taxonID	type	description	language	source
A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	description	Figs 2, 3, 4	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	materials_examined	Material examined. Holotype: MNHN-IA- 2015 - 1608, complete. Paratypes: AM W. 53773, complete; AM W. 53774, complete, some parapodia mounted for SEM; MNHN-IA- 2021 - 725, anterior part only. All material collected from South Pacific Ocean, Papua New Guinea, New Ireland, CP 4254, - 2.483 ° S, 150.66 ° E, depth 273 - 324 m, April 2014.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	description	Description (based on holotype, with variation in parentheses for paratypes). Preserved specimens strongly iridescent (Fig. 2 A, C, D), ~ 230 (220) chaetigers, 112 mm (71 - 157) long, 3.3 mm (2.1 - 5.4) width at chaetiger 10, excluding parapodia. Body elongated and tapered gradually at posterior end (Fig. 2 B). Prostomium rounded anteriorly with two dorsoventrally flattened buccal lips an anterior notch between them, notch more visible ventrally (Fig. 2 A, C, D). Two palps and three antennae slender and tapering, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, longer than palps (same size), shorter (same size) than prostomium (Fig. 2 C). Eyes present, one pair, brownish, very faint, present at posterior base between palps and lateral antennae. First peristomial ring ~ 3 x longer than second one dorsally (Fig. 2 C). Maxillary apparatus (Fig. 3 D, E) partially everted in holotype or paratypes. Formula as follows: MF = 1 + 1, 5 + 5, 6 + 0, 4 + 9, 1 + 1. MI ~ 2 x longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of rounded wings situated at posterolateral margins. MI forceps-like, without attachment lamellae, sub-right-angle falcal arch. Closing system ~ 4 x shorter than MI. Ligament between MI and MII golden. MII without attachment lamella, teeth triangular, distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, longer than left MIV, curved, with equal-sized triangular teeth; short attachment lamella situated in the centre of posterior edge of maxilla, thin, dark. Left MIV short (less than half the size of right MIV) with wide, rounded base, left-most teeth longer than right-most ones; attachment lamella dark, semi-circular. Right MIV long, with teeth triangular, decreasing in size posteriorly; attachment lamella wide, semi-circular, dark. MV, paired, as long as high, with a dorsal curved tooth. Mandibles light brown, concentric stripes not visible; longer than MI; cutting plates whitish, without dorsal teeth (Fig. 3 E). First few parapodia located below middle line of body wall, but gradually positioned dorsally to approximately midline in subsequent segments (Fig. 2 A). Notopodial cirri slender, tapering, slightly longer than ventral cirri, thinner posteriorly (Fig. 3 A-C). Chaetal lobes comprising a low pre-chaetal lip and a globular post-chaetal lobe. Ventral cirri bluntly conical, with rounded tip, shorter than post-chaetal lobes anteriorly, thereafter slightly longer than post-chaetal lobes (Fig. 3 A-C). Branchiae pectinate, commencing from chaetiger 20 (18 - 19) and continuing to near end, very short anteriorly, longer in medium chaetigers but not reaching mid-dorsal line; number of filaments increasing from 1 - 3 anteriorly to 4 - 6 in mid-body, decreasing to 3 - 4 in last several chaetigers (Figs 2 E, 3 A-C). Aciculae black with paler blunt tips, approximately three or four per parapodium in anterior chaetigers, one or two per parapodium in middle chaetigers, and one per parapodium in posterior chaetigers. