to 70, in 3 to 4 rows, 2 - 3 mm long, oblong; connective discoid, pubescent; staminodes absent; carpels free, 3 to 4, ovary 3 - 4 mm long, stigma cylindrical, glabrous. Monocarps stipitate, stipes 5 - 10 mm, ca. 6 mm in diameter, 1 to 4 monocarps, 20 - 40 mm long, 10 - 20 mm in diameter, ellipsoid, apex rounded, glabrous, warty to verrucose, faintly ribbed, color when ripe unknown; seeds 1 to 2 per monocarp, 20 - 25 mm long, 10 - 15 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5D213C87FEC1E9AF33EEF01A833E995.taxon	distribution	Distribution. A central African species, only known from Cameroon and the Democratic Republic of the Congo; in Cameroon known from the Central and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C5D213C87FEC1E9AF33EEF01A833E995.taxon	materials_examined	Specimens examined. South Region: River Ja Bitya, 3.02 ° N, 12.37 ° E, 01 January 1922, Bates G. L. 1699 (K); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1909, Zenker G. A. 3834 (L, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
385642FC49D9EDCB2358504D2AD7F077.taxon	description	Figs 16, 17; Map 2 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
385642FC49D9EDCB2358504D2AD7F077.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Likimi, Malchair L. 274, 20 Avr 1910: holotype: BR [BR 0000008820297]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
385642FC49D9EDCB2358504D2AD7F077.taxon	description	Description. Liana, to 20 m tall, d. b. h. 3 - 5 cm. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches hirsute, with erect hairs. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, hirsute, slightly grooved, blade inserted on top of the petiole; blade 8 - 14 cm long, 3.5 - 5.5 cm wide, oblong to obovate, apex acuminate, acumen 0.5 - 1 cm long, base rounded to subcordate or obtuse, papyraceous, below pubescent when young and old with long appressed hairs, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below pubescent when young and old; secondary veins 9 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence, hook-shaped peduncle 4 - 7 mm long; pedicel 3 - 5 mm long, ca. 1 mm in diameter, pubescent to sparsely hirsute; in fruit 3 - 15 mm long, ca. 2 mm in diameter, pubescent; bracts caduceus, not seen; sepals 3, valvate, free, 5 - 7 mm long, 2 - 4 mm wide, triangular, apex acute, base truncate, green, hirsute outside, glabrous inside, margins flat; petals free, sub equal, green turning yellow; outer petals 3, 10 - 20 mm long, 3 - 4 mm wide, very narrowly elliptic to linear, apex acute, base rounded to broad and concave, green turning bright yellow, margins flat, recurved inwards in vivo, densely appressed-pubescent outside, appressed-pubescent to glabrous inside; inner petals 3, valvate, 12 - 20 mm long, 2 - 3 mm wide, very narrowly elliptic to linear, apex acuminate to acute, base broad and concave, green turning bright yellow, margins flat, recurved inwards in vivo, appressed-pubescent outside, appressed-pubescent inside; stamens 60 to 70, in ca. 5 rows, ca. 1 mm long, broad; connective discoid, glabrous, green; staminodes absent; carpels free, 8 to 10, ovary ca. 1 mm long, stigma capitate, glabrous. Monocarps sessile, 5 to 11, 12 - 15 mm long, 5 - 6 mm in diameter, ellipsoid to fusiform, apex apiculate, glabrous, smooth, not ribbed, red when ripe; seeds 1 to 2 per monocarp, 10 - 12 mm long, 5 mm in diameter, ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
385642FC49D9EDCB2358504D2AD7F077.taxon	distribution	Distribution. A mainly central African species, from Benin to Nigeria and Cameroon to the Republic of the Congo and in the Democratic Republic of the Congo; in Cameroon known from the Central and East regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
385642FC49D9EDCB2358504D2AD7F077.taxon	materials_examined	Specimens examined. Central Region: Febe Mount, 3.91 ° N, 11.48 ° E, 30 March 1962, Breteler F. J. 2717 (A, BR, K, P, WAG, YA); Mont Mbam Minkon on trail 5 km from Nkol Nyada village On top of small hill, 3.97 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 418 (MPU, WAG, YA); Nachtigal, 4.35 ° N, 11.63 ° E, 01 July 1964, de Wilde W. J. J. O 2779 (P, WAG, YA); Nkolbison, 3.88 ° N, 11.45 ° E, 02 November 1964, de Wilde W. J. J. O 3715 (BR, K, P, WAG); Left bank Sanaga river near Ferry Nachtigal ca 20 km N of Obala, 4.34 ° N, 11.64 ° E, 29 April 1965, Leeuwenberg A. J. M. 6011 (B, BR, C, K, MO, P, WAG, YA). East Region: Dimako, 4.38 ° N, 13.57 ° E, 02 August 1961, Breteler F. J. 1752 (P, YA); Bertoua 15 km along road to Deng Deng, 4.66 ° N, 13.63 ° E, 31 August 1961, Breteler F. J. 1825 (WAG); Bamekok (Batouri), 4.2 ° N, 14.15 ° E, 16 April 1962, Breteler F. J. 2825 (P, WAG); 60 km south of Yokadouma 5 km south of Masea village, 3.10 ° N, 14.84 ° E, 06 March 2019, Couvreur T. L. P. 1211 (MPU, WAG, YA); A 25 km au NE de Bange km 75 route Yokadouma-Moloundou, 3.02 ° N, 15.12 ° E, 25 May 1963, Letouzey R. 5147 (P, YA); Mbatika-Malen 20 km de Moloundou route Yokadouma 2.03 ° N, 15.22 ° E, 22 April 1971, Mezili P. 193 (P, YA); Moloundou near Lokomo Bumba and Bange, 2.08 ° N, 15.25 ° E, 26 January 1911, Mildbraed G. W. J. 4350 (B, HBG); Foret inhabitee entre Yokaduma et Assobam, 3.52 ° N, 15.05 ° E, 24 April 1911, Mildbraed G. W. J. 4999 (B, HBG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B61800D4E462234D05FF7D0D133D428E.taxon	description	Figs 17, 18; Map 2 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B61800D4E462234D05FF7D0D133D428E.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Thomson W. C 25, Feb 1863: holotype: K [K 000198871].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B61800D4E462234D05FF7D0D133D428E.taxon	description	Description. Liana, 30 m tall, d. b. h. 10 - 20 cm. Indumentum of simple hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches sparsely pubescent. Leaves: petiole 1 - 10 mm long, ca. 2 mm in diameter, pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 7 - 20 cm long, 5 - 10 cm wide, elliptic to oblong, apex acute to abruptly acuminate, acumen 0.5 - 1 cm long, base rounded to obtuse, coriaceous, below sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above densely pubescent to pubescent when young and old, below sparsely pubescent to glabrous when young and old; secondary veins 7 to 14 pairs, glabrous above; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 30 to 90 per inflorescence, hook-shaped peduncle 20 - 35 mm long; pedicel 10 - 20 mm long, ca. 1 mm in diameter, pubescent; in fruit 10 - 25 mm long, ca. 2 mm in diameter, pubescent; bracts several, basal with one towards the lower half of pedicel, soon falling, basal bracts ca. 1 mm long, ca. 1 mm wide; upper bract ca. 2 mm long, ca. 2 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 2 - 3 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 10 - 20 mm long, 2 - 3 mm wide, elliptic to narrowly elliptic, apex rounded to obtuse, base broad and concave, margins flat, not folded inwards, pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 12 - 16 mm long, 2 - 3 mm wide, narrowly elliptic, apex acute, base broad and concave, margins flat, not folded inwards, pubescent outside, pubescent inside; stamens 30 to 35, in 2 to 3 rows, ca. 1 mm long, cuneiform; connective discoid, glabrous; staminodes absent; carpels free, 4 to 10, ovary ca. 1 mm long, stigma coiled, sparsely pubescent. Monocarps stipitate, stipes 10 - 25 mm long, 2 - 3 mm in diameter; monocarps 1 to 7, 15 - 25 mm long, 12 - 15 mm in diameter, ellipsoid to obovoid, apex rounded, glabrous, smooth, not ribbed, green when ripe; seeds 1 (to 2) per monocarp, 10 - 15 mm long, 8 - 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B61800D4E462234D05FF7D0D133D428E.taxon	distribution	Distribution. A central African species; from Nigeria to Angola (Cabinda) and in the Democratic Republic of the Congo; in Cameroon known from the Central, East, Littoral, South, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B61800D4E462234D05FF7D0D133D428E.taxon	materials_examined	Specimens examined. Central Region: Yaounde, 3.87 ° N, 11.52 ° E, 1896, Zenker G. A. 697 (P). East Region: Near Dimako 28 km SW of Bertoua, 4.38 ° N, 13.57 ° E, 01 August 1961, Breteler F. J. 1725 (BR, K, P, U, WAG, YA); Bertoua 6 km along road to Batouri and Betare-Oya, 4.58 ° N, 13.68 ° E, 30 August 1961, Breteler F. J. 1797 (U, WAG, YA); 67 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.08 ° N, 14.67 ° E, 08 March 2019, Couvreur T. L. P. 1230 (MPU, WAG, YA); Palisco forest consession 15 km along main road into consession, 3.52 ° N, 13.54 ° E, 27 March 2015, Couvreur T. L. P. 751 (WAG, YA); 20 km environ a l'ENE de Mikel village situe a 85 km au N de Moloundou sur la route de Yokadouma 2.81 ° N, 15.24 ° E, 23 February 1971, Letouzey R. 10413 (K, P, YA); Ndongo (Dja-Molundou), 2.58 ° N, 15.29 ° E, 18 March 1973, Letouzey R. 12240 (K, P, YA); Djouo (Somalomo), 3.32 ° N, 12.93 ° E, 26 February 1962, Letouzey R. 4435 (K, P, YA); A 8 km au SSW de Koso (village situe a 60 km au SSW de Batouri), 3.93 ° N, 14.17 ° E, 29 July 1963, Letouzey R. 5533 (P, YA); Entre Badekok et Mpan (50 km ENE de Lomie), 3.22 ° N, 15.02 ° E, 05 August 1963, Letouzey R. 5548 (P, YA). Littoral Region: 8 km W of Massok, 4.13 ° N, 10.47 ° E, 27 March 1965, Leeuwenberg A. J. M. 5201 (B, BR, C, GC, K, MO, P, UC, WAG, YA). South Region: Bitya near R Ja, 3.02 ° N, 12.37 ° E, 01 November 1922, Bates G. L. 1763 (P); Djoum North East Nkout Base of ridge, 2.55 ° N, 12.80 ° E, 05 December 2014, Cheek M. 17781 (K, WAG); Elephant Mont, 2.8 ° N, 10.01 ° E, 24 May 2001, van Andel T. R. 3459 (KRIBI, WAG, YA); Campo-Ma'an area 2.73 ° N, 9.873 ° E, 16 August 2001, van Andel T. R. 3872 (KRIBI, U, WAG); Nkuambe, 3.26 ° N, 10.46 ° E, 01 December 1914, Zenker G. A. 489 (P, WAG). South-West Region: Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.497 ° E, 27 March 2016, Couvreur T. L. P. 1020 (WAG, YA); Ekombe, 4.48 ° N, 10.87 ° E, 16 January 1987, Etuge M. 485 (MO, P, WAG). West Region: Pres Bandounga a 40 km au NW de Ndikinimeki, 4.98 ° N, 10.55 ° E, 12 February 1972, Letouzey R. 11200 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79E3B666E1F52556398A0A1D5A8DE667.taxon	description	Fig. 15; Map 3 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79E3B666E1F52556398A0A1D5A8DE667.taxon	materials_examined	Type. Sierre Leone: Northern Region; Falaba, Scott Elliot G. F. 5137, 5 Mar 1892: holotype: K [K 000198865]; isotype: B [B 10 0154055].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79E3B666E1F52556398A0A1D5A8DE667.taxon	description	Description. Liana, up to 10 m tall, d. b. h. up to 20 cm. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches pubescent. Leaves: petiole 3 - 4 mm long, ca. 1 mm in diameter, pubescent to glabrous, grooved, blade inserted on the side of the petiole; blade 8 - 13 cm long, 4 - 5 cm wide, ovate to elliptic, apex acuminate to acute, acumen 1 - 1.5 cm long, base decurrent to acute, papyraceous, below pubescent to sparsely pubescent when young, glabrous when old, above pubescent when young, sparsely pubescent to glabrous when old, concolorous; midrib impressed, above glabrous when young and old, below densely pubescent to pubescent when young, glabrous to pubescent when old; secondary veins 8 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 5 to 15 per inflorescence, hook-shaped peduncle 10 - 16 mm long; pedicel 2 - 10 mm long, ca. 1 mm in diameter, pubescent with appressed hairs; in fruit 10 - 20 mm long, ca. 1 mm in diameter, pubescent with appressed hairs; bracts 2, all basal, ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 2 - 3 mm wide, triangular, apex acute, base truncate, densely pubescent outside, glabrous inside, yellow and red at the base, margins flat; petals free, sub equal; outer petals 3, 5 - 15 mm long, 1 - 2 mm wide, linear, apex rounded, base broad and concave, margins flat but strongly recurved inwards forming a tube, densely pubescent outside, densely pubescent inside; inner petals 3, valvate, 5 - 15 mm long, 1 - 2 mm wide, linear, apex acute, base broad and concave, yellow and red at the base, margins flat but strongly recurved inwards forming a tube, tomentose outside, tomentose with a glabrous at base inside; stamens 15 to 22, in 2 rows, 1 - 2 mm long, oblong; connective discoid, glabrous; staminodes absent; carpels free, 7 to 12, ovary 1 - 2 mm long, stigma cylindrical, glabrous. Monocarps sessile, 4 to 9, 10 - 20 mm long, 10 mm in diameter, ellipsoid to obovoid, apex rounded, glabrous, smooth, faintly ribbed, color when ripe not seen; seeds 1 to 2 per monocarp, 5 - 7 mm long, 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79E3B666E1F52556398A0A1D5A8DE667.taxon	distribution	Distribution. A west and central African species, from Sierra Leone to Nigeria and Cameroon to the Republic of the Congo and in the Democratic Republic of the Congo; in Cameroon known from the Adamaoua, Central, East, North, South, South-West, and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79E3B666E1F52556398A0A1D5A8DE667.taxon	materials_examined	Specimens examined. Adamaoua Region: Mbibol 40 km W de Ngaoundere, 7.32 ° N, 13.58 ° E, 12 June 1977, Fotius G. 2660 (P, YA); Pres Tekel (60 km NNO de Bagodo), 6.78 ° N, 13.17 ° E, 21 July 1966, Letouzey R. 7481 (P, YA); Boko 14 km Sud-Ouest de Ngaoundere, 7.25 ° N, 13.5 ° E, 06 August 1981, van der Zon A. P. M. 1122 (WAG, YA). Central Region: Pentes orientales du mont Yangba (1473 m) pres Nyafianga (42 km NNE de Bafia), 5.13 ° N, 11.35 ° E, 09 September 1966, Letouzey R. 7826 (K, P, YA). East Region: 27 km ENE de Mopwo (village situe au km 22 route Yokadouma-Batouri), 3.67 ° N, 15.08 ° E, 06 June 1963, Letouzey R. 5248 (K, P, YA). North Region: Mango, 8.42 ° N, 13.25 ° E, 21 July 1974, Fotius G. 2144 (YA). South Region: Rives du Ntem pres du confluent de la Kye 16 km ESE d'Ambam, 2.25 ° N, 11.34 ° E, 01 February 1970, Letouzey R. 10040 (P); Bipindi, 3.08 ° N, 10.42 ° E, 3 April 1897, Zenker G. A. 1222 (B, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1913, Zenker G. A. 231 (P, U, WAG). South-West Region: Edip village to lake edip 2 - 3 km, 4.96 ° N, 9.65 ° E, 11 February 1998, Cheek M. 9143 (K, WAG, YA); Kodmin, 5 ° N, 9.7 ° E, 16 November 1998, Gosline W. G. 149 (K, YA); Bank of river Chide, 4.95 ° N, 9.72 ° E, 04 February 1998, Onana J. M. 523 (K, P, WAG, YA). West Region: Massif du Nkogam (2263 m) 25 km W de Foumban, 5.73 ° N, 10.67 ° E, 12 December 1974, Letouzey R. 13501 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
40EC7AE7ED1E6031AA9AAB536148557F.taxon	description	Description. Same as species. A genus with two species, one widespread and one restricted to northern Angola. One species in Cameroun, not endemic. A genus easily confused with Piptostigma because of the characteristic inner petals being much longer than the outer ones in both genera, a unique feature among Cameroonian Annonaceae (Ghogue et al. 2017). However, sterile, Brieya is mainly distinguished by the lower number of secondary veins (less than 20 versus generally more than 20 in Pipostigma) and discolorous leaves being whitish below (versus concolourous in Piptostigma).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2BA733C1E075141EA6F73F207520AD25.taxon	description	Figs 19, 20; Map 3 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2BA733C1E075141EA6F73F207520AD25.taxon	materials_examined	Type. Democratic Republic of the Congo. Bas-Congo; Kingamu, Ganda sumi, de Briey J. 66, 14 - 16 Oct 1911: lectotype, sheet here designated: BR [BR-S. P. 880319]; isotypes: BR [BR 0000008803252, BR 0000008803245, BR 0000008803191].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2BA733C1E075141EA6F73F207520AD25.taxon	description	Description. Tree, 10 - 25 m tall, d. b. h. 16 - 50 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 2 - 5 mm long, ca. 2 mm in diameter, pubescent, grooved, blade inserted on top of the petiole; blade 12 - 24 cm long, 6 - 8 cm wide, obovate to oblanceolate, apex acuminate to obtuse, acumen 0.5 - 0.8 cm long, base cordate to obtuse, papyraceous, below glabrous to pubescent when young, glabrous to pubescent when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below pubescent when young and old; secondary veins 11 to 17 pairs, glabrous below; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, axillary occurring on short peduncle-like bases 0 - 2 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 1 - 2 mm long, ca. 5 mm in diameter, pubescent; in fruit 15 - 25 mm long, 4 - 5 mm in diameter, glabrous; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 2 - 3 mm long, ca. 2 mm wide; upper bract ca. 1 mm long, ca. 2 mm wide; sepals 3, valvate, free, ca. 2 mm long, ca. 2 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, sepal like, 1.5 - 2 mm long, 1.5 mm wide, ovate, apex acuminate, base truncate, light green, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 38 - 108 mm long, 3 - 7 mm wide, linear, apex acute, base truncate, green, margins flat, pubescent inside, pubescent outside; stamens 30 to 40, in 4 to 5 rows, ca. 1 mm long, broad; connective discoid, glabrous, green; staminodes absent; carpels free, ca. 4, ovary ca. 2 mm long, stigma minute, densely pubescent. Monocarps sessile, 1 to 3, 42 - 46 mm long, 25 - 40 mm in diameter, ellipsoid, apex rounded, glabrous, smooth, fleshy, green when ripe; seeds 18 to 20 per monocarp, ca. 10 mm long, ca. 4 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2BA733C1E075141EA6F73F207520AD25.taxon	distribution	Distribution. From Cote d'Ivoire to Democratic Republic of the Congo and Angola; in Cameroon known from East, South, Centre, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2BA733C1E075141EA6F73F207520AD25.taxon	materials_examined	Specimens examined. Central Region: Pres Ngong (25 km NE d'Edea), 3.75 ° N, 10.98 ° E, 12 December 1973, Letouzey R. 12345 (P, YA). East Region: 68 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.08 ° N, 14.66 ° E, 08 March 2019, Couvreur T. L. P. 1231 (MPU, WAG, YA); Bateka Malen 20 km NE de Moloundou, 2.15 ° N, 15.35 ° E, 23 April 1971, Letouzey R. 10718 (P, YA); Entre Asip et Mang (60 km ENE de Lomie), 3.4 ° N, 14.17 ° E, 13 August 1963, Letouzey R. 5605 (P, YA). North-West Region: Kagwene, 6.10 ° N, 9.744 ° E, 13 June 2009, Ashworth J. 310 (K, YA). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.34 ° E, 11 February 2015, Couvreur T. L. P. 677 (WAG, YA); Abords de la Lobe a 50 km au SSE de Kribi et a 30 km a l'ENE de Campo, 2.51 ° N, 9.82 ° E, 23 March 1968, Letouzey R. 9132 (YA). South-West Region: Mudjika (Wudjika?), 4.29 ° N, 9.41 ° E, 01 January 1933, Aubreville A. 2115 (P); on trail from Ekongo village located 5 km before the entrance to Limbe 7 km on secondary road On flank of Mt Etinde 100 m in Mont Cameroon National Park, 4.07 ° N, 9.132 ° E, 16 October 2013, Couvreur T. L. P. 510 (WAG, YA); on trail from Ekongo village located 5 km before the entrance to Limbe 7 km on secondary road On flank of Mt Etinde 100 m in Mont Cameroon National Park, 4.07 ° N, 9.131 ° E, 16 October 2013, Couvreur T. L. P. 511 (WAG, YA); Rumpi mountains forest trail ca 5 km after Dikome Balue village ca 40 km north of Kumba, 4.93 ° N, 9.240 ° E, 10 January 2016, Couvreur T. L. P. 957 (WAG, YA); Kupe village Muanezum trail = Daniel Ajang's Earthwatch rented area Mt 4.76 ° N, 9.666 ° E, 28 March 1996, Etuge M. 1844 (K); Kupe village, 4.77 ° N, 9.688 ° E, 28 November 1999, Gosline W. G. 234 (K); Njonji, 4.11 ° N, 9.016 ° E, 21 April 1997, Nning J. 385 (K, YA); Cameroon Mountain, 4.12 ° N, 9.028 ° E, 20 June 2001, van Andel T. R. 3732 (U, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D9F7DD46E8A4323229CFD63AAE4C440B.taxon	description	Description. Trees, 15 - 35 m tall, d. b. h. 30 - 80 cm; stilt roots or buttresses absent, trunk white or brown. Indumentum of simple hairs when present, but species generally glabrous. Leaves: petiole 3 - 20 mm long, 1 - 3 mm in diameter, blade 4.5 - 31 cm long, 2 - 6.5 cm wide, elliptic to obovate to oblong, apex acuminate, base acute to rounded, discolorous, whitish below or concolorous; midrib sunken or flat; secondary veins 8 to 24 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary, occurring or not on a short peduncle. Flowers with 9 perianth parts in 3 whorls, 2 to 9 per inflorescence; pedicel 10 - 25 mm long; in fruit 15 - 35 mm long; bracts 2 to 3, all basal, 1 - 2 mm long; sepals 3, valvate, free, 2 - 3 mm long, triangular to ovate, apex acute, sometimes rounded, base truncate; petals free; outer petals longer than inner; outer petals 3, valvate, 7 - 20 mm long, 2 - 7 mm wide, oblong to elliptic to obovate to linear, apex acute to rounded to obtuse, base truncate; inner petals 3, imbricate, 2 - 4 mm long, 2 - 4 mm wide, ovate to suborbicular, apex acute to obtuse, base truncate; stamens 20 to 40, in 3 to 4 rows, ca. 1 mm long, broad; connective discoid, glabrous; staminodes absent; carpels free, 10 to 24, ca. 1 mm long, stigma flat to capitate, glabrous. Monocarps stipitate, stipes 1 - 50 mm long, 3 to 18 monocarps, 15 - 30 mm long, 10 - 25 mm in diameter, globose to ellipsoid to obovoid, apex rounded, smooth, bumpy or constricted around the seeds, glabrous; seeds 1 to 2, 12 - 25 mm long, 8 - 12 mm in diameter, ellipsoid; aril absent. A genus with four accepted species, two widespread, one known only from Cameroon and Gabon and one from Equatorial Guinea and possibly Cameroon. Three (four?) species in Cameroun, none endemic.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E10BB99C9795B68F15D691BD052EC39C.taxon	description	Figs 21, 23; Map 3 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E10BB99C9795B68F15D691BD052EC39C.taxon	materials_examined	Type. Gabon. Estuaire; Libreville, Klaine T. - J. 376, Apr 1896: holotype: B [B 10 0154073]; isotypes: K [K 000198885, K 000198884]; P [P 00362650, P 00362649, P 00362652]; MPU [MPU 011662].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E10BB99C9795B68F15D691BD052EC39C.taxon	description	Description. Tree, 10 - 35 m tall, d. b. h. 80 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 10 - 20 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 5 - 15 cm long, 2 - 5 cm wide, oblong to elliptic, apex acuminate, acumen 0.5 - 1.5 cm long, base decurrent to cuneate, subcoriaceous, below glabrous when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 15 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, axillary, peduncule distinct 2 - 10 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 8 per inflorescence; pedicel 10 - 18 mm long, ca. 1 mm in diameter, sparsely pubescent; in fruit 15 - 35 mm long, 2 - 3 mm in diameter, glabrous; bracts 1 to 3, all basal, basal bracts 1 - 2 mm long, 2 mm wide; sepals 3, valvate, free, ca. 2 mm long, ca. 2 mm wide, triangular to ovate, apex acute, base truncate, green, glabrous outside, glabrous inside, margins flat; petals free; outer petals 3, 10 - 15 mm long, 5 - 7 mm wide, oblong to elliptic, apex rounded, base truncate, green, margins flat, glabrous outside, glabrous inside; inner petals 3, imbricate, 2 - 4 mm long, 2 - 4 mm wide, ovate to orbicular, apex rounded, base truncate, green, margins flat, glabrous outside, glabrous inside; stamens 20 to 30, in 3 to 4 rows, ca. 1 mm long, broad; connective discoid, glabrous, green; staminodes absent; carpels free, 12 to 24, ovary ca. 1 mm long, stigma flat, glabrous. Monocarps stipitate, stipe 18 - 30 mm long, 3 - 4 mm in diameter; monocarps 3 to 8, 18 - 30 mm long, 10 - 15 mm in diameter, obovoid, apex rounded, glabrous, finely warty, not bumpy, green when ripe; seed (1 to) 2 per monocarp, 15 - 25 mm long, 10 - 12 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E10BB99C9795B68F15D691BD052EC39C.taxon	distribution	Distribution. Central Africa; from Cameroon to Democratic Republic of the Congo; in Cameroon known from South, Central, Littoral, South-West and East regions, with one collection from Adamaoua region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E10BB99C9795B68F15D691BD052EC39C.taxon	materials_examined	Specimens examined. Adamaoua Region: A 80 km au SO de Banyo-Plaine Tikar, 6.75 ° N, 11.82 ° E, 27 June 1969, Biholong M. 219 (P, YA). Central Region: Feup (Yaounde), 3.87 ° N, 11.52 ° E, 01 July 1917, Chevalier A. J. B. 33473 (P, WAG); near Ebolbom village 3 km est of Ngoumou 2 km north west of Otele, 3.59 ° N, 11.28 ° E, 02 May 2013, Couvreur T. L. P. 430 (MPU, WAG, YA); Avom, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 57 (P). East Region: Bertoua, 4.58 ° N, 13.68 ° E, 17 February 1960, Letouzey R. 3036 (P, YA); Region de Moloundou, 2.05 ° N, 15.17 ° E, 01 August 1949, SRFK 1372 (P, YA). Littoral Region: 18 km SEof Douala along road to Ndonga (= old to Edea), 4.05 ° N, 9.71 ° E, 20 August 1965, Leeuwenberg A. J. M. 6467 (B, BR, C, GC, K, L, LUAI, MO, P, UC, WAG, YA). South Region: 20 km from Kribi Lolodorf road, 3.05 ° N, 10.05 ° E, 09 June 1969, Bos J. J. 4784 (B, BR, K, LD, LM, MO, P, POZG, WAG, YA); on road between Campo and Kribi, 2.62 ° N, 9.847 ° E, 16 February 2012, Couvreur T. L. P. 389 (WAG, YA); Rocher du Loup km 36 road Kribi-Campo, 2.61 ° N, 9.85 ° E, 06 January 1983, de Kruif A. P. M. 1046 (MO, WAG, YA); Campo-Ma'an area 2.73 ° N, 9.873 ° E, 13 August 2001, van Andel T. R. 3846 (KRIBI, WAG, YA). South-West Region: Mungo River F. R., 4.78 ° N, 9.566 ° E, 02 December 1999, Cheek M. 10229 (K, MO, P, WAG, YA); Ntali, 5.25 ° N, 9.576 ° E, 01 December 2000, Etuge M. 4873 (K); Nyandong-forest above village, 4.98 ° N, 9.585 ° E, 20 March 2003, Etuge M. 4917 (K); Forest and forest relictss near Mundemba, 4.96 ° N, 8.916 ° E, 16 January 1985, Thomas D. W. 4200 (P); Near Mundemba town, 4.96 ° N, 8.916 ° E, 12 May 1986, Thomas D. W. 6121 (MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
87DEACC19B13DB1516A641F573B4B182.taxon	description	Map 3 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
87DEACC19B13DB1516A641F573B4B182.taxon	materials_examined	Type. Cameroon. South Region (?) or Equatorial Guinea. Rio Muni; Campo-Gebiet; Bebao [i?], Weg nach Olonga [Manga?], Tessmann, G. 767, 6 Jan. 1909: holotype: B [B 10 0154074].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
87DEACC19B13DB1516A641F573B4B182.taxon	description	Description. Tree, to 19 m tall, d. b. h. to 16 cm; stilt roots or buttresses absent, trunk brown. Indumentum of simple hairs (?); old leafless branches glabrous, young foliate branches pubescent (?). Leaves: petiole ca. 3 mm long, ca. 1 mm in diameter, pubescent (?), grooved, blade inserted on the side of the petiole; blade 8 - 12 cm long, ca. 4 cm wide, elliptic to oblong, apex acuminate, acumen ca. 1.5 cm long, base cuneate, papyraceous, below pubescent when young (?), glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below pubescent when young (?), glabrous when old; secondary veins 10 to 13 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, axillary, peduncule generally absent. Flowers with 9 perianth parts in 3 whorls, 4 to 9 per inflorescence, pedicel 15 - 27 mm long, ca. 1 mm in diameter, sparsely pubescent (?); in fruit ca. 20 mm long, 2 - 3 mm in diameter, glabrous (?); bracts not seen (soon falling?); sepals 3, valvate, free, 1.5 - 2 mm long, ca. 1.5 mm wide, circular, apex rounded, base rounded, color unknown, pubescence not observed, margins flat; petals free; outer petals 3, valvate, 5 - 7 mm long, 2 - 3 mm wide, broadly elliptic to oblong, apex rounded, base truncate, color unknown, margins flat or wavy when dry, pubescence not observed; inner petals 3, imbricate (?), dimensions and shape not observed; stamens number not counted, row number not counted, ca. 1 mm long, broad; connective discoid, pubescence not observed, color unknown; staminodes absent (?); carpels free, 10 to 15 (?, possibly more based on the number of monocarps counted), ovary ca. 1.5 mm long, stigma flat, glabrous. Monocarps stipitate, stipes 49 - 50 mm long, ca. 1 mm in diameter; monocarps 17 to 18, 10 - 15 mm long, 10 - 15 mm in diameter, globose, apex rounded, glabrous (?), bumpy; seeds not seen.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
87DEACC19B13DB1516A641F573B4B182.taxon	distribution	Distribution. Equatorial Guinea and Cameroon (?); if present in Cameroon then from South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
98FC49D3DA0DB66A50C0A83BCE0CF8DE.taxon	description	Figs 22, 23; Map 3 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
98FC49D3DA0DB66A50C0A83BCE0CF8DE.taxon	materials_examined	Type. Sierra Leone. Northern Region; Bagroo River, Mann G. 828, Apr 1861: lectotype, sheet here designed: K [K 000880416]; isotypes: K [K 000880417]; P [P 00362653].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
98FC49D3DA0DB66A50C0A83BCE0CF8DE.taxon	description	Description. Tree, up to 30 m tall, d. b. h. up to 60 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole (3 -) 10 - 20 mm long, ca. 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 4.5 - 31 cm long, 2.5 - 6 cm wide, oblong to narrowly oblong or oblanceolate to narrowly oblanceolate, apex acute to acuminate, acumen 1 - 1.5 cm long, base cuneate to rounded, coriaceous, above glabrous when young and old, shiny when dry, below glabrous when young and old, green, concolorous; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 10 to 24 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young or old leafless branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 2 to 9 per inflorescence; pedicel 10 - 25 mm long, ca. 1 mm in diameter, glabrous; in fruit 15 - 30 mm long, 2 - 3 mm in diameter, glabrous; bracts 1 to 3, all basal, 1 - 2 mm long, ca. 2 mm wide; sepals 3, valvate, free, 2 - 3 mm long, ca. 2 mm wide, triangular to ovate, apex acute, base truncate, green, glabrous outside, glabrous inside, margins flat; petals free; outer petals longer than inner; outer petals 3, 7 - 12 mm long, 2 - 4 mm wide, obovate to oblong, apex obtuse, base truncate, green, margins flat, glabrous outside, glabrous inside; inner petals 3, imbricate, 3 - 4 mm long, 2 - 3 mm wide, ovate to suborbicular, apex rounded, base truncate, green, margins flat, glabrous outside, glabrous inside; stamens 25 to 30, in 3 to 4 rows, ca. 1 mm long, broad; connective discoid, pubescent, green; staminodes absent; carpels free, ca. 10, ovary ca. 1 mm long, stigma capitate, glabrous. Monocarps stipitate (sometimes shortly so), stipes 3 - 12 mm long, 3 - 4 mm in diameter; monocarps 3 to 6, 15 - 23 mm long, 11 - 25 mm in diameter, ellipsoid to globose, apex rounded, glabrous, finely warty, constricted around seeds, bumpy; seeds 1 to 2 per monocarp, ca. 12 mm long, 8 - 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
98FC49D3DA0DB66A50C0A83BCE0CF8DE.taxon	distribution	Distribution. In West Africa, Senegal, Sierra Leone to Nigeria, and Central Africa from Cameroon to Uganda; in Cameroon known from South, Central, Littoral, South-West and East regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
98FC49D3DA0DB66A50C0A83BCE0CF8DE.taxon	materials_examined	Specimens examined. Central Region: Avome, 3.87 ° N, 11.52 ° E, 13 August 1945, Aubreville A. 41 (P). East Region: 76 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.72 ° E, 05 March 2019, Couvreur T. L. P. 1202 (MPU, WAG, YA); Dja Reserve, 3.17 ° N, 13.18 ° E, 07 October 1994, Fogiel M. K. 947 (P); Deng Deng, 5.2 ° N, 13.51 ° E, 01 July 1939, Jacques-Felix H. 4630 (P, WAG); Rives de la Boumba a 14 km a l'WSW de Kinsassa village situe a 65 km au NNE de Moloundou sur route de Yokadouma 2.58 ° N, 15.26 ° E, 07 March 1971, Letouzey R. 10523 (P, YA). Littoral Region: km 19 Loum-Yabassi 3 km N of Sole, 4.61 ° N, 9.8 ° E, 30 December 1971, Leeuwenberg A. J. M. 9032 (YA, WAG). South-West Region: Southern Bakundu Forest Reserve, 4.55 ° N, 9.433 ° E, 15 June 1960, Adebusuyi J. K. 44049 (WAG); S Bakundu Forest 3 km from Kindongi Camp (8 km from road), 4.49 ° N, 9.374 ° E, 02 May 1972, Leeuwenberg A. J. M. 9785 (B, BR, C, K, M, MO, P, WAG, YA); 2 km W of km 21 Kumba-Victoria road, 4.46 ° N, 9.483 ° E, 04 May 1972, Leeuwenberg A. J. M. 9828 (B, BR, C, K, LD, M, MO, P, WAG, YA); Bibundi, 4.21 ° N, 8.988 ° E, 08 November 1928, Mildbraed G. W. J. 10640 (K); Korup National Park, 4.88 ° N, 8.783 ° E, 22 July 1983, Thomas D. W. 2329 (MO, P, WAG, YA); Limbe (Victoria), 4.01 ° N, 9.133 ° E, 25 October 1997, van der Burgt X. M. 219 (KRIBI, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3B2341A289312719EEC9A975CCBF5D1D.taxon	description	Figs 22, 23; Map 3 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3B2341A289312719EEC9A975CCBF5D1D.taxon	materials_examined	Type. Cameroon. South-West Region; Johann-Albrechtshohe [Kumba], Staudt A. 957, 1896: holotype: B [B 10 0154076]; isotypes: BM [BM 000546890]; K [K 000105343]; LE [LE 00012452]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3B2341A289312719EEC9A975CCBF5D1D.taxon	description	Description. Tree, 15 - 30 m tall, d. b. h. up to 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 10 - 20 mm long, ca. 3 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 9 - 17 cm long, 3 - 6.5 cm wide, obovate, oblong to elliptic, apex acuminate, acumen ca. 1 cm long, base rounded to acute, coriaceous, below glabrous when young and old, above glabrous when young and old, discolorous, whitish below, midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 10 to 12 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 2 to 3 per inflorescence; pedicel 12 - 20 mm long, ca. 1 mm in diameter, glabrous; in fruit 15 - 35 mm long, ca. 3 mm in diameter, glabrous; bracts 1 to 3, all basal, 1 - 2 mm long, 2 mm wide; sepals 3, valvate, free, ca. 2 mm long, ca. 2 mm wide, triangular to ovate, apex acute to rounded, base truncate, green, glabrous outside, glabrous inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 15 - 20 mm long, 2 - 3 mm wide, linear, apex acute, base truncate, green, margins flat, glabrous outside, glabrous inside; inner petals 3, imbricate, 2 - 2.5 mm long, 3 - 3.5 mm wide, ovate, apex acute, base truncate, claw mm long, green, margins flat, glabrous outside, glabrous inside; stamens 30 to 40, in 3 to 4 rows, ca. 1 mm long, broad; connective discoid, pubescent, green; staminodes absent; carpels free, 15 to 22, ca. 1 mm long, stigma capitate, glabrous. Monocarps stipitate to sessile, stipes when present to 10 mm long, 3 - 4 mm in diameter, 3 to 8 monocarps, 15 - 20 mm long, 12 - 15 mm in diameter, globose, apex rounded, glabrous, striate, bumpy, constricted around seeds; seeds 1 to 3 per monocarp, 10 - 15 mm long, 7 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3B2341A289312719EEC9A975CCBF5D1D.taxon	distribution	Distribution. From Cameroon to Gabon; in Cameroon known from South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3B2341A289312719EEC9A975CCBF5D1D.taxon	materials_examined	Specimens examined. Central Region: Ca 50 km NW of Eseka W of Yaounde, 3.65 ° N, 10.78 ° E, 25 November 1963, de Wilde W. J. J. O 1342 (B, BR, C, DES, L, LD, MO, P, U, WAG, YA); Ottotomo Forest Reserve, 3.65 ° N, 11.31 ° E, Service Forestier du Cameroun 32 (P). South Region: ca 15 km from Kribi 1 km S of Ebolowa road, 2.85 ° N, 10.01 ° E, 20 February 1970, Bos J. J. 6383 (P, WAG); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.81 ° N, 11.13 ° E, 02 February 1975, de Wilde J. J. F. E 7947 (B, BR, K, MO, P, U, WAG, YA); Pres de Bella (45 km NE de Kribi), 3.25 ° N, 10.2 ° E, 25 January 1962, Letouzey R. 4149 (P, YA); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1900, Zenker G. A. 2264 (L, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1902, Zenker G. A. 2454 (L, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1902, Zenker G. A. 2495 (L, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 4669 (L, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 4880 (L, P). South-West Region: Ekundu Kundu, 5.16 ° N, 8.874 ° E, 11 April 1996, Cable S. 1825 (K, YA); Kupe Mount Path to Kupe Rock, 4.75 ° N, 9.686 ° E, 24 November 1995, Cheek M. 7915 (K, P, WAG); Muanezum trail from Kupe village towards Daniel Ajang's area 4.77 ° N, 9.708 ° E, 18 July 1996, Etuge M. 2884 (K, MO, P, WAG); Just outside Kupe village going north, 4.77 ° N, 9.688 ° E, 29 November 1999, Gosline W. G. 240 (K, MO, P, WAG, YA); Korup National Park, 5.06 ° N, 8.855 ° E, 05 December 1997, Kenfack D. 984 (MO, P, WAG); Environs of Kumba farmed land and scrub with scattered trees, 4.63 ° N, 9.433 ° E, 01 March 1984, Thomas D. W. 3271 (MO, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50A7E1DA07EE918D544AFFFD29D8E14F.taxon	description	Description. Same as species. A genus with a single widespread species from West Africa (Sierra Leone to Nigeria) and in Cameroon. One species in Cameroon, not endemic. Dennettia was first described by Baker (1913) based on the bisexual flowers and inflorescences occurring on foliate branches. However, Kenfack et al. (2003) recombined the name Dennettia into Uvariopsis as Uvariopsis tripetala (Baker f.) G. E. Schatz based on a number of morphological characters. A molecular phylogenomic analysis of tribe Monodoreae (where this genus bellows) confirmed that Dennettia tripetala did not cluster with other species of Uvariopsis and should be regarded as a genus of its own (Dagallier et al. in prep).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
904A3B32649A2EB360E632244345D65C.taxon	description	Fig. 24; Map 3 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
904A3B32649A2EB360E632244345D65C.taxon	materials_examined	Type. Nigeria. Edo State; Benin City, Dennett R. E. 44, 1 Jan 1907: lectotype, designated by Kenfack et al. (2003), p. 447; sheet here designated: K [K 000040959]; isolectotypes: K [K 000040961]; S [S-G- 9774].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
904A3B32649A2EB360E632244345D65C.taxon	description	Description. Shrub to small tree, 2 - 5 m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to sparsely pubescent. Leaves: petiole 2 - 5 mm long, 1 - 2 mm in diameter, glabrous, slightly grooved, blade inserted on top of the petiole; blade 7.2 - 15.5 cm long, 3 - 6.8 cm wide, elliptic, apex attenuate to acuminate, acumen 0.6 - 1.3 cm long, base acute to decurrent, subcoriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 5 to 10 pairs per side, glabrous above; tertiary venation reticulate. Individuals bisexual, inflorescences ramiflorous on old leafless branches, axillary. Flowers with 6 perianth parts in 2 whorls, 1 to 4 per inflorescence; pedicel 4 - 9 mm long, 1 - 2 mm in diameter, pubescent; in fruit 5 - 15 mm long, 2 - 3 mm in diameter, glabrous to pubescent; bracts 1 to 3, all basal 0.5 - 2 mm long, 1.5 - 2 mm wide; sepals 3 (rarely 2), valvate, basally fused, 1 - 3 mm long, 1.5 - 4 mm wide, triangular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals 3 (rarely 4, see notes), free, 7 - 14 mm long, 6 - 10 mm wide, broadly ovate, apex obtuse, base truncate, margins flat, pubescent outside, glabrous inside; stamens ca. 150, in 10 to 20 rows, 0.5 - 1 mm long, oblong; connective reduced or absent, glabrous; staminodes absent; carpels free, 8 to 30, ovary ca. 2 - 4.5 mm long, stigma globose, pubescent. Monocarps stipitate, stipes 1 - 3 mm long, ca. 1 mm in diameter; monocarps 1 to 8, 11 - 32 mm long, 5 - 15 mm in diameter, ovoid to oblong, apex rounded, glabrous to sparsely pubescent, verrucose, wrinkled; seeds 4 to 12 per monocarp, 4 - 10 mm long, 11 - 14 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
904A3B32649A2EB360E632244345D65C.taxon	distribution	Distribution. Mainly a West African species from Sierra Leone to Cameroon; in Cameroon known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
904A3B32649A2EB360E632244345D65C.taxon	materials_examined	Specimens examined. South-West Region: Missellele, 4.12 ° N, 9.448 ° E, Box H. E. 3556 (BM, K); Limbe (Victoria), 4.07 ° N, 9.189 ° E, 01 April 1929, Maitland T. D. 626 (K); Buea area 4.2 ° N, 9.183 ° E, 01 January 1930, Maitland T. D. s. n. (K [K 000105532]); Likomba-Pflanzung 15 - 35 km NE von Victoria [Limbe], 4.1 ° N, 9.333 ° E, 18 October 1928, Mildbraed G. W. J. 10515 (K); Ngandjo on Kumba Mbonge road, 4.55 ° N, 9.4 ° E, 25 February 1986, Thomas D. W. 5661 (K); Market of Victoria, 4.01 ° N, 9.2 ° E, 04 April 1978, Westphal E. 9932 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C234E099E0F0C575437B44B3973179BD.taxon	description	Description. Trees, 8 - 50 m tall, d. b. h. up to 50 cm; stilt roots or buttresses absent. Indumentum of stellate or fasciculate hairs. Leaves: petiole 1 - 10 mm long, 2 - 6 mm in diameter; blade 7 - 34 cm long, 3 - 8 cm wide, ovate to elliptic to obovate, apex acuminate to acute, acumen 0.5 - 1 cm long, base cordate to acute, discolorous, whitish below or concolorous; midrib sunken or flat; secondary veins 8 to 25 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, leaf opposed or supra-axillary. Flowers with 9 perianth parts in 3 whorls, 2 to 5 per inflorescence; pedicel 3 - 22 mm long; in fruit 1 - 50 mm long; bracts 2, one basal and one upper in the lower half of pedicel, basal bract 7 - 12 mm long, upper bract similar than basal one; sepals 3, valvate, free, 6 - 15 mm long, apex acute, base truncate; petals free, outer petals longer than inner to sub equal; outer petals 3, valvate, free, 10 - 30 mm long, 4 - 10 mm wide, elliptic to ovate, apex acute to acuminate, base truncate; inner petals 3, imbricate, free, 4 - 20 mm long, 4 - 9 mm wide, elliptic to ovate to obovate, apex acute to acuminate, base truncate; stamens numerous, 1 - 2 mm long, broad; connective discoid; staminodes absent; carpels free, 50 to 125, 1.5 - 3.5 mm long, stigma globose. Fruit pseudosyncarpous; carpels sessile, connate or free, 60 to 125 carpels, 15 - 55 mm long, 7 - 30 mm in diameter, globose to ovoid to ellipsoid, apex domed-shaped to acute to apiculate; seed 1, 7 - 20 mm long, 4 - 13 mm in diameter, obovoid to ellipsoid; aril present, rudimentary. A genus of 94 species, with a disjunct distribution, 89 in the Neotropics and 4 in Africa, but absent from Madagascar. All four African species are known from Cameroon, one endemic. This genus of trees is characterized by stellate hairs on its leaves and pseudosyncarpous fruits. The only other tree genus with stellate hairs in Cameroon is Annickia, but the latter has a yellow slash and apocarpous fruits with clearly stipitate monocarps.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E349DB3AAFD87A1EC300916896F74065.taxon	description	Fig. 25; Map 3 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E349DB3AAFD87A1EC300916896F74065.taxon	materials_examined	Type. Nigeria. Anambra state; Onitsha, Barter C. 445, 1858: holotype: K [K 000198875].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E349DB3AAFD87A1EC300916896F74065.taxon	description	Description. Tree, 8 - 40 m tall, d. b. h. 40 - 60 cm; stilt roots or buttresses absent. Indumentum of stellate or fasciculate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches sparsely pubescent to densely pubescent. Leaves: petiole 3 - 7 mm long, 2 - 3 mm in diameter, densely to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 10 - 23 cm long, 3 - 7 cm wide, ovate to elliptic, apex acuminate to acute, acumen ca. 1 cm long, base cordate to acute, subcoriaceous, below densely pubescent when young and old, above glabrous when young and old, concolorous; midrib sunken or flat, not grooved, above glabrous when young and old, below densely pubescent when young and old; secondary veins 9 to 18 pairs, distinct, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, appearing axillary, forming on a short peduncle 4 - 10 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 5 per inflorescence; pedicel 11 - 22 mm long, 3 - 4 mm in diameter, sparsely to densely pubescent; in fruit 1 - 25 mm long, 1 - 4 mm in diameter, sparsely to densely pubescent; bracts 2, one basal and one upper in the lower half of pedicel, basal bracts 7 - 9 mm long; sepals 3, valvate, free, 10 - 15 mm long, 9 - 10 mm wide, ovate, apex acute, base truncate, yellowish green to greyish green, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner to sub equal; outer petals 3, 10 - 22 mm long, 5 - 8 mm wide, elliptic to ovate, apex acute, base truncate, cream to white, margins flat, sparsely pubescent outside, glabrous inside; inner petals 3, imbricate, 15 - 17 mm long, 4 - 5 mm wide, elliptic to ovate, apex acute, base truncate, cream, margins flat, pubescent outside, glabrous inside; stamens numerous, ca. 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 50 to 75, ovary ca. 1.5 mm long, stigma globose, glabrous. Fruit pseudosyncarpous, 40 - 200 mm in diameter, globose to depressed ovoid; individual carpels sessile, 60 to 70 carpels, completely fused, ca. 15 mm long, ca. 7 mm in diameter, globose to ovoid, apex domed-shaped, densely pubescent, longitudinally ribbed with 5 to 6 main ribs, pinkish red when ripe; seed 1 per monocarp, 7 - 15 mm long, 4 - 7 mm in diameter, ellipsoid; aril present, pale yellow.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E349DB3AAFD87A1EC300916896F74065.taxon	distribution	Distribution. A central African species, from Cameroon to Gabon; in Cameroon known from East, South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E349DB3AAFD87A1EC300916896F74065.taxon	materials_examined	Specimens examined. Central Region: Bank of Nyong river 40 km SE of Yaounde, 3.65 ° N, 10.78 ° E, 09 November 1961, Breteler F. J. 2013 (BR, K, P, WAG, YA); Abimoa, 3.57 ° N, 11.62 ° E, 04 April 1962, de Bruijn J. s. n. (WAG [WAG 0175175]); Bordure du Nyong pres du lac de Nkolmaka (pres route Mbalmayo-Akonolinga), 3.51 ° N, 11.82 ° E, 22 April 1954, Letouzey R. 304 (P, YA). Littoral Region: Right bank Ouem river near mouth in Sanaga R 6 km SW of Massok, 4.13 ° N, 10.47 ° E, 04 April 1965, Leeuwenberg A. J. M. 5377 (BR, K, MO, P, WAG, YA); Tissongo study area 3.57 ° N, 9.869 ° E, 01 June 1976, Waterman P. G. 874 (K). South Region: mountain chain Ngovoyang 2 km in forest from Bikiliki village situated between Bipindi and Lolodorf, 3.18 ° N, 10.52 ° E, 18 February 2012, Couvreur T. L. P. 393 (WAG, YA); Campo-Ma'an National Park, 2.38 ° N, 10.06 ° E, 01 July 2001, van Andel T. R. 3810 (KRIBI, WAG, YA). South-West Region: Ekundu Kundu, 5.15 ° N, 8.883 ° E, 30 April 1996, Cheek M. 8274 (K, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C04E0AD3EF0E5B7F975CFB8E25D18DE.taxon	description	Figs 25, 26; Map 3 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C04E0AD3EF0E5B7F975CFB8E25D18DE.taxon	materials_examined	Type. Gabon. Estuaire; Libreville, Klaine T. - J. 217, 10 Oct 1895: lectotype, sheet here designated: P [P 00315819]; isotypes: P [P 00315821, P 00315815].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C04E0AD3EF0E5B7F975CFB8E25D18DE.taxon	description	Description. Tree, 15 - 40 m tall, d. b. h. 40 - 85 cm; stilt roots or buttresses absent. Indumentum of stellate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches densely pubescent. Leaves: petiole 1 - 8 mm long, 2 - 6 mm in diameter, densely pubescent, cylindrical, blade inserted on the side of the petiole; blade 9 - 31 cm long, 3 - 8 cm wide, elliptic, apex acute, acumen 0.5 - 1 cm long, base rounded to acute, coriaceous, below densely pubescent with white erect stellate hairs covering the whole blade when young and old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below densely pubescent when young and old; secondary veins 14 to 25 pairs, indistinct, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, appearing axillary, forming on a short peduncle 2 - 11 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 5 per inflorescence; pedicel 3 - 20 mm long, 2 - 5 mm in diameter, densely pubescent; in fruit 3 - 20 mm long, 2 - 5 mm in diameter, glabrous to densely pubescent; bracts 2, one basal and one towards the upper half of pedicel, basal bracts 5 - 9 mm long, 5 - 10 mm wide; sepals 3, valvate, free, 17 - 22 mm long, 13 - 15 mm wide, ovate, apex acute, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner to sub equal; outer petals 3, 11 - 18 mm long, 5 - 9 mm wide, elliptic, apex acuminate, base truncate, cream to white, margins wavy, densely pubescent outside, glabrous inside; inner petals 3, imbricate, 11 - 18 mm long, 5 - 9 mm wide, elliptic, apex acuminate, base truncate, cream, margins flat, pubescent outside, glabrous inside; stamens 290 to 310, in 7 to 8 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 100 to 125, ovary 1.4 - 2 mm long, stigma globose, glabrous. Fruit pseudosyncarpous, 25 - 50 mm in diameter, ellipsoid; individual carpels sessile, 100 to 125 carpels, completely fused, 15 - 12 mm long, 2 - 5 mm in diameter, obovoid, apex apiculate to acute, densely pubescent, longitudinally ribbed with 5 to 6 main ribs, greyish brown with purplish red or pale brown pulp when ripe; seed 1 per monocarp, 10 - 14 mm long, 4 - 8 mm in diameter, ellipsoid; aril present, red.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C04E0AD3EF0E5B7F975CFB8E25D18DE.taxon	distribution	Distribution. From Cameroon to Gabon and Republic of Congo; in Cameroon known from East, South and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C04E0AD3EF0E5B7F975CFB8E25D18DE.taxon	materials_examined	Specimens examined. Central Region: Est N of Lom near Sanaga river (Yaunde) 250 km from Deng Deng, 4.84 ° N, 13.19 ° E, 01 March 1914, Mildbraed G. W. J. 8558 (K). Littoral Region: Au sud de Ngola (8 km E de l'embouchure de la Sanaga), 3.55 ° N, 9.698 ° E, 05 January 1974, Letouzey R. 12585 (P); Douala-Edea Reserve 10 B, 5.01 ° N, 13.33 ° E, 01 April 1978, Thomas D. W. 1237 (K). South Region: ca 16 km from Kribi Ebolowa road Bidou plantation Kienke forReserve, 2.85 ° N, 10.01 ° E, 03 February 1969, Bos J. J. 3844 (WAG, YA); 20 km From Kribi N of Lolodorf road (SFIA logging road), 3.01 ° N, 10.05 ° E, 15 July 1969, Bos J. J. 5048 (BR, BR, K, WAG, YA); Mt Elephant ca 18 km SE of Kribi, 2.78 ° N, 10.5 ° E, 14 January 1970, Bos J. J. 6128 (BR, P, WAG); Near mouth of, 3.17 ° N, 9.961 ° E, 28 March 1928, Hedin L. 1690 (P, WAG); Campo-Ma'an area 2.71 ° N, 9.866 ° E, 26 October 2001, van Andel T. R. 4205 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3195 (B, BR, E, G, L, M, MO, P, S).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4CFC1C4F756F94E6183C46B40233750B.taxon	description	Map 4 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4CFC1C4F756F94E6183C46B40233750B.taxon	materials_examined	Type. Gabon. Ngounie; new road from Mouila to Yeno, 5 km on either side of Kembele village, Thomas D. W. & Wilks C. M. 6510, 20 Jul 1986: lectotype, sheet here designated: WAG [WAG 0143388]; isotypes: MO [MO- 357359]; P [P 00389133]; WAG [WAG 0027128].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4CFC1C4F756F94E6183C46B40233750B.taxon	description	Description. Tree, up to 30 m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of stellate or fasciculate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches sparsely pubescent. Leaves: petiole 4 - 5 mm long, 2 - 3 mm in diameter, sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 7 - 16 cm long, 2.5 - 6 cm wide, narrowly elliptic to narrowly obovate, apex acuminate to acute, acumen ca. 1 cm long, base acute, subcoriaceous, below sparsely pubescent when young and old, above glabrous when young and old, concolorous; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 8 to 15 pairs, distinct, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed, not forming a peduncle. Flowers with 9 perianth parts in 3 whorls, 2 to 4 per inflorescence; pedicel 10 - 12 mm long, 1 - 2 mm in diameter, sparsely pubescent to densely pubescent; in fruit 10 - 50 mm long, 1 - 15 mm in diameter, sparsely pubescent to densely pubescent; bracts 2, one basal and one towards the lower half of pedicel, basal bracts 5 - 9 mm long, 7 - 9 mm wide; sepals 3, valvate, free, 12 - 15 mm long, 6 - 9 mm wide, elliptic to ovate, apex acute, base truncate, greyish green, pubescent outside, pubescent inside, margins flat; petals free, outer petals longer than inner to sub equal; outer petals 3, 12 - 15 mm long, 4 - 6 mm wide, ovate, apex acute, base truncate, white, margins flat, sparsely pubescent outside, glabrous inside; inner petals 3, imbricate, 13 - 15 mm long, 4 - 5 mm wide, elliptic, apex acute, base truncate, margins flat, pubescent outside, glabrous inside; stamens numerous, 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, ca. 75, 2 - 3.5 mm long, stigma globose, glabrous. Fruit pseudosyncarpous, size and shape unknown; carpels sessile, free to basally fused, unknown number of carpels, 20 - 35 mm long, ovary 10 - 30 mm in diameter, obovoid to deltoid, apex apiculate, pubescent, densely pubescent, longitudinally ribbed with 6 to 7 main ribs, color unknown; seed 1 per monocarp, 12 - 20 mm long, 10 - 13 mm in diameter, ellipsoid; aril present, color unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4CFC1C4F756F94E6183C46B40233750B.taxon	distribution	Distribution. Known from Gabon and Cameroon; in Cameroon known from South and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4CFC1C4F756F94E6183C46B40233750B.taxon	materials_examined	Specimens examined. South Region: Reserve forestiere de la Kienke (Kribi-Ebolowa km 16), 3.1 ° N, 10.25 ° E, 05 January 1968, Bamps P. R. J. 1679 (BR, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 692 (WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6072262267FC183E0310E02A508A383D.taxon	description	Fig. 26; Map 4 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6072262267FC183E0310E02A508A383D.taxon	materials_examined	Type. Cameroon. South Region; near Lolodorf, Staudt A. 133, 1896: holotype: B [B 10 0154084]; isotypes: BM [BM 000843984]; K [K 000198873, K 000198874]; P [P 00315814, P 00315816]; S [S 02 - 94].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6072262267FC183E0310E02A508A383D.taxon	description	Description. Tree, 15 - 50 m tall, d. b. h. 20 - 70 cm; stilt roots or buttresses absent. Indumentum of stellate or fasciculate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches sparsely pubescent to densely pubescent. Leaves: petiole 2 - 10 mm long, 2 - 4 mm in diameter, densely pubescent to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 13 - 34 cm long, 3 - 8 cm wide, narrowly obovate, narrowly oblong to narrowly elliptic, apex acuminate to acute, acumen 0.5 - 1 cm long, base cordate (more rarely acute), coriaceous, below sparsely pubescent when young, pubescent when old, above glabrous when young and old, concolorous; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 10 to 22 pairs, weakly distinct, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless or young foliate branches, leaf opposed or extra axillary, not forming a peduncle. Flowers with 9 perianth parts in 3 whorls, 2 to 4 per inflorescence; pedicel 6 - 12 mm long, 2 - 3 mm in diameter, sparsely pubescent to densely pubescent; in fruit 2 - 30 mm long, 2 - 6 mm in diameter, sparsely pubescent to densely pubescent; bracts 2, one basal and one towards the upper half of pedicel, basal bracts 9 - 12 mm long, 5 - 7 mm wide; sepals 3, valvate, free, 9 - 15 mm long, 7 - 10 mm wide, ovate, apex acute, base truncate, green, densely pubescent outside, glabrescent inside, margins flat; petals free, outer petals longer than inner to sub equal; outer petals 3, 7 - 30 mm long, 4 - 10 mm wide, oblong-elliptic to oblong-obovate, apex acute, base truncate, cream to white, margins flat, sparsely pubescent outside, glabrous inside; inner petals 3, imbricate, 4 - 20 mm long, 4 - 6 mm wide, elliptic to obovate, apex acute to obtuse, base truncate, cream, margins flat, pubescent outside, glabrous inside; stamens 120 to 150, in 5 to 6 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 50 to 100, ovary 1 - 1.5 mm long, stigma globose, glabrous. Fruit pseudosyncarpous, 20 - 55 mm in diameter, globose to depressed ovoid; carpels sessile, basally fused, 50 to 100 monocarps, 20 - 55 mm long, 2 - 10 mm in diameter, globose to ovoid, apex acute, densely pubescent, longitudinally ribbed with 5 to 6 main ribs, red when ripe; seed 1 per monocarp, 7 - 13 mm long, 6 - 8 mm in diameter, ellipsoid; aril present, pale yellow.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6072262267FC183E0310E02A508A383D.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West (Sierra Leone, Liberia, Ivory Coast and Nigeria) and in Central Africa from Cameroon to Democratic Republic of the Congo; in Cameroon known from East, South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6072262267FC183E0310E02A508A383D.taxon	materials_examined	Specimens examined. Central Region: Yaounde, 3.86 ° N, 11.51 ° E, 01 January 1935, Foury P. 69 (P, WAG). East Region: 17 km along road to Deng Deng, 4.58 ° N, 13.68 ° E, 01 September 1961, Breteler F. J. 1841 (WAG); Pres Kinsassa 65 km au NNE de Moloundou sur route Yokadouma 2.63 ° N, 15.37 ° E, 04 March 1971, Letouzey R. 10509 (P, YA); Colline a l'ENE de Mbalam (140 km ESE de Djoum pres Souanke-Congo), 2.22 ° N, 13.82 ° E, 20 January 1973, Letouzey R. 11867 (P, YA). Littoral Region: Douala-Edea Reserve Tissongo study area Transect B, 3.57 ° N, 9.869 ° E, 01 June 1976, Waterman P. G. 879 (U). South Region: Bitye, 3.87 ° N, 11.52 ° E, 01 January 1919, Bates G. L. 1199 (BM, MO); 17 km east from Lele village, 2.28 ° N, 13.32 ° E, 07 September 2013, Couvreur T. L. P. 460 (WAG, YA); 25 km east from Lele village at end of path on Ivindo river, 2.25 ° N, 13.28 ° E, 09 September 2013, Couvreur T. L. P. 489 (WAG, YA); Sud TDC, 2.65 ° N, 9.9 ° E, 06 November 1991, Halle F. 4220 (WAG); Ncolbew 3.28 ° N, 11.2 ° E, 26 April 1928, Hedin L. 1646 (P); Colline Ebon pres Nkobiyo 25 km ENE d'Ambam, 2.45 ° N, 11.5 ° E, 21 March 1970, Letouzey R. 10181 (P, YA); Mvini 35 km east of Campo, 2.37 ° N, 10.09 ° E, 20 December 1983, Mikio K. 5 (P, YA); ca 7 km NE of Ebom, 3.11 ° N, 10.75 ° E, 01 August 1996, Parren M. P. E. 157 (KRIBI, WAG); ca 7 km NE of Ebom, 3.11 ° N, 10.75 ° E, 01 August 1996, Parren M. P. E. 212 (KRIBI, WAG); Lolodorf, 3.23 ° N, 10.73 ° E, 1896, Staudt A. 133 (P); Lolodorf, 3.23 ° N, 10.73 ° E, March 1895, Staudt A. 138 (B); Campo-Ma'an area 2.4 ° N, 10.1 ° E, 02 April 2001, van Andel T. R. 3290 (KRIBI, U, WAG, YA). South-West Region: Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 26 March 2016, Couvreur T. L. P. 1014 (WAG, YA); Near Mamfe, 5.75 ° N, 9.31 ° E, 19 April 1978, Thomas D. W. 384 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
30BFACA9B2D27D9A3BDA6AD7EA36EE1E.taxon	description	Description. Trees, 7 - 45 m tall, d. b. h. 3 - 125 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 2 - 8 mm long, 1 - 3 mm in diameter, blade 6.5 - 16.2 cm long, 2 - 6.7 cm wide, elliptic to oblong, apex acuminate to caudate, base cuneate to rounded, concolorous; midrib sunken or flat; secondary veins 5 to 18 pairs; tertiary venation reticulate. Individuals androdioecious; male and bisexual inflorescences similar in appearance, ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 4 - 6 mm long; in fruit 6 - 13 mm long; bracts 2, one basal and one upper, 1 - 2 mm long; sepals 3, valvate, free, 2 - 4 mm long, ovate, apex acuminate, base truncate; petals free; outer petals longer than inner; outer petals 3, valvate, 8 - 18 mm long, 2.3 - 2.6 mm wide, oblong to elliptic, apex acuminate, base rounded; inner petals 3, valvate, 8 - 18 mm long, 1.3 - 2.6 mm wide, ovate to elliptic, apex acuminate, base rounded; stamens 15 to 25, in 4 to 5 rows, ovary 1 - 2 mm long, elongated; connective tongue-shaped, glabrous; staminodes absent; carpels free, 10 to 20, 1 - 2 mm long, stigma ovoid, pubescent. Fruits apocarpous, monocarps stipitate, stipes 5 - 10 mm long, monocarps 2 to 8, 8 - 21 mm long, 7 - 21 mm in diameter, ellipsoid to globose, apex rounded, smooth, green turning wine red when ripe; seed 1 to 4, 3 - 13 mm long, 3 - 13 mm in diameter, ellipsoid to flattened ellipsoid; aril absent. A genus of six currently described species distributed across Africa. Two species are known from Cameroon, none endemic. Onana (2011) mentions Greenwayodendron oliveri (Engl.) Verdc. from Cameroon, but this is not confirmed here. The latter species is a West African endemic (Lissambou et al. 2018); several specimens from coastal Gabon previously identified as G. oliveri are now separated as a different species: G. littorale Lissambou, Dauby & Couvreur (Lissambou et al. 2018, 2019). Neither species is known from Cameroon to date.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C6D7038B99D65488042BA34716011BEE.taxon	description	Figs 27, 29; Map 4 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C6D7038B99D65488042BA34716011BEE.taxon	materials_examined	Type. Cameroon. South Region; 40 km from Kribi, 5 km. E. of Edea road, tract of Fifinda-Bella road (SFIA), Bos J. J. 6267, 6 Feb 1970: holotype WAG [WAG. 1433854]; isotypes BR [BR 0000014826399]; YA n. v.; WAG [WAG 1433855].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C6D7038B99D65488042BA34716011BEE.taxon	description	Description. Tree, 7 - 30 m tall, d. b. h. 3 - 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 3 - 6 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 6.5 - 16.2 cm long, 2.1 - 5.8 cm wide, elliptic to oblong, apex acuminate to caudate, acumen 0.4 - 2 cm long, base cuneate to rounded, papyraceous, below sparsely pubescent to glabrous when young, glabrous when old, above glabrous when young and old, concolorous; midrib impressed, above completely glabrous when young and old, below glabrous when young and old; secondary veins 5 to 7 pairs, glabrous below; tertiary venation indistinct. Individuals androdioecious; male and bisexual inflorescences similar, ramiflorous on old leafless and young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel ca. 4 mm long, ca. 1 mm in diameter, pubescent to glabrous; in fruit 6 - 13 mm long, 1 - 2 mm in diameter, pubescent to glabrous; bracts 2, one basal and one upper towards the upper half of pedicel, basal bracts 1 - 2 mm long, 2 mm wide; upper bracts 1 - 3 mm long, 1 - 3 mm wide; sepals 3, valvate, basally fused to free, 3 mm long, 3 - 4 mm wide, ovate, apex acuminate, base truncate, green, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 12 - 13 mm long, 2 - 2.5 mm wide, elliptic to ovate, apex acuminate, base rounded, green to light yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 12 - 13 mm long, 2 - 2.5 mm wide, ovate, apex acuminate, base rounded, green to light yellow, margins flat, pubescent outside, glabrous inside; stamens 10 to 15, in 4 to 5 rows, 1 - 2 mm long, elongated; connective tongue shaped, glabrous, green; staminodes absent; carpels free, 10 to 15, ovary ca. 1 mm long, stigma ovoid, pubescent. Monocarps stipitate, stipes 5 - 10 mm long, 1 - 3 mm in diameter; monocarps 2 to 8, 11 - 21 mm long, 11 - 21 mm in diameter, ellipsoid to globose, apex rounded, glabrous, smooth, smooth; seeds 1 to 4 per monocarp, 7 - 13 mm long, 7 - 13 mm in diameter, ellipsoid to flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C6D7038B99D65488042BA34716011BEE.taxon	distribution	Distribution. Known from Cameroon and Gabon; in Cameroon known from the Littoral and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C6D7038B99D65488042BA34716011BEE.taxon	materials_examined	Specimens examined. Central Region: Left bank Nyong R 30 km S of Edea near bridge in road to Kribi, 3.8 ° N, 10.13 ° E, 26 April 1965, Leeuwenberg A. J. M. 5582 (B, BR, C, GC, K, LUAI, MO, P, UC, WAG, YA). Littoral Region: Ndogtima Nyong (Edea), 3.8 ° N, 10.13 ° E, 03 February 1974, Letouzey R. 12869 (BR, P, WAG, YA). South Region: 43 kmN of Kribi 5 km E of Edea road forest track Fifinda-Bella old secondary forest, 3.21 ° N, 10.06 ° E, 06 February 1970, Bos J. J. 6267 (BR, P, WAG, YA); ca 16 km On the road from Ebolowa to Minkok, 2.98 ° N, 11.17 ° E, 12 September 1975, de Wilde J. J. F. E 8465 (B, BR, K, MO, P, WAG, YA); Mvini 35 km East of Campo, 2.39 ° N, 10.04 ° E, 19 December 1983, Kaji M. 4 (YA); Campo-Ma'an region, 2.28 ° N, 9.950 ° E, 17 January 2016, Lissambou B. J. 1745 (BRLU); Campo-Ma'an region, 2.28 ° N, 9.949 ° E, 17 January 2016, Lissambou B. J. 1748 (BRLU); Campo-Ma'an region, 2.28 ° N, 9.948 ° E, 17 January 2016, Lissambou B. J. 1775 (BRLU); Campo-Ma'an region, 2.28 ° N, 9.949 ° E, 17 January 2016, Lissambou B. J. 1788 (BRLU); Campo-Ma'an region, 2.29 ° N, 9.945 ° E, 18 January 2016, Lissambou B. J. 1807 (BRLU); Campo-Ma'an region, 2.40 ° N, 9.895 ° E, 18 January 2016, Lissambou B. J. 1828 (BRLU); Campo-Ma'an region, 2.40 ° N, 9.894 ° E, 18 January 2016, Lissambou B. J. 1830 (BRLU); Campo-Ma'an region, 3.19 ° N, 10.10 ° E, 19 January 2016, Lissambou B. J. 1855 (BRLU); Campo-Ma'an region, 3.19 ° N, 10.10 ° E, 19 January 2016, Lissambou B. J. 1856 (BRLU); Cagnon du Ntem 16 km SW de Nyabessan, 2.32 ° N, 10.28 ° E, 30 November 1982, Nkongmeneck B. A. 400 (YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BE1CCDCCC745F3AAFB14627EA6983642.taxon	description	Fig. 28, 29; Map 4 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BE1CCDCCC745F3AAFB14627EA6983642.taxon	materials_examined	Type. Gabon. Estuaire; Munda, Sibange Farm, Soyaux H. 218, 20 Feb 1881: holotype material presumably destroyed at B; lectotype, designated by Lissambou et al. 2018 (2018), p. 77: P [P 00363356]; isolectotype K [K 000580898].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BE1CCDCCC745F3AAFB14627EA6983642.taxon	description	Description. Tree, 8 - 45 m tall, d. b. h. 10 - 125 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent to sparsely pubescent. Leaves: petiole 2 - 8 mm long, 1 - 3 mm in diameter, pubescent to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 5.1 - 15.6 cm long, 2 - 6.7 cm wide, elliptic to oblong, apex acuminate to caudate, acumen 0.6 - 1.4 cm long, base cuneate to rounded, papyraceous, below pubescent when young, glabrous to pubescent when old, above densely to sparsely pubescent when young, sparsely pubescent when old, concolorous; midrib impressed, above pubescent at least basely when young and old, below densely pubescent when young, sparsely pubescent to densely pubescent when old; secondary veins 5 to 18 pairs, glabrous below; tertiary venation indistinct. Individuals androdioecious; male and bisexual inflorescences similar, ramiflorous on old leafless and young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence, pedicel 3 - 6 mm long, 1 - 2 mm in diameter, pubescent; in fruit 6 - 12 mm long, 2 - 3 mm in diameter, glabrous; bracts 2, one basal and one upper towards the upper half of pedicel, basal bracts 1 - 2 mm long, 2 mm wide; upper bracts 1 - 3 mm long, 1 - 3 mm wide; sepals 3, valvate, basally fused to free, 2 - 4 mm long, 2 - 4 mm wide, ovate, apex acuminate, base truncate, green, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 8 - 18 mm long, 1.3 - 2.6 mm wide, oblong-elliptic to ovate, apex acuminate, base rounded, green to light yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 8 - 18 mm long, 1.3 - 2.6 mm wide, elliptic to ovate, apex acuminate, base rounded, green to light yellow, margins flat, pubescent outside, glabrous inside; stamens 16 to 25, in 4 to 5 rows, 1 - 2 mm long, elongated; connective tongue-shaped, glabrous, green; staminodes absent; carpels free, 12 to 20, ovary 1 - 2 mm long, stigma ovoid, pubescent. Monocarps stipitate, stipes 5 - 10 mm long, 1 - 3 mm in diameter; monocarps 2 to 8, 8 - 18 mm long, 7 - 16 mm in diameter, ellipsoid to globose, apex rounded, glabrous, smooth, green turning wine red when ripe; seeds 1 to 4 per monocarp, 3 - 11 mm long, 3 - 11 mm in diameter, ellipsoid to flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BE1CCDCCC745F3AAFB14627EA6983642.taxon	distribution	Distribution. Known from Nigeria to the Republic of Congo and the Democratic Republic of Congo; in Cameroon known from the East, South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BE1CCDCCC745F3AAFB14627EA6983642.taxon	materials_examined	Selected specimens examined. Central Region: Ndanan 1, 3.62 ° N, 11.58 ° E, 21 October 2002, Cheek M. 11224 (K, YA); Mefou National Park, 3.61 ° N, 11.58 ° E, 13 March 2004, Cheek M. 27 (YA); Mefou National Park, 3.61 ° N, 11.58 ° E, 13 March 2004, Cheek M. 66 (YA); Ca 50 km S of Badjob ca 60 km SW Of Eseka Along the Njong-River, 3.68 ° N, 10.68 ° E, 19 March 1964, de Wilde W. J. J. O 2133 (B, BR, K, MO, P, WAG, YA); Yaounde, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 129 (P); Ngoro, 5.06 ° N, 11.19 ° E, 29 April 2017, Kamdem N. 510 (YA); AYOS, 3.98 ° N, 12.36 ° E, 17 June 2017, Kamdem N. 560 (YA); Ossoessam (Mbalmayo), 3.52 ° N, 11.5 ° E, 01 June 1965, Leeuwenberg A. J. M. 5755 (BR, K, P, WAG). East Region: Palisco forest consession 15 km along main road into consession, 3.48 ° N, 13.59 ° E, 27 March 2015, Couvreur T. L. P. 756 (WAG, YA); Deng Deng, 5.20 ° N, 13.13 ° E, 27 July 2014, Kamdem N. 166 (YA); Mindourou Alpicam, 4.12 ° N, 14.54 ° E, 11 December 2016, Kamdem N. 459 (YA); Colline a l'ENE de Mbalam (140 km ESE de Djoum pres de Souanke-Congo, 2.22 ° N, 13.82 ° E, 20 January 1973, Letouzey R. 11866 (YA); A 6 km au Nord de Mwapak (km 43 piste Yokadouma-Lomie, 3.54 ° N, 14.71 ° E, 22 June 1963, Letouzey R. 5322 (YA). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1180 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 20 June 2014, Couvreur T. L. P. 658 (WAG, YA); Olombe, 3.60 ° N, 9.959 ° E, 05 November 2014, Kamdem N. 175 (YA); Chantier Bakaka km 4 Ebone-EkoMtolo road (Ebone situated on km 11 of Nkongsamba-Loum road), 4.83 ° N, 9.9 ° E, 20 August 1971, Leeuwenberg A. J. M. 8164 (BR, K, L, MO, P, U, WAG, YA). South Region: 20 km from Kribi Lolodorf road, 3.03 ° N, 10.05 ° E, 09 June 1969, Bos J. J. 4769 (B, BR, K, LD, LM, MO, P, POZG, WAG, YA); Mt Elephant ca 18 km SE of Kribi, 2.81 ° N, 10.01 ° E, 08 January 1970, Bos J. J. 6100 (BR, C, K, LD, P, WAG, YA); hill above Nlonacko near village Ebianemeyong, 2.43 ° N, 10.35 ° E, 12 December 1998, de Wilde J. J. F. E 12163 (BR, KRIBI, MO, S, WAG); 16 km on the recently reconstructed road from Ebolowa to Minkok, 2.75 ° N, 11.25 ° E, 30 January 1975, de Wilde J. J. F. E 7940 (BR, K, MO, P, U, WAG, YA); Massif de Ngovayang village de Atog Boga, 3.25 ° N, 10.49 ° E, 30 August 2015, Droissart V. 2050 (BRLU); Massif de Ngovayang village de Atog Boga, 3.25 ° N, 10.49 ° E, 05 September 2015, Droissart V. 2159 (BRLU); A 13 km au N-NW de Djoum (UFA 09 - 007) vers " la tache verte " Foret dense inondee et foret secondaire, 2.77 ° N, 12.74 ° E, 25 April 2011, Droissart V. 834 (BRLU); Ebom, 3.1 ° N, 10.73 ° E, 20 February 1996, Elad M. 443 (KRIBI, WAG); Campo, 2.28 ° N, 9.950 ° E, 05 July 2015, Kamdem N. 329 (YA); Campo, 2.39 ° N, 10.02 ° E, 07 July 2015, Kamdem N. 349 (YA); Ma'an, 2.50 ° N, 10.76 ° E, 11 July 2015, Kamdem N. 397 (YA); Essam (Nanga Eboko), 4.68 ° N, 12.37 ° E, 13 February 1959, Letouzey R. 1106 (P); Essam (Nanga Eboko), 4.68 ° N, 12.37 ° E, 13 February 1959, Letouzey R. 1313 (P); 10 km environ a l'ESE de Campo a Kribi, 2.37 ° N, 9.82 ° E, 26 March 1968, Letouzey R. 9198 (YA); Campo-Ma'an National Park, 2.38 ° N, 10.06 ° E, 01 July 2001, van Andel T. R. 3794 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.41 ° E, 1897, Zenker G. A. 1278 (L, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 1899, Zenker G. A. 2062 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1900, Zenker G. A. 2166 (L, P, WAG). South-West Region: Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 25 March 2016, Couvreur T. L. P. 1002 (WAG, YA); Mokoko Forest Reserve Boa / Likinge (Bousa forest), 4.42 ° N, 8.972 ° E, 05 June 1994, Ekema S. N. 1208 (K, YA); Nguti, 5.34 ° N, 9.496 ° E, 03 June 2017, Kamdem N. 537 (YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
016E5CFA6E4AD54C0E8ED7ABBCC8044E.taxon	description	Description. Trees, 10 - 40 m tall, d. b. h. 35 - 100 cm; stilt roots or buttresses absent, but trunk strongly fluted. Indumentum of simple hairs. Leaves: petiole 1 - 8 mm long, 1 - 4 mm in diameter; blade 3.6 - 36 cm long, 1.2 - 10 cm wide, elliptic or obovate or ovate, apex acuminate or rounded to obtuse, base cuneate or cordate, concolorous; midrib sunken or flat; secondary veins 5 to 17 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 3 per inflorescence; pedicel (0) 1 - 15 mm long, 1 - 5 mm in diameter; in fruit 2 - 30 mm long, 1 - 5 mm in diameter; bracts 5 to 6, several basal and two (sometimes fused) on upper half of pedicel; sepals 3, valvate, free, 4 - 21 mm long, 3 - 14 mm wide, ovate, apex acute, base truncate; petals 6, in a single whorl and basally fused, tube 2 - 10 mm long, inner and outer whorl not differentiated, equal or subequal; lobes 9 - 80 mm long, 3 - 21 mm wide, margins plicate (folded in bud) or wavy; stamens numerous, in 10 to 13 rows, 1 - 8 mm long, elongated; connective discoid or elongated; staminodes absent; carpels free, 2 to 16, ovary 2 - 5 mm long, stigma bilobed or divided into two lobes with margins coiled inwards. Fruit apocarpous, monocarps stipitate or sessile, stipes 0 - 3 mm long; monocarps 1 to 8, 22 - 95 mm long, 13 - 65 mm in diameter, ellipsoid to cylindrical, apex rounded, smooth or rugose or warty, pubescent, orange-brown to medium brown when ripe; seeds 2 to 36, 10 - 40 mm long, 7 - 20 mm in diameter, flattened ellipsoid; aril absent. A genus of five species, distributed across Africa. Four species are known from Cameroon, one endemic. This genus of trees is characterized by thin plicate (folded) petals, a unique character for Annonaceae (Botermans et al. 2011). In addition, the petals are fused at the base and form a short tube with 6 lobes, a character otherwise only seen in Isolona. The trunk of adult trees is strongly fluted, a character also seen in the larger species of the genus Isolona (e. g. I. hexaloba).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
333C062318A62A91B7E3F9B6A72A44B4.taxon	description	Fig. 30; Map 4 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
333C062318A62A91B7E3F9B6A72A44B4.taxon	materials_examined	Type. Cameroon. South Region; Kribi, Busse W. C. O. 3216, 1904: holotype: B [B 10 0154198].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
333C062318A62A91B7E3F9B6A72A44B4.taxon	description	Description. Tree, 20 - 30 m tall, d. b. h. unknown; stilt roots or buttresses absent, trunk slender, not fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 1 - 7 mm long, 3 - 4 mm in diameter, pubescent, grooved, blade inserted on the side of the petiole; blade 15.5 - 36 cm long, 5.5 - 10 cm wide, obovate, apex acuminate, acumen 0.5 - 1 cm long, base cordate, coriaceous, below glabrous when young and old, above sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, concolorous; midrib sunken or flat, above pubescent when young and old, below pubescent when young and old; secondary veins 12 to 17 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 10 - 12 mm long, 4 - 5 mm in diameter, densely pubescent; in fruit 10 - 13 mm long, 2 - 3 mm in diameter, glabrous; bracts 5 to 6, several basal and two (sometimes fused) towards the upper half of pedicel, basal bracts 3 - 9 mm long, 2 - 5 mm wide; upper bracts 6 - 13 mm long, 2 - 3 mm wide; sepals 3, valvate, free, 16 - 20 mm long, 11 - 14 mm wide, ovate, apex acute, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals basally fused, tube 2 - 5 mm long, purple, inner and outer whorl not differentiated, equal, lobes 23 - 40 mm long, 8 - 10 mm wide, elliptic, apex rounded, pale yellow, margins wavy, densely pubescent outside, densely pubescent inside, plicate; stamens numerous, 7 - 8 mm long, elongated; connective elongated, glabrous; staminodes absent; carpels free, 3 to 7, ovary 4 - 5 mm long, stigma elongate, pubescent. Monocarps stipitate, ca. 1 mm long, ca. 3 mm in diameter; monocarps 2 to 4, 53 - 78 mm long, ca. 40 mm in diameter, ellipsoid, apex rounded, sparsely pubescent, rugose, irregularly ribbed in reticulate pattern, orange when ripe; seeds 17 to 19 per monocarp, 23 - 28 mm long, 17 - 19 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
333C062318A62A91B7E3F9B6A72A44B4.taxon	distribution	Distribution. endemic to Cameroon; known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
