identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3A393D28FFE5FFD8FF6DD8C69E43F82D.text	3A393D28FFE5FFD8FF6DD8C69E43F82D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasypoda schwarzi Radchenko et Michez 2022	<div><p>Dasypoda schwarzi Radchenko et Michez sp. nov.</p> <p>Type material. Holotype ♂ with the printed label: " Tunesien, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.683333&amp;materialsCitation.latitude=35.383335" title="Search Plazi for locations around (long 8.683333/lat 35.383335)">Thala</a> 20 km S [35°23'N 8°41'E], 12.4.1981, leg. Max. Schwarz " (MSAA).</p> <p>Paratypes: 2 ♂ with the printed label: " Morocco, 40 km S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-3.3666668&amp;materialsCitation.latitude=33.883335" title="Search Plazi for locations around (long -3.3666668/lat 33.883335)">Guercif</a> [33°53'N 3°22'W], 15.- 17.5.1995, Ma. Halada lgt." (UMONS); Tunesien, Thala 20 km S, 12.4.1981, leg. J. Gusenleitner (OÖLM).</p> <p>Etymology. The species is named after Maximilian Schwarz, an authority on bee systematics.</p> <p>Subgeneric affinity. Dasypoda schwarzi sp. nov. can be regarded as a representative of the subgenus Microdasypoda Michez (in Michez et al. 2004b) due to the combination of its diagnostic features.</p> <p>First, representatives of the subgenus Microdasypoda are characterized by a relatively small body size of approximately 7-12 mm (contrast Megadasypoda and many species of Dasypoda s.str. and Heterodasypoda which are significantly larger). Then, the subgenus is most significantly distinguished from all other Dasypoda subgenera by the structure of the male genitalia and S7: the gonostylus of Microdasypoda is either unilobed with one basal tooth, or bilobed without a membranous structure that connects these lobes (contrast Dasypoda s.str. in which the gonostylus is bilobed with such membranous structure, and Heterodasypoda and Megadasypoda in which it is trilobed), and the S7 does not bear latero-apical processes, which are characteristic of all other subgenera of Dasypoda.</p> <p>The previously cited apomorphies of Microdasypoda should however always be regarded with a combination of other characters that are individually shared by other subgenera, but not all. First, the malar space of Microdasypoda is always much shorter than the pedicel (contrast most Megadasypoda) and their maxillary palpi and galea are of a sub-equal length (contrast Dasypoda s.str. and Megadasypoda). The nervulus (cu-v) of Microdasypoda is strongly antefurcal (contrast most Megadasypoda). The apex of S6 of Microdasypoda has a long, dense pubescence (contrast Dasypoda s.str. and Megadasypoda which have short, sparse ones). Microdasypoda lack lateral hooks at the basal half of S8 (contrast Dasypoda s.str. and some Megadasypoda). The external lobe of the gonostyle of Microdasypoda is not lanceolate (contrast Megadasypoda and some Heterodasypoda), and the inner lobe of the gonostyle of Microdasypoda does not present a scaly surface (contrast Heterodasypoda in which this scaly surface is clearly visible).</p> <p>The new species described in this work, Dasypoda schwarzi sp. nov., clearly corresponds to all the features that characterize the subgenus Microdasypoda.</p> <p>Description. Male (Figs 1, 2): Holotype. Body black, its length (vertex to T7) 11.9 mm (paratypes: 11.5 mm and 10.5 mm).</p> <p>Head slightly wider than long (Fig. 6): L = 2.9 mm (paratypes: 2.8 mm and 2.7 mm), W = 3.4 mm (paratypes: 3.2 and 3.1 mm, respectively); clypeus densely punctured by oblique downward hair-bearing punctures separated by 0.5–1 puncture diameter with abundant long, dense, white adjacent hair obscuring underlying surface, apex of clypeus with narrow depressed impunctate band. Paraocular area irregularly punctate, punctures separated by 1–3 puncture diameter. Paraocular and supraclypeal areas and basal part of frons with long erect pale-white plumose hairs, intermixed on upper part near vertex with dark brown hair. Genal area, apical parts of vertex, and occiput with long, erect pale-white pubescence intermixed with dark brown hair. Central part of frons with narrowly depressed, median stripe, upper part of frons medially polished, unpunctured; laterally irregularly, sparsely punctate and very weakly shagreened. Malar space narrow, 4 times as broad as long, (W = 0.43 mm, L = 0.11 mm) (Fig. 4). Antennal scape anteriorly with long erect white hair. Flagellum moderately long, first flagellomere 1.7 times as long as its apex width and 1.25 times longer than second one (Fig. 3, 44). Labrum polished and shining, apical edge with long dense fringe of golden-yellow or pale-yellow thick hair. Glossa short: L = 0.65 mm, widened at base: W (at base) = 0.23 mm (Fig. 5). 