identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
314F9605EDAA5568B95948406818050D.text	314F9605EDAA5568B95948406818050D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena polycystidiata Z. W. Liu, Y. P. Ge, L. Zou & Q. Na, sp. nov.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mycena polycystidiata Z.W. Liu, Y.P. Ge, L. Zou &amp; Q. Na, sp. nov.</p>
            <p>Figs 2, 3, 4, 5</p>
            <p>Diagnosis.</p>
            <p>Pileus greyish-rose, umbo brownish-orange, hygrophanous. Stipe pubescent. Pleurocystidia polymorphic in shape. Stipitipellis a cutis, with numerous projecting hyphae.</p>
            <p>Holotype.</p>
            <p>China. Heilongjiang Province: Liangshui National Nature Reserve, Yichun City, 47°12'74"N, 128°52'86"E, 20 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0417 (collection number MY0633).</p>
            <p>Etymology.</p>
            <p>Refers to the variable shape of pleurocystidia.</p>
            <p>Description.</p>
            <p> Pileus 14-31 mm in diam., campanulate to hemispherical when young, plano-convex with age, with obtuse umbo at centre, margin slightly revolute, at times cracked at mature; umbo brownish-orange (7C3-7C5), disc purplish-grey (13C2, 14B2, 14C2), reddish-grey (12B2, 12C2) to greyish-rose (12B3), near margin reddish-grey (12D2), greyish-ruby (12D3) or purplish-grey (13D3), margin whitish; striate none or indistinct, greyish-ruby (12E3-12E4), towards the centre up to 1/3 diam.; surface dry and rugose, hygrophanous, generally tomentose. Context white, 2 mm thick, fragile. Lamellae emarginate, slightly decurrent when old, 20-28 reaching the stipe, 1-2 tiers of lamellulae, white, irregularly intervenose, edge concolorous, wavy. Stipe 34-73  × 3-7 mm, central, cylindrical, base occasionally compressed with age; apex violet brown (11E3-11E4, 11F4), greyish-ruby (12E4), lower part brownish-grey (11D2, 11E2) to greyish-brown (11D3, 11E3) or purplish-grey (13C2), fragile, hollow; apex to middle densely pubescent, sparser towards base; whitely villose at base. Odour strongly raphanoid, taste indistinct. </p>
            <p> Basidiospores (130/5/4) (6.4)6.7-7.4-8.3(8.8)  × (3.2)3.5-3.9-4.3(4.6)  μm [Q = (1.62)1.72-2.05(2.18), Q = 1.90  ± 0.11] [holotype (70/2/1) (6.7)6.9-7.6-8.5(8.7)  × (3.4)3.6-4.0-4.4(4.6)  μm , Q = (1.71)1.76-2.05(2.13), Q = 1.90  ± 0.09], elongated ellipsoid to cylindrical, colourless, smooth, thin-walled, amyloid. Basidia 21-31  × 6-8  μm , 4-spored, clavate, hyaline, sterigmata approximately 4  μm in length. Cheilocystidia thin-walled, hyaline, differs in two shapes, mainly utriform, 50-65  × 20-31  μm , some subclavate, 54-78  × 14-19  μm . Pleurocystidia abundant, thin-walled, hyaline, multi-shaped: lanceolate and mostly round to blunt apices, 37-81  × 12-20  μm , lanceolate and acute apices, 51-87  × 14-22  μm , elliptical, 30-86  × 12-31  μm , ovate and acute apices, 49-71  × 15-24  μm , ovate and mostly round to blunt apices, 49-73  × 16-22  μm . Pileipellis a cutis composed of four to five layers cylindrical cells, 51-81  × 4-5  μm , smooth and thin-walled; terminal cells cylindrical or fusiform, 50-69  × 3-22  μm , thin-walled, hyaline.  Hypodermium formed by fusiform to subglobose hyphae, 32-69  × 18-54  μm , thin-walled, hyaline. Lamellar trama subregular, dextrinoid. Stipitipellis a cutis composed of cylindrical hyphae 5-8  μm in diam., smooth, thin-walled, with numbers of projecting hyphae 2-6  μm in diam.; caulocystidia 29-74  × 6-19  μm , clavate or fusiform, thin-walled, smooth, hyaline.  Clamps present in all tissues. </p>