333C062318A62A91B7E3F9B6A72A44B4.taxon	materials_examined	Specimens examined. South Region: S bank of Kienke river just E of Kribi, 3.1 ° N, 10.25 ° E, 12 May 1969, Bos J. J. 4495 (P, WAG); 21 km from Kribi high forest exploitation N of Lolodorf road, 3.03 ° N, 10.05 ° E, 07 August 1969, Bos J. J. 5157 (BR, P, WAG, YA); ca 16 km from Kribi Lolodorf road, 3 ° N, 10.01 ° E, 19 September 1969, Bos J. J. 5370 (BR, C, K, LD, LM, MO, P, WAG, YA); Kribi, 2.95 ° N, 9.916 ° E, 01 September 1904, Busse W. C. O. 3216 (B); 31 km ESE Kribi N of Kienke River Nyabessan, 2.9 ° N, 10.16 ° E, 19 April 1968, Letouzey R. 9387 (P, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 2889 (B, BR, COI, F, G, L, M, MO, P, S, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 26 October 1908, Zenker G. A. 3550 (BR, COI, G, M, MO, P, S); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3592 (G, K); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1909, Zenker G. A. 3889 (BR, COI, G, K, L, M, MO, P, S); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1913, Zenker G. A. 4831 (BR, G, P, S).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C992A4DF28F1D2704318F2CA07A91D02.taxon	description	Figs 31, 32; Map 4 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C992A4DF28F1D2704318F2CA07A91D02.taxon	materials_examined	Type. Guinea. Labe; Fouta D'hiallon [Djallon], Heudelot, J. 865, Apr 1838: lectotype, sheet here designated: P [P 00315839]; isotypes: P [P 00315842, P 00486270, P 00315841]; G [G 00011589].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C992A4DF28F1D2704318F2CA07A91D02.taxon	description	Description. Tree, 25 - 40 m tall, d. b. h. up to 100 cm; stilt roots or buttresses absent, trunk deeply fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 8 mm long, 1 - 3 mm in diameter, densely pubescent, grooved, blade inserted on the side of the petiole; blade 7.2 - 25 cm long, 2.5 - 8.5 cm wide, ovate to obovate, apex acuminate, acumen 0.5 - 1.5 cm long, base cuneate to cordate, coriaceous, below glabrous when young and old, above sparsely pubescent when young, sparsely pubescent when old, concolorous; midrib sunken or flat, above pubescent when young and old, below pubescent when young and old; secondary veins 9 to 19 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 3 per inflorescence; pedicel 12 - 25 mm long, 1 - 2 mm in diameter, glabrous; in fruit 10 - 30 mm long, 4 - 5 mm in diameter, glabrous; bracts 5 to 6, several basal and two (sometimes fused) towards the upper half of the pedicel, basal bracts 3 - 9 mm long, 2 - 5 mm wide; upper bracts 8 - 12 mm long, 4 - 9 mm wide; sepals 3, valvate, free, 12 - 21 mm long, 9 - 12 mm wide, ovate, apex acute, base truncate, brown, densely pubescent outside, densely pubescent inside, margins flat; petals basally fused, tube 4 - 10 mm long, purple, inner and outer whorl not differentiated, sub equal; lobes 37 - 80 mm long, 6 - 21 mm wide, elliptic, apex rounded, green to bright yellow, margins wavy, pubescent outside, pubescent with glabrous base inside, plicate; stamens 190 to 210, in 10 to 13 rows, 3 - 5 mm long, elongated; connective hemispheric, glabrous, cream; staminodes absent; carpels free, 7 to 16, ovary 2 - 5 mm long, stigma bilobed, slightly capitate, pubescent. Monocarps stipitate, ca. 2 mm long, 2 - 3 mm in diameter; monocarps 1 to 8, (42) 50 - 95 mm long, 35 - 65 mm in diameter, ellipsoid to oblong, apex rounded, pubescent to glabrous, smooth, not ribbed, rusty-brown; seeds 12 to 36 per monocarp, 28 - 40 mm long, 17 - 20 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C992A4DF28F1D2704318F2CA07A91D02.taxon	distribution	Distribution. A widespread species, known from Guinea-Bissau to the Democratic Republic of Congo; in Cameroon known from the East, South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C992A4DF28F1D2704318F2CA07A91D02.taxon	materials_examined	Specimens examined. Central Region: Ndanan 2 to Mefou river, 3.62 ° N, 11.56 ° E, 13 October 2002, Cheek M. 11064 (K, YA); Ottotomo Forest Reserve 1 km after reserve base near small loggers road, 3.66 ° N, 11.28 ° E, 25 June 2013, Couvreur T. L. P. 446 (WAG, YA); Ottotomo Forest Reserve 3 km after reserve base near small loggers road, 3.66 ° N, 11.28 ° E, 08 September 2013, Couvreur T. L. P. 506 (WAG, YA); Ottotomo Forest Reserve 45 km South of Yaounde 5 km on path into reserve, 3.66 ° N, 11.28 ° E, 15 January 2015, Couvreur T. L. P. 666 (WAG, YA). East Region: 73 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.16 ° N, 14.71 ° E, 04 March 2019, Couvreur T. L. P. 1197 (MPU, WAG, YA); Sangha R (Ndakan), 2.78 ° N, 16 ° E, 08 March 1988, Fay J. M. 8299 (F, MO, P); West side of Sangha River, 2.35 ° N, 16.13 ° E, 01 November 1988, Harris D. J. 1518 (K, MO); West side of Sangha River, 2.35 ° N, 16.13 ° E, 02 August 1988, Harris D. J. 883 (MO, P); Rives de la Sangha pres Lidjombo 120 km au N de Ouesso, 2.61 ° N, 16 ° E, 08 April 1971, Letouzey R. 10614 (P, YA); A 15 km au Sud de Djouo (20 km E de Somalomo dur le Dja), 3.32 ° N, 12.93 ° E, 25 February 1962, Letouzey R. 4433 (P, YA); South Cameroon Forest Area Molundu District Bange forest between Lokomo Bumba and Bange, 2.83 ° N, 15.25 ° E, 22 February 1911, Mildbraed G. W. J. 4539 (HBG); South Cameroon Forest Area Molundu District between Yokadouma (Post Plehn) and Assobam, 3.4 ° N, 14.38 ° E, 21 April 1911, Mildbraed G. W. J. 4996 (HBG); Reserve de Biosphere du Dja vers 500 m de la station de Bouamir, 3.19 ° N, 12.81 ° E, 27 May 2001, Senterre B. 1641 (BR); Layon nord-sud a partir de Djolimpoun (entre Somalomo-Malen), 3.33 ° N, 12.87 ° E, 13 September 1993, Sonke B. 591 (YA). Littoral Region: km 11 Loum-Sole road, 4.7 ° N, 9.816 ° E, 24 May 1972, Leeuwenberg A. J. M. 9904 (K, MO, WAG, YA). South Region: Hill roughly between Nkolandom and Nkoemvone, 2.8 ° N, 11.16 ° E, 17 January 1975, de Wilde J. J. F. E 7909 (B, BR, C, K, MO, P, U, WAG, YA); Ebolowa, 2.91 ° N, 11.31 ° E, 01 June 1911, Mildbraed G. W. J. 5652 (HBG); Ebom, 3.1 ° N, 10.71 ° E, 24 September 1996, Ndoum D. 129 (KRIBI, WAG); Kribi, 2.92 ° N, 9.900 ° E, 01 October 1997, van der Burgt X. M. 232 (KRIBI, WAG); Bisyang, 2.98 ° N, 9.968 ° E, 04 June 2006, van Velzen R. 90 (BR, G, MO, WAG). South-West Region: Melon to Nyandon ca 2 km, 4.93 ° N, 9.533 ° E, 28 November 1998, Cheek M. 9716 (K, WAG, YA); Nyasoso village on max's trail to Mt 4.82 ° N, 9.701 ° E, 05 April 2016, Couvreur T. L. P. 1062 (WAG, YA); Ambas Bay, 4.01 ° N, 9.2 ° E, 1861, Mann G. 709 (GH, K, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3FC65217829F4CA6B4501955597115C8.taxon	description	Fig. 30; Map 4 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3FC65217829F4CA6B4501955597115C8.taxon	materials_examined	Type. Senegal. Tambacounda; Galam prope Bakel, Leprieur F. M. R. s. n., 1828: lectotype, designated by Botermans et al. (2011), p. 42: G [G 00011597]; isolectotypes: G [G 00011595, G 00011593]; P [P 00315834, P 00315832, P 00315836].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3FC65217829F4CA6B4501955597115C8.taxon	description	Description. Tree to shrub, 10 - 15 m tall, d. b. h. up to 35 cm; stilt roots or buttresses absent, often several stemmed, not fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 1 - 4 mm long, 1 - 2 mm in diameter, densely pubescent, grooved, blade inserted on the side of the petiole; blade 3.6 - 17.5 cm long, 1.2 - 6.5 cm wide, ovate to obovate, apex rounded to obtuse, rarely acuminate, acumen 1 cm long, base cuneate to cordate, coriaceous, below pubescent when young, glabrous when old, above pubescent when young, glabrous when old, concolorous; midrib impressed, above pubescent when young, glabrous when old, below pubescent when young, glabrous when old; secondary veins 6 to 14 pairs, below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, more rarely cauliflorous, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 3 per inflorescence; pedicel sessile or short up to 2 mm long, ca. 2 mm in diameter when present, densely pubescent; in fruit 2 - 4 mm long, 1 - 2 mm in diameter, sparsely pubescent; bracts 5 to 6, several basal and two (sometimes fused) towards the upper half of pedicel, basal bracts ca. 5 mm long, ca. 4 mm wide; upper bracts ca. 5 mm long, ca. 4 mm wide; sepals 3, valvate, free, 4 - 7 mm long, 3 - 6 mm wide, ovate, apex acute, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals basally fused, tube 3 - 4 mm long, inner and outer whorl not differentiated, sub equal; lobes 9 - 27 mm long, 3 - 7 mm wide, elliptic, apex rounded, cream, margins wavy, pubescent outside, pubescent inside, lobes curving inwards at the base and margins reflexed forming a hollow chamber; stamens numerous, 1 - 2 mm long, elongated; connective discoid, glabrous, cream-yellow; staminodes absent; carpels free, 2 to 7, ovary 2 - 3 mm long, stigma divided into two lobes with margins coiled inwards, pubescent. Monocarps stipitate to sessile, stipes <3 mm long, 3 - 4 mm in diameter; monocarps 1 to 5, 22 - 46 mm long, 13 - 22 mm in diameter, ellipsoid to cylindrical, apex rounded, sparsely pubescent, warty, constricted around the seeds, orange when ripe; seeds 2 to 8 per monocarp, 10 - 15 mm long, 7 - 10 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3FC65217829F4CA6B4501955597115C8.taxon	distribution	Distribution. A widespread species, known from Senegal to northern South Africa, with a disjunct population in southern Angola; in Cameroon known from the North and Far-North regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3FC65217829F4CA6B4501955597115C8.taxon	materials_examined	Selected specimens examined. Far-North Region: Route Lara-Guidiguis (15 km ENE de Kaele), 10.1 ° N, 14.45 ° E, 29 August 1964, Letouzey R. 6540 (P, YA); Pres Bourka (65 km SS 0 de Mokolo), 10.3 ° N, 13.56 ° E, 13 October 1964, Letouzey R. 7301 (P, YA); 9 km SE Guili 10 km NE Bourrah, 10.6 ° N, 13.74 ° E, 27 November 1989, Villiers J. - F. 4713 (P, YA). North Region: Ecole de faune de Garoua, 9.3 ° N, 13.4 ° E, 09 August 2000, Dong E. 393 (YA); Sanguere (10 km Garoua), 9.27 ° N, 13.47 ° E, 01 October 1949, Malzy P. 309 (P, YA); Environs village Boulko au pied Hossere Gode 15 km NW de Poli, 8.53 ° N, 13.13 ° E, 24 October 1983, Satabie B. 702 (P, YA); Collines de Tinguelin 10 km N de Garoua, 9.3 ° N, 13.4 ° E, 26 November 1984, Satabie B. 781 (P, YA); map # NC 33 VIII Garoua, 9.93 ° N, 13.86 ° E, 13 August 1983, Thomas D. W. 2432 (MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
783C79A9BE75C82088AFFA0C8453F608.taxon	description	Fig. 31; Map 4 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
783C79A9BE75C82088AFFA0C8453F608.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2268, 1900: holotype: B [B 10 0154199]; isotypes: BM [BM 000546383]; COI [COI 00033180]; G [G 00011622, G 00011623]; GEOT [GOET 005680]; HBG [HBG 518921]; K [K 000582046]; L [L 0049299]; M [M 0089219]; MO n. v.; P [P 00363369, P 00363367]; S [S 12 - 22767]; WAG [WAG 0053744]; WU [WU 0025889]; Z [Z- 000034502].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
783C79A9BE75C82088AFFA0C8453F608.taxon	description	Description. Tree, up to 35 m tall, d. b. h. up to 100 cm; stilt roots or buttresses absent, trunk fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 1 - 3 mm long, 1 - 2 mm in diameter, sparsely pubescent, slightly grooved, blade inserted on top of the petiole; blade 5 - 10 cm long, 1.5 - 3.5 cm wide, elliptic, apex acuminate, acumen 0.5 - 1 cm long, base cuneate, coriaceous, above glabrous when young and old, above sparsely pubescent when young, sparsely pubescent when old, concolorous; midrib sunken or flat, below pubescent when young and old, below pubescent when young and old; secondary veins 5 to 12 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 8 - 15 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 10 - 13 mm long, 2 - 3 mm in diameter, glabrous; bracts 5 to 6, several basal and two (sometimes fused) towards the upper half of pedicel, basal bracts 5 mm long, 3 mm wide; upper bracts 5 mm long, 3 mm wide; sepals 3, valvate, free, 7 - 11 mm long, 4 - 7 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, densely pubescent inside, margins flat; petals basally fused, tube 2 - 4 mm long, purple, inner and outer whorl not differentiated, equal; lobes 17 - 30 mm long, 4 - 8 mm wide, elliptic, apex rounded, cream to yellow, margins wavy, pubescent outside, pubescent inside, plicate; stamens numerous, in 11 to 13 rows, ca. 2 mm long, elongated; connective discoid, glabrous, cream; staminodes absent; carpels free, 3 to 4, ovary 2 - 3 mm long, stigma divided into two lobes with margins coiled inwards, pubescent. Monocarps stipitate, 1 - 2 mm long, ca. 3 mm in diameter; monocarps 2 to 4, 60 - 93 mm long, 40 - 55 mm in diameter, ovoid to oblong, apex rounded, pubescent to glabrous, verrucose, rusty-brown when ripe; seeds 15 to 15 per monocarp, 22 - 31 mm long, 11 - 18 mm in diameter, flattened; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
783C79A9BE75C82088AFFA0C8453F608.taxon	distribution	Distribution. Known from Cameroon, Gabon and the Republic of Congo (one specimen); in Cameroon known from the East, South and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
783C79A9BE75C82088AFFA0C8453F608.taxon	materials_examined	Specimens examined. South Region: South Province 15 km from Kribi, 2.96 ° N, 9.933 ° E, 28 August 1970, Bos J. J. s. n. (WAG [WAG 0162941]); Campakok (Camp Akok), 2.65 ° N, 9.9 ° E, 30 October 1991, Halle F. 4181 (WAG); Colline pres Mezese a 17 km ENE de Sangmeli 2.95 ° N, 12.14 ° E, 19 October 1966, Letouzey R. 8122 (BR, P, YA); Bidou, 2.85 ° N, 9.991 ° E, 28 May 1954, Medou J. SRFK 1703 (P, YA); 22 km NE of Kribi Kribi-Bipindi road Bidou I cultivated fields NE of village, 3.00 ° N, 10.09 ° E, 04 June 2006, van Velzen R. 98 (WAG); 26 km N of Kribi SE of the toll post for Kribi-Edea road forest at Bebamboue II, 3.05 ° N, 9.987 ° E, 26 May 2006, van Velzen R. 99 (WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1900, Zenker G. A. 2268 (COI, G, G, K, L, M, MO, P, S, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1907, Zenker G. A. 3330 (BR, G, UPS).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C082E386300D030606D0D9662F5AED70.taxon	description	Description. Trees, 3 - 30 m tall, d. b. h. 5 - 60 cm; stilt roots or buttresses absent. Indumentum of simple hairs or glabrous. Leaves: petiole 1 - 15 mm long, 1 - 4 mm in diameter, blade 8.5 - 29 cm long, 3 - 15 cm wide, elliptic or obovate or oblong, apex acuminate, base decurrent to rounded or acute, concolorous; midrib raised on upper surface; secondary veins 7 to 20 pairs; tertiary venation reticulate. Inflorescences ramiflorous on old leafless or young foliate branches, axillary. Flowers bisexual with 9 perianth parts in 2 whorls, 1 to 3 per inflorescence; pedicel 1 - 25 mm long, 1 - 2 mm in diameter; in fruit 2 - 29 mm long, 2 - 5 mm in diameter; bracts 2 to 7, several basal and one upper, lower half of pedicel; sepals 3, valvate, free, 1 - 9 mm long, 2 - 5 mm wide, ovate or elliptic, apex acute or acuminate or rounded, base truncate; petals 6, basally fused, tube 3 - 11 mm long, inner and outer whorl not differentiated, equal; lobes 6 - 31 mm long, 2 - 12 mm wide; stamens numerous, in 3 to 4 rows, 1 - 2 mm long, broad; connective discoid; staminodes absent; carpels fused - syncarpous, forming a single visible gynoecium, 1 - 3 mm long, stigma bilobed, slightly capitate or capitate. Fruit syncarpous, forming a single visible fruit, 30 - 90 mm long, 15 - 50 mm in diameter, ovoid or ellipsoid, apex apiculate or rounded or cuspidate; seeds numerous not seriate, 8 - 25 mm long, 5 - 15 mm in diameter, ellipsoid or flattened ellipsoid; aril absent. A genus of trees with 20 known species, 15 in Africa and 5 in Madagascar. In Cameroon nine species are known, none endemic. Isolona, together with its sister genus Monodora, are unique in Annonaceae in having truly syncarpous flowers (fused carpels) and fruits. This translates into single fruits with unordered seeds, in contrast to other genera which have either uni- or biseriate placentation. Petals in Isolona are basally fused forming a clearly visible tube, with six equal lobes of equal length in a single whorl. In the vegetative state, Isolona and Monodora (together with Polyceratocarpus pellegrinii) are characterized by a raised midrib, in contrast to a sunken or flat midrib in all other genera found in Cameroon.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EE1405FC99A1E83954CC9444C9D41800.taxon	description	Fig. 33; Map 4 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EE1405FC99A1E83954CC9444C9D41800.taxon	materials_examined	Type. Cameroon. Northern Region; Bangwe, Conrau G. 93, 17 Oct 1899: holotype: B [B 10 0154200]; isotype: K [K 000198841].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EE1405FC99A1E83954CC9444C9D41800.taxon	description	Description. Tree to shrub, 10 - 15 m tall, d. b. h. 10 - 15 cm; stilt roots or buttresses absent. Indumentum absent; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 3 - 6 mm long, 1 - 2 mm in diameter, glabrous, weakly grooved adaxially, blade inserted on the side of the petiole; blade 10 - 18 cm long, 3 - 7 cm wide, elliptic to obovate, apex acuminate, acumen 1 cm long, base decurrent to cuneate, subcoriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised, above glabrous when young and old, below glabrous when young and old; secondary veins 7 to 12 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 5 - 30 mm long, ca. 1 mm in diameter, glabrous; in fruit 2 - 5 mm long, 2 - 3 mm in diameter, glabrous; bracts 2 to 5, several basal and one upper towards the lower half of pedicel, basal bracts 2 - 3 mm long, 1 - 2 mm wide; upper bract sometimes leaf-like, 2 - 19 mm long, 1 - 4 mm wide; sepals 3, valvate, free, 4 - 9 mm long, 3 - 5 mm wide, elliptic to ovate, apex acute, base truncate, green speckled with red and purple, glabrous inside, glabrous outside, margins flat; petals basally fused, tube 4 - 7 mm long, inner and outer whorl not differentiated, equal; lobes 6 - 20 mm long, 2 - 7 mm wide, triangular, apex acute, green, turning bright yellow, margins flat, glabrous outside, glabrous inside, spreading horizontally; stamens numerous, in 3 to 4 rows, ca. 1 mm long, broad; connective discoid, glabrous, cream; staminodes absent; carpels fused into a single structure, ca. 2 mm long, stigma capitate, glabrous. Fruit syncarpous, sessile, 40 - 75 mm long, 20 - 35 mm in diameter, ovoid, apex rounded, glabrous, smooth, bumpy, green turning deep yellow when ripe; seeds not counted, 8 - 15 mm long, 5 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EE1405FC99A1E83954CC9444C9D41800.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West (Sierra Leone, Liberia, Ivory Coast and Ghana) and Central Africa (Cameroon, Gabon); in Cameroon known from South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EE1405FC99A1E83954CC9444C9D41800.taxon	materials_examined	Specimens examined. Littoral Region: Lombe amp Tissongo Study Area Doula-Edea Reserve, 3.78 ° N, 10.04 ° E, 01 June 1976, Waterman P. G. 830 (K). South Region: ca 15 km SE of Kribi Kienke Forest Res at Bidou II, 2.86 ° N, 10.03 ° E, 30 June 1969, Bos J. J. 4947 (BR, P, WAG, YA); 10 km From Kribi Lolodorf road, 2.96 ° N, 9.966 ° E, 09 July 1970, Bos J. J. 7069 (WAG). South-West Region: 5.46 ° N, 9.883 ° E, 7 October 1899, Conrau G. 93 (B, K); 16 minutes from research station towards rivers, 5.34 ° N, 9.496 ° E, 27 November 2000, Gosline W. G. 289 (K); Ente Ekondo Nene et Loe 15 km NW Ekondo Titi, 4.69 ° N, 8.97 ° E, 03 June 1976, Letouzey R. 15078 (P, YA); Korup National Park, 5.31 ° N, 8.966 ° E, 02 July 1951, Olorunfemi J. 30662 (K); Korup National Park between the Ndian River at PAMOL field and 25 km on transect " P ", 5.01 ° N, 8.833 ° E, 12 April 1985, Thomas D. W. 4763 (BR, K, MO, P, WAG, YA); Takamanda Forest Reserve, 6.21 ° N, 9.433 ° E, 30 April 1987, Thomas D. W. 7354 (MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BCEEEC4EE6E187FF39E2B71E102CCFD8.taxon	description	Figs 33, 34; Map 5 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BCEEEC4EE6E187FF39E2B71E102CCFD8.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Lukandu, Dewevre A. P. 1103, 19 Nov 1896: lectotype, sheet here designated: BR [BR 0000006248932]; isotypes: BR [BR 0000006248857, BR 0000006249588, BR 0000006249250].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BCEEEC4EE6E187FF39E2B71E102CCFD8.taxon	description	Description. Tree, 10 - 30 m tall, d. b. h. 5 - 45 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 4 - 7 mm long, 2 - 3 mm in diameter, densely pubescent, grooved, blade inserted on top of the petiole; blade 13 - 19 cm long, 4 - 5 cm wide, narrowly ovate to narrowly elliptic or oblong, apex acuminate, acumen 0.5 - 1 cm long, base cuneate to rounded, subcoriaceous, below sparsely pubescent when young, glabrous when old, above sparsely pubescent when young, glabrous when old, concolorous; midrib raised, above sparsely pubescent when young, glabrous when old, below sparsely pubescent when young, glabrous when old; secondary veins 11 to 14 pairs, sparsely pubescent to glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 3 per inflorescence; pedicel 10 - 23 mm long, 1 mm in diameter, sparsely pubescent to glabrous; in fruit 2 - 5 mm long, 2 - 3 mm in diameter, glabrous; bracts 3 to 4, several basal and one upper towards the lower half of pedicel, basal bracts 1 mm long, 1 mm wide; upper bracts 1 mm long, 1 mm wide; sepals 3, valvate, free, 3 mm long, 2 mm wide, ovate, apex acute, base truncate, green, pubescent outside, glabrous inside, margins flat; petals basally fused, tube 4 - 10 mm long, inner and outer whorl not differentiated, equal; lobes 9 - 20 mm long, 3 - 7 mm wide, elliptic to ovate, apex rounded, green turning red, margins wavy, glabrous outside, densely pubescent inside, spreading horizontally; stamens numerous, in 3 to 4 rows, 2 mm long, broad; connective discoid, densely pubescent, cream; staminodes absent; carpels fused into a single structure, 2 mm long, stigma bilobed, slightly capitate, densely pubescent. Fruit syncarpous, sessile, 60 - 80 mm long, 40 - 50 mm in diameter, ellipsoid, apex rounded, glabrous, smooth to verrucose, irregularly ribbed, green when ripe; seeds not counted, 15 - 25 mm long, 10 - 15 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BCEEEC4EE6E187FF39E2B71E102CCFD8.taxon	distribution	Distribution. A central African species, with a disjunct distribution between the Cameroon Volcanic Line in Cameroon, and Eastern Democratic Republic of Congo, one collection from Central African Republic; in Cameroon known from the Littoral, North-West and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BCEEEC4EE6E187FF39E2B71E102CCFD8.taxon	materials_examined	Specimens examined. Littoral Region: Manengouba Mt 4 km WNW Of Nkongsamba, 4.98 ° N, 9.900 ° E, 04 April 1972, Leeuwenberg A. J. M. 9550 (B, BR, C, GC, H, K, LD, M, MO, P, WAG, YA). North-West Region: Kagwene, 6.12 ° N, 9.734 ° E, 20 May 2009, Ashworth J. 196 (K, YA); Bamenda Distr Ngong, 6.58 ° N, 10.43 ° E, 01 June 1931, Maitland T. D. 1555 (BM, K); Bamenda Wae, 6 ° N, 10.41 ° E, 01 April 1931, Maitland T. D. 1596 (K). South-West Region: Nyasoso, 4.86 ° N, 9.7 ° E, 04 June 1996, Cable S. 2843 (K, MO, WAG, YA); 2 km north of Nyasoso towards Mpako, 4.84 ° N, 9.679 ° E, 04 April 2016, Couvreur T. L. P. 1054 (WAG, YA); Nyasoso, 4.83 ° N, 9.683 ° E, 19 March 1996, Etuge M. 1794 (K, WAG); White trail (above Kupe village) towards Madam 4.78 ° N, 9.716 ° E, 28 May 1996, Etuge M. 2000 (BR, K, MO, P, WAG, YA); Lake Edip, 4.95 ° N, 9.65 ° E, 21 November 1998, Etuge M. 4488 (K, WAG, YA); Etube Tape from Nyasoso, 4.83 ° N, 9.683 ° E, 06 February 1995, Lane P. 532 (K, YA); Bu 4.15 ° N, 9.233 ° E, 6 June 1898, Lehmbach H. 224 (B). West Region: Bali Ngemba Forest Reserve, 5.82 ° N, 10.08 ° E, 13 April 2004, Etuge M. 5431 (K, WAG, YA); Along the road 6 km W of Dschang on road to Fongo Ndeng, 5.45 ° N, 9.95 ° E, 15 May 1978, Westphal E. 10012 (P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
294CE15D1B400140E36996A2031A003F.taxon	description	Fig. 35; Map 5 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
294CE15D1B400140E36996A2031A003F.taxon	materials_examined	Type. Liberia. Montserrado; near Firestone plantations, along Dukwai road, Cooper G. P. 417, 7 May 1929: lectotype, designated by Couvreur (2009), p. 148: GH [GH 00286760]; isotypes: F [F 0093217]; FHO [FHA 00095994]; G [GH 00286760]; K n. v.; NY [NY 00026103]; WIS [WIS 00000299 MAD].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
294CE15D1B400140E36996A2031A003F.taxon	description	Description. Tree, 6 - 18 m tall, d. b. h. 20 cm; stilt roots or buttresses absent. Indumentum absent; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 1 - 5 mm long, 2 - 3 mm in diameter, glabrous, grooved, blade inserted on top of the petiole; blade 15 - 29 cm long, 6 - 15 cm wide, oblong to obovate, apex acuminate, acumen 1 - 2 cm long, base cuneate to rounded, subcoriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 18 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 14 - 25 mm long, 1 mm in diameter, glabrous; in fruit 16 - 27 mm long, 2 mm in diameter, glabrous; bracts 2 to 4, all basal, 1 mm long, 1 mm wide; sepals 3, valvate, free, 2 mm long, 2 mm wide, ovate, apex rounded, base truncate, green, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 6 - 11 mm long, inner and outer whorl not differentiated, equal; lobes 8 - 15 mm long, 4 - 6 mm wide, oblong, apex acute to rounded, green, margins flat, glabrous outside, glabrous inside, spreading horizontally; stamens numerous, in 3 to 4 rows, 2 mm long, broad; connective discoid, glabrous, green; staminodes absent; carpels fused into a single structure, 3 mm long, stigma capitate, glabrous. Fruit syncarpous, sessile, 30 - 90 mm long, 15 - 30 mm in diameter, ellipsoid, apex apiculate, glabrous, smooth, constricted over seeds in dried material, smooth when fresh, orange with white spots when ripe; seeds not counted, 10 - 15 mm long, 5 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
294CE15D1B400140E36996A2031A003F.taxon	distribution	Distribution. A mainly West African species, from Liberia to Ghana, with a few specimens from Cameroon and one from Gabon; in Cameroon known from South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
294CE15D1B400140E36996A2031A003F.taxon	materials_examined	Specimens examined. South Region: Bipindi, 3.08 ° N, 10.41 ° E, 01 February 1910, Zenker G. A. s. n. (F). South-West Region: Korup National Park, 5.06 ° N, 8.855 ° E, 10 March 1998, Kenfack D. 1063 (MO, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AECC3B815A2C88F5C6026C49552DD7D6.taxon	description	Fig. 36; Map 5 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AECC3B815A2C88F5C6026C49552DD7D6.taxon	materials_examined	Type. Democratic Republic of the Congo. Bas Congo; Lemba-Luki, Dewevre A. P. 365, no date: lectotype, sheet here designated: BR [BR 0000008801050]; isotypes: BR [BR 0000008799968, BR 0000008800725, BR 0000008799630].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AECC3B815A2C88F5C6026C49552DD7D6.taxon	description	Description. Tree to shrub, 8 - 15 m tall, d. b. h. 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 4 - 15 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 10 - 17 cm long, 4 - 7 cm wide, elliptic to obovate, apex acuminate, acumen ca. 1 cm long, base decurrent to cuneate, papyraceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 14 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 per inflorescence; pedicel 2 - 7 mm long, 1 mm in diameter, sparsely pubescent; in fruit 9 - 10 mm long, 2 - 3 mm in diameter, glabrous; bracts 3 to 5, several basal and one upper towards the lower half of pedicel, basal bracts 1 mm long, 0.5 - 1 mm wide; upper bract 1 mm long, 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 3 - 4 mm wide, ovate, apex acuminate, base truncate, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 3 - 4 mm long, inner and outer whorl not differentiated, equal; lobes 7 - 17 mm long, 5 - 7 mm wide, elliptic, apex rounded, green, margins flat, overall glabrous but pubescent towards margins outside and inside; margins curved outwards; stamens ca. 50, in 3 to 4 rows, 2 mm long, broad; connective discoid, glabrous; staminodes absent; carpels fused into a single structure, ca. 1 mm long, stigma capitate, glabrous. Fruit syncarpous, sessile, 60 - 70 mm long, 40 - 50 mm in diameter, ovoid, apex rounded, glabrous, smooth, not ribbed, green when ripe; seeds not counted, 10 - 20 mm long, 10 - 15 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AECC3B815A2C88F5C6026C49552DD7D6.taxon	distribution	Distribution. A widespread species, but with few overall specimens, from Liberia to Nigeria, and from Cameroon to Democratic Republic of Congo; in Cameroon in the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AECC3B815A2C88F5C6026C49552DD7D6.taxon	materials_examined	Specimens examined. South Region: Ngongondje hill near Akonetye 2 30 ' S of Ebolowa, 2.5 ° N, 11.13 ° E, 28 August 1979, Koufani A. 123 (P, YA). South-West Region: Piste Munkep-Gayama 40 km NNW Wum, 6.73 ° N, 9.95 ° E, 08 July 1975, Letouzey R. 13984 (K, MO, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EEF15B7BD01F613CBA37360CC729E77.taxon	description	Figs 37, 38; Map 5 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EEF15B7BD01F613CBA37360CC729E77.taxon	materials_examined	Type. Gabon. Estuaire; Environs de Libreville, Klaine T. - J. 360, 17 Feb 1896: lectotype, sheet here designated: P [P 00363270]; isotypes: B; K [K 00198842]; P [P 00363269, P 00363268]; WAG [WAG 0251603].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EEF15B7BD01F613CBA37360CC729E77.taxon	description	Description. Tree to shrub, 15 - 30 m tall, d. b. h. up to 50 cm; stilt roots or buttresses absent, trunk deeply fluted. Indumentum absent; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 2 - 4 mm long, 2 - 3 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 10 - 28 cm long, 3 - 11 cm wide, ovate to elliptic, apex acuminate, acumen 1 - 2 cm long, base obtuse to acute, coriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 16 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 7 - 30 mm long, ca. 1 mm in diameter, glabrous; in fruit 20 - 29 mm long, 3 - 5 mm in diameter, glabrous; bracts 3 to 5, several basal and one upper towards the lower half of pedicel, basal bracts ca. 1 mm long, 0.5 mm wide; upper bract 2 - 5 mm long, ca. 1 mm wide; sepals 3, valvate, free, 1 - 3 mm long, 2 - 4 mm wide, elliptic, apex acuminate, base truncate, green, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 4 - 10 mm long, inner and outer whorl not differentiated, equal; lobes 6 - 25 mm long, 4 - 12 mm wide, elliptic to ovate, apex acute to rounded, dark red, margins flat to wavy, glabrous outside, glabrous inside, spreading horizontally; stamens numerous, in 3 to 4 rows, 2 mm long, broad; connective discoid, glabrous, green; staminodes absent; carpels fused into a single structure, ca. 1 mm long, stigma bilobed, slightly capitate, glabrous. Fruit syncarpous, sessile, 30 - 70 mm long, 25 - 40 mm in diameter, ovoid, apex rounded, glabrous, smooth, bumpy, irregularly and transversely ribbed, light green to dark purple when ripe; seeds not counted, 8 - 15 mm long, 4 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EEF15B7BD01F613CBA37360CC729E77.taxon	distribution	Distribution. A widespread species from Nigeria to the Democratic Republic of Congo and northern Angola (one specimen); in Cameroon known from Adamaoua, East, South, Central, Littoral and South-West, North-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EEF15B7BD01F613CBA37360CC729E77.taxon	materials_examined	Specimens examined. Central Region: Tibati pres Mbatimbang, 6.14 ° N, 12.48 ° E, 04 December 1959, Letouzey R. 2392 (P, YA). East Region: A 25 km environ a l'ENE de Mikel village situe a 85 km au N de Moloundou sur la route de Yokadouma 2.93 ° N, 15.33 ° E, 24 February 1971, Letouzey R. 10419 (P, WAG, YA); A 14 km a l'Ouest de Yenga Port Gentil village situe a 35 km au NNE de Moloundou, 2.35 ° N, 15.35 ° E, 21 April 1971, Letouzey R. 10703 (BR, COI, K, P, WAG, YA); A 20 km au Sud de Mboy I (45 km a l'Est de Yokadouma), 3.38 ° N, 15.13 ° E, 16 May 1963, Letouzey R. 5072 (K, P, WAG, YA). Littoral Region: Douala (route Razel), 4.05 ° N, 9.71 ° E, 01 January 1955, Endengle E. SRFK 2121 (P, YA); Roue forestiere SNCB (km 36 vers Ndoksom) environ 25 km Sud Yabassi, 4.31 ° N, 9.958 ° E, 11 May 1976, Letouzey R. 14910 (C, K, MO, P, WAG, YA); Tissongo, 3.58 ° N, 9.9 ° E, 01 August 1976, McKey D. B. 245 (K). South Region: ca 15 km east from Lele village, 2.28 ° N, 13.32 ° E, 10 September 2013, Couvreur T. L. P. 495 (WAG, YA); near Bipaga II km 40 road Kribi-Edea, 3.15 ° N, 10.01 ° E, 30 December 1982, de Kruif A. P. M. 998 (MO, WAG, YA); 17 km S of the Lobe river along the road to Campo, 2.81 ° N, 10.13 ° E, 18 March 1975, de Wilde J. J. F. E 8088 (BR, K, MO, P, U, WAG, YA); A l'Ouest d'Alati (100 km SE de Djoum), 2.2 ° N, 13.42 ° E, 13 January 1973, Letouzey R. 11840 (K, P, YA); 22 km on road Kribi to Campo 12 km past Gross Batanga, 2.76 ° N, 9.881 ° E, 24 February 1994, Wieringa J. J. 2327 (MPU, U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 November 1901, Zenker G. A. s. n. (P). South-West Region: Korup National Park, 5.01 ° N, 8.783 ° E, 31 October 2005, van der Burgt X. M. 791 (BR, G, K, MO, P, WAG, YA). West Region: Bali- Ngemba FR, 5.81 ° N, 10.08 ° E, 13 April 2002, Onana J. M. 2030 (K, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
86956CC5940528EE78AFDB13738D5AA5.taxon	description	Fig. 39; Map 5 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
86956CC5940528EE78AFDB13738D5AA5.taxon	materials_examined	Type. Democratic Republic of the Congo. Kasai Oriental; Lualaba, Ledermann C. 11, Mar 1906: holotype: B [B 10 0154216]; isotype: K n. v.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
86956CC5940528EE78AFDB13738D5AA5.taxon	description	Description. Tree, 13 m tall, d. b. h. up to 50 cm; stilt roots or buttresses absent, trunk not fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 3 - 12 mm long, 3 - 4 mm in diameter, densely pubescent, grooved, blade inserted on the side of the petiole; blade 19 - 27 cm long, 6 - 10 cm wide, obovate, apex acuminate, acumen 1 - 2 cm long, base rounded to cordate, papyraceous, below densely pubescent when young, densely pubescent when old, above sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, concolorous; midrib raised above, above sparsely pubescent when young, sparsely pubescent when old, below densely pubescent when young, densely pubescent when old; secondary veins 15 to 20 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 2 - 4 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 2 - 8 mm long, 2 - 3 mm in diameter, pubescent; bracts 2 to 4, all basal, 2 - 4 mm long, 1 - 2 mm wide; sepals 3, valvate, free, 2 - 5 mm long, 2 - 4 mm wide, ovate, apex acuminate, base truncate, densely pubescent outside, glabrous inside, margins flat; petals basally fused, tube 5 - 10 mm long, inner and outer whorl not differentiated, equal; lobes 8 - 13 mm long, 3 - 5 mm wide, elliptic, apex acute, yellow, margins flat, densely pubescent outside, pubescent with base glabrous inside, curving inwards over the receptacle; stamens numerous, in 3 to 4 rows, 2 mm long, broad; connective discoid, glabrous; staminodes absent; carpels fused into a single structure, 2 mm long, stigma bilobed, glabrous. Fruit syncarpous, sessile, 30 - 60 mm long, 20 - 40 mm in diameter, ellipsoid, apex cuspidate, sparsely pubescent, longitudinally ribbed, color unknown; seeds not counted, 13 - 15 mm long, 6 - 8 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
86956CC5940528EE78AFDB13738D5AA5.taxon	distribution	Distribution. A central African species with a disjunct distribution, from Cameroon to Gabon and the Republic of Congo, also present in the Democratic Republic of Congo; in Cameroon known from the extreme East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
86956CC5940528EE78AFDB13738D5AA5.taxon	materials_examined	Specimens examined. East Region: Region near Station Molundu Dscha (Ngoko) Nginda 21 km north Molundu, 2.2 ° N, 15.2 ° E, 07 January 1911, Mildbraed G. W. J. 4193 (HBG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5100371CFE4D0F548E0840C4C1F3D8FA.taxon	description	Fig. 38; Map 5 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5100371CFE4D0F548E0840C4C1F3D8FA.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 3217, Jul 1904: holotype: B [B 10 0154211]; isotypes BR [BR 0000008498991]; G [G 00011576 G 00011761]; K [K 000198837]; S [S 10 - 21236]; WU [WU 0025885].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5100371CFE4D0F548E0840C4C1F3D8FA.taxon	description	Description. Tree, 15 - 30 m tall, d. b. h. up to 60 cm; stilt roots or buttresses absent, trunk fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 4 - 12 mm long, 1 mm in diameter, glabrous, grooved towards the base, blade inserted on the side of the petiole; blade 8.5 - 15.5 cm long, 3 - 6 cm wide, elliptic to obovate, apex acuminate, acumen 1 - 1.5 cm long, base decurrent to cuneate, papyraceous to subcoriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 12 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 per inflorescence; pedicel 1 - 2 mm long, 1 mm in diameter, glabrous; in fruit 30 mm long, 3 - 4 mm in diameter, glabrous; bracts 3 to 7, several basal and one upper towards the lower half of pedicel, basal bracts ca. 1 mm long, ca. 1 mm wide; upper bracts ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 2 - 3 mm wide, ovate, apex acuminate, base truncate, green, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 6 - 15 mm long, red, inner and outer whorl not differentiated, equal; lobes 10 - 23 mm long, 5 - 10 mm wide, ovate, apex acute, bright green-white to yellow, margins wavy, glabrous outside, glabrous inside, spreading horizontally; stamens numerous, in 3 to 4 rows, ca. 2 mm long, broad; connective discoid, glabrous, yellow; staminodes absent; carpels fused into a single structure, ca. 2 mm long, stigma bilobed, slightly capitate, glabrous. Fruit syncarpous, sessile, ca. 50 mm long, ca. 40 mm in diameter, globose, apex rounded, glabrous, rugulose, longitudinally 6 - 8 ribbed, green when unripe; seeds not counted, 8 - 10 mm long, 5 - 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5100371CFE4D0F548E0840C4C1F3D8FA.taxon	distribution	Distribution. Known from Nigeria and Cameroon; in Cameroon known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5100371CFE4D0F548E0840C4C1F3D8FA.taxon	materials_examined	Specimens examined. South Region: NE of Mt Elephant ca 20 km SE of Kribi, 2.8 ° N, 10.03 ° E, 10 February 1970, Bos J. J. 6298 (WAG); mountain chain Ngovoyang 42 km in forest from Bikiliki village situated between Bipindi and Lolodorf, 3.18 ° N, 10.53 ° E, 19 February 2012, Couvreur T. L. P. 402 (WAG, YA); Colline Nkolo Manga (20 km SE Kribi), 2.95 ° N, 9.916 ° E, 16 April 1968, Letouzey R. 9341 (P, WAG); Elephant Mont, 2.8 ° N, 10.01 ° E, 22 October 2001, van Andel T. R. 4177 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.41 ° E, 1895, Zenker G. A. 1716 (B, G, M, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1918, Zenker G. A. 22 (P, WAG); Mbiave, 3.21 ° N, 10.61 ° E, 01 January 1913, Zenker G. A. 267 (A, B, BR, C, G, M, MO, U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 April 1904, Zenker G. A. 3038 (B, COI, G, K, L, M, MO, P, S, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 July 1904, Zenker G. A. 3217 (B, BR, G, K, L, M, S, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3433 (BR, COI, G, G, L, M, M, MO, P, S); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3540 (G, K, L, M, M, MO); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1909, Zenker G. A. 3921 (B, BR, COI, COI, G, L, M, M, MO, P, S); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 4704 (BM, BR, G, K, L, M, P, S); Bipindi, 3.06 ° N, 10.38 ° E, 01 November 1919, Zenker G. A. 95 (BM). South-West Region: Southern Bakundu Forest 3 km from Kindongi Camp, 4.55 ° N, 9.416 ° E, 02 May 1972, Leeuwenberg A. J. M. 9784 (B, BR, C, H, K, LD, M, MO, P, WAG, YA); Korup National Park, 5.06 ° N, 8.783 ° E, 13 April 1978, Thomas D. W. 349 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
916570BCE2ECA7F21C1F82977A36C1B1.taxon	description	Fig. 40; Map 5 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
916570BCE2ECA7F21C1F82977A36C1B1.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Massanga (pres de Monveda), Thonner F. 104, 24 Sep 1896: lectotype, designated by Boutique (1951 b), p. 263, sheet here designated: BR [BR 0000005113330]; isotypes: BR [BR 0000005112715, BR 0000005113040].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
916570BCE2ECA7F21C1F82977A36C1B1.taxon	description	Description. Tree to shrub, 3 - 10 m tall, d. b. h. up to 25 cm; stilt roots or buttresses absent, trunk not fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 3 - 8 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 11 - 20 cm long, 4 - 7.5 cm wide, elliptic to obovate, apex acuminate, acumen 1 - 2 cm long, base decurrent to cuneate, coriaceous to subcoriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 12 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 5 - 18 mm long, ca. 1 mm in diameter, glabrous; in fruit 8 - 10 mm long, 3 - 4 mm in diameter, glabrous; bracts 3 to 7, several basal and one upper towards the lower half of pedicel, basal bracts ca. 1 mm long, ca. 1 mm wide; upper bracts ca. 2 mm long, 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 1 - 2 mm wide, ovate, apex acute, base truncate, dark green, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 3 - 6 mm long, inner and outer whorl not differentiated, equal; lobes 14 - 31 mm long, 3 - 5 mm wide, linear to lorate (strap-shaped), apex acute, green, margins flat, glabrous outside, glabrous inside, pendulous; stamens numerous, in 3 to 4 rows, ca. 2 mm long, broad; connective discoid, glabrous; staminodes absent; carpels fused into a single structure, ca. 2 mm long, stigma bilobed, slightly capitate, sparsely pubescent. Fruit syncarpous, sessile, 40 - 60 mm long, 20 - 35 mm in diameter, ellipsoid, apex rounded, glabrous, smooth, not or faintly ribbed, color unknown; seeds not counted, 15 - 18 mm long, 8 - 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
916570BCE2ECA7F21C1F82977A36C1B1.taxon	distribution	Distribution. A central African species, known from Cameroon to Gabon and the Democratic Republic of Congo; in Cameroon known from the South and East regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
916570BCE2ECA7F21C1F82977A36C1B1.taxon	materials_examined	Specimens examined. East Region: Pres de Banana 10 km ENE de Moloundou, 2.08 ° N, 15.28 ° E, 17 April 1971, Letouzey R. 10682 (P, WAG, YA); near Ndongo ca 45 km WNW of Moloundou, 2.16 ° N, 14.83 ° E, 15 March 1973, Letouzey R. 12085 (K, P); Pres Ndongo a 45 km WNW de Moloundou, 2.15 ° N, 14.86 ° E, 16 March 1973, Letouzey R. 12111 (BR, K, P, WAG, YA); pres Ndongo a 40 km WNW de Moloundou, 2.15 ° N, 14.86 ° E, 16 March 1973, Letouzey R. 12115 (BR, K, P, WAG, YA). South Region: Colline Ongongondje pres Akonekye 15 km NW d'Ambam, 2.46 ° N, 11.16 ° E, 23 March 1970, Letouzey R. 10205 (BR, COI, K, P, WAG, YA); Inselberg d'Akookas pres du village d'Akookas 38 km au sud est d'Ebolowa, 2.71 ° N, 11.27 ° E, 15 March 2001, Parmentier I. 1943 (BRLU, WAG); Inselberg d'Akookas pres du village d'Akookas 38 km au sud est d'Ebolowa, 2.71 ° N, 11.27 ° E, 15 March 2001, Parmentier I. 1961 (BRLU, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F998EB0ABEFF9B3BE202D506E7515B9A.taxon	description	Figs 40, 41; Map 5 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F998EB0ABEFF9B3BE202D506E7515B9A.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 1186, 1896: holotype: B [B 10 0154218]; isotypes: BM n. v., G [G 00011574]; K [K 000199013]; WU [WU 0025863].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F998EB0ABEFF9B3BE202D506E7515B9A.taxon	description	Description. Tree to shrub, 7 - 15 m tall, d. b. h. up to 15 cm; stilt roots or buttresses absent, trunk not fluted. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 2 - 6 mm long, ca. 2 mm in diameter, glabrous, slightly grooved, blade inserted on top of the petiole; blade 16 - 23 cm long, 6.5 - 8.5 cm wide, oblong to oblanceolate, apex abruptly acuminate, acumen 1 - 2 cm long, base rounded to acute, coriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 11 to 13 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary. Flowers with 9 perianth parts in 2 whorls, 1 to 2 per inflorescence; pedicel 3 - 7 mm long, ca. 1 mm in diameter, glabrous; in fruit 5 - 15 mm long, 2 - 3 mm in diameter, glabrous; bracts 2 to 4, all basal, 1 mm long, 1 mm wide; sepals 3, valvate, free, 2 - 5 mm long, 2 - 4 mm wide, ovate, apex acute, base truncate, green to brown-red, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 4 - 7 mm long, inner and outer whorl not differentiated, equal; lobes 15 - 25 mm long, 3 - 4 mm wide, lorate (strap-shaped) to oblong, apex acute, light yellow to light green, margins curved inwards, glabrous outside, glabrous inside, erect over receptacle, verrucose when dried; stamens ca. 40, in 3 to 4 rows, ca. 2 mm long, broad; connective discoid, glabrous, cream; staminodes absent; carpels fused into a single structure, ca. 3 mm long, stigma bilobed, slightly capitate, glabrous. Fruit syncarpous, sessile, 30 - 65 mm long, 15 - 30 mm in diameter, ellipsoid to globose, apex rounded, glabrous, smooth, faintly ribbed to longitudinally ribbed, green turning yellow when ripe; seeds not counted, 15 - 20 mm long, 8 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F998EB0ABEFF9B3BE202D506E7515B9A.taxon	distribution	Distribution. Known from Cameroon to the west of Republic of Congo; in Cameroon known from the South and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F998EB0ABEFF9B3BE202D506E7515B9A.taxon	materials_examined	Specimens examined. South Region: Bipinde, 3.26 ° N, 10.20 ° E, 16 June 1918, Annet E. 359 (WAG); 12 km from Kribi N of Ebolowa road between Kribi aifield and Kienke R, 2.88 ° N, 9.983 ° E, 18 June 1969, Bos J. J. 4866 (BR, K, LM, MO, P, POZG, WAG, YA); 6 km N of km Kribi-Lolodorf, 3.08 ° N, 10.25 ° E, 12 March 1970, Bos J. J. 6522 (BR, K, P, WAG, YA); Campo-Ma'an area Bibabimvoto, 2.25 ° N, 10.36 ° E, 01 February 2000, Elad M. 1269 (WAG); Campo-Ma'an area Ebianemeyong, 2.46 ° N, 10.29 ° E, 24 May 2002, Elad M. 1545 (KRIBI, WAG); 3 km S of Kwambo and 6 km WSW of Bipindi, 3.05 ° N, 10.36 ° E, 19 January 1987, Manning D. 1453 (MO); Campo-Ma'an area Bifa, 2.65 ° N, 10.28 ° E, 12 October 2001, Tchouto Mbatchou G. P. BIFAX _ 2 (WAG); Campo-Ma'an area Bibabimvoto, 2.21 ° N, 10.01 ° E, 13 May 2000, Tchouto Mbatchou G. P. 2855 (KRIBI, WAG, YA); Campo-Ma'an area Bibabimvoto, 2.25 ° N, 10.04 ° E, 24 August 2000, Tchouto Mbatchou G. P. 3009 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.41 ° E, 1896, Zenker G. A. 1186 (B, BM, G, K); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1907, Zenker G. A. 3375 (P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3471 (US); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1912, Zenker G. A. 4405 (G, K, MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A00DA1A50B978E89978BBD5A044DA05D.taxon	description	Description. Lianas, to 40 m tall, d. b. h. up to 4 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 3 - 12 mm long, 1 - 6 mm in diameter, pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 5 - 28 cm long, 3 - 12 cm wide, elliptic to obovate to oblong, apex acuminate to emarginate, base rounded to cordate; secondary veins 11 to 20 pairs; tertiary venation percurrent to indistinct. Individuals bisexual; inflorescences ramiflorous on old leafless branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 3 - 11 mm long, 1 - 7 mm in diameter; in fruit 3 - 7 mm long, 1 - 3 mm in diameter; bracts 2, one basal and one upper towards the lower half of pedicel; sepals 3, valvate, completely fused forming a nearly closed cup but tearing open at anthesis, 10 - 20 mm long; petals free, outer petals longer than inner; outer petals 3, imbricate, 30 - 55 mm long, 15 - 55 mm wide, ovate, apex acute to rounded, base attenuate; inner petals 3, imbricate, 15 - 35 mm long, 10 - 25 mm wide, elliptic to ovate, apex acute, base attenuate to acute; stamens up to 800, in 16 to 20 rows, 2 - 10 mm long, broad; connective flattened, pubescent, red; staminodes absent; carpels free, 150 - 175, 2 - 2.5 mm long, stigma capitate, pubescent. [Fruits only known from L. bella] Fruit pseudosyncarpous ca. 45 mm long, ca. 50 mm in diameter, globose; individual carpels sessile, 125 to 150 carpels, apex rounded to apiculate; seed 1, 15 - 16 mm long, 4 - 8 mm in diameter, ellipsoid; aril present. A genus of lianas with three known species, from Angola (Cabinda), Cameroon, Gabon and Republic of Congo; in Cameroon two species, none endemic. Lestestudoxa is distinguished by the lianescent habit and pseudosyncarpous fruits (carpels fusing after pollination to form a single fruiting unit, similar to those of the genus Duguetia, but the latter being trees) and the sepals completely fused around the floral bud and tearing at anthesis. The only other lianescent Annonaceae liana in Africa with pseudosyncarpous fruits is Pseudartabotrys, a monospecific genus endemic to Gabon (Le Thomas 1969 b).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3C5F071E0211D99EAD9ADAB07B2BA6B3.taxon	description	Figs 42, 43; Map 5 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3C5F071E0211D99EAD9ADAB07B2BA6B3.taxon	materials_examined	Type. Gabon. Nyanga; Midounga, near Tchibanga, Le Testu G. M. P. C. 1637, 24 Oct 1910: holotype: P [P 00364779]; isotype: BM [BM 000546388].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3C5F071E0211D99EAD9ADAB07B2BA6B3.taxon	description	Description. Liana, 16 - 40 m tall, d. b. h. up to 4 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 3 - 10 mm long, 1 - 4 mm in diameter, pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 5 - 18 cm long, 3 - 8 cm wide, elliptic to obovate, apex acuminate to emarginate, acumen 0.5 - 1 cm long, base rounded to cordate, coriaceous, below densely pubescent when young, densely pubescent when old, above glabrous when young and old, concolorous; midrib sunken or flat, above glabrous when young and old, below pubescent when young and old; secondary veins 12 to 20 pairs, pubescent below; tertiary venation percurrent to indistinct. Individuals bisexual; inflorescences ramiflorous on old leafless branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 3 - 6 mm long, 1 - 3 mm in diameter, pubescent; in fruit 3 - 7 mm long, 1 - 3 mm in diameter, pubescent; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 5 - 7 mm long, 3 - 5 mm wide; upper bract 6 - 12 mm long, 4 - 7 mm wide, not clasping the flower bud; sepals 3, valvate, completely fused, tearing at anthesis, 10 - 20 mm long, base truncate, brown, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 30 - 55 mm long, 15 - 55 mm wide, ovate, apex acute, base attenuate, yellow to orange with pinkish margins, margins crisped, densely pubescent outside, pubescent inside; inner petals 3, imbricate, 15 - 30 mm long, 10 - 25 mm wide, elliptic to ovate, apex acute, base attenuate to acute, yellow to orange with red marginal zone, margins wavy, densely pubescent outside, pubescent inside; stamens numerous, 2 - 3 mm long, broad; connective flattened, pubescent, red; staminodes absent; carpels free, around 175, ovary ca. 2 mm long, stigma capitate, pubescent. Fruit pseudosyncarpous, ca. 45 mm long, ca. 50 mm in diameter in total, globose; individual carpels sessile, 125 to 150 carpels, obovoid to obtrulloid, apex rounded to apiculate, sparsely pubescent, 6 to 7 ribbed, green turning red when ripe; seed 1 per monocarp, 15 - 16 mm long, 4 - 8 mm in diameter, ellipsoid; aril present, light brown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3C5F071E0211D99EAD9ADAB07B2BA6B3.taxon	distribution	Distribution. Known from Angola, Cameroon to Gabon and in the Republic of Congo; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3C5F071E0211D99EAD9ADAB07B2BA6B3.taxon	materials_examined	Specimens examined. South Region: 26 km E of confluent Ntem River and Akom River near Ebolowa, 2.29 ° N, 11.86 ° E, 05 March 1970, Letouzey R. 10097 (BR, P, P, YA); Tom (Nyabessan), 2.43 ° N, 10.52 ° E, 04 March 1963, Raynal J. 10195 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DEAA568526C7CB2CD9037E3C958612B7.taxon	description	Figs 42, 43; Map 6 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DEAA568526C7CB2CD9037E3C958612B7.taxon	materials_examined	Type. Gabon. Woleu-Ntem; Ncout, Le Testu G. M. P. C. 9320, 13 Oct 1938: lectotype, sheet here designated: P [P 00364781]; isotypes: P [P 00364782, P 00364783].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DEAA568526C7CB2CD9037E3C958612B7.taxon	description	Description. Liana, 20 m tall, d. b. h. 2 - 3 cm in diameter; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches densely pubescent to tomentose, young foliate branches densely pubescent to tomentose. Leaves: petiole 6 - 12 mm long, 3 - 6 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 15 - 28 cm long, 7 - 12 cm wide, obovate to sometimes oblong, apex rounded or emarginate or obcordate or mucronate, acumen 0.1 - 1.9 cm long, base cordate, coriaceous, below densely pubescent when young, densely pubescent when old, above glabrous when young and old, concolorous; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young, densely pubescent when old; secondary veins 11 to 18 pairs, densely pubescent below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 3 - 11 mm long, 4 - 7 mm in diameter, densely pubescent; in fruit unknown; bracts 2, one basal and one upper towards the upper half of pedicel, basal bract 8 - 13 mm long, 10 - 15 mm wide; upper bract 8 - 18 mm long, 6 - 10 mm wide, clasping the flower bud; sepals 3, valvate, completely fused, tearing at anthesis, 10 - 20 mm long, base truncate, brown, pubescent outside, pubescent inside, margins flat; petals free, outer petals longer than inner; outer petals 3, imbricate, 30 - 50 mm long, 30 - 40 mm wide, ovate, apex rounded to acute, base attenuate, orange to red, margins crisped, densely pubescent inside, pubescent outside; inner petals 3, imbricate, 25 - 35 mm long, 15 - 25 mm wide, elliptic to ovate, apex acute, base attenuate, claw ca. 5 mm long, orange to red, margins wavy, densely pubescent inside, pubescent outside; stamens 750 to 800, in 16 to 20 rows, 7 - 10 mm long, broad; connective flattened, pubescent; staminodes absent; carpels free, 150, ovary 2 - 2.5 mm long, stigma capitate, pubescent. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DEAA568526C7CB2CD9037E3C958612B7.taxon	distribution	Distribution. Known from southern Cameroon to Gabon; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DEAA568526C7CB2CD9037E3C958612B7.taxon	materials_examined	Specimens examined. South Region: Ma'an 3 km on small road towards Ntem river, 2.34 ° N, 10.63 ° E, 25 February 2018, Couvreur T. L. P. 1148 (MPU, WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 697 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 698 (WAG, YA); Campo-Ma'an area Boucle du Ntem, 2.34 ° N, 10.58 ° E, 16 February 2001, Tchouto Mbatchou G. P. 3166 (KRIBI, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
77AD57C4F54CA479105782892D5D98F0.taxon	description	Description. Same as species. A monotypic genus from the Central Atlantic African region (Lower Guinea). Meiocarpidium is characterized by the presence of peltate scale-like hairs, a character unique among Central African Annonaceae.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
82AAB8F0526F0B95F30AB5F6726BE69E.taxon	description	Figs 44, 45; Map 6 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
82AAB8F0526F0B95F30AB5F6726BE69E.taxon	materials_examined	Type. Equatorial Guinea. Rio Muni; Muni River, Mann G. 1774, Aug 1862: lectotype, sheet here designated: K [K 000795931]; isotypes: K [K 000795932]; P [P 00362615].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
82AAB8F0526F0B95F30AB5F6726BE69E.taxon	description	Description. Tree, 8 - 20 m tall, d. b. h. 20 cm; stilt roots or buttresses absent. Indumentum of peltate scale-like hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 9 - 11 mm long, 2 mm in diameter, pubescent, grooved, blade inserted on top of the petiole; blade 15 - 20 cm long, 5 - 7 cm wide, oblong to obovate, apex acuminate to acute, acumen 1 - 2 cm long, base decurrent to acute, coriaceous, below densely pubescent when young giving a silvery color, pubescent when old, above glabrous when young and old, discolorous, whitish below; midrib impressed above, above glabrous when young and old, below pubescent when young and old; secondary veins 13 to 17 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 20 - 30 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 20 - 30 mm long, 2 - 3 mm in diameter, pubescent; bracts 2, one basal and one upper towards the upper half of pedicel, basal bract 8 - 9 mm long, 1 mm wide; upper bract 2 - 3 mm long, 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 1 - 2 mm wide, triangular, apex acute, base truncate, green, pubescent outside, densely pubescent inside, margins flat; petals free, sub equal; outer petals 3, valvate, 15 - 25 mm long, 10 - 15 mm wide, ovate, apex acute, base truncate, cream, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 15 - 25 mm long, 5 - 10 mm wide, ovate, apex acute, base narrowed into a claw 3 - 5 mm long, cream, margins flat, pubescent outside, glabrous inside; stamens 90 to 100, in 5 to 6 rows, 3 - 4 mm long, broad; connective discoid, glabrous, cream; staminodes absent; carpels free, 3 to 5, ovary 7 - 8 mm long, stigma capitate, pubescent. Monocarps sessile, 3 to 4, 40 - 60 mm long, 20 - 30 mm in diameter, cylindrical, apex rounded to apiculate, pubescent, smooth, silver-green when ripe; seeds 14 to 16 per monocarp, 10 - 20 mm long, 5 - 10 mm in diameter, flattened ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
82AAB8F0526F0B95F30AB5F6726BE69E.taxon	distribution	Distribution. Known from Cameroon, Equatorial Guinea Gabon and Republic of Congo, one collection for the Democratic Republic of Congo; in Cameroon known from East, South, Littoral and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
82AAB8F0526F0B95F30AB5F6726BE69E.taxon	materials_examined	Selected specimens examined. Central Region: Bank Nyong River near the new bridge ca 65 km SSW of Eseka, 3.65 ° N, 10.78 ° E, 17 June 1964, de Wilde W. J. J. O 2735 (B, BR, K, MO, P, WAG, YA); Bafia region, 4.75 ° N, 11.23 ° E, 18 December 1957, de Wit H. C. D 7948 (WAG). East Region: Berberati, 4.58 ° N, 13.68 ° E, 01 March 1963, BPFV 2664 (P); 47 km NW of Bertoua near road from Mbang to Abaka, 4.58 ° N, 13.68 ° E, 22 May 1961, Breteler F. J. 1398 (A, BR, K, M, P, UC, WAG, YA); Bertoua 9 km along road to Deng Deng, 4.58 ° N, 13.68 ° E, 31 August 1961, Breteler F. J. 1817 (A, BR, K, M, P, WAG, YA); 70 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.73 ° E, 04 March 2019, Couvreur T. L. P. 1193 (MPU, WAG, YA); Yokadouma 3.52 ° N, 15.05 ° E, 11 September 1939, Jacques-Felix H. 4904 (P); Deng Deng, 5.20 ° N, 13.13 ° E, 27 July 2014, Kamdem N. 169 (YA); sur la route de Esseleke, 4.58 ° N, 13.68 ° E, 18 August 1955, Nana P. 213 (P, YA); km 38 de la route de Deng-Deng, 4.58 ° N, 13.68 ° E, 17 September 1955, Nana P. 257 (P, YA); Environ de Landjwe 25 km SW Yokadouma 3.47 ° N, 14.93 ° E, 16 June 1984, Satabie B. 767 (P, YA). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1182 (MPU, WAG, YA). South Region: 6 km N of km Kribi-Lolodorf, 3.08 ° N, 10.25 ° E, 12 March 1970, Bos J. J. 6523 (WAG); Near village Oveng 27 km from Sangmelima along road to Yaounde, 3.09 ° N, 11.90 ° E, 20 March 1962, Breteler F. J. 2646 (BR, K, L, P, U, WAG, YA); Campo Ma an National Park 26 km after Ntem river, 3.07 ° N, 14.64 ° E, 08 March 2019, Couvreur T. L. P. 376 (WAG, YA); on road from Lolodorf to Mekalat just after 1 sty village Along the Malange river, 3.19 ° N, 10.71 ° E, 20 February 2012, Couvreur T. L. P. 407 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.46 ° N, 10.35 ° E, 14 February 2015, Couvreur T. L. P. 709 (WAG, YA); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.81 ° N, 11.13 ° E, 27 September 1974, de Wilde J. J. F. E 7576 (BR, K, MO, P, WAG, YA); Ebolowa SW of Mbalmayo, 2.96 ° N, 11.28 ° E, 27 February 1964, de Wilde W. J. J. O 1942 (BR, K, MO, P, WAG, YA); Pres Akak 10 km W Sangmeli 2.96 ° N, 11.88 ° E, 11 March 1970, Letouzey R. 10153 (P, YA); Ebom, 3.05 ° N, 10.71 ° E, 29 August 1996, Ndoum D. 88 (KRIBI, WAG); Campo-Ma'an area 2.39 ° N, 10.07 ° E, 12 June 2001, van Andel T. R. 3619 (KRIBI, WAG, YA); Campo-Ma'an area Akom II, 2.8 ° N, 10.53 ° E, 18 August 2001, van Andel T. R. 3931 (U, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 1898, Zenker G. A. 1864 (P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1902, Zenker G. A. 2505 (P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1903, Zenker G. A. 2521 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 2947 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3221 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1911, Zenker G. A. 4209 (P, U); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1918, Zenker G. A. 60 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9E93D0EACC51043148FC6AC69598BE7F.taxon	description	Description. Genus description for Cameroon same as species. A genus of trees or shrubs with five known species (Gosline et al. 2018), from West and Central Africa and one from East Africa (endemic to Tanzania). In Cameroon one species is known, not endemic (previously included in Mischogyne elliotiana). The genus Mischogyne is easily identified when fertile by the presence of a torus, an extended receptacle, and several elongated cylindrical to ovoid carpels, which are divergent from each other.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9301BD4D7295F914D15E3CA602008CE2.taxon	description	Fig. 45; Map 6 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9301BD4D7295F914D15E3CA602008CE2.taxon	materials_examined	Type. Gabon. Nyanga; Mayumba, Le Testu G. M. P. C. 1768, 26 Aug 1914: lectotype, designated by Gosline et al. (2018), p. 28 [P 00315820]; isolectotypes BM [BM 000547338]; BR [BR 0000008801661, BR 0000008802330]; LISC [LISC 000379]; P [P 00315817, P 00315818]; WAG [WAG 0175098, WAG 0175099, WAG 0247284, WAG 0247285, WAG 0247286].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9301BD4D7295F914D15E3CA602008CE2.taxon	description	Description. Tree, 3 - 25 m tall, d. b. h. up to 25 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches shortly pubescent. Leaves: petiole 5 - 10 mm long, ca. 2 mm in diameter, sparsely pubescent to glabrous, cylindrical, blade inserted on the side of the petiole; blade 8 - 16 cm long, 4 - 7 cm wide, obovate, apex acuminate, acumen 1 - 3 cm long, base cuneate, papyraceous to subcoriaceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 6 to 9 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, extra-axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 5 - 12 mm long, 0.5 - 1 mm in diameter, pubescent; in fruit 7 - 15 mm long, 2 - 3 mm in diameter, glabrous; bracts few, reduced to a tuff of hairs, all basal; sepals, valvate, completely fused in bud, tearing at anthesis into 2 or 3 parts, 7 - 13 mm long, 3 - 5 mm wide, triangular, apex acute, base truncate, green, tomentose on both sides, margins flat; petals free, equal; outer petals 3, valvate, 14 - 16 mm long, 3 - 5 mm wide, narrowly elliptic, apex acute, base truncate, bright white, margins flat, densely pubescent outside, tomentose inside; inner petals 3, valvate, 14 - 16 mm long, 3 - 5 mm wide, narrowly elliptic, apex acute, base truncate, bright white, margins flat, densely pubescent outside, tomentose inside; stamens 50 to 60, in 3 to 4 rows, 1 - 3 mm long, elongated; connective tongue shaped, pubescent, cream; staminodes absent; carpels free, 3 - 7, ovary 5 - 6 mm long, stigma bilobed, densely pubescent. Monocarps sessile, 1 to 3, 60 - 65 mm long, 28 - 32 mm in diameter, cylindrical, apex rounded, glabrous, smooth, 1 - ribbed, yellow when ripe; seeds 6 per monocarp, ca. 20 mm long, ca. 15 mm in diameter, ellipsoid, covered with an indumentum of fine white hairs; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9301BD4D7295F914D15E3CA602008CE2.taxon	distribution	Distribution. Cameroon, Gabon and Bioko island (Equatorial Guinea); in Cameroon known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9301BD4D7295F914D15E3CA602008CE2.taxon	materials_examined	Specimens examined. South Region: Environs de Nko'olong 30 km E Campo, 2.40 ° N, 10.03 ° E, 21 November 1992, Satabie B. 978 (YA). South-West Region: Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.054 ° E, 02 April 2016, Couvreur T. L. P. 1033 (WAG, YA); Fako Njonji Njonji along path to the lake Low canal forest, 4.15 ° N, 9.066 ° E, 26 September 2001, Kenfack D. 1545 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
343A917F1C997D6DE6DDA718D0F06465.taxon	description	Description. Scrambling shrubs or lianas, up to 30 m tall, d. b. h. up to 11 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 1 - 11 mm long, 1 - 3 mm in diameter; blade 3 - 35 cm long, 1.4 - 12.5 cm wide, linear to elliptic to ovate to obovate to oblong, apex acuminate to acute, base decurrent to subcordate, discolorous, whitish below; midrib sunken or flat; secondary veins 6 to 23 pairs; tertiary venation percurrent. Individuals unisexual or bisexual; inflorescences cauliflorous or ramiflorous on old or young foliate branches, axillary, leaf opposed or extra axillary. Flowers with (6 -) 9 perianth parts in 2 or 3 whorls, 1 to more than 50 per inflorescence with a peduncle from 1 to 70 mm long; pedicel 1 - 60 mm long; in fruit 2 - 55 mm long; bracts 2 to 3, one or two basal and one inserted on the pedicel at varying levels; sepals 3, valvate, free, 0.5 - 10 mm long, ovate to elliptic to triangular, apex acute or obtuse, base truncate; petals free; outer petals 3 or 6 (when petals in one whorl), valvate, 1 - 50 mm long, 1 - 25 mm wide, ovate to elliptic, apex acute to rounded, base truncate; inner petals (0 -) 3, valvate, 0.5 - 25 mm long, 0.3 - 10 mm wide, ovate to elliptic to rhombic, apex acute to rounded, base truncate; stamens 1 to more than 125, in 1 to 6 rows, 1 - 2 mm long, linear or cuneiform to clavate; anthers sometimes connate apically covering connectives, connective discoid to truncate to absent, glabrous or pubescent; staminodes 6 to 12 (13) or absent; carpels free, 3 to 150, 1 - 4 mm long, stigma bilobed or cylindrical or elongate and flattened at top, pubescent or glabrous. Monocarps sessile or stipitate, stipes 1 - 25 mm long; monocarps 2 to 25, 6 - 60 mm long, 5 - 25 mm in diameter, moniliform, cylindrical to ellipsoid, apex rounded to apiculate, smooth or verrucose or weakly torulose; seeds 6 - 25 mm long, 4 - 11 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6CAE94BC30515DBEAB461A733DA417FC.taxon	description	Map 6 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6CAE94BC30515DBEAB461A733DA417FC.taxon	materials_examined	Type. Democratic Republic of the Congo. Orientale; Yamboa, Louis J. L. P. 8957, 21 Apr 1938: lectotype, chosen by Hoekstra et al. (2021), p. 129: BR [BR 0000008805386]; isolectotypes: BR [BR 6102005255172, BR 0000008805379]; K [K 000913657]; P [P 00362792].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6CAE94BC30515DBEAB461A733DA417FC.taxon	description	Description. Liana, 30 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent with lenticels, young foliate branches yellow-brown, with dense appressed yellow-brown hairs 0.2 - 0.4 mm long. Leaves: petiole 3 - 5 mm long, 0.6 - 1 mm in diameter, densely pubescent, cylindrical, blade inserted on the side of the petiole; blade 6 - 12 cm long, 1.9 - 3.5 cm wide, oblong to elliptic, apex acuminate, acumen 0.5 - 1.5 cm long, base cuneate, papyraceous, below densely pubescent when young, densely pubescent to sparsely pubescent when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above sparsely pubescent to glabrous when young and old, below densely pubescent when young and old; secondary veins 11 to 14 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 15 - 60 mm long, 0.3 - 0.6 mm in diameter, sparsely pubescent to densely pubescent; basal bract ca. 1 mm long, ca. 0.5 mm wide; upper bract 0.5 - 1 mm long, 0.5 - 1 mm wide; sepals 3, valvate, free, ca. 1 mm long, 2 - 3 mm wide, ovate, apex acute, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner; inner petals entirely covered in bud; outer petals 3, 4 - 6.7 mm long, 4.6 - 6 mm wide, ovate, apex obtuse, base truncate, yellow to white, margins flat, densely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 2.5 - 2.8 mm long, 2.2 - 3 mm wide, rhombic, apex obtuse, base truncate, yellow to white, margins flat, pubescent outside, pubescent, base glabrous inside; stamens 15 to 17, in 1 row, ca. 1 mm long, oblong; connective truncate, glabrous, thecae converging apically; staminodes absent; carpels free, 6 to 9, ovary ca. 1 mm long, stigma deeply bilobed, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6CAE94BC30515DBEAB461A733DA417FC.taxon	distribution	Distribution. A central African species, known from Cameroon and the Democratic Republic of the Congo; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6CAE94BC30515DBEAB461A733DA417FC.taxon	materials_examined	Specimens examined. South Region: Route miniere E of Mt Elephant ca 20 km SE of Kribi, 2.78 ° N, 10.01 ° E, 07 April 1970, Bos J. J. 6735 (BR, K, LM, MO, P, WAG, YA); Oveng (Sangmelima), 3.09 ° N, 11.90 ° E, 22 March 1962, Breteler F. J. 2692 (B, BR, K, MO, P, U, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5A749A5FF1A040FA750CDA21CB5EFB60.taxon	description	Fig. 46; Map 6 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5A749A5FF1A040FA750CDA21CB5EFB60.taxon	materials_examined	Type. Democratic Republic of the Congo. no region; no location, Dewevre A. P. 785, no date: lectotype, sheet designated in Hoekstra et al. (2021), p. 133: BR [BR 0000024941433]; isolectotypes: BR [BR 0000008804020, BR 0000008804358].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5A749A5FF1A040FA750CDA21CB5EFB60.taxon	description	Description. Shrub to liana, 6 m tall, d. b. h unknown. Indumentum of simple hairs; old leafless branches glabrescent with lenticels, young foliate branches densely pubescent with erect reddish brown hairs 0.4 - 1.3 mm long. Leaves: petiole 3 - 6 mm long, ca. 1 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 4.5 - 12.5 cm long, 2 - 7 cm wide, oblong to elliptic, apex obtuse, base subcordate, papyraceous to subcoriaceous, below sparsely pubescent when young and old, above sparsely pubescent to glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below pubescent when young and old; secondary veins 7 to 11 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 11 - 24 mm long, ca. 1 mm in diameter, sparsely pubescent to densely pubescent; in fruit 12 - 22 mm long, ca. 1 mm in diameter; basal bracts when present 1 - 2 mm long, ca. 1 mm wide; upper bract 1 - 2 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 3 - 4 mm long, 5 - 7 mm wide, ovate, apex rounded, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 15 - 19 mm long, 8.4 - 11.8 mm wide, ovate, apex rounded to obtuse, base truncate, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, ca. 13 mm long, 4.5 - 5.5 mm wide, linear to elliptic, apex acute, base truncate, margins flat, densely pubescent outside, glabrous but pubescent towards margins inside; stamens 24 to 32, in 3 to 4 rows, ca. 1 mm long, obovate; connective truncate, glabrous; staminodes absent; carpels free, 27 to 38, ovary ca. 3 mm long, stigma cylindrical, glabrous. Monocarps stipitate, stipes 7 - 25 mm long, 2 - 3 mm in diameter; monocarps 1 to 25, 26 - 65 mm long, 5 - 8 mm in diameter, moniliform, cylindrical, apex rounded to apiculate, glabrous, verrucose to weakly torulose, constricted around seeds when more than 1, yellow turning orange when ripe; seeds 1 to 4 per monocarp, 14 - 21 mm long, 4 - 6 mm in diameter, subcylindrical; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5A749A5FF1A040FA750CDA21CB5EFB60.taxon	distribution	Distribution. A central African species, from Cameroon to Gabon, and the Central African Republic and the Democratic Republic of the Congo; in Cameroon known from the Adamaoua, East and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5A749A5FF1A040FA750CDA21CB5EFB60.taxon	materials_examined	Specimens examined. Adamaoua Region: rocher conglomeratique de Mbalarzi dans la vallee de la Mbere (65 km NE de Meiganga), 7 ° N, 14.51 ° E, 16 October 1963, Letouzey R. 6195 (P). East Region: km 17 of Betare Oya-Meiganga road 5 km SE of Ndokayo, 5.55 ° N, 14.1 ° E, 03 February 1966, Leeuwenberg A. J. M. 7716 (BR, K, MO, P, WAG, YA); Kongolo, 5.4 ° N, 14.03 ° E, 01 April 1914, Mildbraed G. W. J. 9010 (K). South-West Region: Piste Akwaya-Mamfe pres Makomono 7 km S Akwaya, 6.31 ° N, 9.557 ° E, 25 July 1975, Letouzey R. 14081 (MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F74570AF885E11F21BC3E8F12F7F048A.taxon	description	Figs 47, 48; Map 6 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F74570AF885E11F21BC3E8F12F7F048A.taxon	materials_examined	Type. Ivory Coast. no region; " l'est l'est du pays d'Attie " ", Angoulvant G. - L. s. n., 1910: lectotype, chosen by Hoekstra et al. (2021), p. 138: P [P 00362786]; isotypes: BM [BM 001125039]; E [E 00624344]; K [K 000041008]; P [P 00362784, P 00362785].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F74570AF885E11F21BC3E8F12F7F048A.taxon	description	Description. Shrub to liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with ascending to erect reddish brown hairs 0.2 - 0.3 mm long. Leaves: petiole 5 - 8 mm long, 1 - 3 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 7.4 - 19.7 cm long, 3.4 - 7.7 cm wide, oblong to oblanceolate, apex acuminate to acute, acumen 1.5 cm long, base cuneate to rounded, subcoriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib depressed, above sparsely pubescent to glabrous when young and old, below sparsely pubescent when young and old; secondary veins 7 to 12 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young and old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 6 - 9 mm long, 0.5 - 1 mm in diameter, densely pubescent; in fruit 11 - 16 mm long, 1 - 2 mm in diameter; basal bracts not seen, upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, ovate, apex obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals partly covered in bud; outer petals 3, 3.5 - 5 mm long, 2 - 3.8 mm wide, ovate, apex acute, base truncate, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 2.5 - 3.5 mm long, 1.5 - 2.4 mm wide, rhombic, apex acute, base truncate, light yellow to white, margins flat, pubescent outside, pubescent inside; stamens 8 to 10, in 1 row, 1 - 2 mm long, clavate, enlarged in the upper part, connective truncate covering the anthers, glabrous; staminodes absent; carpels free, 4 to 6, ovary 7 - 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 3 - 6 mm long, 2 - 3 mm in diameter; monocarps 1 to 6, 20 - 79 mm long, 8 - 10 mm in diameter, moniliform, ellipsoid, apex ellipsoid, pubescent, tuberculate, weakly torulose, color unknown; seeds 1 to 6 per monocarp, ca. 12 mm long, 7 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F74570AF885E11F21BC3E8F12F7F048A.taxon	distribution	Distribution. A west and central African species with a disjunct distribution between Ivory Coast and Cameroon to Gabon, Central African Republic and the Democratic Republic of the Congo; in Cameroon known from Adamaoua and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F74570AF885E11F21BC3E8F12F7F048A.taxon	materials_examined	Specimens examined. Adamaoua Region: A 4 km au SO de Dir pres de Bagodo, 6.42 ° N, 13.38 ° E, 30 July 1966, Letouzey R. 7570 (P, YA). West Region: Pres Kongi (10 km au NO de Kimi bankim sur route Fouman-Banyo), 6.09 ° N, 11.26 ° E, 27 June 1967, Letouzey R. 8738 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E13213F0DDC3A092A87B93CE11A5EAF5.taxon	description	Figs 49, 62; Map 6 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E13213F0DDC3A092A87B93CE11A5EAF5.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Thomson W. C. s. n., 1963: lectotype, chosen by Le Thomas (1969 b), p. 218: K [K 000198911].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E13213F0DDC3A092A87B93CE11A5EAF5.taxon	description	Description. Liana, 25 m tall, d. b. h. up to 4 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent with very short reddish brown hairs 0.05 - 0.1 mm long. Leaves: petiole 5 - 9 mm long, 1 - 2 mm in diameter, densely pubescent, weakly grooved, blade inserted on top of the petiole; blade 12.5 - 15.2 cm long, 4.8 - 5.8 cm wide, oblong to oblanceolate, apex acuminate to acute, acumen ca. 1 cm long, base cuneate to rounded, papyraceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above densely pubescent when young and old, below sparsely pubescent when young, sparsely pubescent to glabrous when old; secondary veins 11 to 16 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals unisexual, monoecious, dimorphic, female inflorescences cauliflorous, male ones axillary, ramiflorous. Flowers with 9 perianth parts in 3 whorls, male and female flowers dimorphic, male inflorescences with 1 to 6 flowers, female inflorescence a condensed panicle to 12 cm in diameter with many flowers; male flowers: pedicel 5 - 6 mm long, ca. 1 mm in diameter, densely pubescent; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, triangular, apex acuminate, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 1.2 - 2.3 mm long, 2 - 3 mm wide, broadly ovate, apex obtuse, base truncate, margins flat, densely pubescent outside, pubescent with short papillose hairs inside; inner petals 3, valvate, 0.5 - 0.7 mm long, 0.3 - 0.5 mm wide, elliptic, apex obtuse, base truncate, margins flat, pubescent outside, pubescent inside; stamens 6, in 1 row but grouped by 2, ca. 1 mm long, oblong; connective truncate, pubescent; staminodes 12 (to 13), in one whorl externally to the stamens, very short 0.2 mm long, sparsely pubescent; female flowers: pedicel 6 - 20 mm long, ca. 1 mm in diameter, densely pubescent; in fruit 30 - 40 mm long, 1 - 2 mm in diameter; basal bract 1 - 1.5 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, ovate to lanceolate, apex acuminate, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 3 - 3.5 mm long, 3.5 - 4.0 (5.0) mm wide, broadly ovate, apex obtuse, base truncate, margins flat, densely pubescent outside, pubescent with short papillose hairs inside; inner petals 3, valvate, 1.3 - 1.4 mm long, 0.5 - 0.6 mm wide, elliptic, apex obtuse, base truncate, margins flat, pubescent outside, pubescent inside;, ca. 1 mm long, stigma elongate, glabrous, carpels free, 48 to 60 (76), ovary 1 - 1.5 mm long, densely hairy. Monocarps stipitate, stipes 6 - 12 mm long, 2 - 3 mm in diameter; monocarps ca. 8, 15 - 57 mm long, 6 - 9 mm in diameter, moniliform, ellipsoid, apex apiculate, pubescent, smooth to verrucose, constricted around seeds when more than 1, color unknown; seeds 1 to 4 per monocarp, 12 - 15 mm long, 5 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E13213F0DDC3A092A87B93CE11A5EAF5.taxon	distribution	Distribution. A central African species, from Nigeria to Gabon and in Equatorial Guinea; in Cameroon known from the Central, Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E13213F0DDC3A092A87B93CE11A5EAF5.taxon	materials_examined	Specimens examined. Central Region: Ndanan I to Ndangan I, 3.62 ° N, 11.58 ° E, 10 March 2004, Cheek M. 11657 (K, YA). Littoral Region: Douala-Edea Reserve Lake Tissongo study area 3.78 ° N, 10.04 ° E, 01 June 1976, Waterman P. G. 840 (E, K). South Region: Kribi-Lolodorf, 3.17 ° N, 10.48 ° E, 05 March 1969, Bos J. J. 4069 (BR, K, P, WAG, YA); 11 km from Kribi Ebolowa road, 2.88 ° N, 9.983 ° E, 02 January 1970, Bos J. J. 6037 (BR, C, K, LD, P, P, WAG, YA); 20 km From Kribi 5 km N of Lolodorf road, 3.05 ° N, 10.05 ° E, 09 February 1970, Bos J. J. 6293 (BR, P, WAG); Mt Elephant SE of Kribi, 2.81 ° N, 10.01 ° E, 28 April 1970, Bos J. J. 6867 (BR, P, WAG); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 11 February 2015, Couvreur T. L. P. 676 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 705 (WAG, YA); C. 15 km South of Ebolowa, 2.49, 11.10, 28 February 1964, de Wilde J. J. F. E 1987 (YA, WAG); Ca 15 km S of Ebolowa, 2.83 ° N, 11.16 ° E, 28 February 1964, de Wilde W. J. J. O 1987 (BR, P, WAG, YA); Aloum, 2.76 ° N, 10.69 ° E, 13 December 2013, Kamdem N. 164 (YA); Pres de Nsola (20 km N Bipindi), 3.16 ° N, 10.38 ° E, 29 January 1974, Letouzey R. 12811 (K, P, P, YA); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 356 (M, P, U, WAG); Bipindi, 3.05 ° N, 10.25 ° E, 01 January 1909, Zenker G. A. 3898 (BR, E, K). South-West Region: Kupe village, 4.76 ° N, 9.691 ° E, 08 November 1995, Cheek M. 7711 (K, MO, WAG, YA); Summit of Mt Loh, 5 ° N, 9.683 ° E, 17 November 1998, Gosline W. G. 175 (K, YA); Korup National Park, 5.01 ° N, 8.833 ° E, 01 January 1985, Thomas D. W. 4324 (K, MO, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E77BE4502AC6C695606D77C77D980A8B.taxon	description	Figs 49, 50; Map 6 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E77BE4502AC6C695606D77C77D980A8B.taxon	materials_examined	Type. Gabon. no region; Ogooue, Thollon F. - R. 813, Jul 1887: lectotype, chosen by Le Thomas (1969 b), p. 258: P [P 00362762, P 00362763, P 00362766]; isolectotypes: K [K 000198992]; MA [MA 630761, MA 698356]; WAG [WAG 0003586, WAG 0003587].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E77BE4502AC6C695606D77C77D980A8B.taxon	description	Description. Shrub to liana, 2 - 5 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with appressed pale brown hairs 0.7 - 1.2 mm long. Leaves: petiole 3 - ca. 8 mm long, ca. 1 mm in diameter, densely pubescent, grooved, blade inserted on top of the petiole; blade 7.5 - 25 cm long, 1.9 - 6 cm wide, oblong to oblanceolate or elliptic, apex acuminate to acute, acumen ca. 1.5 cm long, base cuneate to cordate, papyraceous, below densely pubescent with appressed silky white hairs when young, sparsely pubescent when old, above sparsely pubescent to glabrous when young, glabrous when old, discolorous, whitish below; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 9 to 16 pairs, pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, terminal or leaf opposed. Flowers with 9 perianth parts in 2 whorls, 4 to 10 per inflorescence, in 6 - 13 cm long racemes; pedicel up to ca. 12 mm long, ca. 1 mm in diameter, sparsely pubescent to densely pubescent; in fruit same as in flower; basal bract 2 - 3 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 1 - 3 mm long, 1 - 2 mm wide, triangular to ovate, apex acute, base truncate, densely pubescent outside, pubescent inside, margins flat; petals free, 6, in one whorl, 3 - 4 mm long, 1 - 2 mm wide, elliptic to ovate, apex obtuse, base truncate, green to light yellow, margins flat, densely pubescent outside, pubescent inside; stamens 6, in 1 row, opposite with the petals, ca. 1 mm long, obconic; connective truncate, glabrous; staminodes 6, alternating with stamens, ca. 1 mm long; carpels free, 15 to 21, ovary ca. 1 mm long, stigma bilobed, glabrous. Monocarps stipitate, stipes 2 - 3 mm long, 2 - 3 mm in diameter; monocarps up to 4, 12 - 15 mm long, 9 - 10 mm in diameter, ellipsoid to subglobose, apex rounded, densely pubescent, smooth, dull orange to red when ripe; seed 1 per monocarp; 8 - 9 mm long, 7 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E77BE4502AC6C695606D77C77D980A8B.taxon	distribution	Distribution. A central African species, in Cameroon and Gabon; in Cameroon known from the Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E77BE4502AC6C695606D77C77D980A8B.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On east trail, 4.36 ° N, 10.25 ° E, 15 February 2013, Couvreur T. L. P. 626 (WAG, YA). South-West Region: Etinde Upper Boando footpath to west of village, 4.05 ° N, 9.15 ° E, 06 December 1993, Cable S. 404 (K); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 26 March 2016, Couvreur T. L. P. 1018 (WAG, YA); on trail leading to top of Mt Etinde after Ekonjo village, 4.06 ° N, 9.151 ° E, 01 April 2016, Couvreur T. L. P. 1025 (WAG, YA); Mokoko Forest Reserve Dikome, 4.48 ° N, 9.033 ° E, 05 May 1994, Ekema S. N. 939 (K, YA); Banyang-mbo Sanctuary, 5.35 ° N, 9.541 ° E, 02 December 2000, Gosline W. G. 300 (K); district Buea Upper Boando, 4.03 ° N, 9.106 ° E, 27 March 1992, Kwangue A. T. 22 (K, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B1223BC0071DCEA123F3B1065F5B6095.taxon	description	Fig. 51; Map 6 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B1223BC0071DCEA123F3B1065F5B6095.taxon	materials_examined	Type. Cameroon. Central Region; Ottotomo Forest Reserve, Couvreur T. L. P. 762, 24 Apr 2015: holotype: WAG [WAG. 1576998, WAG. 1576999, WAG. 1577000]; isotypes: MPU [MPU 1374962]; YA n. v.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B1223BC0071DCEA123F3B1065F5B6095.taxon	description	Description. Liana, up to 20 m tall, d. b. h. up to 4 cm. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent with dense ascending reddish brown hairs 0.1 - 0.2 mm long. Leaves: petiole 3 - 5 mm long, ca. 1 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 4.5 - 12 cm long, 1.5 - 4.5 cm wide, oblong to obovate, apex acuminate to acute, acumen ca. 1 cm long, base cuneate to rounded, papyraceous, below sparsely pubescent when young and old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below pubescent when young and old; secondary veins 7 to 11 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous or more rarely ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 (3) per inflorescence when on branches, up to 20 per inflorescence when on main trunk; pedicel 4 - 20 mm long, ca. 1 mm in diameter, sparsely pubescent; in fruit unknown; basal bract not seen, upper bract minute, ca. 0.5 mm, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, 1 mm wide, triangular, apex acute, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals free, subequal, inner petals partly covered in bud; outer petals 3, 3.5 - 5 mm long, 2 - 3.5 mm wide, elliptic to ovate, apex obtuse, base truncate, light yellow to white, margins flat, densely pubescent outside, sparsely pubescent inside; inner petals 3, valvate, 3 - 4.5 mm long, 1.2 - 1.5 mm wide, elliptic to ovate, apex acute, base truncate, light yellow to white, margins flat, pubescent outside, glabrous inside; stamens 13 to 15, in 1 row, basally fused between each them, ca. 1 mm long, linear; connective truncate to rounded, pubescent, cream; staminodes absent; carpels free, 9 to 12, ovary ca. 1 mm long, stigma globose, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B1223BC0071DCEA123F3B1065F5B6095.taxon	distribution	Distribution. endemic to Cameroon; known from the Central Region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B1223BC0071DCEA123F3B1065F5B6095.taxon	materials_examined	Specimens examined. Central Region: Ottotomo Forest Reserve 45 km South of Yaounde ca 5 km on main path into reserve, 3.65 ° N, 11.28 ° E, 24 April 2015, Couvreur T. L. P. 762 (WAG, YA); Reserve Forestiere d'Ottotomo 40 km de Yaounde sur la route de Kribi, 3.64 ° N, 11.27 ° E, 05 May 1970, Farron C. 7266 (P); Reserve forestiere d'Ottotomo Yaounde a environ 40 km au SW sur la route de Makak, 3.64 ° N, 11.27 ° E, 26 May 1970, Farron C. 7359 (YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