2 nd and 3 rd segments of labial palpi apically swollen. Galea weakly shagreened with very sparse and superficial punctation (Fig. 9); maxillary palpi subequal in length with galea.</p> <p>Mesosoma: W (between tegulae) = 2.9 mm (paratypes: 2.9 mm and 2.7 mm). Scutum with hair-bearing punctation, punctures separated by 1–1.5 puncture diameters with exception of sparsely punctate (3–4 puncture diameters) central part (Fig. 7); mesosoma ventrally covered with very long, erect white hair, laterally and dorsally with pale-yellow hair, on scutum intermixed with shorter dark brown hairs; scutellum and metanotum with rufous hair. Propodeal triangle finely shagreened, basally with narrow transverse rugosity; external margins well-defined by lateral lines (Fig. 8). Propodeum laterally very weakly shagreened and sparsely punctured with long erect yellow hair. Wings hyaline with light brown tint; tegulae, venation and stigmata dark brown. Nervulus (cu-v) antefurcal. Legs with long pale-yellow pubescence, apart from greyish pubescence of inner side of tibiae and dense short ginger-yellow pubescence of inner side of metabasitarsus.</p> <p>Metasoma (Figs 10, 11): L = 5.8 mm (paratypes: 5.7 and 5.3 mm); W = 4.3 mm (paratypes: 4.3 mm and 3.8 mm, respectively). T1 basal part with oblique, moderately sparse punctation with long, semierect hair; separated by 1–3 puncture diameters; marginal part of T1 depressed, almost unpunctured and very thinly and densely transversely wrinkled with sparse superficial weakly noticeable punctation bearing short brown hair (Fig. 12); narrow apical margin of all terga light, translucent. Basal parts of T2–T5 moderately dense punctured, punctures separated by 1–2 puncture diameter with long semi-adjacent yellow hair, underlying surface with sparse short brown hair. Apical margins of T2–T5 slightly depressed with entire bands of very short dense white hairs covered from above by apical parts of long yellow hair. Basal part of T6 densely punctate by hair-bearing puncture with semi-adjacent long yellow hair (in holotype specimen this pubescence erased), laterally with long erected pale-yellow hair intermixed with sparse brown hair; apical margin of this tergum polished, impunctate. Basal parts of sterna moderately sparse punctured (denser on S2–S4 and sparser on S5) by very small oblique hair-bearing punctures with very short dark brown hair; underlying surface weakly shagreen (Fig. 13). Sternal margins with sparse, yellow hair fringes longer laterally and shorter centrally, narrowly interrupted at the centre of S1–S4, and widely on S5. Marginal parts of S1–S5 very sparse punctured, polished, its narrow apical margin light, translucent, on S2–S5 medially widely roundly notched. S6 marginal part medially with wide projection deeply triangularly emarginated on apex (Figs 14, 15). S7 (Fig. 17) latero-apically with short, weakly sclerotized semicircular projections bent towards inner side of sternum (Fig. 18). S8 relatively short, stumpy, deeply semicircularly concave laterally before widened basal part (Figs 19, 20); apex of S8 posterodorsal part with whole transverse carina not notched or emarginated in centre (Fig. 16). Genitalia with bilobed gonostyli (Figs 21–34). Inner lobe of gonostylus moderately wide and long, widened in a circular shape at apex and covered with sparse long setae on surface facing penis valves and on margins (Figs 33, 34, 85, 86); external lobe wide, its apex obliquely truncated (Fig. 32), ventral part narrowly, triangularly elongated, inner surface with several hairs, each inserted into crater-like raised alveoli (Figs 35–37).</p> <p>Female unknown.</p> <p>Distribution. Northwest Africa: Atlas Mountains area (Morocco and Tunisia) (Fig. 89).</p> <p>Floral visitation. Pollen grains of the Malvaceae plant family were mainly found on the body of the paratype (Fig. 38), and only a few grains of Asteraceae.</p> <p>Species diagnosis. The main significant differences D. schwarzi sp. nov. from the other known species of Microdasypoda concern the structure of genitalia. Other Microdasypoda species have single-lobed gonostyli with a spine-like tooth on the inner basal part (in D. cingulata, D. crassicornis and D. iberica; Figs 82–84) or bilobed gonostyli with a thickened internal lobe that has a very dense, long pubescence on surface facing penis valves (in D. brevicornis; Figs 75, 80, 81). Unlike other representatives of this subgenus, the genitalia of D. schwarzi have bilobed gonostyli with the inner lobes that are laterally flattened and dorso-ventrally widened in a circular shape at the apical part, and with sparse setae on the surface facing penis valves (Figs 85, 86).