taxonID	type	description	language	source
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	type_taxon	Type species: Wagneromyia aquiri Vieira, Cordeiro, Bravo & Araújo, sp. nov., by present designation.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	etymology	Etymology. The genus name is in honor to Dr. Rüdiger Wagner, dipterologist and specialist in Psychodidae, with the Greek suffix myia meaning “ fly ”. The name is feminine.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Eyes contiguous; antenna with 14 flagellomeres, nodiform (first to 11 th flagellomere), 12 th, 13 th, and 14 th flagellomeres reduced without internodes; setae alveoli of flagellomeres arranged in longitudinal rows; 14 th flagellomere with one to three apical spiniform setula; palpus with four segments, fourth segment equal in length or lesser than the first; second segment of the palpus longer than the other segments; first flagellomere with a dorsal patch of sensilla; wing membrane pilose, with pilosity arranged parallel to the veins; gonocoxites contiguous at base; single aedeagus with lateral folds; epandrial appendage with three apical tenacula.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Description. Male. Head semicircular, eye bridge with four rows of facets; eyes contiguous; group of setae on the frons not reaching the lower limit of the eye bridge; interocular suture present or absent, when present small and U-inverted shape. antenna with cylindrical scape slightly longer than subspherical pedicel; 14 nodiform flagellomeres; pair of ascoids, Y-shaped, in each flagellomere; small additional sensilla near the ascoid insertion (Fig. 5), on flagellomeres I-X; setae alveoli of flagellomeres arranged in longitudinal rows; flagellomeres 12 th, 13 th and 14 th reduced, separated from each other, without internodes; 11 th, 13 th with stout and spiniform setulae and 14 th flagellomere with one to three apical spiniform setula; palpus with four segments, first of one with a group of dorsal sensilla near the base (Fig. 3); length of fourth segment equal to or lesser than the first segment; second segment longer than the other segments. Labrum with convex dorsal margin, slightly expanded (Fig. 3); bulbous labella with robust and spiniform setulae on the inner margin and setae on the lateral margin (Fig. 4). Wing membrane pilose in lines parallel to the veins; C vein with two breaks at the base; Sc slightly curved at the apex, ending near the base of R 1; costal cell slightly infuscated; medial fork usually incomplete, the base of M 2 close to M 1 but always fading before joining M 1; sudden angulation at the end of R 2. Male terminalia with a narrow hypandrium, like a transversal band; gonocoxites usually dorsally contiguous; gonostylus conical, curved, with sparse setae and with an accessory long subapical seta; single aedeagus, narrow plate-like, with apex sometimes bifurcated and lateral folds; parameres of varied shapes, arising from the gonocoxal apodemes and articulated ventrally to the aedeagus at base; external lateral margin of the parameres generally projecting dorsally to the aedeagus forming a parameral sheath; parameral sheath of varied shape, connected to the gonocoxites, never exceeding the apex of the aedeagus; epandrium wider than long, with concave base and a large pseudospiracular opening; club-shaped epandrial appendages, with a group of setulae at the base, a group of subapical papilla on the dorsal surface and bearing three apical tenacula; hypoproct slightly projected and rounded, with pilose margin; lateral region of the hypoproct sclerotized and densely pilose, with setae a little longer than in the medial region; epiproct pilose and reduced. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	distribution	Distribution. South and Central America (Costa Rica and Acre, Brazil). The current distribution of Wagneromyia is restricted to the Neotropical region. Included species: Wagneromyia aquiri Vieira, Cordeiro, Bravo & Araújo, sp. nov., Wagneromyia bicalcarata (Quate, 1996) comb. nov., Wagneromyia deceptrix (Quate, 1996) comb. nov., Wagneromyia dimorpha (Quate, 1996) comb. nov., Wagneromyia majorina (Quate, 1996) comb. nov., Wagneromyia tripetala (Quate, 1996) comb. nov., and Wagneromyia tritenacula (Quate, 1996) comb. nov.