            <p>Habitat.</p>
            <p> Scattered on the litter layers in  Pinus koraiensis and  Larix gmelinii mixed forests. </p>
            <p>Known distribution.</p>
            <p>Heilongjiang Province, China.</p>
            <p>Additional material examined.</p>
            <p>  China. Heilongjiang Province: Liangshui National Nature Reserve,  Yichun City , 47°12'82"N, 128°52'94"E, 20 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0418 (collection number MY0634)  ;  same location, 21 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0421 (collection number MY0659) ;  same location, 21 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0422 (collection number MY0661) . </p>
            <p>Notes.</p>
            <p> Macroscopically,  Mycena luteovariegata Harder &amp;  Læssøe and  M. pura resemble  M. polycystidiata in pileus colour, but the latter possesses more typically utriform cheilocystidia and uncontracted pleuro- and cheilocystidia (Perry 2002; Robich 2003; Harder et al. 2013; Aronsen and  Læssøe 2016; Na 2019).  Mycena pearsoniana also has a rose to violaceous pileus, but differs from  M. polycystidiata in having inamyloid spores and lacking pleurocystidia (Aronsen and  Læssøe 2016; Na 2019). Compared with  M. polycystidiata ,  M. sirayuktha Aravind. &amp; Manim. has similar cheilocystidia, but has an obviously greyish-brown striate pileus, inamyloid spores and slightly glutinous pileipellis with finger-like excrescences (Aravindakshan and Manimohan 2015). </p>
            <p> The pleurocystidia of  M. polycystidiata varied in shape amongst specimens (Fig. 5). In all four specimens, most pleurocystidia were lanceolate and with round to blunt apices, but pleurocystidia with lanceolate and acute apices, elliptical and ovate and acute apices were also observed in FFAAS0417 (Holotype) and FFAAS0418, while elongated lageniform-lanceolate or round apices ovate were detected in FFAAS0421 and FFAAS0422. The multi-shaped pleurocystidia may show a morphological continuum that changes between developmental stages. Nevertheless, the multi-shaped pleurocystidia are unquestionably diagnostic for identification of this species. </p>
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	https://treatment.plazi.org/id/314F9605EDAA5568B95948406818050D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Zewei;Ge, Yupeng;Zeng, Hui;Cheng, Xianhao;Na, Qin	Liu, Zewei, Ge, Yupeng, Zeng, Hui, Cheng, Xianhao, Na, Qin (2022): Four new species of Mycena sect. Calodontes (Agaricales, Mycenaceae) from northeast China. MycoKeys 93: 23-56, DOI: http://dx.doi.org/10.3897/mycokeys.93.86580, URL: http://dx.doi.org/10.3897/mycokeys.93.86580
A5644585F3C2536CB1B0C0ADB1C6B3C4.text	A5644585F3C2536CB1B0C0ADB1C6B3C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena rufobrunnea Z. W. Liu, Y. P. Ge & Q. Na, sp. nov.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mycena rufobrunnea Z.W. Liu, Y.P. Ge &amp; Q. Na, sp. nov.</p>
            <p>Figs 6, 7, 8</p>
            <p>Diagnosis.</p>
            <p>Pileus dark brown at centre, disc gradually turning paler to reddish-brown to greyish-brown, edge white. Lamellae obviously intervenose. Stipe apex to middle greyish-magenta to dull violet, lower part darker to dark purple or dark magenta. Cheilocystidia utriform, sometimes clavate. Pleurocystidia absent. Caulocystidia clavate or fusiform. Pileipellis with fusiform terminal cells.</p>
            <p>Holotype.</p>
