taxonID	type	description	language	source
03D187D04947FFBFFF5CFD80FD3CF926.taxon	type_taxon	Type species: Neoathyreacarus pygmephoroides sp. nov.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBFFF5CFD80FD3CF926.taxon	diagnosis	Diagnosis (adult female only). With all characters of the family (Khaustov & Frolov 2019). Gnathosoma. Gnathosoma dorsally with two pairs of cheliceral setae (cha, chb); palps ventrally with small accessory setigenous structure medially and baculiform solenidion laterally; palp tibiotarsus without lateral seta, distally with short rodlike eupathid and well developed tibial claw, subcapitulum with only one pair of setae (m), setae n absent. Pharynx weakly sclerotized, not clearly subdivided laterally into pharyngeal pumps. Cheliceral stylets very short and thin. Idiosomal dorsum. Prodorsum with three setae (v 1, v 2, sc 2) and alveolar pits sc 1. Bothridia absent. Stigmata located near anterolateral corners of prodorsal shield. Tergite C entire, not subdivided into median and lateral parts. Idiosomal venter. Genital setae absent. Pseudanal segment with three pairs of setae (ps 1 - 3) located ventrally; anal opening small, located ventrally. Legs. Leg I with large tarsal claw ridged on inner surface. Unguinal setae (u’, u ”) on tarsus I separated, modified, spine-like, with weak ridges on inner surface. Tarsi II and III with large claws distinctly thickened near their bases. Claws on tarsus IV not thickened basally. Empodium on tarsi II-IV large, weakly plicate on ventral surface. Leg setation. Leg I: Tr 1 (v’), Fe 5 (d, l’, l ”, v’, v ”), Ge 4 (l’, l ”, v’, v ”), Ti 6 (2) (k, d, l’, l ”, v’, v ”, φ 1, φ 2), Ta 13 (2) (p’, p ”, tc’. tc ”, ft’, ft ”, pl’, pl ”, u’, u ”, s, pv’, pv ”, ω 1, ω 2); leg II: Tr 1 (v’), Fe 3 (d, l’, v ”), Ge 3 (l’, l ”, v’), Ti 4 (1) (d, l’, v’, v ”, φ), Ta 7 (1) (pl ”, tc’, tc ”, u’, u ”, pv’, pv ”, ω); leg III: Tr 1 (v’), Fe 2 (d, v’), Ge 3 (l’, l ”, v’), Ti 4 (1) (d, l’, v’, v ”, φ), Ta 7 (pl ”, tc’, tc ”, u’, u ”, pv’, pv ”); leg IV: Tr 1 (v’), Fe 2 (d, v’), Ge 2 (l ”, v’), Ti 4 (1) (d, l’, v’, v ”, φ), Ta 6 (pl ”, tc’, tc ”, u’, pv’, pv ”). Solenidion φ on tibiae II-IV located in anterior part of segment. Species included. The genus Neoathyreacarus includes one species: N. pygmephoroides sp. nov.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBFFF5CFD80FD3CF926.taxon	distribution	Distribution and hosts. N. pygmephoroides sp. nov. is recorded from Chile on the bolboceratine beetle Bolborhinum tubericeps Fairmaire (Coleoptera: Geotrupidae).	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBFFF5CFD80FD3CF926.taxon	diagnosis	Differential diagnosis. The new genus is most similar to Athyreacarus in having three pairs of pseudanal setae, absence of bothridia and setae sc 1 represented by alveolar pits. The new genus differs from Athyreacarus in having only one pair of subcapitular setae (two in Athyreacarus), entire tergite C (subdivided into median and lateral parts in Athyreacarus), absence of setae d on genu I and p’ on tarsus IV (present in Athyreacarus), pharynx not clearly subdivided into four pharyngeal pumps (pharynx clearly subdivided into four pharyngeal pumps in Athyreacarus), and very small cheliceral stylets (large in Athyreacarus).	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBFFF5CFD80FD3CF926.taxon	etymology	Etymology. The name of the new genus is a combination of two words: Greek neos meaning new, and Athyreacarus, the name of closely related genus.