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	discussion	Remarks. Among the Psychodini, the bulbous labellum and symmetrical aedeagus, as found in Wagneromyia, are features shared with the genera Eurygarka, Feuerborniella, Mucomyia Kvifte & Curler, 2018, Perithreticus Vaillant, 1973, Quatiella Botosaneanu & Vaillant 1970, Soeliella Kvifte, 2015, Trichosepedon Krek, 1999, and some species of Trichopsychoda Tonnoir, 1922 and Philosepedon. Thus, the species of Wagneromyia are easily differentiated from Psychodini genera that include species with laterally-compressed labella and / or an asymmetrical aedeagus, such as Psychoda, Epacretron Quate, 1965, Neomaruina Vaillant, 1963, Rhipidopsychoda Vaillant, 1991, Nielseniella Vaillant, 1972, Threticus Eaton, 1904, and Cookiellocapsa Ježek & Le Pont, 2016. Wagneromyia and Eurygarka share the presence of contiguous eyes but Eurygarka has ascoids with leaf-shaped anterior branches, fourth segment of the palpus greater in length than the other segments, male terminalia with sclerotized and pilose ventral epandrial plate and two tenacula on the epandrial appendages. On the other hand, Wagneromyia species have ascoids with simple posterior branches, fourth segment of the palpus smaller than or equal to the first segment, weakly sclerotized and bare ventral epandrial plate, and three tenacula on the epandrial appendages. Epandrial appendages with three tenacula, as in Wagneromyia can be found in Soeliella and some species transferred to Feuerborniella by Cordeiro et al. (2015), but the new genus can be recognized by the contiguous eye bridge, contrasting with the separate eyes in Feuerborniella and Soeliella, and by the length of the fourth segment of the palpus, smaller or equal to the first segment in the new genus and greater in length than the other segments in the other two genera. Furthermore, in Feuerborniella the terminal flagellomeres can be fused, while in Wagneromyia species they are separated, and in Soeliella, the aedeagus is wide and plate-shaped, being narrow and sometimes apically bifurcated in species of Wagneromyia. The species of the genus Mucomyia, Perithreticus, Philosepedon, Quatiella, Trichopsychoda and Trichosepedon differ from Wagneromyia by the separate eyes, at least in the lower facet rows, and the length of the first segment of the palpus, which is smaller than the others. Also, in Mucomyia the male terminalia has gonostylus without subapical seta, Perithreticus has more than five tenacula, Philosepedon has two tenacula, Quatiella has only one tenaculum in the epandrial appendage, in Trichopsychoda the parameres are absent and in Trichosepedon the gonostylus has a longitudinal groove carrying a row of stout setae and the epandrial appendages have two tenacula. In contrast, in males of Wagneromyia the gonostylus has a subapical seta and no distinct longitudinal groove, the epandrial appendage has three tenacula, and parameres are present. Omelková & Ježek (2012) proposed a monospecific subgenus Philothreticus Krek, 1999 for Philosepedon, with a contiguous eye bridge, but with the lower facet rows separated, ascoids with only two branches and asymmetric aedeagal complex, different from the new genus with all the rows of facets touching in the midline, ascoids with three branches, and the aedeagal complex symmetrical. A feature observed for most species of Wagneromyia is the labrum with a convex and slightly expanded dorsal margin. This characteristic has been only observed in this genus until now, but a review of this structure is still needed regarding the other genera of the tribe, as this structure was rarely described and illustrated in previous works. Furthermore, as described for Quatiella truncata Chagas & Cordeiro, 2015 and species of Feuerborniella auctt (Cordeiro et al. 2015; Pereira et al. 2015), the species of the new genus present an extension of the gonocoxal apodemes (= fused parameres), which surround the aedeagus to form a dorsal sheath, but which do not completely cover the aedeagus as in Brachycera (see Cumming & Wood 2017 for comments). Quate (1996) cited this structure, referring to it as projections of the anterior gonocoxal apodeme, Ibáñez-Bernal (2004) described it as the aedeagal sheath and Kvifte (2019) named these structures as parameral dorsal bridge. Although still not properly named, this is observed for other genera in illustrations (e. g. Kvifte et al. 2016, figs. 4, 8), however, it has gone unnoticed in works and therefore it is necessary to be revised in Psychodini species to understand how diagnostic it can be for the tribe’s genera.