96E735FE11A569D482F0AE8EFECAD7C8.taxon	description	Figs 52, 53; Map 7 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
96E735FE11A569D482F0AE8EFECAD7C8.taxon	materials_examined	Type. Gabon. Estuaire; Gaboon River, Mann G. 960, Jul 1861: lectotype, chosen by Le Thomas (1969 b), p. 216: K [K 000198989].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
96E735FE11A569D482F0AE8EFECAD7C8.taxon	description	Description. Liana, 25 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent with dense pubescence of erect to curling yellow-brown hairs 0.4 - 0.6 mm long. Leaves: petiole 3 - 5 mm long, ca. 2 mm in diameter, densely pubescent, weakly grooved adaxially, blade inserted on top of the petiole; blade 8.5 - 18.5 cm long, 3.5 - 7 cm wide, oblong to obovate, apex acute to rounded, base rounded to subcordate, papyraceous to subcoriaceous, below densely pubescent when young and old with erect yellow hairs, above sparsely pubescent when young and old, discolorous, whitish below; midrib depressed, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 15 to 19 pairs, glabrous above; tertiary venation percurrent. Individuals unisexual, monoecious, dimorphic, male inflorescences ramiflorous and axillary, female ones cauliflorous. Flowers with 9 perianth parts in 3 whorls, male inflorescence with a solitary to a few-flowered fascicle, sometimes a rhipidium with up to 25 flowers per inflorescence, female a many-flowered panicle-like rhipidia; male flowers: pedicel 2 - 5 mm long, ca. 0.5 mm in diameter, densely pubescent; basal bract ca. 0.5 mm long, 0.5 mm wide; upper bract ca. 1 mm long, ca. 0.5 mm wide; sepals 3, valvate, shortly basally fused, ca. 0.5 mm long, ca. 0.5 mm wide, ovate, apex obtuse, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, ca. 1.6 mm long, ca. 2 mm wide, broadly ovate, apex obtuse, base truncate, yellow, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 0.5 - 0.8 mm long, 0.3 - 0.4 mm wide, broadly elliptic, apex rounded, base truncate, margins flat, pubescent outside, pubescent inside; stamens 6, in 1 row, ca. 1 mm long, oblong; connective truncate, pubescent; staminodes 12, in one whorl externally to the stamens, 0.2 - 0.3 mm long, sparsely pubescent. Female flowers: pedicel 15 - 25 mm long, ca. ca. 1 mm in diameter, densely pubescent with erect yellowish brown hairs; in fruit 15 - 45 mm long, 1 - 2 mm in diameter; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract not seen; upper bract 1 - 2 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 1.2 - 1.7 mm long, 1.3 - 1.4 mm wide, broadly ovate to broadly elliptic, apex acuminate, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 3 - 5 mm long, 3.5 - 5.4 mm wide, broadly ovate to circular, apex obtuse or rounded, base truncate, margins flat, densely pubescent with brown hairs outside, pubescent with very short yellowish hairs inside; inner petals 3, valvate, 1.3 - 1.7 mm long, 0.6 - 0.7 mm wide, elliptic, apex obtuse, base truncate, margins flat, pubescent with very short yellowish hairs outside, pubescent with very short yellowish hairs inside; carpels free, 80 to 100, ovary 1 - 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 6 - 14 mm long, 2 - 3 mm in diameter; monocarps up to 14, 23 - 55 mm long, 7 - 8 mm in diameter, moniliform, ellipsoid, apex apiculate, pubescent, verrucose, constricted around seeds when more than 1, green when ripe; seeds 1 to 5 per monocarp, 12 - 15 mm long, 6 - 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
96E735FE11A569D482F0AE8EFECAD7C8.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of the Congo; in Cameroon known from the Littoral and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
96E735FE11A569D482F0AE8EFECAD7C8.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On east trail, 4.36 ° N, 10.25 ° E, 15 February 2013, Couvreur T. L. P. 629 (WAG, YA). South Region: 27 km on an exploitation track leading from Ipono towards Dipikar island Bongala river, 2.28 ° N, 11.13 ° E, 26 June 1975, de Wilde J. J. F. E 8330 (BR, MO, P, WAG, YA); Besou 45 km E of Gross Batanga, 2.85 ° N, 10.3 ° E, 22 July 1911, Mildbraed G. W. J. 6059 (HBG); Bipindi, 3.05 ° N, 10.25 ° E, June 1899, Zenker G. A. 2102 (B, G, K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EBC7EF2780AA9680030689395396138.taxon	description	Map 7 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EBC7EF2780AA9680030689395396138.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2473, Dec 1901: lectotype, chosen by Hoekstra et al. (2021), p. 147: B [B 100154098]; isolectotypes: B [B 100154096, B 100154097, B 100154099]; BM [BM 001125043]; BR [BR 000008801388]; COI [COI 00071518]; E; G [G 00308364]; GOET [GOET 005688, GOET 005689]; HBG [HBG 502481]; K [K 000198948]; KFTA [KFTA 0001554, KFTA 0001555]; L [L 0182291]; M [M- 0240178]; P [P 00363342, P 00363343, P 01988326]; S [S 07 - 13404]; WAG [WAG 0057970]; WU [WU 0025876].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EBC7EF2780AA9680030689395396138.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with orange-brown hairs ca. 0.5 mm long. Leaves: petiole 6 - 11 mm long, ca. 2 mm in diameter, densely pubescent, grooved, blade inserted on top of the petiole; blade 10.5 - 21.5 cm long, 3.5 - 4 cm wide, narrowly oblong to oblanceolate, apex acuminate, acumen 0.5 - 2.5 cm long, base subcordate, subcoriaceous to membranous, below glabrous when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above sparsely pubescent when young and old, below sparsely pubescent when young and old; secondary veins 7 to 14 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence, pedicel 15 - 27 mm long, 1 - 2 mm in diameter, pubescent; in fruit 15 - 30 mm long, ca. 3 mm in diameter, pubescent; basal bract ca. 4 mm long, ca. 4 mm wide; upper bract ca. 5 mm long, ca. 4 mm wide; sepals 3, valvate, free, ca. 4 mm long, ca. 4 mm wide, triangular, apex acute, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 10 - 15 mm long, 8 - 9 mm wide, ovate, apex obtuse, base truncate, margins flat, densely pubescent outside, glabrous inside; inner petals 3, valvate, ca. 8 mm long, ca. 8 mm wide, broadly ovate, apex obtuse, base truncate, margins flat, pubescent outside, glabrous inside; stamens ca. 65, in 3 rows, ca. 0.5 mm long, cylindrical; connective rounded, glabrous; staminodes absent; carpels free, ca. 41, ovary ca. 2 mm long, stigma globose, glabrous. Monocarps stipitate, stipes ca. 3 mm long, ca. 2 mm in diameter; monocarps ca. 6, ca. 45 mm long, 10 - 12 mm in diameter, moniliform, ellipsoid to cylindrical, apex rounded, densely pubescent, rugulose, constricted around seeds when more than 1, brown when unripe; seeds 1 to 2 per monocarp, ca. 15 mm long, ca. 9 mm in diameter, subcylindrical; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EBC7EF2780AA9680030689395396138.taxon	distribution	Distribution. endemic to Cameroon; known from the Central and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2EBC7EF2780AA9680030689395396138.taxon	materials_examined	Specimens examined. Central Region: 60 km SW of Eseka S of Nyong R 12 km W of Songbong, 3.47 ° N, 10.5 ° E, 10 March 1965, Leeuwenberg A. J. M. 5088 (BR, K, MO, P, WAG, YA). South Region: Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1902, Zenker G. A. 2473 (L, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6F175C7098294D1AC00FA8BC3FF8C303.taxon	description	Fig. 49; Map 7 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6F175C7098294D1AC00FA8BC3FF8C303.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 1321, 23 Mar 1897: holotype: B [B 100153023]; isotypes: BM [BM 000553832, BM 000553833]; E [E 00181437]; G [G 00308366]; HBG [HBG- 502506]; K [K 000198990, K 000198991]; M [M- 0198713]; P [P 00362605, P 00362606].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6F175C7098294D1AC00FA8BC3FF8C303.taxon	description	Description. Liana, up to 4 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent with dense appressed to ascending reddish brown hairs 0.1 - 0.2 mm long. Leaves: petiole 3 - 5 mm long, ca. 1 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 4.6 - 15.4 cm long, 2.3 - 4.6 cm wide, narrowly oblong-elliptic to oblong-oblanceolate (to obovate), apex acute to acuminate, acumen ca. 3 cm long, base rounded to subcordate, papyraceous, below sparsely pubescent when young and old, above sparsely pubescent when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 11 to 14 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on foliate branches, leaf opposed to extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 8 per inflorescence, glomerule-like; pedicel 1 - 2 mm long, ca. 1 mm in diameter, densely pubescent with appressed yellowish brown hairs; in fruit 4 - 5 mm long, ca. 1 mm in diameter; basal bract ca. 1 mm long, ca. 0.5 mm wide; upper bract ca. 1 mm long, ca. 0.5 mm wide; sepals 3, valvate, free, ca. 1 mm long, 1 - 2 mm wide, triangular to ovate, apex acute, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 2.6 - 4.3 mm long, 2.6 - 3.5 mm wide, ovate, apex acute, base truncate, yellow to cream, margins flat, densely pubescent outside, glabrous, pubescent towards margins inside; inner petals 3, valvate, ca. 3.5 mm long, ca. 2 mm wide, rhombic, apex acute, base truncate, yellow to cream, margins flat, pubescent towards base outside, pubescent inside; stamens 9, in 1 row, ca. 1 mm long, linear to oblong; connective truncate, glabrous; staminodes 6, alternating with the stamens, minute, glabrous; carpels free, 12 to 20, ovary ca. 1 mm long, stigma globose, glabrous. Monocarps stipitate, stipes ca. 2 mm long, ca. 2 mm in diameter; monocarps 2 to 8, 9 - 16 mm long, 5 - 6 mm in diameter, ellipsoid, apex apiculate, glabrous, verrucose, constricted around seeds when more than 1, color unknown; seeds 1 to 2 per monocarp, 7 - 9 mm long, 5 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6F175C7098294D1AC00FA8BC3FF8C303.taxon	distribution	Distribution. endemic to Cameroon, known from the Central and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6F175C7098294D1AC00FA8BC3FF8C303.taxon	materials_examined	Specimens examined. Central Region: Riverine forest bank Nyong river near the new bridge ca 65 km SSW of Eseka, 3.46 ° N, 10.5 ° E, 16 June 1964, de Wilde W. J. J. O 2709 (B, BR, K, MO, P, WAG, YA); Natchigal ca 20 km N of Obala, 4.35 ° N, 11.63 ° E, 01 July 1964, de Wilde W. J. J. O 2787 (B, BR, K, MO, P, WAG, YA). South Region: Bipinde, 3.26 ° N, 10.20 ° E, 20 June 1918, Annet E. 348 (BR, K, P); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 704 (WAG, YA); Bipindi, 3.08 ° N, 10.41 ° E, 1897, Zenker G. A. 132 (L); Bipindi, 3.08 ° N, 10.41 ° E, 1897, Zenker G. A. 1321 (E, M, P); Lokoundje, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 199 (U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1904, Zenker G. A. 2693 (E, L, M, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1911, Zenker G. A. 4000 (E, L, M, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1912, Zenker G. A. 4477 (E, L, M); Bipindi, 3.05 ° N, 10.25 ° E, Zenker G. A. s. n. (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0F681D80441CA360EAE5D3477B6C2AB.taxon	description	Figs 53, 54; Map 7 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0F681D80441CA360EAE5D3477B6C2AB.taxon	materials_examined	Type. Type. Democratic Republic of the Congo. Nord Kivu; Fort Beni a Semliki, Mildbraed G. W. J. 2213, 1907 - 1908: holotype: B [B 100153056].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0F681D80441CA360EAE5D3477B6C2AB.taxon	description	Description. Liana, up to 15 m tall or up to canopy, d. b. h. to 6 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with dense erect dark-brown hairs 0.9 - 1.4 mm long. Leaves: petiole 3 - 4 mm long, 1 - 2 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 10.8 - 35 cm long, 3.3 - 7.5 cm wide, narrowly oblong to narrowly oblanceolate, apex acuminate to acute, acumen up to 5 cm long, base rounded to subcordate, subcoriaceous to membranous, below whitish blue, densely pubescent to pubescent with erect brown hairs when young, pubescent to glabrous when old, above pubescent when young, sparsely pubescent to glabrous when old, discolorous, whitish below; midrib impressed, above densely pubescent when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 11 to 20 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, (1) 2 to 5 per inflorescence; pedicel 18 - 22 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 18 - 22 mm long, 1 - 2 mm in diameter; basal bract ca. 2 mm long, 2 - 3 mm wide; upper bract 3 - 5 mm long, ca. 4 mm wide; sepals 3, valvate, basally fused, 3 - 5 mm long, 5 - 8 mm wide, ovate, apex rounded, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 12 - 22 mm long, 7 - 14 mm wide, elliptic to ovate, apex obtuse, base truncate, brown-violet, margins flat, densely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 9 - 14 mm long, 8 - 10 mm wide, ovate to rhombic, apex acute, base truncate, brown-violet, margins flat, glabrous but pubescent towards base outside, glabrous but pubescent towards the base inside; stamens 90 to 110, in 3 to 4 rows, ca. 1 mm long, cuneate; connective truncate, glabrous; staminodes absent; carpels free, 40 to 60, ovary ca. 3 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 2 - 5 mm long, 2 mm in diameter; monocarps 5 to 15, 14 - 18 mm long, 34 mm in diameter, moniliform, ellipsoid, apex rounded to apiculate, densely pubescent, smooth, constricted around seeds when more than 1, glaucous green when ripe; seeds 1 to 2 (3) per monocarp, 11 - 12 mm long, 7 - 11 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0F681D80441CA360EAE5D3477B6C2AB.taxon	distribution	Distribution. A widespread west and central African species, from Guinea to Ivory Coast, and from Cameroon to the Republic of the Congo, the Democratic Republic of the Congo, Central African Republic and Uganda; in Cameroon recorded from Adamaoua, Central, East, Littoral, South, South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0F681D80441CA360EAE5D3477B6C2AB.taxon	materials_examined	Selected specimens examined. Adamaoua Region: Mbakaou, 6.00 ° N, 12.88 ° E, 12 January 2017, Kamdem N. 465 (YA). Central Region: Mefou proposed national park Near Mefou town, 3.62 ° N, 11.58 ° E, 08 March 2004, Cheek M. 11499 (K, YA); Ottotomo Forest reserve 7 km north-west from Ngoumou 30 km south west from Yaounde, 3.65 ° N, 11.28 ° E, 24 February 2016, Couvreur T. L. P. 986 (WAG, YA); Ngoro, 5.07 ° N, 11.22 ° E, 29 April 2017, Kamdem N. 499 (YA). East Region: 18 km NW of Doume along road to Nguelemendouka, 4.23 ° N, 13.45 ° E, 24 November 1961, Breteler F. J. 2137 (BR, P, WAG, YA); 75 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.73 ° E, 05 March 2019, Couvreur T. L. P. 1201 (MPU, WAG, YA); Palisco forest consession 15 km along main road into consession, 3.48 ° N, 13.59 ° E, 27 March 2015, Couvreur T. L. P. 754 (WAG, YA); Deng Deng, 5.20 ° N, 13.13 ° E, 27 July 2014, Kamdem N. 167 (YA); Route Bertoua-Deng Deng a 6 km au Sud de Mambaya, 4.91 ° N, 13.76 ° E, 26 January 1961, Letouzey R. 3248 (P, YA); A 15 km au S de Djouo (20 km E de Somalomo sur le Dja), 3.32 ° N, 12.93 ° E, 23 February 1962, Letouzey R. 4359 (P, YA); A 20 km au S de Mvoy I (45 km a l'Est de Yokadouma), 3.31 ° N, 15.51 ° E, 16 May 1963, Letouzey R. 5071 (P, YA). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1176 (MPU, WAG, YA); Ebo Wildlife Reserve Djuma permanent camp On Djashaka trail, 4.35 ° N, 10.24 ° E, 13 February 2014, Couvreur T. L. P. 618 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.91 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 653 (WAG, YA); Ebo forest reserve ca 2500 m on Dicam trail from Bekob camp, 4.34 ° N, 10.40 ° E, 11 March 2007, Wieringa J. J. 5898 (WAG). South Region: 20 km from Kribi 2 km N of Lolodorf road, 3.01 ° N, 10.05 ° E, 12 December 1969, Bos J. J. 5818 (WAG); 24 km from Kribi ca 3 km N of Lolodorf road, 3.03 ° N, 10.08 ° E, 31 March 1970, Bos J. J. 6653 (BR, K, LD, LM, MO, P, WAG, YA); 20 km east from Lele village, 2.27 ° N, 13.33 ° E, 07 September 2013, Couvreur T. L. P. 466 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.33 ° E, 11 February 2015, Couvreur T. L. P. 669 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.33 ° E, 12 February 2015, Couvreur T. L. P. 691 (WAG, YA); Ebom, 3.1 ° N, 10.71 ° E, 27 February 1997, Parren M. P. E. 23 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 May 1913, Zenker G. A. 357 (M, P, U, WAG). South-West Region: Kupe village, 4.76 ° N, 9.694 ° E, 21 May 1996, Cable S. 2523 (K); Gully by Daniel Ajang's saprophyte site, 4.78 ° N, 9.716 ° E, 07 July 1996, Cable S. 3683 (K, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.502 ° E, 25 March 2016, Couvreur T. L. P. 1003 (WAG, YA); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.057 ° E, 02 April 2016, Couvreur T. L. P. 1037 (WAG, YA); on forest trail north of Ngomboku village, 4.91 ° N, 9.730 ° E, 06 April 2016, Couvreur T. L. P. 1065 (WAG, YA); Etome, 4.05 ° N, 9.116 ° E, 31 January 1997, Nning J. 212 (K, MO, YA); Bakingili, 4.06 ° N, 9.033 ° E, 15 February 1997, Nning J. 259 (K, YA); Mahole, 4.81 ° N, 9.615 ° E, 29 November 1999, Onana J. M. 947 (K, MO, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
154CC1B87DAADA73EEFA00356B8B071D.taxon	description	Map 7 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
154CC1B87DAADA73EEFA00356B8B071D.taxon	materials_examined	Type. Angola. Cuanza Norte; Golungo Alto, Welwitsch F. M. J. 761, Jul 1855: lectotype, designated by Paiva (1966), p. 41: LISU [LISU 206061]; isolectotypes: B [B 100153029]; BM [BM 000553834, BM 000553835]; BR [BR 0000008805324]; COI [COI 00004905]; G [G 00308369]; K [K 000198968]; LISU [LISU 206062]; P [P 00362602].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
154CC1B87DAADA73EEFA00356B8B071D.taxon	description	Description. Shrub to liana, 3 - 6 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with dense erect reddish brown hairs 0.5 - 0.9 mm long. Leaves: petiole 4 - 8 mm long, 1 - 2 mm in diameter, densely pubescent, cylindrical, blade inserted on top of the petiole; blade 3.6 - 17.2 cm long, 1.8 - 6.8 cm wide, obovate to oblong-elliptic, apex acute, base subcordate, papyraceous, below pubescent with erect reddish brown hairs when young, sparsely pubescent when old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below densely pubescent when young, pubescent when old; secondary veins 7 to 15 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed to extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 13 - 36 mm long, 0.5 - 1 mm in diameter, pubescent; in fruit 5 - 27 mm long, 1 mm in diameter; bracts 2, one basal, soon falling, and one upper towards the middle of pedicel or lower half of pedicel, basal bract not seen, upper bract 2 - 6 mm long, 1 - 5 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 3 mm wide, ovate to semiorbicular, apex rounded, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 5.8 - 6.7 mm long, 4.5 - 6 mm wide, ovate, apex obtuse, base truncate, pale yellow, margins flat, densely pubescent outside, pubescent towards the margins inside; inner petals 3, valvate, 3 - 5.4 mm long, 2.7 - 2.9 mm wide, elliptic to ovate, apex obtuse, base truncate, pale yellow, margins flat, densely pubescent outside, pubescent with a glabrous base inside; stamens 22 to 25, in 3 rows, 1 - 2 mm long, obovate; connective truncate, glabrous; staminodes absent; carpels free, 12 to 24, ovary 1 - 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 3 - 7 mm long, 2 - 3 mm in diameter; monocarps 2 to 17, 12 - 35 mm long, 6 - 9 mm in diameter, moniliform, ellipsoid, apex rounded to apiculate, glabrous, verrucose to weakly torulose, constricted around seeds when more than 1, orange to red when ripe; seeds 1 to 3 (to 5) per monocarp, 7 - 8 mm long, 5 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
154CC1B87DAADA73EEFA00356B8B071D.taxon	distribution	Distribution. A widespread species in Central and East Africa, with a disjunct distribution between Cameroon, Gabon, the Republic of the Congo and western Democratic Republic of the Congo and eastern Democratic Republic of the Congo, Uganda, Burundi, Rwanda, Ethiopia, and western Tanzania and Kenya; in Cameroon known from the East and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
154CC1B87DAADA73EEFA00356B8B071D.taxon	materials_examined	Specimens examined. East Region: Letta vers Bertoua, 4.55 ° N, 13.49 ° E, 07 February 1960, Letouzey R. 2955 (P, P, YA). South Region: 25 km on the road from N'Koemvone to Akoakas Akoakas rock, 2.71 ° N, 11.28 ° E, 18 July 1975, de Wilde J. J. F. E 8371 (BR, MO, P, WAG); Rocher d'Ako'Akas 25 km SE d'Ebolowa sur piste d'Evindissi, 2.43 ° N, 11.18 ° E, 04 February 1970, Letouzey R. 10007 (IFAN, MO, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D96111349BD066A74F0165DA902D76A6.taxon	description	Figs 55, 56; Map 7 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D96111349BD066A74F0165DA902D76A6.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2985, Apr 1904: holotype: B not seen; isotypes: BR [BR 0000008804297]; E [E 00624352]; GOET [GOET 005686]; HBG [HBG- 502505]; K [K 000198988]; L [L 0038042]; M [M 0107931]; P [P 00362600]; S [S 07 - 13262]; WAG [WAG 0071434]; WU [WU 0025872] ..	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D96111349BD066A74F0165DA902D76A6.taxon	description	Description. Liana, 10 m tall, d. b. h. up to 10 cm. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent with dense erect reddish brown hairs 0.7 - 1.4 mm long. Leaves: petiole 3 - 6 mm long, 1 - 3 mm in diameter, densely pubescent with erect reddish brown hairs, grooved, blade inserted on top of the petiole; blade 12.5 - 28.6 cm long, 4.1 - 12.5 cm wide, obovate to oblanceolate, apex acuminate to acute, acumen 0.5 - 1 cm long, base rounded to subcordate, subcoriaceous, below sparsely pubescent with erect yellowish hairs when young and old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below densely pubescent when young and old; secondary veins 12 to 19 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed to axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 10 per inflorescence; pedicel 1 - 18 mm long, ca. 1 mm in diameter, pubescent with erect reddish brown hairs; in fruit 9 - 31 mm long, 1 - 2 mm in diameter; basal bract ca. 2 mm long, ca. 1 mm wide; upper bract 1 - 8 mm long, 1 - 4 mm wide; sepals 3, valvate, free, 5 - 10 mm long, 3 - 6 mm wide, elliptic to ovate, apex acute, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 8 - 22 mm long, 4.7 - 8.5 mm wide, ovate, apex acute to attenuate, base truncate, yellow-brown bright white towards the base and inside, margins flat, densely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 6.5 - 14 mm long, 3 - 5 mm wide, elliptic to ovate, apex acute to attenuate, base truncate, yellow-brown bright white towards the base and inside, margins flat, densely pubescent outside, pubescent with a glabrous base inside; stamens 17 to 46, in 2 to 4 rows, ca. 2 mm long, linear; thecae convergent apically hiding the connective apically, white, filament 1.3 - 1.9 mm long; staminodes absent; carpels free, 8 to 14, ovary 2 - 3 mm long, stigma elongate, white, glabrous. Monocarps stipitate, stipes 8 - 15 mm long, 2 - 3 mm in diameter; monocarps 1 to 7, 110 - 175 mm long, 7 - 8 mm in diameter, moniliform, cylindrical, apex apiculate, pubescent to sparsely pubescent, verrucose, constricted around seeds when more than 1, yellow to red when ripe; seeds 1 to 7 per monocarp, 17 - 25 mm long, 4 - 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D96111349BD066A74F0165DA902D76A6.taxon	distribution	Distribution. A widespread central African species, from Cameroon to Gabon and Equatorial Guinea Democratic Republic of the Congo and Central African Republic; in Cameroon known from Central, East, Littoral, South, and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D96111349BD066A74F0165DA902D76A6.taxon	materials_examined	Specimens examined. Central Region: Mont Mbam Minkon on trail 5 km from Nkol Nyada village On top of small hill, 3.97 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 417 (MPU, WAG, YA). East Region: 43 km NW of Bertoua Road from Mbang to Ebaka, 4.58 ° N, 13.68 ° E, 19 May 1961, Breteler F. J. 1374 (BR, K, P, U, WAG, YA); Village situe a 35 km au NNE de Moloundou, 2.29 ° N, 15.35 ° E, 21 April 1971, Letouzey R. 10717 (P, YA). Littoral Region: Path above village summit Likombe village Etinde, 4.21 ° N, 9.166 ° E, 28 February 1995, Etuge M. 1191 (K); Nsoung- 1 km from village towards east river Meduya, 4.96 ° N, 9.802 ° E, 21 March 2003, Etuge M. 4919 (K). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 690 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 695 (WAG, YA); 58 km E of Kribi Fenda, 2.8 ° N, 10.4 ° E, 01 January 1911, Mildbraed G. W. J. 5989 (B, HBG); Bipindi, 3.08 ° N, 10.41 ° E, 01 April 1904, Zenker G. A. 2985 (E, L, P, WAG). South-West Region: Nyasoso, 4.81 ° N, 9.683 ° E, 06 February 1995, Cable S. 1185 (K, YA); Upper Boando, 4.06 ° N, 9.15 ° E, 15 March 1995, Cable S. 1581 (K, YA); Kupe village, 4.78 ° N, 9.700 ° E, 23 May 1996, Cable S. 2575 (K, YA); slopes of Mount Cameroon on the Bokwango trail near Bokwango village 4 km south west of Bu 4.12 ° N, 9.186 ° E, 23 March 2016, Couvreur T. L. P. 989 (WAG, YA); Nyasoso, 4.83 ° N, 9.683 ° E, 27 February 1996, Etuge M. 1729 (K, WAG, YA); Nyasoso, 4.83 ° N, 9.683 ° E, 24 June 1996, Etuge M. 2377 (K, WAG, YA); Bime rock face, 5.07 ° N, 9.727 ° E, 12 November 2001, Etuge M. 4531 (K); Entre Ndikoko et Ile 35 km N Kumba, 4.56 ° N, 9.23 ° E, 26 March 1976, Letouzey R. 14591 (BENIN, GC, IFAN, MO, P, TOGO, YA); Cameroon mountain at Bu 4.15 ° N, 9.233 ° E, 01 January 1930, Maitland T. D. 566 (K); Near the path from village Bechati to village Fossimondi, 5.65 ° N, 9.933 ° E, 26 September 2006, Tchiengue B. 2816 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4EDD6FEE7CCEEA06EB56B5FA93B987D3.taxon	description	Fig. 58; Map 7 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4EDD6FEE7CCEEA06EB56B5FA93B987D3.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2050, 1899: lectotype, chosen by Hoekstra et al. (2021), p. 157: B [B 100153030]; isolectotypes: B [B 100153031]; BM [BM 001125038]; BR [BR 0000008820280]; E [E 00624353]; G [G 00014883, G 00014884]; HBG [HBG- 502537]; K [K 000198987]; L [L. 1754335]; M [M- 0205486]; MO n. v., P [P 00362595, P 00362598]; S [S 07 - 13495]; WU [WU 0025871].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4EDD6FEE7CCEEA06EB56B5FA93B987D3.taxon	description	Description. Shrub when young (?) to liana, 4 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches reddish brown to yellowish brown, with dense erect reddish brown hairs 0.2 mm long. Leaves: petiole 5 - 7 mm long, 1 - 2 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 8.8 - 20 cm long, 5.2 - 8.9 cm wide, elliptic to oblong, apex acute to obtuse, base rounded to narrowly cuneate, subcoriaceous, below sparsely to densely pubescent when young, glabrous when old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib depressed, above densely pubescent when young and old, below densely pubescent to sparsely pubescent when young and old; secondary veins 7 to 10 pairs, glabrous above; tertiary venation percurrent, very dense. Individuals bisexual; inflorescences ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 16 per inflorescence; pedicel 6 - 11 mm long, ca. 1 mm in diameter, densely pubescent with dense erect reddish brown hairs; in fruit 7 - 15 mm long, ca. 1 mm in diameter; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract 0.5 - 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, ovate, apex acute to obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals partly covered in bud; outer petals 3, 2.5 - 6.5 mm long, 2 - 4.2 mm wide, ovate, apex obtuse, base truncate, margins flat, densely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 2.4 - 6.1 mm long, 1.4 - 3.4 mm wide, rhombic, apex obtuse, base truncate, margins flat, densely pubescent outside, pubescent with a glabrous base inside; stamens 8 to 9, in 1 row, ca. 1 mm long, clavate; connective truncate, glabrous; staminodes absent; carpels free, 6 to 7, ovary 1 - 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes ca. 4 mm long, ca. 2 mm in diameter; monocarps 1 to 3, 16 - 24 mm long, 9 - 11 mm in diameter, moniliform, cylindrical, apex apiculate, pubescent to glabrous, smooth, constricted around seeds when more than 1, color unknown; seeds 1 to 3 per monocarp, ca. 11 mm long, ca. 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4EDD6FEE7CCEEA06EB56B5FA93B987D3.taxon	distribution	Distribution. A widespread west and central African species, from Ivory Coast to eastern Nigeria and Cameroon to Gabon, with a disjunct distribution in eastern Democratic Republic of the Congo; in Cameroon known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4EDD6FEE7CCEEA06EB56B5FA93B987D3.taxon	materials_examined	Specimens examined. South Region: Bipindi, 3.08 ° N, 10.41 ° E, 1899, Zenker G. A. 2050 (E, L, M, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1904, Zenker G. A. 3001 (B, E, G, K, L, M, MO, P). South-West Region: Permanent Sample Plot on Shrike trail above Nyasoso, 4.81 ° N, 9.683 ° E, 06 February 1995, Cable S. 1169 (K, YA); Saddle of Kupe Rock, 4.79 ° N, 9.686 ° E, 14 November 1995, Cheek M. 7794 (K, YA); Kupe village, 4.79 ° N, 9.701 ° E, 17 November 1995, Cheek M. 7880 (K, WAG, YA); Nyasoso, 4.81 ° N, 9.716 ° E, 27 February 1996, Etuge M. 1748 (K, WAG, YA); Kupe village, 4.76 ° N, 9.7 ° E, 03 December 1999, Gosline W. G. 254 (K); Mapanja, 4.12 ° N, 9.115 ° E, 01 January 1931, Maitland T. D. 1072 (K); Korup National Park, 5.01 ° N, 8.833 ° E, 12 April 1985, Thomas D. W. 4721 (MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
53DB8E747E47DBE51D2A1A1C42A9D086.taxon	description	Map 7 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
53DB8E747E47DBE51D2A1A1C42A9D086.taxon	materials_examined	Type. Nigeria. Cross River State; Oban, Talbot P. A. 403, 1911: holotype: BM [BM 000843988].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
53DB8E747E47DBE51D2A1A1C42A9D086.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches pubescent to glabrescent, young foliate branches densely pubescent with dense appressed to ascending light-brown hairs 0.2 - 0.5 mm long. Leaves: petiole 4 - 7 mm long, 2 - 3 mm in diameter, densely pubescent with light-brown hairs, slightly grooved, blade inserted on top of the petiole; blade 11 - 25.8 cm long, 3.7 - 8.6 cm wide, oblong to obovate, apex acuminate, acumen 0.5 - 2.5 cm long, base subcordate, subcoriaceous to membranous, below sparsely pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 10 to 13 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed to extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 5 - 21 mm long, ca. 1 mm in diameter, densely pubescent; in fruit 5 - 21 mm long, 2 mm in diameter; upper bract ca. 3 mm long, ca. 2 mm wide; sepals 3, valvate, free, ca. 5 mm long, ca. 7 mm wide, ovate, apex obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 30 - 35 mm long, 23 - 25 mm wide, ovate, apex, base truncate, margins flat, pubescent outside, sparsely pubescent to glabrous inside; inner petals 3, valvate, ca. 21 mm long, ca. 26 mm wide, rhombic, apex acute, base truncate, margins flat, pubescent outside, glabrous inside; stamens 100 to 150, in 5 to 6 rows, ca. 1 mm long, cuneate; connective truncate, glabrous; staminodes absent; carpels free, 45 to 50, ovary ca. 2 mm long, stigma globose, glabrous. Monocarps stipitate, stipes 3 - 4 mm long, ca. 2 mm in diameter; monocarps ca. 8, 15 - 26 mm long, 9 - 10 mm in diameter, ellipsoid, apex rounded to apiculate, pubescent, smooth, constricted around seeds when more than 1, color unknown; seeds 1 to 2 per monocarp, ca. 10 mm long, ca. 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
53DB8E747E47DBE51D2A1A1C42A9D086.taxon	distribution	Distribution. Known from Nigeria and Cameroon; in Cameroon known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
53DB8E747E47DBE51D2A1A1C42A9D086.taxon	materials_examined	Specimens examined. South-West Region: Mount 4.78 ° N, 9.683 ° E, 26 November 1999, Cheek M. 10154 (K, MO, WAG, YA); AyinKeh 3 km north of Ngomboku, 4.93 ° N, 9.731 ° E, 17 December 1999, Ghogue J. - P. 500 (K, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AAAD584643ED60B3F4FEBED17F5CF790.taxon	description	Fig. 59; Map 7 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AAAD584643ED60B3F4FEBED17F5CF790.taxon	materials_examined	Type. Sierra Leone. no region; no location, Don G. s. n., no date: holotype: BM [BM 000547066].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AAAD584643ED60B3F4FEBED17F5CF790.taxon	description	Description. Shrub to liana, 4 - 20 m tall, d. b. h. 10 - 20 cm. Indumentum of simple hairs; old leafless branches sparsely pubescent to glabrous, light grey in color, young foliate branches pubescent with very short appressed to ascending hairs 0.1 mm long. Leaves: petiole 2 - 4 mm long, 0.5 - 1 mm in diameter, pubescent, grooved, blade inserted on top of the petiole; blade 4 - 16.5 cm long, 1.4 - 5.7 cm wide, obovate to oblanceolate, apex acuminate to acute, acumen ca. 2 cm long, base cuneate to subcordate, subcoriaceous, below sparsely pubescent to glabrous when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib depressed, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 7 to 10 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed to extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 15 - 45 mm long, ca. 0.5 mm in diameter, pubescent; in fruit 25 - 50 mm long, 0.5 - 1 mm in diameter; basal bract 1 - 2 mm long, 0.5 - 1 mm wide; upper bract 2 - 3 mm long, ca. 1 mm wide, lanceolate in shape; sepals 3, valvate, basally fused, 3 - 6 mm long, 4 - 5 mm wide, ovate, apex obtuse, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 12 - 30 mm long, 7 - 12 mm wide, elliptic, apex obtuse, base truncate, light yellow to light green, margins flat, pubescent outside, pubescent inside; inner petals 3, valvate, 5 - 9 mm long, 3 - 5 mm wide, ovate to rhombic, apex acute, base truncate, margins flat, pubescent outside, sparsely pubescent to glabrous inside; stamens 80 to 125, in 3 to 5 rows, ca. 1 mm long, linear to oblong; connective globose, glabrous; staminodes absent; carpels free, 17 to 24, ovary 1 - 2 mm long, stigma elongate, pubescent. Monocarps stipitate, stipes 4 - 8 mm long, 2 mm in diameter; monocarps 10 to 20, red when ripe, 13 - 35 mm long, 4 - 5 mm in diameter, moniliform, ellipsoid to globose, apex apiculate, densely pubescent, pubescent, constricted around seeds when more than 1, red when ripe; seeds 1 to 5 per monocarp, ca. 6 mm long, ca. 4 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AAAD584643ED60B3F4FEBED17F5CF790.taxon	distribution	Distribution. A mainly west African species, from Sierra Leone to Nigeria, just reaching into Cameroon where known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AAAD584643ED60B3F4FEBED17F5CF790.taxon	materials_examined	Specimens examined. South-West Region: Korup National Park, 5.01 ° N, 8.8 ° E, 15 June 2000, van der Burgt X. M. 614 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D4AA3B1ADA81EC709714693F3A19F27.taxon	description	Fig. 60; Map 8 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D4AA3B1ADA81EC709714693F3A19F27.taxon	materials_examined	Type. Cameroon. South Region; pres de zingui, Letouzey R. G. 10288, 5 Apr 1970: holotype: P [P 01960096]; isotype: YA [YA 0002998].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D4AA3B1ADA81EC709714693F3A19F27.taxon	description	Description. Liana, up to 3 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent with dense appressed to ascending yellowish-white hairs 0.1 - 0.2 mm long. Leaves: petiole 3 - 5 mm long, ca. 1 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 9.5 - 11.5 cm long, 3.3 - 4.1 cm wide, oblong to elliptic, apex acute, acumen 0.5 - 1 cm long, base rounded to subcordate, papyraceous, below sparsely pubescent with yellowish-white hairs when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below pubescent when young, sparsely pubescent when old; secondary veins 11 to 13 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed to axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 15 - 24 mm long, ca. 0.5 mm in diameter, sparsely pubescent; in fruit unknown; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract ca. 1 mm long, ca. 0.5 mm wide; upper bract ca. 1 mm long, ca. 0.5 - 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, ovate, apex acute, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 4.5 - 8 mm long, 3.7 - 5.8 mm wide, ovate, apex rounded to obtuse, base truncate, white, margins flat, sparsely pubescent outside, pubescent, base glabrous inside; inner petals 3, valvate, ca. 5.1 mm long, 2.7 - 2.9 mm wide, elliptic, apex obtuse and shortly acuminate, base truncate, margins flat, densely pubescent outside, pubescent with a glabrous base inside; stamens 6, in 1 row, ca. 2 mm long, linear; connective truncate, pubescent, filament ca. 2 mm long; staminodes absent; carpels free, 6, ovary ca. 2 mm long, stigma globose, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D4AA3B1ADA81EC709714693F3A19F27.taxon	distribution	Distribution. endemic to Cameroon; known only from the type locality in the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA5E0E4D7283AFE446CD557CAACDAA1F.taxon	description	Figs 58, 59; Map 8 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA5E0E4D7283AFE446CD557CAACDAA1F.taxon	materials_examined	Type. Equatorial Guinea. Bioko Norte; Fernando Poo, Mann G. 559, 1860: holotype: K [K 000198950]; isotypes: P [P 00363313; P 00363314].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA5E0E4D7283AFE446CD557CAACDAA1F.taxon	description	Description. Shrub to liana, up to 3 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with erect reddish brown 1.2 - 1.7 mm long hairs. Leaves: petiole 5 - 6 mm long, 2 - 3 mm in diameter, densely pubescent with erect reddish brown hairs, slightly grooved, blade inserted on top of the petiole; blade 8.3 - 28.5 cm long, 4.9 - 7.5 cm wide, oblong to obovate, apex acuminate to acute, acumen 0.5 - 2 cm long, base cordate or subcordate, subcoriaceous to membranous, below pubescent when young and old, above densely pubescent with yellowish hairs when young, sparsely pubescent with yellowish hairs to glabrous when old, discolorous, whitish below; midrib sunken or flat, above sparsely pubescent with erect reddish brown hairs when young, sparsely pubescent with erect reddish brown hairs when old, below densely pubescent when young, pubescent when old; secondary veins 9 to 23 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 7 - 12 mm long, 1 - 3 mm in diameter, densely pubescent with yellow erect hairs; in fruit 13 - 31 mm long, ca. 2 mm in diameter, pubescent; basal bract ca. 5 mm long, ca. 3 mm wide; upper bract ca. 5 mm long, ca. 5 mm wide; sepals 3, valvate, free, 6 - 9 mm long, 5 - 7 mm wide, triangular to ovate, apex acute to obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 21 - 50 mm long, 9 - 17 mm wide, elliptic to ovate, apex acute, base truncate, red to cream, margins flat, densely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 16 - 25 mm long, 6 - 8 mm wide, linear to elliptic, apex acute, base truncate, margins flat, densely pubescent outside, pubescent inside; stamens 100 to 120, in 4 to 5 rows, 1 - 2 mm long, cylindrical to obconic; connective truncate, glabrous; staminodes absent; carpels free, 22 to 24, ovary ca. 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 7 - 11 mm long, 2 - 3 mm in diameter; monocarps 2 to 9, 23 - 52 mm long, 9 - 13 mm in diameter, moniliform, ellipsoid to cylindrical, apex rounded to apiculate, pubescent, densely pubescent with erect hairs, slightly constricted around seeds when more than 1, orange when ripe; seeds 1 to 3, 17 - 27 mm long, 9 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA5E0E4D7283AFE446CD557CAACDAA1F.taxon	distribution	Distribution. A mainly west African species from Guinea to Ivory Coast, and eastern Nigeria to Equatorial Guinea; in Cameroon known from the Central, Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA5E0E4D7283AFE446CD557CAACDAA1F.taxon	materials_examined	Specimens examined. Central Region: Chantier forestier au sud de Song Bong, 3.41 ° N, 10.5 ° E, 08 December 1967, Bamps P. R. J. 1381 (BR, P, YA). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.39 ° E, 28 February 2018, Couvreur T. L. P. 1175 (K, MPU, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECE01948EB8F6B528DFF64FA49A69AA9.taxon	description	Figs 61, 62; Map 8 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECE01948EB8F6B528DFF64FA49A69AA9.taxon	materials_examined	Type. Gabon. Ogooue-Lolo; region de Lastoursville, Le Testu G. M. P. C. 8595, 19 Dec 1930: lectotype, chosen by Le Thomas (1969 b), p. 221: P [P 00362624, P 00362625]; isolectotypes: BM [BM 000553844]; BR [BR 0000008823748, BR 0000008823847].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECE01948EB8F6B528DFF64FA49A69AA9.taxon	description	Description. Liana, 7 m tall, d. b. h. up to 4 cm. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent with very short appressed yellowish to reddish brown hairs 0.1 mm long to glabrous. Leaves: petiole 3 - 9 mm long, 2 - 3 mm in diameter, glabrous, grooved, blade inserted on top of the petiole; blade 9.8 - 26.8 cm long, 4.6 - 10.8 cm wide, obovate, apex acuminate to acute, acumen ca. 1.5 cm long, base rounded to subcordate, papyraceous, below sparsely pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 7 to 10 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 0.5 - 3 mm long, ca. 1 mm in diameter, densely pubescent with short appressed reddish brown hairs; in fruit 1 - 10 mm long, ca. 1 mm in diameter; basal bract 0.5 - 1 mm long, 0.5 - 1 mm wide; upper bract 1 - 1.5 mm long, ca. 0.5 mm wide; sepals 3, valvate, connate, ca. 1 mm long, ca. 1 mm wide, ovate, apex obtuse, base truncate, pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals partly covered in bud; outer petals 3, 4.5 mm long, 2.6 - 2.9 mm wide, elliptic to ovate, apex acute, base truncate, red to yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 3.3 - 3.7 mm long, 1.2 - 1.5 mm wide, elliptic, apex acute, base truncate, margins flat, pubescent outside, glabrous inside; stamens 9, in 1 row, ca. 1 mm long, oblong; thecae on the lateral inner side with transversal dehiscence; connective truncate, glabrous; staminodes 9, in one whorl alternating with the stamens, ca. 0.3 mm long, glabrous; carpels free, 16 to 26, ovary 0.5 - 1 mm long, stigma globose, glabrous. Monocarps stipitate, stipes 1 - 4 mm long, ca. 2 mm in diameter; monocarps 1 to 8, ca. 17 mm long, 7 - 8 mm in diameter, moniliform, ellipsoid, apex rounded to apiculate, pubescent, verrucose, yellow to red when ripe; seed 1 per monocarp, 10 - 12 mm long, 6 - 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECE01948EB8F6B528DFF64FA49A69AA9.taxon	distribution	Distribution. A central African species, from Cameroon to Angola; in Cameroon known from the East and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECE01948EB8F6B528DFF64FA49A69AA9.taxon	materials_examined	Specimens examined. East Region: bezirk Molundu, 2.5 ° N, 15.25 ° E, 19 January 1911, Mildbraed G. W. J. 4286 (HBG). South Region: Bipindi, 3.05 ° N, 10.25 ° E, 01 January 1904, Zenker G. A. 2977 (BM, E).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F267866D5336730E242B711ADEABD3AF.taxon	description	Figs 61, 62; Map 8 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F267866D5336730E242B711ADEABD3AF.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Bikoro, Laurent E. 92, 20 Feb 1896: holotype: BR [BR 0000008805010].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F267866D5336730E242B711ADEABD3AF.taxon	description	Description. Shrub to liana, 20 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, reddish brown, young foliate branches sparsely pubescent with appressed to ascending yellowish hairs 0.2 - 0.3 mm long. Leaves: petiole 3 - 7 mm long, 1 mm in diameter, pubescent, slightly grooved, blade inserted on top of the petiole; blade 4.7 - 18.3 cm long, 2.4 - 7.2 cm wide, obovate to oblanceolate, apex acuminate to acute, acumen up to 2 cm long, base rounded to subcordate, papyraceous, below sparsely pubescent when young and old, above sparsely pubescent when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below pubescent when young and old; secondary veins 7 to 14 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 5 - 11 mm long, ca. 1 mm in diameter, sparsely pubescent; in fruit 15 - 23 mm long, 1 - 2 mm in diameter; basal bract ca. 1 mm long, ca. 0.5 mm wide; upper bracts 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, shortly fused basally or free, 1 - 2 mm long, ca. 3 mm wide, ovate, apex rounded, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 6 - 8 mm long, 5.2 - 8.1 mm wide, ovate, apex obtuse then shortly acuminate, base truncate, green, margins flat, densely to sparsely pubescent outside, glabrous but pubescent towards the margins inside; inner petals 3, valvate, 4.5 - 5.3 mm long, 3.5 - 3.9 mm wide, ovate to rhombic, apex obtuse but then shortly acuminate, base truncate, green, margins flat, densely pubescent outside, pubescent with a glabrous base inside; stamens 23 to 24, in 3 rows, ca. 1 mm long, obovate; connective truncate, glabrous; staminodes absent; carpels free, 9 to 12, ovary ca. 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 10 - 23 mm long, 2 - 3 mm in diameter; monocarps 3 to 11, 25 - 85 mm long, 5 - 6 mm in diameter, moniliform, cylindrical, apex apiculate, glabrous, verrucose, constricted around seeds when more than 1, yellow to orange when ripe; seeds 1 to 6 per monocarp, 14 - 19 mm long, 5 - 6 mm in diameter, subcylindrical; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F267866D5336730E242B711ADEABD3AF.taxon	distribution	Distribution. A widespread West and Central African species, from Sierra Leone to Nigeria and Cameroon to the Democratic Republic of the Congo and Angola; in Cameroon known from the Central, North and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F267866D5336730E242B711ADEABD3AF.taxon	materials_examined	Specimens examined. Central Region: Eloumden 10 km SW of Yaounde, 3.49 ° N, 11.26 ° E, 06 February 1996, Nkongmeneck B. A. 1326 (MO). North Region: Ngoussou, 6.03 ° N, 11.23 ° E, 01 February 1939, Jacques-Felix H. 3226 (P). South-West Region: Nyasoso village on max's trail to Mt 4.82 ° N, 9.694 ° E, 05 April 2016, Couvreur T. L. P. 1056 (WAG, YA); on top of hill near Small Ekombe village 3 km after Kumba on road to Ekondo Titi town, 4.62 ° N, 9.376 ° E, 13 January 2016, Couvreur T. L. P. 981 (WAG, YA); Ndabekim Hill to west of village, 4.91 ° N, 9.716 ° E, 15 December 1999, Etuge M. 4676 (K, MO, P, WAG, YA); Mt Cameroun south slope W of Victoria Transect 5, 4.07 ° N, 9.015 ° E, 16 November 1985, Gentry A. H. 52947 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C90BB550880203A8656D8133F17D0C28.taxon	description	Figs 63, 64; Map 8 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C90BB550880203A8656D8133F17D0C28.taxon	materials_examined	Type. Cameroon. East Region; Nkoum, Letouzey R. G. 3066, 19 Feb 1960: holotype P [P 00362617]; isotype: YA [YA 0002635].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C90BB550880203A8656D8133F17D0C28.taxon	description	Description. Liana, 30 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent completely covered with dense erect reddish brown hairs 0.4 mm lon g. Leaves: petiole 6 - 15 mm long, 2 - 3 mm in diameter, densely pubescent with erect reddish brown hairs, weakly grooved adaxially, blade inserted on top of the petiole; blade 6 - 23 cm long, 6 - 12.2 cm wide, oblong to oblanceolate, apex rounded to emarginate, acumen ca. 1 cm long, base cordate (to rounded), papyraceous, below pubescent when young and old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 14 to 19 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals unisexual, dioecious, male and female inflorescences dimorphic, male inflorescences cauliflorous with 3 to 50 flowers, in a fascicle or glomerule, peduncle 5 - 25 mm long; female inflorescences cauliflorous, in a condensed panicle with many flowers, peduncle 25 - 40 mm long. Flowers with 9 perianth parts in 3 whorls, male and female flowers dimorphic, male flowers: pedicel 6 - 10 mm long, ca. 1 mm in diameter, densely pubescent; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 1 mm wide, broadly ovate, apex rounded, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals shortly connate basally, outer petals longer than inner; outer petals 3, ca. 3 mm long, ca. 5 mm wide, ovate, apex acute, base truncate, margins flat, densely pubescent outside, papillose and sparsely pubescent with a few very short hairs inside; inner petals 3, valvate, 1.5 - 2 mm long, ca. 1 mm wide, elliptic to ovate, apex acute, base truncate, margins flat, papillose, sparsely pubescent towards the apex outside, papillose, sparsely pubescent towards the apex outside inside; stamens 31 to 40, in ca. 4 rows, ca. 1 mm long, oblong; connective truncate, pubescent; staminodes absent. Female flowers: pedicel 8 - 15 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 10 - 30 mm long, 2 mm in diameter; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 1 - 1.5 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 1 - 2 mm long, ca. 1 - 2 mm wide, broadly ovate, apex acuminate, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 5 - 6 mm long, 4.8 - 6 mm wide, broadly ovate, apex obtuse, base truncate, margins flat, densely pubescent with short papillose hairs outside, pubescent with short papillose hairs inside; inner petals 3, valvate, ca. 2 mm long, 1.2 - 1.4 mm wide, spathulate, apex obtuse, base truncate, margins flat, pubescent with short papillose hairs outside, pubescent with short papillose hairs inside; carpels free, 120, ovary ca. 1 mm long, stigma ellipsoid to globose, glabrous. Monocarps stipitate, stipes 7 - 10 mm long, 2 - 3 mm in diameter; monocarps at least 3 (but potentially with many more, up to 100), 20 - 80 mm long, 9 - 10 mm in diameter, moniliform, globose, apex rounded, densely pubescent, smooth, constricted around seeds when more than 1, blackish green when ripe; seeds 1 to 6 per monocarp, ca. 9 mm long, c. 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C90BB550880203A8656D8133F17D0C28.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of the Congo; in Cameroon known from the East and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A05FB9E2D3A2B14760BBF00BD2C77E8.taxon	description	Figs 59, 64; Map 8 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A05FB9E2D3A2B14760BBF00BD2C77E8.taxon	materials_examined	Type. Cameroon. Central Region; Yaounde-Station, Zenker, G. A. 431, 11 Jan 1894: holotype: B [B 100153061].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A05FB9E2D3A2B14760BBF00BD2C77E8.taxon	description	Description. Shrub to liana, up to 7 m tall, d. b. h. up to 1 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with dense appressed to ascending reddish brown hairs 0.2 - 0.3 mm long. Leaves: petiole 3 - 6 mm long, 1 - 2 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 5.5 - 20 cm long, 2.8 - 6.7 cm wide, elliptic to oblanceolate, apex acuminate to acute, acumen 1.5 cm long, base subcordate, subcoriaceous, below pubescent when young and old, above sparsely pubescent to glabrous when young and old, grey when dried, discolorous, whitish below; midrib impressed, above sparsely pubescent when young, glabrous when old, below pubescent when young and old; secondary veins 7 to 13 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 13 - 33 mm long, 0.5 - 1 mm in diameter, sparsely pubescent; in fruit 28 - 38 mm long, 0.5 - 1 mm in diameter; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, reflexed at anthesis, 1 - 2 mm long, 1 - 2 mm wide, broadly elliptic to broadly ovate, apex acute, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 6 - 15 mm long, 5.4 - 6.7 mm wide, ovate, apex obtuse, base truncate, yellow-green and red at the base, margins flat, sparsely pubescent outside, pubescent with a glabrous base inside; inner petals 3, valvate, 5.8 - 7.5 mm long, 4.5 - 9 mm wide, cordate, apex obtuse, base truncate, margins flat, glabrous outside, glabrous inside; stamens 36 to 48, in 5 rows, ca. 1 mm long, linear to oblong; connective truncate, glabrous; staminodes absent; carpels free, 11 to 14, ovary 1 - 2 mm long, stigma globose, glabrous. Monocarps stipitate, stipes 3 - 8 mm long, 2 - 3 mm in diameter; monocarps 4 to 11, 12 - 33 mm long, 4 - 9 mm in diameter, moniliform, ellipsoid, apex rounded to apiculate, pubescent to glabrous, verrucose, constricted around seeds when more than 1, yellow with red stripes or dull red when ripe; seeds 1 to 3 per monocarp, 11 - 12 mm long, 5 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A05FB9E2D3A2B14760BBF00BD2C77E8.taxon	distribution	Distribution. A central African species from Cameroon to the Democratic Republic of the Congo, Central African Republic and Angola; in Cameroon known from the Central and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A05FB9E2D3A2B14760BBF00BD2C77E8.taxon	materials_examined	Specimens examined. Central Region: Yaounde, 3.86 ° N, 11.51 ° E, 1894, Zenker G. A. 431 (B). South-West Region: Njonji, 4.13 ° N, 9.033 ° E, 17 April 1997, Nning J. 360 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A9E27989F39396625FE17DD055FD2A3.taxon	description	Figs 65, 66; Map 8 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A9E27989F39396625FE17DD055FD2A3.taxon	materials_examined	Type. Gabon. Ogooue-Ivindo; north of Koumameyong along SHM lumber roads, McPherson G. D. 16123, 31 Jan 1993: holotype: WAG [WAG 0357246, WAG 0357247]; isotypes: LBV n. v., MO n. v., P [P 01967243].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A9E27989F39396625FE17DD055FD2A3.taxon	description	Description. Liana, 20 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with appressed reddish-brown hairs 0.5 mm long. Leaves: petiole 4 - 8 mm long, ca. 1 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 8.5 - 23.5 cm long, 3.3 - 6.6 cm wide, ovate to oblanceolate, apex acuminate to acute, acumen 0.5 - 2.7 cm long, base cuneate to broadly cuneate with small linear black glands, chartaceous, below densely pubescent with yellowish hairs when young, sparsely pubescent to glabrous when old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 10 to 16 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, axillary or terminal. Flowers with 9 perianth parts in 2 whorls, numerous per inflorescence, peduncle 5.5 - 27 cm long, panicle-like; pedicel 5 - 27 mm long, 0.5 - 1 mm in diameter, densely pubescent with short reddish brown hairs; in fruit 20 - 30 mm long, 2 - 3 mm in diameter, pubescent; bracts 2, one basal and one upper towards the middle or lower half of pedicel, basal bract 1 - 8 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 0.5 - 1 mm long, 1 - 1.5 mm wide, broadly ovate, apex obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, petals 6 in one whorl, ca. 3 mm long, ca 1.5 mm wide, ovate, apex rounded to obtuse, base truncate, margins flat, sparsely pubescent outside, sparsely pubescent inside; stamens 6, in 1 row, inserted on a black hexagonal disc, opposite to the petals, free at the base, ca. 1 mm long, obconic; connective truncate, kidney-shaped in dorsal view, glabrous; staminodes 6, alternating with the stamens, glabrous; carpels free, 14 to 24, ovary ca. 2 mm long, stigma shortly bilobed, acute, glabrous. Monocarps stipitate, stipes 3 - 4 mm long, 3 - 4 mm in diameter; monocarps 2 to 10, 15 - 30 mm long, 10 - 15 mm in diameter, not moniliform, ellipsoid, apex rounded, pubescent, smooth, green when ripe; seed 1 per monocarp, ca. 10 mm long, ca. 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A9E27989F39396625FE17DD055FD2A3.taxon	distribution	Distribution. A central African species from Gabon and Cameroon; in Cameroon known from Littoral and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8A9E27989F39396625FE17DD055FD2A3.taxon	materials_examined	Specimens examined. Littoral Region: Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 651 (WAG, YA). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.34 ° E, 14 February 2015, Couvreur T. L. P. 708 (WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E73457C7AE1C3AD23BAEA0B7927FBBA.taxon	description	Fig. 46; Map 8 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E73457C7AE1C3AD23BAEA0B7927FBBA.taxon	materials_examined	Type. Gabon. Woleu-Ntem; region de Bitam, Le Testu G. M. P. C. 9028, 12 Mar 1933: lectotype, chosen by Le Thomas (1969 b), p. 226: P [P 00362618, P 00362621, P 00362623]; isolectotypes: BM [BM 000553843]; BR [BR 0000008823779, BR 0000008823786, BR 0000008823830].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E73457C7AE1C3AD23BAEA0B7927FBBA.taxon	description	Description. Shrub scrambling when young (?) to liana to ca. 3 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with dense ascending reddish brown hairs 0.1 - 0.3 mm long. Leaves: petiole 3 - 9 mm long, ca. 1 mm in diameter, pubescent to glabrous, grooved, blade inserted on the side of the petiole; blade 6.2 - 22 cm long, 2.8 - 9 cm wide, elliptic, ovate to obovate, apex acuminate to acute, acumen ca. 1.5 cm long, base cuneate to subcordate, papyraceous, below sparsely pubescent when young and old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above pubescent when young, glabrous to pubescent when old, below pubescent when young and old; secondary veins 6 to 11 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 6 per inflorescence; pedicel 16 - 55 mm long, ca. 1 mm in diameter, sparsely pubescent; in fruit; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract ca. 1 mm long, ca. 0.5 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, 2 - 3 mm wide, ovate, apex acute to rounded, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 4.7 - 5.5 mm long, 4.6 - 5.8 mm wide, ovate, apex acute, base truncate, light yellow, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 2.5 - 4 mm long, 2.8 - 3 mm wide, ovate, apex acute, base truncate, margins flat, densely pubescent outside, pubescent inside; stamens 14 to 24, in 1 to 2 rows, 1 - 2 mm long, oblong; connective absent or much reduced, filament much wider, thecae convergent towards apex, glabrous; staminodes absent; carpels free, 10 to 21, ovary ca. 1 mm long, stigma shortly bilobed, acute, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E73457C7AE1C3AD23BAEA0B7927FBBA.taxon	distribution	Distribution. A central African species from Cameroon to Gabon and Central African Republic; in Cameroon known from Adamaoua region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E73457C7AE1C3AD23BAEA0B7927FBBA.taxon	materials_examined	Specimens examined. Adamaoua Region: 21 km NNE de Banyo, 6.77 ° N, 11.81 ° E, 08 June 1967, Letouzey R. 8545 (P, YA); Pres Gandwa (25 km NNO de Banyo), 6.93 ° N, 11.72 ° E, 14 June 1967, Letouzey R. 8648 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2566F48D9E6AC6464F90A7846A55B4EB.taxon	description	Fig. 67; Map 8 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2566F48D9E6AC6464F90A7846A55B4EB.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Mbandaka, Eala, Pynaert L. A. E. J. 852, 20 Dec 1908: lectotype, designated by Hoekstra et al. (2021), p. 191: BR [BR 0000008805348, BR 0000008805355].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2566F48D9E6AC6464F90A7846A55B4EB.taxon	description	Description. Liana, 20 m tall, d. b. h. up to 2 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent with erect reddish brown hairs 0.6 - 1.5 mm long. Leaves: petiole 5 - 7 mm long, 1 - 2 mm in diameter, pubescent with long erect reddish brown hairs, weakly grooved adaxially, blade inserted on top of the petiole; blade 9.5 - 23.2 cm long, 2.9 - 7 cm wide, ovate to oblanceolate, apex acuminate to acute, acumen 0.5 - 1 cm long, base rounded to subcordate, papyraceous to subcoriaceous, below pubescent with erect yellow hairs when young and old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib sunken or flat, above densely pubescent when young and old, below pubescent when young and old; secondary veins 11 to 17 pairs, glabrous above; tertiary venation percurrent. Individuals unisexual, dioecieous, male and female inflorescences dimorphic, male inflorescences axillary, composed of a solitary flower to a few-flowered fascicle with up to 10 flowers; peduncle 4 - 8 mm long with erect hairs, female inflorescences cauliflorous, a condensed panicle with many flowers; peduncle 35 - 50 mm long, densely pubescent with erect hairs; Flowers with 9 perianth parts in 3 whorls, male and female flowers dimorphic. Male flowers: flowering pedicel 2 - 3 mm long, ca. 1 mm in diameter, pubescent; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, c. 1 mm long, ca. 1 mm wide, triangular to ovate, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, ca. 3 mm long, 3 mm wide, circular to broadly ovate, apex rounded to obtuse, base truncate, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, ca. 1 mm long, ca. 0.5 mm wide, ovate, apex rounded, base truncate, margins flat, pubescent outside, pubescent inside; stamens 6 (9), in 1 row, ca. 1 mm long, oblong; connective truncate, glabrous; staminodes 12 to 16, in one whorl externally to the stamens, very short, very sparsely pubescent or glabrous. Female flowers: flowering pedicel 20 - 30 mm long, ca. 1.5 mm in diameter, densely pubescent with erect hairs; in fruit ca. 33 mm long, ca. 2 mm in diameter; basal bract ca. 2 - 3 mm long, 1.5 - 2 mm wide; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 1 - 2 mm long, 1.5 - 2 mm wide, ovate to broadly ovate, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 5 - 6.3 mm long, 5 - 6.3 mm wide, circular to broadly ovate, apex rounded to obtuse, base truncate, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 1.7 - 1.8 mm long, 0.7 - 0.8 mm wide, ovate, apex rounded, base truncate, margins flat, pubescent outside, pubescent inside; carpels free, 95 to 150, ovary ca. 2 mm long, stigma globose to ellipsoid, glabrous. Monocarps stipitate, stipes 9 - 12 mm long, ca. 2 mm in diameter; monocarps up to 20, 20 - 60 mm long, 7 - 8 mm in diameter, moniliform, ellipsoid to cylindrical, apex apiculate, pubescent, verrucose, constricted around seeds when more than 1, color unknown; seeds 1 to 6 per monocarp, 11 - 13 mm long, 7 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2566F48D9E6AC6464F90A7846A55B4EB.taxon	distribution	Distribution. A central African species, from Cameroon to the Democratic Republic of the Congo and the Central African Republic; in Cameroon known from the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79F791D8A866ACCCA22B2FC88D97FCB6.taxon	description	Fig. 68; Map 9 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79F791D8A866ACCCA22B2FC88D97FCB6.taxon	materials_examined	Type. Gabon. Woleu-Ntem; on road from Mitzic to Lalara (N 2), just after the bridge over the Lara, Couvreur T. L. P. 869, 15 Nov 2015: holotype: WAG [WAG. 1575982]; isotypes: LBV; YA.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79F791D8A866ACCCA22B2FC88D97FCB6.taxon	description	Description. Scrambling shrub to liana, up to 6 m tall, d. b. h. up to 2 cm. Indumentum of simple hairs to glabrous; old leafless branches glabrous, young foliate branches pubescent with dense appressed to ascending reddish brown hairs 0.2 - 0.4 mm long. Leaves: petiole 2 - 4 mm long, ca. 1 mm in diameter, pubescent with appressed to ascending reddish brown hairs, slightly grooved, blade inserted on top of the petiole; blade 9.1 - 15.2 cm long, 1.4 - 2.4 cm wide, linear to narrowly elliptic, apex acuminate, acumen 1 - 2 cm long, base cuneate, papyraceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib depressed, above sparsely pubescent when young, glabrous when old, below sparsely pubescent when young, glabrous when old; secondary veins 15 to 20 pairs, almost perpendicular with midrib, straight, but curving halfway, glabrous above; tertiary venation percurrent, hardly visible. Inflorescences, flowers and fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79F791D8A866ACCCA22B2FC88D97FCB6.taxon	distribution	Distribution. A central African species, from Cameroon to the Republic of Congo; in Cameroon known from the Central and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
79F791D8A866ACCCA22B2FC88D97FCB6.taxon	materials_examined	Specimens examined. Central Region: Mefou proposed national park Near Mefou town, 3.62 ° N, 11.58 ° E, 08 March 2004, Cheek M. 11504 (K, WAG, YA). Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On east trail, 4.36 ° N, 10.25 ° E, 15 February 2013, Couvreur T. L. P. 628 (MPU, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
17D022E13BCAFBFC25D28FE21ECC47CA.taxon	description	Fig. 69; Map 9 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
17D022E13BCAFBFC25D28FE21ECC47CA.taxon	materials_examined	Type. Cameroon. Littoral Region; Nlonako, Letouzey R. G. 14476, 17 Mar 1976: holotype: WAG [WAG 0053953]; isotypes: MO [2 sheets]; P [P 01982361].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
17D022E13BCAFBFC25D28FE21ECC47CA.taxon	description	Description. Liana, 6 - 10 m tall, d. b. h. up to 5 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent with short 0.1 - 0.2 mm long appressed to half-erect yellowish hairs. Leaves: petiole 6 - 10 mm long, ca. 1 mm in diameter, pubescent, weakly grooved adaxially, blade inserted on the side of the petiole; blade 7.2 - 14.1 cm long, 2.1 - 3.5 cm wide, oblong to narrowly oblong or elliptic to narrowly elliptic, apex acuminate to acute, acumen ca. 1.5 cm long, base cuneate, papyraceous, below sparsely pubescent when young, sparsely pubescent to glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 11 to 18 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual [although fertile stamens can be absent leading to a female flower, thus possibly individuals gynodioecious], inflorescences cauliflorous, a condensed panicle with many flowers; peduncle up to ca. 70 mm, densely pubescent with appressed to erect reddish brown hairs; Flowers with 9 perianth parts in 3 whorls, pedicel 7 - 55 mm long, ca. 1 mm in diameter, densely pubescent with reddish hairs; in fruit 17 - 55 mm long, 1 - 2 mm in diameter; basal bract 1 - 2 mm long, 1 - 2 mm wide; upper bract 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, free, 3 - 2 mm long, ca. 2 mm wide, ovate, apex acute, base truncate, golden brown, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 3.6 - 5 mm long, 3.6 - 5.7 mm wide, broadly ovate to circular, apex obtuse, base truncate, golden green outside, yellowish inside, margins flat, densely pubescent outside, pubescent inside; inner petals 3, valvate, 0.4 - 1.3 mm long, 0.3 - 1.1 mm wide, broadly elliptic to circular, apex obtuse, base truncate, margins flat, pubescent outside, pubescent inside; stamens 0 to 2, in 1 row near the inner petals, ca. 1 mm long, clavate to oblong; connective reduced hidden by the thecae, pubescent; staminodes 0 to 14; carpels free, 65 to 85, ovary ca. 1 mm long, stigma globose, glabrous. Monocarps stipitate, stipes 7 - 14 mm long, 2 mm in diameter; monocarps up to 18, 23 - 45 mm long, 7 - 9 mm in diameter, moniliform, ellipsoid, apex apiculate, pubescent, smooth to slightly verrucose, constricted around seeds when more than 1, green when ripe; seeds 1 to 4 per monocarp, 13 - 14 mm long, 6 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
17D022E13BCAFBFC25D28FE21ECC47CA.taxon	distribution	Distribution. endemic to Cameroon, known from the South-West and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
17D022E13BCAFBFC25D28FE21ECC47CA.taxon	materials_examined	Specimens examined. Littoral Region: Nlonako Mt, 4.90 ° N, 9.943 ° E, 17 March 1976, Letouzey R. 14476 (MO, P, WAG). South-West Region: Ridge on S side of LOH Mt, 5 ° N, 9.683 ° E, 23 January 1998, Cheek M. 9067 (K, WAG, YA); Kodmin to Nzee Mbeng trail at N'dib river crossing, 5 ° N, 9.716 ° E, 14 February 1998, Cheek M. 9202 (K, WAG, YA); Bakossi Mountains 1 - 8 km NNE of Menyum Village, 5.05 ° N, 9.612 ° E, 22 May 1987, Doumenge C. 554 (MO, P); Nzimbeng road, 5.93 ° N, 9.716 ° E, 04 February 1998, Etuge M. 4122 (K, WAG, YA); Kodmin road towards Mahusom, 5 ° N, 9.683 ° E, 12 November 1998, Etuge M. 4442 (K, WAG, YA); Bakossi Mountains west of Bangem, 5.08 ° N, 9.7 ° E, 01 January 1986, Thomas D. W. 5274 (MO, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
371847843FA54D3AE3EDAF5B005EF164.taxon	description	Figs 70, 71; Map 9 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
371847843FA54D3AE3EDAF5B005EF164.taxon	materials_examined	Type. Cameroon. East Region; 15 km E of Dimako, village halfway Bertoua-Doume, Leeuwenberg A. J. M. 5828, 11 Jun 1965: holotype: WAG [WAG 0110801, WAG 0110802]; isotypes: B [B 100190273]; BR [BR 0000014126253]; C; EA; K; LISC; MO; P [P 01967268]; PRE; YA.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
371847843FA54D3AE3EDAF5B005EF164.taxon	description	Description. Liana, 3 m tall, d. b. h. to 11 cm. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches pubescent with very short appressed reddish brown hairs 0.1 mm long. Leaves: petiole 2 - 8 mm long, 1 - 2 mm in diameter, pubescent, blade inserted on the side of the petiole; blade 4.2 - 16.2 cm long, 1.8 - 5.3 cm wide, oblong to elliptic, apex acuminate to acute, acumen 0.5 - 1 cm long, base rounded, subcoriaceous, below sparsely pubescent when young, glabrous when old, above sparsely pubescent to glabrous when young and old, discolorous, whitish below; midrib impressed, above sparsely pubescent when young, glabrous when old, below pubescent when young, sparsely pubescent when old; secondary veins 7 to 10 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, axillary. Flowers with 6 to 7 perianth parts in 2 to 3 whorls, 1 to 2 per inflorescence, peduncle 0 - 6 mm long, pubescent; pedicel 12 - 20 mm long, 0.5 mm in diameter, pubescent; in fruit 16 - 29 mm long, ca. 2 mm in diameter; basal bract not seen; upper bracts ca. 1 mm long, ca. 0.5 mm wide; sepals 3, valvate, free, ca. 1 mm long, ca. 1 mm wide, ovate, apex obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner (when present), inner petals entirely covered in bud; outer petals 3, 2 - 2.2 mm long, ca. 2.2 mm wide, broadly ovate to circular, apex rounded to obtuse, base truncate, margins flat, pubescent outside, pubescent inside; inner petals absent or more rarely 1 minute, ca. 1.5 mm long, ca. 0.5 - 0.6 mm wide, elliptic, apex acute, base truncate, margins flat, pubescent outside, pubescent and glabrous towards the base inside; stamens 9 to 12, in 1 row, ca. 1 mm long, linear to clavate; connective reduced, slightly acute, glabrous; staminodes absent; carpels free, 9, ovary ca. 1 mm long, stigma flat, glabrous. Monocarps stipitate, stipes 7 - 22 mm long, ca. 2 mm in diameter; monocarps up to 7, 110 - 130 mm long, 8 - 9 mm in diameter, moniliform, ellipsoid, apex apiculate, sparsely pubescent to glabrous, verrucose, constricted around seeds when more than 1, yellow when ripe; seeds 1 to 4 per monocarp, ca. 17 mm long, ca. 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