            <p>China. Jilin Province: Dayangcha, Erdaobaihe Town, Antu County, Yanbian Korean Autonomous Prefecture, 42°20'73"N, 127°56'06"E, 16 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0416 (collection number MY0581).</p>
            <p>Etymology.</p>
            <p>Refers to reddish-brown pileus.</p>
            <p>Description.</p>
            <p> Pileus 12-34 mm in diam., hemispherical to convex when young, then plano-convex, sometimes an unclear umbo at centre, margin slightly revolute, acute to subacute, at times cracked at mature; dark brown (8F6-8F8) at centre, disc gradually turning paler to reddish-brown (8D4-8D5, 8E6-8E8) to greyish-brown (8D3) and turning to whitish at margin; striate reddish-brown (8D4-8D5, 8E6-8E8), towards the centre up to 1/3-1/2 diam.; surface humidus when wet. Context white, 1.5 mm thick, fragile. Lamellae adnexed to emarginate, 20-23 reaching the stipe, 1-3 tiers of lamellulae, white, irregularly intervenose, edge concolorous, slightly serrulate. Stipe 19-62  × 2-6 mm, central, cylindrical, apex to middle greyish-magenta (14E4-14E5) to dull violet (16E3-16E4), lower part darker to dark purple (14F4-14F5) or dark magenta (13F3), fragile, hollow, base slightly swollen with whitish villose. Odour raphanoid, taste indistinct. </p>
            <p> Basidiospores (80/4/3) (7.1)7.6-8.4-9.2(9.6)  × (3.8)4.0-4.5-5.0  μm [Q = (1.73)1.77-1.98(2.05), Q = 1.88  ± 0.07] [holotype (40/2/1) (7.9)8.1-8.6-9.2(9.4)  × 4.2-4.6-5.0  μm , Q = (1.73)1.77-1.96(1.98), Q = 1.87  ± 0.06], elongated ellipsoid to cylindrical, colourless, smooth, thin-walled, amyloid. Basidia 24-34  × 7-10  μm , 4-spored, clavate, hyaline, sterigmata 2-3  μm in length. Cheilocystidia thin-walled, hyaline, utriform, sometimes clavate, 23-44  × 7-17  μm , abundant. Pleurocystidia absent. Pileipellis a cutis composed of four to five slightly interwoven layers of cylindrical cells, 44-70  × 4-7  μm , smooth, thin-walled; terminal cells cylindrical or fusiform, 34-65  × 4-17  μm , thin-walled, hyaline.  Hypodermium formed by fusiform, subcylindrical to subglobose hyphae, 15-50  × 12-37  μm , thin-walled, hyaline. Lamellar trama subregular, dextrinoid. Stipitipellis a cutis composed of hyphae 3-9  μm in diam., smooth, thin-walled; caulocystidia common in the apex, sparse in the middle and base, 23-76  × 6-14  μm , clavate and fusiform, thin-walled, hyaline, smooth.  Clamps present in all tissues. </p>
            <p>Habitat.</p>
            <p> Scattered on the decayed logs of  Acer ,  Larix ,  Pinus ,  Populus ,  Quercus and  Ulmus mixed forests. </p>
            <p>Known distribution.</p>
            <p>Jilin Province, China.</p>
            <p>Additional material examined.</p>
            <p>  China. Jilin Province: Dayangcha,  Erdaobaihe Town , Antu County,  Yanbian Korean Autonomous Prefecture , 42°20'72"N, 127°56'08"E, 16 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0414 (collection number MY0579); same location, 16 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0415 (collection number MY0580)  . </p>
            <p>Notes.</p>