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBDFF5CF8E1FF52F82A.taxon	description	(Figs 1 – 6) Description. Female. Length of idiosoma 300 (290 – 320), width 170 (160 – 180). Gnathosoma (Fig. 2). Gnathosomal capsule, excluding palps, broadly oval, length 32 (32 – 34), width 37 (37 – 46). Dorsal gnathosomal apodeme poorly visible. Two pair of barbed, blunt-tipped cheliceral setae cha 33 (28 – 33) and chb 10 (10 – 11). Postpalpal setae (pp) 8 (8 – 9) blunt-tipped, smooth. Setae m 19 (18 – 20) pointed and smooth. Setae dFe 16 (15 – 16) and dGe 22 (20 – 22) weakly barbed and pointed. Tibial claw slightly hooked, without projection. Cheliceral stylet length 8 (7 – 9). Idiosomal dorsum (Figs 1 A, 5 A, 6 A – C). All dorsal shields with numerous sparsely distributed puncta (Figs 6 A – C). Prodorsal shield without lateral projections. All dorsal setae weakly barbed; setae sc 2, c 2, d, and f pointed, other dorsal setae blunt-tipped;. Cupules ia on tergite D, im and ip on tergite EF and ih on tergite H small, round. Tergites C, D, and EF with few pore-like structures. Stigmata round. Pseudanal segment completely covered dorsally by tergite H. Lengths of dorsal setae: v 1 50 (46 – 52), v 2 20 (19 – 21), sc 2 90 (79 – 90), c 1 53 (46 – 54), c 2 105 (98 – 105), d 110 (105 – 110), e 34 (31 – 40), f 120 (115 – 120), h 1 88 (79 – 88), h 2 73 (70 – 77). Distances between setae: v 1 – v 1 23 (18 – 24), v 2 – v 2 58 (52 – 58), sc 2 – sc 2 63 (55 – 63), c 1 – c 1 70 (66 – 72), c 1 – c 2 31 (27 – 32), d – d 83 (76 – 89), e – e 76 (73 – 80), f – f 83 (82 – 84), h 1 – h 1 29 (28 – 36), h 1 – h 2 25 (22 – 26). Idiosomal venter (Figs 1 B, 5 B, 6 D – F). All ventral plates with numerous puncta (Figs 6 D – F). Setae 3 b, 3 c, 4 a, 4 b, and 4 c smooth; other ventral setae weakly barbed. Setae 1 b, 2 c, 3 a, ag 1 and ps 1 - 3 blunt-tipped, other ventral setae pointed; in most specimens at least one of setae ps 1 and / or ps 3 bifurcate, in other specimens not bifurcate; setae ps 1 - 3 situated in one transverse row. Ap 1 well developed and fused with appr; ap 2 well developed and fused with appr; apsej absent; ap 3 weakly developed and presented by small oval sclerites; ap 4 and ap 5 well developed. Aggenital plate with two pairs of aggenital setae (Fig. 6 F). Length of mid-sternal plate 64 (63 – 67), width 61 (61 – 65); ratio length / width 1.0. Lengths of ventral setae: 1 a 25 (25 – 28), 1 b 15 (15 – 16), 1 c 24 (20 – 24), 2 a 28 (27 – 29), 2 b 28 (25 – 28), 2 c 15 (12 – 20), 3 a 14 (14 – 23), 3 b 28 (27 – 29), 3 c 26 (25 – 27), 4 a 26 (26 – 27), 4 b 34 (28 – 34), 4 c 27 (25 – 28), ag 1 15 (14 – 15), ag 2 33 (29 – 33), ps 1 48 / 53 (47 – 50), ps 2 29 (24 – 41), ps 3 65 / 76 (62 – 72). Legs (Figs 3, 4). Leg I (Fig. 3 A): setae l’, l ”, v’ of femur, v’ of genu, k of tibia, and all of tarsus smooth, other leg setae barbed; seta l’, l ”, v’ of femur, v’ of genu, l ”, v’ and k of tibia and (p), (tc), (ft) of tarsus blunt-tipped, other leg setae pointed; seta l’ of femur spiniform, situated very close to d. Lengths of solenidia ω 1 15 (14 – 15), ω 2 7 (7), φ 1 10 (10 – 11), φ 2 6 (6 – 7); all solenidia digitiform. Leg II (Fig. 3 B): solenidia ω 10 (10 – 14) and φ 3 (3) digitiform; all leg setae weakly barbed; setae d, l’ of femur and (pv) of tarsus blunt-tipped, other leg setae pointed. Leg III (Fig. 4 A) similar in shape and length to leg II; solenidion φ 3 (3 – 4) digitiform; all leg setae weakly barbed; setae v’ of trochanter, d of femur and (pv) of tarsus blunt-tipped, other leg setae pointed. Leg IV (Fig. 4 B): solenidion φ 4 (4) digitiform; all leg setae weakly barbed; setae v’ of femur and genu blunt-tipped, other leg setae pointed.