taxonID	type	description	language	source
C9D5F78A54F95D30A081D3E79EDF2E9D.taxon	description	Figs 3, 4 Verrucous Leaf Litter Toad (in English) / Mi You Zhang Tu Chan (密疣掌突蟾 in Chinese)	en	Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi, Wang, Jian (2022): A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165-178, DOI: http://dx.doi.org/10.3897/herpetozoa.35.e89981, URL: http://dx.doi.org/10.3897/herpetozoa.35.e89981
C9D5F78A54F95D30A081D3E79EDF2E9D.taxon	etymology	Etymology. The specific epithet " Leptobrachella verrucosa " is an adjective meaning " verrucous ", referring to the tuberculate dorsal skin texture of the new species.	en	Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi, Wang, Jian (2022): A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165-178, DOI: http://dx.doi.org/10.3897/herpetozoa.35.e89981, URL: http://dx.doi.org/10.3897/herpetozoa.35.e89981
C9D5F78A54F95D30A081D3E79EDF2E9D.taxon	diagnosis	Diagnosis. (1) Small body size [SVL 23.2 - 25.9 mm in five adult males], (2) iris bicolored, upper half coppery orange and lower half grayish brown, (3) tympanum distinct, (4) supratympanic line black, (5) fingers without lateral fringes, (6) toes with rudimentary webbing, lateral fringes narrow, (7) longitudinal ridges under toes continuous, (8) heels not meeting when adpressed, tibial-tarsal articulation reaching anterior corner of eye, (9) dorsal surface shagreened with numerous conical tubercles, lacking spines, enlarged warts or skin ridges, (10) ventral surface smooth, (11) dorsum grayish brown, with distinct darker brown scattered markings and irregular light orange pigmentations, (12) flanks with several dark spots, (13) ventral surface creamy white with grayish white and dark brown spots.	en	Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi, Wang, Jian (2022): A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165-178, DOI: http://dx.doi.org/10.3897/herpetozoa.35.e89981, URL: http://dx.doi.org/10.3897/herpetozoa.35.e89981
C9D5F78A54F95D30A081D3E79EDF2E9D.taxon	description	Description of holotype. Adult male. Body size small, SVL 23.2 mm. Head length slightly larger than head width, HDW / HDL 0.95; snout slightly protruding, projecting slightly beyond margin of lower jaw; nostril closer to snout than eye; canthus rostralis gently rounded; loreal region slightly concave; interorbital space flat, internarial distance slightly larger than interorbital distance, IND / IOD 1.05; pineal ocellus absent; pupil vertical; snout slightly longer than eye diameter, SNT / EYE 1.03; tympanum distinct, rounded, diameter smaller than that of eye and larger than tympanum-eye distance, TMP / EYE 0.54, TEY / TMP 0.46; upper margin of tympanum in contact with supratympanic ridge; vomerine teeth absent; a single vocal sac; vocal sac openings slit-like, paired, located posterolaterally on floor of mouth, close to margins of mandible; tongue deeply notched distally; supratympanic ridge distinct, extending from posterior corner of eye to supra-axillary gland. Tips of fingers rounded, slightly swollen; relative finger lengths I = II = IV <III; nuptial pad absent; subarticular tubercles absent; inner palmar tubercle large, rounded, distinctly separated from outer palmar tubercle; outer palmar tubercle small, rounded; fingers lacking interdigital webbing and lateral fringes. Tips of toes rounded, slightly swollen; relative toe length I <II <V <III <IV; longitudinal ridges under toes continuous; inner metatarsal tubercle large, oval; outer metatarsal tubercle absent; interdigital webbing between toes rudimentary; narrow lateral fringes present on all toes. Tibia 49 % of snout-vent length; tibiotarsal articulation reaches to anterior corner of eye; heels not meeting when thighs are appressed at right angles with respect to the body. Dorsal surface shagreened with numerous conical tubercles, lacking spines, enlarged warts or skin ridges; ventral skin smooth; pectoral and femoral glands oval; both larger in diameter than tip of fingers; femoral glands larger in diameter than tip of toes; femoral gland situated on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised. Ventrolateral gland distinctly visible, forming a longitudinal uncontinuous series.	en	Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi, Wang, Jian (2022): A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165-178, DOI: http://dx.doi.org/10.3897/herpetozoa.35.e89981, URL: http://dx.doi.org/10.3897/herpetozoa.35.e89981
C9D5F78A54F95D30A081D3E79EDF2E9D.taxon	distribution	Distribution, ecology and conservation. Leptobrachella verrucosa sp. nov. is currently only known from its type locality (Fig. 1; ca. 500 - 600 m a. s. l.) which is located at the boundary of Guangxi Zhuang Autonomous Region, and Guangdong and Hunan provinces. The new species inhabits clear-water rocky streams in primary forests surrounded by broad-leaved forest in granite landforms. Males were observed calling while perched on rocks or leaves of dwarf shrubs near flowing seeps. Data of larvae and females remain unknown. The breeding season lasts from April to June according to our long-term observation. We recommend the new species be listed as LC (Least Concerned) in the IUCN categorization as the forest is well protected and the distribution of the species is potentially wider than currently known, i. e. the adjacent Hunan Province and Guangxi Zhuang Autonomous Region.	en	Lin, Shi-Shi, Li, Yuan-Hang, Lu, Yu-Hong, Su, Hong-Lin, Wu, Shi-Bin, Zhang, Qi-Qi, Mo, Mei-Juan, Xiao, Shao-Jun, Pan, Zhong, Pan, Hu-Jun, Zeng, Zhao-Chi, Wang, Jian (2022): A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165-178, DOI: http://dx.doi.org/10.3897/herpetozoa.35.e89981, URL: http://dx.doi.org/10.3897/herpetozoa.35.e89981