371847843FA54D3AE3EDAF5B005EF164.taxon	distribution	Distribution. A Central African species, from Cameroon to Gabon, in Cameroon known from the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
71D3BF9DF915C9095F98AA87C035C201.taxon	description	Map 9 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
71D3BF9DF915C9095F98AA87C035C201.taxon	materials_examined	Type. Nigeria. Benue State; Abinsi, Dalziel, J. M. 712, 13 Sep 1972: holotype: K [K 000198917, K 000913659]; isotypes: BM [BM 001125037]; MO [2 sheets]; P [P 00362644, P 00362645].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
71D3BF9DF915C9095F98AA87C035C201.taxon	description	Description. Scandent shrub to liana, 3 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches sparsely pubescent with appressed reddish brown hairs 0.1 - 0.2 mm long. Leaves: petiole 2 - 5 mm long, 1 mm in diameter, sparsely pubescent with appressed reddish brown hairs, slightly grooved, blade inserted on top of the petiole; blade 9.7 - 17.5 cm long, 2.5 - 5.6 cm wide, obovate to oblanceolate, apex acuminate to acute, acumen 1.5 cm long, base cuneate to rounded, chartaceous, below sparsely pubescent when young and old, above sparsely pubescent with appressed whitish hairs when young, sparsely pubescent to glabrous when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 7 to 12 pairs, glabrous above; tertiary venation percurrent. Individual bisexual, inflorescences ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 2 to 3 (4) per inflorescence; pedicel 6 - 9 (14) mm long, ca. 1 mm in diameter, densely pubescent with appressed yellow-brown hairs; in fruit 7 - 13 mm long, ca. 1 mm in diameter; upper bract soon falling ca. 2 mm long, ca. 0.7 mm wide; sepals 3, valvate, basally fused, 0.5 - 1 mm long, 1 - 1.5 mm wide, broadly ovate to broadly triangular, apex acute to rounded, base truncate, sparsely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals partly covered in bud; outer petals 3, 2.4 - 3.3 mm long, 2 mm wide, elliptic to ovate, apex obtuse, base truncate, margins flat, sparsely pubescent outside, glabrous inside; inner petals 3, valvate, 2 - 2.7 mm long, 1.3 mm wide, elliptic, apex obtuse, base truncate, margins flat, sparsely pubescent outside, glabrous and sparsely pubescent towards apex inside; stamens (8) 9, in 1 row, 1 mm long, linear to oblong; connective truncate, glabrous; staminodes 6, alternating with the stamens, glabrous; carpels free, 8 to 12, 2 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 2 - 4 mm long, ovary ca. 2 mm in diameter; monocarps 2 to 9, 11 - 15 mm long, 4 - 6 mm in diameter, not moniliform, ellipsoid, apex apiculate, pubescent, smooth, red when ripe; seed 1 per monocarp, ca. 10 mm long, 4 - 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
71D3BF9DF915C9095F98AA87C035C201.taxon	distribution	Distribution. Known from Ghana, Ivory Coast, Benin, Nigeria and just reaching into Cameroon; in Cameroon known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C8248B482E133BF6F085C5506EB63879.taxon	description	Figs 71, 72; Map 9 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C8248B482E133BF6F085C5506EB63879.taxon	materials_examined	Type. Cameroon. South-West Region; NW du Mt Cameroon, Letouzey R. G. 15050, 1 Jun 1976: holotype: P [P 01982551]; isotypes: K [K 001322493]; WAG [WAG. 1576469]; YA [YA 0003005].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C8248B482E133BF6F085C5506EB63879.taxon	description	Description. Liana, to 20 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches sparsely pubescent with appressed yellowish hairs 0.2 mm long to almost glabrous. Leaves: petiole 3 - 6 mm long, ca. 1 mm in diameter, sparsely pubescent with appressed yellowish hairs, slightly grooved, blade inserted on top of the petiole; blade 5.6 - 11.8 cm long, 2.3 - 3.9 cm wide, oblong elliptic, apex acuminate to acute, acumen up to 2 cm long, base rounded to narrowly cuneate or cordate, papyraceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above sparsely pubescent when young and old, below sparsely pubescent when young and old; secondary veins 8 to 12 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 19 - 25 mm long, 0.5 - 1 mm in diameter, sparsely pubescent; in fruit 20 - 37 mm long, ca. 1 mm in diameter; basal bract ca. 1 mm long, ca. 1 mm wide; upper bract, leaf-like, 7 - 15 mm long, 5 - 10 mm wide; sepals 3, valvate, basally fused, ca. 2 mm long, ca. 3 mm wide, ovate, apex acute, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 6.4 - 8 mm long, ca. 6 mm wide, ovate, apex obtuse, base truncate, pale orange, margins flat, sparsely pubescent outside, sparsely pubescent inside; inner petals 3, valvate, 5.3 - 5.7 mm long, ca. 4 mm wide, broadly elliptic, apex acute, base truncate, pale orange, margins flat, pubescent outside, sparsely pubescent inside; stamens 15, in 2 rows, ca. 2 mm long, clavate; connective truncate, larger than base, glabrous; staminodes absent; carpels free, ca. 14, ovary ca. 2 mm long, stigma elongate, flattened at top, glabrous. Monocarps stipitate, stipes 6 - 10 mm long, ca. 2 mm in diameter; monocarps 2 to 13, 12 - 50 mm long, 5 - 7 mm in diameter, moniliform, ellipsoid to cylindrical, apex apiculate, very sparsely pubescent, verrucose, constricted around seeds when more than 1, orange when ripe; seeds 1 to 4 per monocarp, 8 - 15 mm long, 5 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C8248B482E133BF6F085C5506EB63879.taxon	distribution	Distribution. From Nigeria to Cameroon; in Cameroon known from the North-West, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C8248B482E133BF6F085C5506EB63879.taxon	materials_examined	Specimens examined. North-West Region: Mantum, 5.82 ° N, 10.08 ° E, 17 November 2000, Etuge M. 4810 (K, YA). South-West Region: Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.086 ° E, 03 April 2016, Couvreur T. L. P. 1049 (WAG, YA); Pentes NW du Mont vers Efolofo 30 km W, 4.21 ° N, 9.05 ° E, 01 June 1976, Letouzey R. 15050 (K, P, YA); Bamenda Distr at Bambui, 6.02 ° N, 10.21 ° E, 01 June 1931, Maitland T. D. 1618 (K). West Region: Megom, 5.43 ° N, 10.42 ° E, 01 February 1939, Jacques-Felix H. 3078 (P); Massif du Nkogam (2263 m) 25 km W de Foumban, 5.73 ° N, 10.67 ° E, 28 October 1974, Letouzey R. 13046 (P, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BFC1B64E37EAE7B83ADE13AFF23D2EE6.taxon	description	Map 9 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BFC1B64E37EAE7B83ADE13AFF23D2EE6.taxon	materials_examined	Type. Liberia. Sinon; Sinoe Basin, Whyte A. s. n., 1904: holotype: K [K 000198908].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BFC1B64E37EAE7B83ADE13AFF23D2EE6.taxon	description	Description. Shrub to liana, 2 - 50 m tall, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrescent, young foliate branches densely pubescent with appressed to ascending reddish brown hairs 0.2 mm long. Leaves: petiole 1 - 7 mm long, 1 - 2 mm in diameter, densely pubescent with appressed to ascending reddish brown hairs, weakly grooved adaxially, blade inserted on top of the petiole; blade 8.8 - 19.4 cm long, 3.5 - 8.5 cm wide, obovate, apex acuminate to acute, acumen 0.5 - 1 cm long, base subcordate, papyraceous to subcoriaceous, below sparsely pubescent when young, glabrous when old, above sparsely pubescent when young, glabrous when old, discolorous, whitish below; midrib sunken or flat, above sparsely pubescent when young, glabrous when old, below sparsely pubescent when young, glabrous when old; secondary veins 8 to 14 pairs, glabrous above; tertiary venation percurrent, sometimes not visible. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence, peduncle 2 - 8 mm, densely pubescent with yellowish slightly erect hairs; pedicel 7 - 11 mm long, ca. 1 mm in diameter, densely pubescent with yellowish slightly erect hairs; in fruit 13 - 20 mm long, ca. 2 mm in diameter; basal bract 1 - 2 mm long, ca. 1 mm wide; upper bract 1 - 2 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 1 - 2 mm long, 1 - 2 mm wide, broadly ovate, apex obtuse, base truncate, pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 3.4 - 5 mm long, 3.4 - 4.5 mm wide, broadly ovate, apex obtuse, base truncate, yellow to light green, margins flat, pubescent outside, glabrous, pubescent towards margins inside; inner petals 3, valvate, 3 - 4 mm long, 1.5 - 2.2 mm wide, elliptic to ovate, apex obtuse, base truncate, margins flat, pubescent outside, glabrous inside; stamens 9, in 1 row, ca. 1 mm long, obconical; connective truncate, glabrous; staminodes 9, in one external whorl and alternating with the stamens, glabrous; carpels free, 27 to 36, ovary ca. 1 mm long, stigma elongate, glabrous. Monocarps stipitate, stipes 5 - 12 mm long, ca. 2 mm in diameter; monocarps up to 10, 15 - 38 mm long, ca. 8 mm in diameter, moniliform, ellipsoid to globose, apex rounded to apiculate, pubescent to glabrous, verrucose, constricted around seeds, yellow to orange when ripe; seeds 1 to 3 per monocarp, 9 - 10 mm long, 6 - 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BFC1B64E37EAE7B83ADE13AFF23D2EE6.taxon	distribution	Distribution. A mainly West African species, from Sierra Leone to Benin, and Nigeria to Cameroon; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
41469A185974F1C82035586ECBABDFE8.taxon	description	Fig. 73; Map 9 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
41469A185974F1C82035586ECBABDFE8.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 3495 a, Oct 1907: holotype: G [G 00308331]; isotypes: BR [BR 0000013211349]; E [E 00624356]; HBG n. v., K n. v., L [L 1759466]; MO [3726267].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
41469A185974F1C82035586ECBABDFE8.taxon	description	Description. Liana, height unknown, d. b. h. unknown. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent with reddish brown, erect hairs 0.3 - 0.4 mm long. Leaves: petiole 3 - 6 mm long, 1 - 2 mm in diameter, densely pubescent with reddish brown erect hairs, slightly grooved, blade inserted on top of the petiole; blade 4.7 - 20.1 cm long, 2.3 - 9.5 cm wide, elliptic to obovate, apex acute to obtuse, base rounded, papyraceous to subcoriaceous, below sparsely pubescent with short erect yellow-brown hairs when young and old, above densely pubescent with erect yellow-brown hairs when young and old, discolorous, whitish below; midrib impressed, above sparsely pubescent with short erect yellow-brown hairs when young and old, below densely pubescent when young and old; secondary veins 8 to 12 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls 1 to 3 per inflorescence; pedicel 4 - 6 mm long, 0.5 - 1 mm in diameter, densely pubescent; in fruit unknown; basal bract not seen; upper bract ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, ca. 1 mm long, ca. 2 mm wide, ovate to broadly triangular, apex obtuse, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner, inner petals entirely covered in bud; outer petals 3, 2 - 3.1 mm long, 2.1 - 2.5 mm wide, ovate, apex obtuse, base truncate, margins flat, pubescent outside, pubescent towards margins to pubescent and glabrous towards center inside; inner petals 3, valvate, 1.8 - 2.4 mm long, 1.3 - 1.6 mm wide, rhombic, apex obtuse, base truncate, margins flat, densely pubescent outside, pubescent inside; stamens 35, in 3 to 4 rows, ca. 1 mm long, linear; anthers apically pubescent and converging apically and hiding the connective which is reduced or absent, glabrous; staminodes absent; carpels free, ca. 16, ovary ca. 1 mm long, stigma elongate, flattened at top, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
41469A185974F1C82035586ECBABDFE8.taxon	distribution	Distribution. endemic to Cameroon, known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5BF60227D90D2C56B0ECE14A5A993803.taxon	description	Description. Trees, rarely shrubs or rarely lianas, 4 - 40 m tall, d. b. h. 6 - 100 cm; stilt roots or buttresses absent. Indumentum of simple hairs, or absent. Leaves: petiole 2 - 14 mm long, 1 - 2 mm in diameter, blade inserted on the side of the petiole; blade 4 - 50 cm long, 2 - 15 cm wide, elliptic or obovate or oblong or ovate, apex acuminate, acumen ca. 1 cm long; base cuneate to rounded, concolorous; midrib raised above; secondary veins 8 to 23 pairs; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary, 1 per inflorescence; pedicel 8 - 270 mm long, 1 - 2 mm in diameter; in fruit 25 - 350 mm long, 3 - 15 mm in diameter; bract 1, towards the upper half or middle of pedicel; sepals 3, valvate, free, 2 - 40 mm long, 2 - 17 mm wide, ovate or elliptic, apex acute or attenuate or rounded, base truncate, margins flat, undulate, wavy or crisped; petals basally fused, tube 2 - 8 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 17 - 105 mm long, 6 - 40 mm wide, oblong or obovate or ovate, apex truncate, base truncate or with two small lobes, margins flat or wavy or crisped; inner petals 3, valvate, 9 - 35 mm long, 5 - 30 mm wide, cordate or rhombic or triangular to cochleate, apex acuminate to acute, base narrowed into a claw, claw 1 - 9 mm long; receptacle flat to strongly convex; stamens numerous, in 9 to 20 rows, 1 - 2 mm long, broad; connective discoid or elongated; staminodes absent; carpels fused into a syncarpous ovary, 1 - 5 mm long, stigma bilobed, slightly capitate. Fruit syncarpous, sessile, 35 - 150 mm long, 20 - 150 mm in diameter, globose or ovoid or conical, apex apiculate to rounded; seeds numerous, 9 - 22 mm long, 5 - 13 mm in diameter, ellipsoid or flattened ellipsoid; aril absent. A genus of trees with 14 known species, from West, Central and East Africa. In Cameroon six species are known, one endemic. Monodora, together with its sister genus Isolona, are unique in Annonaceae in having truly syncarpous ovaries, producing single fruits with unordered seeds, in contrast to other genera which have either uni- or biseriate placentation. Petals in Monodora are basally fused forming a short (not clearly visible) tube. The petals, however, are differentiated into inner and outer whorls, in contrast to Isolona (which has six equal lobes in a single whorl with a visible tube). In the vegetative state, Monodora and Isolona (together with Polyceratocarpous pellegrinii) are characterized by a raised leaf midrib on the upper side, in contrast to a sunken or flat midrib in all other genera found in Cameroon. Monodora species tend to have a whitish-grey wax indumentum on young leaves and fruits, which is especially noticeable in the common and widespread species M. myristica.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
420C5BA5301C226D33B7EE034CFAF3C7.taxon	description	Fig. 74; Map 9 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
420C5BA5301C226D33B7EE034CFAF3C7.taxon	materials_examined	Type. Angola. Malanje; Pungo Andongo, Welwitsch F. M. J. 774, May 1855: holotype: LISC; isotypes: B [B 10 0190365]; BM [BM 000553856]; COI [COI 00077209]; G [G 00011630]; K [K 000198843, K 000198844].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
420C5BA5301C226D33B7EE034CFAF3C7.taxon	description	Description. Tree, 18 - 20 m tall, d. b. h. 10 - 25 cm; stilt roots or buttresses absent. Indumentum absent; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 2 - 10 mm long, 1 mm in diameter, glabrous, weakly grooved adaxially, blade inserted on the side of the petiole; blade 4 - 20 cm long, 2 - 7.5 cm wide, elliptic, apex acuminate, acumen 0.5 - 1 cm long, base cuneate to obtuse, coriaceous to papyraceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 16 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 8 - 40 mm long, ca. 1 mm in diameter, glabrous; in fruit 25 - 85 mm long, 5 - 7 mm in diameter, glabrous; bract 1, towards the upper half of pedicel, 4 - 17 mm long, 3 - 12 mm wide; sepals 3, valvate, free, 5 - 15 mm long, 2 - 6 mm wide, ovate, apex acute, base truncate, green speckled with red and purple, glabrous outside, glabrous inside, margins undulate; petals basally fused, tube 2 - 3 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 17 - 50 mm long, 10 - 30 mm wide, oblong-obovate, apex truncate, base truncate, red-brown with pale yellow spots towards the apex, margins wavy, glabrous outside, glabrous inside; inner petals 3, valvate, 4 - 11 mm long, 5 - 16 mm wide, cordate to rhombic, apex acuminate, base narrowed into a claw, claw 3.0 - 9 mm long, white tinged with yellow, minutely purple-mottled along margins, margins flat, glabrous outside, glabrous inside; receptacle convex to flat; stamens numerous, in 9 to 11 rows, 1 mm long, broad; connective discoid, glabrous, cream; staminodes absent; carpels fused into a single structure, 1 - 2 mm long, stigma bilobed, slightly capitate, glabrous. Fruit syncarpous, 35 - 80 mm long, 35 - 50 mm in diameter, globose to ovoid, apex apiculate, glabrous, rugose, bumpy, green to brown when ripe; seeds 9 - 13 mm long, 5 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
420C5BA5301C226D33B7EE034CFAF3C7.taxon	distribution	Distribution. A widespread species from eastern Nigeria to northern Angola, Central African Republic, Democratic Republic of Congo, Uganda, western Tanzania and northern Zambia, with one collection from Ivory Coast; in Cameroon known from East, South, Central and Adamaoua regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
420C5BA5301C226D33B7EE034CFAF3C7.taxon	materials_examined	Specimens examined. Central Region: Environs de Ndokalende 10 km SW de Ndikinimeki, 4.72 ° N, 10.78 ° E, 04 March 1984, Satabie B. 754 (P, WAG, YA). East Region: Molunda Sation Ngoko Nginda 21 km north of Molundu, 2.2 ° N, 15.2 ° E, 09 January 1911, Mildbraed G. W. J. 4199 (B, HBG). North Region: Tibati, 6.46 ° N, 12.63 ° E, 01 February 1909, Ledermann C. L. 2462 (B); km 10 Tibati-Mabouka road, 6.43 ° N, 12.7 ° E, 29 June 1972, Leeuwenberg A. J. M. 10034 (BR, K, MO, P, WAG, YA). South Region: 18 km east from Lele village, 2.27 ° N, 13.32 ° E, 07 September 2013, Couvreur T. L. P. 461 (WAG, YA); 26 km east from Lele village, 2.28 ° N, 13.31 ° E, 08 September 2013, Couvreur T. L. P. 477 (WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
176DD89366C919FC2DE86A16FD88C7AF.taxon	description	Fig. 74; Map 9 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
176DD89366C919FC2DE86A16FD88C7AF.taxon	materials_examined	Type. Cameroon. South Region; 9 km N. of Kribi, Bos J. J. 6224, 2 Feb 1970: neotype, designated by Couvreur 2008, p. 206, sheet here designated: WAG [WAG 0158468]; isotypes: BR [BR 0000014056789]; C, K, LD, L MO, P, WAG [WAG 0158469]; YA.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
176DD89366C919FC2DE86A16FD88C7AF.taxon	description	Description. Tree or a more often a woody climber, lianescent, up to 20 m tall, d. b. h. 15 - 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs if present; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 3 - 7 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 5 - 17 cm long, 2.5 - 6 cm wide, elliptic to obovate, apex acuminate, acumen 0.3 - 1 cm long, base rounded, coriaceous to membranous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 13 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 20 - 50 mm long, ca. 1 mm in diameter, glabrous; in fruit 30 - 50 mm long, 4 - 10 mm in diameter, glabrous; bracts 1, upper only, towards the upper half or middle of pedicel, 6 - 15 mm long, 5 - 9 mm wide; sepals 3, valvate, free, 5 - 18 mm long, 3 - 6 mm wide, ovate, apex acute, base truncate, green, glabrous outside, glabrous inside, margins wavy to crisped; petals basally fused, tube 2 - 3 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 35 - 70 mm long, 6 - 20 mm wide, oblong, apex attenuate, base truncate, white or yellow at base and distally yellow with red-brown mottling, margins crisped, glabrous outside, glabrous inside; inner petals 3, valvate, 16 - 25 mm long, 6 - 20 mm wide, cordate to triangular, apex acute, base narrowed into a claw 3 - 8 mm long, white to yellow with red streaks towards margins, margins crisped, glabrous outside, sparsely pubescent inside; receptacle convex to flat; stamens numerous, in 9 to 11 rows, 1 mm long; connective discoid, glabrous, cream; staminodes absent; carpels fused into a single structure, ca. 2 mm long, stigma bilobed, slightly capitate, sparsely pubescent. Fruit syncarpous, 35 - 50 mm long, ca. 20 mm in diameter, long ellipsoid, apex acute, glabrous, 6 - 7 ribbed, otherwise smooth, green when ripe; seeds 10 - 13 mm long, 5 - 9 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
176DD89366C919FC2DE86A16FD88C7AF.taxon	distribution	Distribution. A widespread species present in West and Central Africa, from Sierra Leone to Ghana, and from southeastern Nigeria to Gabon; in Cameroon known from South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
176DD89366C919FC2DE86A16FD88C7AF.taxon	materials_examined	Specimens examined. South Region: 9 km N of Kribi, 4.83 ° N, 9.99 ° E, 02 February 1970, Bos J. J. 6224 (BR, C, K, LD, MO, P, WAG, YA); Just E of Kribi Seca Regrowth veg On S bank of Kienke R, 3.1 ° N, 10.25 ° E, 20 August 1970, Bos J. J. 7199 (BR, P, WAG); bezirk Ebolowa Ekuk 22 km Ebolowa, 2.91 ° N, 11.31 ° E, 01 January 1911, Mildbraed G. W. J. 5650 (HBG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1909, Zenker G. A. 3884 (B); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1909, Zenker G. A. 3935 (BM, BR, G, K, L, MO, P, S). South-West Region: Versant exterieur SSE du cratere du lac Dissoni (= SODEN lac) 20 km WNW Kumba, 4.72 ° N, 9.270 ° E, 20 March 1976, Letouzey R. 14498 (K, MO, WAG, YA); Mt Bakingili-Nja Keta Path, 4.08 ° N, 9.05 ° E, 21 September 1992, Wheatley J. I. 501 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE950688C3804BB15C6298F59E128151.taxon	description	Figs 75, 76; Map 10 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE950688C3804BB15C6298F59E128151.taxon	materials_examined	Type. Jamaica. Cultivated, obtained from Banks: holotype: BM.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE950688C3804BB15C6298F59E128151.taxon	description	Description. Tree, 30 - 40 m tall, d. b. h. 40 - 100 cm; stilt roots or buttresses absent. Indumentum of simple hairs if present; old leafless branches glabrous, young foliate branches glabrous, covered with a whitish wax. Leaves: petiole 8 - 14 mm long, 1 - 2 mm in diameter, glabrous, slightly grooved, blade inserted on the side of the petiole; blade 11 - 50 cm long, 4 - 14 cm wide, obovate, apex acuminate, acumen 1 - 1.5 cm long, base cuneate to cordate, coriaceous to papyraceous, below glabrous when young but covered with a whitish wax, glabrous when old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 13 to 23 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 70 - 270 mm long, 2 mm in diameter, glabrous; in fruit 300 - 350 mm long, 10 - 15 mm in diameter, glabrous; bracts 1, upper only, towards the middle of pedicel, 15 - 40 mm long, 8 - 30 mm wide; sepals 3, valvate, free, 20 - 40 mm long, 7 - 17 mm wide, elliptic to ovate, apex attenuate, base truncate, pale yellow with purple to completely dark red, glabrous outside, glabrous inside, margins wavy; petals basally fused, tube 6 - 8 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 40 - 105 mm long, 20 - 40 mm wide, ovate, apex acute, base truncate, deep yellow, base streaked with dark red, pale yellow spotted dark purple when young, margins wavy, glabrous outside, glabrous inside; inner petals 3, valvate, 25 - 35 mm long, 25 - 30 mm wide, cordate, apex acute to obtuse, base narrowed into a claw 2.0 - 5 mm long, white with red central vein abaxially, white and speckled with red-yellow adaxially, margins flat, pubescent towards base on both sides; receptacle strongly convex; stamens ca. 460 - 470, in 16 to 20 rows, 2 mm long, broad; connective elongated, pubescent, yellow; staminodes absent; carpels fused into a single structure, 4 - 5 mm long, stigma bilobed, slightly capitate, pubescent. Fruit syncarpous, 90 - 150 mm long, 80 - 150 mm in diameter, globose, apex rounded, glabrous, faintly ribbed longitudinally, otherwise smooth, green when ripe; seeds 15 - 22 mm long, 10 - 13 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE950688C3804BB15C6298F59E128151.taxon	distribution	Distribution. A widespread species, from Sierra Leone to northern Angola, Democratic Republic of the Congo, Uganda, western Kenya and Tanzania, also on the Island of Sao Tome; in Cameroon known from the East, South, Central, Littoral, South-West and Adamaoua (one collection) regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE950688C3804BB15C6298F59E128151.taxon	materials_examined	Selected specimens examined. Central Region: Mont Mbam Minkon on trail 2 km from Nkol Nyada village, 3.97 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 412 (WAG, YA); Ottotomo Forest Reserve 1 km after reserve base near small loggers road, 3.65 ° N, 11.28 ° E, 25 June 2013, Couvreur T. L. P. 445 (WAG, YA); on trail to Oveng Lodge hotel near parking just behind the village of Oveng 30 km on road from Mbalmayo to Sangmeli 3.41 ° N, 11.70 ° E, 09 February 2014, Couvreur T. L. P. 608 (WAG, YA); Yaounde, 3.86 ° N, 11.51 ° E, 30 July 1976, Westphal E. 9149 (WAG); Market of Messa Yaounde, 3.87 ° N, 11.52 ° E, 14 February 1978, Westphal E. 9883 (YA, P). East Region: Colline Nkolandjom pres Ngoakele a 25 km a l'W de Ngoulemakong (route Mbalmayo-Ebolowa), 3.38 ° N, 11.81 ° E, 12 July 1972, Letouzey R. 11474 (BR, P, YA); Asia, 3.63 ° N, 13.11 ° E, 21 April 1961, Letouzey R. 3899 (P, WAG, YA); Bezirk Molundu Bange-Busch Lokomo Bumba u Bange, 2.83 ° N, 15.25 ° E, 21 February 1911, Mildbraed G. W. J. 4530 (HBG). Littoral Region: Barombi camp 5 km S of Kumba on Buea-Douala road, 4.58 ° N, 9.45 ° E, 20 May 1983, Thomas D. W. 2118 (K, MO, P, YA). North Region: Kona, 4.6 ° N, 14.53 ° E, 15 December 1960, Letouzey R. 2616 (P, YA). South Region: Longii small marsh near seashore, 3.06 ° N, 9.966 ° E, 22 March 1969, Bos J. J. 4194 (BR, K, LM, MO, P, WAG, YA); Kribi roadside in New Bell, 2.95 ° N, 9.916 ° E, 14 May 1969, Bos J. J. 4526 (BR, K, LD, LM, MO, P, POZG, WAG, YA); on road Lolodorf-Bipindi ca half way near Mbiguiligui village (Mbikiliki), 3.16 ° N, 10.53 ° E, 26 February 2018, Couvreur T. L. P. 1155 (MPU, P, WAG, YA); Ebom, 3.1 ° N, 10.73 ° E, 20 February 1996, Elad M. 444 (WAG). South-West Region: Mungo River F. R., 4.78 ° N, 9.566 ° E, 02 December 1999, Cheek M. 10228 (K, MO, P, WAG, YA); Abang road, 4.92 ° N, 9.733 ° E, 14 December 1999, Cheek M. 10357 (K, MO, P, WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.079 ° E, 03 April 2016, Couvreur T. L. P. 1050 (WAG, YA); 2 km north of Nyasoso towards Mpako, 4.84 ° N, 9.679 ° E, 04 April 2016, Couvreur T. L. P. 1055 (WAG, YA); on trail from Ekongo village located 5 km before the entrance to Limbe 7 km on secondary road On flank of Mt Etinde 100 m in Mont Cameroon National Park, 4.06 ° N, 9.166 ° E, 16 October 2013, Couvreur T. L. P. 514 (WAG, YA); Ngusi, 4.83 ° N, 9.683 ° E, 15 January 1996, Etuge M. 1576 (K, WAG, YA); Nyasoso-Bedume road God-dat trail (opposite Ngusi road), 4.83 ° N, 9.683 ° E, 02 July 1996, Etuge M. 2516 (K, MO, P, WAG, YA); Mbolekang 1 - 2 km N of Nyandong village crossing Ndebessong river, 4.96 ° N, 9.579 ° E, 24 March 2003, Ghogue J. - P. 1487 (K); Entre Okoroba et Mbinda 20 km NW Nguti, 5.45 ° N, 9.272 ° E, 14 June 1975, Letouzey R. 13825 (P, YA); Limbe (Victoria), 4.01 ° N, 9.2 ° E, Preuss P. R. 1303 (BR, K, P, S). West Region: Bayangam, 5.3 ° N, 10.45 ° E, 17 December 1978, Westphal E. 10173 (P, WAG); Bayangam, 5.3 ° N, 10.45 ° E, 28 March 1979, Westphal E. 10204 (P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
155E4192263D7137827FB5CCDBEF367A.taxon	description	Figs 76, 77; Map 10 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
155E4192263D7137827FB5CCDBEF367A.taxon	materials_examined	Type. Nigeria. Lagos State; Eppah, Barter C. 3298, no date: holotype: K [K 000199032].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
155E4192263D7137827FB5CCDBEF367A.taxon	description	Description. Tree, 10 - 20 m tall, d. b. h. up to 60 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 2 - 7 mm long, 1 - 2 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 6 - 21 cm long, 2 - 7.5 cm wide, ovate to elliptic, apex acuminate, acumen 0.5 - 1 cm long, base cuneate, coriaceous to papyraceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 15 pairs, glabrous below; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 25 - 75 mm long, 1 - 2 mm in diameter, glabrous; in fruit 30 - 70 mm long, 3 - 4 mm in diameter, glabrous; bracts 1, upper only, towards the upper half of pedicel, 55 - 65 mm long, 10 - 30 mm wide; sepals 3, valvate, free, 10 - 35 mm long, 4 - 16 mm wide, ovate, apex rounded to attenuate, base truncate, green with red-brown markings, pubescent towards margins outside, pubescent towards margins inside, margins wavy; petals basally fused, tube 2 - 3 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 30 - 90 mm long, 25 - 30 mm wide, ovate, apex acute to rounded, base truncate, yellow-greenish, streaked with red-brown, base shading into bright white, margins wavy, glabrous outside, glabrous inside; inner petals 3, valvate, 10 - 35 mm long, 6 - 10 mm wide, cochleate, with two small oblong and pubescent lateral appendages around mid-height, apex acute to rounded, base narrowed into a claw 1 mm long, green to white streaked with red brown, margins flat, pubescent towards margins on both sides; receptacle convex to flat; stamens 286 to 337, in 10 to 13 rows, ca. 1 mm long, broad; connective discoid, pubescent, white; staminodes absent; carpels fused into a single structure, 2 - 3 mm long, stigma bilobed, glabrous. Fruit syncarpous, 45 - 75 mm long, 20 - 30 mm in diameter, globose, apex rounded, glabrous, covered with a greyish wax when young, faintly ribbed, green when ripe; seeds 12 - 17 mm long, 10 - 13 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
155E4192263D7137827FB5CCDBEF367A.taxon	distribution	Distribution. A widespread species across West and central Africa, from Guinee-Bissau to the Democratic Republic of the Congo; in Cameroon known from Central, East, Littoral, South or South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
155E4192263D7137827FB5CCDBEF367A.taxon	materials_examined	Specimens examined. Central Region: Etwa 125 km NO Jaunde, 4.63 ° N, 12.3 ° E, 01 February 1914, Mildbraed G. W. J. 8301 (K). East Region: 60 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.07 ° N, 14.65 ° E, 07 March 2019, Couvreur T. L. P. 1218 (MPU, WAG, YA); 69 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.07 ° N, 14.64 ° E, 08 March 2019, Couvreur T. L. P. 1233 (MPU, WAG, YA); Somalomo, 3.33 ° N, 12.75 ° E, 17 March 2017, Kamdem N. 483 (YA). Littoral Region: Piste Sole-Koum 20 km NW de Yabassi, 4.52 ° N, 9.773 ° E, 14 March 1976, Letouzey R. 14426 (P, YA). South Region: ca 8 km W of Bipindi 59 km from Kribi near Madoungou, 3.06 ° N, 10.33 ° E, 16 February 1970, Bos J. J. 6365 (B, BR, L, L, M, MO, P, WAG, YA); N´Koemvone, 2.8 ° N, 11.13 ° E, 13 December 1974, de Wilde J. J. F. E 7836 (B, BR, K, MO, P, U, WAG, YA); Nkoemvone, 2.81 ° N, 11.13 ° E, 03 July 1975, de Wilde J. J. F. E 8399 (WAG); Pres Mekomo (8 km SW confluent Dja & Lobo), 3.21 ° N, 12.36 ° E, 19 March 1962, Letouzey R. 4578 (P, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 1899, Zenker G. A. 1938 (A, B, BM, BR, G, K, L, P, S, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1900, Zenker G. A. 2251 (A, B, BM, G, K, L, P, S, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 2727 (BM, BR, K, L, M, P, S, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3793 (G, K); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1912, Zenker G. A. 4317 (B, BM, BR, G, K, L, MO, P, S). South-West Region: between 200 - 600 m alt above Bakingili, 4.08 ° N, 9.05 ° E, 21 December 1993, Cheek M. 5840 (K, YA); West of Kola, 4.83 ° N, 9.733 ° E, 27 October 1998, Cheek M. 9399 (K, WAG, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.503 ° E, 26 March 2016, Couvreur T. L. P. 1019 (WAG, YA); A 11 km au Sud Est de Molobo (village situe a 50 km au Sud de Batouri), 4.02 ° N, 14.32 ° E, 21 July 1963, Letouzey R. 5459 (P, YA); Ambas Bay (Victoria NDLR), 4.01 ° N, 9.226 ° E, 1861, Mann G. 111 (K); Around Masaka-Batanga, 5.1 ° N, 9.17 ° E, 24 March 1988, Thomas D. W. 7750 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4E2472039C718CEC087F1F92E62FFFC1.taxon	description	Figs 78, 79; Map 10 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4E2472039C718CEC087F1F92E62FFFC1.taxon	materials_examined	Type. Nigeria. no region; no location, Palisot de Beauvois A. M. F. J. s. n., no date: holotype: G-DC, scanned image [excluding fruits] [G 00011671].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4E2472039C718CEC087F1F92E62FFFC1.taxon	description	Description. Tree, up to 20 m tall, d. b. h. up to 100 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 2 - 14 mm long, 1 - 2 mm in diameter, glabrous, slightly grooved, blade inserted on the side of the petiole; blade 10 - 40 cm long, 8 - 15 cm wide, oblong to obovate, apex acuminate, acumen 3 - 9 cm long, base rounded to obtuse, coriaceous to papyraceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 17 pairs, glabrous below; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers bisexual with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 30 - 55 mm long, ca. 1 mm in diameter, glabrous; in fruit 40 - 50 mm long, 8 - 10 mm in diameter, glabrous; bracts 1, upper only, towards the upper half or middle of pedicel, 6 - 10 mm long, 7 - 11 mm wide; sepals 3, valvate, free, 7 - 11 mm long, 5 - 10 mm wide, ovate, apex rounded to obtuse, base truncate, green, glabrous, pubescent towards margins outside, glabrous, pubescent towards margins inside, margins wavy; petals basally fused, tube 2 - 3 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 25 - 45 mm long, 15 - 30 mm wide, ovate, apex acute, base truncate, speckled and streaked yellow and purple, base creamy-white, margins wavy, glabrous, pubescent towards margins outside, glabrous, pubescent towards margins inside; inner petals 3, valvate, 17 - 27 mm long, 13 - 20 mm wide, rhombic, apex acute, base narrowed into a claw, claw 2 - 5 mm long, yellow with brown-purple spots, margins flat, glabrous outside, sparsely pubescent inside; receptacle strongly convex; stamens numerous, in 12 to 14 rows, ca. 1 mm long, broad; connective discoid, densely pubescent, cream; staminodes absent; carpels fused into a single structure, 4 mm long, stigma bilobed, slightly capitate, sparsely pubescent. Fruits syncarpous, 60 - 120 mm long, 40 - 60 mm in diameter, ovoid, apex rounded, tomentose, faintly ribbed longitudinally, otherwise smooth, brown when ripe; seeds 9 - 20 mm long, 6 - 11 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4E2472039C718CEC087F1F92E62FFFC1.taxon	distribution	Distribution. A widespread species, disjunct between West Africa (Sierra Leone to Togo) and Central Africa (Nigeria, Cameroon and Sao Tome Island); in Cameroon known from East, South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4E2472039C718CEC087F1F92E62FFFC1.taxon	materials_examined	Specimens examined. Central Region: Reserve forestiere de Makak, 3.48 ° N, 11.01 ° E, 14 December 1967, Bamps P. R. J. 1449 (BR, YA). East Region: ca 9 km from Bertoua near the road to Doume, 4.53 ° N, 13.61 ° E, 09 December 1961, Breteler F. J. 2182 (K, P, WAG, YA). Littoral Region: Ebo Proposed National Park Iboty to Bekob village, 4.46 ° N, 10.46 ° E, 14 February 2006, Cheek M. 12928 (K). South Region: along main road Kribi-Bipindi, 2.99 ° N, 10.01 ° E, 17 February 2012, Couvreur T. L. P. 391 (WAG, YA); Lolodorf, 3.23 ° N, 10.71 ° E, 1896, Staudt A. 40 (G, K, P). South-West Region: Path above Kupe village, 4.76 ° N, 9.694 ° E, 22 May 1996, Cable S. 2544 (K, WAG, YA); Nyangdong volcanic waterfall-whirlpool path, 4.96 ° N, 9.577 ° E, 27 March 2003, Cheek M. 11464 (K); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.07 ° N, 9.051 ° E, 02 April 2016, Couvreur T. L. P. 1042 (WAG, YA); Nyasoso village on max's trail to Mt 4.82 ° N, 9.692 ° E, 05 April 2016, Couvreur T. L. P. 1061 (WAG, YA); Kupe village to Loum State Forest, 4.73 ° N, 9.716 ° E, 29 May 1996, Etuge M. 2017 (K, MO, P, WAG, YA); Korup National Park, 5.06 ° N, 8.855 ° E, 06 February 1998, Kenfack D. 1027 (MO, WAG); 6 km W of Bota rocky coast with volcanic boulders, 4.01 ° N, 9.144 ° E, 31 August 1972, Leeuwenberg A. J. M. 10295 (BR, MO, P, WAG, YA); Limbe (Victoria) Bimbia road, 4.00 ° N, 13.21 ° E, 01 February 1929, Maitland T. D. 408 (K); Johann-Albrechtshohe [Kumba] area 4.63 ° N, 9.416 ° E, 1896, Staudt A. 495 (A); Southern slope of Mount above Batoke, 4.03 ° N, 9.1 ° E, 25 January 1984, Thomas D. W. 3025 (B, BR, MO, P, YA); Small Koto, 4.31 ° N, 9.066 ° E, 07 March 1985, Thomas D. W. 4516 (K, MO, P, YA); Cameroon Mountain, 4.31 ° N, 9.066 ° E, 01 June 1985, Thomas D. W. 4814 (BR, MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6291D9684DB836046FD8FD6E2B23A183.taxon	description	Fig. 79; Map 10 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6291D9684DB836046FD8FD6E2B23A183.taxon	materials_examined	Type. Cameroon. Central Region; Yaounde, Zenker G. A. 776, 1896: holotype: B [B 10 015406]; isotypes: COI [COI 00077202]; G [G 00011721]; K [K 000199027]; NY [NY 00026141]; P [P 00363263].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6291D9684DB836046FD8FD6E2B23A183.taxon	description	Description. Leaning tree [or liana?], to 6 m tall, d. b. h. up to 6 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 2 - 4 mm long, 1 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 10 - 15 cm long, 8 - 14 cm wide, elliptic to obovate, apex acuminate, acumen 3 - 6 cm long, base rounded to obtuse, coriaceous to papyraceous, below glabrous when young and old, above glabrous when young and old, concolorous; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 10 to 13 pairs, glabrous below; tertiary venation intermediate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers bisexual with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 28 - 50 mm long, 1 mm in diameter, sparsely pubescent; in fruit unknown; bracts 1, towards the upper half or middle of pedicel, 11 - 20 mm long, 10 - 13 mm wide; sepals 3, valvate, free, 2 - 5 mm long, 2 - 4 mm wide, ovate, apex acute, base truncate, green to brown-red, glabrous outside, glabrous inside, margins flat; petals basally fused, tube 4 - 7 mm long, inner and outer whorl differentiated, outer petals longer than inner; outer petals 3, 35 - 45 mm long, 20 - 28 mm wide, ovate, apex acute to obtuse, base truncate flanked by two small lobes, speckled and streaked yellow-purple, base creamy-white, margins flat, sparsely pubescent outside, sparsely pubescent to glabrous inside; inner petals 3, valvate, 9 - 13 mm long, 13 - 16 mm wide, triangular, apex acute, base narrowed into a claw, claw 4.0 - 7 mm long, pale green with red spots, margins flat, folded outwards, glabrous inside, pubescent outside; receptacle convex to flat; stamens numerous, in 9 to 10 rows, 1 mm long, broad; connective discoid, glabrous; staminodes absent; carpels fused into a single structure, 1 mm long, stigma bilobed, slightly capitate. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6291D9684DB836046FD8FD6E2B23A183.taxon	distribution	Distribution. endemic to Cameroon; known from Central, East, South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6291D9684DB836046FD8FD6E2B23A183.taxon	materials_examined	Specimens examined. Central Region: ca. 4 km NE of Otele near the road to Yaounde, 3.58 ° N, 11.25 ° E, 28 March 1964, de Wilde J. J. F. E 2249 (BR, P, WAG, YA); Etwa 195 km NO Jaunde, 5.21 ° N, 12.9 ° E, 01 January 1914, Mildbraed G. W. J. 8476 (K); Mt Ngoro a 38 km au N de Bafia (piedmont), 4.93 ° N, 11.38 ° E, 01 May 1978, Ngameni B. K. 113 (P, YA); Yaounde, 3.87 ° N, 11.52 ° E, 1896, Zenker G. A. 776 (B, COI, G, K, P). East Region: Bertoua, 4.58 ° N, 13.73 ° E, 27 April 1961, Breteler F. J. 1319 (WAG); 6 km along road south to Doume, 4.38 ° N, 12.91 ° E, 10 April 1962, Breteler F. J. 2747 (BR, K, P, WAG, YA); Deng Deng, 5.16 ° N, 13.58 ° E, 01 April 1914, Mildbraed G. W. J. 8850 (K); Bertoua, 4.58 ° N, 13.68 ° E, 27 May 1955, Nana P. 98 (P, YA). South Region: Nkoldjobe dans le massif du Mbam-Minkom 18 km Nord-Ouest de Yaounde, 3.93 ° N, 11.35 ° E, 16 April 1984, Achoundong G. 1056 (YA); Bitye river, 3.02 ° N, 12.37 ° E, 01 January 1920, Bates G. L. 1244 (COI); Near village Oveng 27 km from Sangmelima along road to Yaounde, 3.09 ° N, 11.90 ° E, 21 March 1962, Breteler F. J. 2683 (BR, K, P, U, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F1B18027DFAC82E92159B9C3A7BE5DF.taxon	description	Description. Trees to shrubs, 3 - 25 m tall, d. b. h. 3 - 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 2 - 14 mm long, 1 - 3 mm in diameter; blade 6 - 31 cm long, 2 - 10 cm wide, ovate or elliptic or oblong, apex acuminate or acute, acumen 0.5 - 2.4 cm long, base rounded or obtuse or cuneate, discolorous, whitish below or concolorous; midrib sunken or flat; secondary veins 10 to 24 pairs; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 8 per inflorescence; pedicel 5 - 46 mm long, ca. 1 mm in diameter; in fruit 20 - 60 mm long, 2 - 4 mm in diameter; bract 1, basal, 1 - 3 mm long, ca. 1 mm wide; sepals 3, valvate, free, 1 - 4 mm long, 1 - 3 mm wide, triangular or ovate or semi-circular, apex acute or acuminate, base truncate; petals free, valvate, outer petals longer than inner; outer petals 3, 6 - 33 mm long, 1 - 15 mm wide, elliptic or narrowly triangular, apex acute or attenuate, base narrowed and concave; inner petals 3, valvate, 1 - 10 mm long, 0.6 - 5 mm wide, triangular, apex acute, base broad and concave; stamens 30 to 170, in 4 to 6 rows, 2 - 3 mm long, linear or broad; connective tongue-shaped; staminodes absent; carpels free, 20 to 109, ovary 1 - 2.5 mm long, stigma cylindrical or filiform. Monocarps sessile or stipitate, stipes 6 - 23 mm long; monocarps 7 to 144, 10 - 30 mm long, 10 - 20 mm in diameter, ellipsoid, apex apiculate, rounded or pyramidal in shape; seed 1, 4 - 14 mm long, 2 - 11 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BE5D66A4C543D7FE2F4CD56D4CB5826.taxon	description	Figs 80, 81; Map 10 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BE5D66A4C543D7FE2F4CD56D4CB5826.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Thomson W. C. 134, no date: holotype: K [K 000199047].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BE5D66A4C543D7FE2F4CD56D4CB5826.taxon	description	Description. Tree to shrub or scandent shrub, 3 - 5 m tall, d. b. h. 6 - 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 4 - 6 mm long, 1 - 2 mm in diameter, pubescent, slightly grooved to cylindrical, blade inserted on top of the petiole; blade 8.5 - 25.5 cm long, 3.7 - 9 cm wide, oblong to elliptic, apex acuminate to acute, acumen 0.2 - 1.8 cm long, base rounded to obtuse, papyraceous, below pubescent with short appressed straight hairs when young and old, above glabrous to pubescent when young, glabrous to pubescent when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 10 to 18 pairs, glabrous below; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous, on foliate or leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, (1) 2 to 4 per inflorescence; peduncle 1 - 2 mm long, unbranched; pedicel 12 - 38 mm long, ca. 1 mm in diameter, pubescent; in fruit 20 - 40 mm long, 3 - 4 mm in diameter, sparsely pubescent; bract 1, basal, 1 - 3 mm long, ca. 1 mm wide; sepals 3, valvate, free, 1 - 4 mm long, 1 - 3 mm wide, triangular to ovate, apex acute to acuminate, base acute, brown-green, pubescent outside, pubescent inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 6.5 - 29 mm long in total, 2 - 5 (6) mm wide above the claw, narrowly triangular, apex attenuate, base suborbicular, basal part 4 - 6 mm wide, light green to light yellow, margins flat, pubescent to glabrous outside, pubescent inside; inner petals 3, valvate, 3.4 - 10.9 mm long, 1 - 3 mm wide, triangular, apex acute, base broad and concave, 3 - 5 mm wide, margins flat, pubescent to glabrous outside, pubescent inside; stamens 135 - 156, in 4 to 5 rows, ca. 2 mm long, linear; connective tongue shaped, glabrous, red; staminodes absent; carpels free, 67 to 109, ovary ca. 1 mm long, stigma cylindrical, glabrous. Monocarps stipitate, stipes 17 - 23 mm long, ca. 1 mm in diameter; monocarps 15 (22) to 48 (60), 9 - 20 mm long, 6 - 13 mm in diameter, ellipsoid, apex shortly apiculate, sparsely pubescent to glabrous, smooth, not ribbed, brown turning wine red when ripe; seed 1 per monocarp, 4 - 18 mm long, 2 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BE5D66A4C543D7FE2F4CD56D4CB5826.taxon	distribution	Distribution. From Nigeria to Democratic Republic of the Congo; in Cameroon known from East, South, Central, South-West and South West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BE5D66A4C543D7FE2F4CD56D4CB5826.taxon	materials_examined	Specimens examined. Central Region: Between Somalomo and Milon 69 km SE Akonolinga, 3.77 ° N, 12.25 ° E, 18 June 1981, Asonganyi J. N. 310 (P, YA); NE of Ndangan I on logging trail into Mefou NP, 3.62 ° N, 11.59 ° E, 23 March 2004, Cheek M. 11965 (BR, K, K, MO, P, WAG, YA); Mefou National Park just after Ape Action Africa center along road, 3.62 ° N, 11.58 ° E, 24 April 2013, Couvreur T. L. P. 422 (WAG, YA); Banlieu de Yaounde au bord de l'etang Atemengue, 3.87 ° N, 11.52 ° E, 14 April 1960, Endengle E. 116 (P, YA); Goura, 4.55 ° N, 11.4 ° E, 01 November 1938, Jacques-Felix H. 2340 (P); Yaounde, 3.87 ° N, 11.52 ° E, 01 August 1939, Jacques-Felix H. 4803 (P); Kourou (Yoko), 5.03 ° N, 12.42 ° E, 10 July 1959, Letouzey R. 2373 (P, YA); Pres de Mendong (25 km N Akonolinga), 3.77 ° N, 12.25 ° E, 07 March 1962, Letouzey R. 4486 (P, YA); Issandja (Ntui), 4.77 ° N, 11.77 ° E, 19 December 1969, Letouzey R. 9726 (P); Ngolep Mt, 4.95 ° N, 11.38 ° E, 27 April 1978, Ngameni B. K. 44 (P, YA); Pres riviere Malo non loin de Ndikinimeki, 4.77 ° N, 10.83 ° E, 08 November 1983, Nkongmeneck B. A. 518 (P, YA); Mefou National Park, 3.61 ° N, 11.56 ° E, 27 March 2004, Onana J. M. 2870 (BR, G, K, YA); Ngoro (28 km NE Bafia Cameroun), 4.95 ° N, 11.38 ° E, 29 March 1963, Raynal A. 10569 (P, WAG, YA); Nkolbison (8 km W Yaounde Cameroun), 3.88 ° N, 11.45 ° E, 10 February 1963, Raynal J. 9560 (P, YA). East Region: 7 km along road to Batouri and Betare Oya, 4.58 ° N, 13.68 ° E, 16 March 1961, Breteler F. J. 1220 (A, BR, K, M, P, SL, UC, WAG, YA); Bertoua, 4.58 ° N, 13.68 ° E, 13 August 1961, Breteler F. J. 1799 (BR, K, P, WAG, YA); Bertoua, 4.58 ° N, 13.68 ° E, 08 November 1960, Breteler F. J. 644 (BR, K, P, WAG, YA); Doume, 4.23 ° N, 13.45 ° E, 12 November 1960, Breteler F. J. 684 (BR, K, M, P, WAG, YA); Piste Mpoundou-Seglendon marecages, 4.1 ° N, 13.05 ° E, 16 February 1960, Letouzey R. 3021 (P, YA); Pres Goyoum, 5.22 ° N, 13.38 ° E, 01 February 1961, Letouzey R. 3350 (P, YA); 70 km route Bertoua-Ndemba I, 4.75 ° N, 13.43 ° E, 18 July 1955, Nana P. 171 (P, YA); Bertoua, 4.58 ° N, 13.68 ° E, 27 August 1955, Nana P. 240 (P); route Esseleke, 4.58 ° N, 13.68 ° E, 13 February 1956, Nana P. 474 (P); 8 km de Deng Deng, 5.23 ° N, 13.46 ° E, 10 February 2002, Sonke B. 2671 (BR, BRLU, K, WAG, YA). South Region: At Bissombo 59 km SE Akonolinga, 3.28 ° N, 12.47 ° E, 12 June 1981, Asonganyi J. N. 275 (YA); Biyi (Ambam), 3.02 ° N, 12.37 ° E, 12 February 1963, Raynal J. 9806 (P). South-West Region: Korup National Park, 5.33 ° N, 9.42 ° E, 18 September 1990, Harris D. J. 2516 (P, YA). West Region: Touladen en bas du versant oriental du Mbepit 15 km E Foumbot et 25 km SSW Foumban, 5.52 ° N, 10.78 ° E, 26 October 1974, Letouzey R. 13025 (P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BFCCA4C74F3120AA14FDB1E611A2898.taxon	description	Figs 82, 83; Map 10 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BFCCA4C74F3120AA14FDB1E611A2898.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 3105, 1904: lectotype, sheet here designated: B [B 10 0154067]; isotypes: B [B 10 0390251]; BR [BR 0000008824059]; GOET [GOET 005682]; G [G 00014887, G 00014888]; L [L 0038043]; LISC [LISC 000393]; HBG [HBG 502533]; MO [MO- 216971]; M [M 0107916]; P [P 00363312]; S [S 07 - 13457]; WAG [WAG 0000096].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BFCCA4C74F3120AA14FDB1E611A2898.taxon	description	Description. Tree to shrub, up to 7 m tall, d. b. h. 3 - 6 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches densely pubescent. Leaves: petiole 4 - 5 mm long, ca. 1 mm in diameter, densely pubescent, grooved, blade inserted on top of the petiole; blade 15 - 21 cm long, 5 - 8 cm wide, ovate to elliptic, apex acuminate, acumen 1 - 2 cm long, base rounded, papyraceous, below densely pubescent with erect slight curly hairs when young and old, densely pubescent when old, above glabrous when young and old, concolorous or discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below densely pubescent when young, densely pubescent when old; secondary veins 15 to 21 pairs, glabrous below; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; peduncule ca. 2 mm long, unbranched; pedicel 10 - 30 mm long, ca. 1 mm in diameter, densely pubescent; in fruit 20 - 30 mm long, 2 - 3 mm in diameter, densely pubescent; bract 1, basal, ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 1 - 2 mm wide, triangular, apex acute, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 15 - 33 mm long, 8 - 15 mm wide, thick, elliptic, apex acute, base narrowed and concave, light green to light yellow to cream, margins flat, pubescent outside, pubescent inside; inner petals 3, valvate, 10 - 15 mm long, 3 - 5 mm wide, triangular, apex acute, base broad and concave, light green to cream, margins flat, pubescent outside, glabrous inside; stamens 30 to 40, in 5 to 6 rows, 2 - 3 mm long, linear; connective tongue shaped, glabrous, red; staminodes absent; carpels free, 20 to 25, ovary ca. 2 mm long, stigma cylindrical, glabrous. Monocarps sessile, 7 to 15, 20 - 30 mm long, ca. 20 mm in diameter, ellipsoid, apex apiculate, pyramidal in shape, pubescent, wrinkled, brown when ripe; seed 1 per monocarp, 11 - 14 mm long, 10 - 11 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BFCCA4C74F3120AA14FDB1E611A2898.taxon	distribution	Distribution. From Cameroon to Gabon; in Cameroon known from the South, Central Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1BFCCA4C74F3120AA14FDB1E611A2898.taxon	materials_examined	Specimens examined. Central Region: Bank Nyong River near the new bridge ca 65 km SSW of Eseka, 3.53 ° N, 10.41 ° E, 16 July 1964, de Wilde W. J. J. O 2840 (WAG); 16 km SSW d'Eseka, 3.65 ° N, 10.78 ° E, 02 January 1974, Letouzey R. 12550 (P, WAG, YA); Route Vetere (Likouk-Lokoundji), 3.48 ° N, 10.32 ° E, 14 January 1974, Mezili P. 251 (P, YA). Littoral Region: Ebo Bekob abandoned village CRES research station, 4.36 ° N, 10.41 ° E, 21 April 2005, Etuge M. 6447 (K). South Region: Campo Ma an National Park 5 km after main entrance, 2.34 ° N, 10.25 ° E, 15 February 2012, Couvreur T. L. P. 384 (WAG, YA); Campo Ma an National Park 5 km after main entrance, 2.35 ° N, 10.25 ° E, 15 February 2012, Couvreur T. L. P. 386 (WAG, YA); Entre Fenda (60 km ESE de Kribi et riviere Kienke), 2.8 ° N, 10.4 ° E, 22 January 1962, Letouzey R. 4092 (K, P, YA); Elom (Kribi), 2.78 ° N, 10.25 ° E, 25 April 1968, Letouzey R. 9436 (P); 55 km ESE de Kribi 2 km W du village, 3.21 ° N, 10.06 ° E, 12 March 1963, Raynal A. 10389 (P, YA); Assok (Nyabessan), 2.77 ° N, 10.47 ° E, 18 February 1965, Raynal A. 13457 (P, YA); 44 km N de Nyabesssan Reserve forestiere de Kienke-sud 500 m N de la route, 2.78 ° N, 10.37 ° E, 18 February 1965, Raynal A. 13472 (P, YA); Campo-Ma'an area Bibambivoto, 2.24 ° N, 10.26 ° E, 23 August 2000, Tchouto Mbatchou G. P. 2986 (KRIBI, WAG, YA); Campo-Ma'an area Mvini, 2.33 ° N, 10.20 ° E, 25 August 2002, Tchouto Mbatchou G. P. CORIX _ 3 (WAG); Campo-Ma'an area Mvini, 2.27 ° N, 10.18 ° E, 14 September 2000, Tchouto Mbatchou G. P. 3035 (KRIBI, WAG); Campo-Ma'an area Bifa, 2.65 ° N, 10.28 ° E, 12 October 2001, Tchouto Mbatchou G. P. 3298 (KRIBI, WAG); Korup National Park Primary rain forest, 5.03 ° N, 8.833 ° E, 01 March 1987, Thomas D. W. 6891 (MO); Bipindi, 3.08 ° N, 10.42 ° E, 1898, Zenker G. A. 1904 (K); Bipindi, 3.08 ° N, 10.42 ° E, 20 June 1904, Zenker G. A. 2877 (B; Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1918, Zenker G. A. 29 (P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3105 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1908, Zenker G. A. 3749 (L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1909, Zenker G. A. 3819 (E, L, M); Bipindi, 3.08 ° N, 10.41 ° E, Zenker G. A. 3897 (K); Bipindi, 3.08 ° N, 10.41 ° E, 01 December 1913, Zenker G. A. 440 (P); Bipindi, 3.08 ° N, 10.41 ° E, 01 November 1913, Zenker G. A. 440 (P, U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1912, Zenker G. A. 4402 (L). South-West Region: Ekundu Kundu, 5.12 ° N, 8.895 ° E, 27 April 1996, Cable S. 2279 (K, YA); Mundemba, 5.05 ° N, 8.883 ° E, 05 March 1993, Gereau R. E. 5185 (MO, WAG); Between Ikenge ad Esukutang ca 6 km W of Ikenge, 5.27 ° N, 9.1 ° E, 03 April 1988, Thomas D. W. 7554 (MO, P, WAG, YA); Korup National Park, 5.33 ° N, 8.9 ° E, 22 May 1988, Thomas D. W. 7809 (MO, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49A353E1C190B23034C951EACACC33DB.taxon	description	Figs 83, 84; Map 10 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49A353E1C190B23034C951EACACC33DB.taxon	materials_examined	Type. Gabon. Ogooue-Lolo; Lastoursville, Le Testu G. M. P. C. 8635, 28 Dec 1930: lectotype, sheet here designated: BM [BM 000553861]; isotypes: BM [BM 000553860, BM 000553862]; BR [BR 0000008802705, BR 0000008802378]; P [P 00363316, P 00363315].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49A353E1C190B23034C951EACACC33DB.taxon	description	Description. Tree, 5 - 25 m tall, d. b. h. 10 - 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches tomentose, brown. Leaves: petiole 7 - 14 mm long, 1 - 3 mm in diameter, tomentose brown, slightly grooved to cylindrical, blade inserted on the side of the petiole; blade 7.5 - 31 cm long, 2.8 - 10.3 cm wide, oblong to elliptic, apex acuminate to acute, acumen 0.3 - 2.4 cm long, base rounded to obtuse, papyraceous, below densely pubescent with erect slight curly hairs when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young, glabrous when old, below pubescent when young and old; secondary veins 15 to 24 pairs, pubescent below; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 2 to 8 per inflorescence, peduncle arbuscular-like, 8 - 26 mm long; pedicel 22 - 46 mm long, ca. 1 mm in diameter, pubescent; in fruit 30 - 60 mm long, 3 - 4 mm in diameter, pubescent; bract 1, basal, ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, ca. 1 mm long, 1 - 2 mm wide, semiorbicular, apex acuminate, base truncate, brown, pubescent outside, pubescent inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 16.2 - 23.3 mm long, 3 - 4.6 mm wide, narrowly triangular, very thick, apex attenuate, base suborbicular and concave, brown, margins flat, pubescent to glabrous outside, densely pubescent inside; inner petals 3, valvate, 5.6 - 6 mm long, 1.4 - 2.9 mm wide, triangular, apex acute, base broad and concave, margins flat, pubescent outside, glabrous inside; stamens 160 to 170, in 5 to 6 rows, 2 mm long, linear; connective tongue shaped, glabrous, brown; staminodes absent; carpels free, ca. 144, ovary ca. 1 mm long, stigma filiform, glabrous. Monocarps stipitate, stipes 10 - 50 mm long, 1 - 2 mm in diameter; monocarps 19 to 144, 9 - 14 mm long, 5 - 10 mm in diameter, ellipsoid, apex apiculate, densely pubescent, smooth, not ribbed, brown when ripe; seed 1 per monocarp, 7 - 12 mm long, 4 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49A353E1C190B23034C951EACACC33DB.taxon	distribution	Distribution. Southern Cameroon and Gabon; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49A353E1C190B23034C951EACACC33DB.taxon	materials_examined	Specimens examined. South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 11 February 2015, Couvreur T. L. P. 675 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.34 ° E, 11 February 2015, Couvreur T. L. P. 678 (WAG, YA); Efoulan au sud d'Akom II (mi-chemin entre Ebolowa et Kribi), 2.76 ° N, 10.53 ° E, 09 May 2009, Lachenaud O. L. 659 (BR, BR, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0F46348A411CB0578FBE59C4022D38E2.taxon	description	Description. Trees, 1 - 20 m tall, d. b. h. up to 21 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 2 - 11 mm long, 1 - 6 mm in diameter; blade 7 - 66 cm long, 3 - 3 cm wide, ovate or elliptic or obovate or oblong, apex acuminate to emarginate, acumen 0.2 - 2.7 cm long, base cordate to decurrent, blade inserted on top of the petiole, discolorous, whitish below; midrib sunken or flat; secondary veins 14 to 65 pairs; tertiary venation percurrent. Inflorescence variable in length from compact to lax, composed of a peduncle like base and axial internodes. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary, 1 to 28 per inflorescence; pedicel 2 - 20 mm long; in fruit 9 - 25 mm long; bracts 2, one basal and one upper towards the lower half of pedicel; sepals 3, valvate, free, 2 - 12 mm long, apex acute or acuminate, base truncate; petals free, valvate, outer petals shorter than inner; outer petals 3, 5 - 28 mm long, 1.5 - 11 mm wide, elliptic or ovate or obovate, apex acute or acuminate, base truncate; inner petals 3, valvate, 15 - 80 mm long, 4 - 20 mm wide, elliptic or ovate, apex acute, base truncate; stamens numerous, 1 - 2 mm long, broad; connective discoid; staminodes absent; carpels free, 1 to 15, ovary 1 - 3 mm long, stigma globose or lobed. Monocarps sessile, 1 to 6, 13 - 65 mm long, 8 - 40 mm in diameter, globose to oblong, apex acute or rounded or cuspidate; seeds biseriate, 1 - 15 mm long, 2 - 10 mm in diameter, ellipsoid; aril absent. A genus of 13 species from West and Central Africa (no species yet known from East Africa); all species are present in Cameroon, six endemic, making it the center of diversity for this genus. Piptostigma and Brieya are the only Annonaceae genera in Cameroon with longer inner petals than outer. Piptostigma has spectacular inflorescences that are unique among Cameroonian Annonaceae. They appear complex, but are in fact simple and, the different inflorescence types are an important character to identify species. First, inflorescences appear to be " stalked " stalked ", that is they have a short woody peduncle at the based on the inflorescence structure and is probably the result of fallen old inflorescences. This peduncle is variable in length. Then follows the indeterminate part or sympodial rachis, which contains axial internodes variable in length. Different lengths between the internodes will lead to different overall aspects of the inflorescence. Following Ghogue et al. (2017) and Maas et al. (2003) we can distinguish three types: compact: internodes are shorter than 1 cm giving a very compact appearance to the inflorescence. lax or sub-lax: internodes are between 1 and 4 cm long, leading to more or less long inflorescences. This is found in most species. panicle-like: internodes between 5 and 10.5 cm long leading to long pendant inflorescences. We shall refer to these terms in the descriptions below and the key. Finally, we end up with the pedicel and flower per se.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F1E750249F55F35DE69BCE5DC31912BD.taxon	description	Figs 85, 86; Map 10 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F1E750249F55F35DE69BCE5DC31912BD.taxon	materials_examined	Type. Cameroon. South Region; Ebolowa, Mildbraed G. W. J. 5791, 4 Jul 1911: holotype: B [B 100154077]; isotypes: HBG [HBG 502525, HBG 502527, HBG 50252]; L [L 0196773]; P [P 00363307, P 00363307]; YA [YA 0002833].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F1E750249F55F35DE69BCE5DC31912BD.taxon	description	Description. Tree, 8 - 10 m tall, d. b. h. 6 - 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches tomentose. Leaves: petiole 4 - 11 mm long, 3 - 6 mm in diameter, tomentose, cylindrical, blade inserted on top of the petiole; blade 30 - 66 cm long, 14 - 34 cm wide, obovate, apex emarginate to mucronate, acumen 0.7 - 0.8 cm long, base cordate to acute, coriaceous, below densely pubescent when young, densely pubescent when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young, sparsely pubescent when old, below pubescent when young, glabrous when old; secondary veins 30 to 40 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, 2 - 2.5 cm long, peduncle like base 5 - 9 mm, axial internodes 2 - 4 mm long, compact, sympodial rachis 10 - 15 mm long. Flowers with 9 perianth parts in 3 whorls, 3 to 6 per inflorescence; pedicel ca. 6 mm long, ca. 3 mm in diameter, tomentose; in fruit 9 - 11 mm long, 3 - 4 mm in diameter, tomentose; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract ca. 7 mm long, ca. 7 mm wide; upper bract ca. 7 mm long, ca. 7 mm wide; sepals 3, valvate, free, 4 - 5 mm long, 4 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 5 - 7 mm long, 3 - 4 mm wide, ovate, apex acuminate to obtuse, base truncate, light yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 50 - 60 mm long, 6 - 8 mm wide, ovate, apex acute, base truncate, pink to yellow, margins wavy, pubescent outside, pubescent inside; stamens 70 to 150, in 5 to 6 rows, 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 8 to 10, ovary ca. 3 mm long, stigma globose, densely pubescent. Monocarps sessile, 2 to 5, 12 - 35 mm long, 10 - 28 mm in diameter, obovoid, apex cuspidate, tomentose, smooth, not ribbed, brown when ripe; seeds number not counted, ca. 15 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F1E750249F55F35DE69BCE5DC31912BD.taxon	distribution	Distribution. From Cameroon to Gabon; in Cameroon known from the Central, South and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
F1E750249F55F35DE69BCE5DC31912BD.taxon	materials_examined	Specimens examined. Central Region: Edea (Mangombe), 3.8 ° N, 10.13 ° E, 01 January 1963, Letouzey R. s. n. (P). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.39 ° E, 28 February 2018, Couvreur T. L. P. 1167 b (K, MPU, P, WAG, YA); Bassin du Mungo, 4.17 ° N, 9.52 ° E, 01 June 1917, Fleury F. 33400 (P); Edea Mangombe, 3.86 ° N, 10.14 ° E, 01 January 1956, Letouzey R. 1936 (P, YA); Route Douala-Edea pres du km 28, 4.05 ° N, 9.689 ° E, 18 January 1962, Letouzey R. 4011 (P, YA). South Region: Ngovayang, 3.23 ° N, 10.57 ° E, 01 June 2015, Kamdem N. 295 (YA); Pres Akak 10 km W Sangmeli 2.96 ° N, 11.88 ° E, 11 March 1970, Letouzey R. 10157 (P, YA); Piste Meyo Ntem-Evouzok 75 km W Ambam Entre 1 er et 3 e bras du Ntem, 2.28 ° N, 10.52 ° E, 28 November 1979, Letouzey R. 15272 (P, YA); Ebolowa Cameroon, 2.76 ° N, 10.91 ° E, 04 July 1911, Mildbraed G. W. J. 5791 (B, L, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BF888A3CC32EA3E5024971242702FBF7.taxon	description	Fig. 86; Map 10 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BF888A3CC32EA3E5024971242702FBF7.taxon	materials_examined	Type. Ivory Coast: Bas-Sassandra; Cavally basin, Tepos country: Grabo and surrounding villages, at the foot of Mount Cope, Chevallier A. J. B. 19620, 26 - 28 Jun 1907: lectotype, designated by Ghogue et al. (2017), p. 190: P [P 00363296]; isolectotype: P [P 00363297].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BF888A3CC32EA3E5024971242702FBF7.taxon	description	Description. Tree to shrub, 1 - 10 m tall, d. b. h. 6 - 7 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, densely pubescent, cylindrical, blade inserted on top of the petiole; blade 7 - 24 cm long, 3 - 6.5 cm wide, obovate, apex acuminate to acute, acumen 0.3 - 0.7 cm long, base obtuse to acute, papyraceous, below pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below densely pubescent when young, densely pubescent when old; secondary veins 15 to 23 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, axillary, peduncle-like base not apparent, axial internodes absent, compact in aspect, sympodial rachis 5 - 20 mm long. Flowers with 9 perianth parts in 3 whorls; 1 per inflorescence; pedicel 4 - 6 mm long, ca. 2 mm in diameter, densely pubescent; in fruit 11 - 13 mm long, ca. 3 mm in diameter, tomentose; basal bract ca. 3 mm long, ca. 1 mm wide; upper bract ca. 3 mm long, ca. 1 mm wide; sepals 3, valvate, free, ca. 5 mm long, ca. 2 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, ca. 11 mm long, ca. 1.5 mm wide, ovate, apex acute, base truncate, light yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 20 - 40 mm long, 4 - 10 mm wide, ovate, apex acute, base truncate, margins wavy, pubescent outside, sparsely pubescent to glabrous inside; stamens 100 to 120, in 6 to 7 rows, 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 1 to 4, ovary ca. 2 mm long, stigma globose, sparsely pubescent. Monocarps sessile, 1 to 3, ca. 45 mm long, ca. 20 mm in diameter, ellipsoid to ovoid, apex cuspidate, pubescent, smooth, fleshy, white when ripe; seeds 5 per monocarp, ca. 13 mm long, ca. 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BF888A3CC32EA3E5024971242702FBF7.taxon	distribution	Distribution. A mainly West African species from Liberia to Togo and Nigeria; in Cameroon known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
BF888A3CC32EA3E5024971242702FBF7.taxon	materials_examined	Specimens examined. South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.52 ° N, 10.32 ° E, 11 February 2015, Couvreur T. L. P. 681 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 693 (WAG, YA). South-West Region: Ekundu Kundu to Erat ca 25 km, 5.13 ° N, 8.869 ° E, 26 April 1996, Cheek M. 8198 (K, WAG, YA); Edensueh forest, 5.25 ° N, 9.576 ° E, 30 November 2000, Etuge M. 4853 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0DD2EC1971D074C681F31AEE59EC669.taxon	description	Figs 87, 94; Map 11 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0DD2EC1971D074C681F31AEE59EC669.taxon	materials_examined	Type. Equatorial Guinea. Rio Muni; Kongui River, Mann G. 1792, Aug. - Sep 1862: holotype: B [B 100154078]; isotype: K [K 000199000, K 000199706]; P [P 00363305].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0DD2EC1971D074C681F31AEE59EC669.taxon	description	Description. Tree, 8 - 10 m tall, d. b. h. 12 - 15 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 12 - 30 cm long, 4 - 8.5 cm wide, obovate, apex acuminate, acumen 1.4 - 1.7 cm long, base obtuse to acute, papyraceous, below pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below pubescent when young and old; secondary veins 19 to 27 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences cauliflorous, peduncle like base 5 - 20 mm long, axial internodes 10 - 150 mm long, lax to panicle-like, sympodial rachis 20 - 70 mm long. Flowers with 9 perianth parts in 3 whorls, 10 to 28 per inflorescence; pedicel 11 - 13 mm long, 2 mm in diameter, pubescent; in fruit 16 - 20 mm long, 6 mm in diameter, densely pubescent; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 4 - 5 mm long, 3 mm wide; upper bract 2 - 5 mm long, 1 - 2 mm wide; sepals 3, valvate, free, 4 mm long, 2 mm wide, triangular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 5 - 6 mm long, 2 mm wide, ovate, apex acute, base truncate, red, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 30 - 50 mm long, 5 - 10 mm wide, ovate, apex acute, base truncate, red, margins wavy, pubescent outside, sparsely pubescent inside; stamens 90 to 110, in 6 to 7 rows, ca. 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 5 to 8, ovary ca. 1 mm long, stigma globose, pubescent. Monocarps sessile, ca. 8, 20 - 65 mm long, 9 - 21 mm in diameter, oblong to narrowly oblong, apex sometimes mucronate, glabrous or sometimes very sparsely pubescent, puncticulate (covered by small blister-like pimples or wards), irregularly and longitudinally ribbed, color unknown; seeds 1 to 7 per monocarp, ca. 15 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0DD2EC1971D074C681F31AEE59EC669.taxon	distribution	Distribution. From Cameroon to Gabon, one specimen from Republic of Congo; in Cameroon known from the South, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C0DD2EC1971D074C681F31AEE59EC669.taxon	materials_examined	Specimens examined. Littoral Region: Douala (route Razel), 4.05 ° N, 9.71 ° E, 01 January 1955, Endengle E. 2061 (P); Foret de Bakaka 3 km E of Ebone a village on km 11 Nkongsamba-Loum Road, 4.83 ° N, 9.9 ° E, 13 September 1971, Leeuwenberg A. J. M. 8354 (WAG, YA); Hikoa Mahouda (chaine de la L'Hikoa Mandeng- 30 km ENE Edea), 3.87 ° N, 10.39 ° E, 17 December 1973, Letouzey R. 12415 (K); Tissongo, 3.58 ° N, 9.9 ° E, 07 July 1976, McKey D. B. 103 (K). South Region: SE slopes of Mt elephant SE of Kribi, 2.78 ° N, 10.01 ° E, 26 February 1970, Bos J. J. 6415 (BR, C, K, LD, P, WAG, YA); ca 3 km N of Lolodorf road, 3.01 ° N, 10.05 ° E, 18 March 1970, Bos J. J. 6594 (BR, K, LD, LM, MO, P, WAG, YA); 15 km before Kribi on N 7 road from Ed 3.14 ° N, 9.968 ° E, 27 February 2018, Couvreur T. L. P. 1158 (MPU, P, WAG, YA); Nkoemvone, 2.81 ° N, 11.13 ° E, 09 September 1975, de Wilde J. J. F. E 8453 a (BR, K, MO, P, WAG); Pres Bella (45 km NE de Kribi), 3.25 ° N, 10.2 ° E, 25 January 1962, Letouzey R. 4167 (P, YA); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1901, Zenker G. A. 2396 (B, BR, G, K, L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1908, Zenker G. A. 3654 (BR, L, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1911, Zenker G. A. 4006 (K, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1912, Zenker G. A. 4472 (BR, L, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1914, Zenker G. A. 505 (B, G, P, WAG). South-West Region: mountain circular road S of Koto II and the Onge river, 4.32 ° N, 9.064 ° E, 21 October 1993, Cheek M. 5071 (K, YA); Korup National Park nature trail near suspension bridge, 4.98 ° N, 8.85 ° E, 01 February 1995, Cheek M. 7230 (K, YA); Pente septentrionale de Nta Ali (1266) m) descente de la cote 1009 a Mbio 30 km SE de Mamfe, 5.59 ° N, 9.503 ° E, 21 June 1975, Letouzey R. 13912 (P, YA); Barombi F R Station, 4.65 ° N, 9.4 ° E, 1890, Preuss P. R. 251 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B27EEEE4F15306D3DEFDB4CE9BF2529B.taxon	description	Fig. 87; Map 11 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B27EEEE4F15306D3DEFDB4CE9BF2529B.taxon	materials_examined	Type. Cameroon. Littoral Region; around Douala, Fleury F. 33134, Jun 1917: holotype: P [P 02032174]; isotypes: P [P 02032172, P 02032173, P 02032175].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B27EEEE4F15306D3DEFDB4CE9BF2529B.taxon	description	Description. Tree, 8 - 30 m tall, d. b. h. 12 - 15 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 9.5 - 26 cm long, 3.5 - 10 cm wide, obovate, apex acuminate, acumen 0.5 - 1.8 cm long, base obtuse to acute, papyraceous, below sparsely pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below pubescent to tomentose when young, pubescent to tomentose when old; secondary veins 17 to 33 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, peduncle like base 15 - 65 mm long, axial internodes 5 - 35 mm long, compact to lax, sympodial rachis 6 - 38 mm long. Flowers with 9 perianth parts in 3 whorls, (1) 2 to 7 per inflorescence; pedicel ca. 7 mm long, ca. 2 mm in diameter, pubescent; in fruit 10 - 15 mm long, ca. 4 mm in diameter, tomentose; basal bract ca. 8 mm long, ca. 5 mm wide; upper bract ca. 4 mm long, ca. 2 mm wide; sepals 3, valvate, free, ca. 12 mm long, ca. 4 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, ca. 15 mm long, ca. 4 mm wide, ovate, apex acute, base truncate, margins flat, tomentose outside, glabrous inside; inner petals 3, valvate, 38 - 45 mm long, 10 - 12 mm wide, ovate, apex acute, base truncate; stamens numerous, in 6 to 8 rows, ca. 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 4 to 7, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 3 to 4, 20 - 35 mm long, 10 - 20 mm in diameter, ellipsoid to ovoid, apex rounded, sparsely pubescent to glabrous, smooth, bumpy when dry, pink reddish when ripe; seeds 4 to 5 per monocarp, ca. 10 mm long, ca. 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B27EEEE4F15306D3DEFDB4CE9BF2529B.taxon	distribution	Distribution. endemic to Cameroon, known from the Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B27EEEE4F15306D3DEFDB4CE9BF2529B.taxon	materials_examined	Specimens examined. Littoral Region: Aux environs de Douala, 4.05 ° N, 9.7 ° E, 01 June 1917, Fleury F. 33134 (P); Tissongo, 3.58 ° N, 9.9 ° E, 08 July 1976, McKey D. B. 105 (K); Douala-Edea Reserve Lombe Camp, 3.48 ° N, 9.833 ° E, 01 November 1977, Thomas D. W. 510 (K); Lombe Camp Site Douala-Edea Reserve Cameroun, 3.48 ° N, 9.833 ° E, 30 May 1976, Waterman P. G. 801 (K). South-West Region: Nta Ali SE Mamfe, 5.55 ° N, 9.521 ° E, 17 June 1987, Achoundong G. 1267 (YA); Kupe village, 4.78 ° N, 9.683 ° E, 24 January 1995, Cable S. 787 (K, WAG, YA); Mungo River F. R., 4.78 ° N, 9.607 ° E, 30 November 1999, Cheek M. 10197 (K); Kupe village, 4.77 ° N, 9.701 ° E, 16 November 1995, Cheek M. 7849 (K, YA); Kupe village, 4.79 ° N, 9.701 ° E, 19 May 1996, Cheek M. 8328 (K, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 26 March 2016, Couvreur T. L. P. 1017 (WAG, YA); Bayang Mbo Wildlife Sanctuary before Mbu river, 5.34 ° N, 9.487 ° E, 27 March 2016, Couvreur T. L. P. 1023 (WAG, YA); on trail leading to top of Mt Etinde after Ekonjo village, 4.07 ° N, 9.152 ° E, 01 April 2016, Couvreur T. L. P. 1030 (WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.100 ° E, 03 April 2016, Couvreur T. L. P. 1047 (WAG, YA); on top of hill near Small Ekombe village 3 km after Kumba on road to Ekondo Titi town, 4.62 ° N, 9.378 ° E, 13 January 2016, Couvreur T. L. P. 983 (WAG, YA); Ezeze road Nyasoso between shrike and Max's trail following the river upwards, 4.82 ° N, 9.691 ° E, 25 June 1996, Etuge M. 2420 (K, WAG, YA); Kupe village, 4.76 ° N, 9.699 ° E, 09 July 1996, Etuge M. 2698 (K, WAG, YA); Massif Ntali pente NW 30 km SE Mamfe, 5.56 ° N, 9.482 ° E, 15 June 1982, Villiers J. - F. 1448 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FF9307EA03878FB876440F268E1C9D8C.taxon	description	Fig. 88; Map 11 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FF9307EA03878FB876440F268E1C9D8C.taxon	materials_examined	Type. Cameroon. South Region; around Bipinde, Zenker G. A. 1075, 1896: holotype: B [B 100154079]; isotypes: HBG [HBG 502530]; L [L 0183458]; M [M 0107925]; NY [NY 00026195]; P [P 00363294, P 00363295]; S [S 07 - 13478]; WU [WU 0025868].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FF9307EA03878FB876440F268E1C9D8C.taxon	description	Description. Tree, 4 - 16 m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches densely hispid, hairs 4 - 5 mm long. Leaves: petiole 4 - 7 mm long, 2 - 3 mm in diameter, hispid, hairs 1 - 5 mm long, cylindrical, blade inserted on top of the petiole; blade 7.7 - 24.3 cm long, 4 - 10.3 cm wide, obovate, apex acuminate, acumen 0.3 - 1 cm long, base rounded to obtuse, papyraceous, below glabrous when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above densely pubescent when young and old, below pubescent when young and old; secondary veins 14 to 34 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, pubescent all over, especially younger parts, with hairs 1 - 2 mm long, peduncle like base 5 - 20 mm long, axial internodes 14 - 40 mm long, lax to sub-lax, sympodial rachis 45 - 150 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 9 - 12 mm long, ca. 2 mm in diameter, densely pubescent; in fruit 10 - 12 mm long, 4 mm in diameter, pubescent; basal bract 6 - 12 mm long, 4 - 6 mm wide; upper bract 6 - 12 mm long, 4 - 6 mm wide; sepals 3, valvate, free, ca. 6 mm long, 4 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 6 - 10 mm long, 3 - 4 mm wide, elliptic, apex acute, base truncate, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 40 - 80 mm long, 10 - 15 mm wide, elliptic, apex acute, base truncate, margins wavy, densely pubescent outside, pubescent towards base inside; stamens 70 to 90, in 5 to 6 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 3 to 5, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 1 to 3, ca. 15 mm long, ca. 20 mm in diameter, ellipsoid, apex rounded, pubescent, warty, bumpy, orange, pink to yellow when ripe; seeds 10 to 12 per monocarp, ca. 10 mm long, ca. 