identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A6A8C2F07DC45B8BBBC96ED82DA82C0B.text	A6A8C2F07DC45B8BBBC96ED82DA82C0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa banana Lavesque & Daffe & Glasby & Hourdez & Hutchings 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Marphysa banana sp. nov.</p>
            <p>Figs 2, 3, 4</p>
            <p>Material examined.</p>
            <p>
                 Holotype: MNHN-IA-2015-1608, complete.   Paratypes: AM W.53773, complete; AM W.53774, complete, some parapodia mounted for SEM; MNHN-IA-2021-725, anterior part only. All material collected from  
                <a title="Search Plazi for locations around (long 150.66/lat -2.483)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.66&amp;materialsCitation.latitude=-2.483">South Pacific Ocean</a>
                 , Papua New Guinea, New Ireland, CP4254, - 2.483°S, 150.66°E, depth 273-324 m, April 2014  . 
            </p>
            <p>Description</p>
            <p>(based on holotype, with variation in parentheses for paratypes). Preserved specimens strongly iridescent (Fig. 2A, C, D), ~ 230 (220) chaetigers, 112 mm (71-157) long, 3.3 mm (2.1-5.4) width at chaetiger 10, excluding parapodia. Body elongated and tapered gradually at posterior end (Fig. 2B).</p>
            <p> Prostomium rounded anteriorly with two dorsoventrally flattened buccal lips an anterior notch between them, notch more visible ventrally (Fig. 2A, C, D). Two palps and three antennae slender and tapering, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, longer than palps (same size), shorter (same size) than prostomium (Fig. 2C). Eyes present, one pair, brownish, very faint, present at posterior base between palps and lateral antennae. First peristomial ring ~ 3  × longer than second one dorsally (Fig. 2C). </p>
            <p> Maxillary apparatus (Fig. 3D, E) partially everted in holotype or paratypes. Formula as follows: MF = 1+1, 5+5, 6+0, 4+9, 1+1. MI ~ 2  × longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of rounded wings situated at posterolateral margins. MI forceps-like, without attachment lamellae, sub-right-angle falcal arch. Closing system ~ 4  × shorter than MI. Ligament between MI and MII golden. MII without attachment lamella, teeth triangular, distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, longer than left MIV, curved, with equal-sized triangular teeth; short attachment lamella situated in the centre of posterior edge of maxilla, thin, dark. Left MIV short (less than half the size of right MIV) with wide, rounded base, left-most teeth longer than right-most ones; attachment lamella dark, semi-circular. Right MIV long, with teeth triangular, decreasing in size posteriorly; attachment lamella wide, semi-circular, dark. MV, paired, as long as high, with a dorsal curved tooth. Mandibles light brown, concentric stripes not visible; longer than MI; cutting plates whitish, without dorsal teeth (Fig. 3E). </p>
            <p>First few parapodia located below middle line of body wall, but gradually positioned dorsally to approximately midline in subsequent segments (Fig. 2A). Notopodial cirri slender, tapering, slightly longer than ventral cirri, thinner posteriorly (Fig. 3A-C). Chaetal lobes comprising a low pre-chaetal lip and a globular post-chaetal lobe. Ventral cirri bluntly conical, with rounded tip, shorter than post-chaetal lobes anteriorly, thereafter slightly longer than post-chaetal lobes (Fig. 3A-C). Branchiae pectinate, commencing from chaetiger 20 (18-19) and continuing to near end, very short anteriorly, longer in medium chaetigers but not reaching mid-dorsal line; number of filaments increasing from 1-3 anteriorly to 4-6 in mid-body, decreasing to 3-4 in last several chaetigers (Figs 2E, 3A-C).</p>
            <p>Aciculae black with paler blunt tips, approximately three or four per parapodium in anterior chaetigers, one or two per parapodium in middle chaetigers, and one per parapodium in posterior chaetigers. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers (Fig. 3A-C).</p>