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers (Fig. 3 A-C). Pectinate chaetae commencing from approximately chaetiger 20 to near end, three types identified. Type 1 from anterior parapodia to mid-body: isodont-narrow-slender (INS), having ~ 20 short internal teeth, each tooth prolonged by a thin filament (Fig. 4 A, B). Types 2 and 3 from posterior parapodia only (Fig. 4 C, D): type 2 asymmetrical, anodont-wide-thick (AWT), having ~ 10 thick internal teeth, each tooth prolonged by a thin filament; type 3 asymmetrical, anodont-wide-slender (AWS), having ~ 20 internal teeth, each tooth prolonged by a thin filament (Fig. 4 D). Subacicular chaetae with compound spinigers and subacicular hooks (Fig. 3 F). Compound spinigers commencing from first chaetiger to near pygidium, with long, tapered blade. Subacicular hooks transparent, commencing from anterior chaetigers 43 - 52 (range for type material) to near end and inferior to bundle of spinigers, one per parapodium; much thinner than aciculae; subacicular hooks bidentate (Fig. 3 F). Pygidium round, with crenulated margin, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 2 - 3 x length of ventral pair (Fig. 2 B).	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	etymology	Etymology. The species name refers to the decomposing banana leaves among which all the specimens were found.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	distribution	Distribution. Only known from type locality.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	description	Figs 5, 6, 7	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	materials_examined	Material examined. Holotype: MNHN-IA- 2015 - 1559, complete, South Pacific Ocean, Papua New Guinea, New Britain, CP 4264, - 4.6 ° S, 152.4 ° E, depth 430 - 523 m, April 2014. Paratypes: MNHN-IA- 2015 - 1415, complete, South Pacific Ocean, Papua New Guinea, New Britain, CP 4337, - 6.083 ° S, 149.316 ° E, depth 287 - 447 m, May 2014; MNHN-IA- 2015 - 1593, anterior part only, South Pacific Ocean, Papua New Guinea, New Britain, CP 4329, - 6.133 ° S, 149.166 ° E, depth 250 - 500 m, May 2014; AM W. 53770, complete (several parapodia mounted for SEM), South Pacific Ocean, Papua New Guinea, New Britain, CP 4264, 4.6 ° S, 152.4 ° E, depth 430 - 523 m, April 2014; AM W. 53771, anterior part only, mounted for SEM, South Pacific Ocean, Papua New Guinea, New Britain, CP 4334, - 6.116 ° S, 149.166 ° E, depth 430 - 620 m, May 2014; AM W. 53772, complete, gravid, South Pacific Ocean, Papua New Guinea, New Britain, CP 4266, - 4.6166 ° S, 152.416 ° E, depth 575 - 616 m, April 2014.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	materials_examined	Additional material. MNHN-IA- 2015 - 1610, anterior part only, gravid, South Pacific Ocean, Papua New Guinea, New Ireland, CP 4260, - 2.9 ° S, 151.1 ° E, depth 350 - 847 m, April 2014; MNHN-IA- 2015 - 1949, anterior part only, South Pacific Ocean, Papua New Guinea, New Ireland, CP 4434, - 2.25 ° S, 150.8 ° E, depth 1066 - 1200 m, August 2014; MNHN-IA- 2015 - 1615, anterior part only, few parapodia used for molecular analysis, South Pacific Ocean, Papua New Guinea, New Hanover, CP 4482, - 2.683 ° S, 150.116 ° E, depth 761 - 825 m, September 2014.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	description	Description (based on holotype, with variation in parentheses for paratypes). Specimens strongly iridescent (Fig. 5 B), 88 (89) chaetigers, 45 mm (41 - 80) long, 3.6 mm (2.5 - 2.8) width at chaetiger 10, excluding parapodia. Body elongated and tapered gradually at posterior end, anteriorly not flattened (Fig. 5 A). Prostomium bilobed, with buccal lips separated by a ventral notch only (Fig. 5 B). Two palps and three antennae slender and tapering, palpophores not visible, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, slightly longer than palps and prostomium (same size) (Figs 5 C, 7 A). Eyes absent. First peristomial ring ~ 1.8 x longer than second one dorsally (Figs 5 C, 7 A). Maxillary apparatus (Fig. 6 D, E) partially everted in holotype or paratypes. Formula as follows: MF = 1 + 1, 5 (6) + 5 (6), 7 + 0, 4 (3) + 10 (11), 1 + 1. MI ~ 2.5 x longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of rounded wings situated at posterolateral margins. MI forceps-like, without attachment lamellae, sub-right-angle falcal arch. Closing system ~ 4 - 5 x shorter than MI. Ligament between MI and MII rectangular, dark. MII without attachment lamella, teeth