            <p> Species of sect.  Mycena Calodontes that are macroscopically similar to  Mycena rufobrunnea have been recorded in many regions of the world. Most taxa resemble  M. rufobrunnea in pileus colour (Smith 1947; Maas Geesteranus 1992a, 1992b; Grgurinovic 2003; Robich 2003; Chew et al. 2014; Aronsen and  Læssøe 2016).  Mycena dura Maas Geest. &amp; Hauskn., recorded in Europe, also has a dark brown to greyish-brown pileus, but can be distinguished from  M. rufobrunnea in having a white stipe and having pleurocystidia (Robich 2003; Aronsen and  Læssøe 2016).  Mycena kuehneriana A.H. Sm., which is recorded from the United States and Canada, is distinguished from  M. rufobrunnea in that its pileus is pale avellaneous with rose and lilac, almost white when faded and the spores are obviously smaller (5-6  × 2-3  μm ) (Smith 1947; Maas Geesteranus 1992a, 1992b).  Mycena clarkeana Grgur. and  M. nullawarrensis Grgur., described from Australia, are similar to  M. rufobrunnea in having a reddish-brown pileus, but both species have broader spores and possess pleurocystidia (Grgurinovic 2003).  Mycena cahaya A.L.C. Chew &amp; Desjardin,  M. seminau A.L.C. Chew &amp; Desjardin and  M. sinar A.L.C. Chew &amp; Desjardin, known from Malaysia, resemble  M. rufobrunnea owing to the brown pileus, but differ in having adnate to subdecurrent lamellae, a yellowish-grey or brownish-orange stem, mucronate cheilocystidia and lack caulocystidia (Chew et al. 2014). Microscopically, utriform or clavate cheilocystidia and absence of pleurocystidia are key characteristics of  M. rufobrunnea .  Mycena diosma Krieglst. &amp;  Schwöbel has similar cheilocystidia and pleurocystidia are absent or rare, but it has a strongly hygrophanous pileus and a remarkable change in colour (Robich 2003; Aronsen and  Læssøe 2016).  Mycena pura ,  M. sirayuktha and  M. vinacea Cleland have similar cheilocystidia, but are easily distinguished from  M. rufobrunnea by the presence of pleurocystidia (Perry 2002; Grgurinovic 2003; Robich 2003; Aravindakshan and Manimohan 2015; Aronsen and  Læssøe 2016; Na 2019). </p>
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	https://treatment.plazi.org/id/A5644585F3C2536CB1B0C0ADB1C6B3C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Zewei;Ge, Yupeng;Zeng, Hui;Cheng, Xianhao;Na, Qin	Liu, Zewei, Ge, Yupeng, Zeng, Hui, Cheng, Xianhao, Na, Qin (2022): Four new species of Mycena sect. Calodontes (Agaricales, Mycenaceae) from northeast China. MycoKeys 93: 23-56, DOI: http://dx.doi.org/10.3897/mycokeys.93.86580, URL: http://dx.doi.org/10.3897/mycokeys.93.86580
53D69B3C50BD5DF48FBDBF102F7EE427.text	53D69B3C50BD5DF48FBDBF102F7EE427.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena shengshanensis Z. W. Liu, Y. P. Ge & Q. Na, sp. nov.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mycena shengshanensis Z.W. Liu, Y.P. Ge &amp; Q. Na, sp. nov.</p>
            <p>Figs 9, 10, 11, 12</p>
            <p>Diagnosis.</p>
            <p> Pileus brown to violet-brown at centre, disc light brown to dull red. Cheilocystidia clavate with slightly inflated apex, thick-walled. Pleurocystidia absent. Caulocystidia clavate with tapered apices, apex to middle thick-walled. Scattered to gregarious under  Larix gmelinii . </p>
            <p>Holotype.</p>
            <p>China. Heilongjiang Province: Shengshan National Nature Reserve, Heihe City, 49°37'45"N, 126°47'39"E, 23 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0424 (collection number MY0686).</p>
            <p>Etymology.</p>
            <p>Refers to the type locality.</p>
            <p>Description.</p>