</p> <p>By the structure of its genitals and sterna, D. schwarzi is most similar to D. brevicornis, sharing the same structure of S6, S7, dorso-apical part of S8, glossa and galea. In both species, unlike other Microdasypoda, the setae along the lower edge of the galea are longer in the apical half than in the basal half where they are very short and poorly visible (Figs 9, 39) (Michez et al. 2004b indicated the absence of setae in the basal half of galea, probably due to the relatively lower resolution of optics used at that time).</p> <p>Dasypoda schwarzi can also be separated from D. brevicornis by the following features: the external lobe of the gonostyle in D. schwarzi with a strongly oblique apex (Figs 21, 23, 29, 32); the digitus of its volsella shorter and wider (Fig. 28); the 1 st flagellomere relatively longer (Figs 43, 44); the mandibular palpus with thicker segments (Figs 5, 6, 9); the apical projection of the S6 with right angles on the outer edges (rather than rounded lateral corners as in D. brevicornis) (Figs 55, 59); the lateral margins of the S8 deeply semicircularly notched before the apodemes (straight-beveled lateral margins of S 8 in D. brevicornis) (Figs 65, 69); and the lower half of the face with light pubescence without dark brown or black hair (the latter being typical in D. brevicornis).</p> <p>Remarks on the comparative morphology of Microdasypoda. The use of modern optics with a higher resolution made it possible to refine the original diagnosis of the subgenus Microdasypoda Michez (Michez et al. 2004b). In particular, the outer lobe of the gonostyle may have a lanceolate shape (in D. crassicornis), the S 7 in almost all species of the subgenus (with the exception of D. iberica) has a very small almost non-chitinous lateroapical outgrowths (Fig. 18); the dorso-apical part of S8 has an entire transverse carina or two partly or completely separated teeth (Figs 70–74); the S6 apicoventrally may have not only brown pubescence (in most species), but can also present a complete silver-white pubescence (in D. iberica; Fig. 53; Radchenko et al. 2019).</p> <p>This updated clarification of the diagnostic characters of all species belonging to the subgenus Microdasypoda, as well as the detailed morphological characters of the male of D. iberica that was recently published (Radchenko et al. 2019) made it possible to compose a corrected and updated key for the males.</p> </div>	https://treatment.plazi.org/id/3A393D28FFE5FFD8FF6DD8C69E43F82D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Radchenko, Vladimir G.;Ghisbain, Guillaume;Michez, Denis	Radchenko, Vladimir G., Ghisbain, Guillaume, Michez, Denis (2022): A new bee species of the genus Dasypoda Latreille (Hymenoptera, Apoidea) from Northwest Africa with comparative remarks on the subgenus Microdasypoda Michez. Zootaxa 5188 (1): 74-86, DOI: https://doi.org/10.11646/zootaxa.5188.1.4
3A393D28FFEDFFD4FF6DDF8E9FE1FC57.text	3A393D28FFEDFFD4FF6DDF8E9FE1FC57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasypoda (Microdasypoda) Michez 2004	<div><p>Key to the males of Dasypoda (Microdasypoda) Michez, 2004</p> <p>1. Galea black, smooth, polished, shiny, with very sparse punctures (Fig. 40). Glossa narrow and relatively long, 5 times as long as wide (Fig. 46). S6 with brown apical pilosity directed laterally (Figs 51, 56). Marginal edge of S6 strongly bent outside ventrally, especially at the lateral angles (Fig. 56). S7 marginal part deeply notched centrally, its lateral apodemes relatively short (Fig. 61). Dorso-apical part of S8 with two wide, partly separated outgrowths (Fig. 71); lateral apodemes before pregladular area of this sternum rectangularly widened on the sides (Fig. 66). Gonostylus strongly widened and not rounded apically; inner basal part of gonostylus with very short tooth; penis valvae narrower than gonostylus (Figs 76, 82). Mid and hind legs with the last article often yellow. Distribution: species mostly distributed in the western part of the Mediterranean basin (Fig. 88)............................................................................. Dasypoda cingulata Erichson, 1835</p> <p>– S6 with apical pilosity directed radially (Figs 50, 52–54). Latero-apical angles of S6 flat, not curved outward ventrally (Figs 55, 57–59). Dorso-apical part of S8 with entire transverse carina (Figs 70, 72, 74) or with two semicircular, completely separated outgrowths (Fig. 73); lateral apodemes before pregladular area of this sternum more beveled on the sides (Figs 65, 67–69). Inner basal part of gonostylus with longer tooth or lobe-like projection, or penis valvae are wider than gonostylus. Galea brown to dark brown, tightly covered with small tubercles that in some places form the thin wavy striae (Figs 9, 39, 41, 42). Glossa wider and relatively shorter, 2.5-3.5 times as long as wide (Figs 45, 47–49). Mid and hind legs with last article brown to black.................................................................................................... 