            <p>Pectinate chaetae commencing from approximately chaetiger 20 to near end, three types identified. Type 1 from anterior parapodia to mid-body: isodont-narrow-slender (INS), having ~ 20 short internal teeth, each tooth prolonged by a thin filament (Fig. 4A, B). Types 2 and 3 from posterior parapodia only (Fig. 4C, D): type 2 asymmetrical, anodont-wide-thick (AWT), having ~ 10 thick internal teeth, each tooth prolonged by a thin filament; type 3 asymmetrical, anodont-wide-slender (AWS), having ~ 20 internal teeth, each tooth prolonged by a thin filament (Fig. 4D).</p>
            <p>Subacicular chaetae with compound spinigers and subacicular hooks (Fig. 3F). Compound spinigers commencing from first chaetiger to near pygidium, with long, tapered blade. Subacicular hooks transparent, commencing from anterior chaetigers 43-52 (range for type material) to near end and inferior to bundle of spinigers, one per parapodium; much thinner than aciculae; subacicular hooks bidentate (Fig. 3F).</p>
            <p> Pygidium round, with crenulated margin, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 2-3  × length of ventral pair (Fig. 2B). </p>
            <p>Etymology.</p>
            <p>The species name refers to the decomposing banana leaves among which all the specimens were found.</p>
            <p>Type locality.</p>
            <p>South Pacific, Papua New Guinea, New Ireland.</p>
            <p>Distribution.</p>
            <p>Only known from type locality.</p>
            <p>Habitat.</p>
            <p>Between 273 and 324 m, found inside banana leaves that presumably have been entrained from river runoff via coastal waters.</p>
            <p>Remarks.</p>
            <p> With the presence of compound spinigers only and the branchiae present over many chaetigers  Marphysa banana sp. nov. belongs to the group B2, also known as the  Nereis sanguinea -group Quatrefages, 1866. Among the nine species of this group occurring in the Central Indo-Pacific Realm,  M. banana sp. nov. is similar to  M. hongkongensa ,  M. iloiloensis , and  M. mullawa by the presence of subacicular hooks starting from chaetigers 30-50 and the branchiae commencing from chaetigers 14-20. </p>
            <p> However,  M. banana sp. nov. differs from  M. hongkongensa by the presence of pectinate chaetae starting from around chaetiger 20 instead of starting from the first few chaetigers as found for  M. hongkongensa ; and by the presence of three different types of pectinate chaetae instead of four types as found in  M. hongkongensa . Moreover,  M. banana sp. nov. has eyes whereas  M. hongkongensa does not have any. The subacicular hooks of  M. banana sp. nov. are bidentate while those of  M. hongkongensa are unidentate and the maximum number of branchial filaments reaches six for  M. banana sp. nov., while it can be ten for  M. hongkongensa . Finally,  M. hongkongensa lives in the lower intertidal of the Hong Kong region, while  M. banana sp. nov. is a deep-sea species found inside banana leaves. </p>
            <p> Marphysa banana sp. nov. differs from  M. iloiloensis by the presence of four and nine teeth on the maxillary MIV, while  M. iloiloensis has three and five teeth respectively. The two species show three different types of pectinate chaetae but not the same ones, as  M. banana sp. nov. has INS, AWT and AWS with the first ones starting from chaetiger 20 while  M. iloiloensis has INS, IWS and ANT, with first ones commencing from the first few chaetigers. The subacicular hooks are also different as they are bidentate for  M. banana sp. nov. and unidentate for  M. iloiloensis . Finally,  M. iloiloensis lives in the brackish waters of the Philippines region, which is a very different habitat from the deep-sea habitat of  M. banana sp. nov. </p>
            <p> Marphysa banana sp. nov. differs from  M. mullawa by the presence of pectinate chaetae starting from around chaetiger 20 instead of commencing from the first few chaetigers for  M. mullawa , and the anterior chaetae numbering ~ 20 internal teeth instead of 10 for  M. mullawa . The two species differ by their maxillary formulae, especially for maxillary MII (7+7 for  M. mullawa , 5+5 for  M. banana sp. nov.) and MIII (8+0 for  M. mullawa , 6+0 for  M. banana sp. nov.). Another difference concerns the shape of anterior branchiae, which are palmate for  M. mullawa and pectinate for  M. banana sp. nov. Once again, the two species live in very different habitats as  M. mullawa is found in intertidal and shallow waters only, on mud or in seagrass beds. </p>