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 1 – 9)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Type material: holotype ♂. BR [Brazil], Acre, 15 Km a SE do Rio Branco, Embrapa, 170 m [altitude], 10 º 01 ’ S 67 º 41 ’ N, 7 – 16. VII. 2008, Malaise [trap] G. Melo [leg.] (MZFS); paratypes 9 ♂, same date and locality as holotype (MZFS).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	etymology	Etymology. The specific name, aquiri, refers to “ a’quiri ” or “ a’kiru ”, from the word uwákürü, from the dialect of the Ipurinã tribe, the original inhabitants of the region, from which the toponym Acre was derived. Acre is the name of the main river of the region and also the name of the Brazilian state where the species was collected. The name should be treated as a noun in apposition.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Medial fork incomplete; R 2 + 3 1.2 x length of R 2; apex of the gonostylus bent, with a small apical accessory seta and a long subapical seta on the ventral side; aedeagus straight, bending ventrolaterally to form a tubular-like structure that narrows to a small, bifurcated apex; parameres well sclerotized, broad at the base and tapering to a hook-shaped apex, curved outwards; parameral sheath subrectangular, with concave posterior margin; epandrial appendages with a group of five subapical papillae on the dorsal surface; hypoproct pilose with straight posterior margin and rounded vertices.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Description. Male. Head (Fig. 1) as in genus description; eye bridge with a small medial spur on the dorsal margin and five widely spaced supraocular setae; group of setae on frons does not extend to the lower limit of the eye bridge; frontoclypeal suture present, as a sclerotized area; antenna incomplete in all specimens, flagellomeres beyond 10 th lost in holotype; scape, pedicel, flagellomeres, and ascoids as in the genus description; palpus formula 1.0: 1.6: 1.2: 1.0 first segment with a dorsolateral group of sensilla in the middle (Fig. 3); clypeus with four to five prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and three setae on the lateral margin. Wing (Fig. 2) as in genus description; medial fork incomplete; R 2 + 3 1.2 x the length of R 2. Male terminalia (Figs. 6 – 9) with a narrow band-shaped hypandrium, wider medially; gonocoxites as in genus; gonostylus, with a sudden angulation near the apex, bearing a small apical accessory seta and a long subapical seta on the ventral surface (Figs. 6, 7); aedeagus bending ventrally on the sides forming a tubular-like structure with a tapered and bifurcated apex on the ventral view (Figs. 6, 7); parameres well sclerotized, broad at the base and tapering to a hook-shaped apex, curved outwards; parameres meet each other dorsally to the aedeagus forming a band-shaped parameral sheath with a small concavity at the midline (Fig. 6); epandrium as in genus; epandrial appendages as in genus, with a group of five subapical papillae on the dorsal surface and three apical tenacula with half the length of the appendages (Fig. 8); hypoproct pilose with straight posterior margin and rounded vertices (Fig. 8); epiproct as in genus (Fig. 9); subepandrial sclerite membranous. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 10 – 16)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 16). COSTA RICA, Heredia, Pr. Puerto Viejo de Sarapaqui, Est. Biol. LaSelva, Lt. Trp 22 - 25 / VII / 93, L. W. Quate. (INBIO CRI 001468679).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Medial fork incomplete, M 2 not reaching M 1; gonostylus apically bifurcated, like a chela; aedeagus with slightly enlarged bifid apex; subrectangular parameres apically bifurcated with an external arm like a hook and internal arm rounded; parameral sheath absent; hypoproct with medial lobe subtriangular in dorsal view.