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBDFF5CF8E1FF52F82A.taxon	materials_examined	Type material. Female holotype, slide No. ZISP T-Ath- 011, Chile, Termas de Chillán, 15. I. 2010, on Bolborhinum tubericeps. Paratypes: 24 females, same data. Type deposition. The holotype and five paratypes are deposited in the collection of the Zoological Institute of RAS, Saint Petersburg, Russia; other paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04947FFBDFF5CF8E1FF52F82A.taxon	etymology	Etymology. The name of the new species pygmephoroides refers to its morphological (not phylogenetic) similarity to some species of the superfamily Pygmephoroidea sharing the following characters: subcapitulum with only one pair of setae, tergite C not divided into three parts, and similar position of anal opening and pseudanal setae.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFBBFF5CFA92FAA2FA70.taxon	materials_examined	Type species: Athyreacarus pleiotretus Lindquist, Kaliszewski and Rack, 1990, by original designation.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFB2FF5CF9CEFEFDFACA.taxon	description	(Figs 7 – 13) Description. Female. Body broadly oval. Length of idiosoma 635 (565 – 750), width 360 (330 – 445). Gnathosoma (Fig. 8). Gnathosomal capsule, excluding palps, almost round, length 74 (67 – 79), width 75 (66 – 80). Dorsal gnathosomal apodeme not evident. Two pairs of barbed, blunt-tipped cheliceral setae cha 44 (38 – 56) and chb 33 (27 – 41). Postpalpal setae (pp) 17 (15 – 18) blunt-tipped, with some weak barbs in distal half. Setae m 43 (33 – 44) and n 85 (62 – 88) pointed; setae m smooth, n weakly barbed. Setae dFe 37 (27 – 41) and dGe 39 (30 – 40) barbed; setae dFe blunt-tipped, dGe pointed. Tibial claw slightly hooked, with distinct subapical projection. Pharynx typical for the genus. Ass short, with small distal projection. Palpal tibiotarsus with short spiniform seta l ” anterolaterad palpal solenidion, rather long eupathid-like seta distally and very short setiform structure anteriad ass. Cheliceral stylets 11 (10 – 13) relatively short and thin. Idiosomal dorsum (Figs 7 A, 11, 13 A, D). All dorsal shields with numerous sparsely distributed puncta (Figs 13 A, D). Prodorsal shield without lateral projections. Setae sc 1 completely absent, or in some specimens represented by poorly visible alveolar pits (Fig. 13 D). All dorsal setae weakly blunt-tipped and weakly barbed. Cupules ia on tergite D, im, ip on tergite EF and ih on tergite H very small, round. Tergites C, D, and EF with pore-like structures typical for the genus. Stigmata round. Pseudanal segment almost completely covered by tergite H. Lengths of dorsal setae: v 1 94 (70 – 125), v 2 73 (73 – 75), sc 2 160 (135 – 175), c 1 140 (130 – 185), c 2 150 (145 – 195), d 145 (130 – 180), e 120 (105 – 150), f 170 (140 – 180), h 1 170 (145 – 195), h 2 175 (150 – 200). Distances between setae: v 1 – v 1 58 (55 – 78), v 2 – v 2 85 (80 – 