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FF9307EA03878FB876440F268E1C9D8C.taxon	distribution	Distribution. endemic to Cameroon, known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FF9307EA03878FB876440F268E1C9D8C.taxon	materials_examined	Specimens examined. South Region: Route de Bipindi a Dihani, 3.26 ° N, 10.20 ° E, 24 June 1928, Annet E. 420 (P); ca 45 k from Kribi ca 8 km N of Lolodorf road, 3.1 ° N, 10.25 ° E, 02 April 1970, Bos J. J. 6683 (BR, K, LD, LM, MO, P, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 1896, Zenker G. A. 1075 (L, P, WAG). South-West Region: Mokoko Forest Reserve Boa / Mokoko, 4.42 ° N, 8.972 ° E, 31 May 1994, Acworth J. M. 289 (K, YA); Mokoko Forest Reserve Dikome, 4.48 ° N, 9.033 ° E, 02 May 1994, Ekema S. N. 862 (K, YA); Aikoume (?) Banondo (?) forest (Cameroun), 4.45 ° N, 9.016 ° E, 03 May 1994, Sonke B. 1173 (BR, WAG); Mokoko Mount above Bonja village, 4.46 ° N, 9.1 ° E, 23 March 1993, Tchouto Mbatchou G. P. 613 (K); Korup National Park Primary rain forest, 5.03 ° N, 8.833 ° E, 01 March 1987, Thomas D. W. 6906 (MO); Mokoko Forest Reserve Ekombe-Mofako, 4.46 ° N, 9.066 ° E, 21 May 1994, Watts J. 1104 (K, YA); Likingi 35 hrs S on Wonge river E hills, 4.33 ° N, 8.98 ° E, 23 March 1993, Wheatley J. I. 814 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E4B3535A8C63756F0503B50ADB680FC.taxon	description	Figs 88, 89; Map 11 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E4B3535A8C63756F0503B50ADB680FC.taxon	description	Description. Tree, 6 - 18 m tall, d. b. h. 15 - 21 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 2 - 3 mm long, 2 - 3 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 14 - 31 cm long, 7 - 9 cm wide, ovate to oblong, apex acuminate, acumen 0.5 cm long, base rounded to cordate, subcoriaceous, below pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below densely pubescent when young, densely pubescent when old; secondary veins 16 to 30 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, pubescent all over, peduncle like base 6 - 25 mm long, axial internodes 10 - 20 mm long, compact to sub-lax, sympodial rachis 10 - 70 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 4 per inflorescence; pedicel 5 - 20 mm long, 3 - 5 mm in diameter, tomentose; in fruit 20 - 25 mm long, ca. 6 mm in diameter, pubescent; basal bract ca. 10 mm long, ca. 10 mm wide; upper bract ca. 10 mm long, ca. 10 mm wide; sepals 3, valvate, free, ca. 9 mm long, ca. 8 mm wide, triangular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 12 - 28 mm long, 9 - 11 mm wide, obovate, apex acute, base truncate, wine-brown, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 40 - 55 mm long, 10 - 20 mm wide, elliptic, apex acute, base truncate, cream, margins wavy, pubescent outside, pubescent inside; stamens 280 to 320, in 10 to 11 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 9 to 12, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 3 to 6, 30 - 40 mm long, 25 - 40 mm in diameter, obovoid, apex acute, pubescent, muricate with projections up to 10 mm long, not ribbed, projections brown otherwise reddish when mature; seeds 2 to 6 per monocarp, ca. 10 mm long, ca. 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E4B3535A8C63756F0503B50ADB680FC.taxon	distribution	Distribution. From Cameroon to the Republic of Congo; in Cameroon known from the South and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2E4B3535A8C63756F0503B50ADB680FC.taxon	materials_examined	Specimens examined. Central Region: Ca 50 km S of Badjob ca 60 km SW of Eseka, 3.46 ° N, 10.5 ° E, 19 March 1964, de Wilde W. J. J. O 2132 (BR, P, WAG); Akomimbang (Mbalmayo), 3.52 ° N, 11.5 ° E, 13 November 1957, Letouzey R. 1937 (P); Akomimbang Mbalmayo, 3.52 ° N, 11.5 ° E, 13 November 1957, Letouzey R. s. n. (P, YA). South Region: 10 km From Kribi Lolodorf road, 2.98 ° N, 9.966 ° E, 27 May 1969, Bos J. J. 4647 (BR, K, MO, P, U, WAG, YA); Bezirk Kribi Vorland mit einzeln Hugeln bei Adjab 25 km Grand batanga near Eduma and Bidue, 2.93 ° N, 9.92 ° E, 25 July 1911, Mildbraed G. W. J. 6118 (HBG); 51 SE de Campo Nkolmenbegue, 2.39 ° N, 10.3 ° E, 24 November 1992, Satabie B. 1037 (YA); Efoulan, 2.74 ° N, 10.54 ° E, 04 December 2000, Tchouto Mbatchou G. P. 3100 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1903, Zenker G. A. 2528 (B, BR, G, L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 May 1902, Zenker G. A. s. n. (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E0824296D39B8BB2A1819746EFC63498.taxon	description	Fig. 89; Map 11 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E0824296D39B8BB2A1819746EFC63498.taxon	materials_examined	Type. Cameroon. South-West Region; above small Koto village (Mt. Cameroon), Thomas D. W. 4493, 6 Mar 1985: holotype: YA [YA 0002852]; isotypes: MO [MO 3282523]; P [P 00284016].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E0824296D39B8BB2A1819746EFC63498.taxon	description	Description. Tree, 5 - 10 m tall, d. b. h. 10 - 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 6 - 7 mm long, 3 - 4 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 25 - 41 cm long, 9 - 16 cm wide, obovate, apex acuminate, acumen ca. 2.5 cm long, base acute, papyraceous, below glabrous when young and old, above sparsely pubescent when young, sparsely pubescent when old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below sparsely pubescent when young and old; secondary veins 21 to 28 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous or ramiflorous on old leafless branches, axillary, pubescent all over, peduncle like base 5 - 10 mm long, axial internodes 5 - 15 mm long, compact to sub-lax, sympodial rachis up to 70 mm long. Flowers with 9 perianth parts in 3 whorls, 3 to 6 per inflorescence; pedicel 8 - 10 mm long, ca. 2 mm in diameter, tomentose; in fruit 10 - 12 mm long, ca. 4 mm in diameter, glabrous; basal bract ca. 12 mm long, ca. 3 mm wide; upper bract ca. 12 mm long, ca. 3 mm wide; sepals 3, valvate, free, ca. 9 mm long, ca. 3 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 9 - 15 mm long, 2.5 - 3 mm wide, narrowly elliptic, apex acute, base truncate, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 25 - 35 mm long, 4 - 5 mm wide, ovate, apex acute, base truncate, margins wavy, pubescent outside, pubescent inside; stamens numerous, in 6 to 8 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, ca. 5, ovary ca. 3 mm long, stigma globose, densely pubescent. Monocarps sessile, 1 to 3, ca. 33 mm long, ca. 35 mm in diameter, globose, apex rounded, glabrous, verrucose to muricate with short projections, not ribbed, light red pink when ripe; seeds 2 to 4 per monocarp, 10 - 15 mm long, ca. 12 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E0824296D39B8BB2A1819746EFC63498.taxon	distribution	Distribution. endemic to Cameroon, known from the South, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E0824296D39B8BB2A1819746EFC63498.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Forest Reserve Djuma permanent camp on transect 5, 4.33 ° N, 10.23 ° E, 16 February 2013, Couvreur T. L. P. 637 (WAG, YA). South Region: Bord de la Lobe 25 km E Campo, 2.37 ° N, 9.82 ° E, 01 January 1968, Letouzey R. 9156 (P, YA). South-West Region: Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.054 ° E, 02 April 2016, Couvreur T. L. P. 1034 (WAG, YA); on trail through palm oil plantation 3 km before lava flow and Seme Beach hotel when coming from Limbe, 4.06 ° N, 9.079 ° E, 18 October 2013, Couvreur T. L. P. 518 (WAG, YA); Entre DikomeBalue (1200 m) et Ifanga Nalende (650 m) 35 km NNW-Kumba, 4.9 ° N, 9.29 ° E, 25 March 1976, Letouzey R. 14590 (P, YA); Etinde forest reserve Njonji lake, 4.13 ° N, 9.033 ° E, 25 January 1993, Tchouto Mbatchou G. P. 1053 (K); Disturbed forest Bomana and Koto II, 4.3 ° N, 9.05 ° E, 26 April 1996, Tchouto Mbatchou G. P. 1378 (K, YA); Mount Cameroon above small Koto village, 4.3 ° N, 9.1 ° E, 06 March 1985, Thomas D. W. 4493 (YA); Between Ikenge and Esukutang ca 6 kms West of Ikenge, 5.28 ° N, 9.083 ° E, 03 April 1988, Thomas D. W. 7645 (YA); Cameroon Mountain, 4.12 ° N, 9.029 ° E, 20 June 2001, van Andel T. R. 3728 (U, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D7595B396FA537079BDC83F0574FAAE2.taxon	description	Map 11 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D7595B396FA537079BDC83F0574FAAE2.taxon	materials_examined	Type. Cameroon. South-West Region; Korup National Park, van der Burgt R. 689, 25 Aug 2004: holotype: WAG [WAG 0204511]; isotypes: BR [BR 0000013174750]; P [P 06901232].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D7595B396FA537079BDC83F0574FAAE2.taxon	description	Description. Tree, up to 10 m tall, d. b. h. 17 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent. Leaves: petiole 6 - 8 mm long, 2 - 3 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 23.5 - 36.5 cm long, 6.5 - 8.5 cm wide, narrowly to very narrowly elliptic or narrowly obovate (rarely), apex obtuse, base obtuse, papyraceous, below sparsely pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below pubescent when young, glabrous when old; secondary veins 22 to 32 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous or ramiflorous on old leafless branches, axillary, peduncle like base 15 - 30 mm long, axial internodes 1 - 50 mm long, lax or panicle-like, sympodial rachis 30 - 100 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 5 per inflorescence; pedicel 5 - 7 mm long, ca. 2 mm in diameter, densely pubescent; in fruit 15 mm long, 3 mm in diameter, pubescent; basal bract ca. 3 mm long, ca. 3 mm wide; upper bract ca. 2 mm long, ca. 2 mm wide; sepals 3, valvate, free, 2 - 3 mm long, 2 - 3 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 5 - 6 mm long, ca. 5 mm wide, obovate, apex acute, base truncate, pale yellow to cream, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 30 - 40 mm long, 8 - 10 mm wide, elliptic, apex acute, base truncate, pale yellow to cream, margins wavy, densely pubescent outside, pubescent inside; stamens numerous, in 6 to 8 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 4, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 1 to 3, 21 - 43 mm long, 27 - 53 mm in diameter, ellipsoid, apex rounded, pubescent, verrucose with short flat projections, not ribbed, light red all over when ripe; seeds up to 6 per monocarp, ca. 16 mm long, ca. 10 mm in diameter, transversely ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D7595B396FA537079BDC83F0574FAAE2.taxon	distribution	Distribution. endemic to Cameroon, known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D7595B396FA537079BDC83F0574FAAE2.taxon	materials_examined	Specimens examined. South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.52 ° N, 10.32 ° E, 11 February 2015, Couvreur T. L. P. 672 (P, WAG, YA). South-West Region: Korup National Park Ndian, 5.06 ° N, 8.866 ° E, 19 June 2000, Sainge M. 558 (MO); Korup National Park, 5.01 ° N, 8.85 ° E, 10 March 1986, Thomas D. W. 5840 (MO); Korup National Park, 5.01 ° N, 8.8 ° E, 25 August 2004, van der Burgt X. M. 689 (BR, G, K, MO, P, WAG, YA); Korup National Park, 5.01 ° N, 8.8 ° E, 29 October 2005, van der Burgt X. M. 790 (K, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FEDE24A592618745A75BF87065D2DA98.taxon	description	Figs 90, 92; Map 11 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FEDE24A592618745A75BF87065D2DA98.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Dundusana, Mortehan M. G. 626, Oct 1913: lectotype, sheet here designated: BR [BR 0000008802118]; isotype: BR [BR 0000008802798].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FEDE24A592618745A75BF87065D2DA98.taxon	description	Description. Tree, 8 - 10 m tall, d. b. h. 18 cm; stilt roots or buttresses absent, trunk funneled. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent to hirsute, hairs to 2 mm long. Leaves: petiole 3 - 4 mm long, 2 - 3 mm in diameter, hirsute, cylindrical, blade inserted on top of the petiole; blade 11.5 - 20.1 cm long, 3 - 12.5 cm wide, obovate, apex acuminate, acumen 0.2 - 0.6 cm long, base acute, papyraceous, below sparsely pubescent when young and old, above pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below densely pubescent to tomentose when young and old; secondary veins 24 to 34 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, peduncle like base 18 - 30 mm long, axial internodes 10 - 105 mm long, lax to panicle-like, sympodial rachis 40 - 320 mm long. Flowers with 9 perianth parts in 3 whorls, 2 to 8 per inflorescence; pedicel 5 - 8 mm long, ca. 2 mm in diameter, pubescent; in fruit 10 - 15 mm long, ca. 3 mm in diameter, pubescent; basal bract ca. 6 mm long, ca. 4 mm wide; upper bract ca. 6 mm long, ca. 4 mm wide; sepals 3, valvate, free, 4 - 5 mm long, 4 - 5 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 8 - 9 mm long, 4 - 5 mm wide, ovate, apex acute, base truncate, yellow, margins flat, pubescent outside, glabrous to pubescent, base glabrous inside; inner petals 3, valvate, 15 - 20 mm long, 4 - 7 mm wide, elliptic, apex acute, base truncate, greenish yellow, margins wavy, pubescent outside, tomentose inside; stamens 40, in 6 to 7 rows, ca. 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 4 to 5, ovary ca. 2 mm long, stigma lobed, pubescent. Monocarps sessile, 1 to 2, 18 - 23 mm long, ca. 18 mm in diameter, ellipsoid, apex mucronate or rounded, pubescent to tomentose, verrucose, irregularly ribbed, white pink turning wine red to dark sepia when ripe; seeds 2 to 8 per monocarp, ca. 12 mm long, ca. 15 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FEDE24A592618745A75BF87065D2DA98.taxon	distribution	Distribution. From Cameroon to the Democratic Republic of the Congo, one specimen from Ghana and two from Gabon; in Cameroon known from the South, Central, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FEDE24A592618745A75BF87065D2DA98.taxon	materials_examined	Specimens examined. Central Region: Ottotomo Forest Reserve 3 km after reserve base near small loggers road, 3.66 ° N, 11.28 ° E, 02 May 2013, Couvreur T. L. P. 436 (WAG, YA); Ottotomo Forest Reserve 45 km South of Yaounde 5 km on path into reserve, 3.65 ° N, 11.32 ° E, 15 January 2015, Couvreur T. L. P. 667 (WAG, YA); Yaounde, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 73 (P); Route vetere (Likouk-Likoundji), 3.48 ° N, 10.32 ° E, 14 January 1974, Mezili P. 250 (P, WAG, YA). Littoral Region: Ca 40 km NW of Eseka on the other border of Kele River W of Yaounde, 3.82 ° N, 10.48 ° E, 13 December 1963, de Wilde W. J. J. O 1476 (B, BR, MO, P, WAG, YA). South-West Region: Boa, 4.62 ° N, 9.3 ° E, 03 May 1994, Ndam N. 1205 (K); Korup National Park, 4.98 ° N, 8.85 ° E, 01 April 1979, Thomas D. W. 1110 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
474A41299C98CC282707D67EBB3C285D.taxon	description	Figs 90, 91; Map 11 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
474A41299C98CC282707D67EBB3C285D.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2263, Mar 1901: holotype: B [B 100460901]; isotypes: BM [BM 000553960]; G [G 00442259]; GOET [GOET 005683]; HBG [HBG 502522]; K [K 000199004, K 000199005]; P [P 02031265, P 00363277]; WAG [WAG 0065102]; WU [WU 0038180].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
474A41299C98CC282707D67EBB3C285D.taxon	description	Description. Tree, 8 - 10 m tall, d. b. h. 10 - 15 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches densely pubescent. Leaves: petiole 2 - 4 mm long, 2 - 3 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 13 - 21 cm long, 3 - 8 cm wide, obovate, apex acuminate, acumen 0.5 - 1 cm long, base cuneate to obtuse, papyraceous, below pubescent when young and old, above glabrous when young and old, concolorous or discolorous, whitish below; midrib impressed, above glabrous when young and old, below pubescent when young and old; secondary veins 15 to 31 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous and ramiflorous on old leafless branches, axillary, peduncle like base 8 - 20 mm long, axial internodes 5 - 40 mm long, lax or sublax, sympodial rachis 15 - 70 mm long. Flowers with 9 perianth parts in 3 whorls, up to 6 per inflorescence; pedicel 9 - 12 mm long, ca. 2 mm in diameter, pubescent; in fruit ca. 20 mm long, ca. 5 mm in diameter, pubescent; basal bract 5 - 8 mm long, 4 - 5 mm wide; upper bract 5 - 7 mm long, 1 - 2 mm wide; sepals 3, valvate, free, ca. 5 mm long, ca. 4 mm wide, triangular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, ca. 8 mm long, ca. 4 mm wide, ovate, apex acute, base truncate, light yellow to red, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, ca. 35 mm long, ca. 12 mm wide, ovate, apex acute, base truncate, pink, margins wavy, pubescent to pubescent towards base outside, pubescent inside; stamens 70 to 90, in 4 to 5 rows, 1 - 2 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 4 to 6, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 1 to 4, ca. 40 mm long, ca. 28 mm in diameter, ellipsoid, apex rounded, glabrous, verrucose to muricate, not ribbed, white when ripe; seeds 7 to 8 (9) per monocarp, ca. 17 mm long, ca. 11 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
474A41299C98CC282707D67EBB3C285D.taxon	distribution	Distribution. Known from Cameroon and Gabon, and probably present in Equatorial Guinea; in Cameroon known from South, Central and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
474A41299C98CC282707D67EBB3C285D.taxon	materials_examined	Specimens examined. Central Region: Nkolmylon 20 km route Yaounde-Douala, 3.84 ° N, 11.31 ° E, 30 March 1984, Achoundong G. 893 (YA); Colline Nkoldjobe dans le massif Mbaminkom, 3.95 ° N, 11.36 ° E, 15 March 1978, Dang D. 681 (P, YA). Littoral Region: Ebo Forest Reserve Djuma permanent camp on Djashaka trail, 4.35 ° N, 10.23 ° E, 13 February 2014, Couvreur T. L. P. 616 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 649 (WAG, YA); Ebo proposed national park Hospital trail 1430 m from Ndogbayembe trail, 4.14 ° N, 10.18 ° E, 23 March 2006, Mackinnon L. E. 52 (YA); Near Lac Tissongo Mouanko Region, 3.62 ° N, 9.93 ° E, 21 February 1975, McKey D. B. 47 (P, YA); Bekob, 4.61 ° N, 10.70 ° E, 19 February 2006, Tchiengue B. 2555 (K). South Region: 13 km from Kribi S of Lolodorf road, 2.98 ° N, 9.983 ° E, 23 May 1969, Bos J. J. 4625 (BR, MO, P, WAG, YA); 21 km from Kribi Lolodorf road, 3.1 ° N, 10.05 ° E, 16 June 1969, Bos J. J. 4793 (BR, K, LD, LM, MO, P, POZG, WAG, YA); Kribi-Lolodorf, 3.17 ° N, 10.48 ° E, 28 July 1970, Bos J. J. 7163 (BR, P, WAG); Mvini 35 km east of Campo, 2.37 ° N, 10.09 ° E, 27 February 1982, Hoshino J. 359 (YA); Lolodorf, 3.23 ° N, 10.73 ° E, 18 April 1928, Lolo 6 (P); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1914, Zenker G. A. 2079 (WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1918, Zenker G. A. 21 (U); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1900, Zenker G. A. 2263 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 March 1901, Zenker G. A. s. n. (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
20EAEE0E202267367179B8B581C7AD91.taxon	description	Figs 91, 93; Map 11 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
20EAEE0E202267367179B8B581C7AD91.taxon	materials_examined	Type. Gabon. Woleu-Ntem; Oyem, Le Testu G. M. P. C. 9624, 21 Jun 1934: holotype: P [P 00363276]; isotypes: BR [BR 0000008802774, BR 0000008802446]; BM [BM 000553961]; P [P 00363274, P 00363275].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
20EAEE0E202267367179B8B581C7AD91.taxon	description	Description. Tree, 2 - 6 m tall, d. b. h. up to 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 3 - 4 mm long, 1 - 2 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 11 - 27 cm long, 4 - 9 cm wide, elliptic, apex acuminate, acumen 0.2 - 2.5 cm long, base cuneate, papyraceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above glabrous when young and old, below pubescent when young, glabrous when old; secondary veins 21 to 29 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, peduncle like base 7 - 10 mm long, axial internodes 3 - 6 mm long, compact, sympodial rachis 27 - 42 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 11 - 18 mm long, ca. 1 mm in diameter, pubescent; in fruit ca. 12 mm long, ca. 2 mm in diameter, pubescent; basal bract ca. ca. 4 mm long, 2 mm wide; upper bract ca. 4 mm long, ca. 2 mm wide; sepals 3, valvate, free, 5 - 6 mm long, 3 - 4 mm wide, ovate, apex acuminate, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 12 - 14 mm long, 3 - 4 mm wide, obovate, apex acuminate, base truncate, light green, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 40 - 50 mm long, 5 - 10 mm wide, elliptic, apex acute, base truncate, pink, margins wavy, pubescent outside, pubescent inside; stamens numerous, in 6 to 8 rows, ca. 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 3, ovary ca. 2 mm long, stigma globose, pubescent. Monocarps sessile, 1 to 3, 30 - 40 mm long, 25 - 30 mm in diameter, ellipsoid, apex rounded, pubescent when immature, glabrous when mature, verrucose, not ribbed, white with pink when ripe; seeds 3 to 5 per monocarp, 10 - 15 mm long, 5 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
20EAEE0E202267367179B8B581C7AD91.taxon	distribution	Distribution. Known from Cameroon to Gabon; in Cameroon known from South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
20EAEE0E202267367179B8B581C7AD91.taxon	materials_examined	Specimens examined. South Region: Akom II, 2.78 ° N, 10.55 ° E, 12 December 2013, Kamdem N. 159 (YA). South-West Region: Bomana, 4.28 ° N, 9.060 ° E, 05 October 1993, Akogo M. 34 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D600CDEC5E516B8367FB9F80B5D76F6.taxon	description	Figs 92, 94; Map 12 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D600CDEC5E516B8367FB9F80B5D76F6.taxon	materials_examined	Type. Nigeria. Cross River State; Old Calabar, Thomson W. C. 61, no date: holotype: K [K 000199008]; isotype: B [B 100154081].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D600CDEC5E516B8367FB9F80B5D76F6.taxon	description	Description. Tree, up to 12 m tall, d. b. h. 8 - 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 4 mm long, 2 - 3 mm in diameter, densely pubescent, cylindrical, blade inserted on top of the petiole; blade 25 - 41 cm long, 9 - 17.2 cm wide, narrowly elliptic to elliptic or narrowly obovate to obovate, apex acuminate, acumen 1.1 - 2.7 cm long, base rounded to cordate, subcoriaceous, below sparsely pubescent when young and old, above glabrous when young and old, discolorous, whitish below; midrib impressed, above densely pubescent when young and old, below densely pubescent when young, densely pubescent when old; secondary veins 20 to 28 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous or on old leafless branches, axillary, peduncle like base ca. 10 mm long, axial internodes 3 - 23 mm long, compact to sublax, sympodial rachis 35 - 50 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 10 - 25 mm long, ca. 3 mm in diameter, tomentose; in fruit 10 mm long, 3 mm in diameter, tomentose; basal bract ca. 12 mm long, ca. 5 mm wide; upper bract ca. 12 mm long, ca. 5 mm wide; sepals 3, valvate, free, 7 - 8 mm long, 6 - 7 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, 7 - 8 mm long, 3 - 3.5 mm wide, obovate, apex acuminate, base truncate, light yellow cream to red towards the base, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 30 - 60 mm long, 8 - 15 mm wide, elliptic, apex acute, base truncate, pink, margins wavy, pubescent to densely pubescent outside, pubescent inside; stamens numerous, in 6 to 8 rows, 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 5 to 8, ovary ca. 1 mm long, stigma globose, pubescent. Monocarps sessile, 2 to 5, 13 - 30 mm long, 8 - 18 mm in diameter, ellipsoid to ovoid, apex cuspidate, pubescent, finely warty, longitudinally ribbed with ca. 6 ribs, light yellow to orange when ripe; seeds up to 7 per monocarp, 1.5 - 3.5 mm long, 2 - 3 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D600CDEC5E516B8367FB9F80B5D76F6.taxon	distribution	Distribution. A central African species, from south eastern Nigeria to Gabon; in Cameroon known from East, South and mainly South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D600CDEC5E516B8367FB9F80B5D76F6.taxon	materials_examined	Specimens examined. East Region: 78 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.15 ° N, 14.72 ° E, 05 March 2019, Couvreur T. L. P. 1204 (MPU, WAG, YA). South-West Region: Near Bai Kuke SE of Mbonge, 4.53 ° N, 9.11 ° E, 25 January 1958, Keay R. W. J. 37372 (P); Along footpath from Ndian River at PAMOL field 69 and transect P, 5.01 ° N, 8.833 ° E, 24 January 1985, Thomas D. W. 4311 (P, YA); Korup National Park, 5.01 ° N, 8.833 ° E, 12 April 1985, Thomas D. W. 4755 (MO); Limbe W of Njonji Lake, 4.13 ° N, 9.016 ° E, 27 January 1994, Wieringa J. J. 2030 (U, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8DB7A0963AA5DB198DCA1D664323D721.taxon	description	Fig. 92; Map 12 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8DB7A0963AA5DB198DCA1D664323D721.taxon	materials_examined	Type. Cameroon. South-West Region; Rumpi Mountains, between Lokando and Dikome Balue, 30 km NNW Kumba, Letouzey R. G. 14535, 23 Mar 1976: holotype: YA [YA 0002870]; isotype: P [P 02032181].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8DB7A0963AA5DB198DCA1D664323D721.taxon	description	Description. Tree, up to 25 m tall, d. b. h. up to 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches tomentose. Leaves: petiole 2 - 4 mm long, 2 mm in diameter, tomentose, cylindrical, blade inserted on top of the petiole; blade 40 - 49 cm long, 16 - 23 cm wide, obovate, apex acuminate to mucronate, acumen 0.8 - 1.3 cm long, base decurrent to cuneate and narrowly cordate, coriaceous, below densely pubescent when young, densely pubescent when old, above pubescent when young, glabrous when old, discolorous, whitish below; midrib impressed, above pubescent when young and old, below pubescent when young and old; secondary veins 58 to 65 pairs, sparsely pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescence cauliflorous, peduncle like base 10 - 18 mm long, axial internodes 2 - 5 mm long, compact, sympodial rachis 20 - 40 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 2 - 6 mm long, 2 - 3 mm in diameter, tomentose; in fruit ca. 25 mm long, ca. 4 mm in diameter, tomentose; basal bract 7 - 8 mm long, ca. 4 mm wide; upper bract 4 - 6 mm long, ca. 6 mm wide; sepals 3, valvate, free, 5 - 8 mm long, ca. 5 mm wide, ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; petals free, outer petals shorter than inner; outer petals 3, ca. 5 mm long, ca. 4 mm wide, ovate, apex acute, base truncate, light yellow to red, margins flat, densely pubescent outside, glabrous inside; inner petals 3, valvate, 50 - 60 mm long, 5 - 7 mm wide, narrowly elliptic, apex acute, base truncate, margins wavy, densely pubescent outside, glabrous inside; stamens numerous, in 6 to 8 rows, 1 mm long, broad; connective discoid, glabrous, red; staminodes absent; carpels free, 12 to 15, ovary ca. 2 mm long, stigma globose, pubescence not seen. Monocarps sessile, 1 to 2, 20 - 30 mm long, 10 - 25 mm in diameter, obovoid, apex rounded, tomentose, bumpy, brown when ripe; seeds 6 to 8 per monocarp, 6 - 12 mm long, 3 - 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8DB7A0963AA5DB198DCA1D664323D721.taxon	distribution	Distribution. Endemic to Cameroon; known from the South-West and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8DB7A0963AA5DB198DCA1D664323D721.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Forest Reserve Djuma camp Djashaka trail, 4.36 ° N, 10.25 ° E, 15 February 2013, Couvreur T. L. P. 625 (WAG, YA). South-West Region: Edip to Kodmin ca 1 hour's walk, 4.96 ° N, 9.666 ° E, 02 December 1998, Cheek M. 9177 (K, P, WAG, YA); Mount Kupe Kodmin, 4.96 ° N, 9.683 ° E, 21 November 1998, Gosline W. G. 198 (K, P, WAG, YA); Abang road and then right to forest, 4.93 ° N, 9.731 ° E, 11 December 1999, Gosline W. G. 256 (K, MO, WAG, YA); Entre Lokando (900 m) et Dikome Balue (1200 m) 30 km NNW-Kumba, 4.85 ° N, 9.28 ° E, 23 March 1976, Letouzey R. 14535 (P, YA); Rumpi Hills near Madie River, 4.94 ° N, 9.123 ° E, 22 February 1995, Thomas D. W. 10496 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4869CC92D1BAB2D34BDE337E90E51181.taxon	description	Description. Trees, 5 - 12 m tall, d. b. h. up to 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 2 - 8 mm long, 1 - 3 mm in diameter, blade 9 - 34 cm long, 3 - 10 cm wide, oblong to obovate to elliptic, apex acuminate to acute, base cuneate to rounded, discolorous, whitish below; midrib sunken or flat or raised; secondary veins 9 to 17 pairs; tertiary venation intermediate to percurrent. Species unisexual and bisexual, androdioecious, inflorescences ramiflorous on young foliate or old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; male and bisexual flowers similar; pedicel 5 - 10 mm long; in fruit 5 - 15 mm long; bract 1, basal, 2 - 3 mm long; sepals 3, valvate, basally or entirely fused, 1 - 4 mm long, triangular to ovate, apex acute, base truncate; petals free, petals subequal or outer petals longer than inner; outer petals 3, valvate, 10 - 25 mm long, 5 - 9 mm wide, oblong to elliptic to ovate, apex acute, base truncate; inner petals 3, valvate, 5 - 14 mm long, 3 - 7 mm wide, oblong or elliptic to obovate, apex acute, base truncate; stamens 90 to 200 (or numerous), in 7 to 20 rows, 1 - 6 mm long, linear or clavate; connective discoid, glabrous; staminodes absent; carpels free, 2 to 9, ovary 2 - 4 mm long, stigma cylindrical, glabrous or pubescent. Monocarps sessile, 2 to 7, 20 - 110 mm long, 10 - 30 mm in diameter, oblong to obovoid, apex rounded, glabrous, smooth or ribbed; seeds biseriate, 5 - 20 mm long, 3 - 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
063EE49CF928C46B709AEB198C56EACA.taxon	description	Figs 95, 96; Map 12 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
063EE49CF928C46B709AEB198C56EACA.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2899, Mar 1904: holotype: B [B 10 0154089]; isotypes: BR [BR 0000008804808]; K [K 000199038]; M [M 0107917]; P [P 00363351]; S [S 07 - 13391].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
063EE49CF928C46B709AEB198C56EACA.taxon	description	Description. Tree, up to 8 m tall, d. b. h. up to 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 5 - 8 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous, slightly grooved, blade inserted on the side of the petiole; blade 14 - 34 cm long, 3.5 - 8 cm wide, oblong to elliptic, apex acuminate to acute, acumen 0.5 - 1 cm long, base acute, coriaceous, below sparsely pubescent when young, sparsely pubescent to glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 15 pairs, glabrous above; tertiary venation intermediate. Individuals unisexual (?, only male flowers and fruits seen), inflorescences ramiflorous on young foliate or old leafless branches, axillary; flowers with 9 perianth parts in 3 whorls, 2 to 4 per inflorescence; male flowers: pedicel 6 - 8 mm long, 3 - 4 mm in diameter, tomentose; bract 2 - 3 mm long, ca. 1 mm wide; sepals 3, valvate, basally or entirely fused, cup shaped, ca. 4 mm long, 7 - 8 mm wide, triangular to ovate, apex acute, base truncate, tomentose outside, glabrous inside, margins flat; petals free, subequal; outer petals 3, 10 - 14 mm long, 6 - 9 mm wide, oblong to elliptic, apex acute, base truncate, margins flat, densely pubescent outside, pubescent but glabrous towards the base inside; inner petals 3, valvate, 10 - 14 mm long, 5 - 7 mm wide, obovate to oblong, apex acute, base truncate, margins flat, tomentose outside, pubescent inside; stamens 90 to 110, in 7 to 10 rows, 2 - 3 mm long, linear to clavate; connective discoid, glabrous; staminodes absent; female flowers only seen in fruit; fruiting pedicel 10 - 20 mm long, glabrous. Monocarps sessile, 2 to 5, 20 - 30 mm long, 15 - 25 mm in diameter, obovoid, apex rounded, glabrous, smooth, slightly bumpy when dry, not ribbed, white when ripe; seeds 10 to 14 per monocarp, 5 - 10 mm long, 3 - 4 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
063EE49CF928C46B709AEB198C56EACA.taxon	distribution	Distribution. From Cameroon to Gabon, in Cameroon known from the South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
063EE49CF928C46B709AEB198C56EACA.taxon	materials_examined	Specimens examined. South Region: 20 km SE of Kribi route miniere E of Mt Elephant, 2.78 ° N, 10.03 ° E, 01 April 1970, Bos J. J. 6679 (WAG); ca 45 km from Kribi ca 8 km N of Lolodorf road Forest towards Lokoundje R, 3.1 ° N, 10.25 ° E, 02 April 1970, Bos J. J. 6684 (BR, K, LD, LM, MO, P, WAG, YA); Ngovayang, 3.25 ° N, 10.57 ° E, 08 June 2015, Droissart V. 1881 (MO); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 2899 (BR, L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1907, Zenker G. A. 3270 (L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 December 1913, Zenker G. A. 478 (P, U, WAG). South-West Region: Mokoko Forest Reserve Dikome, 4.48 ° N, 9.033 ° E, 05 May 1994, Ekema S. N. 950 (K); Mokoko Forest Reserve ca 6 km W of Mundongo, 4.43 ° N, 9.083 ° E, 22 March 1993, Watts J. 627 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A5850A16E55FCBB7B73BD6C25DA0142E.taxon	description	Figs 95, 96; Map 12 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A5850A16E55FCBB7B73BD6C25DA0142E.taxon	materials_examined	Type. Nigeria. Cross River State; Oban, Talbot P. A. 1607, no date: holotype: K [K 000199039].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A5850A16E55FCBB7B73BD6C25DA0142E.taxon	description	Description. Tree, 5 - 12 m tall, d. b. h. up to 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, glabrous, cylindrical, blade inserted on top of the petiole; blade 9 - 23 cm long, 3 - 10 cm wide, oblong to obovate, apex acuminate, acumen 0.5 - 1 cm long, base acute to rounded, subcoriaceous, below densely pubescent when young, sparsely pubescent when old, above glabrous when young and old, discolorous, whitish below; impressed, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 11 pairs, glabrous above; tertiary venation percurrent. Individuals andromonoecious, male and hermaphrodite inflorescences similar, ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 5 - 6 mm long, 3 - 4 mm in diameter, pubescent; in fruit 5 - 15 mm long, 4 - 7 mm in diameter, glabrous; bract 2 - 3 mm long, 2 - 3 mm wide; sepals 3, valvate, basally or entirely fused, cup shaped, 1 - 2 mm long, 2 - 3 mm wide, ovate, apex acute, base truncate, green to brown-red, tomentose outside, glabrous inside, margins flat; petals free, outer petals longer than inne r; outer petals 3, 10 - 15 mm long, 6 - 8 mm wide, ovate, apex acute, base truncate, white, margins flat, densely pubescent outside, pubescent and glabrous towards base inside; inner petals 3, valvate, 5 - 7 mm long, 3 - 4 mm wide, obovate, apex acute, base truncate, white, margins flat, tomentose outside, glabrous inside; male flowers: stamens 80 to 90, in 6 to 10 rows, 2 - 6 mm long, linear to clavate; connective discoid, glabrous, white; stamens in hermaphrodite flowers: 50 to 60, in 4 to 6 rows, 4 - 6 mm long, connective elongate, glabrous; staminodes absent; carpels free, 2 to 4, ovary 3 - 4 mm long, stigma cylindrical, glabrous. Monocarps sessile, 2 to 4, 35 - 40 mm long, 10 - 15 mm in diameter, oblong, apex rounded, glabrous, irregularly and transversally ribbed, otherwise smooth, light green to white when ripe; seeds 12 to 14 per monocarp, 8 - 10 mm long, 3 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A5850A16E55FCBB7B73BD6C25DA0142E.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West Africa, and central Africa, from eastern Nigeria to Gabon. In Cameroon known from Littoral, South and South West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A5850A16E55FCBB7B73BD6C25DA0142E.taxon	materials_examined	Specimens examined. Littoral Region: At Tissongo 16 km EES of Mouanko, 3.53 ° N, 9.909 ° E, 18 January 1984, Asonganyi J. N. 711 (P, YA); Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1174 (MPU, P, WAG, YA); Ebo Wildlife Reserve Djuma permanent camp On Djuma-Djuma trail, 4.34 ° N, 10.23 ° E, 14 February 2014, Couvreur T. L. P. 623 (WAG, YA); Ebo Wildlife Reserve Djuma permanent camp On transect 5, 4.33 ° N, 10.23 ° E, 16 February 2013, Couvreur T. L. P. 632 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 645 (WAG, YA). South Region: Mvini a Campo, 2.37 ° N, 10.09 ° E, 01 October 1983, Mitani M. 134 (P, YA); ca 7 km NE of Ebom, 3.11 ° N, 10.75 ° E, 01 August 1996, Parren M. P. E. 190 (KRIBI, WAG). South-West Region: 43 km SE of Mamfe Kendem collines, 5.72 ° N, 9.66 ° E, 08 June 1985, Achoundong G. 1332 (YA); Kupe village, 4.78 ° N, 9.716 ° E, 30 May 1996, Cable S. 2721 (K, MO, WAG, YA); Ekundu Kundu, 5.15 ° N, 8.883 ° E, 27 April 1996, Cheek M. 8244 (K, WAG, YA); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.08 ° N, 9.050 ° E, 02 April 2016, Couvreur T. L. P. 1032 (WAG, YA); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.057 ° E, 02 April 2016, Couvreur T. L. P. 1036 (WAG, YA); on trail through palm oil plantation 3 km before lava flow and Seme Beach hotel when coming from Limbe, 4.06 ° N, 9.077 ° E, 18 October 2013, Couvreur T. L. P. 515 (WAG, YA); Parc National de Korup Mt Yuhan entre 700 et 850 m Foret mature de terre ferme transition plaine-submontagnard, 5.15 ° N, 8.860 ° E, 06 March 2012, Droissart V. 1221 (BRLU, MO, YA); Below Kupe rock near Esense river, 4.78 ° N, 9.683 ° E, 25 January 1995, Elad M. 86 (K, YA); Kupe village, 4.77 ° N, 9.688 ° E, 28 November 1999, Gosline W. G. 236 (K); Kupe village, 4.77 ° N, 9.688 ° E, 28 November 1999, Gosline W. G. 237 (K, MO, P, WAG, YA); Pres Kendonge (Reserve forestiere de Bakundu) 13 km SSW Kumba, 4.54 ° N, 9.42 ° E, 20 April 1976, Letouzey R. 14647 (WAG, YA); Korup National Park collected between the Ndian River at Pamol field and 25 km on transect " P " P ", 5.01 ° N, 8.833 ° E, 12 April 1985, Thomas D. W. 4785 (YA); Cameroon Mountain, 4.12 ° N, 9.009 ° E, 20 June 2001, van Andel T. R. 3707 (U, WAG); Korup National Park, 5.01 ° N, 8.806 ° E, 22 March 2004, van der Burgt X. M. 675 (WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ABCF24EAC29B89E0DD0BBE92BD885070.taxon	description	Figs 95, 96; Map 12 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ABCF24EAC29B89E0DD0BBE92BD885070.taxon	materials_examined	Type. Gabon. Ogooue-Lolo; Poungui, Le Testu G. M. P. C. 7754, 10 Dec 1929: lectotype, sheet here designated: P [P 00363348]; isotypes: BR [BR 0000008805041]; P [P 00363347].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ABCF24EAC29B89E0DD0BBE92BD885070.taxon	description	Description. Tree, 5 - 6 m tall, d. b. h. up to 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 5 - 6 mm long, 2 - 3 mm in diameter, pubescent, slightly grooved, blade inserted on top of the petiole; blade 13 - 31 cm long, 3.6 - 5 cm wide, oblong, apex acuminate, acumen 0.5 - 1 cm long, base narrowly cuneate to shortly attenuate, coriaceous, below glabrous when young and old, above glabrous when young and old, (concolorous or) discolorous, whitish below; midrib raised above, above glabrous when young and old, below glabrous when young and old; secondary veins 13 to 17 pairs, glabrous above; tertiary venation intermediate. Individuals andromonoecious, male and hermaphrodite inflorescences similar, ramiflorous on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel ca. 10 mm long, 3 - 4 mm in diameter, pubescent; in fruit 5 - 6 mm long, 5 - 6 mm in diameter, pubescent; bract 2 - 3 mm long, ca. 1 mm wide; sepals 3, valvate, basally or entirely fused, cup shaped, 3 - 4 mm long, ca. 6 mm wide, triangular to ovate, apex acute, base truncate, green to brown-red, densely pubescent outside, glabrous inside, margins flat; petals free, outer petals longer than inner; outer petals 3, 20 - 25 mm long, 5 - 7 mm wide, oblong to elliptic, apex acute, base truncate, yellow, margins flat, pubescent outside, glabrous inside; inner petals 3, valvate, 10 - 14 mm long, 6 - 7 mm wide, elliptic to oblong, apex acute, base truncate, yellow, margins flat, pubescent outside, pubescent inside; stamens 160 to 200, in 8 to 10 rows, 1 - 2 mm long, linear; connective discoid, glabrous, white; staminodes absent; carpels free, 8 to 9, ovary 3 - 4 mm long, stigma cylindrical, pubescent. Monocarps sessile, 5 to 7, 50 - 110 mm long, 20 - 30 mm in diameter, oblong, apex rounded, glabrous, irregularly and transversally ribbed, otherwise smooth, dark green when ripe; seeds 12 to 14 per monocarp, 15 - 20 mm long, ca. 10 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ABCF24EAC29B89E0DD0BBE92BD885070.taxon	distribution	Distribution. From Cameroon to Gabon, and one collection from Democratic Republic of Congo; in Cameroon known from the Littoral and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ABCF24EAC29B89E0DD0BBE92BD885070.taxon	materials_examined	Specimens examined. Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1178 (MPU, WAG). South Region: 20 km From Kribi N of Lolodorf road (SFIA logging road), 3.01 ° N, 10.05 ° E, 15 July 1969, Bos J. J. 5049 (WAG); Ongongondje Hill (Ambam), 2.38 ° N, 11.28 ° E, 27 December 1975, de Wilde J. J. F. E 8718 (BR, K, MO, P, U, WAG, YA); Colline au SE de Ndengue 15 km S d'Ebolowa, 2.78 ° N, 11.12 ° E, 26 March 1970, Letouzey R. 10248 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4974BC6EE633CFE9816BE1329FBE0694.taxon	description	Description. Same as species.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1B4299E652CECE5C2744E9B8E562AC5D.taxon	description	Figs 97, 98; Map 12 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1B4299E652CECE5C2744E9B8E562AC5D.taxon	materials_examined	Type. Gabon. Estuaire; Monts de Cristal, near first bridge after Kinguele, Couvreur T. L. P. 596, 15 Nov 2013: holotype: WAG [WAG 0392343]; isotypes: LBV; P [P 04022675]; YA; WAG [WAG 0392342].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1B4299E652CECE5C2744E9B8E562AC5D.taxon	description	Description. Tree, 4 - 6 m tall, d. b. h. 2 - 4 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent. Leaves: petiole 3 - 4 mm long, 2 - 3 mm in diameter, sparsely pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 20 - 26 cm long, 4.5 - 9 cm wide, ovate to elliptic or obovate, apex long acuminate, acumen 2 - 3 cm long, base obtuse, coriaceous, below glabrous when young and old, above sparsely pubescent to glabrous when young, glabrous when old, concolorous; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 9 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual [possibly androdioecious], inflorescences cauliflorous or ramiflorous on young foliate or old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 2 - 10 mm long, densely pubescent; in fruit 8 - 15 mm long, diameter unknown, glabrous; bracts several basal and one upper towards the lower half of pedicel, bracts ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, 2 - 3 mm long, ca. 2 mm wide, ovate, apex acute, base truncate, pink to light red, densely pubescent outside, glabrous inside, margins flat; petals free, valvate, sub equal; outer petals 3, 4 - 10 mm long, 2.5 - 5 mm wide, elliptic, apex acute, base truncate, red, margins flat, densely pubescent outside, pubescent inside, reflexed; inner petals 3, valvate, 4 - 9 mm long, 2 - 4 mm wide, elliptic, apex acute, base truncate, pink to light red, margins flat, densely pubescent outside, pubescent at least towards margins inside, reflexed; stamens 16 to 19, in 2 to 3 rows, 3 - 4 mm long, linear; connective tongue shaped, glabrous, bright yellow; staminodes absent; carpel 1, ovary 4 - 5 mm long, stigma cylindrical to coiled, sparsely pubescent. Monocarp sessile, 1, 50 - 70 mm long, 15 - 18 mm in diameter, oblong, apex apiculate, sparsely pubescent to glabrous (pubescent when young), smooth, longitudinally 4 to 6 ribbed; seeds not counted, biseriate, 12 - 16 mm long, 6 - 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1B4299E652CECE5C2744E9B8E562AC5D.taxon	distribution	Distribution. Cameroon and Gabon; in Cameroon known from the Littoral region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2A1BC32E9F00FF206DEF5FD5B195AFA0.taxon	description	Description. Same as species. A genus with four species having a disjunct distribution between Africa and South East Asia. Two species are known from Africa, one (S. gracilipes) in Central Africa (Nigeria, Cameroon, Gabon, Equatorial Guinea) and one (S. gracilis) in East Africa (Verdcourt 1971 a). In Cameroon one species, not endemic. The Central African species was initially placed within Friesodielsia (F. gracilipes (Benth.) Steenis) (Le Thomas 1969 b), but a recent molecular phylogeny of Friesodielsia, Monanthotaxis and associated liana genera showed that this species clusters with Sphaerocoryne (Guo et al. 2017 b). Other African species of Friesodielsia have been transferred to Monanthotaxis and Afroguatteria, and thus the genus Friesodielsia does not occur in Africa anymore, being strictly South East Asian.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
68B9EC336E731A5C7B724603D908FAE0.taxon	description	Figs 99, 100; Map 12 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
68B9EC336E731A5C7B724603D908FAE0.taxon	materials_examined	Type. Equatorial Guinea. Bioko, Fernando Po, Mann G. 251, 1860: holotype: K [K 000198951].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
68B9EC336E731A5C7B724603D908FAE0.taxon	description	Description. Liana, 2 - 15 m tall, d. b. h. 1 - 3 cm. Indumentum of simple hairs; old leafless branches glabrous, drying black, young foliate branches sparsely pubescent to glabrous. Leaves: petiole up to 5 mm long, ca. 1 mm in diameter, glabrous, slightly grooved, blade inserted on the side of the petiole; blade 4.5 - 17 cm long, 2 - 6 cm wide, elliptic to ovate, apex acuminate to acute, acumen ca. 1 cm long, base acute (rarely cordate), papyraceous, below glabrous when young and old, above glabrous when young and old, discolorous, whitish below; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 2 to 7 per inflorescence; pedicel 25 - 50 mm long, ca. 1 mm in diameter, glabrous; in fruit 25 - 40 mm long, 2 - 3 mm in diameter, glabrous; bracts 2, one basal and one towards the upper half of pedicel, bracts ca. 1 mm long, ca. 1 mm wide; sepals 3, valvate, free, ca. 2 mm long, ca. 2 mm wide, semiorbicular, apex rounded, base truncate, green, glabrous outside, inside pubescent towards margins but otherwise glabrous, margins flat; petals free, valvate, outer petals larger than inner; outer petals 3, 5 - 15 mm long, 5 - 10 mm wide, elliptic to ovate, apex acute, base truncate, cream to white, margins flat, glabrous outside, inside pubescent towards margins but otherwise glabrous; inner petals 3, valvate, 5 - 7 mm long, 2.5 - 3 mm wide, ovate to oblong, apex acute, base truncate, cream to bright white, margins flat, glabrous outside, glabrous inside; stamens 60 to 80, in 8 to 9 rows, ca. 2 mm long, linear; connective discoid, glabrous, white; staminodes absent; carpels free, 10 to 12, ovary 2 - 3 mm long, stigma filiform, glabrous. Monocarps stipitate, stipes 8 - 10 mm long, 2 - 3 mm in diameter; monocarps 10 to 14, 25 - 30 mm long, 8 - 13 mm in diameter, oblong, apex rounded, glabrous, smooth, sometimes with a prominent rib when dried, red when ripe; seeds 1 to 4 per monocarp, 20 - 25 mm long, 7 - 10 mm in diameter, oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
68B9EC336E731A5C7B724603D908FAE0.taxon	distribution	Distribution. A central African species, from Nigeria to Gabon and Equatorial Guinea; in Cameroon known from the Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
68B9EC336E731A5C7B724603D908FAE0.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On Djashaka trail, 4.35 ° N, 10.24 ° E, 13 February 2014, Couvreur T. L. P. 617 (WAG, YA); Ebo Wildlife Reserve Djuma permanent camp On transect 5, 4.33 ° N, 10.23 ° E, 16 February 2013, Couvreur T. L. P. 633 (WAG, YA). South Region: N bank of Lobe river above Gr Batanga ferry ca. 9 km S of Kribi, 2.87 ° N, 9.893 ° E, 14 October 1968, Bos J. J. 3077 (P, WAG, YA); ca 9 km S of Kribi Lobe R bank E of Gr Batanga ferry, 2.86 ° N, 9.9 ° E, 11 January 1969, Bos J. J. 3602 (P, WAG, YA); Lobe R mouth 7 km S of Kribi, 2.88 ° N, 9.9 ° E, 20 March 1969, Bos J. J. 4180 (BR, K, LD, LM, MO, P, WAG, YA); 10 km From Kribi Lolodorf road, 2.96 ° N, 9.966 ° E, 27 May 1969, Bos J. J. 4655 (BR, P, WAG); Ndoumale 11 km S of Kribi, 2.86 ° N, 9.9 ° E, 29 August 1969, Bos J. J. 5264 (WAG); Campo-Ma'an area Bibabimvoto, 2.28 ° N, 9.950 ° E, 16 August 2002, Tchouto Mbatchou G. P. 3400 (KRIBI, WAG); Campo-Ma'an area 2.28 ° N, 9.866 ° E, 02 October 2001, van Andel T. R. 4095 (KRIBI, U, WAG, YA); Bipindi, 3.08 ° N, 10.41 ° E, 1898, Zenker G. A. 1715 (E, L, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 May 1913, Zenker G. A. 360 (U, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 April 1914, Zenker G. A. 516 (U, WAG). South-West Region: Nyasoso, 4.84 ° N, 9.689 ° E, 02 July 1996, Cable S. 3526 (K, WAG, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 25 March 2016, Couvreur T. L. P. 1001 (WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.090 ° E, 03 April 2016, Couvreur T. L. P. 1043 (WAG, YA); Buea area at Bolifamba, 4.13 ° N, 9.303 ° E, 01 March 1929, Maitland T. D. 536 (K, K, YA); Sousi Forest, 6.36 ° N, 10.61 ° E, 16 February 2006, Onana J. M. 3612 (K); Disturbed forest Bomana and Koto II, 4.3 ° N, 9.05 ° E, 26 April 1996, Tchouto Mbatchou G. P. 1372 (K, YA); Korup National Park, 5.01 ° N, 8.85 ° E, 10 March 1986, Thomas D. W. 5727 (MO); Korup National Park, 5.33 ° N, 8.95 ° E, 25 May 1988, Thomas D. W. 7858 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DF3EB71464F1405C0C43B5CBEAD77CAF.taxon	description	Description. Same as species. A small genus of four species (Luke and Deroin 2005) distributed in central and east Africa, one species occurs in Cameroon, not endemic. Toussaintia is a genus of lianas or scandent shrubs, characterized by the presence of an androgynophore (elongated receptacle on which the stamens and carpels are inserted). All species are rare across their range.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C984DA7367F9163B15B0E1BB5DB103E2.taxon	description	Fig. 101; Map 12 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C984DA7367F9163B15B0E1BB5DB103E2.taxon	materials_examined	Type. Gabon. Ogooue-Ivindo; Belinga, Halle N. 4189, 2 Jul 1966: lectotype, sheet here designated: P [P 00046762]; isotypes: P [P 00046760, P 00046761].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C984DA7367F9163B15B0E1BB5DB103E2.taxon	description	Description. Liana, 10 - 15 m tall, d. b. h. 4 cm. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches tomentose. Leaves: petiole 4 - 7 mm long, ca. 1 mm in diameter, pubescent, grooved, blade inserted on top of the petiole; blade 5 - 14 cm long, 3.5 - 5 cm wide, oblong to elliptic, apex acuminate, acumen ca. 1 cm long, base rounded, subcoriaceous, below pubescent when young and old, above glabrous when young and old, concolorous; midrib impressed, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 7 to 10 pairs, glabrous above; tertiary venation reticulate, very dense and tight. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary. Flowers with 12 to 14 perianth parts in 3 to 4 whorls, 1 to 2 per inflorescence; peduncle short, ca. 2 mm long; pedicel 10 - 17 mm long, 1 - 2 mm in diameter, tomentose; one large bract at the base of the inflorescence, 7 - 13 mm long, 5 - 7 mm wide, elliptic; floral bracts 2 to 4, one basal and several upper towards the lower half of pedicel, basal bract 5 - 9 mm long, 4 - 7 mm wide, elliptic; upper bracts 5 - 8 mm long, 4 - 6 mm wide, elliptic; sepals 3, valvate, free, 10 - 18 mm long, 5 - 7 mm wide, elliptic to ovate, apex acute, base truncate, green, tomentose outside, glabrous inside, margins flat, slightly reflexed in bud; petals free, valvate, sub equal; petals 15 - 30 mm long, 5 - 9 mm wide, narrowly obovate, apex rounded, base truncate, greenish-yellow, streaked with red-brown, base shading into bright white, margins flat, pubescent outside, glabrous inside; staminodes absent; stamens 300 to 350, in 20 to 30 rows arranged on a long conical receptacle (androgynophore), ca. 1 mm long, linear; connective discoid, glabrous, yellow; carpels free, 20 to 22, ovary 4 - 5 mm long, stigma shortly bilobed, glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C984DA7367F9163B15B0E1BB5DB103E2.taxon	distribution	Distribution. A central African species, known from Gabon, Cameroon and Republic of Congo. In Cameroon known from the East region (one collection).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C9872476E2BD0D94438FAE47A32F1526.taxon	description	Description. Lianas, up to 20 (- 30) m tall, d. b. h. up to 20 cm. Indumentum of star, fasciculate and / or simple hairs. Leaves: petiole 2 - 15 mm long, 1 - 2 mm in diameter; blade 5 - 26 cm long, 1 - 3 cm wide, elliptic, ovate, obovate or oblong, apex acuminate to obtuse, base acute to cordate, discolorous, whitish below or concolorous; midrib sunken or flat; secondary veins 6 to 25 pairs; tertiary venation reticulate or percurrent. Individuals bisexual; inflorescences cauliflorous and ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 5 per inflorescence; pedicel 2 - 50 mm long; in fruit 5 - 60 mm long; bracts 2 (or 1), one basal and one upper, 1 - 15 mm long; sepals 3, valvate (or imbricate), free or basally fused or completely fused tearing at anthesis, 2 - 20 mm long, ovate or triangular to semiorbicular, apex acute or acuminate or truncate, base truncate; petals free; outer petals 3, valvate or imbricate, 7 - 35 mm long, 5 - 25 mm wide, ovate to elliptic to obovate to semiorbicular, apex acute to rounded, base truncate; inner petals 3, imbricate, 7 - 35 mm long, 6 - 25 mm wide, ovate to elliptic to obovate to semiorbicular, apex acute to obtuse to rounded, base truncate; stamens 100 to 400, in 5 to 10 rows, 1 - 3 mm long, linear to cuneiform; connective discoid or tongue-shaped, glabrous or pubescent; staminodes absent; carpels free, 15 to 70 (or numerous), 2 - 4 mm long, stigma flat or bilobed or coiled or cylindrical, pubescent or glabrous. Monocarps sessile to stipitate, stipes 5 - 45 mm long; monocarps 8 to 48, 6 - 70 mm long, 6 - 40 mm in diameter, globose or ellipsoid or cylindrical, apex rounded to apiculate, smooth, bumpy, ridged, verrucose or echinate, sometimes strongly ornamented; seeds numerous, bi or uniseriate, 7 - 15 mm long, 4 - 10 mm in diameter, ellipsoid or flattened ellipsoid; aril absent. A diverse genus of ca. 200 species distributed across Africa (west to east), Madagascar and in South East Asia, 17 species occur in Cameroon, one endemic. Uvaria is a genus of lianas or scrambling shrubs, most of which have stellate hairs and numerous seeds per monocarp. The taxonomy of this genus remains complicated and a recent continental revision is still lacking. Differences between the species are mostly based on fruit and leaf characters.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE1FD1F6C315269D222E563F397482C0.taxon	description	Map 12 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE1FD1F6C315269D222E563F397482C0.taxon	materials_examined	Type. Sierra Leone. Southern Province; between Kahreni and Port Lokko, Scott Elliot G. F. 5812, Apr 1891: holotype: K [K 000198777].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE1FD1F6C315269D222E563F397482C0.taxon	description	Description. Liana, ca. 3 m tall, d. b. h. unknown. Indumentum of simple or fasciculate 4 or 3 branched hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches hirsute. Leaves: petiole 3 - 4 mm long, 1 mm in diameter, densely pubescent erect hairs. Leaves: petiole 3 - 4 mm long, 1 mm in diameter, densely hirsute, slightly grooved, blade inserted on top of the petiole; blade 15 - 20 cm long, 4 - 6 cm wide, oblong to ovate, apex acute, base cordate, subcoriaceous, below densely pubescent with fasciculate hairs when young, pubescent when old, above pubescent with simple hairs when young, sparsely pubescent to glabrous when old; midrib sunken or flat, above densely pubescent with simple hairs when young and old, below pubescent with fasciculate hairs when young and old; secondary veins 11 - 15 pairs, sparsely pubescent to glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 35 - 50 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 40 - 55 mm long, 2 mm in diameter, densely pubescent; bracts 2, one basal and one upper towards the lower part of the pedicel, basal bract 3 - 8 mm long, 2 - 3 mm wide; upper bract 3 - 7 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 8 - 10 mm long, 4 - 6 mm wide, ovate, apex obtuse, base truncate, pubescent outside, tomentose inside, margins flat; petals free, sub equal; outer petals 3, 17 - 22 mm long, 13 - 17 mm wide, ovate, apex obtuse, base truncate, yellow, margins flat, pubescent outside, densely pubescent inside; inner petals 3, imbricate, 13 - 20 mm long, 10 - 13 mm wide, ovate, apex obtuse, base attenuate, yellow, margins flat, pubescent outside, tomentose inside, hairs longer towards center; stamens 150 to 200, in 8 to 9 rows, 2 - 3 mm long, linear; connective tongue shaped, pubescent; staminodes absent; carpels free, 40 to 50, ovary 2 - 3 mm long, stigma conical, glabrous. Monocarps stipitate, stipes 35 - 45 mm long, 2 - 3 mm in diameter; monocarps 13 to 25, 13 - 18 mm long, 10 - 15 mm in diameter, club-shaped with the stipes inserted laterally, apex rounded or mucronate, densely pubescent with erect hairs, smooth, constricted around the seeds (2 or 3 bumps), yellow brown to pale orange when ripe; seeds 4 to 6 per monocarp, 12 - 15 mm long, 8 - 10 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
CE1FD1F6C315269D222E563F397482C0.taxon	distribution	Distribution. A mainly west African species, just reaching into Cameroon with a single collection to date, in the North-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
595F7260799A4ED2A3E487BABE67F632.taxon	description	Figs 103, 113; Map 13 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
595F7260799A4ED2A3E487BABE67F632.taxon	materials_examined	Type. Angola. Malanje; Pungo Andongo, Welwitsch F. M. J. 754, Apr 1857: lectotype, sheet here designated: LISU [LISU 206054]; isotypes: B [B 10 0153065]; BM [BM 000554044]; COI [COI 00004859]; K [K 000198826]; P [P 00046766]; LISU [LISU 206055].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
595F7260799A4ED2A3E487BABE67F632.taxon	description	Description. Liana, 2 - 6 m tall, d. b. h. unknown. Indumentum of mixed simple, fasciculate or stellate hairs; old leafless branches sparsely pubescent, young foliate branches densely to sparsely pubescent. Leaves: petiole 3 - 10 mm long, 2 mm in diameter, tomentose to very sparsely pubescent, cylindrical, blade inserted on the side of the petiole; blade 4 - 17 cm long, 2.5 - 7 cm wide, oblong to elliptic, apex acuminate, acumen 1 - 1.5 cm long, base rounded to subcordate, subcoriaceous, below densely to sparsely pubescent when young and old, above sparsely pubescent when young, sparsely pubescent to glabrous when old; midrib sunken or flat, above densely pubescent when young, densely to sparsely pubescent when old, below sparsely pubescent when young, densely to sparsely pubescent when old; secondary veins 6 to 12 pairs, pubescent above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young or old foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 4 - 6 mm long, 1 - 2 mm in diameter, pubescent; in fruit 7 - 10 mm long, 2 mm in diameter, tomentose; bracts 2, one basal and one towards the lower half of pedicel, basal bract 1 - 2 mm long, 1 mm wide; upper bract 2 - 3 mm long, 4 - 5 mm wide; sepals 3, valvate, completely fused, tearing at anthesis, 5 - 9 mm long, 4 - 7 mm wide, ovate, apex truncate, base truncate, green to yellow, tomentose outside, tomentose inside, margins flat; petals free, sub equal; outer petals 3, 12 - 20 mm long, 8 - 11 mm wide, ovate to obovate, apex obtuse, base truncate, green to yellow, margins flat, pubescent outside, pubescent inside; inner petals 3, imbricate, 12 - 19 mm long, 8 - 11 mm wide, obovate to ovate, apex rounded, base truncate, green to yellow, margins flat, pubescent outside, tomentose inside; stamens 190 to 210, in 8 to 9 rows, 3 - 5 mm long, linear; connective tongue shaped, pubescent, yellow to orange; staminodes absent; carpels free, 25 to 35, ovary 4 - 5 mm long, stigma bilobed, slightly capitate, glabrous. Monocarps stipitate, stipes 8 - 12 mm long, 2 - 5 mm in diameter, inserted laterally; monocarps ca. 20, 13 - 40 mm long, 6 - 20 mm in diameter, cylindrical, apex apiculate, tomentose, smooth and slightly constricted around the seeds, brown turning red when ripe; seeds 6 to 16 per monocarp, ca. 10 mm long, 4 - 5 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
595F7260799A4ED2A3E487BABE67F632.taxon	distribution	Distribution. A widespread tropical African species, occurring from Sierra Leone to Central African Republic, Sudan and Ethiopia, and from Kenya to Zambia, one of the few Annonaceae species with such a wide continental distribution (Couvreur et al. 2008 a); in Cameroon known from the Adamaoua, Central, East, North-west, South and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
595F7260799A4ED2A3E487BABE67F632.taxon	materials_examined	Specimens examined. Adamaoua Region: Reserve forestiere de Ngaoundere, 7.32 ° N, 13.58 ° E, 21 July 1977, Fotius G. 2738 (P, YA); Beleldibi (35 km au SSE de Ngaoundere), 7.32 ° N, 13.58 ° E, 22 July 1966, Letouzey R. 7486 (P, YA); Pentes NO de l'Hosere Banyo entre 1100 et 1400 m, 6.75 ° N, 11.82 ° E, 09 June 1967, Letouzey R. 8565 (L, YA). Central Region: Yaounde Mt Eloumden path from foot of mountain (ca 800 m) on Mendong side, 3.81 ° N, 11.43 ° E, 02 May 1996, Cheek M. 8307 (K, WAG, YA); Pentes orientales du mont Yangba (1473 m) pres Nyafianga (42 km NNE de Bafia), 5.13 ° N, 11.35 ° E, 09 September 1966, Letouzey R. 7829 (P, YA); Pres Nyafianga a 46 km SW de Linte, 5.13 ° N, 11.35 ° E, 23 April 1982, Nkongmeneck B. A. 310 (P, YA). East Region: Kumbe, 6 ° N, 14.30 ° E, 01 January 1914, Mildbraed G. W. J. 9224 (COI, K). North Region: Wakwa, 7.23 ° N, 13.58 ° E, 07 October 1960, Breteler F. J. 432 (WAG); Near craterlake ' Lac Tison' ca 12 km SE of Ngaoundere, 7.25 ° N, 13.58 ° E, 30 November 1964, de Wilde W. J. J. O 4361 (WAG). South Region: Rocher de Ako'okas 26 km southeast of Ebolowa, 2.7 ° N, 11.28 ° E, 21 February 1987, Huber H. F. J. 982 (YA). West Region: ca. 6 km NE of Bangangte, 5.15 ° N, 10.52 ° E, 11 May 1964, de Wilde J. J. F. E 2579 (B, BR, K, MO, P, WAG, YA); Mme Vilatte plantations caOca 5 km from Foumbot, 5.55 ° N, 10.61 ° E, 07 July 1972, Leeuwenberg A. J. M. 10160 (BR, K, MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D59822AC6D5A348D709F258312036188.taxon	description	Figs 103, 108; Map 13 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D59822AC6D5A348D709F258312036188.taxon	materials_examined	Type. Gabon. Ogooue-Ivindo; Belinga, Halle N. & Le Thomas A. 484, 14 Aug 1966: holotype: P [P 00362742].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D59822AC6D5A348D709F258312036188.taxon	description	Description. Liana, 5 - 20 m tall, d. b. h. unknown. Indumentum of stellate hairs; old leafless branches sparsely pubescent, young foliate branches tomentose. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, tomentose, grooved, blade inserted on top of the petiole; blade 7 - 12 cm long, 3 - 5 cm wide, ovate, oblong or elliptic, apex acuminate, acumen 1 - 1.5 cm long, base rounded to subcordate, coriaceous, below covered with a continuous persistent layer of small stellate hairs intermingled with larger stellate hairs, above glabrous when young and old; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 18 to 20 pairs, glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches or less often on old leafless branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 10 - 20 mm long, 2 - 3 mm in diameter, tomentose; in fruit unknown; bracts 2, one basal and one towards the lower half of pedicel, basal bract 1 - 2 mm long, 1 mm wide, soon falling; upper bract unknown (soon falling?); sepals 3, valvate, covering in bud, free, 8 - 10 mm long, 8 - 10 mm wide, triangular, apex acute, base truncate, green to yellow, densely pubescent outside, densely pubescent inside, margins reflexed; petals free, sub equal; outer petals 3, 17 - 20 mm long, 10 - 15 mm wide, elliptic to suborbicular, apex obtuse to rounded, base truncate, light yellow, margins flat, pubescent outside, sparsely pubescent inside; inner petals 3, imbricate, 17 - 20 mm long, 10 - 15 mm wide, elliptic, apex obtuse to rounded, base truncate, light yellow, margins flat, tomentose outside, sparsely pubescent inside; stamens 150 to 200, in 8 to 9 rows, 2 - 3 mm long, oblong; connective discoid, sparsely pubescent; staminodes absent; carpels free, 27 to 32, ovary 4 - 5 mm long, stigma coiled, pubescent. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D59822AC6D5A348D709F258312036188.taxon	distribution	Distribution. A central African species, known from Gabon, Cameroon and Democratic Republic of Congo; in Cameroon only known from the South West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D59822AC6D5A348D709F258312036188.taxon	materials_examined	Specimens examined. South-West Region: Monkey forest descending to waterfall at Ndip, 5 ° N, 9.7 ° E, 20 January 1998, Cheek M. 8950 (K, YA); Rumpi mountains forest trail ca 5 km after Dikome Balue village ca 40 km north of Kumba, 4.93 ° N, 9.239 ° E, 10 January 2016, Couvreur T. L. P. 959 (WAG, YA); Kodmin, 5 ° N, 9.666 ° E, 16 November 1998, Gosline W. G. 147 (K, WAG, YA); Above Upper Baondo, 4.06 ° N, 9.15 ° E, 07 April 1992, Tekwe C. F. 49 (K, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
213708699088D35F0C9DBEB79AEF1DE5.taxon	description	Fig. 105; Map 13 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
213708699088D35F0C9DBEB79AEF1DE5.taxon	materials_examined	Type. Nigeria. Cross River State; Oban, Talbot P. A. 1558, 1912: lectotype, sheet here designated: K [K 000198786]; isotypes: K [K 000198786].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
213708699088D35F0C9DBEB79AEF1DE5.taxon	description	Description. Liana, unknown height, d. b. h. unknown. Indumentum of stellate, minute hairs; old leafless branches sparsely pubescent, young foliate branches tomentose. Leaves: petiole 4 - 8 mm long, 1 - 2 mm in diameter, tomentose, grooved, blade inserted on top of the petiole; blade 7 - 25 cm long, 5 - 13 cm wide, obovate to oblong or elliptic, apex acuminate, acumen 1.5 - 2 cm long, base rounded to cordate, coriaceous, below densely pubescent when young, pubescent to sparsely pubescent when old, above densely pubescent quickly becoming glabrous when young, glabrous when old; midrib sunken or flat, above densely pubescent, at least towards the base when young and old, below densely pubescent when young and old; secondary veins 10 to 15 pairs, pubescent to glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches and less often on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 15 - 22 mm long, 1 - 2 mm in diameter, tomentose; in fruit 25 - 30 mm long, 3 - 4 mm in diameter, pubescent; bracts 2, one basal and one towards the lower half of pedicel, basal bract not seen (soon falling?); upper bract 2 - 4 mm long, 3 - 4 mm wide; sepals 3, valvate, fused almost completely, but not tearing, 4 - 5 mm long, 8 - 10 mm wide, suborbicular, apex rounded, base truncate, densely pubescent outside, densely pubescent inside, margins flat; petals free, sub equal; outer petals 3, 17 - 20 mm long, 10 - 15 mm wide, ovate to oblong, apex rounded, base truncate, margins flat, tomentose outside, densely pubescent inside; inner petals 3, imbricate, 17 - 20 mm long, 10 - 15 mm wide, ovate to oblong, apex rounded, base truncate, margins flat, tomentose outside, densely pubescent inside; stamens 150 to 200, in 6 to 7 rows, 1 - 2 mm long, oblong; connective discoid, sparsely pubescent; staminodes absent; carpels free, 20 to 30, ovary 4 - 5 mm long, stigma coiled, densely pubescent. Monocarps stipitate, stipes 20 - 30 mm long, 2 - 3 mm in diameter, centrally inserted; monocarps 8 to 12, 20 - 22 mm long, 20 - 22 mm in diameter, globose, apex rounded, tomentose, verrucose to shortly echinate, not ribbed; seeds not seen.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
213708699088D35F0C9DBEB79AEF1DE5.taxon	distribution	Distribution. A West and Central African species, known from Sierra Leone to Nigeria and Cameroon with a disjunct distribution in eastern Democratic Republic of the Congo; in Cameroon only known from the East and South West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
213708699088D35F0C9DBEB79AEF1DE5.taxon	materials_examined	Specimens examined. East Region: 60 km south of Yokadouma 5 km south of Masea village, 3.10 ° N, 14.84 ° E, 06 March 2019, Couvreur T. L. P. 1208 (MPU, WAG, YA). South-West Region: Munyenge " trouble " trouble " village between liwenyi and Bonja, 4.41 ° N, 9.083 ° E, 20 March 1993, Tchouto Mbatchou G. P. 548 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
32501167ADB547284CADF184FCE4C991.taxon	description	Figs 105, 106; Map 13 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
32501167ADB547284CADF184FCE4C991.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 1116, 1 Jul 1896: holotype: B [B 10 0153072]; isotypes: E [E 00147945]; K [K 000198775]; P [P 00046767]; WU [WU 0025884].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
32501167ADB547284CADF184FCE4C991.taxon	description	Description. Liana, 5 - 20 m tall, d. b. h. unknown. Indumentum of fasciculate hairs; old leafless branches hirsute becoming glabrous, young foliate branches hirsute. Leaves: petiole (2) 4 - 5 mm long, 2 mm in diameter, densely pubescent, slightly grooved, blade inserted on top of the petiole; blade 9 - 27 cm long, 4 - 11 cm wide, obovate to elliptic, apex acuminate, acumen 1.5 - 2 cm long, base cordate to acute or subcordate to rounded, papyraceous, below sparsely hirsute with fasciculate hairs when young and old, above glabrous when young and old; midrib sunken or flat, above densely pubescent with simple hairs when young, pubescent to glabrous when old, below densely pubescent when young, pubescent when old; secondary veins 13 to 20 pairs, pubescent above; tertiary venation percurrent (but some venation network like). Individuals bisexual; inflorescences cauliflorous or occasionally on young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 5 per inflorescence; pedicel 10 - 20 mm long, 1 - 2 mm in diameter, densely pubescent to tomentose; in fruit unknown; bracts 2, one basal and one towards the upper half of pedicel, basal bract not seen (soon falling?); upper bract 2 - 3 mm long, 3 - 5 mm wide; sepals 3, valvate, free, 7 - 8 mm long, 10 - 11 mm wide, suborbicular, apex obtuse, base truncate, brown tomentose outside, densely pubescent inside, margins flat, green-brown; petals free, sub equal; outer petals 3, 16 - 20 mm long, 15 - 16 mm wide, ovate to suborbicular, apex obtuse, base truncate, cream white to cream yellow, margins flat, tomentose outside, densely pubescent towards margins, glabrous towards center inside; inner petals 3, imbricate, 10 - 12 mm long, 10 - 18 mm wide, elliptic to oblong, apex obtuse, base narrowed, cream white to cream yellow, margins flat, tomentose outside, glabrous inside; stamens 150 to 200, in 5 to 7 rows, ca. 4 mm long, linear; connective discoid, glabrous, white; staminodes absent; carpels free, 15 to 25, ovary ca. 4 mm long, stigma coiled, glabrous. Monocarps stipitate, stipes 15 - 20 mm long, 3 - 4 mm in diameter, laterally inserted; monocarps 8 to 13, 20 - 70 mm long, 20 - 40 mm in diameter, cylindrical to oblong, apex rounded, brown tomentose, 4 to 5 ribbed, otherwise smooth, orange-brown when ripe; seeds 5 to 12 per monocarp, ca. 20 mm long, ca. 14 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
32501167ADB547284CADF184FCE4C991.taxon	distribution	Distribution. A central African species, from Cameroon to Gabon and Equatorial Guinea (recently collected in those latter two countries); in Cameroon known from the Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