            <p> Pileus 13-26 mm in diam., when young parabolic to convex, with obtuse umbo at centre, then plano-convex, margin wavy and revolute, at times cracked at mature; centre light brown (7D5-7D6), brown (7E4-7E8), dark brown (8F5-8F6), violet brown (11F4-11F6), disc paler to light brown (7D4-7D5), brown (7E5-7E6), greyish-brown (8D3), reddish-brown (8D4, 8E4), brownish-grey (11C2), dull red (11C3), margin whitish; striate indistinct, brownish-orange (7C3), greyish-brown (7D3), reddish-grey (12D2), greyish-ruby (12E3-12E5), towards the centre up to 1/3-1/2 diam.; surface slightly moist, smooth. Context white, 1-2 mm thick, fragile. Lamellae sinuate to subdecurrent, 19-25 reaching the stipe, 1-3 tiers of lamellulae, white, irregularly intervenose, edge concolorous, wavy and slightly serrulate. Stipe 26-42  × 2-4 mm, central, cylindrical, apex reddish-brown (8E4-8E5), greyish-ruby (12E4-12E5), greyish-brown (11F3), violet brown (11F4-11F5), dark ruby (12F4-12F5), lower part paler to brownish-grey (7C2), brownish-orange (7C3), greyish-brown (11E3), greyish-ruby (12E3), base darker to brown (7E5), reddish-brown (8E4-8E5), greyish-brown (11F3), violet brown (11F4-11F5), greyish-ruby (12E3), dark ruby (12F6-12F8), fragile, hollow, base swollen with white fibrils. Odour raphanoid, taste indistinct. </p>
            <p> Basidiospores (60/3/2) (5.9)6.1-6.9-8.1(8.7)  × 3.4-4.0-4.4(4.7)  μm [Q = 1.62-1.93 (1.97), Q = 1.75  ± 0.09] [holotype (40/2/1) 6.1-7.0-8.2(8.7)  × 3.4-4.0-4.4(4.7)  μm , Q = (1.62)1.65-1.93(1.97), Q = 1.77  ± 0.08], elongated ellipsoid, colourless, smooth, thin-walled, amyloid. Basidia 22-32  × 6-8  μm , 4-spored, clavate, hyaline, sterigmata 3-4  μm in length. Cheilocystidia moderately thick-walled (0.5-0.6  μm ), clavate with slightly inflated apex, 25-63  × 6-12  μm , hyaline. Pleurocystidia absent. Pileipellis a cutis composed of three to four layers of cylindrical cells, 24-57  × 3-5  μm , smooth and thin-walled; terminal cells cylindrical, apically narrow, 28-49  μm in length, apex 1-3  μm and base 2-5  μm in diam., thin-walled, hyaline.  Hypodermium formed by fusiform to subglobose hyphae, 19-53  × 13-30  μm , thin-walled, hyaline. Lamellar trama subregular, weakly dextrinoid to dextrinoid. Stipitipellis a cutis composed of hyphae 4-9  μm in diam., smooth, thin-walled; caulocystidia 22-61  × 5-20  μm , clavate with tapered apices, apex to middle thick-walled, smooth, hyaline.  Clamps present in all tissues. </p>
            <p>Habitat.</p>
            <p> Scattered to gregarious on the litter layer in  Larix gmelinii . </p>
            <p>Known distribution.</p>
            <p>Heilongjiang Province, China.</p>
            <p>Additional material examined.</p>
            <p>  China. Heilongjiang Province: Shengshan National Nature Reserve,  Heihe City , 23 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0425 (collection number MY0687)  . </p>
            <p>Notes.</p>
            <p> In sect.  Mycena Calodontes ,  M. diosma ,  M. pearsoniana and  M. yuezhuoi also have clavate cheilocystidia with a slightly inflated apex and lack pleurocystidia, similar to  M. shengshanensis , but  M. diosma differs in pileus characters,  M. pearsoniana has inamyloid spores and  M. yuezhuoi has a more purple pileus and subcellular lamellar trama (Robich 2003; Aronsen and  Læssøe 2016; Na 2019; Liu et al. 2021). Clavate cheilocystidia are also present in  M. luteovariegata and  M. pura , but these species differ in having pleurocystidia (Perry 2002; Robich 2003; Harder et al. 2013; Aronsen and  Læssøe 2016; Na 2019). Macroscopically, several species in sect.  Mycena Calodontes also have a brown with reddish or violet pileus or stipe (Smith 1947; Maas Geesteranus 1992a, 1992b; Grgurinovic 2003; Robich 2003; Aronsen and  Læssøe 2016).  Mycena dura ,  M. kuehneriana and  M. nullawarrensis are distinguished by basidiospore size (Smith 1947; Maas Geesteranus 1992a, 1992b; Grgurinovic 2003; Robich 2003; Aronsen and  Læssøe 2016). Two species,  M. seminau and  M. sirayuktha , reported from Southeast Asia, are similar to  M. shengshanensis owing to the brown pileus, but they differ in having gelatinised or sometimes mucronate cheilocystidia and caulocystidia have not been observed (Chew et al. 2014; Aravindakshan and Manimohan 2015). Fusiform, obclavate, ovate and clavate cheilocystidia with a subcapitate protuberance were observed occasionally, but clavate cheilocystidia with a slightly inflated apex represented the predominant morphological type in  M. shengshanensis (Fig. 12). </p>