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male (holotype). Head (Fig. 10) as in genus description; eye bridge with small medial dorsal spur and six supraocular setae, the innermost three closest; group of setae on frons extending medially close to the limit of the inferior margin of the eye bridge; frontoclypeal suture as a slightly sclerotized area, incomplete laterally; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and one seta on the lateral margin; antenna (Fig. 11) as in genus; palpus formula 1.0: 1.5: 1.3: 0.8, first segment of palpus with a group of sensilla at the slightly inflated base. Wing (Fig. 12) as in genus; radial fork distal to medial fork; R 2 + 3 1.5 x the length of R 2; M 2 not reaching M 1 (medial fork incomplete). Male terminalia (Figs. 13 – 15) with hypandrium narrow, like a strip; gonocoxites as in genus; gonostylus apically bifurcated (Fig. 13), with two short and strong projections like a chela, and with a long subapical seta on the ventral margin; aedeagus with slightly enlarged bifid apex; parameres subrectangular (Figs. 13, 14), with the apical portion bifurcated into external arm hook-like, positioned dorsally to the aedeagus (Fig. 13), and an internal arm rounded, lateral to the aedeagus (Fig. 14); parameres not developed into a parameral sheath; epandrium (Fig. 15) damaged and in lateral view in the holotype; epandrial appendages (Fig. 15) curved in lateral view, base wider than the apex, with a group of three subapical papillae on the dorsal margin and tree apical tenacula with half the length of the appendices; epiproct short; hypoproct with medial lobe (Fig. 15). Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	discussion	Remarks. Although Quate (1996) described W. bicalcarata with separated gonocoxites, the examination of the holotype shows that they are contiguous as in the other species of the genus.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 17 – 24)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 24). COSTA RICA, Heredia Vara Blanca, 1800 m, 26 / VII / 93 L. W. Quate, 1800 m, Lt Trp, Euparal, LN — 239.500 - 519.000 (INBIO CRI 001469462).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. 14 th flagellomere with two apical spiniform setulae; medial fork complete; subtriangular parameral sheath with a rounded apex; aedeagus straight, with bifid apex, like two ventral lobes; parameral sheath subtriangular with a rounded apex; anterior gonocoxal apodemes with setae on the posteromedial margin, continuous with the ventral setae of the gonocoxite; hypoproct pilose and subovalate.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male (holotype). Head (Fig. 17) as in genus description; eye bridge with a very short dorsal medial spur and five supraocular setae; group of setae on frons not extending medially; frontoclypeal suture only as a slightly less sclerotized area; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin, and four setae on the external lateral margin, the most apical one shorter than the others; antenna (Fig. 19) as in genus; 14 th flagellomere with two apical stout and spiniform setulae; palpus formula: 1.0: 1.6: 1.2: 0.8, first segment of palpus with a group of sensilla near the base. Wing (Fig. 18) as in genus description; radial fork distal to medial fork, both complete; R 2 + 3 1.3 x the length of R 2. Male terminalia (Figs. 20 – 23) with hypandrium as a narrow band, little sclerotized; gonocoxite with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus (Figs. 20, 21) straight, with bilobed apex; parameres (Fig. 21) with a membranous (non-sclerotized) region between them and the gonocoxal apodemes; parameres with a broad base from where it projects dorsally to the aedeagus forming a subtriangular parameral sheath with a rounded apex (Fig. 20); anterior gonocoxal apodemes with setae on the posteromedial margin, continuous with the ventral group of setae of the gonocoxite (Fig. 21); epandrium (Fig. 22) typical for genus; epandrial appendages (Fig. 23) as in genus description, with a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct (Fig. 22) small and rounded; hypoproct (Fig. 23) pilose and subovalate. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 25 – 30)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 30). COSTA RICA, Heredia, Puerto Viejo de Sarapaqui, Est. Biol. LaSelva, Lt. Trp 23 / VII / 93, L. W. Quate, 50 - 100 m. (INBIO CRI 001468641)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Medial fork incomplete; gonostylus with small subapical spines lined on the ventrolateral margin and a long subapical seta; aedeagus with a straight and tapered bifid apex; parameres long, robust, losing sclerotinization as they approach an outward hook-like apex; parameral sheath poorly sclerotized, as a narrow and membranous stripe.