32501167ADB547284CADF184FCE4C991.taxon	materials_examined	Specimens examined. Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1179 (K, MPU, WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 20 June 2014, Couvreur T. L. P. 662 (WAG, YA); Au SE du Lac Tisongo (35 km SW Edea), 3.53 ° N, 9.909 ° E, 09 January 1974, Letouzey R. 12660 (P, YA). South Region: 20 km from Kribi N of Lolodorf road (SFIA logging road), 3 ° N, 10.05 ° E, 10 June 1969, Bos J. J. 4794 (WAG); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.48 ° N, 10.33 ° E, 11 February 2015, Couvreur T. L. P. 673 (WAG, YA); Campo-Ma'an area Onoyong, 2.52 ° N, 10.69 ° E, 18 March 2001, Tchouto Mbatchou G. P. ONOX _ 53 (WAG); Bipindi, 3.08 ° N, 10.42 ° E, 1896, Zenker G. A. 1116 (K, P). South-West Region: Jocteh Andie-Muen, 5.10 ° N, 9.716 ° E, 09 November 2001, Etuge M. 4504 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D3FABB9B1381741A0C9FD05091B7FD6.taxon	description	Map 13 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D3FABB9B1381741A0C9FD05091B7FD6.taxon	materials_examined	Type. Nigeria. no region; no locality, Palisot de Beauvois A. M. F. J. s. n., no date: holotype: G [G 00014882].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D3FABB9B1381741A0C9FD05091B7FD6.taxon	description	Description. Scrambling shrub to liana, 3 - 10 m tall, d. b. h. 3 - 10 cm. Indumentum of minute stellate hairs; old leafless branches glabrous, young foliate branches very sparsely pubescent to glabrous. Leaves: petiole 4 - 10 mm long, ca. 1 mm in diameter, glabrous, grooved, blade inserted on the side of the petiole; blade 9 - 12 cm long, 4 - 6 cm wide, elliptic, apex acuminate, acumen 0.7 - 0.9 cm long, base acute (obtuse), subcoriaceous, below sparsely pubescent to glabrous when young, glabrous when old, above glabrous or sparsely pubescent when young and old; midrib sunken or flat, above pubescent to glabrous when young and old, below glabrous when young and old; secondary veins 12 to 16 pairs, not prominent above, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on foliate branches, extra axillary or terminal. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 14 - 18 mm long, 1 - 2 mm in diameter, pubescent; in fruit 15 - 19 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, one basal and one towards the lower half of pedicel, basal bract 2 - 4 mm long, 3 - 4 mm wide; upper bract 2 - 4 mm long, 3 - 4 mm wide; sepals 3, valvate, completely fused, tearing at anthesis, 9 - 12 mm long, 9 - 12 mm wide, ovate, apex obtuse or acute, base truncate, light green, densely pubescent outside, sparsely pubescent inside, margins flat; petals free, sub equal, reflexed when opened; outer petals 3, 10 - 17 mm long, 5 - 10 mm wide, ovate, apex rounded, base truncate, green to yellow, margins flat, densely pubescent outside, densely pubescent towards margins, glabrous towards center inside; inner petals 3, imbricate, 9 - 15 mm long, 5 - 10 mm wide, ovate, apex rounded, base unguiculate, green to yellow, margins flat, densely pubescent outside, densely pubescent towards margins and glabrous towards center inside; stamens 150 to 200, in 6 to 7 rows, 2 - 3 mm long, linear; connective discoid, pubescent, yellow to cream; staminodes absent; carpels free, 20 to 30, ovary ca. 4 mm long, stigma coiled, glabrous. Monocarps stipitate, stipes 10 - 15 mm long, 2 - 3 mm in diameter, laterally inserted; monocarps 10 to 20, 22 - 27 mm long, 10 - 13 mm in diameter, cylindrical, apex rounded, pubescent with small stellate hairs, slightly constricted over seeds in dried material, otherwise smooth, brown turning orange when ripe; seeds 12 to 16 per monocarp, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D3FABB9B1381741A0C9FD05091B7FD6.taxon	distribution	Distribution. A mainly west African species from Senegal to Cameroon, Central African Republic, and northern Democratic Republic of the Congo; in Cameroon known from the Central, Far-North, North and North-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3D3FABB9B1381741A0C9FD05091B7FD6.taxon	materials_examined	Specimens examined. East Region: Rives de la Kadei pres de Moundia (15 km SSE de Batouri), 4.15 ° N, 14.61 ° E, 31 March 1962, Letouzey R. 4625 (P, YA). Far-North Region: Ziver, 10.8 ° N, 13.77 ° E, 17 May 1974, Fotius G. 1995 (P, WAG, YA); Mogode-Rhumsiki, 10.5 ° N, 13.58 ° E, 26 August 1976, Geerling C. 5666 (WAG). North Region: Koro (Gore), 10.8 ° N, 13.77 ° E, 19 January 1946, Aubreville A. 628 (P); Mango, 8.42 ° N, 13.25 ° E, 25 July 1974, Fotius G. 2173 (P, YA); Parc National de la Benoue Pres du campement du buffle noir, 8.12 ° N, 13.83 ° E, 05 December 1995, Letouzey R. 11695 (P, YA); Parc national de Faro Campement, 8.36 ° N, 12.81 ° E, 05 March 1985, van der Zon A. P. M. 2542 (WAG); Benoue National Park near Buffle Noir Camp, 8.11 ° N, 13.83 ° E, 04 October 1974, Wit P. 2955 (BR, MO, WAG). North-West Region: Piste Munka (= Munkep) 45 km NNW Wum, 6.73 ° N, 9.95 ° E, 09 July 1975, Letouzey R. 13994 (K, P, YA); Munka (= Munkep) 45 km NNW Wum, 6.73 ° N, 9.95 ° E, 09 July 1975, Letouzey R. 13999 (K, P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1090E2B7BC01E9C271523463BE56BDBB.taxon	description	Figs 103, 109, 110; Map 13 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1090E2B7BC01E9C271523463BE56BDBB.taxon	materials_examined	Type. Gabon. Ogooue-Ivindo; Belinga, Halle N. 3267, 20 Nov 1964: holotype: P [P 00046772].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1090E2B7BC01E9C271523463BE56BDBB.taxon	description	Description. Liana, 3 - 5 m tall, d. b. h. 3 - 5 cm. Indumentum of simple or fasciculate hairs; old leafless branches glabrous, young foliate branches tomentose to pubescent. Leaves: petiole 4 - 6 mm long, 1 - 2 mm in diameter, tomentose, grooved, blade inserted on the side of the petiole; blade 6 - 17 cm long, 3 - 6.5 cm wide, oblong, apex acuminate to obtuse, acumen 1 cm long, base rounded to broadly cordate, papyraceous, below pubescent with simple or fasciculate hairs when young and old, above sparsely pubescent with short simple and stellate hairs when young, glabrous when old; midrib sunken or flat, above sparsely pubescent when young and old, below pubescent when young and old; secondary veins 9 to 13 pairs, sparsely pubescent above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 6 - 11 mm long, 1 - 2 mm in diameter, pubescent; in fruit 11 - 16 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, one basal and one towards the lower half of pedicel, basal bract 2 - 4 mm long, 3 - 4 mm wide; upper bract 2 - 4 mm long, 3 - 4 mm wide; sepals 3, valvate, basally fused, 2 - 3 mm long, 3 - 4 mm wide, ovate, apex acuminate to attenuate, base truncate, pale green, tomentose outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 7 - 14 mm long, 6 - 10 mm wide, ovate to oblong, apex obtuse, base truncate, green to light yellow, margins shortly revolute or straight, tomentose outside, pubescent inside; inner petals 3, imbricate, 7 - 14 mm long, 6 - 10 mm wide, ovate, apex obtuse, base shortly unguiculate, green to light yellow, margins flat, tomentose outside, pubescent inside; stamens numerous, in 7 to 8 rows, ca. 2 mm long, linear; connective discoid, sparsely pubescent, yellow; staminodes absent; carpels free, numerous (not clounted), ovary ca. 4 mm long, stigma bilobed, densely pubescent. Monocarps stipitate, stipes 17 - 22 mm long, 1 - 2 mm in diameter, laterally inserted; monocarps 30 to 40, 10 - 15 mm long, 7 - 9 mm in diameter, ellipsoid to subglobose, apex rounded or very shortly apiculate, sparsely pubescent, slightly constricted over seeds in dried material, otherwise smooth, green to brown when ripe; seeds 10 to 14 per monocarp, 7 - 8 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1090E2B7BC01E9C271523463BE56BDBB.taxon	distribution	Distribution. A central African species, known from Cameroon, Gabon and Democratic Republic of the Congo; in Cameroon known from the Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1090E2B7BC01E9C271523463BE56BDBB.taxon	materials_examined	Specimens examined. Littoral Region: Lac Ossa Nord Dizangue, 3.84 ° N, 10.03 ° E, Achoundong G. 786 (YA); Lac Ossa Nord Dizangue, 3.84 ° N, 10.03 ° E, Achoundong G. 790 (YA). South-West Region: North Base, 4.01 ° N, 9.283 ° E, 24 April 1992, Tchouto Mbatchou G. P. 167 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A486D35946681502A9366BB4738C1E90.taxon	description	Fig. 108; Map 13 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A486D35946681502A9366BB4738C1E90.taxon	materials_examined	Type. Gabon. Woleu-Ntem; Oyem, Le Testu G. M. P. C. 9481, 2 Fev 1934: lectotype, sheet here designated: P [P 00363327]; isotypes: BM [BM 000554061]; BR [BR 0000008823953, BR 0000008823960]; LISC [LISC 000396]; P [P 00363326, P 00363328].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A486D35946681502A9366BB4738C1E90.taxon	description	Description. Liana, 3 - 10 m tall, d. b. h. up to 20 cm. Indumentum of simple to stellate and fasciculate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches densely pubescent, with small stellate hairs intermixed with longer simple hairs. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, tomentose, grooved, blade inserted on top of the petiole; blade 7 - 16 cm long, 3 - 6 cm wide, oblong to elliptic, apex acuminate to acute, acumen 1.5 - 2 cm long, base rounded to subcordate, subcoriaceous, below pubescent with short stellate hairs intermixed with long simple hairs when young, sparsely pubescent with short stellate hairs intermixed with long simple hairs when old, above glabrous when young and old; midrib sunken or flat, above densely pubescent when young and old, below pubescent when young and old; secondary veins 10 to 13 pairs, pubescent above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 10 - 12 mm long, 1 - 2 mm in diameter, pubescent; in fruit 40 - 60 mm long, 3 - 4 mm in diameter, sparsely pubescent; bracts 2, one basal and one towards the lower half of pedicel, similar in size, 5 - 6 mm long, 3 - 4 mm wide; sepals 3, valvate, completely fused, tearing at anthesis, 5 - 10 mm long, 7 - 11 mm wide, triangular to ovate, apex acute, base truncate, yellow, pubescent outside, glabrous inside, margins flat; petals free, inner slightly longer than outer to sub equal; outer petals 3, 15 - 25 mm long, 12 - 20 mm wide, ovate to suborbicular, apex rounded, base truncate, yellow, with dark red-brown base, margins flat or revolute, tomentose outside, glabrous inside; inner petals 3, imbricate, 20 - 35 mm long, 10 - 22 mm wide, ovate to suborbicular, apex rounded, base unguiculate, claw 4 - 5 mm long, yellow with dark red-brown base, margins flat or revolute, tomentose outside, glabrous inside; stamens 300 to 320, in 8 to 10 rows, 2 - 3 mm long, narrowly oblong; connective discoid, pubescent; staminodes absent; carpels free, 40 to 50, ovary ca. 3 mm long, stigma coiled, densely pubescent. Monocarps stipitate, stipes 30 - 45 mm long, ca. 1 mm in diameter, laterally inserted; monocarps 42 to 46, 9 - 12 mm long, 7 - 8 mm in diameter, ellipsoid, apex apiculate or mucronate, tomentose, smooth, not ribbed or very faintly ribbed, green to orange when ripe; seeds 1 to 2 per monocarp, 7 - 8 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A486D35946681502A9366BB4738C1E90.taxon	distribution	Distribution. A central African species, known from Cameroon and Gabon; in Cameroon known from the South West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A486D35946681502A9366BB4738C1E90.taxon	materials_examined	Specimens examined. South-West Region: Kupe village, 4.76 ° N, 9.694 ° E, 21 May 1996, Cable S. 2500 (K, MO, P, WAG, YA); Kupe village, 4.78 ° N, 9.700 ° E, 28 May 1996, Cable S. 2674 (K, YA); Mbule, 4.80 ° N, 9.673 ° E, 26 June 1996, Cable S. 3344 (K, MO, WAG, YA); Nyasoso, 4.86 ° N, 9.7 ° E, 01 July 1996, Cable S. 3459 (B, K, MO, P, WAG, YA); Second trail to summit along road to Mbulle from Nyasoso, 4.81 ° N, 9.681 ° E, 03 July 1996, Cable S. 3558 (K, YA); Kupe village, 4.78 ° N, 9.716 ° E, 15 July 1996, Cable S. 3809 (K, WAG, YA); Nyasoso, 4.83 ° N, 9.691 ° E, 23 October 1995, Cheek M. 7457 (K, YA); Bakossi Mt Kodmin 05 km on road to Muawhojun, 4.98 ° N, 9.683 ° E, 17 January 1998, Cheek M. 8863 (K, WAG, YA); Nyasoso, 4.83 ° N, 9.683 ° E, 26 March 1996, Etuge M. 1819 (K, WAG); Nyasoso, 4.81 ° N, 9.683 ° E, 03 June 1996, Etuge M. 2092 (K, WAG, YA); Kupe village, 4.77 ° N, 9.708 ° E, 08 July 1996, Etuge M. 2659 (K, WAG, YA); Pool at summit of waterfall, 4.96 ° N, 9.716 ° E, 05 February 1998, Gosline W. G. 83 (K, WAG, YA); Kupe village, 4.78 ° N, 9.716 ° E, 11 July 1996, Keay R. W. J. 297 (K, YA); km 2 Essosong Estate-Kola N side of Mount Koupe, 4.85 ° N, 9.733 ° E, 20 January 1972, Leeuwenberg A. J. M. 9249 (B, BR, K, MO, P, WAG, YA); Monts Rumpi entre Dikome Balue (1200 m) et Ifanga Nalende (650 m), 4.9 ° N, 9.233 ° E, 25 March 1976, Letouzey R. 14586 (WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECA5A0E6A15A877CBB5E36C6DA75ECEC.taxon	description	Fig. 104; Map 13 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECA5A0E6A15A877CBB5E36C6DA75ECEC.taxon	materials_examined	Type. Togo. Plateaux; Misahohe, Baumann E. 527, 19 Apr 1895: holotype: B [B 10 0153070].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECA5A0E6A15A877CBB5E36C6DA75ECEC.taxon	description	Description. Scrambling shrub to liana, 5 - m tall, d. b. h. 6 cm. Indumentum of small stellate hairs; old leafless branches pubescent to glabrous, young foliate branches tomentose to densely pubescent. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, pubescent, cylindrical, blade inserted on top of the petiole; blade 4.5 - 19 cm long, 2.5 - 6.5 cm wide, oblong to obovate, apex acuminate, acumen 1 - 1.5 cm long, base rounded to subcordate, papyraceous to subcoriaceous, below covered with a persistent continuous layer of small stellate hairs intermingled with sparser larger stellate hairs, whitish, above sparsely pubescent with stellate hairs when young, sparsely pubescent with stellate hairs to glabrous when old; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 17 to 25 pairs, pubescent above; tertiary venation indistinct or percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches and less often on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 10 - 15 mm long, 1 - 2 mm in diameter, tomentose; in fruit 20 - 25 mm long, 2 - 3 mm in diameter, tomentose; bracts 2, one basal and one towards the lower half of pedicel, soon falling, basal bracts 1 - 2 mm long, 1 mm wide; upper bracts 2 - 3 mm long, 3 - 5 mm wide; sepals 3, valvate, completely fused, covering in bud, tearing at anthesis, 5 - 10 mm long, 8 - 10 mm wide, ovate, apex acuminate, base truncate, pubescent with two layers of stellate hairs outside, pubescent with a single layer of minute stellate hairs inside, margins flat; petals free, sub equal; outer petals 3, 12 - 20 mm long, 9 - 15 mm wide, ovate, apex rounded, base truncate, yellow, margins flat, tomentose outside, tomentose inside; inner petals 3, imbricate, 12 - 20 mm long, 10 - 18 mm wide, ovate, apex rounded, base ungulate, yellow, margins flat, tomentose outside, tomentose inside; stamens 100 to 130, in 5 to 6 rows, 2 - 3 mm long, linear; connective discoid, pubescent; staminodes absent; carpels free, 15 to 20, ovary ca. 6 mm long, stigma conical, pubescent. Monocarps stipitate, stipes 30 - 40 mm long, 2 - 3 mm in diameter, inserted laterally; monocarps 30 to 48, 6 - 12 mm long, 6 - 10 mm in diameter, globose, apex generally shortly apiculate, red-tomentose, warty to verrucose, not ribbed, bluish-green when ripe; seeds 4 to 6 per monocarp, 8 - 10 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECA5A0E6A15A877CBB5E36C6DA75ECEC.taxon	distribution	Distribution. A widely distributed species in West and Central Africa, from Sierra Leone to Benin and from Cameroon to the Democratic Republic of the Congo; in Cameroon known from the East, South and South West (?) regions (see notes).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECA5A0E6A15A877CBB5E36C6DA75ECEC.taxon	materials_examined	Specimens examined. East Region: A 14 km de Ntan (80 km ENE de Lomie), 2.7 ° N, 15.13 ° E, 09 August 1963, Letouzey R. 5584 (P, YA). South Region: Campo-Ma'an area Ipono, 2.33 ° N, 9.841 ° E, 18 April 2002, Elad M. 1535 (KRIBI, WAG); Campo-Ma'an area Mamelles Massif, 2.55 ° N, 9.944 ° E, 23 April 2001, Tchouto Mbatchou G. P. 3238 (KRIBI, WAG); Campo-Ma'an area Bibabimvoto, 2.28 ° N, 9.950 ° E, 16 August 2002, Tchouto Mbatchou G. P. 3404 (KRIBI, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E712B967EBFA0D6C566F90C22EF5707D.taxon	description	Fig. 107; Map 13 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E712B967EBFA0D6C566F90C22EF5707D.taxon	materials_examined	Type. Cameroon. West Region; Balong, Buchholz R. W. 103, 26 Jun 1874: holotype: B [B 10 0154065].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E712B967EBFA0D6C566F90C22EF5707D.taxon	description	Description. Scrambling shrub or liana, height unknown, d. b. h. to 25 cm in diameter. Indumentum of simple or fasciculate hairs; old leafless branches glabrous, young foliate branches pubescent becoming quickly glabrous. Leaves: petiole 5 - 9 mm long, ca. 1 mm in diameter, glabrous, grooved, blade inserted on top of the petiole; blade 10 - 24 cm long, 3.5 - 9 cm wide, obovate to oblong, apex acuminate, acumen ca. 1 cm long, base obtuse to subcordate, papyraceous, below sparsely pubescent when young, sparsely pubescent to glabrous when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 10 to 14 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old leafless branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 20 - 25 mm long, 1 - 2 mm in diameter, pubescent; in fruit 25 - 27 mm long, 2 - 3 mm in diameter, glabrous; bracts 2, one basal and one upper towards the upper half of pedicel, basal bract 1 - 2 mm long, 1 mm wide; upper bract 4 - 6 mm long, 3 - 5 mm wide; sepals 3, imbricate, free, 6 - 8 mm long, 6 - 8 mm wide, suborbicular, apex rounded, base truncate, pubescent outside, glabrous inside, margins flat; petals free, inner slight longer than outer; outer petals 3, 10 - 15 mm long, 7 - 10 mm wide, ovate to oblong, apex acute, base truncate, margins revolute, pubescent outside, pubescent inside; inner petals 3, imbricate, 20 - 22 mm long, 14 - 17 mm wide, obovate, apex rounded, base ungulate, margins flat, pubescent outside, glabrous inside; stamens numerous, in 10 rows, 1 - 2 mm long, narrowly oblong; connective discoid, glabrous; staminodes absent; carpels free, numerous, ovary ca. 3 mm long, stigma obpyramidal, glabrous. Monocarps stipitate, stipes 20 - 30 mm long, laterally inserted, 1 - 2 mm in diameter; monocarps 30 to 40, 12 - 15 mm long, 10 - 15 mm in diameter, ellipsoid, apex apiculate or rounded, pubescent, smooth, irregularly ribbed with two main prominent ribs, greyish green turning orange when ripe; seed 1 (more rarely 2) per monocarp, 8 - 10 mm long, 5 - 6 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E712B967EBFA0D6C566F90C22EF5707D.taxon	distribution	Distribution. A central African species, from Cameroon to Gabon; in Cameroon known from the East, South, South West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E712B967EBFA0D6C566F90C22EF5707D.taxon	materials_examined	Specimens examined. East Region: Rocher Ekok Edanbawa a 110 km au SW de Yokadouma 2.81 ° N, 14.48 ° E, 29 March 1973, Letouzey R. 12190 (P, WAG, YA). South Region: Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1913, Zenker G. A. 4926 (L, P); Bipindi, 3.08 ° N, 10.41 ° E, 01 April 1914, Zenker G. A. 597 (U, WAG). South-West Region: Limbe Mt Etinde above Mpanja, 4.08 ° N, 9.174 ° E, 11 July 1990, Cheek M. 3014 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EF261ED84EC2C49BAEB99BD21B50529B.taxon	description	Figs 110, 111; Map 14 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EF261ED84EC2C49BAEB99BD21B50529B.taxon	materials_examined	Type. Gabon. Estuaire; Libreville, Klaine T. - J. 235, 14 Sep 1896: lectotype, sheet here designated: P [P 00362663]; isotypes: B [B 10 0153078]; P [P 00362707, P 00362706, P 00362710, P 00362712, P 00362714, P 00362709, P 00362708, P 00362662].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EF261ED84EC2C49BAEB99BD21B50529B.taxon	description	Description. Scrambling shrub to liana, 3 - 20 (30) m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of stellate hairs; old leafless branches glabrous, young foliate branches tomentose. Leaves: petiole 2 - 5 mm long, 1 - 2 mm in diameter, tomentose covered with minute stellate hairs, grooved, blade inserted on top of the petiole; blade 2 - 9 cm long, 1 - 3 cm wide, oblong to elliptic, apex obtuse, base rounded to obtuse, subcoriaceous, below densely pubescent with short persistent stellate hairs covering the surface, above sparsely pubescent when young, glabrous when old; midrib sunken or flat, above densely pubescent with fasciculate hairs when young, pubescent when old, below densely pubescent when young and old; secondary veins 7 to 12 pairs, pubescent above; tertiary venation reticulate, indistinct. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 5 - 8 mm long, 1 - 2 mm in diameter, tomentose; in fruit 6 - 8 mm long, 2 - 3 mm in diameter, pubescent; bracts 2, one basal and one towards the upper half of pedicel, basal bract 1 - 2 mm long, 1 mm wide; upper bract 4 - 5 mm long, 3 - 4 mm wide; sepals 3, valvate, basally fused, ca. 4 mm long, 5 - 6 mm wide, ovate, apex obtuse, base truncate, light brown, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 15 - 17.5 mm long, 10 - 12 mm wide, ovate to oblong, apex rounded, base truncate, green to light yellow, margins flat becoming revolute, pubescent outside, pubescent to glabrous inside; inner petals 3, imbricate, 15 - 16 mm long, 10 - 12 mm wide, ovate to oblong, apex obtuse, base truncate, green to light yellow, margins flat becoming revolute, pubescent outside, sparsely pubescent to glabrous inside; stamens 130 to 150, in 6 to 7 rows, 2 - 3 mm long, linear; connective discoid, glabrous, yellow to cream; staminodes absent; carpels free, ca. 100, ovary ca. 2 mm long, stigma flat, densely pubescent. Monocarps stipitate, stipes 25 - 40 mm long, 1 - 2 mm in diameter, centrally inserted; monocarps 30 to 50, 12 - 18 mm long, 8 - 15 mm in diameter, globose to ovoid, apex rounded, densely tomentose, verrucose, faintly-one ribbed, green when immature; seeds 8 to 10 per monocarp, 7 - 9 mm long, 4 - 5 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EF261ED84EC2C49BAEB99BD21B50529B.taxon	distribution	Distribution. A central African species, known mainly from Gabon, but also from Cameroon, Equatorial Guinea and Republic of the Congo; in Cameroon known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E326174A51885DF069BFFB41C6C771D6.taxon	description	Fig. 105; Map 14 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E326174A51885DF069BFFB41C6C771D6.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. & Staudt A. 3, no date: lectotype, here designated: K [K 000198767].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E326174A51885DF069BFFB41C6C771D6.taxon	description	Description. Liana (?), unknown height, d. b. h. unknown. Indumentum of fasciculate hairs; old leafless branches sparsely pubescent to glabrous, young foliate branches tomentose. Leaves: petiole 2 - 5 mm long, 1 - 2 mm in diameter, tomentose, slightly grooved, blade inserted on top of the petiole; blade 13 - 16 cm long, 5 - 7 cm wide, oblong to elliptic, apex acuminate, acumen 1.5 - 2 cm long, base subcordate, subcoriaceous, below pubescent with fasciculate hairs when young and old, above glabrous when young and old; midrib sunken or flat, above densely pubescent when young, pubescent when old, below densely pubescent when young or old; secondary veins 14 to 17 pairs, prominent and pubescent above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young branches, leaf-opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 30 - 40 mm long, ca. 2 mm in diameter, tomentose; bracts 2, one basal and one in the upper half of pedicel, basal bract 1 - 2 mm long, 1 - 2 mm wide; upper bract 2 - 3 mm long, 4 - 7 mm wide; sepals 3, valvate, free, 4 - 7 mm long, 5 - 6 mm wide, ovate, apex acute, base truncate, pubescent outside, inside not seen, margins flat; petals free, inner slightly smaller than outer; outer petals 3, 20 - 23 mm long, 11 - 14 mm wide, elliptic, apex rounded, base truncate, margins revolute, pubescent outside, inside not seen; inner petals 3, imbricate, 14 - 16 mm long, 16 - 17 mm wide, ovate to suborbicular, apex rounded, base ungulate, claw ca. 5 mm long, margins flat or revolute, pubescent outside, glabrous inside; stamens 300 to 400, in 6 to 7 rows, 1 - 2 mm long, linear; connective discoid, pubescent; staminodes absent; carpels free, number unknown, ovary ca. 3 mm long, stigma coiled, densely pubescent. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E326174A51885DF069BFFB41C6C771D6.taxon	distribution	Distribution. endemic to Cameroon, known from the Central and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E326174A51885DF069BFFB41C6C771D6.taxon	materials_examined	Specimens examined. Central Region: Yaounde, 3.87 ° N, 11.52 ° E, 1890, Zenker G. A. 249 (B); Yaounde, 3.87 ° N, 11.52 ° E, Zenker G. A. 3 (K); Yaounde, 3.06 ° N, 10.38 ° E, 01 January 1913, Zenker G. A. 475 (NY). East Region: Bitya near R Ja Nsolo, 3.02 ° N, 12.37 ° E, 31 March 1921, Bates G. L. 1818 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3445C79273915110CB2282704AF02DB0.taxon	materials_examined	Type. Gabon. Estuaire; Environ de Libreville, Klaine T. - J. 550, 1896: holotype: B [B 10 0153099]; isotypes: P [P 00362733, P 00362731]. Only var. Uvaria muricata yalingensis is known from Cameroon to date.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA0FE252A0F43CF495B755D0BC0CC398.taxon	description	Figs 112, 113; Map 14 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA0FE252A0F43CF495B755D0BC0CC398.taxon	materials_examined	Type. Central African Republic: Haute-Kotto; Yalinga, Le Testu G. M. P. C. 4241, 25 Oct 1922: holotype: P [P 00362713]; isotype: BM [BM 000554051].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA0FE252A0F43CF495B755D0BC0CC398.taxon	description	Description. Scrambling shrub to liana, 3 - 10 m tall, d. b. h. 6 cm. Indumentum of stellate hairs; old leafless branches glabrous, young foliate branches pubescent to sparsely pubescent. Leaves: petiole 5 - 15 mm long, 1 - 2 mm in diameter, pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 12 - 19 cm long, 2.5 - 9.5 cm wide, elliptic to oblong, apex acuminate, acumen 1 - 1.5 cm long, base cordate, papyraceous to subcoriaceous, below sparsely pubescent when young, glabrous when old, above sparsely pubescent to glabrous when young and old; midrib sunken or flat, above densely pubescent when young, pubescent when old, below pubescent when young and old; secondary veins 12 to 17 pairs, glabrous above; tertiary venation indistinct, slightly percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 10 - 22 mm long, 1 - 2 mm in diameter, pubescent to sparsely pubescent; in fruit 20 - 40 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, one basal and one in lower half of pedicel, basal bract 2 - 4 mm long, 3 - 4 mm wide; upper bract not seen; sepals 3, valvate, distinct in the flower bud, basally fused, 3 - 5 mm long, 4 - 7 mm wide, triangular to ovate to semiorbicular, apex obtuse, base truncate, tomentose with short stellate hairs outside, tomentose with short stellate hairs inside, margins flat; petals free, sub equal; outer petals 3, 12 - 15 mm long, 4 - 7 mm wide, ovate to suborbicular, apex rounded, base truncate, margins flat, tomentose outside, glabrous inside; inner petals 3, imbricate, 12 - 15 mm long, 4 - 7 mm wide, ovate to suborbicular, apex rounded, base ungulate, margins flat, tomentose outside, glabrous inside; stamens numerous, number of rows not seen, 2 - 3 mm long, oblong to cuneiform; connective discoid, sparsely pubescent; staminodes absent; carpels free, 20 to 25, ovary 3 - 4 mm long, stigma obconical, pubescent. Monocarps stipitate, stipes 10 - 15 mm long, 1 - 2 mm in diameter, centrally inserted; monocarps 20 to 25, 10 - 18 mm long, 10 - 18 mm in diameter, globose, apex rounded, pubescent with stellate hairs, finely echinate, not ribbed, green to brown when ripe; seeds 4 to 5 per monocarp, ca. 10 mm long, 5 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA0FE252A0F43CF495B755D0BC0CC398.taxon	distribution	Distribution. In its broad sense Uvaria muricata is a widespread mainly central African species (a few specimens collected from Sierra Leone) known from Cameroon to the Republic of the Congo, the Democratic Republic of the Congo and the Central African Republic; the variety Uvaria yalingensis is known from Central African Republic and Cameroon; in Cameroon known from Adamaoua, Central, North, South, South West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA0FE252A0F43CF495B755D0BC0CC398.taxon	materials_examined	Specimens examined. Adamaoua Region: ca 15 km NE of Meiganga, 6.63 ° N, 14.25 ° E, 24 November 1964, de Wilde W. J. J. O 4049 (WAG); Pres Dota (route Meiganga-Baibokoum a 85 km a vol d'oiseau de Meiganga), 6.97 ° N, 14.93 ° E, 14 October 1963, Letouzey R. 6167 (P, YA). Central Region: Eloumden, 3.83 ° N, 11.46 ° E, 20 October 1961, Breteler F. J. 1977 (WAG); Ca 50 km S of Badjob ca 60 km SW Of Eseka Along the Njong-River, 3.68 ° N, 10.68 ° E, 20 March 1964, de Wilde W. J. J. O 2164 (BR, K, MO, P, WAG, YA). South Region: 24 km from Kribi ca 3 km N of Lolodorf road, 3.03 ° N, 10.08 ° E, 31 March 1970, Bos J. J. 6652 (BR, P, WAG, YA). South-West Region: Ndile waterfall below mission dispensary, 4.91 ° N, 9.745 ° E, 16 December 1999, Cheek M. 10415 (K, MO, P, WAG, YA); Mont versant de Munyenge, 4.37 ° N, 9.256 ° E, 21 January 1985, Nkongmeneck B. A. 954 (YA). West Region: Massif du Mbapit (1988 m) 30 km SW Foumban Versant Sud, 5.5 ° N, 10.63 ° E, 21 October 1974, Letouzey R. 12936 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9EF158E79EF791ED636DF2FA00A5C092.taxon	description	Fig. 113; Map 14 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9EF158E79EF791ED636DF2FA00A5C092.taxon	materials_examined	Type. Type. Nigeria. Cross River State; Oban, Talbot P. A. 1603, 1912: lectotype, sheet here designated: K [K 000198779]; isotypes: BM [BM 000554066]; K [K 000198780, K 000198781].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9EF158E79EF791ED636DF2FA00A5C092.taxon	description	Description. Liana, 3 - 10 m tall, d. b. h. unknown. Indumentum of simple or fasciculate hairs, but overall glabrous; old leafless branches glabrous, young foliate branches sparsely pubescent becoming quickly glabrous. Leaves: petiole 4 - 5 mm long, 2 mm in diameter, glabrous, slightly grooved, blade inserted on top of the petiole; blade 15 - 26 cm long, 5 - 10 cm wide, oblong, apex acuminate to acute, acumen 1.5 - 2 cm long, base rounded to subcordate, coriaceous, below very sparsely pubescent to glabrous when young, glabrous when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 7 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls; 1 to 3 (9) per inflorescence; pedicel 6 - 8 mm long, 2 - 3 mm in diameter, sparsely pubescent to glabrous; bracts 2, one basal and one towards the lower half of pedicel, basal bract 2 - 4 mm long, 3 - 4 mm wide; upper bract 2 - 3 mm long, 3 - 5 mm wide; sepals 3, valvate, free, 4 - 7 mm long, 6 - 8 mm wide, ovate to semiorbicular, apex obtuse, base truncate, green, tomentose outside, pubescent inside, margins flat; petals free, sub equal; outer petals 3, 20 - 30 mm long, 17 - 25 mm wide, ovate to suborbicular, apex rounded, base truncate, green to light yellow, margins flat, tomentose outside, glabrous inside; inner petals 3, imbricate, 20 - 30 mm long, 17 - 25 mm wide, ovate to suborbicular, apex rounded, base unguiculate, green to light yellow, margins flat, tomentose outside, glabrous inside; stamens 120 to 150, in 8 to 10 rows, 2 - 3 mm long, oblong to cuneiform; connective discoid, glabrous; staminodes absent; carpels free, 15 to 20, ovary ca. 3 mm long, stigma coiled, glabrous. Monocarps sessile, monocarps possibly 5 to ca. 8 (but only one seen in McPherson 15524 (LBV )), ca. 40 mm long, ca. 25 mm in diameter, glabrescent to glabrous, ellipsoid, apex rounded, smooth, green when ripe; seeds not seen.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9EF158E79EF791ED636DF2FA00A5C092.taxon	distribution	Distribution. Known from Nigeria to Cameroon, and more recently from Gabon (see below); in Cameroon known from South and South West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9EF158E79EF791ED636DF2FA00A5C092.taxon	materials_examined	Specimens examined. South Region: 9 km N of Kribi, 3 ° N, 9.933 ° E, 30 October 1969, Bos J. J. 5555 (BR, C, K, LD, P, WAG, YA); Ebom, 3.1 ° N, 10.73 ° E, 24 February 1997, Elad M. 564 (KRIBI, WAG). South-West Region: Ekombe-Mofako Mokoko Forest Reserve, 4.47 ° N, 9.092 ° E, 23 April 1994, Acworth J. M. 182 (K, YA); Kumba, 4.63 ° N, 9.433 ° E, 01 December 1908, Busgen M. 191 (B, K); West bank of the Onge River, 4.28 ° N, 8.966 ° E, 07 November 1993, Thomas D. W. 9802 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2C16A6E4FDE12321752D3271ADBFE2A7.taxon	description	Map 14 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2C16A6E4FDE12321752D3271ADBFE2A7.taxon	materials_examined	Type. Cameroon. East Region; Molundu, Mildbraed G. W. J. 4715, 1911: lectotype, sheet here designated: B [B 10 0153106]; isotypes: B [B 10 0153105]; HBG [HBG 502489].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2C16A6E4FDE12321752D3271ADBFE2A7.taxon	description	Description. Liana, 4 - 6 m tall, d. b. h. unknown. Indumentum of stellate or fasciculate hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, densely pubescent to tomentose, slightly grooved, blade inserted on top of the petiole; blade 4.5 - 12 cm long, 2 - 5 cm wide, elliptic to oblong, apex acuminate to acute, acumen 0.9 - 1.5 cm long, base rounded to cordate, papyraceous to subcoriaceous, discolorous, below completely covered with short stipitate stellate hairs when young and old, above pubescent with very short scabrid fasciculate hairs; midrib sunken or flat, above densely to sparsely pubescent when young, sparsely pubescent to glabrous when old, below densely pubescent when young and old; secondary veins 9 to 12 pairs, sparsely pubescent to glabrous above; tertiary venation reticulate, but indistinct. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 10 - 30 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 20 - 30 mm long, 2 - 3 mm in diameter, pubescent; bracts 2, one basal and one towards the lower half of pedicel, basal bract 2 mm long, 2 mm wide; upper bract 2 - 3 mm long, 3 - 5 mm wide; sepals 3, valvate, completely fused, tearing at anthesis, 12 - 15 mm long, 8 - 10 mm wide, ovate, apex acute to acuminate, base truncate, pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 25 - 35 mm long, 10 - 18 mm wide, oblong-obovate to oblong, apex acute, base truncate, green to light yellow, margins flat, densely pubescent outside, pubescent inside; inner petals 3, imbricate, 25 - 35 mm long, 10 - 18 mm wide, obovate to ovate, apex obtuse, base truncate, green to light yellow, margins flat, densely pubescent outside, pubescent inside; stamens 120 to 150, in 6 to 7 rows, 1 - 2 mm long, elongated; connective discoid, sparsely pubescent; staminodes absent; carpels free, 30 to 45, ovary 3 - 4 mm long, stigma coiled, glabrous. Monocarps stipitate, stipes 10 - 15 mm long, 1 - 2 mm in diameter, inserted laterally; monocarps 15 to 20, 15 - 30 mm long, 5 - 12 mm in diameter, cylindrical, apex apiculate, brown-tomentose, verrucose, slightly constricted between the seeds in dried material, brown when ripe; seeds 4 to 8 per monocarp, 7 mm long, 4 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2C16A6E4FDE12321752D3271ADBFE2A7.taxon	distribution	Distribution. A central African species, known from Cameroon, Gabon, Democratic Republic of the Congo and Angola; in Cameroon known from the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
2C16A6E4FDE12321752D3271ADBFE2A7.taxon	materials_examined	Specimens examined. East Region: Prairies 10 km a l'E de Kinsassa village situe a 65 km au NNE de Moloundou par route Yokadouma 2.61 ° N, 15.47 ° E, 04 March 1971, Letouzey R. 10494 (P, YA); Prairie a 30 km WSW de Kinsasa village situe a 65 km NNE de Moloundou sur la route de Yokadouma 2.34 ° N, 15.28 ° E, 10 March 1971, Letouzey R. 10556 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B85F0D1F7D6313C932369933FEAEF390.taxon	description	Map 14 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B85F0D1F7D6313C932369933FEAEF390.taxon	materials_examined	Type. Democratic Republic of the Congo. Kasai Central; Mukenge, Pogge P. 622, 18 Sep 1882: lectotype, designated by Le Thomas (1969 b), p. 73: B [B 10 0153109]; isolectotype: K [K 000198768].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B85F0D1F7D6313C932369933FEAEF390.taxon	description	Description. Liana, 15 - 18 m tall, d. b. h. 3 - 10 cm. Indumentum of minute stellate or simple hairs; old leafless branches sparsely pubescent, young foliate branches tomentose. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, densely pubescent to tomentose, slightly grooved, blade inserted on top of the petiole; blade 8 - 19 cm long, 5 - 9 cm wide, obovate to elliptic, apex acuminate, acumen ca. 1 cm long, base rounded to cordate, papyraceous to membranaceous, below rather densely pubescent with stipitate stellate hairs when young and old, above sparsely pubescent with simple hairs (soft to touch) when young and old; midrib sunken or flat, above sparsely pubescent when young and old, below densely pubescent when young and old; secondary veins 15 to 24 pairs, sparsely pubescent to glabrous above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches, extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 15 - 20 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 25 - 35 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract 2 mm long, 2 mm wide; upper bract 2 - 3 mm long, 3 - 5 mm wide; sepals 3, valvate, free, not covering in bud, 4 - 7 mm long, 5 - 8 mm wide, triangular to ovate, apex obtuse, base truncate, tomentose outside, pubescent inside, margins reflexed; petals free, sub equal; outer petals 3, 15 - 20 mm long, 12 - 15 mm wide, ovate, apex attenuate, base truncate, margins flat, tomentose outside, sparsely pubescent to glabrous inside; inner petals 3, imbricate, 15 - 20 mm long, 12 - 15 mm wide, ovate, apex attenuate, base truncate, margins flat, tomentose outside, sparsely pubescent to glabrous inside; stamens 150 to 200, in 8 to 10 rows, ca. 2 mm long, linear; connective discoid, sparsely pubescent; staminodes absent; carpels free, 15 to 20, ovary 3 - 4 mm long, stigma coiled, pubescent. Monocarps stipitate, stipes 25 - 35 mm long, 1 - 2 mm in diameter, centrally inserted; monocarps 30 to 35, 7 - 15 mm long, 7 - 15 mm in diameter, globose, apex apiculate, densely to sparsely pubescent, smooth, not ribbed, green when ripe; seeds not counted, ca. 7 mm long, 4 - 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B85F0D1F7D6313C932369933FEAEF390.taxon	distribution	Distribution. A central African species, known from Cameroon, Democratic Republic of the Congo, Gabon, Republic of the Congo, Central African Republic and Angola; in Cameroon known from the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B85F0D1F7D6313C932369933FEAEF390.taxon	materials_examined	Specimens examined. East Region: Lobeke Reserve Terra firma forest 1 km east of Djangi Bai, 2.31 ° N, 15.76 ° E, 04 November 1998, Harris D. J. 6200 (E); Lobeke Reserve 3 - 4 km north of Djembe road 5 km northeast of Bolu Bai, 2.21 ° N, 15.78 ° E, 24 November 1998, Harris D. J. 6528 (E).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
27BD8EF754CD5BC89D93097B882C31D6.taxon	description	Figs 114, 116; Map 14 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
27BD8EF754CD5BC89D93097B882C31D6.taxon	materials_examined	Type. Cameroon. Littoral Region; Cameroons River, Mann G. 1433, 1862: lectotype, here designated: K [K 000105336].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
27BD8EF754CD5BC89D93097B882C31D6.taxon	description	Description. Liana, 2 - 20 m tall, d. b. h. unknown. Indumentum of minute stellate or simple hairs; old leafless branches pubescent to glabrescent, young foliate branches densely pubescent. Leaves: petiole 2 - 3 mm long, ca. 1 mm in diameter, pubescent, grooved, blade inserted on top of the petiole; blade 6 - 23 cm long, 2.5 - 10 cm wide, elliptic to oblong, apex acuminate to obtuse, acumen 1 - 1.2 cm long, base rounded to subcordate, papyraceous to coriaceous, below densely pubescent covered with minute stipitate stellate hairs when young and old, above pubescent with short simple scabrid hairs (or glabrous besides veins) when young and old; midrib sunken or flat, above densely pubescent when young and old, below densely pubescent when young and old; secondary veins 9 to 16 pairs, densely pubescent above; tertiary venation percurrent. Individuals bisexual; inflorescences ramiflorous on young foliate branches, leaf opposed or extra axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 10 - 25 mm long, 1 - 2 mm in diameter, densely pubescent to tomentose; in fruit 20 - 40 mm long, 6 mm in diameter, pubescent; bracts 2, soon falling, one basal and one towards the upper half of pedicel, basal bract 10 - 15 mm long, 7 - 10 mm wide; upper bract not seen; sepals 3, valvate, basally fused, 15 - 20 mm long, 7 - 13 mm wide, ovate, apex acute to obtuse, base truncate, yellow brown, densely pubescent outside, pubescent inside, margins flat; petals free, sub equal; outer petals 3, 18 - 30 mm long, 13 - 20 mm wide, elliptic to ovate, apex attenuate, base truncate, yellow-brown, margins flat, densely pubescent outside, pubescent inside; inner petals 3, imbricate, 15 - 25 mm long, 12 - 19 mm wide, ovate, apex obtuse, base truncate, yellow brown, margins flat, pubescent outside, sparsely pubescent to glabrous inside; stamens 180 to 200, in 9 to 10 rows, ovary 1 - 2 mm long, linear; connective discoid, glabrous, cream to yellow; staminodes absent; carpels free, 45 to 70, ca. 3 mm long, stigma coiled, pubescent. Monocarps sessile, 20 to 35 tightly packed together, 15 - 20 mm long, 10 - 13 mm in diameter, pyramidal or four-sided, apex rounded to shortly apiculate, tomentose, verrucose to shortly echinate, not ribbed or 1 - ribbed, green when ripe; seeds 4 to 6 per monocarp, 8 - 12 mm long, 4 - 6 mm in diameter, flattened; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
27BD8EF754CD5BC89D93097B882C31D6.taxon	distribution	Distribution. A widespread species in west and central Africa, in Liberia and Nigeria and in Cameroon, Gabon and Equatorial Guinea to the Democratic Republic of the Congo and Angola; in Cameroon known in the Central, East, Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
27BD8EF754CD5BC89D93097B882C31D6.taxon	materials_examined	Specimens examined. Central Region: Village Nkolbisson 7 km West of Yaounde, 3.88 ° N, 11.45 ° E, 04 May 1962, Breteler F. J. 2880 (U, WAG); Nkolbison, 3.88 ° N, 11.45 ° E, 10 November 1963, de Wilde W. J. J. O 1199 (WAG); Etuk Ebe (Yaounde), 3.87 ° N, 11.52 ° E, 04 May 1971, Mpom B. 544 (P). East Region: Ngoko (Moloundou), 1.67 ° N, 16.04 ° E, 04 April 1971, Letouzey R. 10599 (P, YA). Littoral Region: Tonde (Douala), 4.22 ° N, 9.84 ° E, 09 May 1976, Letouzey R. 14884 (P, WAG, YA). South Region: 65 km S of Kribi Gr Batanga road, 2.89 ° N, 9.905 ° E, 28 October 1968, Bos J. J. 3164 (WAG, YA); 75 km from Kribi Ebolowa road, 2.89 ° N, 9.957 ° E, 13 May 1969, Bos J. J. 4518 (WAG, YA); 15 km from Kribi Ebolowa road Bidou II, 2.85 ° N, 10 ° E, 04 July 1969, Bos J. J. 5011 (BR, P, WAG, YA); 13 km from Kribi Ebolowa road, 2.85 ° N, 10.9 ° E, 16 October 1969, Bos J. J. 5508 (BR, C, K, LD, P, WAG, YA); Ndengue (Ebolowa), 2.78 ° N, 11.12 ° E, 26 March 1970, Letouzey R. 10251 (P, WAG, YA); Sangmeli 2.93 ° N, 11.98 ° E, 26 March 1981, Meijer D. 15270 (MO, WAG); 6 km W de Nyabessan, 2.4 ° N, 10.4 ° E, 26 November 1982, Nkongmeneck B. A. 348 (P, YA); Campo-Ma'an area Medjivini, 2.29 ° N, 10.34 ° E, 31 March 2000, Tchouto Mbatchou G. P. 2721 (KRIBI, WAG, YA); Efoulan, 2.74 ° N, 10.54 ° E, 24 April 2000, Tchouto Mbatchou G. P. 2819 (KRIBI, WAG, YA). South-West Region: on top of hill near Small Ekombe village 3 km after Kumba on road to Ekondo Titi town, 4.62 ° N, 9.376 ° E, 13 January 2016, Couvreur T. L. P. 982 (WAG, YA); Manja village Mundemba, 4.98 ° N, 8.916 ° E, 06 May 1996, Etuge M. 1888 (K).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C01C53D5F1214225CCA12235F884AD95.taxon	description	Description. Trees or shrubs, 4 - 30 m tall, d. b. h. up to 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs. Leaves: petiole 1 - 4 mm long, 1 - 2 mm in diameter; blade 6 - 22 cm long, 2 - 5 cm wide, elliptic or obovate, apex acuminate, acumen 0.7 - 2 cm long, base subcordate to decurrent to rounded to cuneate, concolorous; midrib sunken or flat; secondary veins 7 to 17 pairs, arching well before the margin; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous, or ramiflorous young foliate branches and axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 8 - 50 mm long; in fruit 10 - 50 mm long; bracts 1 to 3, all basal; sepals 3, reduplicate-valvate, free, 10 - 25 mm long, apex acute, base truncate; petals free, sub equal; outer petals 3, valvate, 20 - 40 mm long, 5 - 15 mm wide, ovate to elliptic, apex acute, base truncate or narrowed; inner petals 3, valvate, 10 - 28 mm long, 6 - 15 mm wide, ovate or elliptic, apex acute, base cuneate; stamens numerous (not counted), 2 - 6 mm long, linear; connective elongated to discoid; staminodes absent; carpels free, 5 to 16, 2 - 6 mm long, stigma bilobed or capitate. Monocarps stipitate or sessile, stipes 1 - 6 mm long; monocarps 2 to 8, 30 - 100 mm long, 10 - 50 mm in diameter, globose or ellipsoid, apex apiculate or acute to rounded; seeds 16 to 27, 10 - 25 mm long, 7 - 15 mm in diameter, flattened ellipsoid; aril absent. A genus of five species from West and Central Africa; one species adapted to the drier regions in southern Central Africa (U. hexaloboides (R. E. Fr.) R. E. Fr.); in Cameroon three species, none endemic.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A94E86F9ADD65DE8F6D362F07079FEF0.taxon	description	Fig. 117; Map 14 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A94E86F9ADD65DE8F6D362F07079FEF0.taxon	materials_examined	Type. Cameroon. South-West Region; Johann-Albrechtshohe [Kumba], Staudt A. 740, 1896: lectotype, designated by Couvreur (2014), p. 18: B [B 100153111]; isolectotypes: COI [COI 00077205]; G [G 00011729]; K [K 000105338, K 000105339, K 000105340]; [P 00315828, P 00315829]; S [S 12 - 22768].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A94E86F9ADD65DE8F6D362F07079FEF0.taxon	description	Description. Tree to shrub, 4 - 15 m tall, d. b. h. 3 - 5 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 1 - 4 mm long, 1 mm in diameter, densely pubescent, weakly grooved adaxially, blade inserted on top of the petiole; blade 6 - 14 cm long, 2 - 4 cm wide, elliptic to obovate, apex rounded or acuminate, acumen 1 - 2 cm long, base subcordate, papyraceous to subcoriaceous, below glabrous when young and old, above sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, concolorous; midrib sunken or flat, above pubescent when young, glabrous when old, below sparsely pubescent to densely pubescent when young, sparsely pubescent to densely pubescent when old; secondary veins 8 to 12 pairs, sparsely pubescent to glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous, ramiflorous on young foliate or on old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 8 - 15 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 10 - 20 mm long, 2 - 5 mm in diameter, densely pubescent; bracts 3, all basal, 4 mm long, 3 - 4 mm wide; sepals 3, reduplicate-valvate, free, 10 - 20 mm long, 5 - 8 mm wide, ovate, apex acute, base truncate, green to light green or pale yellow with darker margins, densely pubescent outside, densely pubescent inside, margins slightly revolute; petals free, outer petals longer than inner; outer petals 3, 23 - 35 mm long, 5 - 10 mm wide, ovate, apex acute, base narrowed, light yellow to white, margins flat, pubescent outside, pubescent (sometimes sparsely) pubescent inside; inner petals 3, valvate, 10 - 20 mm long, 6 - 10 mm wide, ovate, apex acute, base cuneate, light yellow, margins flat, densely pubescent outside, sparsely pubescent inside; stamens numerous, 2 - 3 mm long, linear; connective elongated, densely pubescent; staminodes absent; carpels free, 6 to 7, ovary 2 - 3 mm long, stigma bilobed, slightly capitate, glabrous. Monocarps shortly stipitate, stipes 2 - 6 mm long, 2 - 5 mm in diameter; monocarps 2 to 8, 30 - 60 mm long, 10 - 20 mm in diameter, oblong, apex apiculate, densely pubescent, finely warty, longitudinally 4 to 6 ribbed, resembling that of a peanut, light brown when ripe; seeds ca. 20 per monocarp, 10 - 15 mm long, 7 - 9 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A94E86F9ADD65DE8F6D362F07079FEF0.taxon	distribution	Distribution. A mainly West African species, but with a disjunct distribution in West and Central Africa, from Liberia to Cote d'Ivoire, and from Nigeria to Gabon. In Cameroon known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A94E86F9ADD65DE8F6D362F07079FEF0.taxon	materials_examined	Specimens examined. South-West Region: Likomba-Pflanzung 15 - 35 km NE von Victoria [Limbe], 4.1 ° N, 9.333 ° E, 01 December 1928, Mildbraed G. W. J. 10795 (A, K); Johann-Albrechtshohe [Kumba] area 4.63 ° N, 9.416 ° E, 1896, Staudt A. 740 (B, COI, G, K, P, P, S); Johann-Albrechtshohe [Kumba] area 4.63 ° N, 9.416 ° E, 1897, Staudt A. 900 (G).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0481DA67850EE0A3BA782FA66090B048.taxon	description	Figs 117, 118; Map 14 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0481DA67850EE0A3BA782FA66090B048.taxon	materials_examined	Type. Gabon. Estuaire; Libreville, Klaine T. - J. 1091, Oct 1897: lectotype, designated by Le Thomas (1969 b), p. 294: P [P 00315822]; isolectotype: B [B 1001153112].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0481DA67850EE0A3BA782FA66090B048.taxon	description	Description. Tree to shrub, 4 - 25 m tall, d. b. h. 40 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent. Leaves: petiole 2 - 4 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous, weakly grooved adaxially, blade inserted on the side; blade 6 - 16 cm long, 2 - 4.5 cm wide, elliptic to obovate, apex acuminate, acumen 0.7 - 2 cm long, base decurrent to cuneate, papyraceous to subcoriaceous, below sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, above sparsely pubescent to glabrous when young, sparsely pubescent to glabrous when old, concolorous; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 7 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on young foliate or old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 15 - 50 mm long, 1 - 2 mm in diameter, sparsely pubescent to densely pubescent; in fruit 15 - 50 mm long, 4 - 6 mm in diameter, sparsely pubescent to glabrous; bract 1, basal, 6 mm long, 6 mm wide; sepals 3, reduplicate-valvate, free, 15 - 25 mm long, 10 - 20 mm wide, ovate, apex acute, base truncate, grey-green, pubescent towards margins outside, sparsely pubescent inside, margins revolute; petals free, sub equal; outer petals 3, 25 - 40 mm long, 10 - 15 mm wide, elliptic, apex acute, base narrowed, yellow to greyish yellow, margins flat, densely pubescent outside, sparsely pubescent inside; inner petals 3, valvate, 15 - 28 mm long, 6 - 15 mm wide, elliptic, apex acute, base cuneate, yellow to greyish yellow, margins flat, densely pubescent outside, sparsely pubescent inside; stamens numerous, 4 - 6 mm long, linear; connective elongated, pubescent, pinkish red; staminodes absent; carpels free, 5 to 10, ovary 4 - 6 mm long, stigma bilobed, slightly capitate, densely pubescent. Monocarps shortly stipitate to sessile, stipes 0 - 4 mm long, 3 - 5 mm in diameter; monocarps 3 to 5, 90 - 100 mm long, 40 - 50 mm in diameter, ellipsoid to globose, apex rounded, pubescent, velvety, smooth, not ribbed, light green when ripe; seed 16 to 20 per monocarp, 15 - 25 mm long, 1 - 2 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0481DA67850EE0A3BA782FA66090B048.taxon	distribution	Distribution. A widespread species with a disjunct distribution in west and central Africa, from Guinea and Liberia to Ghana, and from Nigeria to Republic of Congo, and northern Democratic Republic of Congo. In Cameroon known from the East, South, Central and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0481DA67850EE0A3BA782FA66090B048.taxon	materials_examined	Specimens examined. Central Region: Mefou National Park, 3.61 ° N, 11.58 ° E, 13 March 2004, Cheek M. 86 (K, YA). East Region: 3 km west of Djembe road head Lobeke Reserve, 2.21 ° N, 16 ° E, 16 October 1998, Harris D. J. 5889 (E); Lobeke Reserve Small Bai, 2.28 ° N, 15.71 ° E, 01 November 1998, Harris D. J. 6135 (E); A 25 km a l'WSW de Kinsassa village situe a 65 km au NNE de Moloundou sur route de Yokadouma 2.56 ° N, 15.16 ° E, 09 March 1971, Letouzey R. 10533 (HGB, P, YA); Foret au Sud de Dimako rive droite de la riviere Mbonda, 4.26 ° N, 13.56 ° E, 18 January 1960, Letouzey R. 2670 (P, YA); A 13 km SSW de Koso (village situe a 60 km au SSW de Batouri), 3.93 ° N, 14.17 ° E, 29 July 1963, Letouzey R. 5529 (P, YA); Approximately 10 km N of Welele between Yokadouma and Molundu, 2.68 ° N, 15.36 ° E, 18 March 1987, Manning D. 1586 (MO, P, YA); Reserve de Biosphere du Dja vers 1175 m sur la piste reliant la station de Bouamir et l'inselberg de Mbasakok, 3.18 ° N, 12.81 ° E, 18 May 2001, Senterre B. 1283 (BR); Reserve de Biosphere du Dja vers 1175 m sur la piste reliant la station de Bouamir et l'inselberg de Mbasakok, 3.18 ° N, 12.81 ° E, 19 May 2001, Senterre B. 1370 (BR). South Region: Bitye, 3.01 ° N, 12.35 ° E, 01 September 1922, Bates G. L. 1764 (K); Campo Ma an National Park 5 km after main entrance, 2.35 ° N, 10.25 ° E, 15 February 2012, Couvreur T. L. P. 385 (WAG, YA); 1 km from main camp near Lele river 16 km East from Lele village, 2.28 ° N, 13.32 ° E, 07 September 2013, Couvreur T. L. P. 454 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 680 (WAG, YA); Ca 30 km S of Mbalmayo, 3.29 ° N, 11.44 ° E, 13 February 1964, de Wilde J. J. F. E 1904 (B, BR, K, MO, P, WAG, YA); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.81 ° N, 11.13 ° E, 18 February 1975, de Wilde J. J. F. E 7972 (B, BR, K, MO, P, U, WAG, YA); Meyo Centre, 2.55 ° N, 11.03 ° E, 24 March 1970, Letouzey R. 10225 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
62AB21AC9CC4A355BE9D0FB74BC4D12D.taxon	description	Figs 118, 119; Map 15 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
62AB21AC9CC4A355BE9D0FB74BC4D12D.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2935, 1904: lectotype, designated by Couvreur (2014), p. 28: B [B 100153114]; isolectotypes: B [B 100190283]; BM, [BM 000554069]; BR [BR 0000014035722]; COI [COI 00077201]; GOET [GOET 005731]; G [G 00011742, G 00011744]; K [K 000198808]; L [L 0191076]; M [M 0089220]; MA [MA 215566 - 3]; P [P 00315826]; S [S 12 - 22789]; WAG [WAG 0057973]; WU [WU 0025789]; Z [Z 000034578, Z 000034577].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
62AB21AC9CC4A355BE9D0FB74BC4D12D.taxon	description	Description. Tree to shrub, 20 - 30 m tall, d. b. h. 20 - 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 2 - 3 mm long, ca. 2 mm in diameter, sparsely pubescent, not grooved, blade inserted on top of the petiole; blade 12 - 22 cm long, 3 - 5 cm wide, elliptic to obovate, apex acuminate, acumen 1 - 2 cm long, base cuneate to rounded, coriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, concolorous; midrib sunken or flat, above pubescent when young, glabrous when old, below sparsely pubescent to densely pubescent when young, sparsely pubescent to densely pubescent when old; secondary veins 11 to 17 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on young foliate or old leafless branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 13 - 30 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous; in fruit 15 - 30 mm long, 2 - 5 mm in diameter, glabrous; bracts 3, all basal, 5 mm long, 3 - 4 mm wide; sepals 3, reduplicate-valvate, free, 15 - 25 mm long, 8 - 15 mm wide, ovate, apex acute, base truncate, light brown, sparsely pubescent to glabrous outside, pubescent and glabrous towards center inside, margins recurved; petals free, sub equal; outer petals 3, 20 - 35 mm long, 8 - 15 mm wide, elliptic to ovate, apex acute, base truncate, yellow, margins flat, densely pubescent outside, pubescent except towards center inside; inner petals 3, valvate, 15 - 25 mm long, 7 - 10 mm wide, ovate, apex acute, base cuneate, claw mm long, white turning bright yellow to orange, light grey when old, margins flat, densely pubescent outside, pubescent except towards center inside; stamens numerous, 2 - 4 mm long, linear; connective discoid, pubescent, red; staminodes absent; carpels free, (1 - 3) 5 to 15, ovary 3 - 5 mm long, stigma capitate, sparsely pubescent to glabrous. Monocarps shortly stipitate, stipes 2 - 5 mm long, ca. 5 mm in diameter; monocarps 2 to 5, 80 mm long, 25 mm in diameter, oblong, apex acute, sparsely pubescent to glabrous, longitudinally 4 to 6 ribbed, sometimes more finely ribbed, light green-grey with dashed of white when ripe; seeds 20 to 27 per monocarp, 15 - 20 mm long, 10 - 15 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
62AB21AC9CC4A355BE9D0FB74BC4D12D.taxon	distribution	Distribution. A widespread species in Central Africa, from extreme southeast of Nigeria to the Republic of Congo, and Democratic Republic of Congo. In Cameroon known from the East, South, Central, South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
62AB21AC9CC4A355BE9D0FB74BC4D12D.taxon	materials_examined	Specimens examined. East Region: Dja reserve, 3.20 ° N, 12.79 ° E, 04 November 1994, Fogiel M. K. 1039 (MO, P). Littoral Region: Ebo Wildlife Reserve Djuma permanent camp, 4.34 ° N, 10.23 ° E, 14 February 2014, Couvreur T. L. P. 624 (WAG, YA); Douala (Route Razel), 4.05 ° N, 9.7 ° E, 01 January 1955, Endengle E. s. n. (P, YA). South Region: 41 kmN of Kribi 5 km E of Edea road forest track Fifinda-Bella old secondary forest, 3.21 ° N, 10.06 ° E, 06 February 1970, Bos J. J. 6266 (BR, C, K, LD, P, WAG, YA); Entre 15 et 25 km au SW de Zingui (soit a 45 km au SSE de Kribi), 2.82 ° N, 10.97 ° E, 22 March 1968, Letouzey R. 9121 (P, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1902, Zenker G. A. 2438 (B, COI, G, K, L, MO, P, S, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 2935 (B, BM, COI, G, K, L, M, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1906, Zenker G. A. 3248 (K); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1907, Zenker G. A. 3289 (B, L, P, S); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1908, Zenker G. A. 3409 (COI, G, L, M, MO, P, S); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1912, Zenker G. A. 4473 (B, COI, G, L, M, P, S); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1914, Zenker G. A. 481 (B, BR, C, F, G, GH, M, P, S, U, WAG). South-West Region: Mabeta Moliwe, 3.96 ° N, 9.233 ° E, 02 April 1992, Bongyu J. 42 (K, P); Korup National Park, 5.06 ° N, 8.855 ° E, 12 January 1998, Kenfack D. 1008 (MO, P, WAG); Boa, 4.4 ° N, 9 ° E, 04 May 1994, Ndam N. 1248 (K); Mokoko Forest Reserve, 4.46 ° N, 9.066 ° E, 19 April 1994, Tchouto Mbatchou G. P. 1108 (K); Ndian River at PAMOL field 69 and transect P, 5.01 ° N, 8.833 ° E, 24 January 1985, Thomas D. W. 4334 (K, MO, P, YA); Korup National Park, 4.98 ° N, 8.85 ° E, 12 January 1979, Thomas D. W. 604 (K); Korup National Park, 5.05 ° N, 8.766 ° E, 10 June 1988, Thomas D. W. 8094 (L, P, WAG, YA); West bank of the Onge River, 4.28 ° N, 8.966 ° E, 07 November 1993, Thomas D. W. 9772 (K, MO, P, WAG, YA); Korup National Park, 5 ° N, 8.8 ° E, 04 February 2000, van der Burgt X. M. 590 (G, WAG, YA); Korup National Park, 5 ° N, 8.8 ° E, 19 March 2004, van der Burgt X. M. 674 (BR, G, K, MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
AE1559319C911694913B8FE6F29A7ADB.taxon	description	Description. Trees, 3 - 20 m tall, d. b. h. up to 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs or absent. Leaves: petiole 4 - 20 mm long, 1 - 9 mm in diameter; blade 13 - 70 cm long, 3.6 - 15 cm wide, ovate or elliptic or obovate or oblong, apex acuminate to emarginate, acumen 0.7 - 2 cm long, base acute to rounded; midrib sunken or flat; secondary veins 11 to 36 pairs per side; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary, 1 to 3 per inflorescence; pedicel 1 - 19 mm long; in fruit 6 - 38 mm long; bracts 2 to 6, several basal and one upper towards the middle of the pedicel; sepals 3, valvate, basally fused, 6 - 35 mm long, apex acuminate, base truncate; petals free, sub equal; outer petals 3, valvate, 12 - 33 mm long, 8 - 25 mm wide, ovate, apex acuminate, base truncate; inner petals 3, valvate, 6.5 - 26.5 mm long, 5 - 17 mm wide, ovate, apex acute, base truncate; stamens 200 to 5000, 2 - 5 mm long, linear to narrowly oblong; connective discoid; staminodes absent; carpels free, 6 to 104, 3 - 7 mm long, stigma bilobed. Monocarps sessile or shortly stipitate; monocarps (1) 5 to 63, 20 - 70 mm long, 12 - 30 mm in diameter, globose or ellipsoid or oblong, apex acute or rounded; seeds 6 - 29 mm long, 4 - 21 mm in diameter, ellipsoid to flattened ellipsoid or oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
555E7885FD3734199128920C1C3A02E3.taxon	description	Fig. 120; Map 15 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
555E7885FD3734199128920C1C3A02E3.taxon	materials_examined	Type. Cameroon. South-West Region; Johann-Albrechtshohe [Kumba], Staudt A. 742 a, 20 Mar 1896; holotype: B [B 10 0153115].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
555E7885FD3734199128920C1C3A02E3.taxon	description	Description. Tree to shrub, 3 - 12 m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to pubescent. Leaves: petiole 3 - 7.5 mm long, 1 - 2 mm in diameter, glabrous to pubescent, grooved, blade inserted on top of the petiole; blade 10 - 19.9 cm long, 3 - 5.8 cm wide, narrowly elliptic, apex acute to acuminate, acumen 1.1 - 1.4 cm long, base acute to cuneate, subcoriaceous, below glabrous to pubescent at base, glabrous when old, above glabrous when young and old, lemon-scented when crushed; midrib sunken or flat, above glabrous when young and old, below glabrous to pubescent at base when young and old; secondary veins 8 to 14 pairs per side, tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 0 - 6 mm long, 2 - 3 mm in diameter, densely pubescent; in fruit ca. 5 mm long, ca. 3.5 mm in diameter; bracts 2 to 6, several basal and one upper towards the middle of pedicel, basal bracts 2 - 3 mm long, 1 - 3 mm wide; upper bract 6 - 11 mm long, 9 - 15 mm wide; sepals 3, valvate, basally fused, imbricate at the middle, 9 - 13 mm long, 9 - 13 mm wide, ovate, apex acuminate, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 15 - 21 mm long, 9 - 15 mm wide, ovate, apex acuminate, base truncate, margins flat, densely pubescent outside, glabrous inside; inner petals 3, valvate, 15 - 22 mm long, 5 - 10 mm wide, ovate, apex acuminate, base truncate, margins flat, densely pubescent outside, glabrous inside; stamens 200 to 300, in 10 to 15 rows, 3 - 3.5 mm long, narrowly oblong; connective discoid; staminodes absent; carpels free, 7 to 30, ovary 3.5 - 4.5 mm long, stigma bilobed, slightly capitate, densely pubescent. Monocarps shortly stipitate, stipes 1 - 3 mm long, ca. 3 mm in diameter; monocarps 2 to 10, 23 - 40 mm long, 17 - 30 mm in diameter, ellipsoid to globose, yellow when ripe emitting strong lemon smell; seeds 9 to 18 per monocarp, 21 - 28 mm long, ca. 10 mm in diameter, ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
555E7885FD3734199128920C1C3A02E3.taxon	distribution	Distribution. A species with a disjunct distribution in West Africa (Ghana, Ivory Coast) and Central Africa (Nigeria, Cameroon); in Cameroon, known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
555E7885FD3734199128920C1C3A02E3.taxon	materials_examined	Specimens examined. South-West Region: Between Bafia and Likoko, 4.37 ° N, 9.324 ° E, 05 February 1958, Keay R. W. J. 37524 (K); Kumba, 4.63 ° N, 9.416 ° E, 1896, Staudt A. 642 (K); Johann-Albrechtshohe [Kumba] area 4.16 ° N, 9.2 ° E, 28 March 1896, Staudt A. 742 (B, K); Lake Barombi Kumba, 4.64 ° N, 9.45 ° E, 01 April 1986, Thomas D. W. 6087 (YA); Along the road between Konye and Bakole, 4.91 ° N, 9.466 ° E, 25 May 1987, Thomas D. W. 7018 (YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
436ED951CE56B644979EFA0FDB0810DA.taxon	description	Fig. 121; Map 15 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
436ED951CE56B644979EFA0FDB0810DA.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 2344, 1901: holotype: B [B 10 0153116]; isotypes: BM [BM 000636669]; G [G 00412241]; GOET [GOET 005732]; HBG [HBG 502513]; K [K 000198797, K 000198796]; M [M 0107940]; P [P 00362661, P 00362658, P 00362659]; S [S 07 - 13396, S 07 - 13393]; WAG [WAG. 1418666].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
436ED951CE56B644979EFA0FDB0810DA.taxon	description	Description. Tree, 2 - 20 m tall, d. b. h. 20 - 35 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches sparsely pubescent, young foliate branches densely pubescent to brown tomentose. Leaves: petiole 4 - 25 mm long, 2 - 9 mm in diameter, densely pubescent to brown tomentose, slightly grooved, blade inserted on top of the petiole; blade 25.8 - 76.5 cm long, 6.1 - 24.8 cm wide, oblong to obovate, apex acuminate to emarginate, acumen 0.6 - 3.2 cm long, base rounded, coriaceous, below glabrous to pubescent at base when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below densely pubescent when young and old; secondary veins 19 to 40 pairs per side, glabrous above, slightly pubescent to pubescent below; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 0 - 9 mm long, 4 - 7 mm in diameter, tomentose; in fruit 1 - 9 mm long, ca. 4 - 7 mm in diameter, tomentose; bracts 2 - 6, several basal and one upper towards the middle of pedicel, basal bracts 2 - 3 mm long, 1 - 3 mm wide; upper bract 10 - 23 mm long, 10 - 40 mm wide; sepals 3, valvate or imbricate, basally fused, 10 - 27 mm long, 10 - 26 mm wide, ovate, apex acuminate, base truncate, brown, densely pubescent to brown tomentose outside, pubescent except towards center inside, margins flat; petals free, sub equal; outer petals 3, 15 - 37 mm long, 10 - 28 mm wide, ovate, apex acuminate, base truncate, greyish yellow, margins flat, densely pubescent to brown tomentose outside, glabrous but pubescent towards margins inside; inner petals 3, valvate, 14 - 34 mm long, 11 - 20 mm wide, ovate, apex acuminate, base truncate, light yellow to cream, margins flat, densely pubescent to brown tomentose outside, pubescent towards margins inside; stamens 3680 to 5256, in 25 to 35 rows, 3.5 - 4.5 mm long, linear; connective discoid, glabrous, cream; staminodes absent; carpels free, 23 to ca. 150, ovary 4 - 5 mm long, stigma bilobed, slightly capitate, densely pubescent. Monocarps sessile to stipitate, stipes 0 - 5 mm long, 2.5 - 3 mm in diameter; 3 to 35 monocarps, 27 - 55 mm long, 9 - 25 mm in diameter, ellipsoid, apex rounded, tomentose, bumpy, otherwise smooth, brown when ripe; seeds 7 to 13 per monocarp, ca. 13 mm long, 8 - 10 mm in diameter, ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
436ED951CE56B644979EFA0FDB0810DA.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West and Central Africa, from Cote d'Ivoire to Ghana, and from Nigeria to Cameroon. In Cameroon known from the South, Central, Littoral, South-West and North-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
436ED951CE56B644979EFA0FDB0810DA.taxon	materials_examined	Specimens examined. Central Region: Sonossi 26 km W of Ndikinimeki, 4.77 ° N, 10.83 ° E, 29 March 1982, Asonganyi J. N. 421 (P, YA); Colline entre Tcherikoy et Sokelle II (30 km NW Eseka), 3.80 ° N, 10.56 ° E, 14 December 1973, Letouzey R. 12352 (P, YA). Littoral Region: 8 km W of Masok, 4.13 ° N, 10.4 ° E, 31 March 1965, Leeuwenberg A. J. M. 5282 (BR, K, MO, P, WAG, YA). North-West Region: Baji-Tumbo (Wum), 6.38 ° N, 10.07 ° E, 12 July 1975, Letouzey R. 14020 (K, P, WAG, YA); Bamenda Prov Wum Distr Nkom-Wum FR, 6.26 ° N, 10.13 ° E, 03 July 1951, Ujor E. U. 29281 (K). South Region: on road Lolodorf-Bipindi ca half way near Mbiguiligui village (Mbikiliki), 3.16 ° N, 10.53 ° E, 26 February 2018, Couvreur T. L. P. 1157 (K, MPU, P, WAG, YA); 31 km east from Lele village, 2.27 ° N, 13.29 ° E, 09 September 2013, Couvreur T. L. P. 486 (WAG, YA); ca 7 km NE of Ebom, 3.11 ° N, 10.75 ° E, 01 August 1996, Parren M. P. E. 223 (KRIBI, WAG); ca 7 km NE of Ebom, 3.11 ° N, 10.75 ° E, 01 August 1996, Parren M. P. E. 68 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 1898, Zenker G. A. 1738 (B, M, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1901, Zenker G. A. 2344 (B, L, M, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 April 1903, Zenker G. A. s. n. (P). South-West Region: Along the path from Pete to Bopo at the right hand side of the road in S Bakundu FR, 4.46 ° N, 9.392 ° E, 23 February 1956, Binuyo A. 35564 (K, WAG); Mungo river forest reserve ca 1 km East of bridge Chained road to S, 4.73 ° N, 9.55 ° E, 24 October 1998, Cheek M. 9337 (K, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.500 ° E, 26 March 2016, Couvreur T. L. P. 1013 (WAG, YA); on top of hill near Small Ekombe village 3 km after Kumba on road to Ekondo Titi town, 4.62 ° N, 9.376 ° E, 13 January 2016, Couvreur T. L. P. 980 (WAG, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.500 ° E, 25 March 2016, Couvreur T. L. P. 999 (WAG, YA); South Bakundu, 4.49 ° N, 9.374 ° E, 19 February 1946, Dundas J. 13989 (K); Entre Babong et Okurikang 35 km WSW Mamfe, 5.58 ° N, 9.05 ° E, 29 May 1975, Letouzey R. 13673 (P, YA); Bolo forest 5 km west of Kumba-Mamfe road near Konye, 4.64 ° N, 9.45 ° E, 25 March 1986, Nemba J. 64 (K, MO, P, WAG, YA); Kumba Distr Mumbo-Southern Bakossi, 4.83 ° N, 9.333 ° E, 09 May 1951, Olorunfemi J. 30561 (K); S Bakundu FR between Bombe Rest House and Mbalange, 4.46 ° N, 9.452 ° E, 19 March 1953, Onochie C. F. A. 30860 (K); Korup National Park, 5.26 ° N, 9.183 ° E, 24 March 1984, Thomas D. W. 3322 (K, P, YA); Takamanda Forest Reserve, 6.23 ° N, 9.316 ° E, 21 March 1985, Thomas D. W. 4549 (YA); 31 km West of Kumba on Mbonge road, 4.51 ° N, 9.366 ° E, 26 March 1986, Thomas D. W. 5965 (K, MO, P, WAG, YA); Lake Barombi Kumba, 4.65 ° N, 9.4 ° E, 01 April 1986, Thomas D. W. 6090 (P, YA); Korup National Park, 5.27 ° N, 9.1 ° E, 01 April 1988, Thomas D. W. 7499 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8254C187545F401103EBFE7FF6A4AE5D.taxon	description	Figs 120, 121; Map 15 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8254C187545F401103EBFE7FF6A4AE5D.taxon	materials_examined	Type. Equatorial Guinea. Bioko Norte; Bioko (Fernando Po), Mann G. 1159, 1861: lectotype, here designated: K [K 000198803]; isolectotypes: K [K 000198804, K 000198805]; P [P 00362655].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8254C187545F401103EBFE7FF6A4AE5D.taxon	description	Description. Tree, 3 - 20 m tall, d. b. h. 2 - 25 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous. Leaves: petiole 4.5 - 21 mm long, 2 - 6 mm in diameter, glabrous, grooved, blade inserted on top of the petiole; blade 25.4 - 63.6 cm long, 7 - 17.7 cm wide, narrowly elliptic to narrowly oblong, apex acuminate, acumen 0.7 - 2.4 cm long, base rounded (sometimes acute, truncate or subcordate), subcoriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 15 to 29 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 5 - 40 mm long, 1.4 - 4 mm in diameter, glabrous to pubescent; in fruit 13 - 31 mm long, 3 - 8 mm in diameter, glabrous; bracts 2 to 5, several basal and one upper towards the upper half of pedicel, basal bracts 2 - 3 mm long, 1 - 3 mm wide; upper bract 4 - 13 mm long, 6 - 14 mm wide; sepals 3, basally fused, valvate or slightly imbricate, 5 - 14 mm long, 7 - 17 mm wide, ovate, apex acuminate, base truncate, green, glabrous outside, pubescent and glabrous towards center inside, margins flat; petals free, sub equal; outer petals 3, 11 - 32 mm long, 4 - 26 mm wide, ovate, apex acuminate, base truncate, purple to wine red, margins flat, glabrous outside, glabrous or pubescent towards the margins inside; inner petals 3, valvate, 9.5 - 30 mm long, 8 - 20 mm wide, ovate, apex acuminate, base truncate, purple to wine red, margins flat, glabrous outside, glabrous to pubescent with glabrous base inside; stamens 1908 to 2456, in 15 to 25 rows, 2 - 5 mm long, linear; connective discoid, glabrous, cream; staminodes absent; carpels free, 6 to 20, ovary 3 - 7 mm long, stigma bilobed, slightly capitate, densely pubescent. Monocarps sessile to stipitate, stipes 0 - 8 mm long, 3 - 10 mm in diameter; monocarps 1 to 10, 22 - 55 mm long, 17 - 32 mm in diameter, ellipsoid to globose, apex rounded, glabrous, smooth, dumpy; seeds 8 to 23 per monocarp, 13 - 29 mm long, 3 - 14 mm in diameter, ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8254C187545F401103EBFE7FF6A4AE5D.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West and Central Africa, from Cote d'Ivoire to Ghana, and from Nigeria to Gabon. In Cameroon known from the Littoral, South, and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
8254C187545F401103EBFE7FF6A4AE5D.taxon	materials_examined	Specimens examined. South Region: ca 6 km S of Kribi 2 - 4 km E of Gr Batanga, 2.88 ° N, 9.916 ° E, 26 September 1969, Bos J. J. 5412 (P, WAG); Campo Ma an National Park 5 km after main entrance, 2.35 ° N, 10.25 ° E, 15 February 2012, Couvreur T. L. P. 383 (WAG, YA); 29 km east from Lele village, 2.27 ° N, 13.29 ° E, 09 September 2013, Couvreur T. L. P. 484 (WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1903, Zenker G. A. 2624 (B, K, L, M, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3204 (K, L, M, MO, P, P); Mimfia, 3.06 ° N, 10.38 ° E, 01 September 1913, Zenker G. A. 358 (M, P, U, WAG). South-West Region: Mabeta 6 km SE Limbe SBL, 3.98 ° N, 9.283 ° E, 10 August 1993, Baker W. J. 294 (K, YA); Ekundu Kundu, 5.12 ° N, 8.895 ° E, 27 April 1996, Cable S. 2230 (K, YA); Dikulu, 3.98 ° N, 9.233 ° E, 17 December 1993, Cable S. 611 (K, YA); Liwenyi, 4.37 ° N, 9.013 ° E, 28 October 1993, Cheek M. 5180 (K, YA); ca 40 minutes walk N then E from Njonji Hunters path to Lake Njonji, 4.13 ° N, 8.993 ° E, 18 November 1993, Cheek M. 5462 (K, YA); Ekundu Kundu, 5.13 ° N, 8.869 ° E, 25 April 1996, Cheek M. 8164 (K, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.078 ° E, 03 April 2016, Couvreur T. L. P. 1051 (WAG, YA); Mokoko Forest Reserve Boa / Likinge, 4.42 ° N, 8.972 ° E, 31 May 1994, Ekema S. N. 1078 (K, YA); Bakolle Bakossi on Kumba-Mamfe road, 5.01 ° N, 9.666 ° E, 24 May 1986, Etuge M. 156 (K, MO, WAG, YA); Nyasoso, 4.81 ° N, 9.683 ° E, 24 June 1996, Etuge M. 2396 (K, YA); Mungo FR, 4.73 ° N, 9.560 ° E, 22 February 2006, Etuge M. 6506 (K); Mahole-Bintulu road, 4.79 ° N, 9.603 ° E, 24 November 1999, Gosline W. G. 209 (K, WAG, YA); Mabeta-Moliwe, 3.98 ° N, 9.25 ° E, 06 April 1992, Jaff B. 73 (K, YA); Rivieres de Mosongosele et de Ndian depuis Mosongosele jusqu'a l'entree amont de la mangrove environ 20 km SW de Mundemba, 4.83 ° N, 8.765 ° E, 13 June 1976, Letouzey R. 15175 (P, YA); Buea are at Bolifamba, 4.13 ° N, 9.303 ° E, 01 January 1929, Maitland T. D. 537 (K, P); Bomana-Koto Road c 500 m Bearing 305 deg towards Onge river, 4.31 ° N, 9.016 ° E, 18 October 1993, Ndam N. 708 (K, YA); Bolo forest 5 kms west of Kumba-Mamfe road near Konye, 4.86 ° N, 9.429 ° E, 25 March 1986, Nemba J. 56 (U); Mont versant de Idenao, 4.24 ° N, 8.99 ° E, 23 January 1985, Nkongmeneck B. A. 959 (YA); Bechati-Fossimondi-Besali forest Path leading from Fossimondi to Besali, 5.64 ° N, 9.966 ° E, 28 April 2005, Tchiengue B. 2204 (K, YA); Above Isobe, 4.16 ° N, 9 ° E, 10 June 1992, Tekwe C. F. 87 (K, YA); Mount above small Koto village, 4.3 ° N, 9.1 ° E, 06 March 1985, Thomas D. W. 4447 (K, P, YA); 3 km N of Limbe-Idenao road, 4.05 ° N, 9.083 ° E, 10 February 1986, Thomas D. W. 5537 (YA); Matene from Mbilishe, 6.25 ° N, 9.37 ° E, 01 March 1987, Thomas D. W. 6928 (P, YA); west of the Onge River and ridges on " Thump Mount ", 4.33 ° N, 8.95 ° E, 09 November 1993, Thomas D. W. 9875 (K, YA); ca 5 km North East of Limbe TB + 5500 m, 4 ° N, 9.25 ° E, 02 June 1992, Watts J. 336 (K, YA); Mabeta-Moliwe TD 5835 m, 4.01 ° N, 9.266 ° E, 24 June 1992, Wheatley J. I. 326 (K, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA6FC380D35E0D38D165BD8DCEB21F23.taxon	description	Fig. 122; Map 15 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA6FC380D35E0D38D165BD8DCEB21F23.taxon	materials_examined	Type. Type. Equatorial Guinea. Bioko Norte; Bioko (Fernando Po), Mann G. 308, 1860: holotype: K [K 000198801]; isotype: P [P 00362657].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA6FC380D35E0D38D165BD8DCEB21F23.taxon	description	Description. Tree, 3 - 15 (20) m tall, up to 35 cm d. b. h.; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to slightly pubescent. Leaves: petiole 4 - 35 mm long, 2 - 8 mm in diameter, glabrous to pubescent with long white hairs, slightly grooved, blade inserted on top of the petiole; blade 15.9 - 70 cm long, 4.3 - 22.5 cm wide, narrowly elliptic to elliptic to narrowly obovate to obovate, apex acuminate, acumen 0.4 - 1.5 cm long, base rounded to acute (sometimes slightly truncate), coriaceous, below densely pubescent with long white hairs to glabrous when young, glabrous when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young, glabrous to slightly pubescent when old; secondary veins 15 to 33 per side, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 0 - 7.5 mm long, ca. 4 mm in diameter, pubescent; in fruit 9 - 15 mm long, 7 - 8 mm in diameter, densely pubescent; bracts 2 to 6, basal bracts 2 - 3 mm long, 1 - 3 mm wide; upper bracts 8 - 22 mm long, 10 - 50 mm wide; sepals 3, valvate, basally fused over 20 - 50 % of the length, 11 - 30 mm long, 13 - 26 mm wide, ovate, apex acuminate, base truncate, densely pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 20 - 40 mm long, 17 - 30 mm wide, ovate, apex acuminate, base truncate, margins flat, densely pubescent outside, glabrous inside, cream with a red streak inside; inner petals 3, valvate, 20 - 42 mm long, 15 - 29 mm wide, ovate, apex acuminate, base truncate, margins flat, pubescent outside, glabrous inside, cream with a red streak inside; stamens 1000 to 3000, in 19 to 25 rows, 3.4 - 5 mm long, 0.1 - 0.5 mm wide, linear; connective discoid, pubescent; staminodes absent; carpels free, 20 to 104, ovary 4 - 7 mm long, stigma bilobed, slightly capitate, densely pubescent. Monocarps [but see comment in notes] sessile to stipitate, stipes 0 - 4 mm long, 2 - 5 mm in diameter; monocarps 6 to 24, 20 - 50 mm long, 11 - 25 mm in diameter, obovoid to oblong, apex rounded or shortly narrowed, pubescent, smooth; seeds 6 to 16 per monocarp, 10 - 15 mm long, 6 - 11 mm in diameter, flattened ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA6FC380D35E0D38D165BD8DCEB21F23.taxon	distribution	Distribution. Nigeria, Cameroon, Equatorial Guinea; in Cameroon known from the South-West region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
EA6FC380D35E0D38D165BD8DCEB21F23.taxon	materials_examined	Specimens examined. South-West Region: Likombe, 4.11 ° N, 9.183 ° E, 22 February 1995, Cable S. 1353 (K, YA); Upper Boando, 4.06 ° N, 9.15 ° E, 14 March 1995, Cable S. 1524 (K, YA); Upper Boando, 4.06 ° N, 9.15 ° E, 16 March 1995, Cable S. 1626 (K, YA); Ekundu Kundu, 5.14 ° N, 8.893 ° E, 26 April 1996, Cable S. 2187 (K, WAG, YA); Liwenyi, 4.37 ° N, 9.013 ° E, 27 October 1993, Cheek M. 5145 (K, YA); on trail leading to top of Mt Etinde after Ekonjo village, 4.06 ° N, 9.152 ° E, 01 April 2016, Couvreur T. L. P. 1026 (WAG, YA); on trail leading to top of Mt Etinde after Ekonjo village, 4.06 ° N, 9.153 ° E, 01 April 2016, Couvreur T. L. P. 1029 (WAG, YA); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.056 ° E, 02 April 2016, Couvreur T. L. P. 1040 (WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.101 ° E, 03 April 2016, Couvreur T. L. P. 1046 (WAG, YA); slopes of Mount Cameroon on the Bokwango trail near Bokwango village 4 km south west of Bu 4.12 ° N, 9.186 ° E, 23 March 2016, Couvreur T. L. P. 990 (WAG, YA); slopes of Mount Cameroon on the Bokwango trail near Bokwango village 4 km south west of Bu 4.12 ° N, 9.170 ° E, 23 March 2016, Couvreur T. L. P. 992 (WAG, YA); Likombe, 4.11 ° N, 9.183 ° E, 02 March 1995, Dahl A. 622 (K, YA); Likombe, 4.11 ° N, 9.183 ° E, 21 February 1995, Groves M. 122 (K, YA); Kumba Distr eastern boundary of Bambuko FR ca 11 miles SSW of Musome, 4.33 ° N, 9.166 ° E, 01 February 1958, Keay R. W. J. 37485 (K); on Shrike Trail leading from Nyasoso to summit, 4.83 ° N, 9.666 ° E, 20 June 1994, Lane P. 142 (K, YA); Bu 4.15 ° N, 9.233 ° E, 1898, Lehmbach H. 178 (B, K); Bu 4.15 ° N, 9.233 ° E, 1898, Lehmbach H. 57 (B, K); Buea above upper farms, 4.15 ° N, 9.233 ° E, 01 March 1929, Maitland T. D. 453 (K); Cameroon Mountain Buea area 4.15 ° N, 9.233 ° E, 01 January 1930, Maitland T. D. s. n. (K [K 000105371]); Likomba-Pflanzung 15 - 35 km NE von Victoria [Limbe], 4.1 ° N, 9.333 ° E, 01 November 1928, Mildbraed G. W. J. 10720 (K); Mt Cameroun flanc d'Ekona Lelu, 4.27 ° N, 9.3 ° E, 14 January 1985, Nkongmeneck B. A. 891 (YA); Between Limbe & Bimbia, 3.96 ° N, 9.25 ° E, 1895, Preuss P. R. 1378 (P); Mount above small Koto village, 4.3 ° N, 9.1 ° E, 06 March 1985, Thomas D. W. 4469 (K, MO, P, YA); Etinde Mont, 4.08 ° N, 9.116 ° E, 29 January 1994, Wieringa J. J. 2058 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4398DDE4FB6EB38F8AA9FD83296D8252.taxon	description	Figs 123, 124; Map 15 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4398DDE4FB6EB38F8AA9FD83296D8252.taxon	materials_examined	Type. Cameroon. Central Region; Yaounde, Zenker G. A. & Staudt A. 108, 1895: lectotype designated by Fries (1930), p. 62, sheet destroyed at B, sheet here designated: P [P 00362654]; isolectotype: COI [COI 00004926].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4398DDE4FB6EB38F8AA9FD83296D8252.taxon	description	Description. Differs from the type variety in having young branches and petioles covered with long soft hairs producing a whitish appearance quickly falling off (vs. young branches and petiole sparsely pubescent to glabrous); leaves 30 - 70 cm long and 8.2 - 22.5 cm wide (vs. 15.9 - 45 cm long and 4.3 - 11.8 cm wide), secondary veins 22 to 33 (vs. 15 to 24), and flowers with sepals 21 - 30 mm long (vs. 11 - 23 mm long); carpels 33 to 104 (vs. 20 to 70).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4398DDE4FB6EB38F8AA9FD83296D8252.taxon	distribution	Distribution. From southern Nigeria to Gabon and Democratic Republic of the Congo; in Cameroon known from the East, South, Central and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4398DDE4FB6EB38F8AA9FD83296D8252.taxon	materials_examined	Selected specimens examined. Central Region: Mont Mbam Minkon on trail 5 km from Nkol Nyada village, 3.96 ° N, 11.40 ° E, 21 March 2013, Couvreur T. L. P. 419 (WAG, YA); Yaounde, 3.87 ° N, 11.52 ° E, 1895, Zenker G. A. 108 (P); Yaunde, 3.86 ° N, 11.51 ° E, Feburary 1895, Zenker G. A. 698 (K). East Region: 80 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.16 ° N, 14.70 ° E, 05 March 2019, Couvreur T. L. P. 1206 (MPU, WAG, YA); 66 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.08 ° N, 14.67 ° E, 08 March 2019, Couvreur T. L. P. 1229 (MPU, WAG, YA). South Region: Pres d'Alati-Ancienne piste Alati-Mintom II, 2.2 ° N, 13.42 ° E, 17 January 1973, Biholong M. 279 (P, YA); Massif de Ngovayang village de Atog Boga, 3.22 ° N, 10.50 ° E, 04 September 2015, Droissart V. 2125 (BRLU); Bipindi, 3.08 ° N, 10.41 ° E, 1898, Zenker G. A. 1438 (L). South-West Region: Nyasoso village on max's trail to Mt 4.82 ° N, 9.699 ° E, 05 April 2016, Couvreur T. L. P. 1057 (WAG, YA); on trail from Ekongo village located 5 km before the entrance to Limbe 7 km on secondary road On flank of Mt Etinde 100 m in Mont Cameroon National Park, 4.07 ° N, 9.133 ° E, 16 October 2013, Couvreur T. L. P. 512 (WAG, YA); Bakossi Mountains 1 - 8 km NNE of Menyum Village, 5.05 ° N, 9.612 ° E, 22 May 1987, Doumenge C. 473 (L, P); Ebondji, 4.76 ° N, 9.598 ° E, 08 June 2017, Kamdem N. 557 (YA); Forest trail 2 km south from Etube-Tape village, 4.85 ° N, 9.7 ° E, 02 February 1995, Lane P. 501 (K, WAG, YA); Cameroon Mountain, 4.12 ° N, 9.187 ° E, 21 June 2001, van Andel T. R. 3761 (U, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DD692A371BA19485136D457E48E42512.taxon	description	Figs 125, 126; Map 15 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DD692A371BA19485136D457E48E42512.taxon	materials_examined	Type. Cameroon. East Region; Sudkameruner Waldgebiet: Bezirk Molundu, ' Bange Busch', unbewohnter Urwald zwischen Lokomo, Bumba und Bange, Mildbraed G. W. J. 4373, 29 Jan 1911: holotype: B [B 10 0153118].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DD692A371BA19485136D457E48E42512.taxon	description	Description. Tree to shrub, 1.2 - 10 m tall, d. b. h. up to 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to slightly pubescent. Leaves: petiole 2 - 17 mm long, 1.5 - 5 mm in diameter, glabrous to slightly pubescent, slightly grooved, blade inserted on top of the petiole; blade 14.1 - 46 cm long, 5.1 - 16 cm wide, narrowly elliptic to oblong to obovate, apex attenuate to acuminate, acumen 0.1 - 3.8 cm long, base rounded to decurrent, papyraceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous to glabrescent when young, glabrous when old; secondary veins 12 to 22 pairs per side, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences cauliflorous or ramiflorous on old or young foliate branches, axillary. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 2 - 14 mm long, 1 - 4 mm in diameter, pubescent; in fruit 3 - 15 mm long, 2 - 6 mm in diameter, pubescent; bracts 2 to 6, several basal and one upper towards the middle of pedicel, basal bracts 2 - 3 mm long, 1 - 3 mm wide; upper bract 3 - 8 mm long, 3 - 10 mm wide; sepals 3, imbricate, free or sometimes fused at base, 5 - 9 mm long, 5.5 - 10 mm wide, ovate, apex acuminate, base truncate, green, densely pubescent outside, glabrous inside, margins flat; petals free, sub equal; outer petals 3, 9 - 30 mm long, 6 - 17 mm wide, ovate, apex acuminate, base truncate, cream-colored with a purple blotch at the base, margins flat, densely pubescent outside, glabrous inside; inner petals 3, valvate, 9 - 27 mm long, 5 - 16 mm wide, ovate, apex acuminate, base truncate, cream-colored with a purple blotch at the base, margins flat, glabrous to densely pubescent outside, glabrous inside; stamens 1000 to 1500, in 15 to 20 rows, 0.8 - 2 mm long, 0.1 - 0.5 mm wide, linear; connective discoid, pubescent, cream; staminodes absent; carpels free, 8 to 36, ovary 2 - 4 mm long, 0.4 - 2 mm wide, stigma bilobed, slightly capitate, densely pubescent. Monocarps shortly stipitate, stipes ca. 0 - 5 mm long, ca. 2 - 5 mm in diameter; monocarps 1 to 12, 20 - 60 mm long, 9 - 30 mm in diameter, ellipsoid to oblong ovoid, apex rounded, bumpy, otherwise smooth, glabrous to sparsely pubescent, greyish green to orange when ripe; seeds 3 to 10 per monocarp, 12 - 15 mm long, 7 - 11 mm wide, oblong, yellow to pale brown; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DD692A371BA19485136D457E48E42512.taxon	distribution	Distribution. From south Nigeria to Gabon to Central African Republic (southeast), and one collection from Democratic Republic of Congo; in Cameroon known from the East, South, Central Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