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	https://treatment.plazi.org/id/53D69B3C50BD5DF48FBDBF102F7EE427	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Zewei;Ge, Yupeng;Zeng, Hui;Cheng, Xianhao;Na, Qin	Liu, Zewei, Ge, Yupeng, Zeng, Hui, Cheng, Xianhao, Na, Qin (2022): Four new species of Mycena sect. Calodontes (Agaricales, Mycenaceae) from northeast China. MycoKeys 93: 23-56, DOI: http://dx.doi.org/10.3897/mycokeys.93.86580, URL: http://dx.doi.org/10.3897/mycokeys.93.86580
E08A7A12E21E5CD0B4D92B1C693B32FC.text	E08A7A12E21E5CD0B4D92B1C693B32FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycena subulata Z. W. Liu, Y. P. Ge & Q. Na, sp. nov.	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Mycena subulata Z.W. Liu, Y.P. Ge &amp; Q. Na, sp. nov.</p>
            <p>Figs 13, 14, 15</p>
            <p>Diagnosis.</p>
            <p>Pileus reddish-grey to dull red, slightly hygrophanous. Cheilocystidia thick-walled, slenderly fusiform with distinctly long and narrow protuberance. Stipitipellis a cutis, with projecting hyphae, caulocystidia thick-walled.</p>
            <p>Holotype.</p>
            <p>China. Heilongjiang Province: Liangshui National Nature Reserve, Yichun City, 47°13'13"N, 128°53'21"E, 21 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0423 (collection number MY0671).</p>
            <p>Etymology.</p>
            <p>Refers to cheilocystidia with distinctly long and narrow protuberance.</p>
            <p>Description.</p>
            <p> Pileus 9-32 mm in diam., convex to campanulate when young, hemispherical to applanate with age, margin sometimes wavy, slightly deflexed; at centre dull red (8C3), brownish-grey (8D2), greyish-brown (8D3), reddish-brown (8D4, 8E4-8E5) and dark brown (8F5), disc paler to reddish-grey (8B2, 9B2), brownish-grey (9C2), dull red (9B3, 9C3), greyish-magenta (13D3), margin light brown (7D4), brown (7E4) or dull red (9C3); striate none or indistinct, reddish-brown (8E4-8E5), towards the centre up to 1/5 diam.; surface dry, unclearly rugose or none, margin slightly hygrophanous. Context white, 1 mm thick, fragile. Lamellae sinuate to subdecurrent, 31-33 reaching the stipe, 1-3 tiers of lamellulae, white, irregularly intervenose, edge concolorous, wavy and slightly serrulate. Stipe 27-75  × 2-5 mm, central, cylindrical, apex to middle brownish-orange (7C3), dull red (8C3), brownish-grey (7D2), greyish-magenta (14D3), lower part brownish-grey (8C2), greyish-brown (8D3), reddish-brown (8D4), fragile, hollow, white granular near apex, base slightly swollen with white fibrils. Odour raphanoid, taste indistinct. </p>