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	https://treatment.plazi.org/id/A6A8C2F07DC45B8BBBC96ED82DA82C0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lavesque, Nicolas;Daffe, Guillemine;Glasby, Christopher;Hourdez, Stephane;Hutchings, Pat	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
EF15540AE13B5319983AEBD0E579298D.text	EF15540AE13B5319983AEBD0E579298D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa papuaensis Lavesque & Daffe & Glasby & Hourdez & Hutchings 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Marphysa papuaensis sp. nov.</p>
            <p>Figs 5, 6, 7</p>
            <p>Material examined.</p>
            <p>
                  Holotype: MNHN-IA-2015-1559, complete,  
                <a title="Search Plazi for locations around (long 152.4/lat -4.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.4&amp;materialsCitation.latitude=-4.6">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4264, - 4.6°S, 152.4°E, depth 430-523 m, April 2014  .   Paratypes: MNHN-IA-2015-1415, complete,  
                <a title="Search Plazi for locations around (long 149.316/lat -6.083)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.316&amp;materialsCitation.latitude=-6.083">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4337, - 6.083°S, 149.316°E, depth 287-447 m, May 2014  ;   MNHN-IA-2015-1593, anterior part only,  
                <a title="Search Plazi for locations around (long 149.166/lat -6.133)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.166&amp;materialsCitation.latitude=-6.133">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4329, - 6.133°S, 149.166°E, depth 250-500 m, May 2014  ;   AM W.53770, complete (several parapodia mounted for SEM),  
                <a title="Search Plazi for locations around (long 152.4/lat -4.6)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.4&amp;materialsCitation.latitude=-4.6">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4264, 4.6°S, 152.4°E, depth 430-523 m, April 2014  ;   AM W.53771, anterior part only, mounted for SEM,  
                <a title="Search Plazi for locations around (long 149.166/lat -6.116)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.166&amp;materialsCitation.latitude=-6.116">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4334, - 6.116°S, 149.166°E, depth 430-620 m, May 2014  ;   AM W.53772, complete, gravid,  
                <a title="Search Plazi for locations around (long 152.416/lat -4.6166)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.416&amp;materialsCitation.latitude=-4.6166">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4266, - 4.6166°S, 152.416°E, depth 575-616 m, April 2014  . 
            </p>
            <p>Additional material.</p>
            <p>
                  MNHN-IA-2015-1610, anterior part only, gravid,  
                <a title="Search Plazi for locations around (long 151.1/lat -2.9)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.1&amp;materialsCitation.latitude=-2.9">South Pacific Ocean</a>
                 , Papua New Guinea, New Ireland, CP4260, - 2.9°S, 151.1°E, depth 350-847 m, April 2014  ;   MNHN-IA-2015-1949, anterior part only,  
                <a title="Search Plazi for locations around (long 150.8/lat -2.25)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.8&amp;materialsCitation.latitude=-2.25">South Pacific Ocean</a>
                 , Papua New Guinea, New Ireland, CP4434, - 2.25°S, 150.8°E, depth 1066-1200 m, August 2014  ;   MNHN-IA-2015-1615, anterior part only, few parapodia used for molecular analysis,  
                <a title="Search Plazi for locations around (long 150.116/lat -2.683)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.116&amp;materialsCitation.latitude=-2.683">South Pacific Ocean</a>
                 , Papua New Guinea, New Hanover, CP 4482, - 2.683°S, 150.116°E, depth 761-825 m, September 2014  . 