DD692A371BA19485136D457E48E42512.taxon	materials_examined	Selected specimens examined. Central Region: Eastern part of park West of Ndangan 1, 3.62 ° N, 11.60 ° E, 18 March 2004, Cheek M. 11839 (K, YA); Colline '' Nkom-Benka'a' ' Nkom-Benka'a' ' dans l'appellation locale 4 - 5 km N du village Ekekam, 3.38 ° N, 11.8 ° E, 09 March 1978, Dang D. 643 (P, YA); Mefou National park 18 km from MFOU, 3.61 ° N, 11.58 ° E, 22 March 2004, Tadjouteu F. 566 (K, YA). East Region: A 30 km au NE de Bange (km 75 route Yokadouma-Moloundou), 3.02 ° N, 15.12 ° E, 25 May 1963, Letouzey R. 5142 (P, YA); Moloundou near Lokomo Bumba and Bange, 2.08 ° N, 15.25 ° E, 29 January 1911, Mildbraed G. W. J. 4373 (B); Camp CFA Bango I Bord de la riviere Bango 36 km SE Bateha Malen village situe a 21 km N de Moloundou, 2.15 ° N, 15.35 ° E, 08 April 1971, Villiers J. - F. 625 (P, YA). Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1172 (MPU, P, WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.91 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 652 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 655 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 20 June 2014, Couvreur T. L. P. 656 (WAG, YA). South Region: 13 km from Kribi Ebolowa road, 2.87 ° N, 9.980 ° E, 13 November 1968, Bos J. J. 3259 (P, WAG, YA); S Bbank Lobe R SE of Grand Batanga ferry riverine forest edge, 2.86 ° N, 9.9 ° E, 11 October 1969, Bos J. J. 5474 (BR, MO, P, WAG, YA); 40 km from Kribi 4 km W of Edea road N bank of lokoundje R, 3.08 ° N, 10.25 ° E, 12 March 1970, Bos J. J. 6521 (WAG); ca 5 km N of km 7 Kribi-Ebolow, 2.93 ° N, 9.95 ° E, 10 July 1970, Bos J. J. 7075 (P, WAG); Massif de Ngovayang village de Ngovayang sommet, 3.25 ° N, 10.57 ° E, 08 June 2015, Droissart V. 1880 (BRLU); Campo-Ma'an area Bibabimvoto, 2.25 ° N, 10.36 ° E, 01 February 2000, Elad M. 1270 (KRIBI, WAG); Meyo Centre, 2.55 ° N, 11.03 ° E, 24 March 1970, Letouzey R. 10222 (P); Region de Kribi, 2.93 ° N, 9.92 ° E, 01 January 1911, Mildbraed G. W. J. 5936 (B); Environs de Mvini 34 km east of Campo, 2.37 ° N, 10.09 ° E, 24 October 1984, Nkongmeneck B. A. 800 (YA); Campo-Ma'an area Bibabimvoto, 2.27 ° N, 10.05 ° E, 26 August 2000, Tchouto Mbatchou G. P. T 8 X _ 70 (WAG); Campo-Ma'an area 2.71 ° N, 9.866 ° E, 26 October 2001, van Andel T. R. 4228 (KRIBI, U, WAG). South-West Region: Mokoko Forest Reserve Ekumbe-Mofako, 4.47 ° N, 9.092 ° E, 21 April 1994, Akogo M. 234 (K, YA); Dikome, 4.92 ° N, 9.240 ° E, 05 May 1994, Sonke B. 1226 (K); Mapanja, 4.08 ° N, 9.15 ° E, 20 April 1992, Tchouto Mbatchou G. P. 136 (K, YA); Mokoko Above Bonja village, 4.41 ° N, 9 ° E, 23 March 1993, Tchouto Mbatchou G. P. 611 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
77ED167318A90B870DFECD6E6738C498.taxon	description	Description. Trees, 3 - 25 m tall, d. b. h. up to 40 cm; stilt roots or buttresses absent. Indumentum of simple hairs or glabrous. Leaves: petiole 2 - 8 mm long, 1 - 6 mm in diameter; blade 10 - 52 cm long, 3.5 - 14 cm wide, ovate, elliptic, obovate or oblong, apex acuminate to abruptly acuminate, acumen 0.7 - 2 cm long, base cuneate to cordate; midrib sunken or flat; secondary veins 6 to 20 pairs; tertiary venation reticulate. Flowers unisexual, monoecious, male and female flowers similar or dissimilar, or bisexual (in one species not present in Cameroon), with (5) 6 perianth parts in 2 whorls. Inflorescences ramiflorous or cauliflorous, flower buds globose or conical, 1 to 6 (50) per inflorescence; pedicel 1 - 450 mm long; in fruit 3 - 450 mm long; bracts 1 to 4, one basal and one upper, or all basal, 1 - 2 mm long; sepals 2, valvate (imbricate), free or basally fused, 1.5 - 10 mm long, ovate to semi-circular, apex acute, acuminate or attenuate, base truncate; petals free or basally fused, (3) 4, valvate, 2 - 45 mm long, 2.5 - 17 mm wide, oblong, ovate, elliptic or linear, apex acuminate or attenuate, base truncate; stamens 100 to 1000, in 9 to 30 rows, 1 - 2 mm long, oblong to elongated; connective reduced or absent, glabrous; staminodes absent; carpels free, 15 to 280, 1 - 4 mm long, stigma ovoid, coiled or flat, glabrous. Monocarps sessile or stipitate, stipes 1 - 10 mm long; monocarps 2 to 25, 15 - 80 mm long, 9 - 55 mm in diameter, cylindrical, ellipsoid or globose, apex rounded or apiculate, smooth, verrucose or bumpy; seeds 4 - 25 mm long, 3 - 15 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
19A7D27AA2D30872D4E1D88786D116AA.taxon	description	Figs 127, 128; Map 15 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
19A7D27AA2D30872D4E1D88786D116AA.taxon	materials_examined	Type. Nigeria. Cross River State; Oban, Talbot P. 1517, 1912: holotype: K [K 000199043]; isotype: BM [BM 000554076].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
19A7D27AA2D30872D4E1D88786D116AA.taxon	description	Description. Tree, 2 - 7 m tall, d. b. h. 3 - 7 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 3 - 5 mm long, 2 - 3 mm in diameter, glabrous to pubescent, slightly grooved, blade inserted on top of the petiole; blade 25 - 34 cm long, 4.5 - 9 cm wide, narrowly oblong to narrowly obovate, apex acuminate, acumen 1 - 3 cm long, base rounded to subcordate, coriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below pubescent when young, glabrous when old; secondary veins 11 to 20 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious, inflorescences cauliflorous. Flowers with 6 perianth parts in 2 whorls, long conical in bud, 1 to 2 per inflorescence, male and female inflorescences similar, but with a tendency of female flowers being located towards the base of the trunk; pedicel 3 - 8 mm long, ca. 1 mm in diameter, pubescent; in fruit 4 - 11 mm long, 2 mm in diameter, pubescent; bracts 2 to 3, all basal or towards the lower half of pedicel, 1 - 2 mm long, 1 - 2 mm wide; sepals 2, valvate, free, 3 - 4 mm long, 2 mm wide, triangular, apex attenuate, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals free, 4, 25 - 45 mm long, 4 - 8 mm wide, linear, apex attenuate, base truncate, wine-brown to dark red, margins flat, pubescent outside, glabrous and highly verrucose inside; male flowers: stamens 400 to 600, in 15 to 20 rows, ca. 1 mm long, broad; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 24 to 40, ovary 2 - 3 mm long, 1 - 1.5 wide, stigma coiled, densely pubescent. Monocarps stipitate, stipes 3 - 6 mm long, ca. 2 mm in diameter; monocarps 4 to 6, 25 - 50 mm long, 10 - 20 mm in diameter, cylindrical, apex rounded, puberulent to glabrous, verrucose, wrinkled, bright red when ripe; seeds 6 to 16 per monocarp, 17 - 22 mm long, 10 - 12 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
19A7D27AA2D30872D4E1D88786D116AA.taxon	distribution	Distribution. From eastern Nigeria to northern Cameroon; in Cameroon known from South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
19A7D27AA2D30872D4E1D88786D116AA.taxon	materials_examined	Specimens examined. South-West Region: Kumba Div S Bakundu FR ca 1 mile west of Bopo village, 4.48 ° N, 9.372 ° E, 12 March 1948, Brenan J. P. M. 9305 (K); Banga (S Bakundu), 4.5 ° N, 9.57 ° E, 13 March 1948, Brenan J. P. M. 9409 (P); Ekundu Kundu, 5.18 ° N, 8.859 ° E, 25 April 1996, Cable S. 2150 (K, YA); Ekundu Kundu, 5.12 ° N, 8.895 ° E, 27 April 1996, Cable S. 2256 (K, YA); Korup National Park nature trail near suspension bridge, 4.98 ° N, 8.85 ° E, 01 February 1995, Cheek M. 7234 (K, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.502 ° E, 25 March 2016, Couvreur T. L. P. 1000 (WAG, YA); Bayang Mbo Wildlife Sanctuary after Mbu river, 5.35 ° N, 9.501 ° E, 26 March 2016, Couvreur T. L. P. 1015 (WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.098 ° E, 03 April 2016, Couvreur T. L. P. 1045 (WAG, YA); Ntali, 5.25 ° N, 9.576 ° E, 30 November 2000, Etuge M. 4860 (K); South Bakundu forest reserve, 4.5 ° N, 9.57 ° E, 14 May 1970, Farron C. 6613 (YA); South Bakundu forest reserve, 4.54 ° N, 9.42 ° E, 01 May 1970, Farron C. 7297 (P); 3 - 25 km S of Six cup Garri Creek, 5.03 ° N, 8.883 ° E, 06 March 1993, Gereau R. E. 5195 (MO, WAG, YA); Entre Etinkem et Nfaitok 16 10 km N Nguti, 5.41 ° N, 9.39 ° E, 15 June 1975, Letouzey R. 13841 (P, YA); Akpasang river camp (Mouanko Region), 4.98 ° N, 8.71 ° E, 07 February 1976, McKey D. B. 70 (P, YA); Etome, 4.1 ° N, 9.05 ° E, 13 February 1997, Tchouto Mbatchou G. P. 1663 (K, YA); Korup National Park, 4.98 ° N, 8.85 ° E, 01 March 1979, Thomas D. W. 1086 (K); Korup National Park, 5.26 ° N, 9.183 ° E, 24 March 1984, Thomas D. W. 3336 (YA); Korup National park Forest along footpath from Ndian River at PAMOL Field 69 and transect P, 5.01 ° N, 8.833 ° E, 24 January 1985, Thomas D. W. 4300 (P, YA); Korup National Park, 5.18 ° N, 8.85 ° E, 16 February 1986, Thomas D. W. 5606 (MO); Korup National Park, 5.33 ° N, 8.9 ° E, 22 May 1988, Thomas D. W. 7835 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76B7F4995B085BAC0EE2E69B75C49027.taxon	description	Fig. 129; Map 15 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76B7F4995B085BAC0EE2E69B75C49027.taxon	materials_examined	Type. Gabon. Estuaire; Monts de Cristal, near first bridge after Kinguele village, 0 ° 46 ' 66 " N, 10 ° 27 ' 81 " E, Couvreur T. L. P 1143, 14 Jun 2016: holotype: WAG; isotypes: LBV, P.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76B7F4995B085BAC0EE2E69B75C49027.taxon	description	Description. Tree, 4 - 10 m tall, d. b. h. 3 - 10 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent. Leaves: petiole 4 - 8 mm long, 3 - 5 mm in diameter, pubescent when young, pubescent to glabrous when old, grooved on top, blade inserted on top of the petiole, strong lemon scent when crushed; blade 31.2 - 50 cm long, 8.8 - 12 cm wide, elliptic to obovate, apex acuminate, acumen 2 - 3 cm long, base subcordate to cordate, coriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 17 to 19 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous, sparsely spaced along the trunk mostly towards the lower half of the trunk. Flowers with 6 perianth parts in 2 whorls, ovoid to conical in bud, 1 to 2 per inflorescence, male and female inflorescences similar; pedicel 0 - 2 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit unknown; bracts up to 3, all basal, 1 - 2 mm long, 4 mm wide; sepals 2, valvate, basally fused, enclosing the petals in bud, 9 - 15 mm long, 4 - 6 mm wide, narrowly ovate, apex acute, base truncate, densely pubescent with hairs appressed outside, densely pubescent or glabrous towards base inside, margins flat; petals 4, 7 - 15 mm long, 5 - 8 mm wide, ovate, apex acute, base truncate, brownish-greenish-yellow, margins flat, pubescent outside, glabrous inside; male flowers: stamen number unknown, 0.5 mm long, oblong, connective truncate, glabrous, pale yellow; female flowers: carpels free, ca. 60, ovary 4 - 5 mm long, ca. 0.5 mm wide, densely pubescent with long appressed hairs, stigma cylindrical coiled. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76B7F4995B085BAC0EE2E69B75C49027.taxon	distribution	Distribution. A species only known from southern Cameroon and two localities in Gabon (Monts de Cristal National Park, Mbe sector); in Cameroon known from the South Region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
76B7F4995B085BAC0EE2E69B75C49027.taxon	materials_examined	Specimens examined. South Region: A 15 km au SSE de Zingui (soit a 50 km au SE de Kribi), 2.82 ° N, 10.97 ° E, 14 March 1968, Letouzey R. 9017 (P, YA); Campo-Ma'an area Bibabimvoto, 2.21 ° N, 10.01 ° E, 13 May 2000, Tchouto Mbatchou G. P. 2869 (KRIBI, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F6D52969AFB0E8EE20A67EE6F56DF26.taxon	description	Figs 130, 140; Map 16 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F6D52969AFB0E8EE20A67EE6F56DF26.taxon	materials_examined	Type. Democratic Republic of the Congo. Bandundu; Lusambo, van Kerkhoven E. s. n., 16 Aug 1913: lectotype, here designated: BR [BR 0000008824318].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F6D52969AFB0E8EE20A67EE6F56DF26.taxon	description	Description. Tree to shrub, 2 - 6 m tall, d. b. h. up to 6 cm; stilt roots or buttresses absent. Indumentum of simple hairs or glabrous; old leafless branches glabrous, young foliate branches sparsely pubescent to glabrous. Leaves: petiole 2 - 6 mm long, 1 - 2.5 mm in diameter, glabrous, slightly grooved, blade inserted on top of the petiole; blade 8 - 17.7 cm long, 2.6 - 7.7 cm wide, elliptic, apex attenuate to acuminate, acumen 0.5 - 1.8 cm long, base acute to decurrent, papyraceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 7 to 12 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences sometimes cauliflorous, mainly ramiflorous on young foliate and old leafless branches, axillary. Flowers with 6 perianth parts in 2 whorls, globose in bud, 1 to 2 per inflorescence, male and female inflorescences dimorphic; male pedicels 2.5 - 6 mm long, 0.5 - 1 mm in diameter, pubescent; female pedicel: 3.5 - 8 mm long, ca. 1 mm in diameter, pubescent; in fruit 5 - 15 mm long, 1 - 3 mm in diameter, pubescent to glabrous; male and female bracts 1 to 4, several basal and one upper towards the lower half of pedicel, similar in size, ca. 1 mm long, ca. 1 mm wide. Male and female sepals 2, valvate, free, 0.7 - 1.5 mm long, 1 - 2 mm wide, triangular, apex attenuate, base truncate, brown, pubescent outside, glabrous inside, margins flat; male and female petals free, 4, 2.5 - 7 mm long, 3 - 5 mm wide, ovate, apex acute, base truncate, margins flat, pubescent outside, glabrous, pubescent towards margins inside, light red; male flowers: stamens 300 to 400, in 15 to 20 rows, 0.5 mm long, linear; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 11 to 40, ovary 1.5 - 3 mm long, stigma globose, densely pubescent. Monocarps stipitate, stipes 0 - 4 mm long, 1 - 3 mm in diameter; monocarps 1 to 15, 13 - 45 mm long, 7 - 18 mm in diameter, cylindrical, apex rounded, glabrous, smooth, bumpy, green to orange to red when ripe; seeds 2 to 10 per monocarp, 9.5 - 14 mm long, 7 - 8 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F6D52969AFB0E8EE20A67EE6F56DF26.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West Africa from Ivory Coast and Ghana, and in Central Africa from eastern Nigeria to the Democratic Republic of the Congo and South Sudan; in Cameroon known from the Central, South and South west regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
3F6D52969AFB0E8EE20A67EE6F56DF26.taxon	materials_examined	Specimens examined. Central Region: Mt Eloumden 6 km Sud ouest de Yaounde, 3.81 ° N, 11.43 ° E, Achoundong G. 872 (YA); Ca 30 km S of Mbalmayo, 3.33 ° N, 11.5 ° E, 13 February 1964, de Wilde W. J. J. O 1914 (P, WAG). East Region: Village Nkolbisson 7 km West of Yaounde, 4.16 ° N, 14.11 ° E, 15 April 1962, Breteler F. J. 2812 (K, P, WAG); West side of Sangha River, 2.21 ° N, 16.09 ° E, 01 November 1988, Harris D. J. 1512 (K, P); Bordure de la Sangha au sud de Lidjombo (pres de Libongo) a 110 km au N de Ouesso, 2.93 ° N, 16.04 ° E, 10 April 1971, Letouzey R. 10641 (P, YA); Pres Ndongo a 45 km WNW de Moloundou, 2.58 ° N, 15.29 ° E, 15 March 1973, Letouzey R. 12086 (K, P, WAG, YA); Rive de la Doume pres Bimba (40 km SW de Batouri), 4.16 ° N, 14.11 ° E, 15 April 1962, Letouzey R. 4755 (K, P, YA); A 11 km au SSW de Kosso (village situe a 60 km au SSW de Batouri, 3.93 ° N, 14.17 ° E, 25 July 1963, Letouzey R. 5491 (K, P, YA); Myko-Malapa (Molundou), 2.05 ° N, 15.17 ° E, 21 April 1971, Villiers J. - F. 683 (P, YA). South Region: Rives du Ntem pres du confluent de la Kye 16 km ESE d'Ambam, 2.30 ° N, 11.38 ° E, 07 February 1970, Letouzey R. 10024 (P, YA); Pres de Ngomebae 70 km ESE d'Ebolowa sur route de Mvangan, 2.72 ° N, 11.73 ° E, 24 January 1970, Letouzey R. 9915 (P, YA). South-West Region: Johann-Albrechtshohe [Kumba] area 4.16 ° N, 9.2 ° E, 1896, Staudt A. 556 (K, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
34E5C985E45DDA1E84DE500ABF8B448E.taxon	description	Figs 130, 131 (no map)	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
34E5C985E45DDA1E84DE500ABF8B448E.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Liboko, Thonner F. 100, 22 Sep 1896: holotype, sheets here designated: BR [BR 0000008824394; BR 0000008824202]; isotypes: BR [BR 0000008824219]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
34E5C985E45DDA1E84DE500ABF8B448E.taxon	description	Description. Tree to shrub, 4 - 10 m tall, d. b. h. 20 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent to glabrous. Leaves: petiole 3 - 8 mm long, 1 - 4 mm in diameter, sparsely pubescent to glabrous, slightly grooved, blade inserted on top of the petiole; blade 16.3 - 39.2 cm long, 5 - 11 cm wide, elliptic to obovate, apex attenuate to acuminate, acumen 1 - 1.5 cm long, base acute to rounded, papyraceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 11 to 22 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous. Flowers with 5 perianth parts in 2 whorls, conical in bud, 2 to 3 per inflorescence, male and female inflorescences dimorphic; male and female pedicels 43 - 450 mm long, 1 - 2 mm in diameter, sparsely pubescent, hanging from the base of the trunk and extending on the ground; in fruit 300 - 450 mm long, 2 - 3 mm in diameter, sparsely pubescent to glabrous; bracts 2, one basal and one upper towards the lower half of pedicel, basal bracts 1 - 2 mm long, 1 mm wide, upper bracts 1 - 3 mm long, 1 - 4 mm wide; male sepals 2, valvate, free, ca. 2 mm long, 5 - 6 mm wide, semiorbicular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; female sepals 2, valvate, free, ca. 4 mm long, ca. 10 mm wide, semiorbicular, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; male petals 3, basally fused, tube 2 - 3 mm long, ca. 20 mm long, 5 - 6 mm wide, broadly ovate, apex acute to attenuate, base truncate, margins flat, pubescent outside, glabrous and papillose towards the center, pubescent towards margins inside; female petals 3, basally fused, tube 2 - 3 mm long, 20 - 30 mm long, 8 - 12 mm wide, ovate, apex acute to attenuate, base truncate, margins flat, pubescent outside, glabrous towards center and pubescent towards margins inside; male flowers: stamens 150 to 200, in 9 to 15 rows, 1 mm long, broad to linear; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 20 to 40, ovary 3 - 4 mm long, stigma flat, densely pubescent. Monocarps shortly stipitate, stipes 3 - 4 mm long, 30 - 80 mm long, 10 - 35 mm in diameter; monocarps 1 to 5, cylindrical, apex apiculate, sparsely pubescent to glabrous, verrucose, longitudinally 4 - 6 ribbed, red when ripe; seeds (2) 6 to 16 per monocarp, 6 - 15 mm long, 3 - 10 mm in diameter, 2 to 16, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
34E5C985E45DDA1E84DE500ABF8B448E.taxon	distribution	Distribution. A central African species, from Cameroon (?) and Gabon to the Democratic Republic of the Congo; unseen and verified specimens were collected in Cameroon from the South-West and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D22740D106DD762A15AD08A2D9B5CF9E.taxon	description	Fig. 132; Map 16 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D22740D106DD762A15AD08A2D9B5CF9E.taxon	materials_examined	Type. Cameroon. Littoral Region, Yabassi, Ebo Forest, Dicam Trail 2000 m from Bekob camp, MacKinnon L. E. 51, 25 March 2008; holotype K [K 001381842]; isotypes MO, YA.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D22740D106DD762A15AD08A2D9B5CF9E.taxon	description	Description. Tree, 3 - 4 m tall, d. b. h. 2 - 3 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate glabrous. Leaves: petiole 4 (5) mm long, ca. 2 mm in diameter, glabrous, blade inserted on top of the petiole; blade 17 - 20 (- 23) cm long, 6 - 8 cm wide, oblanceolate, apex acuminate, acumen 1 - 1.5 cm long, broadly acute but minutely cordate, coriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken, above glabrous when young and old, below glabrous when young and old; secondary veins 5 to 8 (9) pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual [?, female flowers unknown], monoecious [?]; male inflorescences cauliflorous, scattered along the trunk. Male flowers with 6 perianth parts in 2 whorls, narrowly ovoid to pyramidal in bud; 1 to 7 per inflorescence borne on a short peduncle; pedicel ca. 18 - 25 mm long, ca. 1 mm in diameter, glabrescent to glabrous; bracts (1) 2, one basal and one upper towards the lower half of pedicel, similar in size, ca. 1.5 mm long, ca. 0.5 mm wide; sepals 2, valvate, free, ca. 1 - 1.5 mm long, ca. 2 mm wide, semi-orbicular, apex acute, base truncate, glabrous, margins flat; petals free, 4 in one whorl, 14 - 16 mm long, ca. 5 - 9 mm wide, lanceolate-oblong, apex rounded, base truncate, yellow-green, margins flat, sparsely pubescent outside, glabrous inside; stamens ca. 300 in ca. 10 whorls, ca. 0.5 mm long; connective absent; staminodes absent. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
D22740D106DD762A15AD08A2D9B5CF9E.taxon	distribution	Distribution. Endemic to Cameroon; known from the Littoral region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A8903D6B3391BA6F27976E5C3EC1CE73.taxon	description	Figs 127, 133; Map 16 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A8903D6B3391BA6F27976E5C3EC1CE73.taxon	materials_examined	Type. Cameroon. Littoral Region; Ed Winkler H. 909, Nov 1904: lectotype, designated here: B [B 10 0153121].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A8903D6B3391BA6F27976E5C3EC1CE73.taxon	description	Description. Tree to shrub, 6 - 20 m tall, d. b. h. 14 - 40 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches pubescent to glabrous. Leaves: petiole 2 - 5 mm long, 1 - 2.5 mm in diameter, sparsely pubescent to glabrous, narrowly grooved, blade inserted on top of the petiole; blade 11.1 - 24.5 cm long, 3.8 - 9.2 cm wide, elliptic to obovate, apex attenuate to acuminate, acumen 0.7 - 3 cm long, base rounded to acute to decurrent, coriaceous, below sparsely pubescent to glabrous when young, glabrous when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous to pubescent when young, glabrescent when old; secondary veins 6 to 14 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious, dimorphic. Flowers with 6 perianth parts in 2 whorls. Male inflorescences cauliflorous, 5 to 20 - flowered or more, on thick clumps on the lower part of the trunk; male flowers: pedicel 11 - 50 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous; bracts 2, one basal and one upper towards the middle or lower half of pedicel, basal bract minute, ca. 1 mm long, ca. 1 mm wide; upper bract 0.5 - 1.5 mm long, 1 - 2 mm wide; sepals 2, valvate, basally fused, 1.5 - 4.5 mm long, 2.5 - 11 mm wide, circular to broadly ovate, apex acute or rounded, base truncate, purple, pubescent to sparsely pubescent outside, glabrous inside, margins flat; petals free, 4 in one whorl, 6 - 11 mm long, 3.5 - 8 mm wide, ovate to elliptic, apex acute, base truncate, wine red to dark brownish-red, margins flat, pubescent to glabrous outside, glabrous inside; stamens ca. 300, in 20 to 25 rows on a convex receptacle, 0.4 - 0.5 mm long; connective minute, glabrous; staminodes absent. Female inflorescences cauliflorous, axillary, clumps of 6 to 10 flowers or more, on thick clumps on the lower part of the trunk; female flowers: pedicel 10 - 50 mm long; 2 - 3 mm in diameter, sparsely pubescent to glabrous; in fruit 8 - 55 mm long, 2 - 4 mm in diameter, sparsely pubescent to glabrous; bracts 2, one basal and one upper towards the middle or lower half of pedicel, basal bract minute, ca. 1 mm long, ca. 1 mm wide; upper bract 0.5 - 1.5 mm long, 1 - 2 mm wide; sepals 2, valvate, basally fused, 2 - 5 mm long, 2 - 11 mm wide, circular to broadly ovate, apex acute or rounded, base truncate, purple, pubescent to sparsely pubescent outside, glabrous inside, margins flat; petals free, 4, in one whorl, 7 - 20 mm long, 4 - 15 mm wide, ovate to elliptic, apex acute, base truncate, wine red to dark red-brownish outside, margins flat, pubescent to glabrous outside, glabrous inside; carpels free, 100 to 280, ovary 1.5 - 2.5 mm long, stigma coiled bilobed, glabrous. Monocarps stipitate, stipes 0 - 2 mm long, ca. 4 mm in diameter; monocarps 2 to 5, 21 - 60 mm long, 30 - 35 mm in diameter, ovoid to cylindrical, apex rounded, sparsely pubescent to glabrous, smooth, not ribbed, pale brownish grey to yellow to red; seeds 5 to 8 per monocarp, 15 - 22 mm long, ca. 15 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A8903D6B3391BA6F27976E5C3EC1CE73.taxon	distribution	Distribution. A central African species, from Nigeria to Republic of the Congo; in Cameroon known from the Central, East, Littoral, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
A8903D6B3391BA6F27976E5C3EC1CE73.taxon	materials_examined	Specimens examined. Central Region: Reserve forestiere de Makak au bord du Nyong, 3.59 ° N, 11.03 ° E, 14 December 1967, Bamps P. R. J. 1458 (P, YA); Ndiki, 4.77 ° N, 10.83 ° E, 01 November 1938, Jacques-Felix H. 2493 (P); Foret de Mambe pres Boga (30 km N Eseka), 3.9 ° N, 10.78 ° E, 08 December 1973, Letouzey R. 12290 (K, P, YA); Mfiki (Ndo par Esse), 4.31 ° N, 11.96 ° E, 09 November 1969, Letouzey R. 9541 (P, WAG); Etwa 115 km NO Juande, 4.48 ° N, 12.35 ° E, 01 January 1914, Mildbraed G. W. J. 8260 (K). East Region: Reserve de faune du Dja Djolimpoun, 3.17 ° N, 13.18 ° E, 17 April 1995, Sonke B. 1505 (BR, YA). Littoral Region: Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 19 June 2014, Couvreur T. L. P. 654 (WAG, YA); Mambe Massif above Boga village 100 km along road from Yaounde to Ed 3.90 ° N, 10.77 ° E, 20 June 2014, Couvreur T. L. P. 659 (WAG, YA); Au sud de Ngola (8 km Est de l'embouchur de la Sanaga), 3.55 ° N, 9.698 ° E, 05 January 1974, Letouzey R. 12580 (P, YA); Ed 3.81 ° N, 10.13 ° E, 01 January 1904, Winkler H. 909 (B). South Region: N'koladom village 4 km on the road (old road) from Nkoemvone to Ambam, 2.8 ° N, 11.15 ° E, 27 November 1974, de Wilde J. J. F. E 7754 (K, P, WAG); Nkoemvone, 2.81 ° N, 11.13 ° E, 05 June 1975, de Wilde J. J. F. E 8270 a (BR, MO, P, WAG); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.8 ° N, 11.13 ° E, 12 December 1975, de Wilde J. J. F. E 8709 (BR, MO, P, WAG, YA); Nkomo pres Ngoase au S de la rive Lobe, 3.26 ° N, 12.02 ° E, 13 February 1962, Letouzey R. 4219 (K, P, YA); Nkomo pres Ngoase au S de la rive Lobe, 3.26 ° N, 12.02 ° E, 14 February 1962, Letouzey R. 4230 (BR, P, YA); Pres Mevous 50 km SE d'Ebolowa sur piste d'Evindissi, 2.6 ° N, 11.46 ° E, 30 January 1970, Letouzey R. 9934 (BR, K, P, YA). South-West Region: ca 40 minutes walk N then E from Njonji Hunters path to Lake Njonji, 4.13 ° N, 8.993 ° E, 18 November 1993, Cheek M. 5482 (K, YA); ca 40 minutes walk N then E from Njonji Hunters path to Lake Njonji, 4.13 ° N, 8.993 ° E, 19 November 1993, Cheek M. 5501 (K, MO, WAG, YA); Entre Ayong et Baro 20 km SW Nguti, 5.2 ° N, 9.32 ° E, 10 June 1975, Letouzey R. 13790 (P, YA); Bibundi, 4.21 ° N, 8.988 ° E, 01 November 1928, Mildbraed G. W. J. 10647 (K); Pente SW Mt Cameroun ME Bakingili WNW Limbe, 4.07 ° N, 9.04 ° E, 09 December 1984, Villiers J. - F. 2429 (P, YA); Limbe W of Njonji Lake, 4.13 ° N, 9.016 ° E, 27 January 1994, Wieringa J. J. 2029 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	description	Map 16 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	diagnosis	Diagnosis. Uvariopsis etugeana resembles U. peduculosa in being a small tree and in the shape of its leaves, but differs by having a short flowering pedicel (4 - 10 mm versus 25 - 130 mm in U. pedunculosa) and glabrous petals on the outside (versus pubescent in both U. pedunculosa).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	materials_examined	Type. Cameroon. North-West Region; Wum, Letouzey R. 13414, 3 Dec 1974: holotype: P [P 01982826].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	description	Description. Tree, 3 - 6 m tall, d. b. h. unknown; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches slightly pubescent to glabrous. Leaves: petiole 3.5 - 4 mm long, 2.5 - 3.5 mm in diameter, glabrous, blade inserted on top of the petiole; blade 19 - 27 cm long, 7 - 9.5 cm wide, elliptic, apex attenuate to acuminate, acumen 1 - 2 cm long, base acute to slightly decurrent, coriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 10 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious, dimorphic; inflorescences cauliflorous and ramiflorous on leafless branches, axillary. Flowers with 6 perianth parts in 2 whorls; 1 to 3 per inflorescence. Male flowers: pedicel ca. 8 mm long, 1.5 mm in diameter, glabrescent to glabrous; bracts 2, one basal and one upper towards the middle or lower half of pedicel, basal bract minute, ca. 1 mm long, ca. 1 mm wide; upper bract 1 - 1.5 mm long, 1 - 2.5 mm wide; sepals 2, valvate, basally fused, ca. 2 mm long, ca 4. mm wide, broadly ovate, apex acute or rounded, base truncate, sparsely pubescent to glabrous outside, glabrous inside, margins flat; petals free, 4 in one whorl, ca. 10 mm long, ca. 4 mm wide, elliptic to ovate, apex acute, base truncate, color unknown, margins flat, glabrous outside, glabrous inside; stamens numerous (exact number unknown), ca. 0.5 mm long; connective minute, glabrous; staminodes absent. Female flowers: pedicel ca. 4 mm long, ca. 2 mm in diameter, glabrescent to glabrous; in fruit unknown; bracts 2, one basal and one upper towards the middle or lower half of pedicel, basal bract minute, ca. 1 mm long, ca. 1 mm wide; upper bract ca. 1.5 mm long, ca. 2 mm wide; sepals 2, valvate, basally fused, 1.5 mm long, 3.5 mm wide, broadly ovate, apex acute or rounded, base truncate, sparsely pubescent to glabrous outside, glabrous inside, margins flat; petals free, 4, in one whorl, ca. 14 mm long, ca. 8 mm wide, elliptic, apex acute, base truncate, color unknown, margins flat, glabrous outside, glabrous inside; sterile stamens, ca. 5, ca. 1 mm long; carpels free, ca. 20, ovary ca. 3 mm long, ca. 1 mm wide, stigma unknown, glabrescent at base to glabrous. Fruits unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	distribution	Distribution. Endemic to Cameroon, only known from two localities in North-West and South-West Regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
9D8ADADE586B29C9324B5ECBF52A6755.taxon	etymology	Etymology. Named in honor of the late Martin Etuge Ekwoge (1966 - 2020), a passionate Cameroonian horticulturalist, botanist and parataxonomist from Nyassosso village, South-West Cameroon. He was one of the main collectors for the ' Plants of Mount Mwanenguba and the Bakossi Mountains' (Cheek et al. 2004). For a total of 14,538 specimens recorded in the Kew database for Bakossi, 3,170 were collected by him (Cheek et al. 2020) including over 652 (that we have seen) specimens of Annonaceae (representing 28 species).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0BEC0FB34D97D6CE7E64DAC219DE484E.taxon	description	Figs 134, 135; Map 16 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0BEC0FB34D97D6CE7E64DAC219DE484E.taxon	materials_examined	Type. Cameroon. South Region; Korup National Park, Chimpanzee Camp, 5 ° 04 ' N, 8 ° 52 ' E, Kenfack D. 1026, 3 Fev 1998: holotype: YA; isotypes: MO [MO- 022919]; P [P 01817719]; WAG [WAG 0358388].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0BEC0FB34D97D6CE7E64DAC219DE484E.taxon	description	Description. Tree to shrub, 6 - 15 m tall, d. b. h. 12 - 14 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to pubescent. Leaves: petiole 2 - 7 mm long, 2 - 6 mm in diameter, glabrous to pubescent, slightly grooved, blade inserted on top of the petiole; blade 30 - 61.5 cm long, 5.9 - 16.5 cm wide, elliptic to obovate, apex abruptly acuminate, acumen 1.8 - 3.2 cm long, base rounded to cordate, coriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 10 to 26 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous, towards base and up to 3 m on trunk. Flowers with 6 perianth parts in 2 whorls, ovoid to conical in bud, 2 to 3 per inflorescence, male and female inflorescences dimorphic; male pedicels 6 - 35 mm long, 1 - 2 mm in diameter, tomentose; female pedicels 16 - 50 mm long, 1 - 2 mm in diameter, tomentose; in fruit 45 - 90 mm long, 2 mm in diameter, glabrous; bracts 2 to 4, all basal, 1 - 2 mm long, ca. 1 mm wide; male sepals 2, valvate, basally fused, 2 - 7.5 mm long, 3 - 6.5 mm wide, ovate, apex acute, base truncate, greenish, pubescent outside, glabrous inside, margins flat; female sepals 2, valvate, basally fused, 1.5 - 5 mm long, 1.5 - 5 mm wide, ovate, apex acute, base truncate, greenish, pubescent outside, glabrous inside, margins flat; male petals 4, basally fused, tube 1 - 2 mm long, 10 - 38 mm long, 5 - 10 mm wide, elliptic to ovate, apex acute to attenuate, base truncate, greenish-yellow, margins flat, pubescent outside, glabrous inside; female petals 4, basally fused, tube 3 - 5 mm long, 15 - 35 mm long, 8 - 12 mm wide, elliptic, apex acute to attenuate, base truncate, greenish-yellow, margins flat, pubescent outside, glabrous inside; male flowers: stamens 130 to 210, in 9 to 12 rows, 0.5 - 1 mm long, oblong; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 25 to 40, ovary 3 - 4 mm long, stigma flat, glabrous. Monocarps stipitate, stipes 2 - 5 mm long, 2 mm in diameter; monocarps 5 to 9, 30 - 60 mm long, 18 - 30 mm in diameter, ellipsoid to cylindrical, apex rounded, glabrous, smooth, clear orang yellow when ripe; seeds 8 to 14 per monocarp, 10 - 22 mm long, 5 - 14 mm in diameter, ellipsoid to oblong; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0BEC0FB34D97D6CE7E64DAC219DE484E.taxon	distribution	Distribution. endemic to Cameroon; known from the South, South-West and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0BEC0FB34D97D6CE7E64DAC219DE484E.taxon	materials_examined	Specimens examined. Littoral Region: Nkam Yingui Ekem River bank Forest, 4.35 ° N, 10.41 ° E, 05 March 2002, Kenfack D. 1620 (MO). South Region: Campo-Ma'an area Bifa, 2.67 ° N, 10.28 ° E, 13 October 2001, Tchouto Mbatchou G. P. BIFAX _ 25 (WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1911, Zenker G. A. 3971 (K). South-West Region: ca 40 minutes walk N then E from Njonji Hunters path to Lake Njonji, 4.13 ° N, 8.993 ° E, 19 November 1993, Cheek M. 5486 (K, WAG, YA); Ekundu Kundu, 5.15 ° N, 8.883 ° E, 28 April 1996, Cheek M. 8258 (K, WAG, YA); Ekundu kundu, 5.15 ° N, 8.89 ° E, 09 January 1998, Cheek M. 8815 (K, YA); Mount Cameroon National Park Bakinguili trail above Bakinguili village, 4.09 ° N, 9.056 ° E, 02 April 2016, Couvreur T. L. P. 1039 (WAG, YA); Mount Cameroon National Park on the Bomona trail behind Bomona village 10 km NW from Idenau, 4.29 ° N, 9.078 ° E, 03 April 2016, Couvreur T. L. P. 1052 (WAG, YA); Korup National Park, 4.95 ° N, 8.87 ° E, 05 March 1993, Gereau R. E. 5192 (MO, P, WAG); Korup National Park, 5.06 ° N, 8.866 ° E, 03 February 1998, Kenfack D. 1026 (MO, P, WAG, YA); Bakingili, 4.08 ° N, 9.05 ° E, 16 February 1997, Nning J. 284 (K, YA); Bomana, 4.25 ° N, 9.016 ° E, 05 October 1993, Tchouto Mbatchou G. P. 675 (K, YA); Korup National Park, 5.05 ° N, 8.8 ° E, 28 April 1984, Thomas D. W. 3182 (L, P, YA); Korup National Park, 5.05 ° N, 8.8 ° E, 28 February 1984, Thomas D. W. 3210 (BR, YA); Mount Cameroun above small Koto village, 4.3 ° N, 9.1 ° E, 06 March 1985, Thomas D. W. 4477 (MO, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
501D815D9D6500C3CDA1D43FF0FB30B4.taxon	description	Figs 136, 137; Map 16 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
501D815D9D6500C3CDA1D43FF0FB30B4.taxon	materials_examined	Type. Cameroon. South Region, Bipindi, Zenker G. A. 3868, Mar 1906: B [B 10 0153122]; isotypes: BM [BM 000554078]; BR [BR 0000008824196, BR 0000008824226]; HBG [HBG 502486]; K [K 000199041]; M [M 0107937]; P [P 00362599, P 00362601]; US [US 00098850].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
501D815D9D6500C3CDA1D43FF0FB30B4.taxon	description	Description. Shrub to tree, 3 - 8 m tall, d. b. h. 1.5 - 5 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to pubescent. Leaves: petiole 2 - 3 mm long, 1 - 3 mm in diameter, pubescent, slightly grooved, blade inserted on the side of the petiole; blade 17.2 - 29 cm long, 5.7 - 11 cm wide, elliptic to obovate, apex acuminate, acumen 0.3 - 2.3 cm long, base acute, coriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below glabrous when young and old; secondary veins 8 to 15 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous. Flowers with 9 perianth parts in 2 whorls, globose in bud, 1 to 5 per inflorescence, male and female inflorescences dimorphic; male pedicels 25 - 130 mm, 1 - 2 mm in diameter, sparsely pubescent to glabrous; bracts 2, one basal and one upper towards the lower half of pedicel, both bracts 1.5 mm long, ca. 2 mm wide; female pedicels 80 - 320 mm long, 2 - 4 mm in diameter; in fruit mm long, mm in diameter; bracts 2, one basal and one upper towards the lower half of pedicel, both bracts 2.5 - 6 mm long, 5 - 6 mm wide; male sepals 2, valvate, basally fused, 7 mm long, 7 mm wide, very broadly ovate, apex acute, base truncate, brown, pubescent outside, glabrous inside, margins flat; female sepals 2, valvate, basally fused, 7 - 17 mm long, 13 - 15 mm wide, very broadly ovate, apex acute, base truncate, fleshy, brown, pubescent outside, glabrous inside, margins flat; male petals 4, basally fused, tube ca. 3 mm long, 10 - 15 mm long, 8 - 10 mm wide, broadly ovate to broadly elliptic, apex acute, base truncate, wine-brown outside, white cream inside, margins flat, shortly tomentose outside, glabrous inside; female petals 4, basally fused, tube 2.5 - 3 mm long, 14 - 16 mm long, 8 - 12 mm wide, broadly ovate, apex acute, base truncate, fleshy, wine-brown outside, white cream inside, margins flat, pubescent outside, glabrous inside; male flowers: stamens 150 to 250, in 10 to 12 rows, ca. 1 mm long, oblong; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 50 to 140, ovary 2 - 4 mm long, stigma globose, glabrous. Monocarps sessile, 4 to 9, 10 - 17 mm long, 7 - 11 mm in diameter, cylindrical, apex rounded, sparsely pubescent, verrucose, longitudinally 1 ribbed, brown when ripe; seeds 6 to 10 per monocarp, 7 - 10 mm long, 5 - 7 mm in diameter; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
501D815D9D6500C3CDA1D43FF0FB30B4.taxon	distribution	Distribution. A central African species from Cameroon, Equatorial Guinea Gabon and Democratic Republic of the Congo; in Cameroon known from South, South-West and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
501D815D9D6500C3CDA1D43FF0FB30B4.taxon	materials_examined	Specimens examined. Littoral Region: Mapubi 30 km before Edea on Yaounde-Edea road On forestry road 5 km direction to Sanaga river, 3.84 ° N, 10.38 ° E, 28 February 2018, Couvreur T. L. P. 1173 (MPU, P, WAG, YA). South Region: Elephant Mont, 2.79 ° N, 10.02 ° E, 16 October 2001, Tchouto Mbatchou G. P. ELEX _ 15 (WAG); Campo-Ma'an area Onoyong, 2.52 ° N, 10.69 ° E, 18 March 2001, Tchouto Mbatchou G. P. ONOX _ 182 (WAG); Campo-Ma'an area Onoyong, 2.52 ° N, 10.69 ° E, 18 March 2001, Tchouto Mbatchou G. P. ONOX _ 274 (WAG); Campo-Ma'an area Mamelles Massif, 2.56 ° N, 9.949 ° E, 23 April 2001, Tchouto Mbatchou G. P. 3242 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1909, Zenker G. A. 3868 (B, BM, BR, K, L, P). South-West Region: on forest trail north of Ngomboku village, 4.91 ° N, 9.724 ° E, 06 April 2016, Couvreur T. L. P. 1063 (WAG, YA); on forest trail north of Ngomboku village, 4.91 ° N, 9.730 ° E, 06 April 2016, Couvreur T. L. P. 1066 (WAG, YA); on trail through palm oil plantation 3 km before lava flow and Seme Beach hotel when coming from Limbe, 4.07 ° N, 9.085 ° E, 18 October 2013, Couvreur T. L. P. 517 (WAG, YA); Ajang saprophyte plot, 4.76 ° N, 9.683 ° E, 01 December 1999, Gosline W. G. 244 (K, MO, P, WAG, YA); Crete du Nta Ali (1266 m) entre cotes 1009 et 1202 30 km SE Mamfe, 5.59 ° N, 9.502 ° E, 19 June 1975, Letouzey R. 13849 (P, YA); Likomba-Pflanzung 15 - 35 km NE von Victoria [Limbe], 4.1 ° N, 9.333 ° E, 03 December 1928, Mildbraed G. W. J. 10745 (B, K); Environs de Matene RF Takamanda 32 km SW Akwaya, 6.25 ° N, 9.372 ° E, 30 April 1987, Satabie B. 873 (YA); Forest on the southern slope of Mount above Batoke, 4.08 ° N, 9.1 ° E, 29 December 1983, Thomas D. W. 2756 (MO, YA); Takamanda Forest Reserve, 6.21 ° N, 9.433 ° E, 30 April 1987, Thomas D. W. 7364 (P); Massif du Ntali crete sommitale 30 km SE Mamfe, 5.58 ° N, 9.510 ° E, 14 June 1982, Villiers J. - F. 1427 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
559C18F743EA2C280A9617AE034343E9.taxon	description	Map 16 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
559C18F743EA2C280A9617AE034343E9.taxon	materials_examined	Type. Cameroon. East Region; Bezirk Lomie, im grossen Dscha-Bogen, Mildbraed G. W. J. 5239, 1911: holotype: B [B 10 015312]; isotype: HBG [HBG 502485].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
559C18F743EA2C280A9617AE034343E9.taxon	description	Description. Tree, height unknown, d. b. h. unknown; stilt roots or buttresses not observed. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 3 mm long, 1 - 2 mm in diameter, glabrous, slightly grooved, blade inserted on top of the petiole; blade 12 - 18 cm long, 4 - 6 cm wide, elliptic to obovate, apex acute, acumen 0.5 - 0.8 cm long, base acute to obtuse, papyraceous, below densely pubescent when young, densely pubescent when old, above glabrous when young and old; midrib sunken or flat, above densely pubescent when young, glabrous when old, below densely pubescent when young, glabrous when old; secondary veins 10 to 11 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual [?, only female flowers known], monoecious [?]; inflorescences cauliflorous. Flowers with 6 perianth parts in 2 whorls, globose in bud, 2 to 3 per inflorescence, female pedicels 1 - 3 mm long, 1 - 2 mm in diameter, pubescent; in fruit unknown, bracts 2 to 3, all basal or towards the lower half of pedicel, ca. 1 mm long, ca. 1 mm wide; female sepals 2, valvate, free, 2 - 4 mm long, 2 - 4 mm wide, ovate, apex acute, base truncate, pubescent outside, glabrous inside, margins flat; female petals 4, free, 7 - 16 mm long, 6 - 8 mm wide, ovate, apex acute, base truncate, margins flat, pubescent outside, sparsely pubescent and sericeous towards margins inside; male flowers unknown; staminodes unknown; female flowers: carpels free, number unknown, ovary 2 - 3 mm long, stigma unknown,. Monocarps stipitate, 1 seen, ca. 18 mm long, ca. 13 mm in diameter, globose, apex rounded, glabrous, smooth, color unknown; seeds 3 to 4 per monocarp, ca. 8 mm long, ca. 6 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
559C18F743EA2C280A9617AE034343E9.taxon	distribution	Distribution. endemic to Cameroon, known from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6156AEE23F52AC6480593CFAF4228A1F.taxon	description	Figs 135, 137; Map 16 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6156AEE23F52AC6480593CFAF4228A1F.taxon	materials_examined	Type. Type. Democratic Republic of the Congo. Orientale; Yambuya, Solheid A. F. 96, 1906: holotype: BR [BR 0000008824240).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6156AEE23F52AC6480593CFAF4228A1F.taxon	description	Description. Shrub to tree, 3 - 8 m tall, d. b. h. 1.5 - 8 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous to pubescent, young foliate branches tomentose to pubescent. Leaves: petiole 3 - 4 mm long, 1 - 2 mm in diameter, tomentose, slightly grooved, blade inserted on top of the petiole; blade 16.6 - 29 cm long, 5 - 9.5 cm wide, elliptic to obovate, apex acuminate to attenuate, acumen 0.4 - 2 cm long, base rounded to subcordate, papyraceous, below glabrous when young, sparsely pubescent when old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 8 to 13 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous. Flowers with 6 perianth parts in 2 whorls, long conical in bud, 2 to 3 per inflorescence, male and female inflorescences dimorphic; male pedicels up to 22 mm long, 0.5 - 1 mm in diameter, densely pubescent; female pedicels 80 - 160 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 53 - 165 mm long, ca. 2 mm in diameter; bracts 2, one basal and one upper towards the lower half of pedicel, basal bract ca. 1 mm long, ca. 1 mm wide; upper bracts 1 - 2 mm long, 1.5 - 3 mm wide, soon falling; male sepals 2, valvate, free, ca. 2 mm long, 2 - 3 mm wide, ovate, apex acute, base truncate, brown-red, pubescent outside, glabrous inside, margins flat; female sepals 2, valvate, free, 2.5 - 4 mm long, 3 - 5 mm wide, ovate to elliptic, apex acute, base truncate, brown-red, pubescent outside, glabrous inside, margins flat; male petals 4 (sometimes 5), free, 7 - 10 mm long, 2.5 - 4 mm wide, narrowly ovate to narrowly elliptic, apex acute, base truncate, wine-brown, margins flat, pubescent outside, glabrous and finely warty inside; female petals 4, free, 12 - 17 mm long, 4 - 7 mm wide, narrowly ovate to narrowly elliptic, apex acute, base truncate, wine-brown, margins flat, tomentose outside, glabrous and finely warty inside; male flowers: stamens 150 to 200, in 9 to 15 rows, ca. 0.5 mm long, oblong; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 40 to 65, ovary 1.5 - 3.5 mm long, stigma coiled, densely pubescent. Monocarps stipitate, stipes 2 - 4 mm long, ca. 3 mm in diameter; monocarps 5 to 10, 30 - 70 mm long, 13 - 30 mm in diameter, ellipsoid to cylindrical, apex rounded to apiculate, sparsely pubescent to glabrous, longitudinally 4 to 6 ribbed, smooth, red when ripe; seeds 2 to 10 per monocarp, ca. 15 mm long, 8 - 13 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6156AEE23F52AC6480593CFAF4228A1F.taxon	distribution	Distribution. A central African species, from Cameroon to Gabon and Democratic Republic of the Congo; in Cameroon known from the South and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6156AEE23F52AC6480593CFAF4228A1F.taxon	materials_examined	Specimens examined. Central Region: Ndanan 1, 3.62 ° N, 11.58 ° E, 10 March 2004, Cheek M. 11606 (K, WAG, YA); Mont Ngoro a 38 km au Nord de Bafia (pied mont), 5.09 ° N, 11.26 ° E, 29 April 1978, Ngameni B. K. 109 (P, YA); Mont Ngoro a 58 km SW de Linte, 5.09 ° N, 11.26 ° E, 17 April 1982, Nkongmeneck B. A. 273 (P, YA); River N Didoumou, 3.61 ° N, 11.56 ° E, 26 March 2004, Onana J. M. 2848 (K). South Region: Zingui, 2.85 ° N, 10.98 ° E, 21 July 1975, de Wilde J. J. F. E 8373 (WAG); Ngongonjie (Akonetye), 2.67 ° N, 12.87 ° E, 30 August 1978, Koufani A. 154 (P); Efoulan, 2.74 ° N, 10.54 ° E, 06 December 2000, Tchouto Mbatchou G. P. ONOX _ 191 (WAG); Efoulan, 2.74 ° N, 10.53 ° E, 25 April 2000, Tchouto Mbatchou G. P. 2843 (KRIBI, WAG); Efoulan, 2.74 ° N, 10.54 ° E, 04 December 2000, Tchouto Mbatchou G. P. 3089 (KRIBI, WAG, YA); Campo-Ma'an area Akom II, 2.8 ° N, 10.53 ° E, 19 August 2001, van Andel T. R. 3951 (MO); Colline de Nkoltsia a 23 km NW de Bipindi, 3.17 ° N, 10.27 ° E, 27 April 1974, Villiers J. - F. 893 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
18EFD8C845B0986504A16BB9E096C1F5.taxon	description	Figs 138, 139; Map 16 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
18EFD8C845B0986504A16BB9E096C1F5.taxon	materials_examined	Type. Cameroon. South-West Region; Rumpi Hills, Kenfack D. 1334, 6 Feb 2000: holotype: YA; isotypes: K [000683145]; MO; SCA.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
18EFD8C845B0986504A16BB9E096C1F5.taxon	description	Description. Tree, 8 - 25 m tall, d. b. h. up to 30 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches glabrous to pubescent. Leaves: petiole 3 - 8 mm long, 3 - 5 mm in diameter, glabrous to sparsely pubescent, grooved, blade inserted on top of the petiole; blade 16 - 38 cm long, 5 - 11 cm wide, oblong to obovate, apex attenuate to acuminate, acumen 2 - 3 cm long, base rounded to subcordate, papyraceous to subcoriaceous, below glabrous when young and old, above glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below sparsely pubescent when young, glabrous when old; secondary veins 9 to 18 pairs per side, glabrous above; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous, with hundreds (up to 500) of flowers packed at the base and then more sparsely distributed up to 6 m. Flowers unisexual, monoecious, with 6 perianth parts in 2 whorls, conical to pyramidal in bud, 6 to 50 per inflorescence, male and female inflorescences dimorphic; male pedicel: 25 - 50 mm, ca. 1 mm in diameter, pubescent; female pedicel: 30 - 60 mm long, ca. 1 mm in diameter, pubescent; in fruit 25 - 90 mm long, 3 - 7 mm in diameter, glabrous; bracts 2 to 4, all basal, ca. 1 mm long, ca. 1 mm wide; male sepals 2, connate, splitting at maturity, 5 - 10 mm long, 6 - 12 mm wide, triangular, apex acute, base truncate, brown-red, pubescent outside, glabrous inside, margins flat; female sepals 2, connate, splitting at maturity, 6 - 8 mm long, 6 - 9 mm wide, broadly ovate, apex acute, base truncate, brown-red, pubescent outside, glabrous inside, margins flat; male petals, 4, basally fused, tube 4 - 9 mm long, 8 - 18 mm long, 5 - 8 mm wide, elliptic to narrowly elliptic, apex acute, base truncate, wine red, margins flat, tomentose outside, pubescent inside; female petals, 4, basally fused, tube 3 - 4 mm long, 15 - 17 mm long, 5 - 7 mm wide, elliptic to narrowly elliptic, apex acute, base truncate, wine red, margins flat, tomentose outside, pubescent inside; male flowers: stamens 700 to 1000, in 25 to 30 rows, 0.5 - 1 mm long, oblong; connective reduced or absent, glabrous, cream; staminodes absent; female flowers: carpels free, 60 to 100, ovary 15 - 24 mm long, stigma globose, pubescent. Monocarps shortly stipitate, stipes ca. 1 mm long, ca. 1 mm in diameter; monocarps 9 to 25, 17 - 80 mm long, 13 - 55 mm in diameter, ovoid to oblong, apex apiculate, sparsely pubescent to glabrous, smooth, slightly constricted over seeds in dried material, pale green turning dark yellow at maturity; seeds 6 to 12 per monocarp, 18 - 25 mm long, 8 - 13 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
18EFD8C845B0986504A16BB9E096C1F5.taxon	distribution	Distribution. endemic to Cameroon, known from the South, Littoral and South West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
18EFD8C845B0986504A16BB9E096C1F5.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Wildlife Reserve Djuma permanent camp On east trail, 4.36 ° N, 10.25 ° E, 15 February 2013, Couvreur T. L. P. 627 (WAG, YA); Ebo Bekob abandoned village CRES research station, 4.37 ° N, 10.41 ° E, 22 April 2005, Etuge M. 6482 (K); Nkam Yingui Bataba, 4.53 ° N, 10.24 ° E, 20 February 2002, Kenfack D. 1602 (MO). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 682 (WAG, YA). South-West Region: Nyasoso, 4.81 ° N, 9.7 ° E, 08 February 1995, Cable S. 1221 (K, MO, WAG, YA); Kupe village, 4.78 ° N, 9.716 ° E, 30 May 1996, Cable S. 2736 (K, WAG, YA); Esense river near farm of Philip Taza, 4.76 ° N, 9.683 ° E, 19 January 1995, Cheek M. 7034 (K, WAG, YA); Kupe village, 4.76 ° N, 9.683 ° E, 24 January 1995, Cheek M. 7131 (K, WAG); Nyasoso village on max's trail to Mt 4.82 ° N, 9.692 ° E, 05 April 2016, Couvreur T. L. P. 1059 (WAG, YA); Rumpi mountains forest trail ca 5 km after Dikome Balue village ca 40 km north of Kumba, 4.93 ° N, 9.241 ° E, 10 January 2016, Couvreur T. L. P. 965 (WAG, YA); Below Kupe rock near Esense river, 4.78 ° N, 9.683 ° E, 23 January 1995, Elad M. 69 (K, YA); South of Nyasoso (end of village) Trail at the end of the village left hand side Nyasoso-Mbulle road, 4.81 ° N, 9.683 ° E, 03 July 1996, Etuge M. 2562 (K, YA); Rumpi Hills, 4.95 ° N, 9.033 ° E, 06 February 2000, Kenfack D. 1334 (K, MO, YA); Forest trail 2 km south from Etube-Tape village, 4.85 ° N, 9.7 ° E, 01 February 1995, Lane P. 490 (K, WAG, YA); Nyasoso, 4.82 ° N, 9.666 ° E, 23 October 1995, Sebsebe D. 5035 (K, YA); 7 km WNW of Bomana 34 km NW of Limbe, 4.27 ° N, 9.112 ° E, 15 December 1984, Villiers J. - F. 2490 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E9D01316E8120A0137B5EE3B67364E0F.taxon	description	Figs 139, 140; Map 17 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E9D01316E8120A0137B5EE3B67364E0F.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 1117, 1896: holotype: B [B 10 0153124]; isotypes: BM [BM 000554075]; GOET [GOET 005734]; HBG [HBG 502515]; P [P 00362604, P 00362603]; K [K 000199042]; M [M 0107936]; S [S 07 - 11029]; WU [WU 0025790]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E9D01316E8120A0137B5EE3B67364E0F.taxon	description	Description. Shrub to tree, 2 - 7 m tall, d. b. h. 1.5 - 11 cm; stilt roots or buttresses absent. Indumentum of simple hairs; old leafless branches glabrous, young foliate branches densely pubescent. Leaves: petiole 2 - 5.5 mm long, 1 - 2 mm in diameter, densely to sparsely pubescent, grooved, blade inserted on top of the petiole; blade 11 - 15.8 cm long, 3.5 - 5.8 cm wide, elliptic to obovate, apex long acuminate, acumen 1 - 2.4 cm long, base acute to decurrent, papyraceous, above glabrous when young and old, below glabrous when young and old; midrib sunken or flat, above glabrous when young and old, below pubescent when young, glabrous when old; secondary veins 6 to 13 pairs, glabrous above and below; tertiary venation reticulate. Individuals unisexual, monoecious; inflorescences cauliflorous or ramiflorous on old leafless branches, axillary. Flowers with 6 perianth parts in 2 whorls, globose in bud, 1 per inflorescence, male and female inflorescences similar; pedicel 0 - 7 mm long, 1 - 1.5 mm in diameter, densely pubescent; in fruit ca. 10 mm long, 2 - 3 mm in diameter, glabrous; bracts 1 to 2, all basal 1 - 4 mm long, 2.5 - 5 mm wide; sepals 2, valvate, basally fused, 3 - 7 mm long, 4.5 - 7 mm wide, circular, apex obtuse, base truncate, brown, densely pubescent outside, glabrous inside, margins flat; petals 4, basally fused, tube 1 - 4 mm long, 6 - 13 mm long, 4 - 7 mm wide, triangular, apex acute, base truncate, greenish-white, margins flat, densely pubescent outside, glabrous inside; male flowers: stamens 100 to 150, in 10 to 12 rows, 0.5 mm long, broad; connective reduced or absent, glabrous, pinkish red; staminodes absent; female flowers: carpels free, 13 to 22, ovary 1.5 - 3 mm long, 1 - 1.5 wide, stigma coiled, densely pubescent. Monocarps stipitate, stipes ca. 1 mm long, ca. 2 mm in diameter; monocarps 1 to 3, ca. 37 mm long, ca. 17 mm in diameter, cylindrical, apex rounded, tomentose, bumpy when dry, green when ripe; seeds 10 to 12 per monocarp, ca. 13 mm long, ca. 7 mm in diameter, ellipsoid; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E9D01316E8120A0137B5EE3B67364E0F.taxon	distribution	Distribution. endemic to Cameroon; known from the South, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E9D01316E8120A0137B5EE3B67364E0F.taxon	materials_examined	Specimens examined. Littoral Region: Ebo Forest proposed National Park Ebo Forest Research Station-Bekob, 4.36 ° N, 10.41 ° E, 27 October 2006, Osborne J. 200 (K). South Region: Bipindi, 3.26 ° N, 10.20 ° E, 20 June 1918, Annet E. 351 (P); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.34 ° E, 14 February 2015, Couvreur T. L. P. 707 (WAG, YA); Efoulan, 2.74 ° N, 10.53 ° E, 25 April 2000, Tchouto Mbatchou G. P. 2846 (KRIBI, WAG); Campo-Ma'an area Mamelles Massif, 2.56 ° N, 9.949 ° E, 23 April 2001, Tchouto Mbatchou G. P. 3252 (KRIBI, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 1896, Zenker G. A. 1117 (K, L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3045 (K, L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1904, Zenker G. A. 3228 (L); Bipindi, 3.08 ° N, 10.41 ° E, 01 March 1914, Zenker G. A. 515 (U, WAG). South-West Region: on trail leading to top of Mt Etinde after Ekonjo village, 4.06 ° N, 9.153 ° E, 01 April 2016, Couvreur T. L. P. 1027 (WAG, YA); on top of hill near Small Ekombe village 3 km after Kumba on road to Ekondo Titi town, 4.62 ° N, 9.374 ° E, 13 January 2016, Couvreur T. L. P. 978 (WAG, YA); Mokoko Forest Reserve Dikome, 4.48 ° N, 9.033 ° E, 05 May 1994, Ekema S. N. 944 (K); Mount above Batoke, 4.08 ° N, 9.083 ° E, 24 April 1984, Thomas D. W. 3455 (BR, LBV, MO, P, WAG, YA); Takamanda Forest Reserve, 6.21 ° N, 9.433 ° E, 30 April 1987, Thomas D. W. 7372 (MO, P, WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
49B73FDCFDA20879292ED9A1836C4E24.taxon	description	Description. Trees or shrubs, 2 - 50 m tall, d. b. h. up to 90 cm; stilt roots or buttresses absent or present. Indumentum of simple hairs. Leaves: petiole 1 - 12 mm long, 1 - 2 mm wide; blade 3.6 - 21.3 cm long, 1.2 - 8.4 cm wide, lanceolate, ovate, elliptic, obovate, oblong, or oblanceolate, apex acuminate or acute or obtuse or cuspidate, acumen 0.2 - 2.1 cm long, base rounded, cordate, cuneate, obtuse, or truncate; midrib sunken or flat above, rarely slightly raised; secondary veins 7 to 20 pairs; tertiary venation reticulate. Inflorescences axillary, plants ramiflorous on young foliate or older leafless branches, rarely cauliflorous, 1 - 32 - flowered; pedicel 1 - 12 mm long; in fruit 1 - 30 mm long; bracts 1 - 6, basal or inserted along the pedicel. Flowers bisexual with 9 perianth parts in 3 whorls; sepals 3, valvate, free or basally fused, 1 - 7 mm long, apex acute or acuminate or rounded or apiculate, base truncate; petals free, outer petals longer than inner; outer petals 3, 5.8 - 64 mm long, 1.2 - 6 mm wide, linear, lanceolate, ligulate-lanceolate, or ovate, apex acute, rounded, or obtuse, base broad and concave; inner petals 3, valvate, 3.5 - 48 mm long, 1 - 5.4 mm wide, linear, ovate, oblong, rhombic, or lanceolate, apex acute, obtuse, or acuminate, base broad and concave,; stamens 40 - 300, 1 - 2 mm long, oblong or clavate; connective apex capitate, shieldlike, or conical, filaments connate at base to form a cone surrounding the carpels, or staminal cone absent; staminodes present, in one outer and one inner whorl, the inner whorl absent in X. aurantiiodora, X. mildbraedii, and X. quintasii; carpels free, 3 to 50, 1 - 3 mm long, stigma filiform, cylindrical, oblong, linear-falcate or ellipsoid. Monocarps dehiscent, stipitate, subsessile, or sessile; monocarps 1 to 36, 19 - 98 mm long, 6 - 40 mm wide, narrowly oblong to oblong, sometimes falciform, torulose, or moniliform, apex acute, rounded or mucronate; seeds 5 - 22 mm long, 3 - 17 mm wide, ellipsoid to oblong, somewhat flattened, seed coat with a fleshy outer layer (sarcotesta) and hard inner layer, or sarcotesta absent; aril absent or present.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5DD43446DD3488885986FEA50E88F056.taxon	description	Figs 141, 142; Map 17 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5DD43446DD3488885986FEA50E88F056.taxon	materials_examined	Type. Sierre Leone. without definite locality, Smeathman H. s. n., no date: lectotype, designated by Verdcourt (1971 a), p. 77: G, secondary lectotype, designated here: G-DC [00201441 on 2 sheets]; isolectotypes: BM [BM 000510763, branch on right-hand side of sheet]; FI-W [005603].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5DD43446DD3488885986FEA50E88F056.taxon	description	Description. Tree, up to 46 m tall, d. b. h. 30 - 58 cm; plank buttresses present. Old branches glabrous, young branches glabrous to sparsely pubescent, the hairs 0.1 - 0.4 mm long. Leaves: petiole 4 - 9 mm long, ca. 1 mm wide, glabrous to sparsely pubescent, slightly grooved, blade inserted on the side of the petiole; blade 7.3 - 16.3 cm long, 2.1 - 6.6 cm wide, lanceolate-ovate to elliptic, occasionally oblong, narrowly elliptic, oblanceolate, or ovate, apex acuminate, acumen 0.6 - 2.0 cm long, base cuneate to broadly cuneate, short-decurrent, sometimes asymmetrical, subcoriaceous, below glabrous to sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous, whitish below; midrib sunken, above glabrous when young and old, below glabrous when young and old; secondary veins 9 to 13 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate or old leafless branches, axillary, peduncle 1.5 - 2.8 (- 4) mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 7 per inflorescence; pedicel 5 - 15 mm long, 1 - 2 mm in diameter, pubescent; in fruit 10 - 18 mm long, 2 - 6 mm in diameter, glabrous; bracts 1 to 2, towards the middle of pedicel, 2 - 3 mm long, 3 mm wide; sepals 3, valvate, basally to 2 / 3 fused, forming a cup, 2 - 4 mm long, 3 - 6 mm wide, triangular, apex acute to rounded, base truncate, greenish, glabrous to sparsely pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 28 - 64 mm long, 3.8 - 6 mm wide at base, linear, apex obtuse, base broad and concave, yellow to light green or white, pubescent outside, pubescent inside; inner petals 3, valvate, 18.7 - 51 mm long, 3.8 - 6 mm wide at base, linear, apex acute, base broad and concave, cream to greenish yellow, pubescent, base glabrous outside, pubescent inside; stamens 140 to 300, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, pubescent to glabrous, cream; carpels free, 45 to 50, ovary 1 - 2 mm long, stigmas connivent, linear, 3.5 - 4.7 mm long, pubescent to glabrous. Monocarps stipitate, stipe 2 - 8 mm long, 2 - 4 mm in diameter; monocarps 20 to 30 (36), 32 - 82 mm long, 3 - 8 mm wide, narrowly oblong, weakly torulose, apex with a blunt beak 1.5 - 3.5 mm long ,, glabrous, verrucose and wrinkled when dried, green to red outside, endocarp red; seeds 4 to 12 per monocarp, in a single row, 5 - 6 mm long, 3 - 4 mm wide, ellipsoid; sarcotesta absent; aril flat, bilobed, orange to pale yellow.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5DD43446DD3488885986FEA50E88F056.taxon	distribution	Distribution. A widespread species in West, Central and East Africa, from Senegal to Kenya and south to Angola and northern Mozambique; in Cameroon known from East, South, Central, Littoral, South-West, and West and regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5DD43446DD3488885986FEA50E88F056.taxon	materials_examined	Selected specimens examined. Central Region: Ndanan 1 abandoned enclosure on river before bridge, 3.62 ° N, 11.58 ° E, 09 March 2004, Cheek M. 11579 (K, YA); NE of Ndangan I on logging trail into Mefou NP, 3.62 ° N, 11.59 ° E, 23 March 2004, Cheek M. 11999 (K, YA); East side of park, 3.60 ° N, 11.59 ° E, 16 October 2002, Gosline W. G. 423 (K, YA); 60 km S of Edea S of Mboke 11 km E of km 58 of road Edea-Kribi, 3.47 ° N, 10.5 ° E, 22 April 1965, Leeuwenberg A. J. M. 5504 (BR, K, MO, P, WAG, YA). East Region: 64 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.09 ° N, 14.66 ° E, 07 March 2019, Couvreur T. L. P. 1223 (MPU, WAG, YA); Roue Mintom I (70 km E de Djoum-Alati (100 km SE de Djoum) PK 15, 2.83 ° N, 13.35 ° E, 01 January 1973, Letouzey R. 11773 (K, P, WAG, YA); Piste du massif de Fessimi sur la carte de Deng-Deng, 5.2 ° N, 13.52 ° E, 09 February 1961, Letouzey R. 3433 (YA); Deng Deng, 5.2 ° N, 13.52 ° E, 01 April 1914, Mildbraed G. W. J. 8826 (K). Littoral Region: Yabassi, 4.34 ° N, 10.41 ° E, 27 October 2007, Fenton E. 163 (K); Duala, 4.05 ° N, 9.71 ° E, 01 June 1917, Fleury F. 33338 (P); Olombe, 3.60 ° N, 9.958 ° E, 05 November 2014, Kamdem N. 177 (YA). South Region: Okala (Batanga), 3.2 ° N, 11.02 ° E, 20 September 1945, Aubreville A. 133 (P); Ebolowa, 2.96 ° N, 11.28 ° E, 20 November 1968, Bos J. J. 3341 (P, WAG, YA); ca 15 km from Kribi 1 km S of Ebolowa road, 3.1 ° N, 10.25 ° E, 20 February 1970, Bos J. J. 6382 (B, BR, C, K, LD, M, P, WAG, YA); 23 km east from Lele village, 2.28 ° N, 13.32 ° E, 08 September 2013, Couvreur T. L. P. 471 (WAG, YA); 16 km on the recently reconstructed road from Ebolowa to Minkok, 2.75 ° N, 11.25 ° E, 29 January 1975, de Wilde J. J. F. E 7931 (B, BR, K, MO, P, U, WAG, YA); Station de cacaoyer de N'koemvone 14 km on the road from Ebolowa to Ambam, 2.81 ° N, 11.13 ° E, 17 February 1975, de Wilde J. J. F. E 7970 (BR, K, MO, P, U, WAG, YA); 18 km S of the Lobe River along the road to Campo, 2.71 ° N, 9.866 ° E, 18 March 1975, de Wilde J. J. F. E 8085 (BR, MO, P, WAG, YA); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.8 ° N, 11.13 ° E, 10 December 1975, de Wilde J. J. F. E 8707 (BR, K, MO, P, U, WAG); Mebemonko (20 km NO d'Oveng), 2.55 ° N, 12.17 ° E, 24 October 1966, Letouzey R. 8186 (YA); km 48 route Kribi-Ndjabilobi (village Elone), 3.1 ° N, 10.25 ° E, 23 July 1957, Mpom B. 282 (P, YA); Canon du Ntem 16 km SW de Nyabessan, 2.32 ° N, 10.28 ° E, 30 November 1982, Nkongmeneck B. A. 399 (P); Campo-Ma'an area Nsengou, 2.18 ° N, 10.58 ° E, 05 February 2001, Tchouto Mbatchou G. P. 3126 (KRIBI, WAG); Campo-Ma'an area Ebodje, 2.56 ° N, 9.833 ° E, 16 April 2001, van Andel T. R. 3343 (KRIBI, WAG, YA). South-West Region: Bayang Mbo Wildlife Sanctuary before Mbu river, 5.34 ° N, 9.495 ° E, 27 March 2016, Couvreur T. L. P. 1021 (WAG, YA); on trail through palm oil plantation 3 km before lava flow and Seme Beach hotel when coming from Limbe, 4.06 ° N, 9.078 ° E, 18 October 2013, Couvreur T. L. P. 516 (WAG, YA); Bakolle Bakossi on Kumba-Mamfe road, 5.01 ° N, 9.666 ° E, 24 May 1986, Etuge M. 148 (MO, WAG, YA); A 15 km au Sud Est de Molobo (village situe a 50 km au Sud de Batouri), 4.02 ° N, 14.32 ° E, 21 July 1963, Letouzey R. 5456 (P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C933E27BB3F6C3E341EA561FFBECE22.taxon	description	Fig. 142; Map 17 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C933E27BB3F6C3E341EA561FFBECE22.taxon	materials_examined	Type. Cameroon. South-West Region; Mount Cameroon, Mann G. 1193, Feb 1862: lectotype, designated by Johnson and Murray (2018), p. 42: K [K 000105591]; isolectotypes: GH; K [K 000105592]; P [P 00169118]; U [U 0095511].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C933E27BB3F6C3E341EA561FFBECE22.taxon	description	Description. Tree, up to 20 m tall, d. b. h. up to 30 cm; stilt roots present, to 2 m above the base. Old branches glabrous, brown, young branches pubescent, with appressed hairs 0.2 - 0.3 mm long. Leaves: petiole 6 - 9 mm long, ca. 2 mm wide, pubescent to glabrous, grooved, blade inserted on the side of the petiole; blade 6.3 - 15.4 cm long, 2.9 - 7.9 cm wide, oblong to obovate, sometimes elliptic, apex acuminate to cuspidate, acumen 0.2 - 1.0 cm long, base cuneate and decurrent, coriaceous to subcoriaceous, below pubescent when young, glabrous when old, above glabrous when young and old, discolorous, much paler below; midrib flat to sunken, above glabrous when young and old, below glabrous to sparsely pubescent when young, glabrous to sparsely pubescent when old; secondary veins 10 to 13 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle not visible. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 6 - 11 mm long, 1 - 2 mm in diameter, pubescent; in fruit 10 - 18 mm long, 4 - 5 mm in diameter, glabrous; bracts 2, towards the middle of pedicel, 1 - 3 mm long, 1 - 3 mm wide; sepals 3, valvate, basally fused, 5 - 7 mm long, 5 - 6 mm wide, triangular to ovate, apex acute, base truncate, green, pubescent outside, glabrous inside; petals free; outer petals 3, 8.2 - 9.5 mm long, 3 - 4.5 mm wide at base, ovate, apex acute to rounded, base truncate, yellow, minutely pubescent outside, minutely pubescent inside; inner petals 3, valvate, 5.9 - 8.3 mm long, 1 - 2.6 mm wide at base, elliptic to rhombic, apex acuminate to acute, base truncate, yellow, pubescent, nearly glabrous outside, minutely pubescent inside; stamens 100 to 120, in 7 to 8 rows, 2 - 3 mm long, clavate to oblong; connective apex shield-like, pubescent, cream; carpels 9 to 15, ovary 2 mm long, stigmas connivent, linear, 3.2 - 4.6 mm long, glabrous. Monocarps stipitate, stipe 10 - 18 mm long, 3 - 5 mm in diameter; monocarps 1 to 10, 66 - 100 mm long, 10 - 14 mm wide, narrowly oblong to cylindrical, somewhat falciform and torulose, apex with a beak 1 - 1.5 mm long, glabrous, verrucose and longitudinally wrinkled when dried, smooth when fresh, reddish to purplish green outside, endocarp red; seeds 1 to 5 per monocarp, in a single row, 13 - 15 mm long, 9 - 11 mm wide, ellipsoid; sarcotesta absent; aril brushlike, unlobed, blood-red.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C933E27BB3F6C3E341EA561FFBECE22.taxon	distribution	Distribution. Eastern Nigeria and Cameroon, also on the Islands of Bioko (Equatorial Guinea) and Sao Tome (Sao Tome & Principe); in Cameroon known from Central, South-West, Northwest and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4C933E27BB3F6C3E341EA561FFBECE22.taxon	materials_examined	Specimens examined. Central Region: Chaine de Nkohom a 42 km SSW de Ndiki, 4.77 ° N, 10.83 ° E, 14 November 1983, Nkongmeneck B. A. 580 (P). North-West Region: High ridge on boundary of Bali-Ngemba, 5.80 ° N, 10.09 ° E, 14 November 2000, Cheek M. 10527 (K, MO, P, WAG, YA); Gazette Bali Ngemba F R, 5.81 ° N, 10.08 ° E, 05 October 2001, Onana J. M. 1825 (K); Gazette Bali Ngemba F R, 5.81 ° N, 10.08 ° E, 05 October 2001, Onana J. M. 1835 (K). South-West Region: Nyasoso, 4.86 ° N, 9.7 ° E, 04 June 1996, Cable S. 2870 (K, YA); Kupe village, 4.78 ° N, 9.716 ° E, 15 July 1996, Cable S. 3814 (K, YA); Mungo River F. R., 4.78 ° N, 9.607 ° E, 30 November 1999, Cheek M. 10203 (K); Peak 1, 4.80 ° N, 9.708 ° E, 01 November 1995, Cheek M. 7605 (K, YA); Kodmin Mt above the village, 4.99 ° N, 9.701 ° E, 13 February 1998, Cheek M. 9192 (K, P, WAG, YA); Rumpi mountains forest trail ca 5 km after Dikome Balue village ca 40 km north of Kumba, 4.93 ° N, 9.242 ° E, 10 January 2016, Couvreur T. L. P. 949 (WAG, YA); Rumpi mountains forest trail ca 5 km after Dikome Balue village ca 40 km north of Kumba, 4.93 ° N, 9.225 ° E, 11 January 2016, Couvreur T. L. P. 967 (WAG, YA); slopes of Mount Cameroon on the Bokwango trail near Bokwango village 4 km south west of Buea, 4.12 ° N, 9.183 ° E, 23 March 2016, Couvreur T. L. P. 993 (WAG, YA); Buea, 4.15 ° N, 9.24 ° E, 01 January 1904, Deistel H. 454 (U); Below Kupe rock near Esense river, 4.78 ° N, 9.683 ° E, 25 January 1995, Elad M. 90 (K, YA); Likombe, 4.21 ° N, 9.166 ° E, 22 February 1995, Etuge M. 1160 (K, YA); Kupe village, 4.78 ° N, 9.7 ° E, 08 November 1995, Etuge M. 1441 (K, WAG, YA); Nyasoso, 4.81 ° N, 9.716 ° E, 27 February 1996, Etuge M. 1740 (K, WAG, YA); 5 km west of village spring in Kodmin, 4.98 ° N, 9.7 ° E, 22 January 1998, Gosline W. G. 75 (K, WAG, YA); Monts Rumpi-Rata Mount (1788 m) 2 km au SW de Dikome Balue, 4.89 ° N, 9.23 ° E, 24 March 1976, Letouzey R. 14551 (P, YA); Buea, 4.15 ° N, 9.233 ° E, 01 January 1929, Maitland T. D. 233 (K); Cameroon Mountain, Buea area, 4.15 ° N, 9.233 ° E, 01 January 1930, Maitland T. D. s. n. (K [K 000105602]); Cameroon Mountain, 4.2 ° N, 9.183 ° E, Feburary 1862, Mann G. 1193 (K); southern slope of Mount Cameroon above Batoke, 4.13 ° N, 9.08 ° E, 09 January 1984, Thomas D. W. 2981 (K, MO); Forest in the Rumpi Hills near Dikome Balue, 4.90 ° N, 9.253 ° E, 01 March 1984, Thomas D. W. 3305 (K, MO, P, WAG, YA); Aguosho 10 km SSW of Akwaya, 6.3 ° N, 9.466 ° E, 19 March 1985, Thomas D. W. 4554 (YA); Limbe District Fako Division Mt Etinde N face of N ridge, 4.07 ° N, 9.11 ° E, 24 October 1992, Wheatley J. I. 605 (K, P). West Region: Banna, 5.15 ° N, 10.27 ° E, 01 January 1904, Deistel H. 151 (A, BM, P); Route Batcha-Batchingou (22 km ESE Bafang), 5.15 ° N, 10.18 ° E, 23 November 1974, Letouzey R. 13300 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E92C79F7CB2AF6807880157BDC5E45D5.taxon	description	Map 17 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E92C79F7CB2AF6807880157BDC5E45D5.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Coquilhatville [= Mbandaka], Dewevre A. P. 660, 24 Jan 1896: lectotype, sheet here designated: BR [BR 0000008824691]; isotypes: BR [BR 0000008824271, BR 0000008824271].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E92C79F7CB2AF6807880157BDC5E45D5.taxon	description	Description. Tree to shrub, 2 - 10 m tall, d. b. h. 30 cm; stilt roots and buttresses absent. Old branches glabrous, young branches glabrous to sparsely pubescent, the hairs 0.2 - 0.3 mm long. Leaves: petiole 3 - 7 mm long, ca. 1 mm wide, glabrous to sparsely pubescent, slightly grooved, blade inserted on the side of the petiole; blade 6.3 - 11.1 cm long, 2.5 - 4.7 cm wide, elliptic to oblong, occasionally oblanceolate, apex acute to obtuse, base cuneate and decurrent, papyraceous to subcoriaceous, below glabrous when young, glabrous to sparsely pubescent when old, above glabrous when young and old, discolorous, tan-colored below; midrib sunken or flat, above glabrous to sparsely pubescent when young, glabrous to sparsely pubescent when old, below glabrous when young, sparsely pubescent when old; secondary veins 10 to 14 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 1.5 - 2 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 5 per inflorescence; pedicel 4 - 7 mm long, ca. 1 mm in diameter, sparsely pubescent to glabrous; in fruit 7 - 10 mm long, 1 - 2 mm in diameter, sparsely pubescent to glabrous; bracts 2 to 3, basal and towards the upper half of pedicel, all similar, 1 - 2 mm long, 1 - 2 mm wide; sepals 3, slightly imbricate at base, free, 2 mm long, 2 mm wide, triangular, apex apiculate, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 8.9 - 12 mm long, 2.4 - 3.5 mm wide at base, ligulate-lanceolate, apex obtuse, base broad and concave, light yellow to light green, pubescent outside, pubescent but glabrous towards center inside; inner petals 3, valvate, 9.3 - 11.2 mm long, 1.2 - 2.3 mm wide at base, narrowly oblong, apex obtuse, base broad and concave with a pronounced internal tooth overhanging the concavity and fleshy glandlike margins, light yellow to greenish yellow, densely pubescent outside, pubescent inside; stamens 40 to 60, in 5 to 6 rows, ca. 2 mm long, oblong; connective apex capitate, glabrous; carpels 3 to 5, ovary 1 mm long, stigmas slightly separate, narrowly oblong, ca. 1 mm long, glabrous. Monocarps stipitate, stipe 6 - 10 mm long, 2 - 3 mm in diameter; monocarps 1 to 3, 33 - 65 mm long, 7 - 9 mm wide, narrowly oblong, sometimes falciform, strongly torulose, apex with an acute beak up to 5 mm long, glabrous, wrinkled or striate when dried, green outside, endocarp color unknown; seeds 1 to 4 per monocarp, in a single row, 13 - 21 mm long, 6 - 8 mm wide, ellipsoid, pointed at one end; sarcotesta absent; aril present, fimbriate, extending over the length of the seed, membranous, smooth, red to orange-brown when dried.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E92C79F7CB2AF6807880157BDC5E45D5.taxon	distribution	Distribution. From southern Cameroon to the Democratic Republic of the Congo; in Cameroon known from the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
