identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03EFB311FFB1FF94FF79FE89FC80FE4D.text	03EFB311FFB1FF94FF79FE89FC80FE4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus punctatus Frolov & Scholtz 2003	<div><p>Sarophorus punctatus Frolov &amp; Scholtz, 2003</p> <p>(Figs 1–5)</p> <p>Type locality. Keurboomstrand [Garden Route District Municipality, Western Cape Province, South Africa].</p> <p>Type material examined. Holotype, male (TMSA, aedeagus extracted): “S.Afr., S. Cape [Western Cape Province] | Keurboomstrand | 34[°].00[′]S-23[°].27[′]E” || “ 8.12.1976,E-Y:1301| groundtraps [pitfall traps] 8days | leg.EndrödyYounga” || “ SAROPHORUS | tuberculatus | Cast. | det.Y.Camberfort” || “ HOLOTYPUS | Sarophorus | punctatus | A. Frolov det. 2002” || “ TM SOUTH Africa | TMSC 08252”. Paratypes, 5 females (TMSA): same data as holotype.</p> <p>Frolov &amp; Scholtz (2003) originally described this species from two males and six females, all with the same data. We studied five female paratypes as well as the holotype male.</p> <p>Material examined representing new distribution records (21 specimens). SOUTH AFRICA. Eastern Cape Province. 3♂, 1♀ Olifantskop Pass, 33°19′08″S 25°56′23″E, 25-27.i.2021, GM Daniel &amp; WP Strümpher, site 1, forest (deep valley), pitfall trap baited with pig dung + chicken livers, (BMSA; BMSA(C) 121989, 121990, 121993, 121998). 6♂, 2♀ Zuurberg Pass, 33°22′46″S 25°42′37″E, 25-27.i.2021, GM Daniel &amp; WP Strümpher, site 1, forest (deep valley), pitfall trap baited with pig dung + chicken livers (BMSA; BMSA(C) 117000–11705, BMSA(C) 122112). 4♂, 1♀ idem (TMSA, preserved in 99% ethanol). 2♂, 2♀ Makhanda [formerly Grahamstown], Signal Hill, 33°19′57″S 26°32′27″E, 26-28.i.2021, GM Daniel &amp; WP Strümpher, site 2, indigenous forest (deep valley), pitfall trap baited with pig dung + chicken livers (TMSA).</p> <p>Distribution and broad habitat associations. The new distribution records show that Sarophorus punctatus is more widespread than previously thought with a disjunct range that now comprises Keurboomstrand (Western Cape Province - type locality), Olifantskop Pass, Zuurberg Pass, and Makhanda (Eastern Cape Province - new provincial distribution records) (Fig. 5). The considerable expansion of the species’ known range, further inland to the east of the type locality, suggests that forest patches of the Western and Eastern Cape have been under sampled. Such under sampling of isolated forest fragments could explain the apparent rarity of S. punctatus and some other forest associated members of the genus (e.g., S. frolovi Roets, 2017). Additional sampling is needed in the two provinces to get a better understanding of its distribution, rarity and habitat preferences. These efforts may help to retrieve critical data on additional dung beetle species, which are known to be associated with woody vegetation, in the Forest, Fynbos and Renosterveld biomes.</p> <p>Biological notes. As a generality, species of the genus Sarophorus seem to show a bias to vegetation offering shade (Davis et al. 2020). Over 12 months in Gauteng, quantitative data for the widespread S. costatus (Fåhraeus in Boheman, 1857) (costatus group) recorded 271 individuals in shaded thickets but only seven in grassland (Davis 1996). Several, other, poorly-known species of the tuberculatus group have only been recorded from type- or other localities within patches of dense, shaded, savanna woodland (S. carinatus Frolov &amp; Scholtz, 2003), renosterveld (S. diabolus Roets, 2017) or forest (S. frolovi Roets, 2017) (Davis et al. 2020). As regards S. punctatus, it may be particularly cool-adapted as it was recorded in coastal areas or in deep forested valleys (new records) under cool, rainy conditions.