            </p>
            <p>Description</p>
            <p>(based on holotype, with variation in parentheses for paratypes). Specimens strongly iridescent (Fig. 5B), 88 (89) chaetigers, 45 mm (41-80) long, 3.6 mm (2.5-2.8) width at chaetiger 10, excluding parapodia. Body elongated and tapered gradually at posterior end, anteriorly not flattened (Fig. 5A).</p>
            <p> Prostomium bilobed, with buccal lips separated by a ventral notch only (Fig. 5B). Two palps and three antennae slender and tapering, palpophores not visible, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, slightly longer than palps and prostomium (same size) (Figs 5C, 7A). Eyes absent. First peristomial ring ~ 1.8  × longer than second one dorsally (Figs 5C, 7A). </p>
            <p> Maxillary apparatus (Fig. 6D, E) partially everted in holotype or paratypes. Formula as follows: MF = 1+1, 5(6)+5(6), 7+0, 4(3)+10(11), 1+1. MI ~ 2.5  × longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of rounded wings situated at posterolateral margins. MI forceps-like, without attachment lamellae, sub-right-angle falcal arch. Closing system ~ 4-5  × shorter than MI. Ligament between MI and MII rectangular, dark. MII without attachment lamella, teeth triangular, distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, longer than left MIV, curved, with equal-sized triangular teeth; short attachment lamella situated in the centre of posterior edge of maxilla, oval, dark. Left MIV short (half the size of right MIV) with wide, rounded base, two left teeth longer than right-most ones; attachment lamella dark, semi-circular. Right MIV long, with teeth triangular, decreasing in size posteriorly; attachment lamella wide, semi-circular, dark. MV, paired, longer than wide, with a long tooth pointed ventrally, and a rounded dorsal margin (Fig. 6D). Mandibles dark with golden tips, with fine concentric stripes visible dorsally and ventrally, same size as MI; cutting plates whitish, with distinct growth rings, with three dorsal teeth (Fig. 6E). </p>
            <p> Notopodial cirri very long, slender and, tapering (Fig. 6A-C), 2-3  × longer than ventral cirri in all chaetigers. Pre-chaetal lobe inconspicuous. Post-chaetal lobe digitiform in the two or three first chaetigers, triangular with tapering tip from chaetiger 4, reducing in size from chaetiger 17, almost inconspicuous from chaetiger 28 (21) (Fig. 6A-C). Ventral cirri (Fig. 6A-C) bluntly conical, with slightly expanded bases and rounded tips from chaetigers 1-22, subconical and thinner thereafter. Branchiae pectinate (Figs 5C, 6A, 7A), starting from chaetiger 7 (7) and continuing for a limited number of segments, until chaetiger 16 (14); with 8-16 long filaments. </p>
            <p>Aciculae black with paler blunt tips, 2-4 per parapodium along the body. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers. Pectinate chaetae commencing from first few chaetigers to near end, one type identified as heterodont-narrow-slender (HNS; Fig. 7B, D, E), one outer tooth very long (Fig. 7E), having nine or ten short internal teeth, each tooth prolonged by a thin filament.</p>
            <p>Subacicular chaetae with compound falcigers and subacicular hooks (Figs 6F, G, 7B, C). Compound falcigers bidentate, with short blade and large teeth, commencing from first chaetiger to near pygidium, with more than 50 chaetae within a parapodium in anterior part, with ~ 10 chaetae in mid-body and ~ 3-5 in last chaetigers (Fig. 6F). Subacicular hooks black with pale yellow tip, commencing from anterior chaetiger 20 (24) to end, most of the body with one hook per parapodia, but some posterior chaetigers with two, subacicular hooks bidentate (Figs 6G, 7B, C).</p>
            <p> Pygidium round and crenulated, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 2 (3)  × length of ventral pair (Fig. 5D). </p>
            <p>Etymology.</p>
            <p>This species name refers to the type locality and geographical distribution of this species.</p>
            <p>Type locality.</p>
            <p>Papua New Guinea, Solomon Sea, New Britain.</p>
            <p>Distribution.</p>
            <p>Papua New Guinea, Solomon Sea (New Britain) and Bismarck Sea (New Ireland).</p>
            <p>Habitat.</p>
            <p>Between 250 and 1200 m, mostly found inside sunken wood.</p>
            <p>Remarks.</p>