2</p> <p>2. Marginal part of S6 very slightly notched centrally and with white pilosity (Figs 53, 58). Apical third of S7 strongly widened, its marginal part slightly notched centrally (Fig. 63); dorso-apical part of S8 with two deeply separated semicircular outgrowths (Fig. 73). Penis valves narrower than gonostyle; inner basal part of gonostylus with a long spine-like tooth (Figs 78, 84). Pilosity of the body silver-white. Distribution: species endemic to central and southern Spain (Fig. 88)......................................................................................... Dasypoda iberica Warncke, 1973</p> <p>– Marginal part of S6 strongly triangularly notched centrally, forming two separate projections covered with a brown pilosity (Figs 50, 52, 54). Apical third of S7 narrowed, its marginal part strongly notched centrally (Figs 60, 62, 64); dorso-apical part of S8 with entire transverse carina very slightly notched centrally (Figs 70, 72, 74). Inner basal part of gonostylus with a small tooth or bilobed without a long spine-like tooth. Penis valves as wide as the gonostyle (Figs 75, 77, 79). Pilosity of the body mostly yellowish to brownish............................................................................ 3</p> <p>3. Face with a patch of pure white hairs surrounded by a pure black pilosity. S6 apical projections with right angles on the inner edges (Figs 52, 57). Gonostylus narrowed at apex, lanceolate, with a small tooth at the base (Figs 77, 83). Distribution: species bound to the western part of the Mediterranean basin (Italy, France, Spain, Portugal and Morocco; Fig. 87)...................................................................................... Dasypoda crassicornis Friese, 1896</p> <p>– Hair of the face with a different colour pattern. Apical projections of S6 with rounded angles on the inner edges (Figs 50, 54). Gonostyli bilobed without spine-like tooth at the base (Figs 75, 79)............................................. 4</p> <p>4. 1 st flagellomere relatively shorter, conical, 1.4 times as long as wide at the apex (Fig. 43). Face with a yellowish pilosity intermixed with black hairs especially on the paraocular areas. Apical projections of S6 with rounded angles on the outer edges (Fig. 55). S8 not notched laterally before the widened basal part (Fig. 65). Gonostylus with a well-developed internal lobe strongly thickened in the middle and with very dense long pubescence on the side facing the penis valvae; external lobe rounded at the top (Figs 75, 80, 81). Distribution: species only recorded in the northern parts of Algeria and Tunisia (Fig. 87)............................................................................... Dasypoda brevicornis Pérez, 1895</p> <p>– 1 st flagellomere relatively longer, partly cylindrical in apical half, 1.7 times as long as wide at the apex (Fig. 44). Face with a white pilosity except vertex and adjacent parts that intermixed with black hairs. Apical projections of S6 with right angles on the outer edges (Fig. 59). S8 deeply notched laterally before the widened basal part (Fig. 69). Inner lobe of gonostylus widened in a circular shape and flat at the apical half, racquet-like with a widened base, and covered with very sparse long setae on surface facing penis valves; external lobe of gonostylus wide, its apex oblique truncated (Figs 79, 85, 86). Distribution: species currently known from Morocco and Tunisia (Fig. 89)................ Dasypoda schwarzi Radchenko &amp; Michez sp. nov.</p></div> 	https://treatment.plazi.org/id/3A393D28FFEDFFD4FF6DDF8E9FE1FC57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Radchenko, Vladimir G.;Ghisbain, Guillaume;Michez, Denis	Radchenko, Vladimir G., Ghisbain, Guillaume, Michez, Denis (2022): A new bee species of the genus Dasypoda Latreille (Hymenoptera, Apoidea) from Northwest Africa with comparative remarks on the subgenus Microdasypoda Michez. Zootaxa 5188 (1): 74-86, DOI: https://doi.org/10.11646/zootaxa.5188.1.4