</p> <p>Conservation status. For many of the Sarophorus species in South Africa, basic information on their distributions, biology, ecology, and natural history is severely lacking. Therefore, from a conservation standpoint, it is important to document such baseline data through further field work. For instance, S. punctatus was recently assessed as endangered (EN) (Davis et al. 2020) based on its known occurrence at a single, disturbed forest patch and no further records over a period of 44 years. This assessment may remain valid as the Extent of Occurrence (EOO) is still less than 5000 km 2 at less than five localities. However, addition of further localities might demand a reassessment although this should be tempered by any increased threat to the woody vegetation of the Western and Eastern Cape.</p> <p>Additional Sarophorus material used for comparison:</p> <p>Sarophorus carinatus Frolov &amp; Scholtz, 2003. Holotype, female (TMSA): “Z.A.49 | Lydenburg D. [district] | Ohrigstad [24°45’S 30°34’E]” || “Humus | III-1962 ” || “N. LELEUP leg.” || “ HOLOTYPUS | Sarophorus | carinatus | A. Frolov det 2002” || “TM SOUTH Africa | TMSC08181 ”.</p> <p>Sarophorus frolovi Roets, 2017. Holotype, male (SANC: SANC-TYPH-02478, aedeagus extracted): “ SOUTH AfRICA: KwaZulu- | Natal Province, Midlands, | Weza forestry plantations, | –30.61680 29.683626, | 02-2011, F. Roets &amp; J. Pryke” || “100 ml pig dung and | chicken liver mix (8:1 ratio) | left in the field for 2 days” || “ HOLOTYPE | Sarophorus frolovi | Roets 2017 ”.</p> <p>Sarophorus tuberculatus (Laporte de Castelnau, 1840). SOUTH AFRICA: Eastern Cape Province: Algoa Bay, Brauns (2 specimens, TMSA); Addo Park, 16-20.xii.1996, R. Wolmarans (2 specimens, TMSA). Western Cape Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.766666&amp;materialsCitation.latitude=-33.4" title="Search Plazi for locations around (long 22.766666/lat -33.4)">Swartberge Hagas Farm</a>, 33°24’S 22°46’E, 2.iii.1979, sandy valley, S. Endrödy-Younga (1 specimen, TMSA); Little Karoo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.316668&amp;materialsCitation.latitude=-33.416668" title="Search Plazi for locations around (long 22.316668/lat -33.416668)">Raubenheimer Dam</a>, 33°25’S 22°19’E, 21.x.1993, donkey dung, S. Endrödy-Younga (2 specimens, TMSA); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.183332&amp;materialsCitation.latitude=-33.25" title="Search Plazi for locations around (long 18.183332/lat -33.25)">8 km W of Ysterfontein</a>, 33°15’S 18°11’E, 28.viii.1983, groundtraps, 70 days, Endrödy-Younga and Penrith (1 specimen, TMSA); Ysterfontein, 14.ix. 1976, N.J. Duke (5 specimens, TMSA).</p> </div>	https://treatment.plazi.org/id/03EFB311FFB1FF94FF79FE89FC80FE4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79F949FCB1F8BD.text	03EFB311FFB3FF94FF79F949FCB1F8BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus angolensis Frolov 2004	<div><p>Sarophorus angolensis Frolov, 2004: 95</p> <p>Type locality: Tchivinguire Huíla [15°10′S 13°21′E], SE 1513 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=13.35&amp;materialsCitation.latitude=-15.166667" title="Search Plazi for locations around (long 13.35/lat -15.166667)">Ab</a> [Huíla, Angola].</p> <p>Distribution: Angola, Namibia (Frolov 2004; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79F949FCB1F8BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79FD3DFC72FC81.text	03EFB311FFB3FF94FF79FD3DFC72FC81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus bidentatus Frolov & Scholtz 2003	<div><p>Sarophorus bidentatus Frolov &amp; Scholtz, 2003: 187</p> <p>Type locality: Namaqualand, Kamieskroon [30°12’S 17°56’E, Northern Cape, South Africa].</p> <p>Distribution: South Africa (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79FD3DFC72FC81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79FC8DFC72FC71.text	03EFB311FFB3FF94FF79FC8DFC72FC71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus carinatus Frolov & Scholtz 2003	<div><p>Sarophorus carinatus Frolov &amp; Scholtz, 2003: 187</p> <p>Type locality: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.566668&amp;materialsCitation.latitude=-24.75" title="Search Plazi for locations around (long 30.566668/lat -24.75)">Lydenburg Distr.</a>, Ohrigstad [24°45’S 30°34’E, Mpumalanga, South Africa].