105), sc 2 – sc 2 120 (115 – 145), c 1 – c 1 145 (135 – 180), c 1 – c 2 77 (65 – 87), d – d 180 (145 – 190), e – e 185 (145 – 205), f – f 200 (175 – 245), h 1 – h 1 145 (120 – 150), h 1 – h 2 25 (24 – 35). Idiosomal venter (Figs 7 B, 112, 13 B, C). All ventral plates with numerous puncta (Figs 13 B, C). Setae ps 1 and ps 2 weakly barbed; other ventral setae smooth. Setae 1 c pointed; other ventral setae weakly blunt-tipped; setae 1 a, 1 b, 2 a, 2 b, 2 c, 3 a, 3 b, 3 c, 4 a, 4 b, 4 c, and ag 1 swollen basally and attenuate distally (Figs 13 B, C). Ap 5 long, ap 3 and ap 4 fused; apsej not evident. Aggenital plate with two pairs of aggenital setae. Anal opening ventral. Pseudanal setae located ventrally Length of mid-sternal plate 128 (96 – 128), width 145 (120 – 145); ratio length / width 0.8. Lengths of ventral setae: 1 a 39 (34 – 43), 1 b 34 (29 – 37), 1 c 96 (73 – 99), 2 a 45 (43 – 49), 2 b 39 (38 – 50), 2 c 40 (36 – 53), 3 a 29 (27 – 35), 3 b 43 (42 – 51), 3 c 40 (34 – 46), 4 a 44 (38 – 55), 4 b 45 (39 – 61), 4 c 43 (40 – 52), ag 1 41 (40 – 60), ag 2 79 (79 – 96), ps 1 110 (79 – 140), ps 2 125 (105 – 145), ps 3 43 (31 – 59). Legs (Figs 9, 10). Leg I (Fig. 9 A): setae v’ of trochanter and d of femur and genu barbed, other setae smooth; setae d of femur, v’ of genu, d, l’, v ” of tibia pointed, other leg setae weakly blunt-tipped; lengths of solenidia ω 1 15 (13 – 16), ω 2 14 (11 – 14), φ 1 11 (10 – 11), φ 2 12 (10 – 12); all solenidia digitiform. Tarsal claw very thick and blunttipped. Leg II (Fig. 9 B): solenidia ω 14 (12 – 14) and φ 13 (11 – 14) digitiform; seta d of femur barbed, other setae smooth; seta u ” very short, spiniform; seta v’ of trochanter with swollen basal half and attenuate distally; setae (tc) with widened and flattened tips; setae (pv) weakly blunt-tipped, other setae pointed. Tarsal claws distinctly smaller than in other Athyreacarus species. Empodium with distinct dorsal longitudinal striation. Leg III (Fig. 10 A) similar in shape and length to leg II; solenidion φ 12 (11 – 14) digitiform; setae d of femur, l ” of genu and d of tibia barbed, other setae smooth; seta u ” very short, spiniform; seta v’ of trochanter with swollen basal half and attenuate distally; setae (tc) with widened and flattened tips; setae v’ of trochanter, d of femur and (pv) weakly blunt-tipped, other setae pointed. Claws and empodium as on tarsus II. Leg IV (Fig. 10 B): solenidion φ 14 (14 – 16) digitiform; setae d of femur, l ” of genu, d of tibia and p’ barbed, other setae smooth; seta v’ of trochanter with swollen basal half and attenuate distally; seta v’ of femur thickened in basal half; seta tc’ with widened and flattened tip; setae v’ of trochanter, d of femur, v’ of genu, and p’ weakly blunt-tipped, other setae pointed. Claws and empodium as on tarsi II and III.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFB2FF5CF9CEFEFDFACA.taxon	materials_examined	Type material. Female holotype, slide No. ZISP T-Ath- 012, Argentina, Catamarca Prov., Londres, 7 - 17. II. 2015, G. L. Agnoli, on Zefevazia cantisani Martinez. Paratypes: 7 females, same data; 17 females, Argentina, Catamarca Prov., Punta de Balasto, II. 2015, G. L. Agnoli, on Zefevazia cantisani; 1 female, Argentina, Catamarca Prov., Belen, II. 2013, G. L. Agnoli, on Zefevazia cantisani. Type deposition. The holotype and five paratypes are deposited in the