            <p> Within the Central Indo-Pacific Realm, a single species with only compound falcigers present and branchiae restricted in a short region (group C1) has been described:  M. bernardi Rullier, 1972 (type locality in New Caledonia). However, this species differs from  M. papuaensis sp. nov. by the presence of a prostomium that is not bilobed, of antennae that are articulated and the absence of eyes. In contrast,  M. papuaensis sp. nov. has smooth antennae, no eyes and a bilobed prostomium. The branchiae of  M. bernardi are present from chaetiger 3, instead of chaetiger 7 for  M. papuaensis sp. nov. and apparently  M. bernardi has no pectinate chaetae, while  M. papuaensis sp. nov. has pectinates commencing from first few chaetigers to near end. Finally,  M. bernardi was collected in a bay from 7-8 m depth while  M. papuaensis sp. nov. occurs only in deep sea, at 1200 m depth. </p>
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	https://treatment.plazi.org/id/EF15540AE13B5319983AEBD0E579298D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lavesque, Nicolas;Daffe, Guillemine;Glasby, Christopher;Hourdez, Stephane;Hutchings, Pat	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
95959A1EFC26545C96F39DE2447356BB.text	95959A1EFC26545C96F39DE2447356BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa Quatrefages 1866	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Marphysa Quatrefages, 1866</p>
            <p>Type species.</p>
            <p> Nereis sanguinea Montagu, 1813. </p>
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	https://treatment.plazi.org/id/95959A1EFC26545C96F39DE2447356BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lavesque, Nicolas;Daffe, Guillemine;Glasby, Christopher;Hourdez, Stephane;Hutchings, Pat	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
3E74F0EF84D85754AB0199910B65B639.text	3E74F0EF84D85754AB0199910B65B639.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa zanolae Lavesque & Daffe & Glasby & Hourdez & Hutchings 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Marphysa zanolae sp. nov.</p>
            <p>Figs 8, 9</p>
            <p>Material examined.</p>
            <p>
                  Holotype: MNHN-IA-2015-1519, entire, few parapodia used for molecular analysis,  
                <a title="Search Plazi for locations around (long 151.1/lat -2.9)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.1&amp;materialsCitation.latitude=-2.9">South Pacific Ocean</a>
                 , Papua New Guinea, New Ireland, CP4260, - 2.9°S, 151.1°E, depth 350-847 m, April 2014  .   Paratype: MNHN-IA-2015-1607, anterior part only,  
                <a title="Search Plazi for locations around (long 152.25/lat -4.616)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.25&amp;materialsCitation.latitude=-4.616">South Pacific Ocean</a>
                 , Papua New Guinea, New Britain, CP4266, - 4.616°S, 152.25°E, depth 575-616 m, April 2014  . 
            </p>
            <p>Description</p>
            <p>(based on holotype, with variation in parentheses for paratype). Preserved specimens 197 (85 ant. part only) chaetigers, 101 mm (36 mm) long, 4.1 mm (2.8 mm) wide at chaetiger 10, excluding parapodia. Body elongated, slightly tapering at posterior end.</p>
            <p>Prostomium strongly bilobed with two dorsoventrally flattened buccal lips and an anterior notch between them (Fig. 8B, C). Two palps and three antennae slender and tapering, arranged in an arc on posterior margin of prostomium. Antennae more or less smooth, of equal length, shorter than prostomium (slightly longer for PNG012), slightly longer than palps (palps very short for paratype PNG12, but probably broken) (Fig. 8C). Eyes absent. First peristomial ring approximately the same size as second one dorsally (Fig. 8C).</p>
            <p> Maxillary apparatus yellow to golden brown, partially everted in holotype and paratype. Maxillae with carriers and four paired elements and one single one, formula as follows (Fig. 9F): MF = 1+1, 4+4, 5+0, 3+6, 1+1. MI ~ 2  × longer than maxillary carrier, rectangular anteriorly, triangular posteriorly, with a pair of oval wings situated at posterolateral margins. MI forceps-like, without attachment lamellae; well-developed falcal arch. Closing system ~ 4-5  × shorter than MI. MII wide, without attachment lamella, teeth triangular, recurved, and distributed in less than half of plate length. Ligament between MII and MIII absent (or not sclerotized). MIII, single, slightly shorter than right MIV, curved forming part of distal arc; with left four teeth recurved, equal-sized and triangular, two right teeth shorter and blunt, without attachment lamella. Left MIV short (half the size of right MIV) with wide, triangular base, left 2 teeth longer than right-most one; attachment lamella dark, semi-circular. Right MIV with teeth triangular, recurved, decreasing in size posteriorly; attachment lamella large, wide, best developed centrally. MV, paired, rectangular (longer than wide), with a broad cutting edge, and no clearly defined teeth (but following tradition to score as 1+1). Mandibles (Fig. 9G) yellow to golden brown, slightly shorter than MI plus carriers; cutting plates whitish, with distinct growth rings. </p>