triangular, distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, longer than left MIV, curved, with equal-sized triangular teeth; short attachment lamella situated in the centre of posterior edge of maxilla, oval, dark. Left MIV short (half the size of right MIV) with wide, rounded base, two left teeth longer than right-most ones; attachment lamella dark, semi-circular. Right MIV long, with teeth triangular, decreasing in size posteriorly; attachment lamella wide, semi-circular, dark. MV, paired, longer than wide, with a long tooth pointed ventrally, and a rounded dorsal margin (Fig. 6 D). Mandibles dark with golden tips, with fine concentric stripes visible dorsally and ventrally, same size as MI; cutting plates whitish, with distinct growth rings, with three dorsal teeth (Fig. 6 E). Notopodial cirri very long, slender and, tapering (Fig. 6 A-C), 2 - 3 x longer than ventral cirri in all chaetigers. Pre-chaetal lobe inconspicuous. Post-chaetal lobe digitiform in the two or three first chaetigers, triangular with tapering tip from chaetiger 4, reducing in size from chaetiger 17, almost inconspicuous from chaetiger 28 (21) (Fig. 6 A-C). Ventral cirri (Fig. 6 A-C) bluntly conical, with slightly expanded bases and rounded tips from chaetigers 1 - 22, subconical and thinner thereafter. Branchiae pectinate (Figs 5 C, 6 A, 7 A), starting from chaetiger 7 (7) and continuing for a limited number of segments, until chaetiger 16 (14); with 8 - 16 long filaments. Aciculae black with paler blunt tips, 2 - 4 per parapodium along the body. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers. Pectinate chaetae commencing from first few chaetigers to near end, one type identified as heterodont-narrow-slender (HNS; Fig. 7 B, D, E), one outer tooth very long (Fig. 7 E), having nine or ten short internal teeth, each tooth prolonged by a thin filament. Subacicular chaetae with compound falcigers and subacicular hooks (Figs 6 F, G, 7 B, C). Compound falcigers bidentate, with short blade and large teeth, commencing from first chaetiger to near pygidium, with more than 50 chaetae within a parapodium in anterior part, with ~ 10 chaetae in mid-body and ~ 3 - 5 in last chaetigers (Fig. 6 F). Subacicular hooks black with pale yellow tip, commencing from anterior chaetiger 20 (24) to end, most of the body with one hook per parapodia, but some posterior chaetigers with two, subacicular hooks bidentate (Figs 6 G, 7 B, C). Pygidium round and crenulated, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 2 (3) x length of ventral pair (Fig. 5 D).	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	etymology	Etymology. This species name refers to the type locality and geographical distribution of this species.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.taxon	distribution	Distribution. Papua New Guinea, Solomon Sea (New Britain) and Bismarck Sea (New Ireland).	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.taxon	description	Figs 8, 9	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.taxon	materials_examined	Material examined. Holotype: MNHN-IA- 2015 - 1519, entire, few parapodia used for molecular analysis, South Pacific Ocean, Papua New Guinea, New Ireland, CP 4260, - 2.9 ° S, 151.1 ° E, depth 350 - 847 m, April 2014. Paratype: MNHN-IA- 2015 - 1607, anterior part only, South Pacific Ocean, Papua New Guinea, New Britain, CP 4266, - 4.616 ° S, 152.25 ° E, depth 575 - 616 m, April 2014.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.taxon	description	Description (based on holotype, with variation in parentheses for paratype). Preserved specimens 197 (85 ant. part only) chaetigers, 101 mm (36 mm) long, 4.1 mm (2.8 mm) wide at chaetiger 10, excluding parapodia. Body elongated, slightly tapering at posterior end. Prostomium strongly bilobed with two dorsoventrally flattened buccal lips and an anterior notch between them (Fig. 8 B, C). Two palps and three antennae slender and tapering, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, shorter than prostomium (slightly longer for