collection of the Zoological Institute of RAS, Saint Petersburg, Russia; other paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFB2FF5CF9CEFEFDFACA.taxon	diagnosis	Differential diagnosis. The new species considerably differs from other Athyreacarus species in having very short spiniform setae u ” on tarsi II and III (always seta-like in other Athyreacarus), setae 1 a, 1 b, 2 a, 2 b, 2 c, 3 a, 3 b, 3 c, 4 a, 4 b, 4 c, and ag 1 swollen basally and attenuate distally (not modified in other Athyreacarus), palp tibiotarsus ventrally with short spiniform seta l ” (absent in other Athyreacarus).	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFB2FF5CF9CEFEFDFACA.taxon	etymology	Etymology. The name of the new species is a Latin word staturosus meaning gigantic and refers to extremely large body size.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D04943FFB2FF5CF9CEFEFDFACA.taxon	discussion	Remarks. The considerable differences in the new species compared to other Athyreacarus may represent another genus. However, the most sufficient difference (presence of seta l ” on palpal tibiotarsus) is a plesiomorphic character state and not useful for creating a new taxon. However, apomorphic characters such as very short spiniform setae u ” on tarsi II and III, unusually small cheliceral stylets, and modifications of most ventral idiosomal setae could be helpful. The discovery of other species with similar character states may result in creation of another supraspecific taxon.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D0494AFFAFFF5CFA95FD20FD42.taxon	description	(Figs 14 – 20) Description. Female. Body broadly fusiform. Length of idiosoma 405 (350 – 405), width 230 (190 – 255). Gnathosoma (Fig. 15). Gnathosomal capsule, excluding palps, broadly oval, length 64 (61 – 64), width 66 (64 – 69). One pair of barbed, blunt-tipped cheliceral setae cha 29 (27 – 29). Postpalpal setae (pp) 12 (11 – 12) blunt-tipped, with tiny subapical barbs. Setae m 10 (9 – 10) smooth and blunt-tipped, n 53 (51 – 54) pointed and barbed. Setae dFe 14 (14 – 16) and dGe 36 (32 – 36) weakly barbed; setae dFe blunt-tipped, dGe pointed. Tibial claw very small, with tiny basal projection. Pharynx typical for the genus. Ass almost three times shorter than palpal solenidion. Tiny eupathid-like seta on palpal tibiotarsus located dorsally. Cheliceral stylet length 20 (18 – 22). Idiosomal dorsum (Figs 14 A, 18, 20 A). All dorsal shields with tiny puncta (Fig. 20 A). Prodorsal shield without lateral projections. Setae sc 1 present, strongly reduced to alveolar remnants (Fig. 20 A). All dorsal setae weakly barbed; setae v 2 and e blunt-tipped, other dorsal setae pointed. Cupules ia on tergite D, im, ip on tergite EF and ih on tergite H small, round. Tergites C, D, and EF with pore-like structures typical for the genus. Stigmata round. Pseudanal segment shorter than tergite H. Lengths of dorsal setae: v 1 73 (72 – 74), v 2 34 (30 – 35), sc 2 140 (140 – 145), c 1 125 (120 – 125), c 2 150 (145 – 150), d 140 (130 – 145), e 37 (33 – 39), f 145 (120 – 145), h 1 115 (95 – 115), h 2 110 (100 – 110). Distances between setae: v 1 – v 1 47 (39 – 48), v 2 – v 2 75 (68 – 80), sc 1 – sc 1 78 (68 – 84), sc 2 – sc 2 89 (79 – 89), c 1 – c 1 87 (66 – 88), c 1 – c 2 49 (46 – 50), d – d 95 (77 – 96), e – e 88 (76 – 89), f – f 91 (76 – 94), h 1 – h 1 52 (47 – 52), h 1 – h 