            <p>First two parapodia located below middle line of body wall, but gradually positioned dorsally to approximately midline in subsequent segments (Fig. 8A). Notopodial cirri with large base and slender, tapering tip from anterior to mid-body chaetigers, digitiform cirri in posterior chaetigers; same size as neuropodial cirri, but shorter than post-chaetal lobe in anterior chaetigers (Fig. 9A-C). Chaetal lobes comprising a low pre-chaetal lip and a large tongue-like post-chaetal lobe from first chaetiger to approximately chaetiger 25, almost inconspicuous thereafter. Ventral cirri bluntly conical until chaetiger 25, digitiform with bulbous base thereafter (Fig. 9A-C).</p>
            <p>Branchiae with a long single filament (Figs 8A, 9B), commencing from chaetiger 31 (32) and continuing to mid-body (i.e., chaetiger 118 for holotype).</p>
            <p>Aciculae black with paler blunt tips, ~ four per parapodium in anterior chaetigers, two or three per parapodium in middle chaetigers, and one or two per parapodium in posterior chaetigers. Supra-acicular chaetae with limbate capillaries and pectinates; capillaries present from first chaetiger to near pygidium, numbering up to 20 in anterior chaetigers. Pectinate chaetae commencing from first few chaetigers to near end, one type only (Fig. 9H, I), with two or three pectinate chaetae per parapodium in anterior body, up to seven from posterior chaetigers, isodont-wide-thick (IWT) having 11-20 long teeth (Fig. 9H, I).</p>
            <p>Subacicular chaetae compound falcigers and subacicular hooks (Fig. 9D, E). Compound falcigers bidentate, with long blades and short teeth, commencing from first chaetiger to near pygidium, with more than 30 chaetae within a parapodium in anterior part, with ~ 20 chaetae in mid-body and ~ 5-7 in last chaetigers (Fig. 9D). Subacicular hooks amber to black, with much paler tip, commencing from anterior chaetiger 28 (chaetiger 30) to near end, one per parapodium in anterior and posterior parts, few chaetigers with two hooks in middle body; slightly thinner than aciculae; subacicular hooks unidentate, with blunt tip (Fig. 9E).</p>
            <p> Pygidium round, dorsally positioned, with two pairs of tapering pygidial cirri attached at ventral edge, dorsal pair 3  × length of ventral pair (Fig. 8D). </p>
            <p>Etymology.</p>
            <p> This species is dedicated to Joana Zanol for her great contributions to the knowledge of  Eunicidae and  Marphysa , and her friendship to PH. </p>
            <p>Type locality.</p>
            <p>Solomon Sea, Papua New Guinea, New Britain and New Ireland.</p>
            <p>Distribution.</p>
            <p>Only known from type locality.</p>
            <p>Habitat.</p>
            <p>Between 350 to 616 m depth, among pumice rocks, inside sunken wood.</p>
            <p>Remarks.</p>
            <p> Within the Central Indo-Pacific Realm, only one species having only compound falcigers present and branchiae present in a long region (group C2) occurs:  M. soembaensis Augener, 1933 (type locality in Pulau Sumba, South Indonesia). However, this species differs from  M. zanolae sp. nov. by the presence of poorly developed branchiae with two or three branchial filaments instead of well-developed branchiae with a single long filament only for  M. zanolae sp. nov. These branchiae start from chaetiger 40 for  M. soembaensis and from chaetiger 31 for  M. zanolae . Moreover,  M. soembaensis has bidentate subacicular hooks while they are unidentate for  M. zanolae sp. nov. Finally,  M. zanolae sp. nov. has pectinate chaetae with very long outer teeth, which are not present in  M. soembaensis . The blade of the compound falcigers is very short for  M. soembaensis compared to those of  M. zanolae sp. nov. Finally, specimens of  M. soembaensis were sampled intertidally in a bay in Indonesia while  M. zanolae sp. nov. occurs in deep-sea environments in Papua New Guinea. </p>
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	https://treatment.plazi.org/id/3E74F0EF84D85754AB0199910B65B639	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Lavesque, Nicolas;Daffe, Guillemine;Glasby, Christopher;Hourdez, Stephane;Hutchings, Pat	Lavesque, Nicolas, Daffe, Guillemine, Glasby, Christopher, Hourdez, Stephane, Hutchings, Pat (2022): Three new deep-sea species of Marphysa (Annelida, Eunicida, Eunicidae) from Papua New Guinea (Bismarck and Solomon seas). ZooKeys 1122: 81-105, DOI: http://dx.doi.org/10.3897/zookeys.1122.89990, URL: http://dx.doi.org/10.3897/zookeys.1122.89990