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male. Head (Fig. 25) as in genus description; eye bridge with a small medial spur and five supraocular setae, the two or three innermost closest; group of setae on frons extending medially; frontoclypeal suture only as a slightly less sclerotized area; antenna incomplete in holotype, flagellomeres beyond 7 th lost; ascoids lost in holotype; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin and one seta on the external lateral margin; palpus formula 1.0: 1.4: 1.3: 0.9, first segment of palpus as in genus description. Wing (Fig. 26) as in genus description; complete radial fork distal to an incomplete medial fork; R 2 + 3 1.3 x the length of R Male terminalia (Figs. 27 – 29) with hypandrium as a narrow band; gonocoxites as in genus description; gonostylus with small subapical spines lined on the ventrolateral margin and a long subapical seta; aedeagus straight and bifid on distal half; parameres (Figs. 27, 28) reaching the apex of aedeagus, robust and wide at the base, losing sclerotinization as they approach an outward hook-like apex; parameral sheath (Fig. 27) poorly sclerotized, like a narrow and membranous stripe; epandrium (Fig. 29) as in genus description, but damaged; epandrial appendages with a group of short setulae on the base, a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct (Fig. 29) short; hypoproct not very pronounced. 2. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 31 – 37)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 37). COSTA RICA, Heredia, S Rafael de Vara Blanca, Rio Santo Dominigo, 18 - 26 / VII / 93 L. W. Quate, 1700 m, Lt Trp, Euparal. (INBIO CRI 001468528).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Frontoclypeal suture absent; medial fork complete; aedeagus with bifid and straight apex; parameres broad at the base, not extending posteriorly; parameral sheath bilobed.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male. Head (Fig. 31) as in genus description; eye bridge with dorsal medial spur and five to six supraocular setae; group of setae on frons not extending medially to the base of eye bridge; frontoclipeal suture absent; antenna (Fig. 32) as in genus description; clypeus with four prominent lateral alveoli and slightly concave apical margin; labella with robust spiniform setulae on the inner margin; palpus formula 1.0: 1.4: 1.2: 0.9. Wing (Fig. 33) as in genus description; radial fork complete, distal to complete medial fork; R 2 + 3 1.5 x the length of R 2. Male terminalia (Figs. 34 – 36) with hypandrium reduced to a narrow band wider medially, poorly sclerotized; gonocoxite with setae on the posteromedial dorsal margin, lined up near the base; gonostylus as in genus description; aedeagus (Fig. 34) with bifid and straight apex; parameres (Fig. 35) united to the gonocoxal apodemes by a membranous area; parameres broad at the base, not extending posteriorly, projecting dorsally to the aedeagus to form a bilobed parameral sheath (Fig. 34); anterior gonocoxal apodemes without setae; epandrium and appendages (Fig. 36) as in genus description; epandrial appendages with a group of three subapical papillae on the dorsal margin and with three apical tenacula longer than half the length of the appendices; epiproct inconspicuous and hypoproct slightly pronounced, wider than long. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 38 – 44)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 44). COSTA RICA, Heredia Vara Blanca, 1800 m, 26 / VII / 93 L. W. Quate, 1800 m, Lt Trp, Euparal, LN — 239.500 - 519.000 (INBIO CRI 001469454).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. Medial fork incomplete; gonocoxites fused at the base; posterior gonocoxal apodemes forming a highly sclerotized ring, dorsal to the aedeagus and the parameral sheath; parameral sheath membranous and rounded; parameres well sclerotized, broad at based tapering to a hook-shaped apex; hypoproct little pronounced, ellipsoid, wider than long.