2 20 (15 – 21). Idiosomal venter (Figs 14 B, 19, 20 B – D). All ventral plates with tiny puncta (Figs 20 B – D). All ventral setae weakly barbed. Setae 1 b and 2 c blunt-tipped, other ventral setae pointed. Ap 5 long, extending beyond bases of setae 4 a. Aggenital plate with three pairs of aggenital setae (Fig. 20 D). Female holotype with additional pair of setae on coxisternal fields III (3 x), other specimens with normal three pairs of setae (Fig. 20 C). Anal opening terminal. Length of mid-sternal plate 68 (66 – 68), width 42 (42 – 46); ratio length / width 1.4 – 1.6. Lengths of ventral setae: 1 a 39 (39 – 45), 1 b 27 (23 – 30), 1 c 27 (27 – 30), 2 a 53 (51 – 55), 2 b 41 (40 – 42), 2 c 33 (25 – 34), 3 a 35 (26 – 35), 3 b 48 (41 – 48), 3 c 47 (43 – 48), 4 a 47 (46 – 48), 4 b 56 (55 – 57), 4 c 41 (35 – 42), ag 1 50 (42 – 50), ag 2 62 (47 – 63), ag 3 58 (48 – 58), ps 1 64 (59 – 65), ps 2 69 (61 – 70), ps 3 42 (41 – 48). Legs (Figs 16, 17). Leg I (Fig. 16 A): setae l’, v’ of femur, v’ of genu, k of tibia, and s, pv’, pv ”, pl’, (p), (tc), (ft) of tarsus smooth, other leg setae barbed; setae l’, v’ of femur, v’ of genu, v’ and k of tibia and s, pv ”, (p), (tc), (ft) of tarsus blunt-tipped, other leg setae pointed; lengths of solenidia ω 1 12 (11 – 12), ω 2 10 (9 – 10), φ 1 11 (11 – 12), φ 2 9 (8 – 9); all solenidia digitiform. Leg II (Fig. 16 B): solenidia ω 12 (11 – 12) and φ 10 (9 – 10) digitiform; setae (u) smooth, other setae weakly barbed; setae l’ of femur blunt-tipped, other leg setae pointed. Leg III (Fig. 17 A) similar in shape and length to leg II; solenidion φ 10 (9 – 10) digitiform; setae (u) smooth, other setae weakly barbed; all setae pointed. Leg IV (Fig. 17 B): solenidion φ 10 (9 – 10) digitiform; all leg setae weakly barbed; seta p’ blunttipped, other leg setae pointed.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D0494AFFAFFF5CFA95FD20FD42.taxon	materials_examined	Type material. Female holotype, slide No. ZISP T-Ath- 013, Republic of South Africa, Northern Cape Prov., Brandvlei, 12 - 14. VI. 1997, Namibiobolbus helgae Gussmann and Scholtz. Paratypes: 8 females, same data. Type deposition. The holotype and two paratypes are deposited in the collection of the Zoological Institute of RAS, Saint Petersburg, Russia; other paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D0494AFFAFFF5CFA95FD20FD42.taxon	diagnosis	Differential diagnosis. The new species is most similar to A. pseudoindicus Khaustov and Frolov, 2021 in having similar setal lengths on idiosoma, presence of one pair of cheliceral setae, presence of cupuli ih and presence of setae 1 c. The new species clearly differs from A. pseudoindicus only by the presence of tiny setae sci (completely absent in A. pseudoindicus). Also females of the new species are phoretic on beetles of the genus Namibiobolbus, while A. pseudoindicus phoretic on beetles of the genus Mimobolbus.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
03D187D0494AFFAFFF5CFA95FD20FD42.taxon	etymology	Etymology. The new species named after Anne M. Camerik who passed away in 2021, for her contribution to study of heterostigmatic mites of South Africa.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): New taxa of Athyreacaridae (Acari: Heterostigmata) from Neotropical and Afrotropical realms. Zootaxa 5188 (6): 501-520, DOI: https://doi.org/10.11646/zootaxa.5188.6.1