            <p> Basidiospores (100/5/4) 6.0-6.7-7.3(7.9)  × 3.3-3.8-4.3(4.6)  μm [Q = (1.61)1.65-1.87(1.90), Q = 1.76  ± 0.07] [holotype (40/2/1) (6.0)6.2-6.6-7.1  × 3.4-3.7-4.0(4.2)  μm , Q = 1.65-1.87(1.90), Q = 1.78  ± 0.07], elongated ellipsoid, colourless, smooth, thin-walled, amyloid. Basidia 23-34  × 5-6  μm , 4-spored, clavate, hyaline, sterigmata 2-3  μm in length. Cheilocystidia moderately thick-walled (0.5-0.6  μm ), hyaline, narrowly fusiform with long and narrow protuberance, 43-82  × 4-11  μm , protuberance 14-36  × 1-2  μm . Pleurocystidia absent. Pileipellis a cutis composed of three to four layers of cylindrical cells, 20-89  × 3-7  μm , smooth and thin-walled; terminal cells cylindrical, apically narrow, 24-61  μm in length, the apex 3-4  μm and base 4-7  μm in diam., thin-walled, hyaline.  Hypodermium formed by fusiform to subglobose hyphae, 21-41  × 17-25  μm , thin-walled, hyaline. Lamellar trama subregular, dextrinoid. Stipitipellis a cutis composed of hyphae 3-8  μm in diam., smooth, thin-walled, with projecting hyphae 3-8  μm in diam.; caulocystidia 29-47  × 6-12  μm , clavate and apices tapered, thick-walled (0.5-0.6  μm ), smooth.  Clamps present in all tissues. </p>
            <p>Habitat.</p>
            <p> Scattered on the litter layer in  Pinus koraiensis ,  Larix gmelinii and  Tilia sp. mixed forests. </p>
            <p>Known distribution.</p>
            <p>Heilongjiang Province, China.</p>
            <p>Additional material examined.</p>
            <p>  China. Heilongjiang Province: Liangshui National Nature Reserve,  Yichun City , 21 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0419 (collection number MY0654); same location, 21 August 2021, Zewei Liu, Yupeng Ge, Qin Na and Shixin Wang, FFAAS0420 (collection number MY0657)  ;   Heilongjiang Province: Taipinggou National Nature Reserve,  Hegang City , 3 September 2021, Shixin Wang, FFAAS0426 (collection number MY0795)  . </p>
            <p>Notes.</p>
            <p> Cheilocystidia with a long and narrow protuberance is the key microscopic character that distinguishes  M. subulata and is uncommon in sect.  Mycena Calodontes (Smith 1947; Maas Geesteranus 1992a, 1992b; Grgurinovic 2003; Robich 2003; Chew et al. 2014; Aravindakshan and Manimohan 2015; Aronsen and  Læssøe 2016; Na 2019; Liu et al. 2021).  Mycena lammiensis Harmaja and  M. pelianthina (Fr.)  Quél . have similar cheilocystidia, but differ from  M. subulata by their broader cheilocystidia with purplish-brown contents and having pleurocystidia (Smith 1947; Robich 2003; Aronsen and  Læssøe 2016). The cheilocystidia of  M. subcorticalis (Cooke &amp; Massee) Sacc. with a protuberance are similar to those of  M. subulata . However,  M. subcorticalis has larger and inamyloid spores, a gelatinised pileipellis and a stipitipellis with sparse excrescences (Grgurinovic 2003). More rarely, mucronate cheilocystidia and absence of pleurocystidia have been described for  M. pearsoniana and its clay pink pileus is similar to that of  M. subulata , but  M. pearsoniana differs in having a slightly glutinous pileus when wet and inamyloid spores (Aronsen and  Læssøe 2016; Na 2019). Other species that are macroscopically similar to  M. subulata , namely  M. luteovariegata ,  M. nullawarrensis and  M. pura , can be distinguished by cheilocystidia shape and presence of pleurocystidia (Perry 2002; Robich 2003; Grgurinovic 2003; Harder et al. 2013; Aronsen and  Læssøe 2016; Na 2019). </p>
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	https://treatment.plazi.org/id/E08A7A12E21E5CD0B4D92B1C693B32FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Zewei;Ge, Yupeng;Zeng, Hui;Cheng, Xianhao;Na, Qin	Liu, Zewei, Ge, Yupeng, Zeng, Hui, Cheng, Xianhao, Na, Qin (2022): Four new species of Mycena sect. Calodontes (Agaricales, Mycenaceae) from northeast China. MycoKeys 93: 23-56, DOI: http://dx.doi.org/10.3897/mycokeys.93.86580, URL: http://dx.doi.org/10.3897/mycokeys.93.86580