</p> <p>Distribution: South Africa (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79FC8DFC72FC71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79F8D9FB7AF82D.text	03EFB311FFB3FF94FF79F8D9FB7AF82D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus cicatricosus (Peringuey 1901)	<div><p>Sarophorus cicatricosus (Péringuey, 1901: 280)</p> <p>Type locality: Mashunaland, Salisbury [Harare: Zimbabwe].</p> <p>Distribution: Democratic Republic of the Congo, Zambia, Zimbabwe (Frolov &amp; Scholtz 2003).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79F8D9FB7AF82D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79FC1DFCB3FBE1.text	03EFB311FFB3FF94FF79FC1DFCB3FBE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus diabolus Roets 2017	<div><p>Sarophorus diabolus Roets, 2017: 266</p> <p>Type locality: Riebeeck-Kasteel, Porseleinberg, –33.45995[°], 18.88627[°] (Western Cape, South Africa).</p> <p>Distribution: South Africa (Roets et al. 2017; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79FC1DFCB3FBE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79FB6DFCB3FB51.text	03EFB311FFB3FF94FF79FB6DFCB3FB51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus frolovi Roets 2017	<div><p>Sarophorus frolovi Roets, 2017: 268</p> <p>Type locality: Midlands, Weza forestry plantations, –30.61680[°] 29.683626[°] (KwaZulu-Natal, South Africa).</p> <p>Distribution: South Africa (Roets et al. 2017; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79FB6DFCB3FB51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB3FF94FF79FA4DFC72F9B1.text	03EFB311FFB3FF94FF79FA4DFC72F9B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus tuberculatus (Laporte de Castelnau 1840)	<div><p>Sarophorus tuberculatus (Laporte de Castelnau, 1840: 88)</p> <p>Type locality: Afrique [Africa].</p> <p>Distribution: South Africa (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB3FF94FF79FA4DFC72F9B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB4FF93FF79FF7DFB88FF41.text	03EFB311FFB4FF93FF79FF7DFB88FF41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus costatus (Fahraeus 1857)	<div><p>Sarophorus costatus (Fåhraeus in Boheman, 1857: 204)</p> <p>Type locality: Terra Natalensi [KwaZulu-Natal, South Africa].</p> <p>Distribution: Botswana, South Africa, Zimbabwe (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB4FF93FF79FF7DFB88FF41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB4FF93FF79FECDFC72FE31.text	03EFB311FFB4FF93FF79FECDFC72FE31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus latus Frolov & Scholtz 2003	<div><p>Sarophorus latus Frolov &amp; Scholtz, 2003: 195</p> <p>Type locality: Farm Rhenosterpoort [25°43’S 28°56’E, Gauteng, South Africa].</p> <p>Distribution: South Africa (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB4FF93FF79FECDFC72FE31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB4FF93FF79FE5DFC8AFDA1.text	03EFB311FFB4FF93FF79FE5DFC8AFDA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus nitidus Frolov & Scholtz 2003	<div><p>Sarophorus nitidus Frolov &amp; Scholtz, 2003: 96.</p> <p>Type locality: Tanganyika Terr., Longido, Masai Distr., [Arusha Region, Tanzania].</p> <p>Distribution: Kenya, Tanzania (Frolov &amp; Scholtz 2003; Frolov 2004).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB4FF93FF79FE5DFC8AFDA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
03EFB311FFB4FF93FF79FDADFC72FD11.text	03EFB311FFB4FF93FF79FDADFC72FD11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sarophorus striatus Frolov & Scholtz 2003	<div><p>Sarophorus striatus Frolov &amp; Scholtz, 2003: 193</p> <p>Type locality: 23 km S of Avontuur [Eastern Cape, South Africa].</p> <p>Distribution: South Africa (Frolov &amp; Scholtz 2003; Davis et al. 2020).</p></div> 	https://treatment.plazi.org/id/03EFB311FFB4FF93FF79FDADFC72FD11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Strümpher, Werner P.;Daniel, Gimo M.;Davis, Adrian L. V.	Strümpher, Werner P., Daniel, Gimo M., Davis, Adrian L. V. (2022): Woody plant communities of southern South Africa and new distribution records for the rare dung beetle species Sarophorus punctatus Frolov & Scholtz, 2003 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5188 (4): 396-400, DOI: https://doi.org/10.11646/zootaxa.5188.4.7