PNG 012), slightly longer than palps (palps very short for paratype PNG 12, but probably broken) (Fig. 8 C). Eyes absent. First peristomial ring approximately the same size as second one dorsally (Fig. 8 C). Maxillary apparatus yellow to golden brown, partially everted in holotype and paratype. Maxillae with carriers and four paired elements and one single one, formula as follows (Fig. 9 F): MF = 1 + 1, 4 + 4, 5 + 0, 3 + 6, 1 + 1. MI ~ 2 x longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of oval wings situated at posterolateral margins. MI forceps-like, without attachment lamellae; well-developed falcal arch. Closing system ~ 4 - 5 x shorter than MI. MII wide, without attachment lamella, teeth triangular, recurved, and distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, slightly shorter than right MIV, curved forming part of distal arc; with left four teeth recurved, equal-sized and triangular, two right teeth shorter and blunt, without attachment lamella. Left MIV short (half the size of right MIV) with wide, triangular base, left 2 teeth longer than right-most one; attachment lamella dark, semi-circular. Right MIV with teeth triangular, recurved, decreasing in size posteriorly; attachment lamella large, wide, best developed centrally. MV, paired, rectangular (longer than wide), with a broad cutting edge, and no clearly defined teeth (but following tradition to score as 1 + 1). Mandibles (Fig. 9 G) yellow to golden brown, slightly shorter than MI plus carriers; cutting plates whitish, with distinct growth rings. First two parapodia located below middle line of body wall, but gradually positioned dorsally to approximately midline in subsequent segments (Fig. 8 A). Notopodial cirri with large base and slender, tapering tip from anterior to mid-body chaetigers, digitiform cirri in posterior chaetigers; same size as neuropodial cirri, but shorter than post-chaetal lobe in anterior chaetigers (Fig. 9 A-C). Chaetal lobes comprising a low pre-chaetal lip and a large tongue-like post-chaetal lobe from first chaetiger to approximately chaetiger 25, almost inconspicuous thereafter. Ventral cirri bluntly conical until chaetiger 25, digitiform with bulbous base thereafter (Fig. 9 A-C). Branchiae with a long single filament (Figs 8 A, 9 B), commencing from chaetiger 31 (32) and continuing to mid-body (i. e., chaetiger 118 for holotype). Aciculae black with paler blunt tips, ~ four per parapodium in anterior chaetigers, two or three per parapodium in middle chaetigers, and one or two per parapodium in posterior chaetigers. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers. Pectinate chaetae commencing from first few chaetigers to near end, one type only (Fig. 9 H, I), with two or three pectinate chaetae per parapodium in anterior body, up to seven from posterior chaetigers, isodont-wide-thick (IWT) having 11 - 20 long teeth (Fig. 9 H, I). Subacicular chaetae compound falcigers and subacicular hooks (Fig. 9 D, E). Compound falcigers bidentate, with long blades and short teeth, commencing from first chaetiger to near pygidium, with more than 30 chaetae within a parapodium in anterior part, with ~ 20 chaetae in mid-body and ~ 5 - 7 in last chaetigers (Fig. 9 D). Subacicular hooks amber to black, with much paler tip, commencing from anterior chaetiger 28 (chaetiger 30) to near end, one per parapodium in anterior and posterior parts, few chaetigers with two hooks in middle body; slightly thinner than aciculae; subacicular hooks unidentate, with blunt tip (Fig. 9 E). Pygidium round, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 3 x length of ventral pair (Fig. 8 D).	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.taxon	etymology	Etymology. This species is dedicated to Joana Zanol for her great contributions to the knowledge of Eunicidae and Marphysa, and her friendship to PH.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.taxon	distribution	Distribution. Only known from type locality.	en	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