E92C79F7CB2AF6807880157BDC5E45D5.taxon	materials_examined	Specimens examined. East Region: West bank of Sangha River opposite Ndakan camp and 2 km S, 2.35 ° N, 16.13 ° E, 12 February 1989, Harris D. J. 1846 (MO); Sangha R, 2.78 ° N, 16 ° E, 22 May 1988, Harris D. J. 752 (P); Mbekou pres Moloundou, 2.05 ° N, 15.22 ° E, 01 January 1959, Letouzey R. 1374 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7D8C0171A33AF4508A6DE0CB2639A23F.taxon	description	Map 17 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7D8C0171A33AF4508A6DE0CB2639A23F.taxon	materials_examined	Type. Cameroon. South Region; Bipindi, Zenker G. A. 4747, 1903: holotype: BM [000511011]; isotypes: BR [BR 0000014581892]; G; K; L [0191105]; M; MO [751089]; P.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7D8C0171A33AF4508A6DE0CB2639A23F.taxon	description	Description. Tree, height unknown, d. b. h. up to 30 cm; buttresses present. Old branches glabrous, young branches pubescent, the hairs 0.3 - 0.4 mm long. Leaves: petiole 3 - 10 mm long, 2 mm wide, sparsely pubescent, slightly grooved, blade inserted on top of the petiole; blade 10 - 17.2 cm long, 3.6 - 6.5 cm wide, elliptic to oblong, apex acuminate, acumen 0.3 - 2.1 cm long, base cuneate to rounded, papyraceous to subcoriaceous, below sparsely pubescent when young, sparsely pubescent when old, above glabrous when young, sparsely pubescent when old; concolorous or slightly glaucous abaxially; midrib sunken, above sparsely pubescent when young and old, below sparsely pubescent when young and old; secondary veins 8 to 15 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle branched, 1 - 3 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 10 per inflorescence; pedicel 5 - 8 mm long, 1 - 2 mm in diameter, pubescent; in fruit 6 - 7 mm long, 2 - 3 mm in diameter, rusty pubescent; bracts 1 to 2, one basal and one upper towards the middle of pedicel, 2 - 3 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 4 - 5 mm long, 4 - 5 mm wide, ovate, apex acute to obtuse, base truncate, densely pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 13 - 23 mm long, 3.8 - 6 mm wide at base, oblong-lanceolate, apex obtuse, base broad and concave, light yellow, pubescent, base glabrous outside, pubescent inside; inner petals 3, valvate, 10 - 17.5 mm long, 2.5 - 4 mm wide at base, oblong-lanceolate, apex acute, base broad and concave, yellow with red on the inner base, pubescent outside, glabrous inside except at apex; stamens ca. 200, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous, red; carpels ca. 9, ovaries 2 mm long, stigmas connivent at base, linear, slightly thickened at the midpoint, ca. 2.2 mm long, pubescent. Monocarps stipitate, stipe ca. 3 mm long, ca. 7 mm in diameter; monocarps ca. 8, ca. 32 mm long, ca. 17 mm wide, ellipsoid, apex rounded, pubescent, verrucose and slightly wrinkled when dried, color unknown; seeds 7 to 8 per monocarp, in two rows, ca. 9 mm long, ca. 7 mm wide, ellipsoid to flattened ellipsoid, sarcotesta unknown; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7D8C0171A33AF4508A6DE0CB2639A23F.taxon	distribution	Distribution. From southern Nigeria to Cameroon; in Cameroon known only from the South region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7D8C0171A33AF4508A6DE0CB2639A23F.taxon	materials_examined	Specimens examined. South Region: Pres des chutes du Ntem ou de Menve'ele pres Nyabessan (60 km Est de Campo), 2.39 ° N, 10.35 ° E, 08 April 1970, Letouzey R. 10306 (P, YA); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1913, Zenker G. A. 4747 (BM, BR, G, K, L, M, MO, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FC09C298BC283CAF2D319836BA985366.taxon	description	Fig. 143; Map 17 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FC09C298BC283CAF2D319836BA985366.taxon	materials_examined	Type. Cameroon. East Region; Deng-Deng, Mildbraed G. W. J. 8649, Mar 1914: holotype B [B 10 0153141]; isotype: BM [fragment].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FC09C298BC283CAF2D319836BA985366.taxon	description	Description. Tree, up to 35 m tall, d. b. h. up to 50 cm; buttresses present, small. Old branches glabrous, young branches glabrous to pubescent, with fine appressed hairs 0.1 - 0.3 mm long. Leaves: petiole 2 - 5 mm long, ca. 1 mm wide, sparsely pubescent, slightly grooved, blade inserted on the side of the petiole; blade 4.6 - 7.8 cm long, 1.3 - 2.4 cm wide, lanceolate to elliptic or elliptic-oblong, apex acuminate, acumen 0.6 - 1.1 cm long, base cuneate to broadly cuneate, usually asymmetrical, papyraceous, below sparsely to densely golden-sericeous (appressed) when young, sparsely pubescent to densely pubescent when old, above glabrous when young and old, concolorous; midrib slightly raised, above glabrous to sparsely pubescent when young, glabrous when old, below pubescent when young and old; secondary veins 10 to 15 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle rarely present, up to ca. 4 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 5 - 12 mm long, ca. 1 mm in diameter, sparsely pubescent; in fruit 6 - 15 mm long, 3 - 10 mm in diameter, glabrous; bracts 2, towards the middle of pedicel, ca. 2 mm long, ca. 2 mm wide; sepals 3, valvate, 1 / 2 - 3 / 4 fused, forming a cup, 2 - 3 mm long, 2 - 3 mm wide, broadly ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 15.5 - 36 mm long, 2.2 - 3.4 mm wide at base, linear to linear-lanceolate, apex acute, base broad and concave, yellow to cream, sericeous outside, pubescent inside; inner petals 3, valvate, 13.5 - 17.7 mm long, 2.4 - 3.5 mm wide at base, linear, apex acute, base broad and concave, yellow to cream, pubescent with glabrous base on both surfaces; stamens 160 to 200, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 12 to 20, ovary ca. 1 mm long, 3.5 - 3.8 mm long, stigmas connivent, filiform, 2.5 - 2.8 mm long, pubescent. Monocarps stipitate, stipe 5 - 24 mm long, 2 - 5 mm in diameter; monocarps 11 to 18, 23 - 54 mm long, 14 - 19 mm wide, obovoid to oblongoid, apex rounded, glabrous, verrucose and wrinkled when dried, reddish green outside, endocarp pink to dark red; seeds 6 to 8 per monocarp, in two rows, 10 - 13 mm long, 6 - 10 mm wide, ellipsoid; sarcotesta glaucous blue or gray; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FC09C298BC283CAF2D319836BA985366.taxon	distribution	Distribution. A widespread species across Central Africa, from southeastern Nigeria to northeastern Democratic Republic of the Congo and south to Angola; in Cameroon known from the East, South, Littoral and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
FC09C298BC283CAF2D319836BA985366.taxon	materials_examined	Specimens examined. East Region: IK 30 Road Mintom 1 (70 km E de Djoum) - Alati (100 km SE de Djoum), 2.83 ° N, 13.35 ° E, 08 January 1973, Letouzey R. 11801 (P, WAG); 21 km ENE of Moloundou-Nguilili Chantier, 2.05 ° N, 15.17 ° E, 10 March 1975, Mbenkum T. F. 310 (P, YA); Deng Deng, 5.2 ° N, 13.52 ° E, 01 March 1914, Mildbraed G. W. J. 8649 (B). Littoral Region: 6 km NE of Dibombe a village on km 11 of Loum-Yabassi road, 4.68 ° N, 9.816 ° E, 26 May 1972, Leeuwenberg A. J. M. 9915 (WAG, YA); Douala (route Razel), 4.02 ° N, 9.71 ° E, Mbarga A. 58 (P, YA). South Region: 30 km east from Lele village, 2.27 ° N, 13.29 ° E, 09 September 2013, Couvreur T. L. P. 485 (WAG, YA). South-West Region: Likomba-Pflanzung 15 - 35 km NE von Victoria [Limbe], 4.09 ° N, 9.33 ° E, 01 November 1928, Mildbraed G. W. J. 10629 (A, K); Mount Mabeta peninsula Mabeta, 3.98 ° N, 9.233 ° E, 19 October 1997, Tchouto Mbatchou G. P. 1756 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6C39D8EF108C036B6B1ABBB7D106B44C.taxon	description	Map 17 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6C39D8EF108C036B6B1ABBB7D106B44C.taxon	materials_examined	Type. Guinea. Farana Region; Farana, Scott Elliot G. F. 5325, Mar 1924: lectotype, designated by Johnson and Murray (2018), p. 165: B [B 100153142]; isolectotypes: BM [BM 000510796, lower half of sheet]; GH; K [K 000199071]; P [P 00169156]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6C39D8EF108C036B6B1ABBB7D106B44C.taxon	description	Description. Tree, 10 - 18 m tall, d. b. h. unknown; stilt roots and buttresses absent. Old branches glabrous, young branches densely pubescent with erect reddish brown hairs 0.2 - 0.8 mm long. Leaves: petiole 4 - 6 mm long, ca. 1 mm wide, pubescent, slightly grooved, blade inserted on the side of the petiole; blade 4.5 - 9.5 cm long, 1.8 - 5.6 cm wide, lanceolate-ovate to elliptic, oblong, or oblong-lanceolate, apex obtuse, emarginate, or acute, rarely with an acumen ca. 0.5 cm long, base cuneate to rounded, subcoriaceous, below finely appressed-pubescent when young, sparsely pubescent to glabrous when old, above glabrous when young and old, discolorous; midrib sunken, above pubescent when young and old, below pubescent when young and old; secondary veins 10 to 16 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 3 - 6 mm long, 1 mm in diameter, pubescent; in fruit 10 - 12 mm long, 1 - 6 mm in diameter, glabrous; bracts 2, towards the middle of pedicel, 2 - 3 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 2 - 5 mm long, 3 - 4 mm wide, ovate, apex acute to apiculate, base truncate, sericeous outside, glabrous inside; petals free, subequal; outer petals 3, 12.6 - 32 mm long, 1.2 - 2.5 mm wide at base, linear, apex obtuse, base broad and concave, white, tinged purple at base, densely pubescent outside, pubescent inside; inner petals 3, valvate, 16 - 24 mm long, 2.3 - 3.9 mm wide at base, linear, apex acute, base broad and concave, white, tinged with purple at the base, pubescent, base glabrous outside, pubescent and glabrous towards base inside; stamens ca. 120, in 7 to 8 rows, 1 - 2 mm long, clavate to oblong; connective apex shield-like, glabrous; carpels 9 to 10, ovary 1 - 2 mm long, stigmas connivent, lanceolate, 1.3 - 2.3 mm long, pubescent to glabrous. Monocarps usually sessile or stipitate, stipe 3 - 6 mm long, 2 - 3 mm in diameter; monocarps 3 to 8, 20 - 38 mm long, 9 - 13 mm wide, obovoid to oblong, apex rounded, glabrous to sparsely pubescent, verrucose, slightly wrinkled when dried, red to purple tinged outside, endocarp red; seeds up to 9 per monocarp, in two rows, 9 - 12 mm long, 7 - 8 mm wide, flattened ellipsoid to oblong; sarcotesta present, color in vivo unknown; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6C39D8EF108C036B6B1ABBB7D106B44C.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West and Central Africa, from Guinea-Bissau to Togo and from Cameroon to Central African Republic; in Cameroon known from Adamaoua and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
6C39D8EF108C036B6B1ABBB7D106B44C.taxon	materials_examined	Specimens examined. Adamaoua Region: Falls in the Vina river ca 15 km SE of Ngaoundere, 7.2 ° N, 13.71 ° E, 01 December 1964, de Wilde W. J. J. O 4447 (K, WAG); Mbalang- 16 km E Ngaoundere, 7.32 ° N, 13.58 ° E, 27 January 1978, Fotius G. 2984 (YA); Tchabal-Mbabo, 7.05 ° N, 12.25 ° E, 16 March 1978, Fotius G. 3108 (YA); Ngaou Loumou (1700 m), 7.52 ° N, 13.85 ° E, 20 October 1967, Jacques-Felix H. 8724 (YA); Mayo Darle, 6.47 ° N, 11.55 ° E, 10 November 1967, Jacques-Felix H. 9110 (YA); Sadolkoulay (36 km E Ngaoundere), 7.31 ° N, 13.87 ° E, 05 December 1964, Raynal J. 12228 (P, YA); Pres Katil-Foulbe 50 km SE Ngaoundere, 7.02 ° N, 13.94 ° E, 20 October 1983, Satabie B. 687 (P). West Region: Ndop Plain road to French Map ref No 23, 5.97 ° N, 10.40 ° E, 30 March 1962, Brunt M. A. 261 (K); Bayangam, 5.3 ° N, 10.45 ° E, 01 January 1939, Jacques-Felix H. 2965 (P); Kontchankap, 5.58 ° N, 10.80 ° E, 01 February 1939, Jacques-Felix H. 3039 (P); a 20 km E de Foumbot, 5.56 ° N, 10.8 ° E, 26 October 1974, Satabie B. 15 (MO, P); Bayangam, 5.3 ° N, 10.45 ° E, 17 May 1978, Westphal E. 10047 (P, WAG); Bayangam, 5.3 ° N, 10.45 ° E, 17 May 1978, Westphal E. 10048 (WAG, YA); Bayangam, 5.3 ° N, 10.45 ° E, 17 May 1978, Westphal E. 10049 (K, WAG, YA); Plants collected in the region of Bayangam, 5.3 ° N, 10.45 ° E, 17 December 1978, Westphal E. 10172 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5AD78677744E4855FCB27D7FAD8CFD2D.taxon	description	Map 17 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5AD78677744E4855FCB27D7FAD8CFD2D.taxon	materials_examined	Type. Democratic Republic of the Congo. Tshopo Region; Yangambi, plateau de la Luweo, Louis J. L. P. 6777, Nov 1937: holotype: BR [BR 0000008824738]; isotypes: B [B 100249561]; BM K [K 000542217]; P [P 00169152]; US.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5AD78677744E4855FCB27D7FAD8CFD2D.taxon	description	Description. Tree, up to 40 m tall, d. b. h. up to 40 cm; buttresses present, small. Old branches sparsely pubescent with erect hairs, young branches pubescent with persistent erect hairs 0.3 - 0.8 mm long. Leaves: petiole 3 - 4 mm long, ca. 1 mm wide, densely erect pubescent, slightly grooved, blade inserted on the side of the petiole; blade 4.2 - 9.7 cm long, 2 - 3.3 cm wide, elliptic-oblong to lanceolate, occasionally oblong or ovate, apex acute to rounded, base cuneate to rounded, papyraceous to subcoriaceous, below sparsely pubescent with erect hairs when young and old, above glabrous when young, glabrous to pubescent when old, slightly discolorous; midrib sunken or flat, above densely pubescent when young and old, below pubescent when young and old; secondary veins 7 to 14 pairs, pubescent above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 1.5 - 2.5 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 6 per inflorescence; pedicel 1 - 2 mm long, 1 - 2 mm in diameter, densely pubescent; in fruit 5 - 10 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, evenly spaced, 2 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, free to basally fused, ca. 2 mm long, 3 - 4 mm wide, reniform to semicircular, apex acute to rounded, base truncate, densely pubescent outside, glabrous inside; petals free, subequal, fawn-olive, brown-violet, or purple; outer petals 3, 6.6 - 11 mm long, 2.4 - 4.4 mm wide at base, lanceolate to ovate, apex obtuse, base broad and concave, densely pubescent outside, pubescent but glabrous towards center inside; inner petals 3, valvate, 5.8 - 9.3 mm long, 3 - 4 mm wide at base, lanceolate, apex acuminate to obtuse, base broad and concave, with round glandlike thickenings on margins, pubescent outside, pubescent but glabrous towards center inside; stamens 80 - 130, in 6 to 7 rows, ca. 1 mm long, clavate to oblong; connective apex shield-like to capitate, glabrous; carpels 6 to 11, ovary 1 - 2 mm long, stigmas connivent, linear, 1.5 - 2.2 mm long, sparsely pubescent. Monocarps stipitate, stipe 5 - 13 mm long, 1 - 4 mm in diameter; monocarps 1 to 10, 25 - 37 mm long, 10 - 13 mm wide, oblong, apex rounded or with a blunt beak 1 - 1.5 mm long, glabrous to sparsely pubescent with erect hairs, sparsely verrucose and weakly wrinkled when dried, reddish or purplish green outside, endocarp color unknown; seeds up to 6 per monocarp, in a single row, 7 - 10 mm long, 4 - 7 mm wide, ellipsoid; sarcotesta red in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5AD78677744E4855FCB27D7FAD8CFD2D.taxon	distribution	Distribution. From southern Cameroon to northern Gabon and Central African Republic and east to the northeastern Democratic Republic of the Congo; in Cameroon known from the East and South regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
5AD78677744E4855FCB27D7FAD8CFD2D.taxon	materials_examined	Specimens examined. East Region: A 9 km a l'ouest de Yenga Port Gentil village situe a 25 km au NNE de Moloundou, 2.34 ° N, 15.26 ° E, 21 April 1971, Letouzey R. 10701 (P, YA). South Region: Pres Akonetye PK 95 sur route Mintom I (70 km E de Djoum) - Mbalam (140 km ESE de Djoum), 2.67 ° N, 12.87 ° E, 22 January 1973, Letouzey R. 11876 (P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7712CE1CCADE45950D9DC8C78EE9F72A.taxon	description	Figs 144, 145; Map 17 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7712CE1CCADE45950D9DC8C78EE9F72A.taxon	materials_examined	Type. Cameroon. East Region; Lomie, Mildbraed G. W. J. 5183, 13 May 1911: lectotype, designated by Le Thomas (1969 b), p. 181: HBG [HBG 502479].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7712CE1CCADE45950D9DC8C78EE9F72A.taxon	description	Description. Tree, 30 - 40 m tall, d. b. h. up to 80 cm; stilt roots and buttresses absent. Old branches glabrous, young branches glabrous to pubescent, with fine matted hairs 0.2 - 0.4 mm long. Leaves: petiole 5 mm long, 1 - 2 mm wide, pubescent, grooved, blade inserted on the side of the petiole; blade 5.7 - 7.5 cm long, 1.4 - 1.9 cm wide, lanceolate to elliptic, apex acuminate to acute, acumen 1.2 cm long, base cuneate to rounded, subcoriaceous, below densely silvery-sericeous when young and old, above glabrous when young and old, discolorous; midrib sunken or flat, above glabrous when young and old, below densely pubescent when young and old; secondary veins 10 to 18 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 2 - 4 mm long, ca. 1 mm in diameter, pubescent; in fruit 3 - 6 mm long, 3 - 5 mm in diameter, glabrous; bracts 3 to 4, evenly spaced, ca. 2 mm long, 1 - 2 mm wide; sepals 3, valvate, basally fused, 2 - 3 mm long, 2 - 3 mm wide, triangular, apex acute, base truncate, green, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 25 - 28.7 mm long, 2.5 - 2.6 mm wide at base, linear, apex obtuse, base broad and concave, yellow to light green, pubescent with base glabrous to sericeous outside, pubescent and glabrous towards base inside; inner petals 3, valvate, 16 - 31 mm long, 1.9 - 2.5 mm wide at base, linear, apex acute, base broad and concave, yellow to light green, pubescent outside, pubescent inside; stamens ca. 100, in 7 to 8 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous, red; carpels 7 to 12, ovary ca. 1 mm long, stigmas connivent, sometimes free at tips, filiform, 1.6 - 2.5 mm long, pubescent. Monocarps sessile; monocarps 6 to 8, 26 - 41 mm long, 11 - 22 mm wide, obovoid, ellipsoid or oblongoid, becoming slightly ridged and sunken between the ridges when dried, apex rounded, glabrous, greenish brown, flecked with pale brown lenticels outside, endocarp pink-red; seeds up to 10 per monocarp, in two rows, 7 - 11 mm long, 6 - 8 mm wide, ellipsoid; sarcotesta greenish white in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7712CE1CCADE45950D9DC8C78EE9F72A.taxon	distribution	Distribution. Central Cameroon to southwestern Central African Republic and south to southwestern Democratic Republic of the Congo; in Cameroon known from East, South, Central and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
7712CE1CCADE45950D9DC8C78EE9F72A.taxon	materials_examined	Specimens examined. Central Region: Mefou proposed national park near Mefou town, 3.62 ° N, 11.58 ° E, 08 March 2004, Cheek M. 11487 (K, YA); Mefou National Park just after Ape Action Africa center along road, 3.62 ° N, 11.58 ° E, 24 April 2013, Couvreur T. L. P. 420 (WAG, YA); Ottotomo Forest Reserve 1 km after reserve base near small loggers road, 3.66 ° N, 11.28 ° E, 25 June 2013, Couvreur T. L. P. 450 (WAG, YA); Yaounde, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 101 (OWU, P); Banlieu de Yaounde sur une piste apres la borne 9, 3.87 ° N, 11.52 ° E, 26 November 1959, Mpom B. 362 (P, YA). East Region: 74 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.16 ° N, 14.71 ° E, 04 March 2019, Couvreur T. L. P. 1198 (MPU, WAG, YA); A 30 km au NE de Bange (km 75 route Yokadouma-Moloundou), 3.02 ° N, 15.12 ° E, 25 May 1963, Letouzey R. 5139 (P, YA); entre Bidjum et Dscha-Posten, 3.15 ° N, 13.61 ° E, 13 May 1911, Mildbraed G. W. J. 5183 (HBG); Deng Deng, 5.2 ° N, 13.52 ° E, 01 January 1914, Mildbraed G. W. J. 8827 (BM, K). South Region: 21 km east from Lele village, 2.27 ° N, 13.33 ° E, 07 September 2013, Couvreur T. L. P. 467 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.47 ° N, 10.33 ° E, 13 February 2015, Couvreur T. L. P. 694 (WAG, YA); 17 km on the road from Ebolowa to Minkok, 2.81 ° N, 11.13 ° E, 06 February 1975, de Wilde J. J. F. E 7963 (B, BR, K, MO, P, U, WAG, YA); Bipindi - Ebolowa, 3.00 ° N, 10.92 ° E, 01 December 1913, Mildbraed G. W. J. 7618 (B, K); Mvie (Akom II), 2.88 ° N, 10.58 ° E, 28 January 1998, van der Burgt X. M. 364 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B9B32ECA896B3DD1418AB80B6F710B57.taxon	description	Map 18 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B9B32ECA896B3DD1418AB80B6F710B57.taxon	materials_examined	Type. Democratic Republic of the Congo. Katanga; Lukafu, Verdick E. 503, May 1900: holotype: BR [BR 0000024941556].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B9B32ECA896B3DD1418AB80B6F710B57.taxon	description	Description. Tree, up to 13 m tall, d. b. h. 60 cm; buttresses present. Old branches glabrous, young branches sparsely pubescent, the hairs ca. 0.2 mm long. Leaves: petiole 4 - 9 mm long, ca. 1 mm wide, glabrous, slightly grooved, blade inserted on the side of the petiole; blade 7.1 - 9.4 cm long, 2.4 - 3.3 cm wide, lanceolate to lanceolate-elliptic, apex acuminate to acute, acumen 0.4 - 1.1 cm long, base cuneate, subcoriaceous, below glabrous, rarely sparsely pubescent when young, glabrous when old, above glabrous when young and old, slightly discolorous; midrib slightly raised or flat, above glabrous when young and old, below glabrous when young, glabrous when old; secondary veins 10 to 15 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate or old leafless branches, axillary, peduncle 1 - 3.5 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 12 per inflorescence; pedicel 6 - 9 mm long, 0.5 - 1 mm in diameter, pubescent to sparsely pubescent; in fruit 8 - 14 mm long, 2 - 3 mm in diameter, glabrous; bracts 2 to 4, several basal and one upper towards the upper half of pedicel, bract 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, basally fused, 1 - 2 mm long, 2 - 3 mm wide, triangular to ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 19 - 37 mm long, 2.7 - 3.2 mm wide at base, linear, apex obtuse, base broad and concave, yellow-green and red at the base, sparsely pubescent outside, pubescent inside; inner petals 3, valvate, 11.4 - 29 mm long, 2.3 - 2.9 mm wide at base, linear and rigid, apex acute, base broad and concave, yellow-green and red at the base, pubescent outside, pubescent, base glabrous inside; stamens ca. 90, in 6 to 7 rows, 1 mm long, oblong; connective apex shield-like, glabrous; carpels 3 to 4, ovary 1 mm long, stigmas loosely connivent with tips free, linear, 2.5 - 4.4 mm long, glabrous. Monocarps stipitate, stipe ca. 4 mm long, 3 - 4 mm in diameter; monocarps 2 to 4, 19 - 34 mm long, 12 - 15 mm wide, oblongoid, apex obtuse, glabrous, weakly torulose, verrucose and longitudinally wrinkled when dried, green outside, endocarp red; seeds 1 to 5 per monocarp, in a single or two irregular rows, 9 - 11 mm long, 7 - 8 mm wide, ellipsoid; sarcotesta gray or light green in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B9B32ECA896B3DD1418AB80B6F710B57.taxon	distribution	Distribution. From north-central Nigeria to southern Democratic Republic of the Congo and south to northeastern Zambia; in Cameroon known from a single collection in the Central region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0357A8E4902C6CA37A81737860CB0EF2.taxon	description	Fig. 146; Map 18 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0357A8E4902C6CA37A81737860CB0EF2.taxon	materials_examined	Type. Gabon. Nyanga; Mayombe Bayaka, Tono-Sanga Le Testu G. M. P. C. 1760, 9 Aug 1914: holotype: P [P 00169125]; isotypes: BM; K [K 000199054]; BR [BR 0000008825322]; EA [EA 000002492]; LISC [LISC 000403]; P [P 00169126, P 00169127].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0357A8E4902C6CA37A81737860CB0EF2.taxon	description	Description. Tree, up to 40 m tall, d. b. h. up to 30 cm; buttresses present, narrow and thin. Old branches sparsely pubescent to glabrous, young branches densely pubescent, the hairs 0.4 - 1 mm long, branches sinuous or zigzagging. Leaves: petiole 1 - 2 mm long, ca. 2 mm wide, pubescent, grooved, blade inserted on top of the petiole; blade 4.7 - 10.9 cm long, 1.2 - 2.7 cm wide, lanceolate to lanceolate-oblong, apex acute, base truncate and often slightly asymmetrical, subcoriaceous, below densely pubescent when young, sparsely pubescent to densely pubescent when old, the hairs dull grayish brown, above glabrous when young and old, discolorous; midrib flat to slightly raised, above densely pubescent when young, glabrous to densely pubescent when old, below densely appressed-pubescent when young, sparsely appressed-pubescent to densely appressed-pubescent when old; secondary veins 8 to 14 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle ca. 2.5 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; pedicel 1 - 7 mm long, ca. 1 mm in diameter, pubescent; in fruit 8 - 14 mm long, 2 - 3 mm in diameter, glabrous; bracts 2, evenly spaced, 2 - 6 mm long, 2 - 6 mm wide; sepals 3, valvate, basally to 2 / 3 fused, forming a cup, 4 - 6 mm long, ca. 3 mm wide, elliptic to ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 13 - 22 mm long, 2.3 - 3.2 mm wide at base, lanceolate-ligulate, linear-lanceolate, or narrowly triangular, elliptic, apex acute, base broad and concave, cream-colored with a purple blotch at the base, pubescent, base glabrous to densely pubescent outside, pubescent towards margins inside; inner petals 3, valvate, 10.3 - 18.1 mm long, 2.4 - 3.2 mm wide at base, linear-lanceolate, apex acute, base broad and concave with conspicuous hairy tufts on the margins, cream, densely pubescent outside, pubescent, and glabrous towards the base inside; stamens numerous, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 7 to 10, ovary 1 mm long, stigmas loosely connivent with tips free, linear, sinuate, 3 - 3.5 mm long, pubescent. Monocarps sessile; monocarps 4 to 7, 27 - 45 mm long, 21 - 34 mm wide, nearly spherical to ellipsoid, apex rounded or obtuse, pubescent, smooth to verrucose, wrinkled when dried, green outside, endocarp bright red; seeds up to 6 per monocarp, in two rows, 8 - 19 mm long, 8 - 13 mm wide, ellipsoid to flattened ellipsoid; sarcotesta grayish white in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0357A8E4902C6CA37A81737860CB0EF2.taxon	distribution	Distribution. A widespread species with a disjunct distribution between West Africa, from Sierra Leone to Ghana, and Central Africa from eastern Nigeria to western Democratic Republic of the Congo; in Cameroon known from the East, South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0357A8E4902C6CA37A81737860CB0EF2.taxon	materials_examined	Specimens examined. East Region: Dja Reserve (Reserve de Faune du Dja) Bouamir Research Area 90 km southeast of Akonolinga, 3.18 ° N, 12.79 ° E, 23 May 1997, Fogiel M. K. 2098 (MO, WAG); Entre Kondon I et Mbalam pres Ngoila (axe Lomie Souanke), 2.85 ° N, 13.93 ° E, 22 December 1972, Letouzey R. 11707 (P, YA); A 15 km au Sud de Djouo (20 km E de Somalomo sur le Dja), 3.32 ° N, 12.93 ° E, 23 February 1962, Letouzey R. 4361 (P, YA). South Region: 24 km east from Lele village, 2.28 ° N, 13.31 ° E, 08 September 2013, Couvreur T. L. P. 475 (WAG, YA); Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 688 (WAG, YA); Sud Cameroun TDC, 2.65 ° N, 9.9 ° E, 15 November 1991, Halle F. 4247 (P, WAG). South-West Region: Korup National Park P transect P plot subplot 4 E, 5.01 ° N, 8.8 ° E, 24 February 2008, van der Burgt X. M. 1129 (BR, G, K, MO, P, WAG, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECB33CD0E3F12F2C0FC2599D0B706267.taxon	description	Fig. 147; Map 18 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECB33CD0E3F12F2C0FC2599D0B706267.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Bangala, Dewevre A. P. 876, May 1896: holotype: BR [BR 0000008825360]; isotype: BR [BR 0000008825377].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECB33CD0E3F12F2C0FC2599D0B706267.taxon	description	Description. Tree to shrub, up to 15 m tall, d. b. h. up to 40 cm; stilt roots and buttresses absent. Old branches glabrous, young branches glabrous to sparsely pubescent with erect hairs 0.1 - 0.5 mm long. Leaves: petiole 3 - 6 mm long, ca. 1 mm wide, pubescent to glabrous, slightly grooved, blade inserted on the side of the petiole; blade 4.2 - 8.8 cm long, 1.7 - 3.7 cm wide, oblong or elliptic, occasionally lanceolate, lanceolate-ovate, or ovate-oblong, apex acuminate to obtuse, acumen 0.4 - 0.8 cm long, base rounded, papyraceous to subcoriaceous, below sparsely pubescent when young, sparsely pubescent to glabrous when old, above glabrous when young and old, slightly discolorous; midrib sunken, above sparsely pubescent when young, glabrous to sparsely pubescent when old, below sparsely pubescent when young, sparsely pubescent when old; secondary veins 9 to 14 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 1 - 1.5 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 4 per inflorescence; peduncle 1 - 1.5 mm long; pedicel 2 - 3 mm long, ca. 2 mm in diameter, sparsely pubescent to glabrous; in fruit 10 - 17 mm long, 7 - 11 mm in diameter, sparsely pubescent to glabrous; bracts 2 to 3, towards the middle of pedicel, 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, basally fused, reflexed at anthesis, 2 - 3 mm long, ca. 2 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside; petals free, outer petals longer than inner; outer petals 3, 16 - 62 mm long, 2.7 - 5 mm wide at base, linear, lax and ribbonlike, apex obtuse, base broad and concave, pale yellow to yellow-green with purple to red base, densely pubescent outside, sparsely pubescent inside; inner petals 3, valvate, 19 - 48 mm long, 3.1 - 5.2 mm wide at base, linear, lax and ribbonlike, crinkled when dried, apex acute to obtuse, base broad and concave, yellow-green with red base, pubescent at least towards base outside, glabrous to sparsely pubescent inside, densely hairy in the basal concavity; stamens 80 to 100, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 5 to 7, ovary 1 mm long, stigmas loosely connivent with tips free, linear and sometimes falcate, 3.8 - 7 mm long, pubescent to glabrous. Monocarps stipitate, stipe 1 - 5 mm long, 3 - 8 mm in diameter; monocarps 3 to 6, 30 - 44 mm long, 11 - 19 mm wide, oblongoid, apex rounded or with a beak ca. 2 mm long, glabrous, longitudinally ridged, verrucose and wrinkled when dried, green, sometimes purple or red tinged outside, endocarp bright red; seeds 7 to 12 per monocarp, in two rows, 10 - 12 mm long, 6 - 9 mm wide, ellipsoid; sarcotesta white to green in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECB33CD0E3F12F2C0FC2599D0B706267.taxon	distribution	Distribution. Occurs from Senegal to southern Chad and south to northern Angola and the northeastern Democratic Republic of the Congo; in Cameroon known from East, Central and Adamaoua regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
ECB33CD0E3F12F2C0FC2599D0B706267.taxon	materials_examined	Specimens examined. Adamaoua Region: Rives du Djerem pres Mbakaou, 6.32 ° N, 12.82 ° E, 08 December 1959, Letouzey R. 2459 (BR, K, P, YA). Central Region: left bank Sanaga River near ferry Nachtigal ca 20 km N of Obala, 4.35 ° N, 11.63 ° E, 11 June 1964, de Wilde W. J. J. O 2676 (B, BR, K, MO, P, WAG); Left bank Sanaga R near Ferry Nachtigal ca 20 km N of Obala, 4.21 ° N, 11.38 ° E, 19 November 1965, Leeuwenberg A. J. M. 7033 (B, BR, C, GC, K, LUAI, MO, P, UC, WAG, YA); Yangafok II 25 km ENE de Bafia, 4.93 ° N, 11.37 ° E, 26 November 1969, Letouzey R. 9614 (P, YA). East Region: On island in the Sangha River adjacent to the Ndakan gorilla study site, 2.20 ° N, 16.15 ° E, 13 March 1988, Fay J. M. 8307 (MO, WAG); Eastern Province W bank of Sangha River, 2.63 ° N, 16.06 ° E, 22 May 1988, Harris D. J. 757 (K); Rives du Dja pres Ndongo a 40 km WNW de Moloundou, 2.58 ° N, 15.29 ° E, 18 March 1973, Letouzey R. 12138 (K, P, YA); Djerem R (Niadaba), 5.93 ° N, 12.88 ° E, 27 June 1959, Letouzey R. 2261 (P, YA); Bertoua-Batouri, 4.58 ° N, 13.68 ° E, 01 January 1962, Tchinaye V. 103 (P, YA); 7 km NW du confluent Boumba Dja Ngoba, 2.07 ° N, 15.23 ° E, 17 April 1971, Villiers J. - F. 666 (P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
357372A17FB052D77B7FAD3421296094.taxon	description	Fig. 148; Map 18 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
357372A17FB052D77B7FAD3421296094.taxon	materials_examined	Type. Cameroon. South Region; 45 km west of Grand Batanga, Mildbraed G. W. J. 6055, 22 Jul 1911: holotype: B [B 10 0153148]; isotypes: HBG [HBG 502478, HBG 502477].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
357372A17FB052D77B7FAD3421296094.taxon	description	Description. Tree up to 9 m tall, rarely a shrub, d. b. h. up to 9 cm; stilt roots and buttresses absent. Old branches glabrous, young branches pubescent with appressede hairs 0.1 - 0.5 mm long. Leaves: petiole 2 - 5 mm long, 1 mm wide, sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 9.5 - 17.5 cm long, 3.2 - 5.6 cm wide, elliptic to oblong, occasionally oblanceolate or lanceolate, apex acuminate, acumen 0.8 - 1.9 cm long, base cuneate, papyraceous, below pubescent when young, glabrous to sparsely pubescent when old, above glabrous when young and old, discolorous but sometimes concolorous; midrib sunken, above pubescent when young and old, below pubescent when young, glabrous when old; secondary veins 12 to 20 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 4 - 9 mm long, 1 - 2 mm in diameter, pubescent; in fruit 5 - 7 mm long, 5 - 7 mm in diameter, glabrous; bracts 3 to 4, evenly spaced, 2 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 4 - 7 mm long, 4 - 5 mm wide, triangular, apex acute, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 45 - 79 mm long, 3.6 - 5.5 mm wide at base, linear, apex obtuse, base broad and concave, cream, pubescent outside, pubescent towards base inside; inner petals 3, valvate, 35 - 61 mm long, 4.9 - 5.4 mm wide at base, linear, apex acute, base broad and concave with a tuft of long hairs at the apex of the concavity, cream, pubescent outside, pubescent inside; stamens ca. 100, in 5 to 6 rows, 1 - 2 mm long, clavate to oblong; connective apex shield-like, glabrous; carpels 10 to 11, ovary 1 - 2 mm long, stigmas connivent, linear, 4.5 - 6 mm long, pubescent. Monocarps stipitate, stipe 9 - 15 mm long, 3 - 5 mm in diameter; monocarps 1 to 14, 42 - 65 mm long, 13 - 15 mm wide, oblong, apex with a curved beak ca. 1 mm long, glabrous to sparsely pubescent, verrucose and wrinkled when dried, pale green to yellowish green outside, endocarp red to pink-red; seeds 5 to 9 per monocarp, in a single row, 13 - 14 mm long, 7 - 9 mm wide, ellipsoid; sarcotesta greenish blue, greenish white, or gray; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
357372A17FB052D77B7FAD3421296094.taxon	distribution	Distribution. From Cameroon to Gabon; in Cameroon known from the South and Littoral regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
357372A17FB052D77B7FAD3421296094.taxon	materials_examined	Specimens examined. Littoral Region: Foret de Bakaka 3 km E of Ebone a village on km 11 Nkongsamba-Loum Road, 4.83 ° N, 9.9 ° E, 13 September 1971, Leeuwenberg A. J. M. 8362 (BR, MO, P, WAG, YA). South Region: 20 km from Kribi Lolodorf road, 3.03 ° N, 10.05 ° E, 09 June 1969, Bos J. J. 4772 (B, BR, K, LD, LM, MO, P, WAG, YA); Campo-Ma'an area Bibabimvoto, 2.25 ° N, 10.09 ° E, 01 February 2000, Elad M. 1253 (KRIBI, WAG); Beson 45 km E Kribi, 2.79 ° N, 10.27 ° E, 22 July 1911, Mildbraed G. W. J. 6055 (B, HBG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
75C3F918F796F90DF6DCF20F3672BCE2.taxon	description	Map 18 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
75C3F918F796F90DF6DCF20F3672BCE2.taxon	materials_examined	Type. Nigeria. Taraba [" N. E. State "] State, Sardauna Province, Kurmin Kugapa, below Kurmin Dodo below the western edge of Cabbal Wade [" Chappal Waddi "], ca. 5500 ft, 28 Feb 1975, Chapman J. D. 3755 Holotype: K.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
75C3F918F796F90DF6DCF20F3672BCE2.taxon	description	Description. Tree up to 10 m (- 20 m) tall, d. b. h. unknown; stilt roots and buttresses absent. Old branches glabrate, young branches pubescent with erect dull gray to brown hairs 0.1 - 1.2 mm long. Leaves: petiole 3.5 - 9 mm long, pubescent, grooved; blades 7.8 - 10.9 cm long, 2.7 - 5.1 cm wide, lanceolate, lanceolate-oblong, elliptic to oblong or oblong-oblanceolate, apex blunt-acuminate, acumen 0.4 - 1.6 cm long, base broadly cuneate to nearly rounded, papyraceous to subcoriaceous, below sparsely pubescent to glabrate, above glabrous, concolorous; midrib flat, above pubescent, below pubescent; secondary veins 7 - 13 pairs; tertiary venation conspicuously reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 5.2 - 8.1 mm long, 1 - 1.3 mm in diameter; in fruit 6 - 8.7 mm long, 1.5 - 2.8 mm thick, sparsely pubescent, sometimes with bracts or sepals persistent; bracts 3 - 4, evenly spaced, 2 - 3.1 mm long, ovate; sepals 3, valvate, basally fused, 2.7 - 3.5 mm long, 2.9 - 3.1 mm wide, triangu-lar to ovate, apex acute, pubescent outside; outer petals 3, (15 -) 36 - 52 mm long, 2.7 - 3.6 mm wide at base, linear, apex acute, base concave, cream, pubescent inside, sericeous outside; inner petals 3, valvate, (15.5 -) 29 - 37 mm long, 2 - 2.6 mm wide at base, linear, apex obtuse, base concave, cream, pubescent on both surfaces except for glabrous base; stamens 160 - 200, 1.8 - 2.2 mm long, narrowly oblong, connective apex shield-like, glabrous; carpels 7 to 8, ovary 1.2 - 1.3 mm long, stigma loosely connivent, linear, 2 - 3.4 mm long, glabrous except for apical hair tuft. Monocarps stipitate, stipe 8 - 13 mm long, 2.1 - 2.6 mm in diameter, monocarps 4 to 8, 34 - 40 mm long, 6 - 10 mm wide, narrowly oblong and slightly falciform, torulose, apex obtuse or with a beak 0.5 - 2 mm long, glabrate, verrucose, green outside, endocarp red; seeds up to 5 per monocarp, in a single row, 11 - 13.1 mm long, 5.8 - 6.5 mm wide, oblongoid; sarcotesta " glaucous " glaucous "; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
75C3F918F796F90DF6DCF20F3672BCE2.taxon	distribution	Distribution. Easternmost Nigeria and adjoining Cameroon (South-West region).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
75C3F918F796F90DF6DCF20F3672BCE2.taxon	materials_examined	Specimens examined. South-West Region: Near Aguosho 10 km SSW of Akwaya, 6.3 ° N, 9.466 ° E, 20 March 1985, Thomas D. W. 4558 (MO); Takamanda Forest Reserve along footpath from Malishi to Kalu 6.15 ° N, 9.433 ° E, 01 May 1987, Thomas D. W. 7400 (B, K, MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0366F727636BB015720414F3DFB1F0D1.taxon	description	Map 18 F	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0366F727636BB015720414F3DFB1F0D1.taxon	materials_examined	Type. Angola. Cabinda; Belize, Mayumbe, Gossweiler J. 6988, no date: holotype: BM; isotypes: COI [COI 00004886]; LISC [LISC 000323, LISC 000324, LISC 000322, LISC 000321]	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0366F727636BB015720414F3DFB1F0D1.taxon	description	Description. Tree, up to 35 m tall, d. b. h. up 35 cm; stilt roots or small buttresses present. Old branches glabrous, young branches glabrous to pubescent with loosely appressed hairs 0.4 - 0.9 mm long. Leaves: petiole 2 - 3 mm long, 1 mm wide, pubescent, grooved, blade inserted on the side of the petiole; blade 7.2 - 11.5 cm long, 1.8 - 2.7 cm wide, narrowly oblong to elliptic, apex acuminate, acumen 0.2 - 0.7 cm long, base decurrent to cuneate, papyraceous to subcoriaceous, below sparsely pubescent when young, sparsely pubescent when old, above glabrous when young and old, somewhat shiny above, slightly discolorous; midrib raised to slightly sunken, above sparsely pubescent when young, glabrous when old, below pubescent when young, sparsely pubescent when old; secondary veins 8 to 16 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 2 - 3 per axil, highly branched, 1 - 4.5 mm long, sometimes with a longer floriferous axis emerging from among the cluster of flowers in an axil. Flowers with 9 perianth parts in 3 whorls, up to 32 per inflorescence; pedicel 2 - 3 mm long, ca. 1 mm in diameter, pubescent; in fruit 23 - 30 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 1 or 2, at or above the middle of pedicel, 2 - 3 mm long, ca. 1 mm wide; sepals 3, valvate, basally fused, 2 - 3 mm long, 2 - 3 mm wide, ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, inner and outer whorl subequal; outer petals 3, 10 - 19.7 mm long, 2.3 - 3.6 mm wide at base, linear-lanceolate, apex obtuse, base broad and concave, yellow-green with red base, densely pubescent outside, pubescent and glabrous towards center inside; inner petals 3, valvate, 9.7 - 13.1 mm long, 2.2 - 2.5 mm wide at base, linear, apex acute, base broad and concave, yellow-green with red base, pubescent, base glabrous outside, pubescent, base glabrous inside; stamens 120 to 130, in 5 to 6 rows, 1 - 2 mm long, narrowly oblong; connective apex capitate, glabrous; carpels 3 to 6, ovary ca. 1 mm long, stigmas connivent, filiform, 2.5 - 2.7 mm long, sparsely pubescent. Monocarp stipitate, stipe 2 - 3 mm long, 2 - 3 mm in diameter; monocarp 1, 56 - 85 mm long, 31 - 40 mm wide, obovoid to oblongoid, apex rounded, pubescent, longitudinally ridged and wrinkled when dried, green outside, endocarp carmine, red or pink-red; seeds 3 to 4 per monocarp, in a single row, 21 - 22 mm long, 16 - 17 mm wide, flattened ellipsoid; sarcotesta grayish blue or greenish blue in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
0366F727636BB015720414F3DFB1F0D1.taxon	distribution	Distribution. A sparsely distributed species from Cameroon to Angola (Cabinda); in Cameroon known from a single collection in the East region.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1414A0B38776CE8C11629F5F8CFC774F.taxon	description	Figs 148, 149; Map 18 G	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1414A0B38776CE8C11629F5F8CFC774F.taxon	materials_examined	Type. Cameroon. South Region; between Bipindi and Ebolowa near Malakat, Mildbraed G. W. J. 7592, Dec 1913: lectotype, designated by Le Thomas (1969 b), p. 185: B [B 10 0153152]; isolectotype: K [K 000199053].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1414A0B38776CE8C11629F5F8CFC774F.taxon	description	Description. Tree, 30 - 35 m tall, d. b. h. up to 80 cm; buttresses present, up to 1.5 m high, stilt roots rarely present. Old branches glabrous, young branches glabrous to densely pubescent with appressed hairs 0.1 - 0.4 mm long. Leaves: petiole 4 - 8 mm long, ca. 2 mm wide, pubescent, grooved, blade inserted on the side of the petiole; blade 5.7 - 17.2 cm long, 1.9 - 5.9 cm wide, elliptic, oblong-elliptic or lanceolate, apex acuminate to obtuse, acumen 0.5 - 1.0 cm long, base cuneate to subcordate, subcoriaceous, below sparsely pubescent when young, sparsely pubescent when old, above glabrous when young and old, concolorous or slightly discolorous; midrib sunken, above glabrous when young, glabrous to sparsely pubescent when old, below pubescent when young, sparsely pubescent when old; secondary veins 10 to 16 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate or old leafless branches, axillary, peduncle 2 - 4 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 10 per inflorescence; pedicel 3 - 6 mm long, ca. 1 mm in diameter, pubescent; in fruit 5 - 16 mm long, 5 - 8 mm in diameter, glabrous; bracts 2, towards or above the middle of pedicel, 2 - 3 mm long, 2 - 3 mm wide; sepals 3, valvate, basally to 2 / 3 fused, forming a cup, 2 - 4 mm long, 3 mm wide, triangular to ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 19 - 23 mm long, 3 - 4.3 mm wide at base, linear-lanceolate, apex obtuse, base broad and concave, cream-colored or pale yellow with a purple blotch at the base, sericeous outside, pubescent with base glabrous inside; inner petals 3, valvate, 12.5 - 21 mm long, 2 - 3 mm wide, linear, apex acute, base broad and concave, light yellow to cream, pubescent with glabrous base on both sides; stamens 120, in 5 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 5 to 8, ovary 1 - 2 mm long, stigmas connivent, linear, slightly widened at midpoint, 2.1 - 3 mm long, glabrous to sparsely pubescent. Monocarps sessile; monocarps 4 to 9, 27 - 43 mm long, 17 - 24 mm wide, ovoid, broadly ellipsoid or oblongoid, apex rounded, glabrous, rugose to verrucose, conspicuously lenticellate, wrinkled when dried, brown outside, endocarp light pink; seeds 7 to 12 per monocarp, in two rows, 16 - 21 mm long, 7 - 10 mm wide, ellipsoid; sarcotesta orange, fleshy in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1414A0B38776CE8C11629F5F8CFC774F.taxon	distribution	Distribution. A central African species, from southern Nigeria to Republic of Congo and northern Democratic Republic of the Congo; in Cameroon known from East, South and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1414A0B38776CE8C11629F5F8CFC774F.taxon	materials_examined	Specimens examined. Central Region: Bank Nyong River near the new bridge ca 65 km SSW of Eseka, 3.65 ° N, 10.78 ° E, 16 July 1964, de Wilde J. J. F. E 2838 A (P); Bank Nyong River near the new bridge ca 65 km SSW of Eseka, 3.65 ° N, 10.78 ° E, 16 July 1964, de Wilde J. J. F. E 2838 B (B, BR, K, L, MO, P, P, WAG, YA); Edea, 3.8 ° N, 10.13 ° E, 26 August 1955, Mpom B. 121 (P, YA). East Region: 63 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.07 ° N, 14.65 ° E, 07 March 2019, Couvreur T. L. P. 1222 (MPU, WAG, YA); Ndakan Sango River, 2.36 ° N, 16.11 ° E, 10 May 1988, Gentry A. H. 62649 (MO); A 25 km environ a l'ENE de Mikel village situe a environ 85 km au N de Moloundou sur route de Yokadouma, 2.93 ° N, 15.33 ° E, 24 February 1971, Letouzey R. 10418 (P, YA); Nkonbong II sur axe Lomie-Ngoila-Swanke a 15 km au SSW de Ngoila, 2.50 ° N, 14.00 ° E, 22 February 1973, Letouzey R. 12026 (P, WAG, YA); Pres Ngola (30 km a l'Est de Yokadouma), 3.47 ° N, 15.32 ° E, 11 May 1963, Letouzey R. 5027 (P, YA); A 24 km a l'Ouest de Masea (village situe a 50 km au SSW de Yokadouma), 3.15 ° N, 14.87 ° E, 04 July 1963, Letouzey R. 5401 (P, YA). Littoral Region: Tissongo strip, 3.57 ° N, 9.869 ° E, 26 July 1976, McKey D. B. 138 (K); Douala-Edea Reserve Tissongo study area Transect C, 3.57 ° N, 9.869 ° E, 21 June 1976, Waterman P. G. 880 (K). South Region: 17 km before Kribi on N 7 road from Ed 3.14 ° N, 9.954 ° E, 27 February 2018, Couvreur T. L. P. 1164 (K, MPU, P, WAG, YA); Bezirk Kribi Vorland mit einzeln Hugeln bei Adjab 35 km ostlich Gross-Batanga, 2.82 ° N, 10.2 ° E, 24 July 1911, Mildbraed G. W. J. 6090 (B); Bipindi - Ebolowa, 3.00 ° N, 10.92 ° E, 01 December 1913, Mildbraed G. W. J. 7592 (B).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50FF5926A5288FA3E1E02D0FC0A77824.taxon	description	Fig. 146; Map 18 H	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50FF5926A5288FA3E1E02D0FC0A77824.taxon	materials_examined	Type. Democratic Republic of the Congo. Equateur; Eala, Pynaert L. A. E. J. 567, 15 Oct 1906: lectotype, designated by Johnson and Murray (2018), p. 201: BR [BR 0000024941426]; isolectotypes: BR [BR 0000008825339, BR 0000008825346].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50FF5926A5288FA3E1E02D0FC0A77824.taxon	description	Description. Tree, up to 35 m tall, d. b. h. up to 40 cm; buttresses present, ca. 0.5 m tall, upper bark red, rough, scaly. Old branches glabrous, young branches pubescent, the hairs 0.4 - 1.5 mm long. Leaves: petiole 1 - 3 mm long, 1 mm wide, pubescent, grooved, blade inserted on the side of the petiole; blade 3.6 - 8.7 cm long, 1.2 - 2.3 cm wide, lanceolate, ovate or elliptic, apex acute to obtuse, base cuneate, papyraceous to subcoriaceous, below pubescent when young, sparsely pubescent when old, above glabrous when young and old, discolorous or occasionally concolorous; midrib sunken, above glabrous to pubescent when young, glabrous to pubescent when old, below pubescent when young and old; secondary veins 8 to 13 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle ca. 1 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 2 per inflorescence; pedicel 4 - 5 mm long, ca. 1 mm in diameter, pubescent; in fruit 3 - 24 mm long, 2 - 3 mm in diameter, sparsely pubescent; bracts 2, evenly spaced, 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, basally to ½ / 2 fused, forming a cup, 2 - 3 mm long, ca. 2 mm wide, triangular to ovate, apex acute, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 15.2 - 20.5 mm long, 2.4 - 3 mm wide at base, linear, apex acute to obtuse, base broad and concave, light yellow to white, sericeous outside, pubescent but glabrous towards the base inside; inner petals 3, valvate, 12.3 - 16.8 mm long, 2.2 - 3.2 mm wide at base, linear, apex acute, base broad and concave, light yellow to white, pubescent with base glabrous on both sides; stamens ca. 140, in 4 to 5 rows, ca. 1 mm long, oblong; connective apex shield-like, glabrous; carpels 9 to 11, ovary ca. 1 mm long, stigmas connivent, linear, 2.5 - 3.8 mm long, pubescent. Monocarps sessile, or stipitate with the stipe 2 - 8 mm long, 4 - 6 mm in diameter; monocarps 4 to 8, 26 - 48 mm long, 13 - 18 mm wide, obovoid to oblongoid, apex rounded, glabrous, verrucose, wrinkled when dried, green outside, endocarp red; seeds 5 to 6 per monocarp, in two rows, 9 - 13 mm long, 6 - 7 mm wide, ellipsoid to flattened ellipsoid; sarcotesta white to grayish blue in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50FF5926A5288FA3E1E02D0FC0A77824.taxon	distribution	Distribution. Sparsely occurring from eastern Nigeria to the Democratic Republic of the Congo; in Cameroon known from two collections in the South and Central regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
50FF5926A5288FA3E1E02D0FC0A77824.taxon	materials_examined	Specimens examined. Central Region: Pont Kelle (20 km N d'Eseka), 3.65 ° N, 10.78 ° E, 09 December 1973, Letouzey R. 12317 (K, P, YA). South Region: Massif de Ngovayang village de Atog Boga, 3.15 ° N, 10.29 ° E, 30 August 2015, Droissart V. 2049 (BRLU, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
566F0A9E50A97AED8FE6EB355393E059.taxon	description	Fig. 150; Map 18 I	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
566F0A9E50A97AED8FE6EB355393E059.taxon	materials_examined	Type. Sao Tome and Principe. Insel St. Thome, bei Angolares, Quintas F. 3, Jan 1886: lectotype, designated by Johnson and Murray (2018), p. 68: K [K 000199059]; isolectotype: COI [COI 00004887].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
566F0A9E50A97AED8FE6EB355393E059.taxon	description	Description. Tree, 10 - 42 m tall, d. b. h. up to 75 cm; buttresses present, narrow and thin. Old branches glabrous, young branches glabrous to sparsely pubescent, the hairs 0.1 - 0.2 mm long. Leaves: petiole 3 - 7 mm long, ca. 1 mm wide, glabrous to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 6 - 11.9 cm long, 2.6 - 5.5 cm wide, obovate to oblanceolate, occasionally elliptic, apex acuminate, acumen 0.2 - 0.6 cm long, base cuneate and decurrent, papyraceous to subcoriaceous, below glabrous when young, glabrous to sparsely pubescent when old, above glabrous when young and old, discolourous, often tan-colored below; midrib sunken, above glabrous when young and old, below glabrous when young, glabrous when old; secondary veins 7 to 12 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate or old leafless branches, axillary, peduncle sometimes present, 2.6 - 2.8 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 7 per inflorescence; pedicel 7 - 9 mm long, ca. 1 mm in diameter, pubescent; in fruit 8 - 19 mm long, 2 mm in diameter, glabrous; bracts (2) 3 (4), evenly spaced, 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, free to basally fused, ca. 2 mm long, 2 - 1 mm wide, triangular to ovate, apex acute to obtuse, base truncate, green, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 8 - 15 mm long, 1.8 - 2.8 mm wide at base, ligulate, apex obtuse, base broad and concave, cream to light green, pubescent outside, pubescent and glabrous towards the base inside; inner petals 3, valvate, 7.1 - 13.2 mm long, 0.7 - 1.5 mm wide, linear, apex obtuse, base broad and concave, lacking an internal tooth overhanging the concavity, with fleshy glandlike margins, cream to light green, pubescent outside, pubescent but glabrous towards the base inside; stamens 50 to 80, in 3 to 4 rows, 1 - 2 mm long, oblong; connective apex conical, glabrous; carpels 3 to 5, ovary 1 - 2 mm long, stigmas separate, clavate, ca. 0.5 mm long, glabrous. Monocarps stipitate, stipe 7 - 11 mm long, 2 - 4 mm in diameter; monocarps 1 to 4, 35 - 64 mm long, 7 - 10 mm wide, narrowly oblongoid, apex with a beak 1.5 - 4 mm long, glabrous, obliquely striate, occasionally verrucose when dried, slightly constricted around seeds, green outside tinged with brown, endocarp green; seeds 3 (5) monocarp, in a single row, 10 - 13 mm long, 6 - 7 mm wide, ellipsoid; sarcotesta absent; aril present, fimbriate, extending over the length of the seed, membranous, red to deep orange.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
566F0A9E50A97AED8FE6EB355393E059.taxon	distribution	Distribution. A widespread species in West and Central Africa, from Sierra Leone to northwestern Democratic Republic of the Congo and south to northern Angola, also on the island of Sao Tome; in Cameroon known from East, South, Central, Littoral, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
566F0A9E50A97AED8FE6EB355393E059.taxon	materials_examined	Selected specimens examined. Central Region: Chantier forestier au sud de Song Bon (SE d'Eseka), 3.42 ° N, 10.49 ° E, 08 December 1967, Bamps P. R. J. 1376 (P, YA); Ca 5 O km NW of Eseka W of Yaounde on opposite of the Kele River, 3.65 ° N, 10.78 ° E, 23 November 1963, de Wilde J. J. F. E 1320 (BR, K, P, WAG, YA); ca 50 km NW of Eseka W of Yaounde on opposite [side?] of the Kele-river, 3.75 ° N, 10.61 ° E, 01 November 1963, de Wilde W. J. J. O 1320 (K, P, WAG); Yaounde, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 36 (P); M´balmayo, 3.52 ° N, 11.5 ° E, 28 October 1959, Mpom B. 353 (P, YA). East Region: 28 km SW of Bertoua near Toungrelo, 4.58 ° N, 13.68 ° E, 12 January 1961, Breteler F. J. 2398 (A, BR, G, K, M, P, SL, U, UC, WAG, YA); A 26 km au SSW de Koso (village situe a 60 km au SSW de Batouri), 3.93 ° N, 14.17 ° E, 27 July 1963, Letouzey R. 5510 (K, P, YA); Moloundou, 2.05 ° N, 15.17 ° E, 06 December 1910, Mildbraed G. W. J. 4003 (P); south of Sanaga between Yaounde and Deng Deng, between the union of the Lom (Sanaga) and Djerem rivers about, 125 km NE of Yaounde, 5.33 ° N, 13.4 ° E, 01 February 1914, Mildbraed G. W. J. 8294 (K). South Region: Bipindi, 3.26 ° N, 10.20 ° E, 23 June 1918, Annet E. 319 (P); 22 km from Kribi Lolodorf road, 3.05 ° N, 10.05 ° E, 09 June 1969, Bos J. J. 4773 (BR, K, LD, LM, MO, P, POZG, WAG, YA); 28 km east from Lele village, 2.27 ° N, 13.29 ° E, 09 September 2013, Couvreur T. L. P. 483 (WAG, YA); ca 15 km east from Lele village, 2.29 ° N, 13.33 ° E, 10 September 2013, Couvreur T. L. P. 500 (WAG, YA); Hill facing the village of N'koladom, 2.80 ° N, 11.16 ° E, 03 January 1975, de Wilde J. J. F. E 7871 (B, BR, K, MO, P, U, WAG, YA); Hill roughly between Nkolandom and Nkoemvone, 2.8 ° N, 11.15 ° E, 09 January 1975, de Wilde J. J. F. E 7889 (B, BR, K, MO, P, U, WAG, YA); TDC Sud Cameroun, 2.65 ° N, 9.9 ° E, 14 November 1991, Halle F. 4240 (WAG); Bipindi - Ebolowa, 3.00 ° N, 10.92 ° E, 01 December 1913, Mildbraed G. W. J. 7613 (K); [Locality not legible- but should be Bedai], 2.17 ° N, 11.12 ° E, December 1908 [?], Tessmann G. 760 (K); Campo-Ma'an area 2.56 ° N, 9.833 ° E, 16 April 2001, van Andel T. R. 3345 (KRIBI, WAG, YA); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1900, Zenker G. A. 2080 (L, P, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1914, Zenker G. A. 2094 (B, GH, M, MO, P, WAG); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1903, Zenker G. A. 2655 (L, WAG); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1903, Zenker G. A. 2663 (P); Mimfia, 3.06 ° N, 10.38 ° E, 01 June 1913, Zenker G. A. 359 (P, U); Bipindi, 3.08 ° N, 10.41 ° E, 01 October 1913, Zenker G. A. 408 (P, U); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1911, Zenker G. A. 4096 (L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1913, Zenker G. A. 4738 (L, P); Mimfia, 3.06 ° N, 10.38 ° E, 01 May 1914, Zenker G. A. 580 (P, U). South-West Region: forest near Ngusi village north of Nyassosso, 4.88 ° N, 9.7 ° E, 26 April 1986, Etuge M. 56 (B, K, MO, WAG, YA); Korup National Park, 5.02 ° N, 8.8 ° E, 01 January 1998, Kenfack D. 1187 (MO); mile 12 Mamfe road between Kumba and Badu 4.75 ° N, 9.48 ° E, 01 October 1986, Nemba J. 293 (MO, P, WAG). West Region: Piste Santchou-Bale 18 km SSW de Dschang, 5.26 ° N, 10.04 ° E, 26 November 1974, Letouzey R. 13327 (P, YA).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
31B9710F6BE54931C5FD3183C70B0803.taxon	description	Fig. 151; Map 19 A	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
31B9710F6BE54931C5FD3183C70B0803.taxon	materials_examined	Type. Nigeria. Rivers state; Old Calabar, Thomson W. C. 53, no date: holotype: K [K 000199073].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
31B9710F6BE54931C5FD3183C70B0803.taxon	description	Description. Tree, up to 30 m tall, d. b. h. up to 70 cm; stilt roots present, up to 2 m on the trunk. Old branches glabrous, turning grayish white, young branches glabrous to pubescent with appressed golden hairs 0.2 - 0.5 mm long. Leaves: petiole 5 - 12 mm long, ca. 2 mm wide, glabrous to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 7.3 - 21.3 cm long, 3.6 - 8.4 cm wide, oblong, elliptic, or oblanceolate, apex acuminate to cuspidate, acumen 0.2 - 1.5 cm long, base cuneate to rounded, decurrent, papyraceous to subcoriaceous, below sparsely appressed-pubescent, rarely glabrous, above glabrous, strongly discolorous; midrib flat to sunken, above glabrous when young and old, below glabrous to sparsely pubescent when young, glabrous when old; secondary veins 8 to 16 pairs, glabrous above; tertiary venation reticulate. Flowers bisexual with 9 perianth parts in 3 whorls. Individuals bisexual; inflorescences ramiflorous on young foliate or more often clustered on older leafless branches, axillary, peduncle 1.5 - 2.5 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 4 - 9 mm long, ca. 1 mm in diameter, pubescent; in fruit 9 - 15 mm long, 2 - 7 mm in diameter, glabrous; bracts (2) 3 (5), evenly spaced, 1 - 3 mm long, 1 - 3 mm wide; sepals 3, valvate, basally fused, 2 - 4 mm long, 3 - 4 mm wide, ovate to broadly triangular, apex acute, base truncate, green, sericeous outside, glabrous inside; petals free, outer petals much longer than inner; outer petals 3, 17.6 - 35 mm long, 2.5 - 5.1 mm wide at base, linear, apex acute, base broad and concave, yellow, pubescent outside, pubescent and glabrous towards the base inside; inner petals 3, valvate, 3.5 - 6.7 mm long, 2.2 - 4.6 mm wide, ovate to rhombic, apex acuminate, base broad and concave, red with cream base and apex, pubescent outside, pubescent and glabrous towards the base inside; stamens 70 to 77, in 4 to 5 rows, ca. 2 mm long, clavate to oblong; connective apex shield-like, glabrous; carpels 4 to 12, ovary 1 - 2 mm long, stigmas loosely connivent, linear, 1.4 - 2.9 mm long, with glandular appendages. Monocarps stipitate, stipe 4 - 20 mm long, 2 - 6 mm in diameter; monocarps 1 to 15, 41 - 16 mm long, 6 - 12 mm wide, narrowly oblong, somewhat falciform and strongly torulose to moniliform, apex with a distinct beak up to 5 mm long, glabrous, verrucose and wrinkled when dried, black or dark purple outside, endocarp pink to scarlet; seeds 1 to 7 per monocarp, in a single row, 10 - 20 mm long, 6 - 11 mm wide, ellipsoid; sarcotesta absent; aril present, brushlike, unlobed, red to orange.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
31B9710F6BE54931C5FD3183C70B0803.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West Africa, from Guinea-Bissau to Ghana, and in Central Africa from Nigeria to southern South Sudan and south to northern Angola and Mozambique; in Cameroon known from East, South, Littoral and regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
31B9710F6BE54931C5FD3183C70B0803.taxon	materials_examined	Specimens examined. East Region: Marecage du Niagoul entre Koumbou et Miambo, 4.27 ° N, 12.9 ° E, 13 April 1959, Letouzey R. 1611 (P, YA). Littoral Region: Duala, 4.05 ° N, 9.71 ° E, 01 June 1917, Fleury F. 33135 (P). South Region: Batanga, 2.86 ° N, 9.889 ° E, 20 September 1945, Aubreville A. 125 (P); Bitye Yaunde, 3.02 ° N, 12.37 ° E, 01 January 1919, Bates G. L. 1317 (BM, MO); N´Koemvone, 2.8 ° N, 11.13 ° E, 11 April 1975, de Wilde J. J. F. E 8166 (B, BR, K, MO, P, WAG, YA); Pres Akok Bikele, 3.8 ° N, 12.7 ° E, 03 March 1962, Letouzey R. 4464 (YA); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1904, Zenker G. A. 2827 (G, L, M, WAG). Unknown region: Vuneli, 01 February 1928, Hedin L. 1668 (P, OWU).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0FA2AF7BBFCB9FBB5BD9850F2476C66.taxon	description	Figs 150, 152; Map 19 B	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0FA2AF7BBFCB9FBB5BD9850F2476C66.taxon	materials_examined	Type. Cameroon. South-West Region; Johann-Albrechtshohe [Kumba], Staudt A. 530, 1896: holotype: B; isotypes K [K 000105614]; P [00169112, 00169113].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0FA2AF7BBFCB9FBB5BD9850F2476C66.taxon	description	Description. Tree, up to 30 (50) m tall, d. b. h. up to 80 cm; stilt roots and small buttresses present. Old branches glabrous, young branches glabrous to pubescent with appressed hairs 0.3 - 0.4 mm long. Leaves: petiole 3 - 9 mm long, 2 mm wide, glabrous to sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 5.1 - 11.8 cm long, 2 - 5.6 cm wide, oblanceolate to obovate, occasionally elliptic, apex acuminate, acumen 0.2 - 0.3 cm long, base cuneate and decurrent, coriaceous to subcoriaceous, below sparsely pubescent when young, glabrous when old, above glabrous when young and old, discolorous; midrib sunken, above glabrous when young and old, below glabrous when young, glabrous when old; secondary veins 7 to 11 pairs, above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 0.5 - 2.4 mm long. Flowers with 9 perianth parts in 3 whorls, 1 to 3 per inflorescence; pedicel 3 - 8 mm long, ca. 1 mm in diameter, sparsely pubescent to glabrous; in fruit 7 - 13 mm long, 3 - 8 mm in diameter, sparsely pubescent to glabrous; bracts 2 to 4, evenly spaced, 1 - 2 mm long, 1 - 2 mm wide; sepals 3, valvate, basally to ½ / 2 fused, forming a cup, 2 - 3 mm long, 3 - 4 mm wide, ovate, apex acute to obtuse, base truncate, green, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 5.8 - 9.6 mm long, 4 - 5.2 mm wide at base, ovate, apex acute, base truncate, yellow, pubescent outside, pubescent but glabrous towards the base inside; inner petals 3, valvate, 4.1 - 8.4 mm long, 2 - 3.2 mm wide at base, elliptic to rhombic, apex acute, base truncate, yellow, pubescent outside, pubescent towards base inside; stamens 100 to 120, in 5 to 6 rows, 2 mm long, clavate; connective apex shield-like with a central conical point, pubescent, bright yellow; carpels 3 to 11, ovary 1 - 2 mm long, stigmas loosely connivent or separate, linear, 2.6 - 4.6 mm long, tuberculate and sparsely pubescent. Monocarps stipitate, stipe 7 - 15 mm long, 3 - 6 mm in diameter; monocarps 3 to 5, 37 - 98 mm long, 12 - 21 mm wide, cylindrical, apex rounded to sometimes with a curved beak 1.3 - 3 mm long, glabrous to sparsely pubescent, sometimes slightly constricted around seeds and verrucose and wrinkled when dried, green outside, endocarp scarlet; seeds 1 to 5 per monocarp, in a single row, 14 - 19 mm long, 9 - 13 mm wide, ellipsoid; sarcotesta absent; aril present, brushlike, bright yellow to orange.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0FA2AF7BBFCB9FBB5BD9850F2476C66.taxon	distribution	Distribution. A widespread species with a disjunct distribution in West Africa, from Sierra Leone to Ghana, and in Central Africa from eastern Nigeria to the Democratic Republic of the Congo, western Uganda and Cabinda (Angola); in Cameroon known from East, South, Central, Littoral, South-West and West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
B0FA2AF7BBFCB9FBB5BD9850F2476C66.taxon	materials_examined	Specimens examined. Central Region: Yaounde, 3.87 ° N, 11.52 ° E, 01 January 1935, Foury P. 46 (P); 21 km NO d'Oveng, 3.7 ° N, 11.37 ° E, 24 October 1966, Letouzey R. 8178 (L, P, YA). East Region: Dja Reserve (Reserve de Faune du Dja) Bouamir Research Area 90 km southeast of Akonolinga, 3.41 ° N, 12.96 ° E, 24 October 1994, Fogiel M. K. 864 (MO, YA); Colline a l'ENE de Mbalam (140 km ESE de Djoum pres Souanke-Congo), 2.22 ° N, 13.82 ° E, 20 January 1973, Letouzey R. 11865 (P, YA); 23 km a l'ouest de Masea (village situe a 50 km au SSW de Yokadouma), 3.15 ° N, 14.87 ° E, 04 July 1963, Letouzey R. 5404 (P, YA). Littoral Region: Makak, 3.87 ° N, 10.62 ° E, 22 January 1945, Letouzey R. 1126 (P). South Region: hill above Nlonacko near village Ebianemeyong, 2.43 ° N, 10.35 ° E, 12 December 1998, de Wilde J. J. F. E 12161 (BR, KRIBI, MO, S, WAG); Station de cacaoyer de N'koemvone 14 km On the road from Ebolowa to Ambam, 2.81 ° N, 11.13 ° E, 31 January 1975, de Wilde J. J. F. E 7941 (B, BR, K, MO, P, U, WAG, YA); N´Koemvone, 2.81 ° N, 11.13 ° E, 04 March 1975, de Wilde J. J. F. E 8029 (B, BR, K, MO, P, U, WAG, YA); Pres Nteigne PK 108 sur route Mintom I (70 km E de Djoum), 2.2 ° N, 13.81 ° E, 19 January 1973, Letouzey R. 11854 (P, YA); Bebai- Camposgebiet- Weg u [locality reported as being in Equatorial Guinea], 2.17 ° N, 11.12 ° E, 18 November 1908, Tessmann G. 644 (K); Mimfia, 3.06 ° N, 10.38 ° E, 01 March 1913, Zenker G. A. 246 (P, U); Bipindi, 3.08 ° N, 10.41 ° E, 01 January 1908, Zenker G. A. 3653 (L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1911, Zenker G. A. 3953 (L, P); Bipindi, 3.08 ° N, 10.42 ° E, 01 January 1913, Zenker G. A. 4862 (L, P). South-West Region: Johann-Albrechtshohe [Kumba] area 4.16 ° N, 9.2 ° E, 1896, Staudt A. 530 (K, P); Mount above Batoke, 4.08 ° N, 9.083 ° E, 24 April 1984, Thomas D. W. 3463 (MO, P, WAG, YA); Takamanda Forest Reserve footpath from Mbilishi to Kalu 6.25 ° N, 9.43 ° E, 01 May 1987, Thomas D. W. 7401 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C40394580136DF88CBBA2A2F971F8B16.taxon	description	Map 19 C	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C40394580136DF88CBBA2A2F971F8B16.taxon	materials_examined	Type. Nigeria. Cross River State; Oban, Talbot P. A. 1601, 1912: holotype: BM; isotype: K [000199068].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C40394580136DF88CBBA2A2F971F8B16.taxon	description	Description. Tree, up to 5 m tall, d. b. h. unknown; stilt roots and buttresses absent. Old branches glabrous to pubescent, young branches rusty-pubescent, with sparser erect hairs 1.5 - 2 mm long mixed with denser erect hairs 0.2 - 0.4 mm long. Leaves: petiole 3 - 5 mm long, ca. 1 mm wide, rusty-pubescent, grooved, blade inserted on the side of the petiole; blade 9.2 - 12.5 cm long, 3 - 4.2 cm wide, lanceolate-oblong to elliptic, rarely oblanceolate, apex acuminate, acumen 0.9 - 1.7 cm long, base cuneate to rounded, papyraceous, below pubescent when young, densely pubescent to pubescent when old, above glabrous when young and old, concolorous to discolorous; midrib sunken or flat, above densely pubescent when young, pubescent when old, below pubescent when young and old; secondary veins 11 to 15 pairs, glabrous, above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 per inflorescence; pedicel 3 - 6 mm long, 1 - 2 mm in diameter, rusty-pubescent; in fruit 3 - 6 mm long, 1 - 2 mm in diameter, glabrous; bracts 2 to 3, evenly spaced, 3 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 4 - 6 mm long, 3 - 4 mm wide, triangular to ovate, apex acute to acuminate, base truncate, densely rusty-pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 32 - 38 mm long, 3.1 - 3.3 mm wide at base, linear, apex acute to obtuse, base broad and concave, color unknown, sericeous outside, pubescent and glabrous towards the base inside; inner petals 3, valvate, 16 - 33 mm long, 2.1 - 3.2 mm wide at base, linear, apex acute, base truncate, color unknown, pubescent, base glabrous outside, pubescent and glabrous towards the base inside; stamens ca. 140, in rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels ca. 8, ovary 1 mm long, stigmas connivent, linear, bent at midpoint, 2.5 - 3 mm long, glabrous. Monocarps (only immature ones seen) sessile; monocarps ca. 8, 10 - 30 mm long, 9 - 25 mm wide, apex with a small curved beak, sparsely pubescent; seeds unknown.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C40394580136DF88CBBA2A2F971F8B16.taxon	distribution	Distribution. Eastern Nigeria and Cameroon; in Cameroon known from South and South-West regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
C40394580136DF88CBBA2A2F971F8B16.taxon	materials_examined	Specimens examined. South Region: ca 10 km From Kribi S of Lolodorf road, 2.96 ° N, 9.966 ° E, 27 May 1969, Bos J. J. 4650 (BR, P, WAG, YA); Bebai - Camposgebiet - Weg u [locality reported as being in Equatorial Guinea], 2.17 ° N, 11.12 ° E, December 1908, Tessmann G. 747 (K). South-West Region: Korup National Park, 5.05 ° N, 8.8 ° E, 03 April 1984, Thomas D. W. 3204 (K, MO, P).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1D5CABBDB6DB4929A23FB1CDDEF788CF.taxon	description	Fig. 153; Map 19 D	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1D5CABBDB6DB4929A23FB1CDDEF788CF.taxon	materials_examined	Type. Nigeria. Rivers state; Old Calabar, Thomson W. C. 53, no date: holotype: K [000199064].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1D5CABBDB6DB4929A23FB1CDDEF788CF.taxon	description	Description. Scandent shrub up to 10 m tall or rarely an upright tree, d. b. h. up to 17 cm; stilt roots and buttresses absent. Old branches pubescent, young branches pubescent with erect dull gray to brown hairs 0.1 - 1.3 mm long. Leaves: petiole 2 - 6 mm long, ca. 1 mm wide, sparsely pubescent, grooved, blade inserted on the side of the petiole; blade 8.4 - 13.7 cm long, 2.5 - 4.7 cm wide, lanceolate, elliptic to oblong, or oblong-oblanceolate, apex acuminate, acumen 0.3 - 1.6 cm long, base cuneate to rounded, papyraceous, below glabrous to sparsely pubescent when young, glabrous to sparsely pubescent when old, above glabrous when young and old, generally discolorous; midrib sunken, above pubescent when young, glabrous when old, below pubescent when young and old; secondary veins 11 to 17 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate or old leafless branches, axillary, peduncle absent. Flowers with 9 perianth parts in 3 whorls, 1 (2) per inflorescence; pedicel 3 - 8 mm long, ca. 1 mm in diameter, pubescent; in fruit 5 - 12 mm long, 2 - 8 mm in diameter, sparsely pubescent; bracts 3 to 6, overlapping toward pedicel base, 2 - 4 mm long, 2 - 3 mm wide; sepals 3, valvate, basally fused, 2 - 4 mm long, 3 mm wide, triangular to ovate, apex acute to obtuse, base truncate, pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 14.6 - 49 mm long, 2.4 - 3.8 mm wide at base, linear, apex acute to obtuse, base broad and concave, cream to white, pubescent towards base outside, pubescent with glabrous base inside; inner petals 3, valvate, 16 - 33 mm long, 2.1 - 3.2 mm wide at base, linear, apex acute, base broad and concave, cream to white, pubescent with glabrous base on both sides; stamens ca. 200, in 4 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 7 to 13, ovary ca. 1 mm long, stigmas connivent, linear-falcate, 1.8 - 3.4 mm long, pubescent. Monocarps stipitate, stipe 3 - 11 mm long, 2 - 3 mm in diameter; monocarps 5 to 10 (12), 21 - 65 mm long, 6 - 12 mm wide, narrowly oblong, weakly torulose, apex obtuse or with a beak 1 - 4 mm long, glabrous, verrucose or lenticellate (in vivo) and smooth to longitudinally wrinkled when dried, green outside, endocarp red to red-purple; seeds up to 9, commonly 4 - 8 per monocarp, in a single row, 9 - 12 mm long, 6 - 7 mm wide, ellipsoid; sarcotesta thin, waxy, green in vivo; aril absent.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1D5CABBDB6DB4929A23FB1CDDEF788CF.taxon	distribution	Distribution. A widespread species in central Africa, from Nigeria to South Sudan and south to northeastern Angola; in Cameroon known from East, South, Central, Littoral, South-West, West and Adamaoua regions.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
1D5CABBDB6DB4929A23FB1CDDEF788CF.taxon	materials_examined	Selected specimens examined. Adamaoua Region: At Sabal Haleo 62 km NE de Tibati, 6.78 ° N, 13.17 ° E, 11 April 1983, Asonganyi J. N. 632 (P, YA); A 10 km a l'ouest de Bagodo, 6.42 ° N, 13.38 ° E, 28 July 1966, Letouzey R. 7563 (YA); Banyo, 6.75 ° N, 11.82 ° E, 08 June 1967, Letouzey R. 8552 (P, YA). Central Region: Ndanan I to Ndangan I Forest on path left of road after bridge, site of 25 m × 25 m plot, 3.61 ° N, 11.59 ° E, 16 March 2004, Cheek M. 11690 (K, YA); Ca 50 km S of Badjob ca 60 km SW of Eseka Along the Nyong-River, 3.68 ° N, 10.68 ° E, 20 March 1964, de Wilde W. J. J. O 2172 (B, BR, K, MO, P, WAG, YA); Syzygeraie pres Malandi, 3.57 ° N, 11.22 ° E, 06 December 1959, Letouzey R. 2423 (P, YA); Pres des lacs de Boubala, 5.75 ° N, 11.9 ° E, 13 December 1960, Letouzey R. 2566 (P, YA); Rive boisee de la Sanaga pres Mbargue, 4.9 ° N, 13.08 ° E, 27 January 1960, Letouzey R. 2824 (P, YA). East Region: Betare Oya 5 km along road to Bertoua, 5.5 ° N, 14.1 ° E, 17 February 1960, Breteler F. J. 1063 (P, WAG, YA); Goyoum, 5.22 ° N, 13.38 ° E, 27 January 1961, Breteler F. J. 931 (BR, K, P, WAG); Mbussa, 6 ° N, 14.38 ° E, 29 April 1914, Mildbraed G. W. J. 9069 (K); Route Esseleke, 4.58 ° N, 13.68 ° E, 15 February 1956, Nana P. 479 (P); Sangha River (international frontier) collected 30 km from Libongo southwards, 2.5 ° N, 16.06 ° E, 27 July 1987, Thomas D. W. 7281 (YA). Littoral Region: 8 km W of Massok, 4.13 ° N, 10.47 ° E, 27 March 1965, Leeuwenberg A. J. M. 5217 (B, BR, C, GC, K, LUAI, MO, P, WAG, YA). North-West Region: Anyajua, 6.18 ° N, 10.36 ° E, 12 December 1998, Cheek M. 9923 (K, WAG, YA); Bamenda Distr Fonfuka, 6.53 ° N, 10.48 ° E, 01 May 1931, Maitland T. D. 1731 (K). South Region: Bitya near R Ja, 3.02 ° N, 12.37 ° E, 01 January 1921, Bates G. L. 1852 (P); S bank of Lobe R SE of Gr Batanga ferry, 2.88 ° N, 9.892 ° E, 11 October 1969, Bos J. J. 5475 (BR, C, K, LD, P, WAG, YA); Ca 10 km SW of Ambam S of Ebolowa On N bank of Ntem-River, 2.38 ° N, 11.28 ° E, 02 March 1964, de Wilde W. J. J. O 2043 (B, BR, K, MO, P, WAG, YA); Colline de Mill (5 km NE Lolodorf), 3.23 ° N, 10.73 ° E, 26 January 1974, Letouzey R. 12797 (P, YA); Ngoasik (10 km SSE Ambam) rive du Nt, 2.28 ° N, 11.33 ° E, 01 March 1963, Raynal J. 10130 (P, YA); Campo-Ma'an National Park, 2.33 ° N, 10.5 ° E, 24 August 2001, van Andel T. R. 4023 (KRIBI, WAG). South-West Region: Menyum (Bakossi), 4.6 ° N, 9 ° E, 28 February 1987, Doumenge C. 313 (MO, P, WAG); Mai Idoanu, 4.24 ° N, 8.99 ° E, 12 February 1955, Latilo M. G. 34468 (P); Onge River, 4.3 ° N, 8.95 ° E, 20 October 1993, Tchouto Mbatchou G. P. 896 (K, YA); Disturbed forest between Liwenyi and big Koto, 4.3 ° N, 8.95 ° E, 22 October 1993, Tchouto Mbatchou G. P. 942 (K, YA); Korup National Park, 5.05 ° N, 8.8 ° E, 28 February 1984, Thomas D. W. 3242 (MO, YA); Korup National Park, 4.98 ° N, 8.85 ° E, 12 January 1979, Thomas D. W. 595 (K). West Region: Baleng, 5.53 ° N, 10.38 ° E, 10 May 1964, de Wilde W. J. J. O 2565 (WAG).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4B05EA585A44114FEF4AB76CAB4E76D6.taxon	description	Fig. 153; Map 1 E	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4B05EA585A44114FEF4AB76CAB4E76D6.taxon	materials_examined	Type. Nigeria. Lagos State; Ibadan Forest Reserve, Lagos, Punch C. 119, Dec 1913: lectotype, designated by Johnson and Murray (2018), p. 215: K [000199069].	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4B05EA585A44114FEF4AB76CAB4E76D6.taxon	description	Description. Tree, up to 30 m tall, d. b. h. up to 90 cm; buttresses narrowly concave ca. 1 m high. Old branches sparsely pubescent to glabrous, young branches densely villous, with erect orange or reddish brown hairs 0.5 - 1.3 mm long. Leaves: petiole 2 - 4 mm long, ca. 2 mm wide, pubescent, grooved, blade inserted on top of the petiole; blade 8.6 - 12.6 cm long, 2.6 - 4.1 cm wide, lanceolate to lanceolate-elliptic or oblong-lanceolate, apex acuminate to acute, acumen 0.4 - 1.3 cm long, base cuneate to rounded, subcoriaceous, below golden-sericeous when young, sparsely golden-sericeous to golden-sericeous when old, above glabrous when young and old, discolorous; midrib impressed, above pubescent when young, glabrous to pubescent when old, below glabrous to pubescent when young, pubescent when old; secondary veins 10 to 15 pairs, glabrous above; tertiary venation reticulate. Individuals bisexual; inflorescences ramiflorous on young foliate branches, axillary, peduncle 8.5 - 17 mm. Flowers with 9 perianth parts in 3 whorls, 1 to 8 per inflorescence; pedicel 2 - 5 mm long, ca. 2 mm in diameter, densely pubescent; in fruit 6 - 42 mm long, 7 - 11 mm in diameter, glabrous; bracts 2 to 4, evenly spaced, 3 - 4 mm long, 3 - 4 mm wide; sepals 3, valvate, basally to ½ / 2 fused, forming a cup, 4 - 6 mm long, 3 - 4 mm wide, ovate, apex acute, base truncate, densely pubescent outside, glabrous inside; petals free, subequal; outer petals 3, 22 - 34 mm long, 3.5 - 3.9 mm wide at base, linear, apex acute to obtuse, base broad and concave, light yellow, pubescent, base glabrous outside, pubescent, base glabrous inside; inner petals 3, valvate, 17.4 - 23 mm long, 2.5 - 3.6 mm wide at base, linear, apex acute, base broad and concave, light yellow, pubescent with glabrous base outside, pubescent towards base to pubescent and glabrous towards center inside; stamens ca. 200, in 4 to 6 rows, 1 - 2 mm long, oblong; connective apex shield-like, glabrous; carpels 10 to 12, ovary 1 - 2 mm long, stigmas connivent with tips free, linear, 2.5 - 4 mm long, pubescent to sparsely pubescent. Monocarps stipitate, stipe ca. 3 mm long, ca. 9 mm in diameter; monocarps 1 to 10, ca. 46 mm long, ca. 23 mm wide, oblongoid, apex rounded, sparsely pubescent, verrucose and wrinkled when dried, green outside, endocarp color unknown; seeds unknown (sarcotesta blue?).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4B05EA585A44114FEF4AB76CAB4E76D6.taxon	distribution	Distribution. A widespread species in West Africa from Liberia to Ghana, and in Central Africa from southern Nigeria to Cameroon; in Cameroon known from East, South, Central and South-West regions. Given the past confusion with X. letestui, it is difficult to state the full distribution of X. villosa precisely.	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
4B05EA585A44114FEF4AB76CAB4E76D6.taxon	materials_examined	Specimens examined. Central Region: pres Nkomeyo 10 km d'Esse, 4.07 ° N, 11.97 ° E, 07 November 1969, Letouzey R. 9524 (P, YA). East Region: 65 km south of Yokadouma 30 km after Ngato 15 km after river ALPICAM ' base de vie' then 40 km on forestry road starting 4 km before Masea village, 3.08 ° N, 14.67 ° E, 08 March 2019, Couvreur T. L. P. 1227 (MPU, WAG, YA). South Region: Campo Ma'an National Park 11 km on trail from Ebinanemeyong village on road 7 km from Nyabessan to Campo town, 2.49 ° N, 10.34 ° E, 12 February 2015, Couvreur T. L. P. 686 (WAG, YA); 3 km E of km 58 of road Edea-Kribi, 3.72 ° N, 10.3 ° E, 05 October 1965, Leeuwenberg A. J. M. 6815 (BR, K, MO, P, WAG, YA). South-West Region: Korup National Park, 5.26 ° N, 9.2 ° E, 08 April 1988, Thomas D. W. 7703 (MO).	en	Couvreur, Thomas L. P., Dagallier, Leo-Paul M. J., Crozier, Francoise, Ghogue, Jean-Paul, Hoekstra, Paul H., Kamdem, Narcisse G., Johnson, David M., Murray, Nancy A., Sonke, Bonaventure (2022): Flora of Cameroon - Annonaceae Vol 45. PhytoKeys 207: 1-532, DOI: http://dx.doi.org/10.3897/phytokeys.207.61432, URL: http://dx.doi.org/10.3897/phytokeys.207.61432