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male. Head (Fig. 38) as in genus description; eye bridge without medial dorsal spur and with seven supraocular setae; fourth row of facet nearly separated; group of setae on frons not extending medially toward base of eye bridge; frontoclipeal suture only as a slightly less sclerotized area; clypeus with nine prominent lateral alveoli and slightly concave apical margin; palpus formula 1.0: 1.5: 1.4: 0.9, first segment of palpus with a group of sensilla in the medial region; labella with four robust spiniform setulae on the inner margin, and six setae on the external lateral margin; antenna (Fig. 39) as in genus description. Wing (Fig. 40) as in genus description; radial fork complete, distal to incomplete medial fork; R 2 + 3 1.4 x the length of R 2. Male terminalia (Figs. 41 – 43) with hypandrium as a narrow band, a little wider medially; gonocoxites (Fig. 41) fused at the base, with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus narrow, tapering to a single apex; posterior gonocoxal apodeme fused medially, forming a highly sclerotized ring — parameral sclerite, dorsal to the aedeagus and the parameral sheath (Fig. 41); parameres continuous with the posterior gonocoxal apodemes; parameres (Figs. 41, 42) well sclerotized, broad at based tapering to a hook-shaped apex; membranous parameral sheath with a rounded apex (Fig. 41); anterior gonocoxal apodemes (Fig. 42) with setae on the posteromedial margin, continuous with the ventral setae of the gonocoxite; epandrium (Fig. 43) as in genus description; epandrial appendages as in genus description with a group of three subapical papillae on the dorsal margin and three apical tenacula half the length of the appendices; epiproct inconspicuous and hypoproct (Fig. 43) not very pronounced, ellipsoid, wider than long, both pilose. Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	(Figs 45 – 51)	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	materials_examined	Material examined: holotype ♂ (Fig. 51). COSTA RICA, Guanac. Estacion Pitilla, 700 m. 9 Km S Santa Cecilia, 10 - 14 / VII / 93, L. W. Quate, Light Trap Euparal. (INBIO CRI 001469215).	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	diagnosis	Diagnosis. 14 th flagellomere with three apical stout and spiniform setulae; medial fork complete; aedeagus with lateral folds subapically, forming an abrupt bottleneck anterior to a bifurcated apex; parameres broad at the base, not extending posteriorly; parameral sheath trapezoidal; gonocoxal apodemes with four setae posteriorly.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	description	Redescription. Male. Head (Fig. 45) as in genus description; eye bridge without medial dorsal spur and with five to six supraocular setae; group of setae on frons extending medially near the inferior limit of the eye bridge; frontoclipeal suture only as a slightly less sclerotized area; clypeus with three prominent lateral alveoli and slightly concave apical margin; labella with four robust spiniform setulae on the inner margin and three setae on the external lateral margin; antenna (Fig. 46) as in genus description; palpus formula 1.0: 1.5: 1.2: 0.8, first segment of palpus with a group of sensilla near the base. Wing (Fig. 47) as in genus description; radial fork complete, distal to the complete medial fork; R 2 + 3 1.4 x the length of R 2. Male terminalia (Figs. 48 – 50) with hypandrium reduced to a narrow band; gonocoxite with setae on the posteromedial dorsal margin lined up near the base; gonostylus as in genus description; aedeagus (Figs. 48, 49) with subapical lateral folds, forming a bottleneck anterior to the bifurcated apex; parameres united to the gonocoxal apodemes by a membranous area; parameres (Fig. 49) broad at the base, not extending posteriorly, projecting dorsally to the aedeagus to form a trapezoidal parameral sheath (Fig. 48); gonocoxal apodema with four setae posteriorly; epandrium (Fig. 50) as in genus description; epandrial appendages with a group of short setulae on the base, a group of three subapical papillae on the dorsal margin and three apical tenacula of half the length of the appendices; epiproct inconspicuous and hypoproct (Fig. 50) slightly pronounced, wider than long Female. Unknown.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
CB6675115E61FFB98CE5BFA0EAC1100D.taxon	discussion	Identification key for males of Wagneromyia species The species of the new genus are present in the key of Quate (1996) to the genus Philosepedon. A new key is provided below to identify the new described species and the species transferred to the new genus.	en	Vieira, Gabriel Santos, Cordeiro, Danilo Pacheco, Bravo, Freddy, Araújo, Maíra Xavier (2022): A new genus of Psychodini (Diptera: Psychodidae) from the Neotropical region. Zootaxa 5188 (4): 330-346, DOI: https://doi.org/10.11646/zootaxa.5188.4.2
